Morocco

Morocco
Location
Region	Africa
Subregion	Northern Africa
BioRegion	Palaearctic Region
Diversity
Subfamilies	8
Genera	34
Endemic Genera	0
% Endemic Genera	0.0%
Species/Subspecies	192
Endemic Species	91
% Endemic Species	47.4%
Introduced Species	1

The following valid species and subspecies of ants are known to occur in Morocco. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Morocco ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Morocco Ants web page. Please advise if any corrections are warranted.

Aenictus

• Aenictus mauritanicus Santschi, 1910
• Aenictus vaucheri Emery, 1915

Anochetus

• Anochetus ghilianii Spinola, 1851

Aphaenogaster

• Aphaenogaster afra Santschi, 1933
• Aphaenogaster atlantis Santschi, 1929
• Aphaenogaster baronii Cagniant, 1988
• Aphaenogaster crocea André, 1881
• Aphaenogaster curiosa Santschi, 1933
• Aphaenogaster dejeani Cagniant, 1982
• Aphaenogaster depilis Santschi, 1911
• Aphaenogaster espadaleri Cagniant, 1984
• Aphaenogaster fallax Cagniant, 1992
• Aphaenogaster gemella (Roger, 1862)
• Aphaenogaster gemella marocana Forel, 1903
• Aphaenogaster gibbosa (Latreille, 1798)
• Aphaenogaster koniari Cagniant & Galkowski, 2013
• Aphaenogaster mauritanica Dalla Torre, 1893
• Aphaenogaster miniata Cagniant, 1990
• Aphaenogaster nadigi Santschi, 1923
• Aphaenogaster pallida (Nylander, 1849)
• Aphaenogaster praedo Emery, 1908
• Aphaenogaster praedo ellipsoida Santschi, 1933
• Aphaenogaster praenoda Santschi, 1933
• Aphaenogaster praenoda confinis Santschi, 1933
• Aphaenogaster rifensis Cagniant, 1994
• Aphaenogaster rupestris Forel, 1909
• Aphaenogaster sardoa Mayr, 1853
• Aphaenogaster senilis Mayr, 1853
• Aphaenogaster senilis disjuncta Santschi, 1933
• Aphaenogaster sicardi Cagniant, 1990
• Aphaenogaster splendida (Roger, 1859)
• Aphaenogaster testaceopilosa (Lucas, 1849)
• Aphaenogaster testaceopilosa cabylica Stitz, 1917
• Aphaenogaster theryi Santschi, 1923
• Aphaenogaster tinauti Cagniant, 1992
• Aphaenogaster torossiani Cagniant, 1988
• Aphaenogaster weulersseae Cagniant, 1989
• Aphaenogaster wilsoni Cagniant, 1988

Bajcaridris

• Bajcaridris theryi Santschi, 1936

Bothriomyrmex

• Bothriomyrmex meridionalis marocanus Santschi, 1923

Camponotus

• Camponotus atlantis maroccanus Emery, 1925
• Camponotus cruentatus Latreille, 1802
• Camponotus foreli tingitanus Santschi, 1921
• Camponotus hoelldobleri Cagniant, 1991
• Camponotus lateralis (Olivier, 1792)
• Camponotus magister Santschi, 1925
• Camponotus micans asniensis Santschi, 1939
• Camponotus obscuriventris Cagniant, 1991
• Camponotus pexus Santschi, 1929
• Camponotus pupillus Santschi, 1939
• Camponotus serotinus Cagniant, 1996
• Camponotus spissinodis Forel, 1909
• Camponotus vagus ifranensis Cagniant, 1987

Cardiocondyla

• Cardiocondyla batesii
• Cardiocondyla mauritanica Forel 1890
• Cardiocondyla nigra

Cataglyphis

• Cataglyphis bicolor pubens Santschi, 1929
• Cataglyphis cana Santschi, 1925
• Cataglyphis cubica Forel, 1903
• Cataglyphis espadaleri Cagniant, 2009
• Cataglyphis fossilis Cagniant, 2009
• Cataglyphis gaetula Santschi, 1929
• Cataglyphis marroui Cagniant, 2009
• Cataglyphis mauritanica occidentalis Cagniant, 2009
• Cataglyphis mauritanica tonsilis Santschi, 1936
• Cataglyphis otini Santschi, 1929
• Cataglyphis pilisquamis Santschi, 1929
• Cataglyphis theryi Santschi, 1921
• Cataglyphis vaucheri Emery, 1906
• Cataglyphis viatica (Fabricius, 1787)

