Liometopum microcephalum

AntWiki: The Ants --- Online
Liometopum microcephalum
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Dolichoderinae
Genus: Liometopum
Species: L. microcephalum
Binomial name
Liometopum microcephalum
(Panzer, 1798)

Liometopum microcephalum casent0249747 p 1 high.jpg

Liometopum microcephalum casent0249747 d 1 high.jpg

Specimen Labels

Synonyms

Liometopum microcephalum is a rare arboricolous ant, which forms large colonies of high ecological importance and ranks at the top position in the hierarchy of ant assemblages. Many aspects of the species' biology remain unknown due to its scattered occurrence and difficult nest accessibility. It is uncommon in Greece, being recorded from mainland Greece, Aegean Islands and Ionian Islands. In Achaia, a large column of thousands of workers was observed on the ground and in trees inside high-growing oak forest (Borowiec & Salata, 2021).

Identification

Borowiec and Salata (2022) - Within Greece: Liometopum microcephalum is a very distinct species. Its bicolored body and mesosoma regularly convex in profile are unique characters in the subfamily Dolichoderinae. In field, members of this species look like large, bicolored workers of Lasius emarginatus or Lasius illyricus but both members of the subfamily Formicinae well differ in conical propodeum and deep mesonotal groove.

Distribution

This is moderately common species, recorded from the Aegean Islands, Epirus, the ionian Islands, Macedonia, Peloponnese, Sterea Ellas, Thessaly and Thrace (Borowiec et al., 2022).

Latitudinal Distribution Pattern

Latitudinal Range: 49.933334° to 33.22143333°.

   
North
Temperate 		North
Subtropical 		Tropical South
Subtropical 		South
Temperate

Distribution based on Regional Taxon Lists

Palaearctic Region: Albania, Austria (type locality), Balearic Islands, Bulgaria, Croatia, Czechia, Georgia, Greece, Hungary, Iran, Israel, Italy, Montenegro, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Switzerland, Türkiye.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
pChart

Habitat

Borowiec and Salata (2022) - In Greece, L. microcephalum inhabits mainly oak forests, olive plantations and stream valleys with plane forests. Few samples were collected in pastures with large oaks or oak shrubs, small gorge with oak shrubs, area near a small lake in a moist, shaded valley of a small creek, gardens in tourist resorts, mountain pastures with limestones rocks, only one sample was from coniferous forest. Most records are from low altitude, the highest locality was placed at altitude of 1078 m.

Biology

Borowiec and Salata (2022) - Often forms large agglomerations of thousands of workers foraging on litter, tree trunks and shrubs, but does not belong to a group of true army ants.

Flight Period

X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

Explore-icon.png Explore: Show all Flight Month data or Search these data. See also a list of all data tables or learn how data is managed.

Association with Other Organisms

Explore-icon.png Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.

Pekar (2020) studied the ant-like spider Micaria sociabilis in the Czech Republic. This spider occurred exclusively on tree trunks where L. microcephalum workers occurred. The abundance of M. sociabilis increased positively with the abundance of L. microcephalum. Experiments using an olfactometer and Y-maze with volatile and contact cues obtained from the two most abundant ant species in the area, L. microcephalum and Lasius fuliginosus, were performed to find whether Micaria preferred any cue. Micaria sociabilis did not respond to volatile cues obtained from the gaster of the two ant species. In contrast, it avoided contact cues from L. fuliginosus and was attracted to contact cues from L. microcephalum ants and its gaster extract in hexane. The results thus show that M. sociabilis associates exclusively with L. microcephalum and is attracted to contact cues from this ant while avoiding cues from the competing ant. This study reveals that Batesian mimics may use kairomones to associate with visual models.

