Anoplolepis gracilipes

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Anoplolepis gracilipes
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Plagiolepidini
Genus: Anoplolepis
Species: A. gracilipes
Binomial name
Anoplolepis gracilipes
(Smith, F., 1857)

Anoplolepis gracilipes casent0125111 profile 1.jpg

Anoplolepis gracilipes casent0125111 dorsal 1.jpg

Specimen Label

Synonyms


Common Name
Yellow crazy ant
Language: English
Longlegged Ant
Language: English
Ashinaga-ki-ari
Language: Japanese

The Yellow Crazy Ant has the dubious distinction of being among the 100 worst invasive species in the world (IUCN/SSC Invasive Species Specialist Group). Its distribution has been expanded by human commerce, so that it is now widespread in tropical and subtropical regions of the globe. It is a well-known pest species, because it protects aphids and coccids which injure tropical crops. In this role it has been rated a secondary agricultural pest (Reimer et. al., 1990, etc.).

At a Glance • Supercolonies  • Highly invasive  

Identification

Keys including this Species

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 28.08333333° to -8.5°.

   
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Afrotropical Region: United Arab Emirates.
Australasian Region: Australia, New Caledonia.
Indo-Australian Region: Borneo, Cook Islands, Fiji, Guam, Hawaii, Indonesia, Krakatau Islands, Malaysia, Marshall Islands, Micronesia (Federated States of), New Guinea, Niue, Northern Mariana Islands, Palau, Philippines, Samoa, Singapore (type locality), Solomon Islands, Tokelau, Tonga, Wallis and Futuna Islands.
Malagasy Region: Mauritius, Réunion, Seychelles.
Neotropical Region: Chile, Mexico.
Oriental Region: Bangladesh, Cambodia, India (type locality), Laos, Nicobar Island, Sri Lanka, Thailand, Vietnam.
Palaearctic Region: China, Greece, Japan, Türkiye.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
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Biology

Bertelsmeier et al. (2015) examined elements of interspecific aggression between this species and several other highly invasive ants. In laboratory assays Anoplolepis gracilipes was adept at avoiding aggressive interactions. When confronted by workers of other invasive ant species A. gracilipes either acted indifferently or moved away.

Milar et al. (2017) found in an experimental test, simulating being threatened with entrapment in sand (as might happen if falling in an ant lion pit or if subjected to a collapse of a ground nest), that this species did not exhibit rescue behaviour. This was in agreement with their hypothesis that species that do not face entrapment situations would not show such a response. Anoplolepis gracilipes natural occur in open areas with less friable soils.

Anoplolepis gracilipes, together with Dolichoderus thoracicus and Oecophylla smaragdina, is one of the most common ant species which tends honeydew-producing hemipterans in Indonesia. Fanani et al. (2020) examined the influence of these species on the introduced parasitoid Anagyrus lopezi, a species used to control the invasive cassava mealybug Phenacoccus manihoti (Hemiptera: Pseudococcidae). They found that when ants were absent the average time spent foraging by individual parasitoids was significantly longer (27.39 minutes) compared to when ants were present (2.47- 4.68 minutes). As a result, parasitoids spent less time in finding hosts and a longer time in handling hosts. This resulted in more oviposition activities and a 2-3 fold increase in parasitism and the number of wasps that emerged from their hosts.

Flight Period

X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

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Association with Other Organisms

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Other Insects

  • This species is a mutualist for the aphid Aphis gossypii (a trophobiont) (Idechiil et al., 2007; Saddiqui et al., 2019).
  • This species is a mutualist for the aphid Pentalonia nigronervosa (a trophobiont) (Idechiil et al., 2007; Saddiqui et al., 2019).
  • This ant has been observed tending larvae of Lampides boeticus (Obregon et al. 2015).

The following myrmecophilous crickets are known to be associated with this species (Hsu et al., 2020):

  • Myrmecophilus albicinctus
  • Myrmecophilus antilucanus
  • Myrmecophilus dubius
  • Myrmecophilus hebardi
  • Myrmecophilus mayaealberti
  • Myrmecophilus pallidithorax
  • Myrmecophilus quadrispina
  • Myrmophilellus pilipes

This species is a prey for the tiger beetle Cicindela duponti (a predator) in Western Ghats, India (Sinu et al., 2006).

