Taxa Described in 2003

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The following 2 genera were described in 2003 (see all years)

Taxon Name Subfamily Tribe Author Type Species Fossil Fossil Age Species Count Notes
Tyrannomyrmex Myrmicinae Solenopsidini Fernández Tyrannomyrmex rex 4
Zhangidris Myrmicinae Bolton Heteromyrmex atopogaster, now Zhangidris atopogaster Yes Early Miocene
Shanwang, China
1

The following 494 species/subspecies were described in 2003 (see all years)

Taxon Name Subfamily Tribe Genus Species Subspecies Author Type Locality Country Fossil Fossil Age Notes
Adelomyrmex betoi Myrmicinae Solenopsidini Adelomyrmex betoi Fernández Mexico
Adelomyrmex brevispinosus Myrmicinae Solenopsidini Adelomyrmex brevispinosus Fernández Synonym, see Adelomyrmex tristani
Adelomyrmex costatus Myrmicinae Solenopsidini Adelomyrmex costatus Fernández Colombia
Adelomyrmex cristiani Myrmicinae Solenopsidini Adelomyrmex cristiani Fernández Colombia
Adelomyrmex foveolatus Myrmicinae Solenopsidini Adelomyrmex foveolatus Fernández Costa Rica
Adelomyrmex grandis Myrmicinae Solenopsidini Adelomyrmex grandis Fernández Colombia
Adelomyrmex laevigatus Myrmicinae Solenopsidini Adelomyrmex laevigatus Mackay, W.P. Panama
Adelomyrmex longinoi Myrmicinae Solenopsidini Adelomyrmex longinoi Fernández Costa Rica
Adelomyrmex mackayi Myrmicinae Solenopsidini Adelomyrmex mackayi Fernández Mexico
Adelomyrmex micans Myrmicinae Solenopsidini Adelomyrmex micans Fernández Mexico
Adelomyrmex microps Myrmicinae Solenopsidini Adelomyrmex microps Fernández Costa Rica
Adelomyrmex minimus Myrmicinae Solenopsidini Adelomyrmex minimus Fernández Costa Rica
Adelomyrmex robustus Myrmicinae Solenopsidini Adelomyrmex robustus Fernández Mexico
Adelomyrmex striatus Myrmicinae Solenopsidini Adelomyrmex striatus Fernández Brazil
Adelomyrmex vaderi Myrmicinae Solenopsidini Adelomyrmex vaderi Fernández Colombia
Baracidris pilosa Myrmicinae Solenopsidini Baracidris pilosa Fernández Uganda
Bradoponera electrina Proceratiinae Proceratiini Bradoponera electrina De Andrade & Baroni Urbani Yes Bartonian, Middle to Late Eocene Baltic amber, Baltic Sea region
Bradoponera wunderlichi Proceratiinae Proceratiini Bradoponera wunderlichi Baroni Urbani & De Andrade Yes Bartonian, Middle to Late Eocene Baltic amber, Baltic Sea region
Camponotus fergusoni Formicinae Camponotini Camponotus fergusoni McArthur Australia
Camponotus longifacies Formicinae Camponotini Camponotus longifacies McArthur Australia
Camponotus pawseyi Formicinae Camponotini Camponotus pawseyi McArthur Australia
Camponotus pitjantjatarae Formicinae Camponotini Camponotus pitjantjatarae McArthur Australia
Camponotus rudis Formicinae Camponotini Camponotus rudis McArthur Australia
Camponotus scotti Formicinae Camponotini Camponotus scotti McArthur Australia
Camponotus simpsoni Formicinae Camponotini Camponotus simpsoni McArthur Australia
Camponotus xingdoushanensis Formicinae Camponotini Camponotus xingdoushanensis Wang, W. & Chen China
Cardiocondyla bicoronata Myrmicinae Crematogastrini Cardiocondyla bicoronata Seifert Jordan
Cardiocondyla brachyceps Myrmicinae Crematogastrini Cardiocondyla brachyceps Seifert Iran
Cardiocondyla breviscapa Myrmicinae Crematogastrini Cardiocondyla breviscapa Seifert Synonym, see Cardiocondyla minutior