Colobopsis

• Colobopsis imitans Schifani, Giannetti, Csősz, Castellucci, Luchetti, Castracani, Spotti, Mori & Grasso, 2021

Crematogaster

• Crematogaster auberti Emery, 1869
• Crematogaster auberti regilla Santschi, 1937
• Crematogaster laestrygon Emery, 1869
• Crematogaster laestrygon theryi Santschi, 1921
• Crematogaster laestrygon vivax Santschi, 1937
• Crematogaster sordidula marocana Santschi, 1921

Dorylus

• Dorylus fulvus juvenculus Shuckard, 1840

Formica

• Formica cunicularia Latreille, 1798
• Formica fusca maura Santschi, 1929

Goniomma

• Goniomma hispanicum André, 1883
• Goniomma kugleri Espadaler, 1986
• Goniomma otini Santschi, 1929

Hypoponera

• Hypoponera eduardi (Forel, 1894)
• Hypoponera punctatissima Roger, 1859

Lasius

• Lasius rabaudi Bondroit, 1917
• Lasius lasioides Emery, 1869
• Lasius mauretanicus Seifert, 2020
• Lasius myops Forel, 1894
• Lasius tibialis Santschi, 1936

Lepisiota

• Lepisiota frauenfeldi marocana Santschi, 1936

Leptanilla

• Leptanilla ortunoi López, Martínez & Barandica, 1994
• Leptanilla revelierii Emery, 1870
• Leptanilla vaucheri Emery, 1899

Linepithema

• Linepithema humile (introduced)

Messor

• Messor abdelazizi Santschi, 1921
• Messor aegyptiacus (Emery, 1878)
• Messor antennatus Emery, 1908
• Messor antennatus personatus Santschi, 1931
• Messor barbarus Linnaeus, 1767
• Messor barbarus gallaoides Santschi, 1929
• Messor boyeri Cagniant, 2006
• Messor brevispinosus Santschi, 1923
• Messor capitatus nigricans Santschi, 1929
• Messor caviceps (Forel, 1902)
• Messor erectus Espadaler, 1998
• Messor foreli Santschi, 1923
• Messor grandinidus Emery, 1912
• Messor hispanicus Santschi, 1919
• Messor hoggarensis Santschi, 1929
• Messor lobicornis Forel, 1894
• Messor marocanus Santschi, 1927
• Messor medioruber Santschi, 1910
• Messor minor (André, 1883)
• Messor picturatus Santschi, 1927
• Messor sanctus Emery, 1921
• Messor striatulus (Emery, 1891)
• Messor striativentris Emery, 1908
• Messor vaucheri Emery, 1908

Monomorium

• Monomorium salomonis bedui Santschi, 1936
• Monomorium salomonis obscuriceps Santschi, 1921
• Monomorium salomonis transversale Santschi, 1921
• Monomorium salomonis volubile Santschi, 1936

Myrmica

• Myrmica cagnianti Espadaler, 1996

Oxyopomyrmex

• Oxyopomyrmex saulcyi Emery, 1889

Plagiolepis

• Plagiolepis barbara Santschi, 1911
• Plagiolepis pygmaea
• Plagiolepis schmitzii Forel, 1895
• Plagiolepis schmitzii gaetula Santschi, 1936

Ponera

• Ponera coarctata Latreille, 1802
• Ponera testacea

Proceratium

• Proceratium algiricum Forel, 1899
• Proceratium numidicum Santschi, 1912

Solenopsis

• Solenopsis lusitanica Emery, 1915
• Solenopsis lusitanica gaetula Santschi, 1936

Stenamma

• Stenamma punctiventre Emery, 1908

Stigmatomma

• Stigmatomma denticulatum Roger, 1859
• Stigmatomma emeryi (Saunders, 1890)
• Stigmatomma gaetulicum (Baroni Urbani, 1978)