Castes

Worker

Images from AntWeb

Anoplolepis gracilipes casent0064816 head 1.jpgAnoplolepis gracilipes casent0064816 profile 1.jpgAnoplolepis gracilipes casent0064816 dorsal 1.jpgAnoplolepis gracilipes casent0064816 label 1.jpg
Worker. Specimen code casent0064816. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Anoplolepis gracilipes casent0188536 head 1.jpgAnoplolepis gracilipes casent0188536 profile 1.jpgAnoplolepis gracilipes casent0188536 dorsal 1.jpgAnoplolepis gracilipes casent0188536 label 1.jpg
Worker. Specimen code casent0188536. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Anoplolepis gracilipes casent0172799 head 1.jpgAnoplolepis gracilipes casent0172799 profile 1.jpgAnoplolepis gracilipes casent0172799 dorsal 1.jpgAnoplolepis gracilipes casent0172799 label 1.jpg
Worker. Specimen code casent0172799. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MCZ, Cambridge, MA, USA.
Anoplolepis gracilipes casent0102951 head 1.jpgAnoplolepis gracilipes casent0102951 head 2.jpgAnoplolepis gracilipes casent0102951 profile 1.jpgAnoplolepis gracilipes casent0102951 dorsal 1.jpgAnoplolepis gracilipes casent0102951 dorsal 2.jpgAnoplolepis gracilipes casent0102951 dorsal 3.jpgAnoplolepis gracilipes casent0102951 label 1.jpg
Syntype of Anoplolepis gracilipesWorker. Specimen code casent0102951. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by OUM, Oxford, UK.
Anoplolepis gracilipes casent0102965 head 1.jpgAnoplolepis gracilipes casent0102965 profile 1.jpgAnoplolepis gracilipes casent0102965 profile 2.jpgAnoplolepis gracilipes casent0102965 dorsal 1.jpgAnoplolepis gracilipes casent0102965 label 1.jpg
Worker. Specimen code casent0102965. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMUK, London, UK.
Anoplolepis gracilipes casent0103001 head 1.jpgAnoplolepis gracilipes casent0103001 head 2.jpgAnoplolepis gracilipes casent0103001 profile 1.jpgAnoplolepis gracilipes casent0103001 dorsal 1.jpgAnoplolepis gracilipes casent0103001 dorsal 2.jpgAnoplolepis gracilipes casent0103001 label 1.jpg
Syntype of Anoplolepis gracilipesWorker. Specimen code casent0103001. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by OUM, Oxford, UK.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • gracilipes. Formica gracilipes Smith, F. 1857a: 55 (w.) SINGAPORE.
    • Type-material: syntype workers (number not stated).
    • [Note: Donisthorpe, 1932c: 444, cites 2w syntypes OXUM (confirmed by Bolton unpublished notes 1978).]
    • Type-locality: Singapore: “Sing. 30” (A.R. Wallace).
    • Type-depository: OXUM.
    • Replacement name for Formica longipes Jerdon, 1851: 122. [Junior primary homonym of Formica longipes Latreille, 1802c: 233.]
    • [Note: gracilipes junior synonym of longipes Jerdon (synonymy by Emery, 1887a: 247, etc.); hence gracilipes first available replacement name for longipes Jerdon, 1851: 122 (Bolton, 1995b: 67).]
    • Mayr, 1867a: 73 (q.).
    • Combination in Prenolepis: Mayr, 1862: 698;
    • combination in Plagiolepis: Mayr, 1867a: 73;
    • combination in Anoplolepis: Bolton, 1995b: 67.
    • Junior synonym of longipes: Emery, 1887a: 247; Dalla Torre, 1893: 173; Pergande, 1894: 163; Forel, 1894c: 415; Mayr, 1895: 149; Forel, 1899c: 124; Wheeler, W.M. 1919e: 101; Wheeler, W.M. 1922a: 933; Emery, 1925b: 17; Donisthorpe, 1932c: 444; Baltazar, 1966: 266; Kempf, 1970a: 30; Kempf, 1972a: 22; Snelling, R.R. & Hunt, 1976: 114.
    • Status as species: Smith, F. 1858b: 22; Smith, F. 1859a: 136; Smith, F. 1860a: 68; Mayr, 1862: 698; Roger, 1863b: 10; Mayr, 1863: 451; Mayr, 1865: 50; Mayr, 1867a: 73 (redescription); Smith, F. 1871a: 305; Mayr, 1872: 144; Mayr, 1876: 78; Emery, 1883: 147; Rothney, 1889: 373; Mayr, 1893: 197; Bolton, 1995b: 67; Zhou, 2001b: 174; Wetterer, 2002: 129; Blard, et al. 2003: 129; Bolton, 2003: 267; Imai, et al. 2003: 82; Wetterer & Vargo, 2003: 417; Jaitrong & Nabhitabhata, 2005: 13; Wetterer, 2005: 77; Wetterer, 2006: 415; Don, 2007: 201; Clouse, 2007b: 223; Framenau & Thomas, 2008: 61; Terayama, 2009: 206; Mohanraj, et al. 2010: 6; Collingwood, et al. 2011: 448; Pfeiffer, et al. 2011: 36; Ran & Zhou, 2011: 66; Borowiec, L. & Salata, 2012: 463; Guénard & Dunn, 2012: 27; Sarnat & Economo, 2012: 43; Sarnat, et al. 2013: 69; Borowiec, L. 2014: 8; Ramage, 2014: 156; Bharti, Guénard, et al. 2016: 23; Jaitrong, Guénard, et al. 2016: 26; Dias, R.K.S. et al. 2020: 29; Khachonpisitsak, et al. 2020: 39; Borowiec, L. & Salata, 2022: 56; Wang, W.Y., Soh, et al. 2022: 35.
    • Senior synonym of trifasciata: Mayr, 1867a: 73; Mayr, 1872: 144; Dalla Torre, 1893: 173; Bolton, 1995b: 67; Zhou, 2001b: 174.
    • Distribution [tramp species]
    • Afrotropical: Tanzania (Zanzibar).
    • Austral: Australia, New Caledonia, New Zealand.
    • Malagasy: Mauritius, Réunion, Rogrigues I., Seychelles.
    • Malesian: Brunei, Christmas I., Cocos-Keeling Is, Cook Is, Fiji Is, French Polynesia, Hawaii Is, Indonesia (Aru, Irian Jaya, Java, Sulawesi, Sumatra), Line Is, Malaysia (Peninsula, Sabah, Sarawak), Marianna Is, Marshall Is, Micronesia, Niue, Palau, Papua New Guinea, Philippines (Luzon, Mindanao, Negros, Palawan), Samoa, Singapore, Solomon Is, Tokelau Is, Tonga, Tuvalu, Vanuatu, Wallis & Futuna Is.
