Temnothorax parvulus
Temnothorax parvulus | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | Myrmicinae |
Tribe: | Crematogastrini |
Genus: | Temnothorax |
Species group: | nylanderi |
Species complex: | parvulus |
Species: | T. parvulus |
Binomial name | |
Temnothorax parvulus (Schenck, 1852) |
Photo Gallery
Identification
Csösz et al. (2015) - A member of the parvulus species-complex. Temnothorax parvulus can be separated easily from members of other species-complexes by its homogenously areolate microsculpture on the head dorsum. The head sculpture of other species complexes may vary from smooth to coarsely rugulo-reticulate but is never homogenously areolate. In exceptional cases, long-spined species of the Temnothorax lichtensteini complex might exhibit a homogenous areolate sculpture on an extended area of the head dorsum, but T. parvulus is distinguished from T. lichtensteini and Temnothorax laconicus by its more erect propodeal spines (38–42° vs. ca. 20–25°). Temnothorax parvulus can also be safely separated from weakly sculptured, lightly colored Temnothorax tergestinus samples using slightly overlapping NOH/CS, SPTI/CS and SPWI/CS ratios (see S4 Table).
Temnothorax parvulus shares most its surface sculpture characters with other species belonging to this complex, Temnothorax ariadnae and Temnothorax helenae. Spine length ratio (SPST/CL) helps distinguish Temnothorax parvulus from T. ariadnae and T. helenae.
Non-overlapping values of SPST/CL for nest sample means of T. parvulus (mean: 0.282 [min: 0.259, max: 0.303]); and T. ariadnae (mean: 0.216 [min: 0199, max: 0.227]) provide a prefect means of separation. As 5% of nest samples of T. parvulus and T. helenae (mean: 0.234 [min: 0.189, max: 0.274] overlap in this ratio, the safest determination is provided by a discriminant (D5 = 0.0393_CL -0.0616_SPST +0.0892_PEW -0.0580_PPW -0.0338_SPWI -8.7356) formula, which separates 97.1% of single individuals and all nest samples of T. parvulus and T. helenae.
Keys including this Species
Distribution
Western Europe to the Black see coast and Turkey and from Italy and the Balkans to Central Europe.
Latitudinal Distribution Pattern
Latitudinal Range: 52.91666667° to 36.105°.
North Temperate |
North Subtropical |
Tropical | South Subtropical |
South Temperate |
- Source: AntMaps
Distribution based on Regional Taxon Lists
Palaearctic Region: Andorra, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Czechia, Georgia, Germany (type locality), Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Montenegro, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Türkiye, Turkmenistan.
Distribution based on AntMaps
Distribution based on AntWeb specimens
Check data from AntWeb
Countries Occupied
Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species. |
Estimated Abundance
Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species. |
Biology
Association with Other Organisms
- Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.
Cestoda
- This species is a host for the cestode Dilepididae (type 1(?)) (a parasite) in Germany (Buschinger, 1973; Laciny, 2021).
Flight Period
X | X | ||||||||||
Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
Source: antkeeping.info.
- Check details at Worldwide Ant Nuptial Flights Data, AntNupTracker and AntKeeping.
- Explore: Show all Flight Month data or Search these data. See also a list of all data tables or learn how data is managed.
Life History Traits
- Queen number: monogynous (Frumhoff & Ward, 1992)
Castes
Nomenclature
The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.
- parvulus. Myrmica parvula Schenck, 1852: 103 (w.) GERMANY.
- [Also described as new by Schenck, 1853: 187.]
- Mayr, 1855: 448 (q.); Novák, in Kratochvíl, Novák & Šnoflák, 1944: 116 (m.).
- Combination in Leptothorax: Mayr, 1855: 448.
- Combination in Temnothorax: Bolton, 2003: 271.
- Junior synonym of cingulata: Nylander, 1856b: 93.
- Junior synonym of nylanderi: Smith, F. 1858b: 8; Roger, 1863b: 26.
- Subspecies of tuberum: André, 1883a: 300.