Cardiocondyla gallilaeica Myrmicinae Crematogastrini Cardiocondyla gallilaeica Seifert Israel
Cardiocondyla goa Myrmicinae Crematogastrini Cardiocondyla goa Seifert India
Cardiocondyla israelica Myrmicinae Crematogastrini Cardiocondyla israelica Seifert Israel
Cardiocondyla littoralis Myrmicinae Crematogastrini Cardiocondyla littoralis Seifert Kazakhstan
Cardiocondyla longiceps Myrmicinae Crematogastrini Cardiocondyla longiceps Seifert Yemen
Cardiocondyla melana Myrmicinae Crematogastrini Cardiocondyla melana Seifert Yemen
Cardiocondyla nana Myrmicinae Crematogastrini Cardiocondyla nana Seifert Borneo
Cardiocondyla opaca Myrmicinae Crematogastrini Cardiocondyla opaca Seifert India
Cardiocondyla opistopsis Myrmicinae Crematogastrini Cardiocondyla opistopsis Seifert Kuwait
Cardiocondyla paranuda Myrmicinae Crematogastrini Cardiocondyla paranuda Seifert Australia
Cardiocondyla persiana Myrmicinae Crematogastrini Cardiocondyla persiana Seifert Iran
Cardiocondyla rugulosa Myrmicinae Crematogastrini Cardiocondyla rugulosa Seifert Yemen
Cardiocondyla semirubra Myrmicinae Crematogastrini Cardiocondyla semirubra Seifert Türkiye
Cardiocondyla shagrinata Myrmicinae Crematogastrini Cardiocondyla shagrinata Seifert India
Cardiocondyla tenuifrons Myrmicinae Crematogastrini Cardiocondyla tenuifrons Seifert Jordan
Cardiocondyla tibetana Myrmicinae Crematogastrini Cardiocondyla tibetana Seifert Tibet
Cardiocondyla unicalis Myrmicinae Crematogastrini Cardiocondyla unicalis Seifert Iran
Carebara acutispina Myrmicinae Crematogastrini Carebara acutispina Xu, Z. China
Carebara altinodus Myrmicinae Crematogastrini Carebara altinodus Xu, Z. China
Carebara bihornata Myrmicinae Crematogastrini Carebara bihornata Xu, Z. China
Carebara curvispina Myrmicinae Crematogastrini Carebara curvispina Xu, Z. China
Carebara obtusidenta Myrmicinae Crematogastrini Carebara obtusidenta Xu, Z. China
Carebara rectidorsa Myrmicinae Crematogastrini Carebara rectidorsa Xu, Z. China
Carebara reticapita Myrmicinae Crematogastrini Carebara reticapita Xu, Z. China
Carebara striata Myrmicinae Crematogastrini Carebara striata Xu, Z. China
Cataglyphis gadeai Formicinae Formicini Cataglyphis gadeai De Haro & Collingwood Spain
Chelaner punctulatus Myrmicinae Solenopsidini Chelaner punctulatus Heterick Australia
Chelaner sublamellatus Myrmicinae Solenopsidini Chelaner sublamellatus Heterick Australia
Crematogaster bryophilia Myrmicinae Crematogastrini Crematogaster bryophilia Longino Costa Rica
Crematogaster flavomicrops Myrmicinae Crematogastrini Crematogaster flavomicrops Longino Costa Rica
Crematogaster flavosensitiva Myrmicinae Crematogastrini Crematogaster flavosensitiva Longino Venezuela
Crematogaster foliocrypta Myrmicinae Crematogastrini Crematogaster foliocrypta Longino Costa Rica
Crematogaster jardinero Myrmicinae Crematogastrini Crematogaster jardinero Longino Costa Rica
Crematogaster levior Myrmicinae Crematogastrini Crematogaster levior Longino Brazil
Crematogaster monteverdensis Myrmicinae Crematogastrini Crematogaster monteverdensis Longino Costa Rica
Crematogaster raptor Myrmicinae Crematogastrini Crematogaster raptor Longino Costa Rica
Crematogaster snellingi Myrmicinae Crematogastrini Crematogaster snellingi Longino Costa Rica
Crematogaster sotobosque Myrmicinae Crematogastrini Crematogaster sotobosque Longino Costa Rica