Castes

Worker

Images from AntWeb

Liometopum microcephalum casent0906112 h 1 high.jpgLiometopum microcephalum casent0906112 p 1 high.jpgLiometopum microcephalum casent0906112 d 1 high.jpgLiometopum microcephalum casent0906112 l 1 high.jpg
Worker. Specimen code casent0906112. Photographer Estella Ortega, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Liometopum microcephalum casent0906113 h 1 high.jpgLiometopum microcephalum casent0906113 p 1 high.jpgLiometopum microcephalum casent0906113 d 1 high.jpgLiometopum microcephalum casent0906113 l 1 high.jpg
Worker (major/soldier). Specimen code casent0906113. Photographer Estella Ortega, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • microcephalum. Formica microcephala Panzer, 1798: no. 1652 (m.) AUSTRIA. Mayr, 1861: 39 (w.q.). Combination in Liometopum: Mayr, 1861: 39. Senior synonym of austriaca: Mayr, 1861: 39. See also: Baroni Urbani, 1971c: 158; Atanassov & Dlussky, 1992: 189; Shattuck, 1994: 129; Del Toro, et al. 2009: 325.
  • austriaca. Formica austriaca Mayr, 1853a: 144 (w.) AUSTRIA. Junior synonym of microcephalum: Mayr, 1861: 39.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Worker

Borowiec and Salata (2022) - Polymorphic, with strongly marked size-variation: the smallest worker HLmin: 0.825; HWmin: 0,810; SLmin: 0.683; EL: 0.174; MLmin: 1.04-1.03; MWmin: 0.346; the largest worker HLmax: 1.508; HWmax: 1,540; SLmax: 1.175; EL: 0.302; MLmax: 1.94-1.03; MWmax: 0.946. Color. Head bicolor, clypeus, anterior third of gena and antennal sockets yellowish to yellowish red, clypeus often with brown spot in the middle, rest of head brown to dark brown, mesosoma yellowish to yellowish red, pronotum sometimes with two small obscure spots of diffused borders, occasionally pronotum mostly brown or whole mesosoma partly obscure, petiolar scale and gaster dark brown to black, antennae from yellow to completely brown, often scapus darker than funicle, coxa and femora usually brown, tibiae pale brown and tarsi yellowish, often femora and tibiae dark brown and tarsi yellowish brown. Head. Stout, as long as wide or slightly wider than long, on sides softly rounded, occipital corners rounded, occipital margin distinctly concave. Clypeus with diffused microreticulation but shiny, trapezoidal, lateral lobes slightly pronounced anterad, anterior margin straight or in major workers with shallow median excavation, sides concave, in the middle with deep excavation for antennal scrobes, posterior margin rounded; Clypeus with moderately long and moderately dense appressed hairs not covering clypeal background, anterior margin with 2-4 long setae in the middle and a row of short setae laterally, oblique row of three long setae in front of antennal scrobes and 2-4 short setae laterally, anterior long setae with length up to 0.142. Head with subtle microreticulation, shiny, with moderately long and moderately dense appressed pubescence not covering head surface, with a row of long erected setae on sides of frons, and several long erected setae in centre of frons, interocular and ocellar area and occipital margin, gena with only few long erected setae, sides of head with very short semierect hairs, ventral side of head with numerous erected setae. Scape short, 0.76- 0.84 times as long as width of head, stout, distinctly widened from base to apex, its surface microreticulate but shiny, with short and sparse appressed pubescence, upper margin with moderately dense, short erected setae, ventral surface with very short erected setae. Funicular segments 1-3 elongate, 4-10 only slightly longer than wide, thin, first segment 2.5 times as long as wide, second segment 0.75 times as long as first and 1.3 times as third segment. Eyes big, placed in frontal surface of head, almost circular, 0.2 length of head. Mandibles stout, distinctly sculptured tending to form a striation. Mesosoma. Elongate, 2.1-2.3 times as long as wide, dorsally and laterally with diffused microreticulation, shiny, with moderately long to long and moderately dense and dense appressed pubescence, partly covering body surface, especially on pronotum. In lateral view mesosoma forms regular arch, without mesonotal groove only with mesonotal suture, propodeum with softly declivous posterior slope. Pronotal dorsum with very long erected setae, the longest with length up to 0.286, metanotum anteriorly with short erected setae, posteriorly with a group of long setae but shorter than in pronotum, propodeum only with short erected setae anteriorly. Waist and gaster. Petiolar scale narrow, narrowed from base to narrowly rounded apex, on anterior surface with dense pubescence on posterior surface smooth and shiny. Gaster longer than mesosoma, tergites distinctly microreticulated, covered with long and dense appressed pubescence mostly. First tergite with few erected setae on anterior slope, and several setae close to posterior margin, second tergite with several erected setae anteriorly and posteriorly and third tergite with several setae on the whole surface, sternites with median keel. Legs. Moderately long and stout, first segment of tarsi not or slightly longer than subsequent segments combined, surface covered with moderately long and dense appressed hairs, erected setae absent, inner margin of tibiae lacking row of thorns.

