Plagiolepis pallescens

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Plagiolepis pallescens
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Plagiolepidini
Genus: Plagiolepis
Species: P. pallescens
Binomial name
Plagiolepis pallescens
Forel, 1889

Plagiolepis pallescens casent0906084 p 1 high.jpg

Plagiolepis pallescens casent0906084 d 1 high.jpg

Specimen Labels

Subspecies
Synonyms

This minute species lives in small isolated colonies nesting under flat stones usually with several queens (Collingwood 1979). In Russia it occurs in steppe habitats on rocky slopes with nests in soil (Zryanin & Zryanina, 2007). It is common in Greece, being known from all Greek provinces except Epirus. It is a thermophilous species that prefers open and warm habitats. In Achaia and Aetolia-Acarnania, its nests were observed under stones in rural sites in urban areas, alpine pastures and stream valleys with plane trees (Borowiec & Salata, 2021).

At a Glance • Ergatoid queen  

Identification

Distribution

This is a common species, recorded from all Greek provinces except Epirus (Borowiec et al., 2022).

Latitudinal Distribution Pattern

Latitudinal Range: 50.513° to 28.133333°.

   
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Palaearctic Region: Albania, Armenia, Austria (type locality), Azerbaijan, Belgium, Bulgaria, Channel Islands, China, Croatia, Cyprus, Czech Republic, Democratic Peoples Republic of Korea, France, Germany, Greece (type locality), Hungary, Iran, Israel, Kyrgyzstan, Mongolia, Montenegro, Republic of Korea, Romania, Russian Federation, Slovenia, Spain, Switzerland, Türkiye (type locality), Ukraine (type locality), United Kingdom of Great Britain and Northern Ireland.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Biology

Association with Other Organisms

Explore-icon.png Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.
  • This species is a host for the ant Plagiolepis ampeloni (a workerless inquiline).
  • This species is a host for the ant Plagiolepis delaugerrei (a workerless inquiline) in Corsica, France (Casevitz-Weulersse, J. 2014).
  • This species is associated with the aphids Acyrthosiphon pisum, Aphis craccivora, Aphis fabae, Aphis frangulae, Aphis spiraecola, Brachycaudus (Appelia) tragopogonis, Brachycaudus tragopogonis, Callaphis juglandis, Hyalopterus amygdali, Staegeriella necopinata, Tinocallis platani and Toxoptera aurantii (Saddiqui et al., 2019 and included references).

Fungi

  • This species is a host for the fungus Myrmicinosporidium durum (a parasitoid) in Austria (Sanchez-Peña et al., 1993; Espadaler & Santamaria, 2012) (as Plagiolepis vindobonensis).
  • This species is a host for the fungus Myrmicinosporidium durum (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission within nest).

Flight Period

X X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

Castes

Ergatoid queens reported by Espadaler (2007)

