Formicoxenus

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Formicoxenus
Formicoxenus nitidulus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Myrmicinae
Tribe: Crematogastrini
Alliance: Formicoxenus genus group
Genus: Formicoxenus
Mayr, 1855
Type species
Myrmica nitidula, now Formicoxenus nitidulus
Diversity
8 species
(Species Checklist, Species by Country)

Formicoxenus nitidulus casent0173159 profile 1.jpg

Formicoxenus nitidulus

Formicoxenus nitidulus casent0173159 dorsal 1.jpg

Specimen Label

Synonyms

Identification

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Keys including this Genus

 

Keys to Species in this Genus

Distribution

Distribution and Richness based on AntMaps

Species by Region

Number of species within biogeographic regions, along with the total number of species for each region.

Afrotropical Region Australasian Region Indo-Australian Region Malagasy Region Nearctic Region Neotropical Region Oriental Region Palaearctic Region
Species 0 0 0 0 5 0 0 3
Total Species 2840 1735 3042 932 835 4378 1740 2862

Biology

Association with Other Organisms

All Associate Records for Genus

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Taxon Relationship Associate Type Associate Taxon Associate Relationship Locality Source Notes
Formicoxenus chamberlini xenobiont ant Manica invidia host Wheeler (1904)
Formicoxenus diversipilosus xenobiont ant Formica obscuripes host
Formicoxenus hirticornis xenobiont ant Formica ciliata host
Formicoxenus hirticornis xenobiont ant Formica integroides host needs confirmation
Formicoxenus hirticornis xenobiont ant Formica obscuripes host
Formicoxenus hirticornis xenobiont ant Formica ravida host
Formicoxenus nitidulus xenobiont ant Formica aquilonia host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica exsecta host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica lugubris host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica pisarskii host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica polyctena host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica pratensis host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica rufa host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica truncorum host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Formica uralensis host Holldobler & Wilson 1990; Busch 2001; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Polyergus rufescens host Wilson 1971; Martin et al. 2007
Formicoxenus nitidulus xenobiont ant Tetramorium caespitum host Wilson 1971; Martin et al. 2007
Formicoxenus provancheri xenobiont ant Myrmica fracticornis host
Formicoxenus provancheri xenobiont ant Myrmica incompleta host
Formicoxenus quebecensis guest ant Myrmica alaskensis host
Formicoxenus sibiricus xenobiont ant Formica clara xenobiont Seifert, 2023
Formicoxenus sibiricus xenobiont ant Formica pisarskii xenobiont Seifert, 2023 misidentified as ''Formica pressilabris''

Life History Traits

  • Mean colony size: "a few" to ~100 (Greer et al., 2021)
  • Compound colony type: xenobiosis (Greer et al., 2021)
  • Nest site: hypogaeic (Greer et al., 2021)
  • Diet class: omnivore (Greer et al., 2021)
  • Foraging stratum: subterranean/leaf litter (Greer et al., 2021)

Castes

Morphology

Worker Morphology

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• Antennal segment count: 11 • Antennal club: 3 • Palp formula: 5,3; 4,3 • Total dental count: 5-6 • Spur formula: 0,0 • Eyes: 11-100 ommatidia • Pronotal Spines: absent • Mesonotal Spines: absent • Propodeal Spines: dentiform • Petiolar Spines: absent • Caste: none or weak • Sting: present • Metaplural Gland: present • Cocoon: absent

Male Morphology

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 • Antennal segment count 12 • Antennal club 0 in alates; gradual in ergatoids • Palp formula 5,3; 4,3 • Total dental count 0-4 • Spur formula 0, 0 • Caste alates/intermediates/ergatoids

Karyotype

All Karyotype Records for Genus

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Taxon Haploid Diploid Karyotype Locality Source Notes
Formicoxenus chamberlini 28 USA Buschinger & Francoeur, 1983; Francoeur et al., 1985; Fischer, 1987 as ''Symmyrmica chamberlini''
Formicoxenus nitidulus 15 France Buschinger et al., 1980; Francoeur et al., 1985; Fischer, 1987
Formicoxenus provancheri 11 22 Canada Buschinger et al., 1980; Fischer, 1987 as ''Leptothorax provancheri''
Formicoxenus quebecensis 28 Canada Francoeur et al., 1985; Fischer, 1987

Phylogeny

Myrmicinae
Myrmicini
Pogonomyrmecini
Stenammini
Solenopsidini
Attini

Ochetomyrmex  (2 species, 0 fossil species)

Tranopelta  (2 species, 0 fossil species)

Diaphoromyrma  (1 species, 0 fossil species)

Lachnomyrmex  (16 species, 0 fossil species)

Blepharidatta  (4 species, 0 fossil species)

Allomerus  (8 species, 0 fossil species)

Wasmannia  (11 species, 0 fossil species)

Pheidole  (1,294 species, 7 fossil species)

Cephalotes  (123 species, 16 fossil species)

Procryptocerus  (44 species, 0 fossil species)

Strumigenys  (879 species, 4 fossil species)

Phalacromyrmex  (1 species, 0 fossil species)

Pilotrochus  (1 species, 0 fossil species)

Protalaridris  (7 species, 0 fossil species)

Rhopalothrix  (19 species, 0 fossil species)

Basiceros  (9 species, 0 fossil species)

Octostruma  (35 species, 0 fossil species)

Eurhopalothrix  (54 species, 0 fossil species)

Talaridris  (1 species, 0 fossil species)

Acanthognathus  (7 species, 1 fossil species)

Daceton  (2 species, 0 fossil species)

Lenomyrmex  (7 species, 0 fossil species)

Microdaceton  (4 species, 0 fossil species)