Crematogaster urvijae Myrmicinae Crematogastrini Crematogaster urvijae Bharti Synonym, see Crematogaster biroi
Crematogaster wardi Myrmicinae Crematogastrini Crematogaster wardi Longino Costa Rica
Cryptomyrmex boltoni Myrmicinae Solenopsidini Cryptomyrmex boltoni Fernández Paraguay
Cryptomyrmex longinodus Myrmicinae Solenopsidini Cryptomyrmex longinodus Fernández & Brandão Brazil
Cyphomyrmex maya Myrmicinae Attini Cyphomyrmex maya De Andrade Yes middle Miocene
Mexican amber, Chiapas, Mexico
Cyphomyrmex taino Myrmicinae Attini Cyphomyrmex taino De Andrade Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Holcoponera aspera Ectatomminae Ectatommini Holcoponera aspera Morgan, Mackay & Pachero Bolivia
Klondikia whiteae Uncertain Klondikia whiteae Dlussky & Rasnitsyn Yes Ypresian, Early Eocene Klondike Formation, Republic, Washington, United States
Lasius austriacus Formicinae Lasiini Lasius austriacus Schlick-Steiner Austria
Leptanilloides mckennae Dorylinae Leptanilloides mckennae Longino Costa Rica
Leptogenys khammouanensis Ponerinae Ponerini Leptogenys khammouanensis Roncin & Deharveng Laos
Megalomyrmex wettereri Myrmicinae Solenopsidini Megalomyrmex wettereri Brandão Costa Rica
Myrmica afghanica Myrmicinae Myrmicini Myrmica afghanica Radchenko & Elmes Afghanistan
Myrmica pelops Myrmicinae Myrmicini Myrmica pelops Seifert Greece
Ochetomyrmex neopolitus Myrmicinae Attini Ochetomyrmex neopolitus Fernández Colombia
Perissomyrmex nepalensis Myrmicinae Crematogastrini Perissomyrmex nepalensis Radchenko Synonym, see Perissomyrmex monticola
Pheidole acamata Myrmicinae Attini Pheidole acamata Wilson Nicaragua
Pheidole aciculata Myrmicinae Attini Pheidole aciculata Wilson Peru
Pheidole aculifera Myrmicinae Attini Pheidole aculifera Wilson Panama
Pheidole ademonia Myrmicinae Attini Pheidole ademonia Wilson Peru
Pheidole aenescens Myrmicinae Attini Pheidole aenescens Wilson Brazil
Pheidole agricola Myrmicinae Attini Pheidole agricola Wilson Mexico
Pheidole alayoi Myrmicinae Attini Pheidole alayoi Wilson Cuba
Pheidole albipes Myrmicinae Attini Pheidole albipes Wilson Mexico
Pheidole alexeter Myrmicinae Attini Pheidole alexeter Wilson Brazil
Pheidole allarmata Myrmicinae Attini Pheidole allarmata Wilson Brazil
Pheidole alpestris Myrmicinae Attini Pheidole alpestris Wilson Ecuador
Pheidole alticola Myrmicinae Attini Pheidole alticola Wilson Colombia
Pheidole amabilis Myrmicinae Attini Pheidole amabilis Wilson Costa Rica
Pheidole amazonica Myrmicinae Attini Pheidole amazonica Wilson Peru
Pheidole ambigua Myrmicinae Attini Pheidole ambigua Wilson Brazil
Pheidole angulifera Myrmicinae Attini Pheidole angulifera Wilson Costa Rica
Pheidole angusticeps Myrmicinae Attini Pheidole angusticeps Wilson Costa Rica
Pheidole anima Myrmicinae Attini Pheidole anima Wilson Honduras
Pheidole arachnion Myrmicinae Attini Pheidole arachnion Wilson Costa Rica
Pheidole araneoides Myrmicinae Attini Pheidole araneoides Wilson Brazil
Pheidole arboricola Myrmicinae Attini Pheidole arboricola Wilson Costa Rica
Pheidole arctos Myrmicinae Attini Pheidole arctos Wilson Synonym, see Pheidole simonsi
Pheidole ariel Myrmicinae Attini Pheidole ariel Wilson Mexico
Pheidole arietans Myrmicinae Attini Pheidole arietans Wilson Synonym, see Pheidole walkeri
Pheidole aripoensis Myrmicinae Attini Pheidole aripoensis Wilson Trinidad and Tobago
