Myrmica schencki

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Myrmica schencki
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Myrmicinae
Tribe: Myrmicini
Genus: Myrmica
Species: M. schencki
Binomial name
Myrmica schencki
Viereck, 1903

Myrmica schencki casent0172731 profile 1.jpg

Myrmica schencki casent0172731 dorsal 1.jpg

Specimen labels

Synonyms

Host for the Mountain Alcon Blue (Phengaris rebeli) butterfly.


At a Glance • Polygynous  

Photo Gallery

  • Myrmica schencki workers on their woven nest entrance, Ratingen, Germany. Photo by Kajetan Mszk.

Identification

Radchenko and Elmes (2010) - A member of the schencki group. Its workers are characterized by the presence of a big vertical lobe at the scape base, by the well-developed petiolar peduncle and not-angled petiolar node (seen in profile) and especially, by the very narrow frons (mean FI 0.23); after Myrmica ravasinii, it has the second narrowest frons among all known Myrmica species in the World. M. schencki is fairly stable morphologically. It most resembles Myrmica koreana,Myrmica caucasicola and Myrmica obscura, but differs from them by its narrower frons. Also the males of all these forms are fairly well distinguishable (see also Radchenko, Elmes and. Alicata 2006; Elmes et al. 2009). In Europe, M. schencki quite often was confused with Myrmica lobicornis, though workers of the latter have the wider frons (FI > 0.27), shorter propodeal spines, and the short petiole with weakly developed peduncle and almost straight, quite steep anterior face that meets with the dorsal one at a sharp right or even acute angle. When males are available such confusions should not occur because M. lobicornis males have a much longer scape.

Collingwood (1979) - Brownish red with gaster and sometimes head darker. Frontal triangle striate. Antennal scape sharply angled near base, with an upright flange at the bend fitted closely into the thin divergent frontal ridge. Frons very narrow, about 1/4 head width. Mesopropodeal furrow shallow and postpetiole low, somewhat cubical in profile and spherical from above. Head Index: 88.4; Frons Index: 24.5; Frontal Laminae Index: 63.3. Length: 4.0-5.5 mm.

Keys including this Species

Distribution

South Europe to South Scandinavia.

Latitudinal Distribution Pattern

Latitudinal Range: 65.256706° to 40.897222°.

 
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Palaearctic Region: Andorra, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Canary Islands, China, Croatia, Czechia, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kyrgyzstan, Latvia, Lithuania, Montenegro, Netherlands, Norway, Poland, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Türkiye, Ukraine, United Kingdom of Great Britain and Northern Ireland.

Distribution based on AntMaps

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Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Biology

Throughout most of its northern distribution, M. schencki is associated with hot, dry grassland habitats including calcareous grassland, glacial sand deposits and seaboard sand-dunes. In its southern distribution it is associated with hot sub-alpine grassland. Much of its distribution seems to coincide with species of Tetramorium and Tapinoma, which are dominant species in such habitats; we have often recorded and excavated M. schencki nests from the middle of Tetramorium caespitum territories: T. caespitum can defend discrete territories that contain several polycalic nest centres connected by tunnels (e.g. see Brian and Elmes 1974). The common factor appears to be the need for relatively high soil temperatures but not completely arid conditions. In biotopes where the soil is potentially very arid, e.g. the sand-dunes of NW Europe, the sub-alpine pastures of the Southern Alps and limestone pavement as in the Burren, Ireland, M. schencki only occurs if there is a relatively high rainfall, which stops complete desiccation of the soils. Myrmica specioides is the only European Myrmica species that can tolerate similarly hot habitats. In many places its main competitors are ants from other genera and we believe that this contributes to its habit of building a small woven funnel at its nest entrance in which a “guard-worker” sits with its head more or less blocking the entrance. Tarbinsky (1981) showed that in the eastern part of its range M. schencki has rather limited foraging distances which generally agrees with our observations.

A population of M. schencki living on sandy soil In Jutland, Denmark has been studied by several authors. Elmes and Abbott (1981) showed that colonies averaged about 1 queen and 300 workers, and the overall mean head widths were 1.19 mm and 1.04 mm for queens and workers respectively. The workers were generally bicoloured but in some colonies a proportion of workers were a uniform gingery orange colour (not to be confused with the pale yellow of callow workers); no queens of this colour were noted but a few queens were almost completely dusky black. Thus colour is quite variable in this species. On the same site nest densities of 0.13 nests per m2 with an estimated 47 workers per m2 (Jensen 1976), giving an estimated colony size of about 250 workers which accords well with the later estimate. This population was shown to have a distinct diurnal foraging activity and was most active when soil surface temperatures were in the range 15-30°C, relative humidity did not affect foraging activity (Nielsen 1981a), a finding confirmed at another site (Nielsen 1981b).