Images from AntWeb

Plagiolepis pallescens casent0906253 h 1 high.jpgPlagiolepis pallescens casent0906253 d 1 high.jpgPlagiolepis pallescens casent0906253 p 1 high.jpgPlagiolepis pallescens casent0906253 l 1 high.jpg
Worker. Specimen code casent0906253. Photographer Estella Ortega, uploaded by California Academy of Sciences. Owned by NHMUK, London, UK.
Plagiolepis taurica cfh000040 l 1 high.jpgPlagiolepis taurica cfh000040 h 1 high.jpgPlagiolepis taurica cfh000040 d 1 high.jpgPlagiolepis taurica cfh000040 p 1 high.jpg
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Plagiolepis vindobonensis casent0281165 d 1 high.jpgPlagiolepis vindobonensis casent0281165 h 1 high.jpgPlagiolepis vindobonensis casent0281165 p 1 high.jpgPlagiolepis vindobonensis casent0281165 l 1 high.jpg
Worker. Specimen code casent0281165. Photographer Cerise Chen, uploaded by California Academy of Sciences. Owned by PSWC, Philip S. Ward Collection.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • pallescens. Plagiolepis pygmaea var. pallescens Forel, 1889: 265 (w.) GREECE.
    • [Misspelled as pallecens by Santschi, 1920g: 168.]
    • Santschi, 1911i: 287 (q.); Finzi, 1928c: 790 (m.).
    • Subspecies of pygmaea: Forel, 1890a: lxvii; Forel, 1889: 265; Emery, 1891b: 16; Dalla Torre, 1893: 173; Forel, 1902a: 155; Ruzsky, 1902d: 18; Ruzsky, 1903b: 308; Ruzsky, 1905b: 467; Forel, 1911d: 350; Forel, 1911h: 457; Santschi, 1911i: 287; Santschi, 1920g: 168; Ruzsky, 1923: 3.
    • Status as species: Emery, 1921d: 315; Emery, 1925b: 21; Karavaiev, 1926e: 188; Wheeler, W.M. 1927g: 114; Karavaiev, 1927a: 294; Finzi, 1928c: 790; Karavaiev, 1931a: 311; Menozzi, 1936d: 297; Finzi, 1936: 184; Arnol'di, 1948: 212 (in list); Baroni Urbani, 1971c: 168; Tarbinsky, 1976: 127 (redescription); Aktaç, 1977: 124; Dlussky, 1981a: 17; Agosti & Collingwood, 1987a: 57; Agosti & Collingwood, 1987b: 280 (in key); Dlussky, Soyunov & Zabelin, 1990: 162; Collingwood, 1993b: 195; Arakelian, 1994: 78; Bolton, 1995b: 335; Mei, 1995: 767; Radchenko, 1996a: 184; Schultz, R. et al. 2006: 205; Gratiashvili & Barjadze, 2008: 134; Csösz, et al. 2011: 58; Legakis, 2011: 25; Borowiec, L. & Salata, 2012: 528; Borowiec, L. & Salata, 2013: 364; Borowiec, L. 2014: 146 (see note in bibliography); Salata, Borowiec & Radchenko, 2018: 812 (redescription); Salata & Borowiec, 2018c: 48.
    • Senior synonym of ancyrensis: Salata, Borowiec & Radchenko, 2018: 812.
    • Senior synonym of satunini: Salata, Borowiec & Radchenko, 2018: 812 (by implication).
    • Senior synonym of taurica: Salata, Borowiec & Radchenko, 2018: 812.
    • Senior synonym of vindobonensis: Salata, Borowiec & Radchenko, 2018: 812.
    • Current subspecies: nominal plus atlantis, elguerrhae, isis, kabyla, polygyna, sordida.
  • ancyrensis. Plagiolepis maura var. ancyrensis Santschi, 1920g: 171, fig. 1 (w.q.) TURKEY.
    • [Misspelled as angyrensis by Santschi, 1921a: 114.]
    • Finzi, 1936: 184 (m.).
    • Subspecies of maura: Santschi, 1921a: 114; Emery, 1921b: 208.
    • Subspecies of pallescens: Emery, 1925b: 21; Menozzi, 1933b: 78; Finzi, 1936: 184.
    • Status as species: Tohmé, G. 1969: 10; Kugler, J. 1988: 258; Bolton, 1995b: 334; Legakis, 2011: 25; Borowiec, L. & Salata, 2012: 528; Borowiec, L. 2014: 145; Pashaei Rad et al., 2018: 7.
    • Junior synonym of pallescens: Salata, Borowiec & Radchenko, 2018: 812.
  • satunini. Plagiolepis (Plagiolepis) satunini Karavaiev, 1931a: 314, fig. 4 (w.) ARMENIA.
    • Status as species: Arakelian, 1994: 80; Bolton, 1995b: 336.
    • Junior synonym of taurica: Radchenko, 1996a: 185; Radchenko, 2016: 343.
    • Junior synonym of pallescens: Salata, Borowiec & Radchenko, 2018: 812 (by implication).
  • taurica. Plagiolepis maura var. taurica Santschi, 1920g: 171 (w.) RUSSIA (Crimea), TURKMENISTAN.
    • Subspecies of pallescens: Emery, 1921d: 314; Santschi, 1922a: 67; Emery, 1925b: 21; Karavaiev, 1927a: 294.
    • Subspecies of pygmaea: Karavaiev, 1931a: 309.
    • Status as species: Radchenko, 1989b: 155; Bolton, 1995b: 336; Radchenko, 1996a: 185; Markó, Sipos, et al. 2006: 69; Schultz, R. et al. 2006: 205; Zryanin & Zryanina, 2007: 232; Casevitz-Weulersse & Galkowski, 2009: 485; Lapeva-Gjonova, et al. 2010: 34; Csösz, et al. 2011: 58; Legakis, 2011: 25; Borowiec, L. & Salata, 2012: 529; Borowiec, L. & Salata, 2013: 364; Borowiec, L. 2014: 147; Bračko, et al. 2014: 20; Casevitz-Weulersse, 2014: 28; Radchenko, 2016: 343; Borowiec, L. & Salata, 2018: 7.
    • Junior synonym of pallescens: Salata, Borowiec & Radchenko, 2018: 812.
  • vindobonensis. Plagiolepis vindobonensis Lomnicki, 1925c: 77, figs. 1-4 (w.) AUSTRIA.
    • Santschi, 1926f: 293 (q.); Kratochvíl, in Kratochvíl, et al. 1944: 84 (m.).
    • Subspecies of pygmaea: Bernard, 1967: 275 (in key).
    • Status as species: Santschi, 1926f: 288; Stärcke, 1936: 278; Novák & Sadil, 1941: 99 (in key); Kratochvíl, in Kratochvíl, et al. 1944: 82; van Boven, 1946: 9; van Boven, 1947: 181 (in key); Kutter, 1968a: 60; Baroni Urbani, 1971c: 172; Bolton & Collingwood, 1975: 5 (in key); van Boven, 1977: 129; Kutter, 1977c: 189; Collingwood, 1979: 110; Agosti & Collingwood, 1987a: 57; Agosti & Collingwood, 1987b: 280 (in key); Casevitz-Weulersse, 1990c: 427; Atanassov & Dlussky, 1992: 203; Collingwood, 1993b: 195; Seifert, 1994: 29; Bolton, 1995b: 336; Csösz, & Markó, 2005: 227; Bračko, 2006: 151; Bračko, 2007: 20; Seifert, 2007: 259; Werner & Wiezik, 2007: 142; Boer, 2010: 46; Legakis, 2011: 25.
    • Junior synonym of pygmaea: Bernard, 1967: 277 (in taxonomic synopsis and footnote).
    • Junior synonym of taurica: Radchenko, 1989b: 155; Radchenko, 1996a: 185; Markó, Sipos, et al. 2006: 69; Casevitz-Weulersse & Galkowsky, 2009: 485; Lapeva-Gjonova et al. 2010: 34; Csösz, et al. 2011: 58; Borowiec, L. 2014: 147; Casevitz-Weulersse, 2014: 28; Radchenko, 2016: 343.
    • Junior synonym of pallescens: Salata, Borowiec & Radchenko, 2018: 812.