    • Neotropical: Chile, Mexico.
    • Oriental: Cambodia, China, India (+ Andaman Is, Nicobar Is), Japan, Myanmar, Sri Lanka, Taiwan, Thailand, Vietnam.
    • Palaearctic: Greece, Portugal (Azores), Turkey, United Arab Emirates.
    • [Note: distribution based on Wetterer, 2005: 77.]
  • longipes. Formica longipes Jerdon, 1851: 122 (w.) INDIA (no state data).
    • Type-material: syntype workers (number not stated).
    • Type-locality: India: (no further data), “found in all the forests of India” (T.C. Jerdon).
    • Type-depository: unknown (no material known to exist).
    • [Duplicated in Jerdon, 1854b: 105.]
    • [Junior primary homonym of Formica longipes Latreille, 1802c: 233 (Emery, 1921a: 25).]
    • Wheeler, G.C. & Wheeler, J. 1953c: 132 (l.); Imai, et al. 1984: 68 (k.).
    • Combination in Oecophylla: Roger, 1863b: 10;
    • combination in Plagiolepis: Emery, 1887a: 247;
    • combination in Plagiolepis (Anoplolepis): Santschi, 1914b: 123;
    • combination in Anoplolepis (Anoplolepis): Emery, 1925b: 17; Santschi, 1926a: 13.
    • Junior synonym of Oecophylla smaragdina: Smith, F. 1857a: 53; Mayr, 1872: 144; Forel, 1885b: 176.
    • Status as species: Smith, F. 1858b: 19; Mayr, 1863: 417; Roger, 1863b: 10; Smith, F. 1871a: 304; Emery, 1887a: 247; Emery, 1888a: 530; Emery, 1889b: 510; André, 1892b: 51; Dalla Torre, 1893: 173; Emery, 1893e: 194; Emery, 1893f: 250; Emery, 1893g: 267; Pergande, 1894: 163; Forel, 1894c: 415; Forel, 1895a: 49; Emery, 1895c: 334; Emery, 1895k: 476; Mayr, 1897: 432; Emery, 1897d: 573; Forel, 1899c: 124; Emery, 1900d: 697; Emery, 1901f: 121; Emery, 1901g: 567; Forel, 1901b: 24; Dahl, 1901: 12; Rothney, 1903: 99; Bingham, 1903: 320; Forel, 1903d: 407; Forel, 1905c: 24; Forel, 1907e: 18; Forel, 1908a: 4; Wheeler, W.M. 1908c: 166; Forel, 1909d: 229; Wheeler, W.M. 1909d: 336, 342; Forel, 1910d: 127; Yano, 1910: 421; Emery, 1911b: 532; Forel, 1911b: 207; Forel, 1912a: 72; Forel, 1912d: 109; Stitz, 1912: 510; Wheeler, W.M. 1912a: 46; Forel, 1913f: 197; Forel, 1913k: 100; Wheeler, W.M. 1913e: 237; Viehmeyer, 1914a: 114; Emery, 1914f: 420; Forel, 1915a: 34; Viehmeyer, 1916a: 145; Santschi, 1919a: 326; Wheeler, W.M. 1919e: 101; Mann, 1919: 366; Santschi, 1920h: 172; Mann, 1921: 474; Wheeler, W.M. 1922a: 933, 1036; Wheeler, W.M. 1923b: 4; Wheeler, W.M. 1924b: 253; Santschi, 1924c: 112; Emery, 1925b: 17; Stitz, 1925: 123; Mann, 1925b: 6; Mukerjee & Ribeiro, 1925: 208; Stärcke, 1926: 118 (in key); Wheeler, W.M. 1927b: 46; Wheeler, W.M. 1927d: 9; Wheeler, W.M. 1927h: 100; Santschi, 1928a: 53; Santschi, 1928c: 70; Cheesman & Crawley, 1928: 523; Wheeler, W.M. 1928c: 31; Wheeler, W.M. 1929g: 57; Karavaiev, 1930a: 213; Mukerjee, 1930: 157; Menozzi, 1930a: 3; Stärcke, 1930: 376; Wheeler, W.M. 1930h: 74; Wheeler, W.M. 1932c: 162; Menozzi, 1932d: 9; Wheeler, W.M. 1932d: 18; Wheeler, W.M. 1933a: 63; Wheeler, W.M. 1933f: 144; Wheeler, W.M. 1934a: 178; Donisthorpe, 1935: 634; Karavaiev, 1935a: 109; Goetsch & Menozzi, 1935: 99; Wheeler, W.M. 1935g: 37; Wheeler, W.M. 1936f: 15; Wheeler, W.M. 1937a: 24; Teranishi, 1940: 59; Yasumatsu, 1940a: 69; Donisthorpe, 1942d: 457; Donisthorpe, 1946e: 32; Donisthorpe, 1949b: 501; Donisthorpe, 1950a: 340; Chapman & Capco, 1951: 213; Wilson, 1962c: 18; Baltazar, 1966: 266; Wilson & Taylor, 1967: 84; Taylor, 1967b: 1094; Kempf, 1970a: 30; Kempf, 1972a: 22; Baker, 1976: 253; Snelling, R.R. & Hunt, 1976: 96; Taylor, 1976a: 87; Haines & Haines, 1978: 109; Onoyama, 1980: 199; Taylor & Brown, 1985: 108; Taylor, 1987a: 8; Rao & Veeresh, 1991: 261; Morisita, et al. 1991: 18; Dlussky, 1994: 55; Wu, J. & Wang, 1995: 132; Dorow, 1996a: 84; Tiwari, 1999: 82; Mathew & Tiwari, 2000: 348; Lin & Wu, 2003: 62.
    • Senior synonym of gracilipes: Emery, 1887a: 247; Dalla Torre, 1893: 173; Pergande, 1894: 163; Forel, 1894c: 415; Mayr, 1895: 149; Forel, 1899c: 124; Wheeler, W.M. 1919e: 101; Wheeler, W.M. 1922a: 933, Emery, 1925b: 17; Donisthorpe, 1932c: 444; Baltazar, 1966: 266; Kempf, 1970a: 30; Kempf, 1972a: 22.
    • Replacement name: Formica gracilipes Smith, F. 1857a: 55.
    • [Note: gracilipes Smith, F. oldest junior synonym of longipes Jerdon (synonymy by Emery, 1887a: 247, confirmed by examination of syntypic workers; hence gracilipes first available replacement name (Bolton, 1995b: 67).]
  • trifasciata. Formica trifasciata Smith, F. 1858b: 27 (q.) INDONESIA (Java).
    • Type-material: holotype (?) queen.
    • [Note: no indication of number of specimens is given.]
    • Type-locality: Indonesia: Java (no collector’s name).
    • Type-depository: unknown (“Coll. East India House”).
    • Combination in Camponotus: Roger, 1863b: 3.
    • Status as species: Mayr, 1863: 421; Roger, 1863b: 3; Smith, F. 1871a: 306.
    • Junior synonym of longipes: Pergande, 1894: 163; Forel, 1894c: 415; Forel, 1899c: 124; Wheeler, W.M. 1919e: 102; Wheeler, W.M. 1922a: 933; Emery, 1925b: 17.
    • Junior synonym of gracilipes: Mayr, 1867a: 73; Mayr, 1872: 144; Dalla Torre, 1893: 173; Bolton, 1995b: 67.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Worker