Mountain Alcon Blue (Phengaris rebeli)

M. schencki has become important to nature conservation study in recent decades because it is a primary host of the larvae of the parasitic butterfly Phengaris rebeli (Hir.) (Settele et al. 2005). Numerous nests have been excavated by different teams from many parts of Europe, and the' general consensus is that the population structure outlined above is typical. This work has led to the development of a spatially explicit simulation model to describe the interaction between M. schencki and M. sabuleti, its main Myrmica competitor on many P. rebeli sites. These models suggest that underlying heterogeneity of “habitat quality” colony fission and the ability to pre-empt a nest site is important to the spatial pattern of nest distribution (e.g. see Elmes et al. 1996; Clarke et al. 1997 and references therein).

An important aspect of the interaction between M. schencki and the larvae of P. rebeli is the ability of the caterpillars to mimic various ant signals. Consequently the cuticular chemicals of the ants have been identified (e.g. Elmes et al. 2002) and how they interact with caterpillar recognition (e.g. Elmes et al. 2004; Schrbnrogge et al. 2004). One of the most interesting discoveries was that P. rebeli has a species-specific parasitoid that can detect both the M. schencki host ants and P. rebeli larvae by their odour, and itself has a suite of chemicals that can provoke fighting in the host ants (e.g. Thomas and Elmes 1993; Thomas et al. 2002). Recently the stridulations of queens and workers of M. schencki have been recorded and were shown to be similar to the sounds made by P. rebeli caterpillars which respond to the recorded ant sounds (Barbero et al. 2009). In the laboratory Starcke (1949) made quite an important contribution to the morphology of larvae. The chemistry of some glandular secretions of M. schencki workers have been identified (e.g. Evershed et al. 1983; Cammaerts et al. 1982; Attygalle et al. 1983). Workers have been shown to be quite prolific egg-layers while queens regularly lay some trophic eggs (Wardlaw and Elmes 1995, 1998). Mating swarms occur from the end of July to early September, though often slightly earlier than in some sympatric Myrmica species, we have observed mixed swarms and a mating swarm was described by Pontin (1986).

Collingwood (1979) - This is an interesting species somewhat resembling a large paler Myrmica lobicornis but distinguished by the lower more cubical postpetio1e. According to the Danish myrmecologist Chr. Skott, this species differs from other European Myrmica in having no winter brood, is mainly nocturnal and derives much of its food from the glandular excretions of low herbage such as Hypochaeris and Hieracium spp. The entrance to the nest is frequently built up as a collar of vegetable detritus (Bisgaard, 1944). Colonies are single queened and isolated, situated in sandy banks and dry pasture. Alatae are found in August, mating occurring on the ground near the nest.

Flight Period

X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

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Association with Other Organisms

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Other Insects

This ant is parasitized by the butterflies Phengaris rebeli (Settele et al., 2005; Sielezniew et al., 2010a), Phengaris arion (Sielezniew et al. 2010b) and Maculinea rebeli (Barbero et al., 2009).

Nematode

  • This species is a host for the nematode Mermithidae (unspecified "Mermix") (a parasite) in Germany (Gosswald, 1930; Laciny, 2021).

Life History Traits

  • Queen number: polygynous (Rissing and Pollock, 1988; Frumhoff & Ward, 1992)

Castes

Queen

Images from AntWeb

Myrmica schencki casent0172732 head 1.jpgMyrmica schencki casent0172732 profile 1.jpgMyrmica schencki casent0172732 profile 2.jpgMyrmica schencki casent0172732 dorsal 1.jpgMyrmica schencki casent0172732 label 1.jpg
Queen (alate/dealate). Specimen code casent0172732. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Male

