Greece

Greece
Location
Region	Europe
Subregion	Southern Europe
BioRegion	Palaearctic Region
Diversity
Subfamilies	7
Genera	45
Endemic Genera	1
% Endemic Genera	2.2%
Species/Subspecies	328
Endemic Species	54
% Endemic Species	16.5%
Introduced Species	1

The following is a list of ant species found in Greece. There is a second webpage about this ant fauna that provides more information about the Ants of Greece.

The following valid species and subspecies of ants are known to occur in Greece. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Greece ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Greece Ants web page. Please advise if any corrections are warranted. You might also be interested in the Ants of Crete.

Acropyga

• Acropyga paleartica Menozzi, 1936

Aenictus

• Aenictus rhodiensis Menozzi, 1936

Anoplolepis

• Anoplolepis gracilipes (F. Smith, 1857)

Aphaenogaster

• Aphaenogaster aktaci Kiran & Tezcan, 2008
• Aphaenogaster balcanica (Emery, 1898)
• Aphaenogaster balcanicoides Boer, 2013
• Aphaenogaster cecconii Emery, 1894
• Aphaenogaster charesi Salata & Borowiec, 2016
• Aphaenogaster epirotes (Emery, 1895)
• Aphaenogaster festae Emery, 1915
• Aphaenogaster graeca Schulz, 1994
• Aphaenogaster illyrica Bračko, Lapeva-Gjonova, Salata, Borowiec & Polak, 2019
• Aphaenogaster jolantae Borowiec & Salata, 2014
• Aphaenogaster karpathica Boer, 2013
• Aphaenogaster ledouxi Tohmé, 1969
• Aphaenogaster lesbica Forel, 1913
• Aphaenogaster olympica Borowiec & Salata, 2014
• Aphaenogaster ovaticeps (Emery, 1898)
• Aphaenogaster peloponnesiaca Salata, Karaman, Kiran & Borowiec, 2021
• Aphaenogaster rugosoferruginea Forel, 1889
• Aphaenogaster sangiorgii (Emery, 1901)
• Aphaenogaster simonellii Emery, 1894
• Aphaenogaster splendida (Roger, 1859)
• Aphaenogaster sporadis Santschi, 1933
• Aphaenogaster subcostata Viehmeyer, 1922
• Aphaenogaster subterranea (Latreille, 1798)
• Aphaenogaster subterraneoides Emery, 1881

Bothriomyrmex

• Bothriomyrmex communista Santschi, 1919
• Bothriomyrmex corsicus Santschi, 1923
• Bothriomyrmex syria Forel, 1910

Camponotus

• Camponotus aegaeus Emery, 1915
• Camponotus aethiops (Latreille, 1798)
• Camponotus aktaci
• Camponotus anatolicus
• Camponotus atricolor (Nylander, 1849)
• Camponotus baldaccii Emery, 1908
• Camponotus boghossiani Forel, 1911
• Camponotus candiotes Emery, 1894
• Camponotus cecconii Emery, 1908
• Camponotus dalmaticus (Nylander, 1849)
• Camponotus dalmaticus rhodius Santschi, 1934
• Camponotus fallax (Nylander, 1856)
• Camponotus festai Emery, 1894
• Camponotus gestroi Emery, 1878
• Camponotus heidrunvogtae Seifert, 2019
• Camponotus herculeanus (Linnaeus, 1758)
• Camponotus ionius Emery, 1920
• Camponotus jaliensis Dalla Torre, 1893
• Camponotus kiesenwetteri (Roger, 1859)
• Camponotus laconicus Emery, 1920
• Camponotus lateralis (Olivier 1792)
• Camponotus libanicus André, 1881
• Camponotus ligniperda (Latreille, 1802)
• Camponotus nitidescens Forel, 1889
• Camponotus oertzeni Forel, 1889
• Camponotus piceus (Leach 1825)
• Camponotus rebeccae Forel, 1913
• Camponotus samius Forel, 1889
• Camponotus sanctus Forel, 1904
• Camponotus sannini Tohmé &]] Tohmé, 2000
• Camponotus schulzi
• Camponotus tergestinus Müller
• Camponotus vagus (Scopoli 1763)

Cardiocondyla

• Cardiocondyla bulgarica Forel, 1892
• Cardiocondyla dalmatica Soudek, 1925
• Cardiocondyla mauritanica Forel, 1890
• Cardiocondyla stambuloffii Forel, 1892