Taxonomic Notes

Pashaei Rad et al. (2018) treat the junior synonyms Plagiolepis ancyrensis and Plagiolepis taurica as a valid species although they provide no evidence to support their treatment of either name as representing distinct species. Note that Bolton (in his unpublished An online catalogue of the ants of the world, 15 Feb. 2020) does not recognise the species status proposed by Pashaei Rad et al. (2018), noting that "the entries in this publication are omitted from the catalogue as it contains numerous nomenclatural errors; it also ignores previous taxonomic decisions, and entirely fails to apply ICZN rules that govern the availability of names in the species-group." Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Worker

Sharaf et al. (2011) - Syntype. TL 1.49, HL 0.45, HW 0.39, SL 0.39, EL 0.12, CI 87, SI 100, PRW 0.25, ML 0.46.

Yellowish brown, gaster yellowish paler than rest of body, legs and antennae yellow, propodeum dorsum characteristically reddish brown. Body smooth and shining. Head distinctly longer than broad with clearly convex sides. Eyes, in full-face view, fail to break head sides. Antennae with appressed pubescence. Second funicular segment as long as or little broader than long. Third funicular segment little longer than broad. Second and third funicular segments nearly subequal. Fourth funicular segment distinctly longer than broad, and little longer than broad. Scape surpassing posterior margin of head by 2-3 X its thickeness. Eyes large (EL 0.12), 0.3 x HW, with eleven ommatadia in the longest row. Anterior clypeal margin with several pairs of long hairs. Occiput nearly straight or feebly concave with strongly rounded posterior corners. Promesonotal suture indistinct. Metanotal groove feebly impressed. Erect hairs limited to anterior margin of clypeus and a small number on the gaster. Dorsum of head with abundant but widely spaced pubescence; other pubescence sparse.

Queen

Sharaf et al. (2011) - Syntype (gaster is broken) HL 0.63, HW 0.56, SL 0.53, EL 0.19, CI 89, SI 95, PRW 0.25, ML 0.92.

Body brown, legs and antennae yellowish brown. Eyes very large (EL 0.19) occupying about half head sides. Anterior clypeal margin with two pairs of long hairs. Palp formula 6,4. Second funicular segment about twice broader than long. Third funicular segment twice longer than broad, and twice longer than second. Fourth funicular segment clearly longer than broad and nearly as long as, or little shorter than, second and third segments combined. Occiput nearly straight or feebly concave with rounded posterior corners. Mesosoma robust, distinctly dorsoventrally compressed. Propodeal spiracle circular. Preonotal humeri with one pair of long hairs. Body with abundant appressed pubescence.