Images from AntWeb

Myrmica schencki casent0172721 head 1.jpgMyrmica schencki casent0172721 profile 1.jpgMyrmica schencki casent0172721 profile 2.jpgMyrmica schencki casent0172721 dorsal 1.jpgMyrmica schencki casent0172721 label 1.jpg
Male (alate). Specimen code casent0172721. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • schencki. Myrmica rubra var. schencki Viereck, 1903: 72 (w.q.m.) EUROPE (see Baroni Urbani, 1971c: 35 (note)). [First available use of Myrmica rubra subsp. scabrinodis var. schencki Emery, 1895c: 315; unavailable name.] Wheeler, G.C. & Wheeler, J. 1953a: 120 (l.). Subspecies of scabrinodis: Emery, 1908a: 178; Forel, 1915d: 29. Status as species: Bondroit, 1911: 11; Bondroit, 1912: 351; Donisthorpe, 1915c: 265; Bondroit, 1918: 103; Müller, 1923: 44; Finzi, 1926: 109; Karavaiev, 1926b: 95; Stärcke, 1927: 84; Santschi, 1931b: 351; Karavaiev, 1934: 93; Bernard, 1967: 118; Arnol'di, 1970b: 1843; Tarbinsky, 1976: 42; Kutter, 1977c: 70; Arnol'di & Dlussky, 1978: 535; Collingwood, 1979: 56; Seifert, 1988b: 35; Kupyanskaya, 1990: 109; Atanassov & Dlussky, 1992: 99. Senior synonym of kutteri: Bernard, 1967: 118; of subopaca: Arnol'di, 1970b: 1843; of betuliana: Radchenko, 1994f: 77; of schenckioides: Radchenko & Elmes, 2010: 266. See also: Seifert, 2003b: 145; Radchenko, Elmes & Alicata, 2006: 513; Radchenko & Elmes, 2010: 266.
  • kutteri. Myrmica schencki var. kutteri Finzi, 1926: 111 (w.m.) SWITZERLAND. Junior synonym of schencki: Bernard, 1967: 118.
  • subopaca. Myrmica schencki nat. subopaca Arnol'di, 1934: 172, figs. 36, 37 (w.) UKRAINE. [Also described as new by Arnol'di, in Karavaiev, 1934: 96.] [Unresolved junior primary homonym of subopaca Smith, F. 1858b: 127, above.] Junior synonym of schencki: Arnol'di, 1970b: 1843.
  • betuliana. Myrmica betuliana Ruzsky, 1946: 70 (w.) RUSSIA. Junior synonym of schencki: Radchenko, 1994f: 77.
  • schenckioides. Myrmica schenckioides Boer & Noordijk, 2005: 120 figs. 1-4 (q.) NETHERLANDS. Junior synonym of schencki: Radchenko & Elmes, 2010: 266.

Type Material

Radchenko, Elmes and Alicata (2006) - Type material of M. schencki: lectotype worker (upper specimen on the pin with 2 workers, designated by Seifert 2003), “Graten?”, “Myrmica lobicornis Nyl.”, “M. scabrinodis var. Schencki Emery (lobicornis Schencki)”, “Lectotype (top specimen) Myrmica rubra scabrinodis var. schencki Emery, 1895, designated by Seifert”, “Museo Civico di Genova” (Musee d'Histoire Naturelle Genève); paralectotypes: 1 worker (bottom specimen) on the pin with lectotype; 1 worker and 1 queen (damaged) on the same pin, “w(Queen)Ex”, “Paralectotypus var. schencki Emery, 1895”; 1 worker and 1 male (damaged); “(Male)wEx”, “Paralectotypus var. schencki Emery, 1895” (Musee d'Histoire Naturelle Genève) Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Seifert (2003) - Lectotype worker (top specimen, by present designation) and paralectotype worker (bottom specimen on the same pin), labelled by Emery himself "Graten" (?, poorly legible) and "Jl4yrmica scabrinodis var. schencki Emery", MCSN Genova.

Karyotype

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  • n = 23, 2n = 46 (Switzerland) (Hauschteck-Jungen & Jungen, 1983).

Etymology

Radchenko and Elmes (2010) - named for the notable 19th century German myrmecologist, Prof. Carl Friedrich Schenck (1803-1878) of Weilberg, who first recognised this form.