- Subspecies of nylanderi: Forel, 1874: 86; Dalla Torre, 1893: 126; Ruzsky, 1902d: 23; Stitz, 1914: 64; Forel, 1915d: 22 (in key); Emery, 1916b: 175; Menozzi, 1918: 85; Bondroit, 1918: 123; Menozzi, 1918: 85; Müller, 1923b: 95; Emery, 1924d: 255; Novák & Sadil, 1941: 91 (in key).
- Status as species: Mayr, 1855: 448; Bernard, 1956a: 161; Bernard, 1959: 349; Bernard, 1967: 203 (redescription); Baroni Urbani, 1971c: 119; van Boven, 1977: 105; Arnol'di & Dlussky, 1978: 541 (in key); Agosti & Collingwood, 1987b: 274 (in key); Dlussky, Soyunov & Zabelin, 1990: 191; Casevitz-Weulersse, 1990c: 418 (in key); Atanassov & Dlussky, 1992: 140; Arakelian, 1994: 56; Radchenko, 1994d: 153 (in key); Bolton, 1995b: 242; Radchenko, 1995d: 4; Czechowski, et al. 2002: 52; Seifert, 2007: 241; Boer, 2010: 52; Czechowski, et al. 2012: 150; Csösz, Heinze & Mikó, 2015: 47 (redescription).
Type Material
Csösz et al. (2015) - Lectotype: “L. parvulus # Sch” and “Lectotype Leptothorax parvulus Schenck, 1852 det. A. Schulz &M. Verhaagh 1999” (1# / SMF), [GER:parvulus-TYPE]. Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Description
Worker
Csösz et al. (2015) - Body color: brown; yellow. Body color pattern: mesosoma, antenna and legs, waist and anterior region of 1st gastral tergite lighter than head dorsum and posterior region of gaster. Antenna color pattern: clava concolorous funicle. Absolute cephalic size: 488–586 μm (mean = 550, n = 29). Cephalic length vs. Maximum width of head capsule (CL/CWb): 1.147–1.214 (mean = 1.184). Postocular distance vs. cephalic length (PoOc/CL): 0.392–0.413 (mean = 0.405). Postocular sides of cranium contour frontal view orientation: converging posteriorly. Postocular sides of cranium contour frontal view shape: strongly convex. Vertex contour line in frontal view shape: straight. Vertex sculpture: main sculpture absent, ground sculpture areolate; main sculpture dispersed forked costate sculpture, ground sculpture areolate. Genae contour from anterior view orientation: converging. Gena contour line in frontal view shape: convex. Gena sculpture: rugoso-reticulate with areolate ground sculpture. Median region of antennal rim vs. frontal carina in frontal view structure: not fully overlapped by frontal carina. Concentric carinae laterally surrounding antennal foramen count: present. Eye length vs. absolute cephalic size (EL/CS): 0.237–0.262 (mean = 0.250). Frontal carina distance vs. absolute cephalic size (FRS/CS): 0.353–0.376 (mean = 0.361). Longitudinal carinae on median region of frons count: absent. Smooth median region on frons count: present. Antennomere count: 12. Scape length vs. absolute cephalic size (SL/CS): 0.763–0.796 (mean = 0.778). Facial area of the scape absolute setal angle: 0–15°. External area of the scape absolute setal angle: 30°. Ground sculpture of submedian area of clypeus: smooth. Median carina of clypeus count: present. Lateral carinae of clypeus count: present. Median anatomical line of propodeal spine angle value to Weber length in lateral view: 38–42°. Spine length vs. absolute cephalic size (SPST/CS): 0.278–0.331 (mean = 0.306). Minimum spine distance vs. absolute cephalic size (SPBA/CS): 0.273–0.312 (mean = 0.292). Maximum spine distance vs. absolute cephalic size (SPWI/CS): 0.353–0.415 (mean = 0.384). Apical spine distance vs. absolute cephalic size (SPTI/CS): 0.332–0.395 (mean = 0.364). Maximum mesosoma width vs. absolute cephalic size (MW/CS): 0.599–0.636 (mean = 0.618). Metanotal depression count: present. Metanotal depression shape: shallow. Dorsal region of mesosoma sculpture: rugulose with areolate ground sculpture. Lateral region of pronotum sculpture: areolate ground sculpture, main sculpture forked costate. Mesopleuron sculpture: areolate ground sculpture superimposed by dispersed rugulae. Metapleuron sculpture: areolate ground sculpture superimposed by dispersed rugulae. Frontal profile of petiolar node contour line in lateral view shape: concave. Dorsal profile of petiolar node contour line angle value to frontal profile of petiole contour line in lateral view: 105–110°. Anterodorsal rim of petiole count: absent medially.