Male

Sharaf et al. (2011) - Syntype. TL 1.42-1.49, HL 0.38, HW 0.35-0.36, SL 0.34-0.38, EL 0.17, CI 92-95, SI 97-106, PRW 0.35-0.36, ML 0.61-0.63.

Body pale brown, legs yellow, all covered with relatively few sparse pubescence. Head distinctly broader than long. Occiput straight. Third funicular segment about twice longer than second segment. Fourth funicular segment as long as or little longer than second and third segments combined. Anterior clypeal margin with one pair of long hairs. Eyes prominent occupying more than half of head sides.

Type Material

Sharaf et al. (2011) - Syntype worker and two syntype males and one syntype queen, Morocco: Magador, iv.1905 (Vaucher) (Naturhistorisches Museum, Basel) [examined].

References

References based on Global Ant Biodiversity Informatics

  • Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
  • Aktaç, N. "Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon." Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41 (1977): 115-135.
  • Alvarado M., and L. Galle. 2000. Ant assemblages associated with lowland forests in the southern part of the great Hungarian plain. Acta Zoologica Academiae Scientarum Hungaricae 46(2): 79-102.
  • AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
  • Antarea (at www.antarea.fr on June 11th 2017)
  • Antonova V., and L. Penev. 2006. Change in the zoogeographical structure of ants (Hymenoptera: Formicidae) caused by urban pressure in the Sofia region (Bulgaria). Myrmecologische Nachrichten 8: 271-276.
  • Antonova V., and L. Penev. 2008. Classification of assemblages of ants in the green areas in Sofia City. Acta Zoologica Bulgarica 60(2): 103-110.
  • Bakr R. F. A., H. H. Fadl, R. M. Badawy, and M. R. Sharaf. 2007. Myrmecophile insects associated with some ant species (Hymenoptera : Formicidae )in Egypt. The second international conference of economic entomology (Entomological Society of Egypt) ,Cairo, Egypt, 8-11 December, (1): 205-233.
  • Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
  • Barrett K. E. 1967. Ants in South Brittany. Entomologist's Record and Journal of Variation 79:112-116.
  • Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
  • Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
  • Baugnee J. Y. 2002. Nouvelles observations de Plagiolepis vindobonensis Lomnicki, 1925 en Belgique (Hymenoptera, Formicidae). Bulletin Société Royale Belge d’Entomologie 138-133-134.
  • Baugnee J. Y., and J. F. Godeau. 2000. Signalement de Monomorium monomorium Bolton et de quelques autres fourmis à Belle-Ile-en-Mer, en Bretagne (Hymenoptera, Formicidae). Bulletin de la Société entomologique de France 105(2): 205-208.
  • Bernard, F. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
  • Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
  • Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
  • Boer P., W. Dekoninck, A. J. Van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten (Amsterdam) 63: 54-58.
  • Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
  • Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
  • Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
  • Borowiec L., C. Galkowski, and S. Salata. 2015. What is Tetramorium semilaeve André, 1883? (Hymenoptera, Formicidae). ZooKeys 512:39-62.
  • Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
  • Borowiec L., and S. Salata. 2013. Ants of Greece – additions and corrections (Hymenoptera: Formicidae). Genus (Wroclaw) 24(3-4): 335-401.
  • Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
  • Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) of Samos Island, Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-13.
  • Borowiec M. L., L. Borowiec, S. Csősz, and A. Radchenko. 2009. Ants collected during 2006 Polish expedition to Kyrgyzstan (Hymenoptera: Formicidae). Genus (Wroclaw) 20: 367-379.
  • Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
  • Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
  • Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
  • Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
  • Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
  • Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
  • Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