Carebara

• Carebara oertzeni Forel, 1886

Cataglyphis

• Cataglyphis cretica (Forel, 1910)
• Cataglyphis hellenica (Forel, 1886)
• Cataglyphis minos Borowiec & Salata, 2022
• Cataglyphis nodus (Brullé, 1833)
• Cataglyphis viaticoides (André, 1881)

Colobopsis

• Colobopsis truncata (Spinola, 1808)

Crematogaster

• Crematogaster auberti Emery, 1869
• Crematogaster erectepilosa Salata & Borowiec, 2015
• Crematogaster gullukdagensis Salata & Borowiec, 2015
• Crematogaster ionia Forel, 1911
• Crematogaster jehovae Forel, 1907
• Crematogaster lorteti Forel, 1910
• Crematogaster lorteti hellenica Forel, 1911
• Crematogaster ourea Salata, Ziecina & Borowiec, 2024
• Crematogaster schmidti (Mayr, 1853)
• Crematogaster scutellaris (Olivier 1792)
• Crematogaster sordidula (Nylander, 1849)

Cryptopone

• Cryptopone ochracea (Mayr, 1855)

Dolichoderus

• Dolichoderus quadripunctatus (Linnaeus, 1771)

Formica

• Formica bruni Kutter, 1967
• Formica cinerea Mayr, 1853
• Formica clara Forel, 1886
• Formica cunicularia Latreille, 1798
• Formica exsecta Nylander, 1846
• Formica fusca Linnaeus, 1758
• Formica gagates Latreille, 1798
• Formica lemani Bondroit, 1917
• Formica lugubris Zetterstedt, 1838
• Formica picea Nylander, 1846
• Formica polyctena Förster, 1850
• Formica pratensis Retzius, 1783
• Formica rufa Linnaeus, 1761
• Formica rufibarbis Fabricius, 1793
• Formica sanguinea Latreille, 1798

Formicoxenus

• Formicoxenus nitidulus (Nylander, 1846)

Hypoponera

• Hypoponera eduardi (Forel, 1894)
• Hypoponera punctatissima (Roger, 1859)

Lasius

• Lasius alienus (Förster 1850)
• Lasius balcanicus
• Lasius bicornis (Förster, 1850)
• Lasius bombycina Seifert & Galkowski, 2016
• Lasius brunneus (Latreille, 1798)
• Lasius carniolicus Mayr, 1861
• Lasius citrinus Emery, 1922
• Lasius creticus Seifert, 2020
• Lasius distinguendus (Emery, 1916)
• Lasius emarginatus (Olivier, 1792)
• Lasius flavus (Fabricius, 1782)
• Lasius fuliginosus (Latreille, 1798)
• Lasius illyricus Zimmermann, 1935
• Lasius jensi Seifert, 1982
• Lasius karpinisi Seifert, 1992
• Lasius kritikos Seifert, 2020
• Lasius lasioides (Emery, 1869)
• Lasius meridionalis Bondroit, 1920
• Lasius mixtus (Nylander, 1846)
• Lasius myops Forel, 1894
• Lasius neglectus Van Loon, Boomsma & Andrasfalvy, 1990
• Lasius niger (Linnaeus, 1758)
• Lasius nitidigaster Seifert, 1996
• Lasius paralienus Seifert, 1992
• Lasius precursor
• Lasius platythorax Seifert, 1991
• Lasius psammophilus Seifert, 1992
• Lasius reginae
• Lasius sabularum (Bondroit)
• Lasius tapinomoides Salata & Borowiec, 2018
• Lasius turcicus Santschi, 1921
• Lasius umbratus (Nylander, 1846)
• Lasius viehmeyeri Emery, 1922

Lepisiota

• Lepisiota frauenfeldi (Mayr, 1855)
• Lepisiota melas (Emery, 1915)
• Lepisiota nigra (Dalla Torre, 1893)
• Lepisiota syriaca

Leptanilla

There are 3 undescribed species.

Leptothorax

• Leptothorax acervorum (Fabricius, 1793)
• Leptothorax gredleri Mayr, 1855
• Leptothorax muscorum (Nylander, 1846)

Linepithema

• Linepithema humile (Mayr, 1868)

Liometopum

• Liometopum microcephalum (Panzer, 1798)

Manica

• Manica rubida (Latreille, 1802)