Dorsal profile of petiolar node contour line in lateral view shape: slightly convex. Dorsal region of petiole sculpture: ground sculpture areolate, main sculpture dispersed rugose; ground sculpture areolate, main sculpture absent. Dorso-caudal petiolar profile contour line in lateral view shape: straight; concave. Dorsal region of postpetiole sculpture: ground sculpture areolate, main sculpture dispersed rugose; ground sculpture areolate, main sculpture absent.
Karyotype
- See additional details at the Ant Chromosome Database.
- Explore: Show all Karyotype data or Search these data. See also a list of all data tables or learn how data is managed.
- n = 14 (France, Italy, Switzerland) (Fischer, 1987) (as Leptothorax parvulus).
References
- Csösz, S., Heinze, J. and Mikó, I. 2015. Taxonomic synopsis of the Ponto-Mediterranean ants of Temnothorax nylanderi species-group. PLoS ONE. 10(11):62pp. e0140000 (doi:10.1371/journal.pone.0140000).
- Agosti, D.; Collingwood, C. A. 1987a. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitt. Schweiz. Entomol. Ges. 60: 51-62 (page 56, Revived status as species)
- Atanassov, N.; Dlussky, G. M. 1992. Fauna of Bulgaria. Hymenoptera, Formicidae. Fauna Bûlg. 22: 1-310 (page 140, Revived status as species)
- Baroni Urbani, C. 1971c. Catalogo delle specie di Formicidae d'Italia (Studi sulla mirmecofauna d'Italia X). Memorie della Societa Entomologica Italiana 50: 5-287 (page 120, Revived status as species)
- Bernadou, A., Fourcassié, V., Espadaler, X. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. ZooKeys 277, 13–23 (doi:10.3897/zookeys.277.4684).
- Bernard, F. 1956a. Révision des Leptothorax (Hyménoptères Formicidae) d'Europe occidentale, basée sur la biométrie et les genitalia mâles. Bull. Soc. Zool. Fr. 81: 151-165 (page 161, Revived status as species)
- Bernard, F. 1967a [1968]. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp. (page 203, Revived status as species)
- Bolton, B. 2003. Synopsis and Classification of Formicidae. Mem. Am. Entomol. Inst. 71: 370pp (page 271, Combination in Temnothorax)
- Borowiec, L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
- Borowiec, M.L., Borowiec, L. 2013. New data on the occurrence of ants (Hymenoptera: Formicidae) in Lower Silesia and other regions of Poland. Wiadomosci Entomologiczne 32: 49-57.
- Boven, J. K. A. van. 1977. De mierenfauna van België (Hymenoptera: Formicidae). Acta Zool. Pathol. Antverp. 67: 1-191 (page 105, Revived status as species)
- Bračko, G. 2019. New data on the ant fauna (Hymenoptera: Formicidae) of Azerbaijan. Caucasian Entomological Bulletin 15, 165–175 (doi:10.23885/181433262019151-165175).
- Bracko, G., Wagner, H.C., Schulz, A., Gioahin, E., Maticic, J., Trantnik, A. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10: 10-24.
- Bulter, I. 2020. Hybridization in ants. Ph.D. thesis, Rockefeller University.
- Buschinger, A. 1973. Ameisen des Tribus Leptothoracini (Hym. Formicidae) als Zwischenwirte von Cestoden. Zoologischer Anzeiger 191: 369-380.