Strongylognathus

• Strongylognathus afer Emery, 1884

Strumigenys

• Strumigenys baudueri Emery, 1875
• Strumigenys rogeri Emery, 1890
• Strumigenys tenuipilis

Tapinoma

• Tapinoma insularis Seifert, Kaufmann & Fraysse, 2024
• Tapinoma magnum Mayr, 1861
• Tapinoma nigerrimum
• Tapinoma simrothi Krausse-Heldrungen 1911

Technomyrmex

• Technomyrmex vexatus (Santschi, 1919)

Temnothorax

• Temnothorax algerianus (Cagniant, 1968)
• Temnothorax algiricus Forel, 1894
• Temnothorax anacanthus Santschi, 1912
• Temnothorax anacanthus maximus (Santschi, 1931)
• Temnothorax atlantis silvanus (Forel, 1907)
• Temnothorax atlantis suturalis (Santschi, 1931)
• Temnothorax atomus Cagniant & Espadaler, 1997
• Temnothorax brunneus (Cagniant, 1985)
• Temnothorax bucheti Santschi, 1909
• Temnothorax bucheti mixtus (Santschi, 1929)
• Temnothorax convexus Forel, 1894
• Temnothorax curtulus Santschi, 1929
• Temnothorax ditifet Espadaler, 1997
• Temnothorax formosus Santschi, 1909
• Temnothorax gaetulus Santschi, 1923
• Temnothorax kiudiria Espadaler, 1997
• Temnothorax laurae antoniae Cagniant, 1997
• Temnothorax longipilosus (Santschi, 1912)
• Temnothorax marocanus Santschi, 1909
• Temnothorax mauritanicus Santschi, 1909
• Temnothorax mimeuri Cagniant, 1997
• Temnothorax mirabilis Espadaler & Cagniant, 1996
• Temnothorax neminan Espadaler, 1997
• Temnothorax nitidiceps (Dalla Torre, 1893)
• Temnothorax obscurior Dalla Torre, 1893
• Temnothorax pan Santschi, 1936
• Temnothorax pardoi
• Temnothorax personatus Cagniant, 1987
• Temnothorax productus Santschi, 1918
• Temnothorax santschii Forel, 1905
• Temnothorax simesno Espadaler, 1997
• Temnothorax spinosus (Forel, 1909)
• Temnothorax tamriensis Ajerrar, Gómez, Bouharroud, Zaafrani & Cagniant, 2018
• Temnothorax theryi Santschi, 1921
• Temnothorax tramieri (Cagniant, 1983)
• Temnothorax tricolor Santschi, 1925
• Temnothorax turritellus Espadaler, 1997
• Temnothorax tyndalei Forel, 1909
• Temnothorax unifasciatus Latreille, 1798

Tetramorium

• Tetramorium alternans Santschi, 1929
• Tetramorium atlante Cagniant, 1970
• Tetramorium biskrense Forel, 1904
• Tetramorium exasperatum Emery, 1891
• Tetramorium forte Forel, 1904
• Tetramorium semilaeve jugurtha Menozzi, 1934

Uncertain species

The following species was described from Morocco but its true identity is currently unknown.

• Neivamyrmex maroccanus (Santschi, 1926)

References

• Cagniant, H. 1997. Le genre Tetramorium au Maroc (Hymenoptera: Formicidae): clé et catalogue des espèces. Ann. Soc. Entomol. Fr. (N.S.) 33: 89-100.
• Cagniant, H. 2006. Liste actualisée des fourmis du Maroc. Myrmecologische Nachrichten 8: 193-200.
• Cagniant, H.; Espadaler, X. 1997. Les Leptothorax, Epimyrma et Chalepoxenus du Maroc (Hymenoptera: Formicidae). Clé et catalogue des espèces. Ann. Soc. Entomol. Fr. (n.s.) 33: 259-284.
• Taheri, A., Reyes-Lopez, L. 2015. Five new records of ants (Hymenoptera: Formicidae) From Morocco. Journal of Insect Science, 15(37): 1-3 (doi:10.1093/jisesa/iev022).
• Taheri, A., Reyes-Lopez, J.L. 2023. New and additional records for the ant fauna (Hymenoptera, Formicidae) of Morocco. Journal of the Entomological Research Society 25(1): 1-10 (doi:10.51963/jers.v25i1.1995).