Orectognathus  (29 species, 0 fossil species)

Colobostruma  (16 species, 0 fossil species)

Epopostruma  (20 species, 0 fossil species)

Mesostruma  (9 species, 0 fossil species)

Paleoattina

Apterostigma  (44 species, 2 fossil species)

Mycocepurus  (6 species, 0 fossil species)

Myrmicocrypta  (31 species, 0 fossil species)

Neoattina

Cyatta  (1 species, 0 fossil species)

Kalathomyrmex  (1 species, 0 fossil species)

Mycetarotes  (4 species, 0 fossil species)

Mycetosoritis  (2 species, 0 fossil species)

some Cyphomyrmex  (23 species, 2 fossil species)

some Cyphomyrmex

Paramycetophylax  (1 species, 0 fossil species)

Mycetophylax  (21 species, 0 fossil species)

Mycetagroicus  (4 species, 0 fossil species)

Mycetomoellerius  (31 species, 1 fossil species)

Sericomyrmex  (11 species, 0 fossil species)

Xerolitor  (1 species, 0 fossil species)

Paratrachymyrmex  (9 species, 0 fossil species)

Trachymyrmex  (9 species, 0 fossil species)

Amoimyrmex  (3 species, 0 fossil species)

Atta  (20 species, 1 fossil species)

some Acromyrmex  (56 species, 0 fossil species)

some Acromyrmex

Pseudoatta  (2 species, 0 fossil species)

Crematogastrini

Rostromyrmex  (1 species, 6 fossil species)

Cardiocondyla  (90 species, 0 fossil species)

Ocymyrmex  (34 species, 0 fossil species)

Nesomyrmex  (84 species, 2 fossil species)

Xenomyrmex  (5 species, 0 fossil species)

Terataner  (14 species, 0 fossil species)

Atopomyrmex  (3 species, 0 fossil species)

Cataulacus  (65 species, 3 fossil species)

Carebara  (249 species, 9 fossil species)

Diplomorium  (1 species, 0 fossil species)

Melissotarsus  (4 species, 1 fossil species)

Rhopalomastix  (14 species, 0 fossil species)

Calyptomyrmex  (38 species, 0 fossil species)

Strongylognathus  (27 species, 0 fossil species), Tetramorium  (598 species, 2 fossil species)

Cyphoidris  (4 species, 0 fossil species)

Dicroaspis  (2 species, 0 fossil species)

Aretidris  (2 species, 0 fossil species)

Vollenhovia  (83 species, 3 fossil species)

Dacetinops  (7 species, 0 fossil species)

Indomyrma  (2 species, 0 fossil species)

Crematogaster  (783 species, 3 fossil species)

Meranoplus  (91 species, 0 fossil species)

Lophomyrmex  (13 species, 0 fossil species)

Adlerzia  (1 species, 0 fossil species)

Recurvidris  (12 species, 0 fossil species)

Stereomyrmex  (3 species, 0 fossil species)

Trichomyrmex  (29 species, 0 fossil species)

Eutetramorium  (3 species, 0 fossil species)

Royidris  (15 species, 0 fossil species)

Malagidris  (6 species, 0 fossil species)

Vitsika  (16 species, 0 fossil species)

Huberia  (2 species, 0 fossil species)

Podomyrma  (62 species, 1 fossil species)

Liomyrmex  (1 species, 0 fossil species)

Metapone  (31 species, 0 fossil species)

Kartidris  (6 species, 0 fossil species)

Mayriella  (9 species, 0 fossil species)

Tetheamyrma  (2 species, 0 fossil species)

Dacatria  (1 species, 0 fossil species)

Proatta  (1 species, 0 fossil species)

Dilobocondyla  (22 species, 0 fossil species)

Secostruma  (1 species, 0 fossil species)

Acanthomyrmex  (19 species, 0 fossil species)

Myrmecina  (106 species, 0 fossil species)

Perissomyrmex  (6 species, 0 fossil species)

Pristomyrmex  (61 species, 3 fossil species)

some Lordomyrma  (36 species, 0 fossil species)

Propodilobus  (1 species, 0 fossil species)

Lasiomyrma  (4 species, 0 fossil species)

some Lordomyrma

Ancyridris  (2 species, 0 fossil species)

some Lordomyrma

Paratopula  (12 species, 0 fossil species)

Poecilomyrma  (2 species, 0 fossil species)

Romblonella  (10 species, 0 fossil species)

Rotastruma  (3 species, 0 fossil species)

Gauromyrmex  (3 species, 0 fossil species)

Vombisidris  (19 species, 0 fossil species)

Temnothorax  (504 species, 7 fossil species)

Harpagoxenus  (4 species, 0 fossil species)

Formicoxenus  (8 species, 0 fossil species)

Leptothorax  (20 species, 0 fossil species)

See Phylogeny of Myrmicinae for details.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • FORMICOXENUS [Myrmicinae: Formicoxenini]
    • Formicoxenus Mayr, 1855: 413. Type-species: Myrmica nitidula, by monotypy.
    • Formicoxenus junior synonym of Stenamma: Mayr, 1863: 422; Roger, 1863b: 25.
    • Formicoxenus revived status as genus: André, 1882c: 273.
    • Formicoxenus senior synonym of Symmyrmica: Francoeur, Loiselle & Buschinger, 1985: 347.
  • SYMMYRMICA [junior synonym of Formicoxenus]
    • Symmyrmica Wheeler, W.M. 1904a: 3. Type-species: Symmyrmica chamberlini, by monotypy.
    • Symmyrmica junior synonym of Formicoxenus: Francoeur, Loiselle & Buschinger, 1985: 347.

References