- Castracani, C., Spotti, F.A., Schifani, E., Giannetti, D., Ghizzoni, M., Grasso, D.A., Mori, A. 2020. Public engagement provides first insights on Po Plain ant communities and reveals the ubiquity of the cryptic species Tetramorium immigrans (Hymenoptera, Formicidae). Insects 11, 678. (doi:10.3390/insects11100678).
- Csősz, S., Báthori, F., Gallé, L., Lőrinczi, G., Maák, I., Tartally, A., Kovács, É., Somogyi, A.Á., Markó, B. 2021. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: Survey of ant species with an annotated synonymic inventory. Insects 16;12(1):78 (doi:10.3390/insects12010078).
- Csosz, S., Marko, B., Galle, L. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
- Czechowski, W., Radchenko, A., Czechowska, W. 2002. The ants (Hymenoptera, Formicidae) of Poland. MIZ PAS Warsaw.
- Dekoninck, W., Ignace, D., Vankerkhoven, F., Wegnez, P. 2012. Verspreidingsatlas van de mieren van België. Bulletin de la Société royale belge d’Entomologie 148: 95-186.
- Emery, C. 1924f [1922]. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [concl.]. Genera Insectorum 174C: 207-397 (page 255, Variety/subspecies of mylanderi)
- Forel, A. 1874. Les fourmis de la Suisse. Systématique, notices anatomiques et physiologiques, architecture, distribution géographique, nouvelles expériences et observations de moeurs. Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 26: 1-452 (page 86, Revived from synonymy as variety of nylanderi)
- Forel, A. 1915d. Fauna insectorum helvetiae. Hymenoptera. Formicidae. Die Ameisen der Schweiz. Mitt. Schweiz. Entomol. Ges. 12(B Beilage: 1-77 (page 22, Variety/subspecies of mylanderi)
- Gonzalez, J.A. 2021. Description of Temnothorax estel sp. nov. (Hymenoptera: Formicidae), with a review of the Iberian species of the sordidulus species-complex. Zootaxa 5005, 145–160 (doi:10.11646/zootaxa.5005.2.2).
- Karaman, C., Kiran, K. 2022. Additional records of parasitic Camponotus Mayr (Hymenoptera: Formicidae) species from Turkey with queen description of Camponotus ruseni Karaman, 2012. Zoology in the Middle East 68(2), 156–164 (doi:10.1080/09397140.2022.2051918).
- Kiran, K., Karaman, C. 2020. Additions to the ant fauna of Turkey (Hymenoptera, Formicidae). Zoosystema 42(18), 285-329 (doi:10.5252/zoosystema2020v42a18).
- Kiran, K., Karaman, C., Lapeva-Gjonova, A., Aksoy, V. 2017. Two new species of the "ultimate" parasitic ant genus Teleutomyrmex KUTTER, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic. Myrmecological News 25, 145-155.
- Lapeva-Gjonova, A., Antonova, V. 2022. An updated checklist of ants (Hymenoptera, Formicidae) of Bulgaria, after 130 years of research. Biodiversity Data Journal 10, e95599 (doi:10.3897/bdj.10.e95599).
- Lapeva-Gjonova, A., Antonova, V., Ljubomirov, T. 2021. Ants (Hymenoptera, Formicidae) of Sarnena Sredna Gora Mountains (Bulgaria). Fauna of Sarnena Sredna Gora Mts, Part 2 ZooNotes, Supplement 10: 18-27.
- Lapeva-Gjonova, A., Kiran, K. 2012. Ant fauna (Hymenoptera, Formicidae) of Strandzha (Istranca) Mountain and adjacent Black Sea coast. North-Western Journal of Zoology 8(1), 72-84.