Pheidole astur Myrmicinae Attini Pheidole astur Wilson Peru
Pheidole aurea Myrmicinae Attini Pheidole aurea Wilson United States
Pheidole azteca Myrmicinae Attini Pheidole azteca Wilson Mexico
Pheidole bajaensis Myrmicinae Attini Pheidole bajaensis Wilson Mexico
Pheidole bellatrix Myrmicinae Attini Pheidole bellatrix Wilson Colombia
Pheidole beloceps Myrmicinae Attini Pheidole beloceps Wilson Costa Rica
Pheidole bidens Myrmicinae Attini Pheidole bidens Wilson Brazil
Pheidole binasifera Myrmicinae Attini Pheidole binasifera Wilson Colombia
Pheidole bison Myrmicinae Attini Pheidole bison Wilson Bolivia
Pheidole boliviana Myrmicinae Attini Pheidole boliviana Wilson Bolivia
Pheidole boltoni Myrmicinae Attini Pheidole boltoni Wilson Costa Rica
Pheidole boruca Myrmicinae Attini Pheidole boruca Wilson Costa Rica
Pheidole brachyops Myrmicinae Attini Pheidole brachyops Wilson Costa Rica
Pheidole brandaoi Myrmicinae Attini Pheidole brandaoi Wilson Colombia
Pheidole browni Myrmicinae Attini Pheidole browni Wilson Costa Rica
Pheidole bufo Myrmicinae Attini Pheidole bufo Wilson Brazil
Pheidole bulliceps Myrmicinae Attini Pheidole bulliceps Wilson Ecuador
Pheidole bureni Myrmicinae Attini Pheidole bureni Wilson United States
Pheidole calimana Myrmicinae Attini Pheidole calimana Wilson Colombia
Pheidole caltrop Myrmicinae Attini Pheidole caltrop Wilson Panama
Pheidole camilla Myrmicinae Attini Pheidole camilla Wilson Ecuador
Pheidole caracalla Myrmicinae Attini Pheidole caracalla Wilson Peru
Pheidole cardiella Myrmicinae Attini Pheidole cardiella Wilson Peru
Pheidole cardinalis Myrmicinae Attini Pheidole cardinalis Wilson Brazil
Pheidole carinata Myrmicinae Attini Pheidole carinata Wilson Costa Rica
Pheidole cataphracta Myrmicinae Attini Pheidole cataphracta Wilson Colombia
Pheidole caulicola Myrmicinae Attini Pheidole caulicola Wilson Costa Rica
Pheidole ceibana Myrmicinae Attini Pheidole ceibana Wilson Honduras
Pheidole celaena Myrmicinae Attini Pheidole celaena Wilson Costa Rica
Pheidole cerina Myrmicinae Attini Pheidole cerina Wilson Costa Rica
Pheidole chalcoides Myrmicinae Attini Pheidole chalcoides Wilson Synonym, see Pheidole nitidicollis
Pheidole charazana Myrmicinae Attini Pheidole charazana Wilson Colombia
Pheidole chiapasana Myrmicinae Attini Pheidole chiapasana Wilson Synonym, see Pheidole deceptrix
Pheidole chocoensis Myrmicinae Attini Pheidole chocoensis Wilson Colombia
Pheidole chrysops Myrmicinae Attini Pheidole chrysops Wilson Brazil
Pheidole cielana Myrmicinae Attini Pheidole cielana Wilson Synonym, see Pheidole nubicola
Pheidole citrina Myrmicinae Attini Pheidole citrina Wilson Costa Rica
Pheidole coracina Myrmicinae Attini Pheidole coracina Wilson Mexico
Pheidole cornicula Myrmicinae Attini Pheidole cornicula Wilson Bolivia
Pheidole coveri Myrmicinae Attini Pheidole coveri Wilson Trinidad and Tobago
Pheidole crinita Myrmicinae Attini Pheidole crinita Wilson Ecuador
Pheidole crozieri Myrmicinae Attini Pheidole crozieri Wilson Peru
Pheidole cuevasi Myrmicinae Attini Pheidole cuevasi Wilson Peru
Pheidole cuprina Myrmicinae Attini Pheidole cuprina Wilson Peru
Pheidole cursor Myrmicinae Attini Pheidole cursor Wilson Peru
Pheidole cyrtostela Myrmicinae Attini Pheidole cyrtostela Wilson Brazil
Pheidole daphne Myrmicinae Attini Pheidole daphne Wilson Costa Rica