  • Casevitz-Weulersse J. 2014. A propos des fourmis de la Corse: Plagiolepis delaugerrei n. sp., parasite de Plagiolepis taurica Santschi, 1920 (Insecta: Hymenoptera: Formicidae: Formicinae). Le bulletin d’Arthropoda 47: 27-33.
  • Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
  • Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
  • Casevitz-Weulersse, J. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
  • Choi B.-M. 1987. Taxonomic study on ants (Formicidae) in Korea (1). On the genus Monomorium. Journal of the Institute of Science Education (Cheongju National Teachers' College) 11:17-30.
  • Choi B.M. 1986. Studies on the distribution of ants (Formicidae) in Korea. Journal of Chongju National Teacher College 23: 317-386.
  • Choi B.M. 1988. Studies on the distribution of ants (Formicidae) in Korea (5) Ant fauna in Is. Kanghwado. Chongju Sabom Taehak Nonmunjip (Journal of Chongju National Teacher' College) 25: 217-231.
  • Choi B.M. 1996. Distribution of ants (Formicidae) in Korea (16) - Ant fauna from Chollabukdo. Korean J. Soil. Zoology 1(1): 5-23.
  • Choi B.M. 1996. Distribution of ants (Formicidae) in Korea (16): Ant fauna from Chollabukdo. Korean Journal of Soil Zoology 1(1): 5-23.
  • Choi B.M. 1996. Studies on the distribution of ants (Formicidae) in Korea (15) -Ant fauna islands Ullungdo and Dokdo. Journal of Chongju National University of Education 33: 201-219.
  • Choi B.M. 1997. Distribution of Ants (Formicidae) in Korea (18). Ants Fauna in island Paekryongdo and Taechongdo. Journal of Chongju National University of Education 34: 119-138.
  • Choi B.M. 1998. Distribution of ants (Formicidae) in Korea (19) Ants fauna from Chungcheongbukdo Province. Cheongju Tea. Coll. 35: 213-266.
  • Choi B.M. and C.H. Kim, 1987, Studies on the distribution of ants (Formicidae) in Korea (4). Ant fauna in Is. Hongdo and Is. Taehukusando. Journal of Chongju National Teacher College 24: 357-370.
  • Choi B.M., Bang, J.R. 1992. Studies on the distribution of ants (Formicidae) in Korea (9). Ant fauna in Mt. Togyusan. Korean Journal of Applied Entomology 31:101-112.
  • Choi B.M., Kim, C.H., Bang, J.R. 1993. Studies on the distribution of ants (Formicidae) in Korea (13). A checklist of ants from each province (Do), with taxonomic notes. Cheongju Sabom Taehakkyo Nonmunjip (Journal of Cheongju National University of Education) 30: 331-380.
  • Choi B.M.; Bang, J.R. 1992. Studies on the distribution of ants (Formicidae) in Korea (10). Ant distribution in Gangweon Do. Journal of the Institute of Science Education (Cheongju National Teachers' College) 14:12-30.
  • Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
  • Collingwood, C. A. 1993. A Comparitive Study of the Ant Fauna of Five Greek Islands. Biologia Gallo-hellenica. 20,1:191-197
  • Collingwood, C. A. "A comparative study of the ant fauna of five Greek islands." Biologia Gallo-Hellenica 20 (1993): 191-197.
  • Collingwood, C. A. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
  • Consani M. 1947. Reperti corologici sulle formiche italiane. Redia 32: 179-182.
  • Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
  • Csősz S. and Markó, B. 2005. European ant species (Hymenoptera: Formicidae) in the ant collection of the Natural History Museum of Sibiu (Hermannstadt/Nagyszeben), Romania II. Subfamily Formicinae. Annales Historico-Naturales Musei Nationalis Hungarici 97: 225-240.
  • Csősz S., B. Markó, and L. Gallé. 2001. Ants (Hymenoptera: Formicidae) of Stana Valley (Romania): Evaluation of the effectiveness of a myrmecological survey. – Entomologica Romanica 6 : 121-126.
  • Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
  • Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
  • Dlussky G. M., O. S. Soyunov, and S. I. Zabelin. 1990. Ants of Turkmenistan. Ashkabad: Ylym Press, 273 pp.
  • Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
  • Dvorak, L., P. BOGUSCH, I. MALENOVSKÝ, P. BEZDÌÈKA, K. BEZDÌÈKOVÁ, K. HOLÝ, P. LIŠKA, J. MACEK, L. ROLLER, M. RÍHA et al. "Hymenoptera of Hády Hill, near the city of Brno (Czech Republic), collected during the Third Czech-Slovak Hymenoptera meeting." Acta Musei Moraviae, Scientiae biologicae (Brno) 93 (2008): 53-92.