- Mayr, G. 1855. Formicina austriaca. Beschreibung der bisher im österreichischen Kaiserstaate aufgefundenen Ameisen, nebst Hinzufügung jener in Deutschland, in der Schweiz und in Italien vorkommenden Arten. Verh. Zool.-Bot. Ver. Wien 5: 273-478 (page 448, queen described, Combination in Leptothorax)
- Menozzi, C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti Soc. Nat. Mat. Modena (5) 4: 81-88 (page 85, Variety/subspecies of mylanderi)
- Müller, G. 1923b. Le formiche della Venezia Guilia e della Dalmazia. Boll. Soc. Adriat. Sci. Nat. Trieste 28: 11-180 (page 95, Variety/subspecies of mylanderi)
- Novák, V. 1944c. K taxonomii mravencu rodu Bothriomyrmex a Leptothorax. Pp. 103-132 in: Kratochvíl, J., Novák, V., Snoflák, J. Mohelno. Soubor práci venoványch studiu vyznamne památky prírodní. 5. Hymenoptera - Aculeata. Formicidae - Apidae - Vespidae. (page 116, male described)
- Nylander, W. 1856b. Synopsis des Formicides de France et d'Algérie. Ann. Sci. Nat. Zool. (4) 5: 51-109 (page 93, Junior synonym of cingulata)
- Purkart, A., Kollár, J., Goffová, K. 2019. Fauna of ants (Hymenoptera: Formicidae) of selected sand habitats in Podunajsko Region, Slovakia. Naturae Tutela 23(1): 101-111.
- Ruzsky, M. 1902d. Material on the ant fauna of the Caucasus and the Crimea. Protok. Obshch. Estestvoispyt. Imp. Kazan. Univ. 206(su suppl: 1-33 (page 23, Variety/subspecies of mylanderi)
- Salata, S., Borowiec, L. 2014. New localities of some rare ant species (Hymenoptera: Formicidae). Wiadomości Entomologiczne 33, 77-79.
- Salata, S., Borowiec, L. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131 (doi:10.3897/zookeys.877.36320).
- Schenck, C. F. 1852. Beschreibung nassauischer Ameisenarten. Jahrb. Ver. Naturkd. Herzogthum Nassau Wiesb. 8: 1-149 (page 103, worker described)
- Schifani, E., Nalini, E., Gentile, V., Alamanni, F., Ancona, C., Caria, M., Cillo, D., Bazzato, E. 2021. Ants of Sardinia: An updated checklist based on new faunistic, morphological and biogeographical notes. Redia 104, 21–35 (doi:10.19263/redia-104.21.03).
- Schultner, E., Pulliainen, U. 2020. Brood recognition and discrimination in ants. Insectes Sociaux 67, 11–34 (doi:10.1007/s00040-019-00747-3).
- Smith, F. 1858b. Catalogue of hymenopterous insects in the collection of the British Museum. Part VI. Formicidae. London: British Museum, 216 pp. (page 8, Junior synonym of nylanderi)
- Snegovaya, N., Shigayev, C. 2021. A checklist of the ants (Insecta, Formicidae) of Azerbaijan Republic. Iranian Journal of Animal Biosystematics 17(2): 179-207 (doi:10.22067/ijab.2022.67343.1000).
- Wegnez, P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie153, 46–49.
References based on Global Ant Biodiversity Informatics
- AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
- Antarea (at www.antarea.fr on June 11th 2017)
- Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
- Asociacion Iberica de Mirmecologia. 2012. List of species collected during the Taxomara Tres Cantos 2012. Iberomyrmex 4: 17-18.
- Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
- Baugnée J. Y. 2004. Observations récentes de la fourmi Leptothorax parvulus (Schenck, 1852) en Belgique (Hymenoptera Formicidae). Bulletin de la Société Royale Belge d'Entomologie 140: 101-103.
- Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
- Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:5971
- Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
- Bernard F. 1959. Les fourmis de l'île de Port-Cros. Contribution à l'écologie des anciennes forêts méditerranéennes. Vie Milieu 9: 340-360.
- Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
- Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
- Bernard F. 1976. Écologie des fourmis des grès d'Annot, comparées à celles de la Provence calcaire. Annales du Muséum d'Histoire Naturelle de Nice 3: 33-54.