Pheidole dasypyx Myrmicinae Attini Pheidole dasypyx Wilson Panama
Pheidole davidsonae Myrmicinae Attini Pheidole davidsonae Wilson Peru
Pheidole deima Myrmicinae Attini Pheidole deima Wilson Brazil
Pheidole delicata Myrmicinae Attini Pheidole delicata Wilson Bolivia
Pheidole demeter Myrmicinae Attini Pheidole demeter Wilson Peru
Pheidole diabolus Myrmicinae Attini Pheidole diabolus Wilson Colombia
Pheidole dinophila Myrmicinae Attini Pheidole dinophila Wilson Brazil
Pheidole dolon Myrmicinae Attini Pheidole dolon Wilson Peru
Pheidole dorsata Myrmicinae Attini Pheidole dorsata Wilson Colombia
Pheidole dossena Myrmicinae Attini Pheidole dossena Wilson Costa Rica
Pheidole drepanon Myrmicinae Attini Pheidole drepanon Wilson Dominican Republic
Pheidole dryas Myrmicinae Attini Pheidole dryas Wilson Costa Rica
Pheidole dumicola Myrmicinae Attini Pheidole dumicola Wilson Mexico
Pheidole ebenina Myrmicinae Attini Pheidole ebenina Wilson Synonym, see Pheidole laselva
Pheidole ectatommoides Myrmicinae Attini Pheidole ectatommoides Wilson Costa Rica
Pheidole ecuadorana Myrmicinae Attini Pheidole ecuadorana Wilson Ecuador
Pheidole eparmata Myrmicinae Attini Pheidole eparmata Wilson Venezuela
Pheidole epetrion Myrmicinae Attini Pheidole epetrion Wilson Ecuador
Pheidole erethizon Myrmicinae Attini Pheidole erethizon Wilson Mexico
Pheidole eriophora Myrmicinae Attini Pheidole eriophora Wilson Colombia
Pheidole erratilis Myrmicinae Attini Pheidole erratilis Wilson Costa Rica
Pheidole euryscopa Myrmicinae Attini Pheidole euryscopa Wilson Colombia
Pheidole excubitor Myrmicinae Attini Pheidole excubitor Wilson Costa Rica
Pheidole exquisita Myrmicinae Attini Pheidole exquisita Wilson Ecuador
Pheidole fariasana Myrmicinae Attini Pheidole fariasana Wilson Synonym, see Pheidole insipida
Pheidole fissiceps Myrmicinae Attini Pheidole fissiceps Wilson Peru
Pheidole flavifrons Myrmicinae Attini Pheidole flavifrons Wilson Peru
Pheidole floricola Myrmicinae Attini Pheidole floricola Wilson Honduras
Pheidole fowleri Myrmicinae Attini Pheidole fowleri Wilson Brazil
Pheidole fracticeps Myrmicinae Attini Pheidole fracticeps Wilson Trinidad and Tobago
Pheidole fullerae Myrmicinae Attini Pheidole fullerae Wilson Ecuador
Pheidole furtiva Myrmicinae Attini Pheidole furtiva Wilson United States
Pheidole gagates Myrmicinae Attini Pheidole gagates Wilson Peru
Pheidole galba Myrmicinae Attini Pheidole galba Wilson Peru
Pheidole gangamon Myrmicinae Attini Pheidole gangamon Wilson Synonym, see Pheidole simonsi
Pheidole geminata Myrmicinae Attini Pheidole geminata Wilson Ecuador
Pheidole gemmula Myrmicinae Attini Pheidole gemmula Wilson Peru
Pheidole gigaflavens Myrmicinae Attini Pheidole gigaflavens Wilson Argentina
Pheidole gigas Myrmicinae Attini Pheidole gigas Wilson Brazil
Pheidole gilva Myrmicinae Attini Pheidole gilva Wilson Peru
Pheidole globularia Myrmicinae Attini Pheidole globularia Wilson Ecuador
Pheidole glomericeps Myrmicinae Attini Pheidole glomericeps Wilson Mexico
Pheidole glyphoderma Myrmicinae Attini Pheidole glyphoderma Wilson Synonym, see Pheidole walkeri
Pheidole gnomus Myrmicinae Attini Pheidole gnomus Wilson Ecuador
Pheidole gradifer Myrmicinae Attini Pheidole gradifer Wilson Synonym, see Pheidole angusticeps
Pheidole grandinodus Myrmicinae Attini Pheidole grandinodus Wilson Brazil