References

References based on Global Ant Biodiversity Informatics

  • Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
  • Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 47–54. doi: 10.3897/zookeys.106.139
  • Alvarado M. 2000. Habitat correlates of ant assemblages in different forests of the South Pannonian Plain. Tiscia 32: 35-42.
  • Alvarado M., and L. Galle. 2000. Ant assemblages associated with lowland forests in the southern part of the great Hungarian plain. Acta Zoologica Academiae Scientarum Hungaricae 46(2): 79-102.
  • AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
  • Antonov I. A. 2013. Ant Assemblages (Hymenoptera: Formicidae) of Cities of the Temperate Zone of Eurasia. Russian Journal of Ecology 44(6): 523–526.
  • Antonova V., and L. Penev. 2008. Classification of assemblages of ants in the green areas in Sofia City. Acta Zoologica Bulgarica 60(2): 103-110.
  • ArtDatabanken Bugs (via GBIG)
  • Assing V. 1989. Die Ameisenfauna (Hym.: Formicidae) nordwestdeutscher Calluna-Heiden. Drosera 89: 49-62.
  • Babik H., C. Czechowski, T. Wlodarczyk, and M. Sterzynska. 2009. How does a strip of clearing affect the forest community of ants (Hymenoptera: Formicidae)? Fragmenta Faunistica 52(2): 125-141?
  • Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
  • Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
  • Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
  • Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
  • Behr D., S. Lippke, and K. Colln. 1996. Zur kenntnis der ameisen von Koln (Hymenoptera, Formicidae). Decheniana-Beihefte (Bonn) 35: 215-232.
  • Behr D., and K. Colln. 1993. Zur ameisenfauna (Hymenoptera, Formicidae) von Gonnersdorf (Kr. Daun). Dendrocopos 20: 148-160.
  • Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
  • Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:59–71
  • Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
  • Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
  • Bernard F. 1973. Etudes ecologiques sur les fourmis de Breuil-Cervinia (Val d"Aoste). Revue Ecol Biol Sol, 102: 237-269.
  • Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
  • Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
  • Boer P., W. Dekoninck, A. J. Van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten (Amsterdam) 63: 54-58.
  • Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
  • Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
  • Boer P., and J. Noordijk. 2004. De ruige gaststeekmier Myrmica hirsuta nieuw voor Nederland (Hymenoptera: Formicidae). Ned. Faun. Meded. 20: 25-32.
  • Boer P., and J. Noordijk. 2005. Myrmica schenckioides nov. sp., a new socially parasitic ant species (Hymenoptera: Formicidae). Entomol. Ber. (Amst.) 65(4): 120-123.
  • Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
  • Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
  • Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
  • Boven J. van 1949. Notes sur la faune des Hautes-Fagnes en Belgique. Bulletin et Annales de la Société Entomologique de Belgique 85: 135-143.
  • Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
  • Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
  • Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
  • Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
  • Braschler, B. and B. Baur. 2005. Experimental Small-Scale Grassland Fragmentation Alters Competitive Interactions among Ant Species. Oecologia 143(2):291-300
  • Carniel A. 1998. Ricerche sulla mirmecofauna delle Prealpi Orobiche (Lombardia) (Insecta, Hymenoptera, Formicidae). Atti. Mus. Civ. Stor. Nat. Morbegno 9: 29-39.
  • Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
  • Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
  • Chen Z. L., S. Y. Zhou, and J. H. Huang. 2016. Seven species new to science and one newly recorded species of the ant genus Myrmica Latreille, 1804 from China, with proposal of a new synonym (Hymenoptera: Formicidae). ZooKeys 551: 85–128.
  • Cherix D., and S. Higashi. 1979. Distribution verticale des fourmis dans le Jura vaudois et recensement prelimaire des bourdons (Hymenoptera, Formicidae et Apidae). Bull. Soc. Vaud. Sc. Nat. 356(74): 315-324.
  • Colindre L. 2015. Les fourmis en Picardie: bilan 2014 (Hymenoptera/ Formicidae). Entomologiste Picard 26, 15 pages.
  • Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
  • Collingwood C. A. 1951. The distribution of ants in north-west Scotland. Scottish Naturalist 63: 45-49
  • Collingwood C. A. 1951. The distribution of ants in north-west Scotland. Scottish Naturalist 63: 45-49.
  • Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
  • Collingwood C. A. 1961. Ants in the Scottish Highlands. Scotish Naturalist 70: 12-21.
  • Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
  • Collingwood C.A. 1959. Scandinavian Ants. Entomol. Rec. 71: 78-83
  • Collingwood C.A. 1961. Ants in Finland. Entomol. Rec. 73: 190-195
  • Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
  • Collingwood, C. A. 1958. The ants of the genus Myrmica in Britain. Proc. R. Entomol. Soc. Lond. Ser. A 33: 65-75
  • Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
  • Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
  • Collingwood, C. A. 1974. A revised list of Norwegian ants (Hymenoptera: Formicidae). Norsk Entomologisk Tidsskrift 21: 31-35.
  • Collingwood, C. A. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
  • Collingwood, C.A. 1958. A survey of Irish Formicidae. Proceedings of the Royal Irish Academy 59B:213-219
  • Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
  • Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
  • Cuesta-Segura, D., F. García, and X. Espadaler. "The westernmost locations of Lasius jensi Seifert, 1982 (Hymenoptera: Formicidae): first records in the Iberian Peninsula." Myrmecological News 16 (2012): 35-38.
  • Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
  • Czechowski W., B. Marko, A. Radchenko, and P. Slipinski. 2013. Long-term partitioning of space between two territorial species of ants (Hymenoptera: Formicidae) and their effect on subordinate species. Eur. J. Entomol. 110(2): 327–337.
  • Czekes Z., Radchenko, A. G., Csősz, S. Szász-Len, A., Tăuşan, I., Benedek, K., and Markó, B. 2013. The genus Myrmica Latreille, 1804 (Hymenoptera: Formicidae) in Romania: distribution of species and key for their identification. Entomologica Romanica 17: 29-50.
  • Dekoninck W., F. Hendrickx, M. Dethier, and J. P. Maelfait. 2010. Forest Succession Endangers the Special Ant Fauna of Abandoned Quarries along the River Meuse (Wallonia, Belgium). Restoration Ecology 18(5): 681–690.
  • Dekoninck W., H. De Koninck, J. Y. Baugnee, and J. P. Maelfait. 2007. Ant biodiversity conservation in Belgian calcareous grasslands: active management is vital. Belg. J. Zool. 137 (2): 137-146.
  • Dekoninck W., K. Desender, and P. Grootaert. 2008. Establishment of ant communities in forests growing on former agricultural fields: Colonisation and 25 years of management are not enough (Hymenoptera: Formicidae). Eur. J. Entomol. 105: 681–689.
  • Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
  • Della Santa E. 2000. L'identification des espèces du genre Myrmica Latreille (Formicidae) de Suisse; essai de présentation synoptique. Bull. Romand Entomol. 18: 169-187.
  • Dewes E. 2005. Ameisenerfassung im „Waldschutzgebiet Steinbachtal/Netzbachtal“. Abh. Delattinia 31: 89-118.
  • Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
  • Dvorak, L., P. BOGUSCH, I. MALENOVSKÝ, P. BEZDÌÈKA, K. BEZDÌÈKOVÁ, K. HOLÝ, P. LIŠKA, J. MACEK, L. ROLLER, M. RÍHA et al. "Hymenoptera of Hády Hill, near the city of Brno (Czech Republic), collected during the Third Czech-Slovak Hymenoptera meeting." Acta Musei Moraviae, Scientiae biologicae (Brno) 93 (2008): 53-92.
  • Elmes G. W., A. G. Radchenko, E. Balleto, and F. Barbero. 2008. Myrmica obscura Finzi - a good ant species (Hymenoptera: Formicidae) endemic to Italy. Fragmenta Faunistica (Warsaw) Volume: 51 Issue: 2 Page(s): 91-100.
  • Elmes, G.W., T. Akino, J.A. Thomas, R.T. Clarke and J.J. Knapp. 2002. Interspecific differences in cuticular hydrocarbon profiles of Myrmica ants are sufficiently consistent to explain host specificity by Maculinea (large blue) butterflies. Oecologia 130:525-535
  • Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
  • Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
  • Entomological Society of Latvia. 2003. http://leb.daba.lv/Formicidae.htm (Accessed on December 1st 2013).
  • Espadaler X., X. Roig, K. Gómez, and F. García. 2011. Formigues de les Planes de Son i mata de València (Hymenoptera, Formicidae) Treballs de la Institució Catalana d'Història Natural 16: 609-627.
  • Espadaler, X., and C. Ascaso. "Adición a las hormigas del Montseny (Barcelona)." Orsis 5 (1990): 141-147.
  • Espadaler, X. "Contribución al conocimiento de los formícidos (Hymenoptera, Formicidae) del Pirineo catalán." Tesis Universida (1979): 285 pp.
  • Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants – assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
  • Finzi B. 1926. Le forme europee del genere Myrmica Latr. Primo contributo. Bollettino della Società Adriatica di Scienze Naturali in Trieste. 29: 71-119.
  • Formidabel Database
  • Fowles, A.P. 1996. A provisional checklist of the invertebrates recorded from Wales. 2. Aculeate wasps, bees and ants (Hymenoptera: Aculeata). Countryside Council for Wales