References

References based on Global Ant Biodiversity Informatics

  • Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
  • Aktaç, N.. "Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon." Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41 (1977): 115-135.
  • Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 47–54. doi: 10.3897/zookeys.106.139
  • Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
  • Baroni Urbani, C.. "Catalogo delle specie di Formicidae d'Italia (Studi sulla mirmecofauna d'Italia X)." Memorie della Società Entomologica Italiana Volume 50 (1971): 5-287.
  • Baroni Urbani, C.. "Formiche dell'Italia appenninica (Studi sulla mirmecofauna d'Italia, III)." Memorie del Museo Civico di Storia Naturale di Verona 12 (1964): 149-172.
  • Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
  • Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
  • Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
  • Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
  • Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
  • Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
  • Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
  • Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
  • Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
  • Bracko, G.. "New species for the ant fauna of Slovenia (Hymenoptera:Formicidae)." Natura Sloneniae 5 (1) (2003): 17-25.
  • Collingwood, C. A. 1993. A Comparitive Study of the Ant Fauna of Five Greek Islands. Biologia Gallo-hellenica. 20,1:191-197
  • Collingwood, C. A.. "A comparative study of the ant fauna of five Greek islands." Biologia Gallo-Hellenica 20 (1993): 191-197.
  • Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
  • Csősz S. and Markó, B. 2005. European ant species (Hymenoptera: Formicidae) in the ant collection of the Natural History Museum of Sibiu (Hermannstadt/Nagyszeben), Romania II. Subfamily Formicinae. Annales Historico-Naturales Musei Nationalis Hungarici 97: 225-240.
  • Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
  • Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
  • De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
  • Del Toro I., J. A. Pacheco, W. P. MacKay. 2009. Revision of the ant genus Liometopum (Hymenoptera: Formicidae). Sociobiology 53: 299-369.
  • Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
  • Emery C. 1869. Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli con descrizioni di specie nuove o meno conosciute. Ann. Accad. Aspir. Nat. Secunda Era 2: 1-26.
  • Emery C. 1891. Zur Biologie der Ameisen. Biologisches Centralblatt 11: 165-180.
  • Emery C. 1913. Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae. Genera Insectorum 137: 1-50.
  • Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
  • Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants – assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
  • Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
  • Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
  • Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
  • Forel, A.. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
  • Forel, A.. "Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen." Annales de la Société Entomologique de Belgique 30 (1886): clix-clxviii.
  • Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
  • Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae). — In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
  • Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
  • Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
  • Karaman M. G. 2004. Checklist of known species of ants (Hymenoptera, Formicidae) in the fauna of Montenegro. Natura Montenegrina 3: 83-92.
  • Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
  • Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
  • Karaman M. G., and G. S. Karaman. 2003. Contribution to the knowledge of the ants (Hymenoptera, Formicidae) from Serbia. The Montenegrin Academy of Sciences and Arts. Glasnik of the Section of Natural Sciences 15: 39-58. 