Pheidole gravida Myrmicinae Attini Pheidole gravida Wilson Venezuela
Pheidole grex Myrmicinae Attini Pheidole grex Wilson Colombia
Pheidole guajirana Myrmicinae Attini Pheidole guajirana Wilson Colombia
Pheidole guayasana Myrmicinae Attini Pheidole guayasana Wilson Ecuador
Pheidole guerrerana Myrmicinae Attini Pheidole guerrerana Wilson Mexico
Pheidole gulo Myrmicinae Attini Pheidole gulo Wilson Nicaragua
Pheidole hamtoni Myrmicinae Attini Pheidole hamtoni Wilson Mexico
Pheidole harlequina Myrmicinae Attini Pheidole harlequina Wilson Dominican Republic
Pheidole harrisonfordi Myrmicinae Attini Pheidole harrisonfordi Wilson Honduras
Pheidole haskinsorum Myrmicinae Attini Pheidole haskinsorum Wilson Bolivia
Pheidole hasticeps Myrmicinae Attini Pheidole hasticeps Wilson Costa Rica
Pheidole hazenae Myrmicinae Attini Pheidole hazenae Wilson Ecuador
Pheidole hector Myrmicinae Attini Pheidole hector Wilson Costa Rica
Pheidole hedlundorum Myrmicinae Attini Pheidole hedlundorum Wilson Venezuela
Pheidole hierax Myrmicinae Attini Pheidole hierax Wilson Panama
Pheidole hispaniolae Myrmicinae Attini Pheidole hispaniolae Wilson Dominican Republic
Pheidole hizemops Myrmicinae Attini Pheidole hizemops Wilson Costa Rica
Pheidole hoelldobleri Myrmicinae Attini Pheidole hoelldobleri Wilson Costa Rica
Pheidole hoplitica Myrmicinae Attini Pheidole hoplitica Wilson United States
Pheidole horribilis Myrmicinae Attini Pheidole horribilis Wilson Ecuador
Pheidole hortonae Myrmicinae Attini Pheidole hortonae Wilson Venezuela
Pheidole huacana Myrmicinae Attini Pheidole huacana Wilson Peru
Pheidole huilana Myrmicinae Attini Pheidole huilana Wilson Colombia
Pheidole humeridens Myrmicinae Attini Pheidole humeridens Wilson Argentina
Pheidole humida Myrmicinae Attini Pheidole humida Wilson Synonym, see Pheidole boltoni
Pheidole inca Myrmicinae Attini Pheidole inca Wilson Peru
Pheidole indagatrix Myrmicinae Attini Pheidole indagatrix Wilson Costa Rica
Pheidole infernalis Myrmicinae Attini Pheidole infernalis Wilson Bolivia
Pheidole iracunda Myrmicinae Attini Pheidole iracunda Wilson Synonym, see Pheidole arachnion
Pheidole jaculifera Myrmicinae Attini Pheidole jaculifera Wilson Brazil
Pheidole jeannei Myrmicinae Attini Pheidole jeannei Wilson Brazil
Pheidole jivaro Myrmicinae Attini Pheidole jivaro Wilson Ecuador
Pheidole juniperae Myrmicinae Attini Pheidole juniperae Wilson United States
Pheidole kugleri Myrmicinae Attini Pheidole kugleri Wilson Peru
Pheidole kukrana Myrmicinae Attini Pheidole kukrana Wilson Nicaragua
Pheidole kuna Myrmicinae Attini Pheidole kuna Wilson Panama
Pheidole kusnezovi Myrmicinae Attini Pheidole kusnezovi Wilson Argentina
Pheidole laelaps Myrmicinae Attini Pheidole laelaps Wilson Colombia
Pheidole lancifera Myrmicinae Attini Pheidole lancifera Wilson Brazil
Pheidole lanigera Myrmicinae Attini Pheidole lanigera Wilson Peru
Pheidole laselva Myrmicinae Attini Pheidole laselva Wilson Costa Rica
Pheidole laticornis Myrmicinae Attini Pheidole laticornis Wilson Synonym, see Pheidole idiota
Pheidole lattkei Myrmicinae Attini Pheidole lattkei Wilson Venezuela
Pheidole laudatana Myrmicinae Attini Pheidole laudatana Wilson Dominica
Pheidole lemnisca Myrmicinae Attini Pheidole lemnisca Wilson Colombia
Pheidole leonina Myrmicinae Attini Pheidole leonina Wilson Peru
Pheidole leptina Myrmicinae Attini Pheidole leptina Wilson Peru