Borowiec and Salata (2022) - Moderately large: HL: 0.889-1.024 (mean 0.959); HW: 0.579-0.667 (mean 0.633); SL: 1.820-2.030 (mean 1.897); EL: 0.222-0.270 (mean 0.245); ML: 1.64-1.84; MW: 0.48-0.59. Color. Head, mesosoma, petiolar node, antennae and legs yellow, often anterior half of head lighter yellow colored than posterior half, occasionally antennal scape slightly infuscate, gaster mostly yellowish brown to brown, anterior slope of first tergite usually yellow, paler than rest of tergites. Head. Elongate, 1.5-1.6 times as long as wide, in front of eyes parallelsided, behind eyes regularly rounded, occipital margin convex. Clypeus with diffused microreticulation but shiny, slightly trapezoidal, its anterior margin convex, sides convergent posterad, posterior margin concave in the middle, mostly without appressed hairs, anterior margin with a row of 5-6 long setae and few short setae, close to base a pair of long erected setae, the longest anterior seta with length 0.206. Head with diffused microreticulation, shiny, mostly without appressed pubescence only on genaand sides of head with short and sparse hairs, interantennal and interocular area with three pairs of long erected setae, ocellar area with a pair of long erected setae and sometimes with 1-2 additional short setae, gena lacking erected setae, ventral side of head lacking erected setae. Scape very long, 3.0- 3.1 times longer than width of head, very thin, only slightly widened from base to apex, its surface microreticulate but shiny, with short and sparse appressed pubescence. Funicular segments elongate, thin, first segment 2.9-3.0 times as long as wide, second segment as long as first and shorter than third, the rest of funicular segments more than three times longer than broad. Eyes big, elongate oval, 0.26 length of head. Mandibles long, without striation. Mesosoma. Elongate, in dorsal view distinctly constricted in the middle, 3.0-3.4 times as long as wide, dorsally and laterally with diffused microreticulation, without appressed pubescence, shiny. In lateral view pronotum and mesonotum almost flat, propodeum regularly convex, mesonotal groove moderately deep. Mesosomal dorsum lacking erected setae except a single long setae behind propodeal spiracle. Waist and gaster. Petiolar scale in form of high node, on top with shallow longitudinal impression, in lateral view rounded apically. Gaster shorter than mesosoma, tergites with diffused microreticulation sometimes tending to form transverse sculpture, surface shiny, covered with short and sparse appressed pubescence, distance between hairs several times longer than length of hairs. First gastral tergite only with few very long erected setae close to posterior margin, subsequent tergites also with a row of very long setae close to posterior margin and usually with a pair of long setae centrally. Legs. Very long and slim, covered with short and sparse appressed to slightly decumbent hairs, without setae, inner margin of tibiae lacking row of thorns.