  • Emery C. 1921. Notes critiques de myrmécologie. [IX-X.]. Annales de la Société Entomologique de Belgique 61: 313-319.
  • Emery, C. "Exploration scientifique de la Tunisie. Zoologie. - Hyménoptères. Révision critique des fourmis de la Tunisie." Explor. Scient. De la Tunisie Zoll. Hym. (Folleto) Paris. Imp (1891): iii + 21 pp.
  • Espadaler X., F. Garcia, X. Roig, and R. Vila. 2013. Ants (Hymenoptera, Formicidae) from the Castell de Montesquiu park (Osona, north-east of the Iberian Peninsula). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 53: 223-227.
  • Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants – assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
  • Finzi, B. "Risultati scientifici della spedizione di S. A. S. il Principe Alessandro della Torre e Tasso nell'Egitto e peninsola del Sinai. XI. Formiche." Bulletin de la Société Entomologique d'Egypte 20 (1936): 155-210.
  • Forel A. 1890. Fourmis de Tunisie et de l'Algérie orientale. Annales de la Société Entomologique de Belgique 34: lxi-lxxvi.
  • Forel A. 1905. Miscellanea myrmécologiques II (1905). Ann. Soc. Entomol. Belg. 49: 155-185.
  • Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
  • Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
  • Forel A. 1911. Sur le genre Metapone n. g. nouveau groupe des Formicides et sur quelques autres formes nouvelles. Rev. Suisse Zool. 19: 445-459.
  • Forel, A. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
  • Forel, A. "Fourmis de Tunisie et de l'Algerie orientale recoltees et decrites par Auguste Forel." Annales de la Société Entomologique de Belgique 34 (1890): lxi-lxxvi.
  • Forel, A. "Les fourmis du Sahara algérien récoltées par M. le Professeur A. Lameere et le Dr. A. Diehl." Annales de la Société Entomologique de Belgique 46 (1902): 147-158.
  • GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
  • Galle L. 1972. Study of ant-populations in various grassland ecosystems. Acta Biologica Szeged 18(1-4): 159-164.
  • Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
  • Galle L. 1993. Data to the ant fauna of the Bukk (Hymenoptera: Formicidae). Natural history of the national parks of Hungary 7: 445-448.
  • Galle L., and G. Szonyi. 1988. A check list of ants (Hymenoptera: Formicoidea) of a sandy grassland in Kiskunsag National Park (Hungary). Acta Biol. Szeged 34: 167-168.
  • Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae). — In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
  • Gholami M., A. Afshari, and S. A. Mafi Pashakolaei. 2012. The fauna and frequency of cottony cushion scale (Icerya purchasi Maskell) related ants (Hymenoptera: Formicidae) community in citrus orchards of Sari region, northern Iran. 20th Iranian Plant Protection Congress, 25-28 August 2012.
  • Glaser F., A. Freitag, and H. Martz. 2012. Ants (Hymenoptera: Formicidae) in the Münstertal (Val Müstair) – a hot spot of regional species richness between Italy and Switzerland. Gredleriana 12: 273 - 284.
  • Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
  • Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
  • Gulzar A. 2014. Classification and Distribution of ants in Kashgar, Xinjiang. Master's Thesis Shaanxi Normal University, 75 pages.
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Huang Ren-Xing, Ouyang Tong, Wu Wei, and Fan Zhao-tian. 2004. Forty two new record species of Family Formicidae (Hymenoptera: Formicoidea) from Xinjiang, China. Entomotaxonomia 26 (2): 156-160.
  • Izhaki I., B. Idelovich, R. Laster, and Y. Ofer. 2009. The impact of macro- vs micro environmental factors on the structure of ant communities inhabiting East-Mediterranean Aleppo pine forests. Israel Journal of Entomology 39: 129-146.
  • Kanizsai O., R. Gallé, and L. Gallé. 2009. Perception of spatial patchiness by ant assemblages (Hymenoptera: Formicidae). Tiscia 37: 3-7.
  • Karaman C., K. Kiran, V. Aksoy, and Y. Camlitepe. 2017. A new species of the genus Camponotus (Mayr) (Hymenoptera, Formicidae) from Turkey. Turkish Journal of Zoology 41: 998-1004.
  • Karaman G. S., and M. G. Karaman 2005. Contribution to the knowledge of the ants (Hymenoptera: Formicidae) of Crna Gora (Montenegrao). Natura Montenegrina 5: 91-108.
  • Karaman M. G. 2004. Checklist of known species of ants (Hymenoptera, Formicidae) in the fauna of Montenegro. Natura Montenegrina 3: 83-92.
  • Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
  • Karavaiev V. 1927. Übersicht der Ameisenfauna der Krim nebst einigen Neubeschreibungen. Konowia 5: 281-303.
  • Karavaiev V. 1931. Myrmekologische Fragmente, III. Zool. Anz. 92: 309-317.
  • Khalili-Moghadam A., L. Borowiec, and A. Nemati. 2019. New records of ants (Hymenoptera: Formicidae) from the Chaharmahal va Bakhtiari Province of Iran with taxonomic comments. Polish Journal of Entomology 88 (2): 163–182.
  • Kim B.J.; Kim, K.G.; Lim, K.H. 1993. Systematic study of ants from Chejudo province. Korean Journal of Entomology 23: 117-138
  • Kim C.H., B.M. Choi, and J.R. Bang. 1992. Studies on the distribution of ants (Formicidae) in Korea (8)-Ant fauna in 10 islands, Chollanam-do. Korean J. Appl. Entomol. 31(4): 345-359.
  • Kim K.I., C.H. Kim, and B. Choi. 1989. The ant fauna of the southern shore in Gyeongsangnamdo, Korea. Journal of Gyeongsang Nat. Univ. 28(2): 213-226.
  • Kim et al. 1993. Systematic study of ants from Chejudo Province. Koran Journal of Entomology 23(3): 117-141.
  • Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
  • Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
  • Kiran, K., and N. Aktaç. "The social parasitic ant Plagiolepis xene Stärcke, 1936 new to Turkey (Hymenoptera: Formicidae)." Myrmecological News 10 (2007): 110.
  • Kratochvíl J., V. Novák, and J. Snoflák. 1944. Mohelno. Soubor práci venoványch studiu vyznamne památky prírodní. 5. Hymenoptera - Aculeata. Formicidae - Apidae - Vespidae. Arch. Svazu Ochr. Prír. Domov. Moravé 6: 1-155.
  • Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
  • Kuznetsov G. T. 1990. Comparative analysis of Hymenoptera (Formicidae) population on altitudinal zones of central Kopetdag. Izvestiya Akademii Nauk Turkmenskoi SSR. Seriya Biologicheskikh Nauk 1990(3): 64-67.
  • Lapeva-Gjonova A., and K. Kiran. 2012. Ant fauna (Hymenoptera, Formicidae) of Strandzha (Istranca) Mountain and adjacent Black Sea coast. North-western journal of Zoology 8(1): 72-84.
  • Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
  • Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24 – 33
  • Legakis Collection Database
  • Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
  • Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
  • Livory A. 2003. Les fourmis de la Manche. L'Argiope 39: 25-49.
  • Lomnicki J. 1925. Plagiolepis vindobonensis n. sp. (Hym. Formicidae). Pol. Pismo Entomol. 4: 77-79.
  • Mei M. 1995. Arthropoda di Lampedusa, Linosa e Pantelleria (Canale di Sicilia, Mar Mediterraneo). Hymenoptera Formicidae (con diagnosi di due nuove specie). Il Naturalista Siciliano (4)19(suppl.): 753-772.
  • Menozzi C. 1928. Note sulla mirmecofauna paleartica. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici. 21: 126-129.
  • Meyer M., C. Braunert, R. Gerend, and I. Schrankel. 2003. Résultats de l'excursion annuelle du groupe de travail entomologique de la Société des naturalistes luxembourgeois. Bull. Soc. Nat. luxemb. 103: 103-120.
  • Mokrousov M. V., and V.A. Zryanin. 2015. Materials on the early spring wasps and ants fauna of Uzbekistan (Hymenoptera: Vespomorpha: Chrysidoidea, Scolioidea, Pompiloidea, Vespoidea, Apoidea [Spheciformes], Formicoidea). Entomological research Russia and its neighboring regions 5: 36–48.
  • Mortazavi Z. S., H. Sadeghi, N. Aktac, L. Depa, L. Fekrat. 2015. Ants (Hymenoptera: Formicidae) and their aphid partners (Homoptera: Aphididae) in Mashhad region, Razavi Khorasan Province, with new records of aphids and ant species for fauna of Iran. Halteres 6:4-12.
  • Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
  • Oussalah N., F. Marniche, X. Espadaler, and M. Biche. 2019. Exotic ants from the Maghreb (Hymenoptera, Formicidae) with first report of the Hairy Alien Ant Nylanderia jaegerskioeldi (Mayr) in Algeria. Arxius de Miscel·lania Zoologica 17: 45–58.
  • Paik W.H. 1984. A checklist of Formicidae (Hymenoptera) of Korea. Korean J. Plant Prot. 23(3): 193-195.
  • Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
  • Pashaei Rad S., B. Taylor, R. Torabi, E. Aram, G. Abolfathi, R. Afshari, F. Borjali, M. Ghatei, F. Hediary, F. Jazini, V. Heidary Kiah, Z. Mahmoudi, F. Safariyan, and M. Seiri. 2018. Further records of ants (Hymenoptera: Formicidae) from Iran. Zoology in the Middle East 64(2): 145-159.
  • Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae,Hymenoptera) of Vojvodina (Serbia). Arch. Biol. Sci., Belgrade, 54, (3–4)27P–28P.
  • Ran H., and S. Y. Zhou. 2013. Checklist of Chinese Ants: Formicomorph Subfamilies ( Hymenoptera: Formicidae) (III). Journal of Guangxi Normal University : Natural Science Edition 31(1): 104-111.
  • Ruzsky, M. 1923. Ants of Cheleken Island. Izv. Tomsk. Gos. Univ. 72(2nd p part: 1-6
  • Salata S., L. Borowiec, and A. G. Radchenko. 2018. Description of Plagiolepis perperamus, a New Species from East Mediterranean and Redescription of Plagiolepis pallescens Forel, 1889 (Hymenoptera: Formicidae). Annales Zoologici (Warsaw) 68(4): 809-824.
  • Salata S., L. Borowiec, and A.Trichas. 2018. Taxonomic Revision of the Cretan Fauna of the Genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with Notes on the Endemism of Ant Fauna of Crete. Annales Zoologici (Warsaw) 68(4): 769-808.
  • Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
  • Salata S., and L. Borowiec. 2018. A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae). ZooKeys 789: 139–159.
  • Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
  • Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
  • Santschi F. 1926. Travaux scientifiques de l'Armée d'Orient (1916-1918). Fourmis. Bulletin du Muséum National d'Histoire Naturelle 32: 286-293.
  • Santschi F. 1928. Nouvelles fourmis de Chine et du Turkestan Russe. Bulletin et Annales de la Société Entomologique de Belgique. 68: 31-46.
  • Santschi, F. 1911. Formicides de diverses provenances. Annales de la Société Entomologique de Belgique 55: 278-287
  • Santschi, F. "Formicides de diverses provenances." Annales de la Société Entomologique de Belgique 55 (1911): 278-287.
  • Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
  • Schultz, R., A. G. Radchenko, and B. Seifert. "A critical checklist of the ants of Kyrgyzstan (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 201-207.
  • Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
  • Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
  • Shiran E., M. S. Mossadegh, and M. Esfandiari. 2012. Mutualistic ants (Hymenoptera: Formicidae) associated with aphids in central and southwestern parts of Iran. J. Crop Prot. 2013, 2 (1): 1-12.
  • Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
  • Stukalyuk S. V. 2013. Daily activity in multi-species ant (Hymenoptera, Formicidae) assemblages in the mountain Crimea. Proceedings of the Russian Entomological Society. St. Petersburg 84 (2): 114–128.
  • Stukalyuk S. V. 2016. Crematogaster subdentata Mayr, 1877, a potentially invasive species of ant (Hymenoptera, Formicidae) new to the fauna of Crimea. Entomological Review 95(8)
  • Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
  • Ticha, K. 2005. Inventarizacní pruzkum mravencu (Hymenoptera: Formicidae) NPP Svarec. 2005. Acta Rerum Naturalium 1:127-130.
  • Tohme G., and H. Tohme. 2014. Nouvelles liste des especes de fourmis du Liban (Hymenoptera, Formicoidea). Lebanese Science Journal 15(1): 133-141.
  • Vagalinski B., and A. Lapeva-Gjonova. 2012. The ants (Hymenoptera, Formicidae) of Plana Mountain (Bulgaria). Historia naturalis bulgarica 20: 87-101.
  • Vassilev I. 1987. The ants (Formicidae, Hymenoptera) from the valley of the river Racene. Annuaire de l'Universite de Sofia Kliment Ohridski Faceulte de Biologie 78: 74-79.
  • Vonshak M., and A. Ionescu-Hirsch. 2009. A checklist of the ants of Israel (Hymenoptera: Formicidae). Israel Journal of Entomology 39: 33-55.
  • Wiezik M., A. Wiezikova, and M. Svitok. 2011. Vegetation structure, ecological stability, and low-disturbance regime of abandoned dry grasslands support specific ant assemblages in Central Slovakia. Tuexenia 31: 301–315.
  • Yusupov Z. M. 2009. Ants (Hymenoptera, Formicidae) in the arid hollow of the Cherek-Balkarskiy River. Materials of the 13th All-Russian Myrmecological Symposium, N. Novgorod: 123-125.
  • Zryanin V. A., and T. A. Zryanina. 2007. New data on the ant fauna Hymenoptera, Formicidae in the middle Volga River Basin. Uspekhi Sovremennoi Biologii 127(2): 226-240.