  • Freitag A. 2013. Biodiversite 2010 en ville de Neuchatel: les fourmis (Hymenopteres, Formicidae). Bulletin de la societe Neuchateloise des sciences naturelles 133: 183-200.
  • GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
  • Gadeau de Kerville H. 1922. Materiaux pour la Faune des Hymenopteres de la Normandie. Bull. Soc. Amis Sc. Nat. Rouen 1916-1921, 1922: 217-225.
  • Galle L., and G. Szonyi. 1988. A check list of ants (Hymenoptera: Formicoidea) of a sandy grassland in Kiskunsag National Park (Hungary). Acta Biol. Szeged 34: 167-168.
  • Gallé L. 1991. Structure and succession of ant assemblages in a north European sand dune area. Holarctic Ecology 14: 31-37.
  • Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae). — In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
  • Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
  • Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
  • Gaspare Charles. 1965. Étude myrmécologique d'une région naturelle de Belgique: la Famenne Survey des Fourmis de la Région (Hymenoptera, Formicidae). Institut agronomique de l'Etat a' Gembloux. 32(4): 427-434.
  • Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
  • Glaser F., A. Freitag, and H. Martz. 2012. Ants (Hymenoptera: Formicidae) in the Münstertal (Val Müstair) – a hot spot of regional species richness between Italy and Switzerland. Gredleriana 12: 273 - 284.
  • González Martín J. D., and X. Espadaler. 2011. Formicidos del País Vasco (Hymenoptera: Formicidae): Nuevas aportaciones. Heteropterus Revista de Entomologia 11(1): 109-122.
  • Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
  • Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
  • Groc S., J. H. C. Delabie, R. Cereghino, J. Orivel, F. Jaladeau, J. Grangier, C. S. F. Mariano, and A. Dejean. 2007. Ant species diversity in the ‘Grands Causses’ (Aveyron, France): In search of sampling methods adapted to temperate climates. C. R. Biologies 330: 913–922.
  • Grzes I. M. 2009. Ant species richness and evenness increase along a metal pollution gradient in the Boles?aw zinc smelter area. Pedobiologia 53: 65-73.
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Hauschteck-Jungen E., and H. Jungen. 1983. Ant chromosomes. II. Karyotypes of western palearctic species. Insectes Soc. 30: 149-164.
  • Holgersen H. 1940. Myrmekologiske notiser I. Nor. Entomol. Tidsskr. 5: 183-187.
  • Holgersen H. 1943. Ant studies in Rogaland (south-western Norway). Avhandlingar utgitt av det Norske Videnskaps-Akademi i Oslo. I. Matematisk-Naturvidenskapelig Klasse 1943(7): 1-75.
  • Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-203.
  • Hågvar S. 2005. Altitudinal zonation of ants (Formicidae) in a steep fjord landscape in Sogndal, Western Norway. Norw. J. Entomol. 52: 3-12.
  • Jansen G., R. Savolainen, K. Vespalainen. 2010. Phylogeny, divergence-time estimation, biogeography and social parasite–host relationships of the Holarctic ant genusMyrmica(Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution 56: 294-304.
  • Kanizsai O., R. Gallé, and L. Gallé. 2009. Perception of spatial patchiness by ant assemblages (Hymenoptera: Formicidae). Tiscia 37: 3-7.
  • Karaman C., K. Kiran, and V. Aksoy. 2014. New records of the genus Strumigenys Smith, 1860 (Hymenoptera, Formicidae) from Black Sea region of Turkey. Trakya University Journal of Natural Sciences, 15(2): 59-63.
  • Karaman G. S., and M. G. Karaman 2005. Contribution to the knowledge of the ants (Hymenoptera: Formicidae) of Crna Gora (Montenegrao). Natura Montenegrina 5: 91-108.
  • Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
  • Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
  • Karavaiev V. 1912. Ameisen aus dem paläarktischen Faunengebiete. Rus. Entomol. Obozr. 12: 581-596.
  • Karavaiev V. 1926. Beiträge zur Ameisenfauna des Kaukasus, nebst einigen Bemerkungen über andere palaearktische Formen. Konowia 5: 93-109.
  • Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
  • Kvamme T. 1982. Atlas of the Formicidae of Norway (Hymenoptera: Aculeata). Insecta Norvegiae 2: 1-56.
  • Kvamme T. 1999. Notes on Norwegian ants (Hymenoptera, Formicidae). Norwegian Journal of Entomology 46: 19-23.
  • Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur – Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
  • Laeger T., and R. Schultz. 2005. Ameisen (Hymenoptera: Formicidae) als Beifänge in Bodenfallen – wie genau spiegeln sie reale Abundanzverhältnisse wider? Myrmecologische Nachrichten 7: 17-24.
  • Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
  • Le Moli F., and A. Zaccone. 1995. Ricerche sulla mirmecofauna del Cansiglio (Prealpi Carniche). Soc. Ven. Sc. Nat. 20: 33-52.
  • Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24 – 33
  • Lillig M., and E. Dewes. 2015. The former Siegfried Line as habitats for ants (Hymenoptera: Formicidae). Abh. Delattinia 37: 117 - 136
  • Lorite P., J. A. Carrillo, M. F. García, and T. Palomeque. 2000. Chromosome numbers in Spanish Formicidae. III. Subfamily Myrmicinae (Hymenoptera). Sociobiology 36: 555-570.
  • Maavara V. 1953. Ants of Estonian SSR. ABIKS loodusevaatlejale 10: 1-44.
  • Malozemova L. A. 1972. Ants of steppe forests, their distribution by habitats, and perspectives of their utilization for protection of forests (north Kazakhstan). [In Russian.]. Zoologicheskii Zhurnal 51: 57-68.
  • Marikovsky P. I. 1979. Ants of the Semireche Desert. [In Russian.]. Alma Ata: Nauka, 263 pp.
  • Marko B. 2008. Ants (Hymenoptera: Formicidae) of the “Sand Dunes of Foieni” Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
  • Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
  • Monteserín Real, S. "Invertebrados de la Reserva Natural Integral de Muniellos, Asturias: Formicidae." Consejería de Medio Ambiente, Ordenación del Territorio e Infraestructuras del Principado de Asturias KRK edicio (2003): 269.
  • Moscaliuc L. 2008. Notes on the ant fauna (Hymenoptera: Formicidae) of Rarau mountain. Analele Facult??ii de Biologie, Univ A.I. Cuza Ia?i 54: 53-55.
  • Nagy C., A. Tartally, F. Villisics, O. Merkl, E. Szita, A. Podlussany, D. Redei, S. Csosz, G. Pozgai, A. Orosz, G. Szovenyi, and V. Marko. 2009. Effects of the invasive garden ant, Lasius neglectus VAN LOON, BOOMSMA & ANDRÁS-FALVY, 1990 (Hymenoptera: Formicidae), on arthropod assemblages: pattern analyses in the type supercolony. Myrmecological News 12: 171-181.
  • Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
  • O'Rourke F. J. 1948. The distribution and general ecology of the Irish Formicidae. Proceedings of the Royal Irish Academy. Section B: Biological, Geological, and Chemical Science 52: 383-410.
  • Paraschivescu D. 1972. Fauna mirmecologica din zonele saline ale Romaniei. Studii si Cercetari de Biologie. Seria Zoologie 24: 489-495.
  • Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
  • Petal J. M. 1967. Contribution a la connaissance des fourmis (Formicidae, Hymenoptera) de la region de Lublin. VII. Associations des fourmis des milieux de tourbieres, de forets et de dunes aux environs de Libiszow (dist. De Parczew). Annales Universitatis Mariae Curie-Sklodowska Lublin-Polonia 22(9): 117-130.
  • Petal J., and A. Kusinska. 1994. Fractional composition of organic matter in the soil of anthills and of the environement of meadows. Pedobiologia 38: 493-501.
  • Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(1–2): 57-64.
  • Petrov I. Z., B. Petrov, D. Milicic, T. Karan-Znidarsic. 2007. Contribution to the Myrmecofauna (Hymenoptera: Formicidae) of East and South Serbia. Acta Zoologica Bulgarica 59(3): 295-299.
  • Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
  • Pisarski B. 1953. The ants from the vicinity of Kazimierz. Fragmenta Faunistica Mus Zool Polonici 18(6): 465-500.
  • Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
  • Pusvaskyte O. 1979. Myrmecofauna of the Lituanian SSR. Acta Entomologica Lituanica 4: 99-105.
  • Radchenko A. G., and G. W. Elmes. 2010. Myrmica ants (Hymenoptera: Formicidae) of the Old World. Fauna Mundi 3. Warsaw: Natura Optima Dux Foundation, 790 pp.
  • Radchenko A.G., G. W. Elmes, and A. Alicata. 2006. Taxonomic revision of the schencki-group of the ant genus Myrmica from the Palaearctic region. Annales Zoologici (Warszawa) 56: 499-538.
  • Reznikova Z. I. 2003. Distribution patterns of ants in different natural zones and landscapes in Kazakhstan and West Siberia along a meridian trend. Euroasian Entomological Journal 2(4): 235-342.
  • Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
  • Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
  • Saaristo M. I. 1990. New provincial records for twelve ant species (Hymenoptera, Formicidae) from Finland. Entomologica Fennica 1: 191-192.
  • Saaristo M. I. 1995. Distribution maps of the outdoor myrmicid ants (Hymenoptera, Formicidae) of Finland, with notes on their taxonomy and ecology. Entomol. Fennica 6: 153-162.
  • Sadil J. V. 1952. A revision of the Czechoslovak forms of the genus Myrmica Latr. (Hym.). Sb. Entomol. Oddel. Nár. Mus. Praze 27: 233-278.
  • Salata S. 2014. Ants (Hymenoptera: Formicidae) of the National Park of the Stołowe Mts. Przyroda Sudetow 17: 161-172.
  • Salata S., and L. Borowiec. 2014. New localities of some rare ant species (Hymenoptera: Formicidae). WIAD. ENTOMOL., 33 (1): 77-79.
  • Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