Type Material

The following notes on F. Smith type specimens have been provided by Barry Bolton (details):

Formica gracilipes

Two worker syntypes in Oxford University Museum of Natural History. Labelled “Sing. 30.” Also specimens det. as gracilipes from “Aru” and “N” (= New Guinea).

Karyotype

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  • n = 17, 2n = 34 (India; Indonesia; Malaysia; Sarawak) (Goni et al., 1982; Tjan et al., 1986; Imai et al., 1983; Imai et al., 1984; Imai et al., 1985) (as Anoplolepis longipes).

References

References based on Global Ant Biodiversity Informatics

  • Abe T., S. Yamane, and K. Onoyama. Ants collected on the Krakatau Islands 100 years after the great eruptions. Biogeography 14: 65-75.
  • Alcantara M. J., S. Modi, T. C. Ling, J. Monkai, H. Xu, S. Huang, and A. Nakamura. 2019. Differences in geographic distribution of ant species (Hymenoptera: Formicidae) between forests and rubber plantations: a case study in Xishuangbanna, China, and a global meta-analysis. Myrmecological News 29: 135-145.
  • Amarasinghe H. E. 2010. Species composition and nesting habits of ants in a hill-country home garden in Sri Lanka. Asian Myrmecology 3: 9-20.
  • André E. 1892. Voyage de M. Chaper à Bornéo. Catalogue des fourmis et description des espèces nouvelles. Mém. Soc. Zool. Fr. 5: 46-55.
  • Asfiya W., L. Lach, J. D. Majer, B. Heterick, and R. K. Didham. 2015. Intensive agroforestry practices negatively affect ant (Hymenoptera: Formicidae) diversity and composition in southeast Sulawesi, Indonesia. Asian Myrmecology 7: 87-104.
  • Asfiya W., R. Ubaidillah, and Sk. Yamane. 2008. Ants (Hymenoptera: Formicidae) of the Krakataus, and Sebesi and Sebuku islands. Treubia 36: 1-9.
  • Azuma, S. and M. Kinjo. 1987. Family Formicidae, In Checklist of the insects of Okinawa. The Biological Society of Okinawa, Nishihara. Pages 310-312.
  • Bluthgen N., D. Mezger, and K.E. Linsenmair. 2006. Ant-hemipteran trophobioses in a Bornean rainforest- diversity, specificity and monopolisation. Insectes Sociaux 53: 194-203.
  • Borowiec L., and S. Salata. 2019. Next step in the invasion: Trichomyrmex mayri (Forel, 1902) new to the Philippines (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 28(3): 1-3.
  • CSIRO Collection
  • Chen Y. Q., Q. Li, Y. L. Chen, Z. X. Lu, X. Y. Zhou. 2011. Ant diversity and bio-indicators in land management of lac insect agroecosystem in Southwestern China. Biodivers. Conserv. 20: 3017-3038.
  • Chen Y., C. W. Luo, H. W. Li, Y. J. Liu, H. F. Zheng, and F. C. Yang. 2013. Investigation of ant species and distribution on Wuliang Mountain. Journal of Henan Agricultural Sciences 42(5): 118-122.
  • Chhotani O. B., and P. K. Maiti. 1977. Contribution to the knowledge of Formicidae of the Andaman Islands. Zoological Survey of India 3(1): 17-20.
  • D'Cunha P., and V. Mala Grover Nair. 2013. Diversity and Distribution of Ant Fauna in Hejamadi Kodi Sandspit, Udupi District, Karnataka, India. Halteres 4: 33-47.
  • Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
  • Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
  • Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
  • Dias R. K. S. 2013. Diversity and importance of soil-dweeling ants. Proceedings of the National Symposium on Soil Biodiversity, chapt 4, pp 19-22.
  • Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
  • Dias R. K. S., and K. R. K. A. Kosgamage. Systematics and community composition of foraging workers ants (Family: Formicidae) collected from three habitats in a dry zone region of Sri Lanka. Proceedings of the Annual Research Symposium 2008. Faculty of Graduate Studies, University of Kelaniya.
  • Dias R. K. S., and K. R. K. Anuradha Kosgamage. 2012. Occurrence and species diversity of ground-dwelling worker ants (Family: Formicidae) in selected lands in the dry zone of Sri Lanka. J. Sci. Univ. Kelaniya 7: 55-72.
  • Dias R. K. S., and R. P. K. C. Rajapaksa. 2016. Geographic records of subfamilies, genera and species of ants (Hymenoptera: Formicidae) in the four climatic zones of Sri Lanka: a review. J. Sci. Univ. Kelaniya 11(2): 23-45.
  • Dias R. K. S.; Perera K. A. M. 2011. Worker ant community observed by repeated sampling and information on endemic Aneuretus simoni Emery in the Gilimale Forest Reserve in Sri Lanka. Asian Myrmecology 4: 69-78.
  • Dias, R.K.S. 2006. Overview of ant research in Sri Lanka: 2000-2004. ANeT Newsletter 8:7-10
  • Donisthorpe H. 1942. Ants from the Colombo Museum Expedition to Southern India, September-October 1938. Annals and Magazine of Natural History (11)9: 449-461.
  • Eguchi K., T. V. Bui, S. Yamane, H. Okido, and K. Ogata. 2004. Ant faunas of Ba Vi and Tam Dao, North Vietnam (Insecta: Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 27: 77-98.
  • Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia. [part]. Ann. Mus. Civ. Stor. Nat. 24(4): 209-258.
  • Emery C. 1888. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza . Formiche raccolte dal sig. Elio Modigliani in Sumatra e nell'isola Nias. Annali del Museo Civico di Storia Naturale 25: 528-534.
  • Emery C. 1893. Formicides de l'Archipel Malais. Revue Suisse de Zoologie 1: 187-229.
  • Emery C. 1893. Voyage de M. E. Simon à l'île de Ceylan (janvier-février 1892). Formicides. Annales de la Société Entomologique de France 62: 239-258.
  • Emery C. 1895. Beiträge zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8: 257-360.
  • Emery C. 1899. Intorno alle larve di alcune formiche. Mem. R. Accad. Sci. Ist. Bologna (5) 8: 3-10.
  • Emery C. 1901. Ameisen gesammelt in Ceylon von Dr. W. Horn 1899. Deutsche Entomologische Zeitschrift 1901: 113-122.
  • Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 14:565-580.
  • Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
  • Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 4, no. 24 (1887): 209-258.
  • Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
  • Emery, C. "Voyage de MM. Bedot et Pictet dans l'Archipel Malais. Formicides de l'Archipel Malais." Revue Suisse de Zoologie 1 (1893): 187-229.