Messor

• Messor alexandri Tohmé & Tohmé 1981
• Messor antennatus fodorii Röszler, 1942
• Messor atanassovii Atanassov, 1982
• Messor caducus (Motschoulsky 1839)
• Messor capitatus (Latreille, 1798)
• Messor carpathous Menozzi, 1936
• Messor concolor Santschi, 1927
• Messor creticus Salata & Borowiec, 2019
• Messor danaes Salata, Georgiadis & Borowiec, 2023
• Messor hellenius Agosti & Collingwood, 1987
• Messor ibericus Santschi, 1931
• Messor intermedius Santschi, 1927
• Messor kardamenae Salata & Borowiec, 2023
• Messor laboriosus Santschi, 1927
• Messor mcarthuri Steiner et al., 2018
• Messor meridionalis (Andre, 1883)
• Messor oertzeni Forel, 1910
• Messor orientalis (Emery, 1898)
• Messor veneris Salata, Georgiadis & Borowiec, 2023
• Messor wasmanni Krausse, 1910

Metalasius

• Metalasius myrmidon (Mei, 1998)

Monomorium

• Monomorium creticum Emery, 1895
• Monomorium exiguum Forel, 1894
• Monomorium monomorium Bolton, 1987
• Monomorium nitidiventre Emery, 1893
• Monomorium pharaonis (Linnaeus, 1758)
• Monomorium phoenicum Santschi, 1927
• Monomorium subopacum (F. Smith, 1858)

Myrmecina

• Myrmecina graminicola (Latreille, 1802)

Myrmica

• Myrmica constricta Karavaiev, 1934
• Myrmica gallienii Bondroit, 1920
• Myrmica hellenica Finzi, 1926
• Myrmica hirsuta Elmes, 1978
• Myrmica karavajevi (Arnoldi, 1930)
• Myrmica lobicornis Nylander, 1846
• Myrmica lonae Finzi, 1926
• Myrmica pelops Seifert, 2003
• Myrmica ravasinii Finzi, 1923
• Myrmica rubra (Linnaeus, 1758)
• Myrmica ruginodis Nylander, 1846
• Myrmica rugulosa Nylander, 1849
• Myrmica sabuleti Meinert, 1861
• Myrmica scabrinodis Nylander, 1846
• Myrmica schencki Viereck, 1903
• Myrmica specioides Bondroit, 1918
• Myrmica sulcinodis Nylander, 1846
• Myrmica tulinae Elmes, Radchenko & Aktaç, 2002

Nylanderia

• Nylanderia jaegerskioeldi (Mayr, 1904)
• Nylanderia vividula (Nylander, 1846)

Oxyopomyrmex

• Oxyopomyrmex krueperi Forel, 1911
• Oxyopomyrmex laevibus Salata & Borowiec, 2015
• Oxyopomyrmex polybotesi Salata & Borowiec, 2015
• Oxyopomyrmex saulcyi Emery, 1889

Paratrechina

• Paratrechina longicornis

Pheidole

• Pheidole balcanica Seifert, 2016
• Pheidole indica Mayr, 1879
• Pheidole koshewnikovi Ruzsky, 1905
• Pheidole megacephala (Fabricius, 1793)
• Pheidole pallidula (Nylander, 1849)

Plagiolepis

• Plagiolepis karawajewi Radchenko, 1989
• Plagiolepis pallescens Forel, 1889
• Plagiolepis perperamus Salata, Borowiec & Radchenko, 2018
• Plagiolepis pygmaea (Latreille, 1798)
• Plagiolepis xene Stärcke, 1936

Polyergus

• Polyergus rufescens (Latreille, 1798)

Ponera

• Ponera coarctata (Latreille, 1802)
• Ponera testacea Emery, 1895

Prenolepis

• Prenolepis nitens (Mayr, 1853)

Proceratium

• Proceratium algiricum Forel, 1899
• Proceratium melinum (Roger, 1860)
• Proceratium numidicum

Proformica

• Proformica borowieci Lebas, Galkowski, Lenoir & Perdereau, 2023
• Proformica chelmosensis Lebas & Galkowski, 2019
• Proformica korbi (Emery)
• Proformica lebasi Borowiec & Salata, 2022
• Proformica oculatissima (Forel, 1886)
• Proformica striaticeps (Forel, 1911)

Solenopsis

• Solenopsis crivellarii Menozzi, 1936
• Solenopsis fugax Latreille, 1798
• Solenopsis geminata (Fabricius, 1804)
• Solenopsis latro Forel, 1894
• Solenopsis wolfi Emery, 1915