  • Schultz, R., A. G. Radchenko, and B. Seifert. "A critical checklist of the ants of Kyrgyzstan (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 201-207.
  • Seifert B. 1988. A taxonomic revision of the Myrmica species of Europe, Asia Minor, and Caucasia (Hymenoptera, Formicidae). Abhandlungen und Berichte des Naturkundemuseums Görlitz 62(3): 1-75. 
  • Seifert B. 2003. The Palaearctic members of the Myrmica schencki group with description of a new species (Hymenoptera: Formicidae). Beiträge zur Entomologie 53: 141-159.
  • Seifert, B. "The Paleartic members of the Myrmica schenki group with description of a new species." Beitr. Entomol. 53 (1) (2003): 141-159.
  • Sellier Y., C. Galkowski, C. Lebas, and P. Wegnez. 2016. Découverte de Temnothorax pardoi (Tinaut, 1987) dans la réserve naturelle nationale du Pinail (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 106-113.
  • Sielezniew M., D. Patricelli, I. Dziekanska, F. Barbero, S. Bonelli, L. P. Casacci, M. Witek, and E. Balletto. 2010. The frst record of Myrmica lonae (Hymenoptera: Formicidae) as a host of the socially parasitic large blue butterfly Phengaris (Maculinea) arion (Lepidoptera: Lycaenidae). Sociobiology 56(2): 465-.
  • Slipinski P., M. Zmihorski, and W. Czechowski. 2012. Species diversity and nestedness of ant assemblages in an urban environment. Eur. J. Entomol. 109: 197–206.
  • Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
  • Stankiewicz A. M., M. Sielezniew, and G. Svitra. 2005. Myrmica schencki (Hymenoptera: Formicidae) rears Maculinea rebeli (Lepidoptera: Lycaenidae) in Lithuania: new evidence for geographical variation of host-ant specificity of an endangered butterfly. Myrmecologische Nachrichten 7: 51-54.
  • Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
  • Stitz, H. "Ameisen aus dem westlichen Mittelmeergebiet und von den Kanarischen Inseln." Mitteilungen aus den Zoologischen Museum in Berlin 8 (1917): 333-353.
  • Stumper R. 1953. Etudes myrmecologiques. XI. Fourmis luxembourgeoises. Bulletin Soc. Nat. luxemb. 57: 122-135.
  • Stärcke A. 1942. Ants, collected by Dr. C. F. Engelhard at Stockmarknes (Island Hadsekoy, Vesterale Archipelago circa 68°50' Lat. N. Norway) and between 64° and 66° Lat. N. on the western coast of Norway, July 1932. Entomologische Berichten (Amsterdam) 11: 21-23.
  • Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
  • Tartally, A., D.R. Nash, S. Lengyel and Z. Varga. 2008. Patterns of host ant use by sympatric populations of Maculinea alcon and M. rebeli in the Carpathian Basin. Insectes Sociaux 55:370-381
  • Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
  • Vagalinski B., and A. Lapeva-Gjonova. 2012. The ants (Hymenoptera, Formicidae) of Plana Mountain (Bulgaria). Historia naturalis bulgarica 20: 87-101.
  • Vepsalainen K., H. Ikonene, and M. J. Koivula. 2008. The structure of ant assembalges in an urban area of Helsinki, southern Finland. Ann. Zool. Fennici 45: 109-127.
  • Wegnez P. 2014. Premières captures de Lasius distinguendus Emery, 1916 et de Temnothorax albipennis (Curtis, 1854) au Grand-Duché de Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société royale belge d’Entomologie 150 (2014) : 168-171.
  • Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
  • Wegnez P. 2018. Premières decouvertes de Myrmica bibikoffi Kutter, 1963 et de Ponera testacea Emery, 1895, au Luxembourg (Hymenoptera: Formicidae)Bulletin de la Société royale belge d’Entomologie 154: 263–272.
  • Wegnez P., and A. Ronk. 2017. Découverte de Camponotus herculeanus (Linnaeus, 1758) et signalement de quelques autres espèces rares de fourmis au Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société des naturalistes luxembourgeois 119 : 153–159.
  • Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
  • Wheeler W. M. 1905. Ants from the summit of Mount Washington. Psyche (Cambridge) 12: 111-114.
  • Wieziková A. 2008. Epigaeic activity of ants (Hymenoptera: Formicidae) at hemerobic gradient: arable land – green forage – permanent grassland. Acta Facultatis Ecologiae 18: 69-76.
  • Wlodarczyk T. 2010. Ant species composition in relation to forest associations in Szczecin Landscape Park. Polish Journal of Entomology 79: 91-99.
  • Wu wei, Li Xiao Mei, Guo Hong. 2004. A primary study on the fauna of Formicidae in Urumqi and its vicinities. Arid Zone Research 21(2): 179-182
  • Zryanin V. A., and T. A. Zryanina. 2007. New data on the ant fauna Hymenoptera, Formicidae in the middle Volga River Basin. Uspekhi Sovremennoi Biologii 127(2): 226-240.
  • de Biseau, J.-C., Y. Quinet, L. Deffernez and J.M. Pasteels. 1997. Explosive food recruitment as a competitive strategy in the ant Myrmica sabuleti (Hymenoptera, Formicidae). Insectes Sociaux 44:59-73