  • Fayle T.M., Bakker, L., Cheah, C., Ching, T.M., Davey, A., Dem, F., Earl, A., Huaimei, Y., Hyland, S., Johansson, B., Ligtermoet, E., Lim, R., Lin, L.K., Luangyotha, P., Martins, B.H., Palmeirim, A.F., Paninhuan, S., Rojas, S.K., Sam, L., Sam, P.T.T., Susanto, D., Wahyudi, A., Walsh, J., Weigl, S., Craze, P.G., Jehle, R., Metcalfe, D. & Trevelyan, R. 2011. A positive relationship between ant biodiversity (Hymenoptera: Formicidae) and rate of scavenger-mediated nutrient redistribution along a disturbance gradient in a south-east Asian rain forest. Myrmecological News 14: 5-12.
  • Field Museum Collection, Chicago, Illinois (C. Moreau)
  • Forel A. 1886. Études myrmécologiques en 1886. Annales de la Société Entomologique de Belgique. 30: 131-215.
  • Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. J. Bombay Nat. Hist. Soc. 8: 396-420.
  • Forel A. 1903. Les fourmis des îles Andamans et Nicobares. Rapports de cette faune avec ses voisines. Rev. Suisse Zool. 11: 399-411.
  • Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
  • Forel A. 1909. Ameisen aus Java und Krakatau beobachtet und gesammelt von Herrn Edward Jacobson. Notes Leyden Mus. 31: 221-232.
  • Forel A. 1911. Ameisen aus Ceylon, gesammelt von Prof. K. Escherich (einige von Prof. E. Bugnion). Pp. 215-228 in: Escherich, K. Termitenleben auf Ceylon. Jena: Gustav Fischer, xxxii + 262 pp.
  • Forel A. 1911. Ameisen aus Java beobachtet und gesammelt von Herrn Edward Jacobson. II. Theil. Notes Leyden Mus. 33: 193-218.
  • Forel A. 1912. Ameisen aus Java beobachtet und gesammelt von Edward Jacobson. III. Theil. Notes Leyden Mus. 34: 97-112
  • Forel A. 1913k. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
  • Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
  • Forel, A. 1908. Fourmis de Ceylan et d'Égypte récoltées par le Prof. E. Bugnion. Lasius carniolicus. Fourmis de Kerguelen. Pseudandrie? Strongylognathus testaceus. Bull. Soc. Vaudoise Sci. Nat. 44: 1-22
  • Gillison A.N. 2000. Above ground biodiversity assesment working group summary report 1996-99: Impact of different land uses on biodiversity and social indicators. ASB Working Group Report, ICRAF, Nairobi, 160pp. http://www.asb.cgiar.org/PDFwebdocs/ASB Biodiversity Report.pdf
  • Gumawardene, N.R., J.D. Majer and J.P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2:71-83
  • Gunawardene N. R., J. D. Majer, and J. P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2: 71-83.
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Hannan M.A. 2007. Ant records from Savar, Dhaka, Bangladesh. Asian Myrmecology 1: 101-104.
  • Hannan M.A. 2008. New ant records fro Bangladesh. Asian Myrmecology 2: 95-98.
  • Hatter S. J. S., Sen, M. Nibedita, R. Mathew, and S. Sharma. 2004. Faunal diversity of Saipung wild life sanctuaryl Narpuh Reserve forest, laintia Hills, Meghalaya, Conservation Area Series 21 : 1-66.
  • Helms J. A., S. M. Helms, N. I. Fawzi, Tarjudin, F. Xaverius. 2017. Ant community of an Acacia mangium forest in Indonesian Borneo. Serangga 22(1): 147-159.
  • Herwina H., N. Nasir, Jumjunidang, and Yaherwandi. 2013. The composition of ant species on banana plants with Banana Bunchy-top Virus (BBTV) symptoms in West Sumatra, Indonesia. Asian Myrmecology 5: 151-161.
  • Herwina H., R. Satria, Yaherwandi, and Y. Sakamaki. 2018. Subterranean ant species diversity (Hymenoptera: Formicidae) in educational and biological research forest of universitas andalas, Indonesia. Journal of Entomology and Zoology Studies 6(1): 1720-1724.
  • Herwina H., and K. Nakamura. 2007. Ant species diversity study using pitfall traps in a small yard in Bogor Botanic garden, West Java, Indonesia. Treubia 35: 99-116.
  • Imai H. T., C. Baroni Urbani, M. Kubota, G. P. Sharma, M. H. Narasimhanna, B. C. Das, A. K. Sharma, A. Sharma, G. B. Deodikar, V. G. Vaidya, and M. R. Rajasekarasetty. 1984. Karyological survey of Indian ants. Japanese Journal of Genetics 59: 1-32.
  • Imai H. T., M. Kubota, W. L. Brown, Jr., M. Ihara, M. Tohari, and R. I. Pranata. 1985. Chromosome observations on tropical ants from Indonesia. Annu. Rep. Natl. Inst. Genet. Jpn. 35: 46-48.
  • Imai H. T.; Brown, W. L., Jr.; Kubota, M.; Yong, H.-S.; Tho, Y. P. 1984. Chromosome observations on tropical ants from western Malaysia. II. Annual Report. National Institute of Genetics, Japan 34:66-69.
  • Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
  • Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
  • Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
  • Kaleeswaran B., S. Ezil, B. P. Ganesh, and S. Bhavatarini. 2008. Biodiversity and niches of Ants in Alagar hills, Tamil nadu Wildlife Biodiversity Conservation Published by Day publishing house pp 188-208.
  • Karavaiev V. 1930. Ameisen von den Molukken und Neuguinea. (Ergebnisse der Sunda-Expedition der Notgemeinschaft der deutschen Wissenschaft 1929/30.). Zool. Anz. 92: 206-214.
  • Karmaly K. A.; S. Sumesh, T. P. Rabeesh, and L. Kishore. 2010. A checklist of ants of Thirunelli in Wayanad, Kerala. J. of the Bombay Natural History Society 107(1): 64-67.
  • Kishimoto-Yamata K., F. Hyodo, M. Matsuoka, Y. Hashimoto, M. Kon, T. Ochi, S. Yamane, R. Ishii, and T. Itioka. 2012. Effects of remnant primary forests on ant and dung beetle species diversity in a secondary forest in Sarawak, Malaysia. Journal of Insect Conservation DOI 10.1007/s10841-012-9544-6
  • Kitching R.L. 1987. Aspects of the natural history of the lycaenid butterfly Allotinus major in Sulawesi. Journal of Natural History 21: 535-544.
  • Kosgamage K. R. K. A., and R. K. S. Dias 2008. Systematics and community composition of Foraging worker ants ((Order: Hymenoptera, Family: Formicidae) collected from three habitats on a dry zone region of Sri Lanka.Proceedings of Postgraduate Symposium of Kelaniya University. 115pp.
  • Latumahina F., M. Borovanska, N. S. Putra, and M. Janda. 2015. Ants of Ambon Island – diversity survey and checklist. ZooKeys 472: 43–57.
  • Leong C. M., S. F. Shiao, and B. Guenard. 2017. Ants in the city, a preliminary checklist of Formicidae (Hymenoptera) in Macau, one of the most heavily urbanized regions of the world. Asian Myrmecology 9: e009014.
  • Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
  • Lopwichan S., and S. Khachonpisitsak. 2015. Ant diversity in Nong Tha Yu Arboretum, Si Racha District, Chon Buri Province. Proceedings The 7 th National Science Research Conference. 30-31 March 2015. Naresuan University.