Stenamma

• Stenamma debile (Förster, 1850)
• Stenamma striatulum Emery, 1895

Stigmatomma

• Stigmatomma denticulatum Roger, 1859
• Stigmatomma impressifrons Emery, 1869

Strongylognathus

• Strongylognathus bulgaricus Pisarski
• Strongylognathus huberi dalmaticus Baroni Urbani, 1969
• Strongylognathus karawajewi Pisarski
• Strongylognathus silvestrii Menozzi, 1936
• Strongylognathus testaceus (Schenck, 1852)

Strumigenys

• Strumigenys argiola (Emery, 1869)
• Strumigenys baudueri (Emery, 1875)
• Strumigenys membranifera (Emery, 1869)
• Strumigenys tenuipilis (Emery, 1915)
• Strumigenys tenuissima (Brown, 1953)

Tapinoma

• Tapinoma erraticum (Latreille, 1798)
• Tapinoma festae Emery, 1925
• Tapinoma magnum Mayr, 1861
• Tapinoma simrothi Krausse, 1911
• Tapinoma subboreale

Temnothorax

• Temnothorax aeolius Forel, 1911
• Temnothorax adlerzi (Douwes, Jess en & Buschinger, 1988)
• Temnothorax affinis (Mayr, 1855)
• Temnothorax angulinodis Csösz, Heinze & Mikó, 2015
• Temnothorax angustifrons Csösz, Heinze & Mikó, 2015 (introduced)
• Temnothorax antigoni Forel, 1911
• Temnothorax ariadnae Csösz, Heinze & Mikó, 2015
• Temnothorax arkasi Salata & Borowiec, 2022
• Temnothorax brackoi Salata & Borowiec, 2019
• Temnothorax bulgaricus (Forel, 1892)
• Temnothorax clypeatus (Mayr, 1853)
• Temnothorax corticalis (Schenck, 1852)
• Temnothorax crasecundus Seifert & Csősz, 2015
• Temnothorax crassispinus (Karavaiev, 1926)
• Temnothorax crassistriatus Salata, Borowiec & Trichas, 2018
• Temnothorax curtisetosus Salata & Borowiec, 2015
• Temnothorax daidalosi Salata, Borowiec & Trichas, 2018
• Temnothorax dessyi (Menozzi, 1936)
• Temnothorax euboeae Salata & Borowiec, 2022
• Temnothorax exilis (Emery, 1869)
• Temnothorax flavicornis (Emery, 1870)
• Temnothorax gordiagini (Ruzsky, 1902)
• Temnothorax graecus (Forel, 1911)
• Temnothorax helenae Csösz, Heinze & Mikó, 2015
• Temnothorax ikarosi Salata, Borowiec & Trichas, 2018
• Temnothorax incompletus Salata, Borowiec & Trichas, 2018
• Temnothorax kemali (Santschi, 1934)
• Temnothorax kraussei (Emery, 1915)
• Temnothorax laconicus Csösz et al., 2013
• Temnothorax lichtensteini (Bondroit, 1918)
• Temnothorax lucidus Csösz, Heinze & Mikó, 2015
• Temnothorax messiniaensis Salata & Borowiec, 2019
• Temnothorax minotaurosi Salata, Borowiec & Trichas, 2018
• Temnothorax morea Csősz, Salata & Borowiec, 2018
• Temnothorax muellerianus (Finzi, 1922)
• Temnothorax mytilenes Salata, Srodon & Boroweic, 2023
• Temnothorax niger (Forel, 1894)
• Temnothorax nigriceps (Mayr, 1855)
• Temnothorax parnonensis Salata & Borowiec, 2022
• Temnothorax parvulus (Schenck, 1852)
• Temnothorax phaetoni Salata, Srodon & Boroweic, 2023
• Temnothorax proteii Salata, Borowiec & Trichas, 2018
• Temnothorax ravouxi (André, 1896)
• Temnothorax recedens (Nylander, 1856)
• Temnothorax rogeri Emery, 1869
• Temnothorax rottenbergii (Emery, 1870)
• Temnothorax saxonicus (Seifert, 1995)
• Temnothorax semiruber (André, 1881)
• Temnothorax smyrnensis (Forel, 1911)
• Temnothorax solerii (Menozzi, 1936)
• Temnothorax sordidulus (Müller, 1923)
• Temnothorax strymonensis Csősz, Salata & Borowiec, 2018
• Temnothorax subtilis Csösz, Heinze & Mikó, 2015
• Temnothorax tauricus (Ruzsky, 1902)
• Temnothorax tergestinus (Finzi, 1928)
• Temnothorax triangularis Salata & Borowiec, 2019
• Temnothorax turcicus (Santschi, 1934)
• Temnothorax unifasciatus (Latreille, 1798)
• Temnothorax variabilis Salata, Borowiec & Trichas, 2018