  • Karaman M. G., and G. S. Karaman. 2007. Contribution to the Knowledge of the ants (Hymenoptera, Formicidae) from Special nature reserve Zasavica , Serbia. Zbornik “Naucno-strucni skup Zasavica 2007, Sremska Mitrovica, 67-75.
  • Karaman, M. G.. "Zoogeography, diversity and altitudinal distribution of ants (Hymenoptera: Formicidae) in the Mediterranean and the oro-Mediterranean parts of Montenegro." NORTH-WESTERN JOURNAL OF ZOOLOGY 7, no. 1 Article No.: 101105 (2010): 1-9.
  • Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
  • Korlevic, A.. "Prilozi fauni hrvatskih opnokrilaca." Glasn. Hrv. Narav. Dr. 5 (1890): 189-250.
  • Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
  • Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
  • Le Moli F., and M. R. Rose. 1991. Atti della Societa Italiana di Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milan. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 132:25-40.
  • Legakis A. 1983. First contribution to the study of the ants (Hymenoptera, Formicidae) of the Zagori Region (Epirus, Greece): an annotated list of species Entomologia Hellenica 1: 3-6.
  • Legakis A. 1983. The Zoological Museum of the University of Athens 2. The collection of ants from Greece. Biologia Gallo-Hellenica 11(1): 85-87.
  • Legakis Collection Database
  • Marko B. 2008. Ants (Hymenoptera: Formicidae) of the “Sand Dunes of Foieni” Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
  • Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
  • Markó B., and S. Csősz. 2002. Die europäischen Ameisenarten (Hymenoptera: Formicidae) des Hermannstädter (Sibiu, Rumänien) Naturkundemuseums I.: Unterfamilien Ponerinae, Myrmicinae und Dolichoderinae. Annales Historico-Naturales Musei Nationalis Hungarici 94: 109-121.
  • Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
  • Müller, G.. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
  • Omid P., and H. G. Kami. 2007. New and additional records for the formicid fauna (Insecta: Hymenoptera) of Iran. Zoology in the Middle East 40(1): 85-90.
  • Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
  • Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
  • Petrakova L., 2016. Populations of Liometopum microcephalum (Panzer, 1798) at different spatial scales. PhD Thesis Masaryk University. 156 pages
  • Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(1–2): 57-64.
  • Petrov I. Z. 2010. Contribution to the myrmecofauna (Hymenoptera: Formicidae) of Krusevac and its surroundings. Acta entomologica serbica, 2010, 15(1): 145-148.
  • Petrov I. Z. 2012. Preliminary data on ants (Formicidae, Hymenoptera) on Mountain Avala (Belgrade, Serbia). Bulletin of the Natural History Museum 5: 95-99.
  • Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
  • Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
  • Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
  • Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
  • Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
  • Santschi F. 1926. Travaux scientifiques de l'Armée d'Orient (1916-1918). Fourmis. Bulletin du Muséum National d'Histoire Naturelle 32: 286-293.
  • Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
  • Shattuck S. O. 1994. Taxonomic catalog of the ant subfamilies Aneuretinae and Dolichoderinae (Hymenoptera: Formicidae). University of California Publications in Entomology 112: i-xix, 1-241.
  • Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
  • Tohme G. 1996. Formicidae. Etude de la diversité biologique n° 4 . Ministère de l’Agriculture à Beyrouth (Eds.). P85-87.
  • Tohme G., and H. Tohme. 2014. Nouvelles liste des especes de fourmis du Liban (Hymenoptera, Formicoidea). Lebanese Science Journal 15(1): 133-141.
  • Vogrin, V.. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
  • Vonshak M., and A. Ionescu-Hirsch. 2009. A checklist of the ants of Israel (Hymenoptera: Formicidae). Israel Journal of Entomology 39: 33-55.
  • Zimmermann, S.. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
Retrieved from "https://antwiki.org/wiki/index.php?title=Liometopum_microcephalum&oldid=721101"