  • Lu Z. X., Y. Q. Chen, Q. Li, S. M. Wang, C. J. Liu, and W. Zhang. 2012. Effect of lac insect honeydew on the diversity of ground-dwelling ants in lac plantation. Chin. J. Appl. Ecol. 23(4): 1117-1122.
  • Lu Z., B. D. Hoffmann, and Y. Chen. 2016. Can reforested and plantation habitats effectively conserve SW China’s ant biodiversity? Biodivers. Conserv. DOI 10.1007/s10531-016-1090-1
  • Lu Z., Y. Chen, Q. Li, S. Wang, C. Liu, and W. Zhang. 2012. Effect of population of Kerria yunnanensis on diversity of ground dwelling ant. Acta Ecologica Sinica 32(19): 6195-6202.
  • Lu Z., and Y. Chen. 2016. Effects of habitat on ant functional groups: a case study of Luchun County, Yunnan Province, China. Chinese Journal of Eco-Agriculture 24(5): 801-810.
  • Masayuki, Morita et al. 200?. Implication of predation incidences by ant species on green turtle nests in Chagar Hutang, Redang Island. Proc. 3rd. Intl. Symp. SEASTAR and Asian Bio-logging Science. Part I: Sea Turtle. 21-26.
  • Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
  • Mathew, R. 2000. Insecta : Hymenoptera : Formicidae. State Fauna Series. , Fauna of Tripura , Zool. Surv. India. 7 (3) : 347-354.
  • Menozzi C. 1930. Formiche di Sumatra raccolte dal Prof. J. C. van der Meer Mohr. Miscellanea Zoologica Sumatrana 47: 1-5.
  • Menozzi, C. 1932. Formiche dell'Isola di Nias. Misc. Zool. Sumatr. 65: 1-13
  • Mezger D., and M. Pfeiffer. 2011. Influence of the arrival of Anoplolepis gracilipes (Hymenoptera: Formicidae) on the composition of an ant community in a clearing in Gunung Mulu National Park, Sarawak, Malaysia. Asian Myrmecology 4: 89-98.
  • Mohamed M. 1995. A preliminary list of ants (Hymenoptera: Formicidae) of the Tawau Hills Park, Sabah. In: Ghazally Ismail et al. (eds.), Tawau Hills Park, Sabah, Pelanduk Pub. Pp. 205-213.
  • Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
  • Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
  • Mukherji D., and S. Ribeiro. 1925. On a collection of ants (Formicidae) from the Andaman Islands. Records of the Indian Museum 27: 205-209.
  • Musthak Ali T. M. 1992. Ant Fauna of Karnataka-2. Newsletter of IUSSI Indian Chapter 6(1-2): 1-9.
  • Narenda, A. and T.V. Ramachandra. 2008. Remote detection and distinction of ants using nest-site specific LISS-derived Normalised Difference Vegetation Index. Asian Myrmecology 2:51-62.
  • Narendra A., H. Gibb, and T. M. Ali. 2011. Structure of ant assemblages in Western Ghats, India: role of habitat, disturbance and introduced species. Insect Conservation and diversity 4(2): 132-141.
  • Ngoc Anh L., K. Ogata, and S. Hosoishi. 2010. Ants of agricultural fields in Vietnam (Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 33: 1-11.
  • Nie L., M. Ni, D. Ning, H. Ran, B. Hassan, and Y. Xu. 2019. Comparing mechanisms of competition among introduced and resident ants in China: from behavior to trophic position (Hymenoptera: Formicidae). Myrmecological News 29: 125-133.
  • Ogata K. 2005. Asian ant inventory and international networks. Report on Insect inventory Project in Tropic Asia TAIIV: 145-170.
  • Overbeck, H. (1924). A List of Ants from Singapore and Neighbouring Places. Journal of the Malayan Branch of the Royal Asiatic Society 2 (1):25-40.
  • Parui A. K., S. Chatterjee, and P. Basu. 2015. Habitat characteristics shaping ant species assemblages in a mixed deciduous forest in Eastern India. Journal of Tropical Ecology: 1-14. doi:10.1017/S0266467415000036
  • Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
  • Pfeiffer, M., H. Cheng Tuck, and T. Chong Lay. 2008. Exploring arboreal ant community composition and co-ccurrence patterns in plantations of oil palm Elaeis guineensis in Borneo and Peninsular Malaysia. Ecography 31(1): 21-32.
  • Philpott S.M., P. Bichier, R.A. Rice, and R. Greenberg. 2008. Biodiversity conservation, yield, and alternative products in coffee agroecosystems in Sumatra, Indonesia. Biodivers. Conserv. 17: 1805-1820. Data obtained from Stacy Philpott
  • Rajagopal T., S. P. Sevarkodiyone, and A. Manimozhi. 2005. Ant diversity in some selected localities of Sattur Taluk, Virudhunagar district, Tamil Nadu. Zoos' Print Journal 20(6): 1887-1888.
  • Ran H., and S. Y. Zhou. 2011. Checklist of Chinese Ants: the Formicomorph Subfamilies (Hymenoptera: Formicidae) (I). Journal of Guangxi Normal University: Natural Science Edition. 29(3): 65-73.
  • Rickson F.R., and M.M. Rickson. 1998. The cashew nut, Anacardium occidentale (Anacardiaceae), and its perennial association with ants: extrafloral nectary location and the potential for ant defense. American Journal of Botany 85(6): 835-849.
  • Rizali A., A. Rahim, B. Sahari, L.B. Prasetyo, and D. Buchori. 2011. Impact of invasive ant species in shaping ant community structure on small islands in Indonesia. Jurnal Biologi Indonesia 7(2): 221-230.
  • Rizali A., D. J. Lohman, D. Buchori, L. Budi Prasetyo, H. Triwidodo, M. M. Bos, S. Yamane, and C. H. Schulze. 2009. Ant communities on small tropical islands: effects of island size and isolation are obscured by habitat disturbance and tramp ant species. Journal of Biogeography 37(2): 229-236.
  • Rizali A., M. M. Bos, D. Buchori, Sk. Yamane, and C. H. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 15(2): 77-84.
  • Rizali A., M.M. Bos, D. Buchori, Sk. Yamane, C. Hans, and J. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 77-84.
  • Rizali A., Y. Clough, D. Buchori, M. L . A. Hosang, M. M. Bos, and T. Tscharntke. 2012. Long-term change of ant community structure in cacao agroforestry landscapes in Indonesia. Insect Conservation and Diversity doi: 10.1111/j.1752-4598.2012.00219.x
  • Rothney G. A. J. 1889. Notes on Indian ants. Trans. Entomol. Soc. Lond. 1889: 347-374.
  • Rothney G. A. J. 1903. The aculeate Hymenoptera of Barrackpore, Bengal. Trans. Entomol. Soc. Lond. 1903: 93-116.
  • Rubiana R., A. Rizali, L. H. Denmead, W. Alamsari, P. Hidaya, Pudjianto, D. Hindayana, Y. Clough, T. Tscharntke, and D. Buchori. 2015. Agricultural land use alters species composition but not species richness of ant communities. Asian Myrmecology 7, .
  • Santschi F. 1924. Fourmis d'Indochine. Opuscules de l'Institut Scientifique de l'Indochine 3: 95-117