Tetramorium

• Tetramorium albenae Salata, van Delft & Borowiec, 2023
• Tetramorium atratulum (Schenck, 1852)
• Tetramorium biskrense Forel, 1904
• Tetramorium breviscapus Wagner, Arthofer, Seifert, Muster, Steiner & Schlick-Steiner, 2017
• Tetramorium chefketi Forel, 1911
• Tetramorium davidi Forel, 1911
• Tetramorium diomedeum Emery, 1908
• Tetramorium ferox Ruzsky, 1903
• Tetramorium hippocratis Agosti & Collingwood, 1987
• Tetramorium hungaricum Röszler, 1935
• Tetramorium indocile Santschi, 1927
• Tetramorium kephalosi Salata & Borowiec, 2017
• Tetramorium meridionale Emery, 1870
• Tetramorium moravicum Kratochvíl, 1941
• Tetramorium punctatum Santschi, 1927
• Tetramorium punicum (F. Smith, 1861)
• Tetramorium rhodium Emery, 1924
• Tetramorium sahlbergi Finzi, 1936
• Tetramorium schmidti Forel, 1904
• Tetramorium splendens Ruzsky, 1902

Trichomyrmex

• Trichomyrmex perplexus (Radchenko, 1997)

Fossils known from Greece

Epoch	Formation	Locality	Species
Miocene	Varvara Formation Fossil	Apomarma, Messara Basin, Crete	Polyrhachis annosa

References

• Borowiec, L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2):1-340.
• Borowiec, L.; Salata, S. 2012. Ants of Greece – checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus (Wroclaw) 23:461-563.
• Borowiec, L. & Salata, S. 2013. Ants of Greece – additions and corrections (Hymenoptera Formicidae). Genus (Wroclaw) 24, 335-401.
• Borowiec, L., Salata, S. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86, 193–236 (doi:10.1515/PJEN-2017-0013).
• Borowiec, L., Salata, S. 2017. New records of ants (Hymenoptera: Formicidae) from Sterea Ellas, Greece. Acta entomologica silesiana 25: 1-3.
• Borowiec, L., Salata, S. 2018. Ants from Thessaly, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 87, 217–248 (doi:10.2478/PJEN-2018-0016).
• Borowiec, L., Salata, S. 2018. New records of ants (Hymenoptera: Formicidae) from Epirus, Greece. Acta entomologica silesiana 26: 1-22 (doi:10.5281/ZENODO.1169150).
• Borowiec, L., Salata, S. 2018. Notes on ants (Hymenoptera: Formicidae) of the Euboea Island, Central Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-15 (doi:10.5281/ZENODO.1485235).
• Borowiec, L., Salata, S. 2018. Notes on ants (Hymenoptera: Formicidae) of Zakynthos Island, Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-13 (doi:10.5281/ZENODO.1481794).
• Borowiec, L., van Delft, J.P.L., van Delft, J.J.C.W., Salata, S. 2023. Five ant species (Hymenoptera: Formicidae) new to the Greek fauna with notes on ants from Greek Thrace. Annales of the Upper Silesian Museum in Bytom, Entomology 32 (online 008), 1-13 (doi:10.5281/ZENODO.10101028).
• Bračko, G., Kiran, K., Karaman, C., Salata, S., Borowiec, L. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4, e7945 (doi:10.3897/BDJ.4.E7945).
• Legakis, A. 1983. First Contribution to the Study of the Ants (Hymenoptera, Formicidae) of the Zagori Region (Epirus, Greece): an Annotated List of Species. Entomologica Hellenica 1: 3-6.
• Legakis, A. (1987). Aspects of the zoogeography of the ants of Crete (Greece). Chemistry and biology of social insects. J. Eder and H. Rembold. München. xxxv + 757 p., Verlag J. Peperny: 77.
• Legakis, A. (1990 (1989)). Aspects of the distribution of ants in an insular mediterranean ecosystem (Cyclades is., Greece). Rev. Ecol. Biol. Sol 26: 363-369.
• Legakis, A. (1984). The zoological museum of the University of Athens 2. The collection of ants from Greece. Biol. Gallo-Hell. 11: 85-87.
• Legakis, A. (1985). Contribution to the zoogeography of ants (Hymenoptera, Formicidae) in the Greek Islands. Biol. Gallo Hell. 10: 253-257.
• Legakis A. 2011. Annotated list of the ants (Hymenoptera, Formicidae) of Greece. Hellenic Zoological Archives. 7:1-55.