- Bernard F. 1978. Fourmis et milieu dans le massif des Maures. Vie et Milieu. Série C. Biologie Terrestre 27: 83-118.
- Bernard, F. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
- Berville L., M. Renucci, and E. Provost. 2012. Mise en place de protocoles de contrôle de la fourmi dArgentine (Linepithema humile) sur les îles de Port-Cros et de Porquerolles (Var, France). Sci. Rep. Port-Cros natl. Park, 26: 91-108.
- Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
- Bezdecka P., and K. Bezdeckova. 2012. Updated list of the ants of the Czech Republic. Pp 7-12. Bezd??ka P. & Bezd??ková K. (eds) 2012: Blanok?ídlí v ?eských zemích a na Slovensku 8, Chaloupky, 1.-3. ?ervna 2012, sborník abstrakt? z konference. MVJ Jihlava, 37 pp.
- Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
- Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
- Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
- Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
- Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
- Borowiec M. L., and L. Borowiec. 2013. New data on the occurrence of ants (Hymenoptera: Formicidae) in Lower Silesia and other regions of Poland. Wiadomo?ci Entomologiczne 32: 49-57.
- Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
- Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
- Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
- Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
- Buschinger A. 1975. Die ameisenfauna des Bausenberges, der nordostlichen Eifel und Voreifel (Hym., Formicidae) mit einer quantitativen Auswertung von Fallenfangen. Beitrage Landespflege Rhid-Pfalz Beiheft 4: 251-273.
- Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
- Casevitz-Weulersse, J. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
- Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
- Colindre L. 2016. Les fourmis en Picardie: bilan des decouvertes et redecouvertes 2015 (Hymenoptera/ Formicidae). Entomologiste Picard 28: 8-10.
- Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
- Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
- Csősz S, Heinze J, and I. Mikó. 2015. Taxonomic synopsis of the Ponto-Mediterranean ants of Temnothorax nylanderi species-group. PLoS ONE 10(11): e0140000. doi:10.1371/journal.pone.0140000
- Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
- Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
- Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
- Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
- Diniz, M. A. "Estado actual do conhecimento dos himenópteros de Portugal." Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 259 (1959): 1-42.
- Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
- Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
- Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
- Espadaler X., F. Garcia, X. Roig, and R. Vila. 2013. Ants (Hymenoptera, Formicidae) from the Castell de Montesquiu park (Osona, north-east of the Iberian Peninsula). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 53: 223-227.
- Espadaler X., M. Boieiro, R. Azedo, C. Bernardes, D. Figueiredo, and A. R. M. Serrano. 2008. Additions to the ant fauna of Portugal (Hymenoptera, Formicidae). Boletin Sociedad Entomologica Aragonesa 42: 349-351.
- Espadaler, X., A. R. Azedo, C. Bernardes, D. Figueiredo, A. R. M. Serrano, and M. Boeiro. "Additions to the Ant fauna of Portugal (Hymenoptera: Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 349-351.
- Foitzik S., and Heinze, J. 1999. Non-random size differences between sympatric species of the ant genus Leptothorax (Hymenoptera: Formicidae). Entomologia Generalis 24: 65-74.
- Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
- Forel, A. "Fourmis de Tunisie et de l'Algerie orientale recoltees et decrites par Auguste Forel." Annales de la Société Entomologique de Belgique 34 (1890): lxi-lxxvi.
- Francois J. 1958. Contribution a l'etude ecologique des Formicides (Insectes, Hymenopteres) de la region Dijonnaise. Travaux du laboratoire de Zoologie et de la Station Aquicole Grimaldi de la Faculte des Sciences de Dijon 25, 39 pages.
- GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
- Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
- Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
- Garcia F., X. Espadaler, and K. Gomez. 2009. First record of Amblyopone impressifrons (Emery, 1869) from the Iberian Peninsula and of Proceratium melinum (Roger, 1860) from Catalonia (Hymenoptera, Formicidae). Boletín Sociedad Entomológica Aragonesa 45: 357-360.
- Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
- Ghahari H., and C. A. Collingwood. 2013. A study on the ants (Hymenoptera: Vespoidea: Formicidae) from Western Iran. Acta Phytopathologica et Entomologica Hungarica 48 (1): 155164.
- Giacalone I., and M. Moretti. 2001. Contributo alla conoscenza della mirmecofauna (Hymenoptera: Formicidae) dei castagneti al Sud delle Alpi (ticino, Svizzera). Bollettino della Societa ticinese di Scienze naturali 89(1-2): 51-60.
- Glaser F. 2004. Distribution and conservation of ants (Hymenoptera, Formicidae) in the floodplain of the river Etsch / Adige (Southern Tyrol, Italy). Gredleriana 4: 203-246.
- Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
- Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
- Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
- Kiran K. C. Karaman, A. Lapeva-Gjonova, and V. Aksoy. 2017. Two new species of the "ultimate" parasitic ant genus Teleutomyrmex KUTTER, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic. Myrmecological News 25: 145-155.
- Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
- Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24 33
- Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
- Legakis Collection Database
- Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
- Marko B. 2008. Ants (Hymenoptera: Formicidae) of the Sand Dunes of Foieni Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
- Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
- Mei, M. "Nuovi reperti de formicidi per l'Italia centrale (Hymenoptera, Formicidae)." Bollettino dell'Associazione Romana di Entomologia 37 (1984): 49-58.
- Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
- Meyer M., C. Braunert, R. Gerend, and I. Schrankel. 2003. Résultats de l'excursion annuelle du groupe de travail entomologique de la Société des naturalistes luxembourgeois. Bull. Soc. Nat. luxemb. 103: 103-120.
- Moretti M. P. Duelli, and M. K. Obrist. 2006. Biodiversity and resilience of arthropod communities after fire disturbance in temperate forests. Oecologia 149: 312327.
- Ottonetti L., L. Tucci, and G. Santini. 2006. Recolonization Patterns of Ants in a Rehabilitated Lignite Mine in Central Italy: Potential for the Use of Mediterranean Ants as Indicators of Restoration Processes. Restoration Ecology 14(1): 6066.
- Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
- Petrov I. Z. 2004. A list of currently known ant species (Formicidae, Hymenoptera) of Serbia. Arch. Biol. Sci., Belgrade 56(3-4): 121-125.
- Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
- Plateaux L. 2000. Les fourmis de Lorraine. Bull. Soc. Ent. Lorr. 11-16.
- Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
- Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
- Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
- Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
- Salata S., P. Zurawlew, and J. K. Kowalczyk. 2018. New distribution records of several ant species (Hymenoptera: Formicidae) in Poland. Wiadomosci Entomologiczne 37(1): 46-53.
- Salata S., and L. Borowiec. 2014. New localities of some rare ant species (Hymenoptera: Formicidae). WIAD. ENTOMOL., 33 (1): 77-79.
- Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
- Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
- Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
- Schmitz H. 1950. Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit. Brotéria Ser. Cienc. Nat. 19: 12-16.
- Schmitz, H. "Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit." Brotéria, Ciências Naturais 19 (1950): 12-16.
- Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
- Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
- Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
- Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
- Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
- Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
- Tausan I. 2014. Temnothorax parvulus (Schenck, 1852)(Hymenoptera: Formicidae) in Romania. Brukenthal, Acta Musei 9: 511-514.
- Vogrin, V. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
- Wegnez P. 2014. Premières captures de Lasius distinguendus Emery, 1916 et de Temnothorax albipennis (Curtis, 1854) au Grand-Duché de Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société royale belge d’Entomologie 150 (2014) : 168-171.
- Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
- Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
- Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
- Pages using DynamicPageList3 parser function
- Photo Gallery
- North temperate
- Cestode Associate
- Host of Dilepididae (type 1(?))
- FlightMonth
- Karyotype
- Species
- Extant species
- Formicidae
- Myrmicinae
- Crematogastrini
- Temnothorax
- Temnothorax parvulus
- Myrmicinae species
- Crematogastrini species
- Temnothorax species
- Need Overview