  • Schmidt F. A., C. R. Ribas, T. G. Sobrinho, R. Ubaidillah, J. H. Schereder, Y. Clough, and T. Tscharntke. 2017. Similar alpha and beta diversity changes in tropical ant communities, comparing savannas and rainforests in Brazil and Indonesia. Oecologia DOI 10.1007/s00442-017-3960-y
  • Sinu P. A., M. Nasser, and P. D. Rajan. 2006. Feeding fauna and foraging habits of tiger beetles found in agro-ecosystems in Western Ghats, India. Biotropica 38(4): 500-507.
  • Sitthicharoenchai D., and N. Chantarasawat. 2006. Ant species diversity in the establishing area for Advanced Technology Institute at Lai-Nan Sub-district, Wiang Sa district, Nan Province, Thailand. The Natural History Journal of Chulalongkorn University 6(2): 67-74
  • Song Y., Z Xu, C Li, N. Zhang, L. Zhang, H. Jiang, and F Mo. 2013. An analysis on the ant fauna of the Nangun River Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
  • Song Y., Z. Xu, C. Li, N. Zhang, L. Zhang, H. Jiang, and F. Mo. 2013. An Analysis on the Ant Fauna of the Nangun river Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
  • Stitz H. 1912. Ameisen aus Ceram und Neu-Guinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1912: 498-514.
  • Stitz H. 1925. Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1923: 110-136.
  • Tak N., and S. I. Kazmi. 2013. On some ants (Insecta: Hymenoptera: Formicidae) from Nagaland, India. Rec. zool. Surv. India: 113(1): 169-182.
  • Thomas S. K., and P. Aswathi. 2012. Potential of rubber litter dwelling ants as biocontrol agent of home invading nuisance pest, Luprops tristis. Jbiopest 5: 188-191.
  • Tiwari R. N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4): 1-96.
  • Tiwari R. N., B. G. Kundu, S. Roy Chowdhury, and S. N. Ghosh. 2003. Insecta: Hymenoptera: Formicidae. Fauna of Sikkim. Part 4. State Fauna Series. 9.Zool.Surv.India. i-iii, 1-512. Chapter pagination: 467-506.
  • Tiwari R.N., B.G. Kundu, S. Roychowdhury, S.N. Ghosh. 1999. Insecta: Hymenoptera: Formicidae. Pp. 211-294 in: Director; Zoological Survey of India (ed.) 1999. Fauna of West Bengal. Part 8. Insecta (Trichoptera, Thysanoptera, Neuroptera, Hymenoptera and Anoplura). Calcutta: Zoological Survey of India, iv + 442 pp.
  • Tiwari R.N., B.G. Kundu, S. Sheela, and S. N. Ghosh. 2004. Insecta: Hymenoptera: Formicidae. Alfred, J.R.B. [Ed.], Fauna of Manipur, part - 2 (insects). State fauna series 10. Zool.Surv.India.i-v, 1-625. Chapter pagination: 605-625.
  • Tiwari, R.N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4):1-96
  • Tjan K. N., H. T. Imai, M. Kubota, W. L., Jr., Brown, W. H. Gotwald, H.-S. Yong, and C. Leh. 1986. Chromosome observations of Sarawak ants. Annu. Rep. Natl. Inst. Genet. Jpn. 36: 57-58.
  • Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
  • Viehmeyer H. 1914. Ameisen aus Perak, Bali und Ceram (Hym.) (Freiburger Molukken-Expedition), gesammelt von E. Streesemann. Entomologische Mitteilungen. Berlin-Dahlem 3: 112-116.
  • Wang W. R., S. Q. Zhang, and Z. H. Xu. 2012. A faunistic and taxonomic study of ants (Hymenoptera: Formicidae) in Shenzhen Municipality. Journal of Southwest Forestry University 32(1): 64-73.
  • Way M. J., and B. Bolton. 1997. Competition between ants for coconut palm nesting sites. Journal of Natural History 31: 439-455.
  • Wetterer J. K. 2005. Woldwide distribution and potential spread of the long-legged ant, Anoplolepis gracilipes (Hymenoptera: Formicidae). Sociobiology 45: 77-97.
  • Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
  • Wheeler W. M. 1921. Chinese ants collected by Prof. C. W. Howard. Psyche (Cambridge) 28: 110-115.
  • Wheeler W. M. 1927. Ants collected by Professor F. Silvestri in Indochina. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 20: 83-106.
  • Wheeler W. M. 1930. A list of the known Chinese ants. Peking Natural History Bulletin 5: 53-81.
  • Wheeler W. M. 1937. Additions to the ant-fauna of Krakatau and Verlaten Island. Treubia 16: 21-24.
  • Wheeler, William Morton. 1924. Ants of Krakatau and Other Islands in the Sunda Strait. Treubia. 5(1-3):1-20.
  • Woodcock P., D. P. Edwards, R. J. Newton, C. Vun Khen, S. H. Bottrell, and K. C. Hamer. 2013. Impacts of Intensive Logging on the Trophic Organisation of Ant Communities in a Biodiversity Hotspot. PLoS ONE 8(4): e60756. doi:10.1371/journal.pone.0060756
  • Woodcock P., D. P. Edwards, T. M. Fayle, R. J. Newton, C. Vun Khen, S. H. Bottrell, and K. C. Hamer. 2011. The conservation value of South East Asia's highly degraded forests: evidence from leaf-litter ants. Phil. Trans. R. Soc. B. 366: 3256-3264.
  • Wu B., Y. Lu, G. Liang, and L. Zeng. 2010. Influence of the red inported fire ant, Solenopsis invicta Buren (Hymenoptera: Formicidae) on the diversity of ant communities in a newly infested longan orchard and grass areas nearby. Acta Ecologica Sinica 30(8): 2075-2083.
  • Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
  • Yamane S., T. Itino, and A.R. Nona. 1996. Ground ant fauna in a Bornean dipterocarp forest. Raffles Bulletin of Zoology 44(1): 253-262.
  • Yamane S.; Bui T. V.; Ogata K.; Okido H.; Eguchi K. 2002. Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture Kyushu University 25: 51-62.
  • Yamane S.; Nona, A. R. 1994. Ants from Lambir Hills National Park, Sarawak. Pp. 222-226 in: Inoue, T.; Hamid, A. A. (eds.) 1994. Plant reproductive systems and animal seasonal dynamics. Long-term study of dipterocarp forests in Sarawak. Kyoto: Center for Ecological Research, Kyoto University, vii + 255 pp. 
  • Yamane Sk. 2005. Krakatau in 1982, and the commencement of myrmecological research. The nature and Insects (Konchu to shizen) 40: 27-33.
  • Zhang N. N., Y. Q. Chen, Z. X. Lu, W. Zhang, and K. L. Li. 2013. Species diversity, community structure difference and indicator species of leaf-litter ants in rubber plantations and secondary natural forests in Yunnan, southwestern China. Acta Entomologica Sinica 56(11): 1314-1323.
  • Zhou S.-Y. 2001. Ants of Guangxi. Guangxi Normal University Press, Guilin, China, Guilin, China. 255 pp.
  • Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.
  • Zryanin V. A., and M. V. Mokrousov. 2015. Contribution to the ant fauna of Lombok Island. Proceedings of the 10th ANeT International Conference, 23-26 October 2015, University of Kelaniya, Sri Lanka. 34