Pheidole longinoi Myrmicinae Attini Pheidole longinoi Wilson Costa Rica
Pheidole longiseta Myrmicinae Attini Pheidole longiseta Wilson Peru
Pheidole lourothi Myrmicinae Attini Pheidole lourothi Wilson Costa Rica
Pheidole lovejoyi Myrmicinae Attini Pheidole lovejoyi Wilson Brazil
Pheidole lucaris Myrmicinae Attini Pheidole lucaris Wilson Brazil
Pheidole lupus Myrmicinae Attini Pheidole lupus Wilson Peru
Pheidole lustrata Myrmicinae Attini Pheidole lustrata Wilson Costa Rica
Pheidole machetula Myrmicinae Attini Pheidole machetula Wilson Peru
Pheidole mackayi Myrmicinae Attini Pheidole mackayi Wilson Mexico
Pheidole macracantha Myrmicinae Attini Pheidole macracantha Wilson Panama
Pheidole macromischoides Myrmicinae Attini Pheidole macromischoides Wilson Cuba
Pheidole macrops Myrmicinae Attini Pheidole macrops Wilson Mexico
Pheidole madrensis Myrmicinae Attini Pheidole madrensis Wilson Mexico
Pheidole mallota Myrmicinae Attini Pheidole mallota Wilson Panama
Pheidole mantilla Myrmicinae Attini Pheidole mantilla Wilson Costa Rica
Pheidole manuana Myrmicinae Attini Pheidole manuana Wilson Peru
Pheidole medialis Myrmicinae Attini Pheidole medialis Wilson Synonym, see Pheidole radoszkowskii
Pheidole meinertopsis Myrmicinae Attini Pheidole meinertopsis Wilson Peru
Pheidole melastomae Myrmicinae Attini Pheidole melastomae Wilson Panama
Pheidole mera Myrmicinae Attini Pheidole mera Wilson United States
Pheidole metana Myrmicinae Attini Pheidole metana Wilson Colombia
Pheidole micon Myrmicinae Attini Pheidole micon Wilson Colombia
Pheidole micridris Myrmicinae Attini Pheidole micridris Wilson Peru
Pheidole microps Myrmicinae Attini Pheidole microps Wilson Peru
Pheidole midas Myrmicinae Attini Pheidole midas Wilson Trinidad and Tobago
Pheidole minax Myrmicinae Attini Pheidole minax Wilson Peru
Pheidole mincana Myrmicinae Attini Pheidole mincana Wilson Synonym, see Pheidole boliviana
Pheidole mirabilis Myrmicinae Attini Pheidole mirabilis Wilson Colombia
Pheidole mittermeieri Myrmicinae Attini Pheidole mittermeieri Wilson Bolivia
Pheidole mixteca Myrmicinae Attini Pheidole mixteca Wilson Mexico
Pheidole moffetti Myrmicinae Attini Pheidole moffetti Wilson French Guiana
Pheidole monstrosa Myrmicinae Attini Pheidole monstrosa Wilson Brazil
Pheidole monteverdensis Myrmicinae Attini Pheidole monteverdensis Wilson Costa Rica
Pheidole mooreorum Myrmicinae Attini Pheidole mooreorum Wilson Synonym, see Pheidole insipida
Pheidole morelosana Myrmicinae Attini Pheidole morelosana Wilson Mexico
Pheidole mosenopsis Myrmicinae Attini Pheidole mosenopsis Wilson Bolivia
Pheidole multispina Myrmicinae Attini Pheidole multispina Wilson Panama
Pheidole napoensis Myrmicinae Attini Pheidole napoensis Wilson Ecuador
Pheidole nasifera Myrmicinae Attini Pheidole nasifera Wilson Peru
Pheidole naylae Myrmicinae Attini Pheidole naylae Wilson Cuba
Pheidole nebulosa Myrmicinae Attini Pheidole nebulosa Wilson Costa Rica
Pheidole nesiota Myrmicinae Attini Pheidole nesiota Wilson Brazil
Pheidole nigricula Myrmicinae Attini Pheidole nigricula Wilson Costa Rica
Pheidole nitella Myrmicinae Attini Pheidole nitella Wilson Costa Rica
Pheidole noar Myrmicinae Attini Pheidole noar Wilson Dominican Republic
Pheidole nubicola Myrmicinae Attini Pheidole nubicola Wilson Mexico
Pheidole oaxacana Myrmicinae Attini Pheidole oaxacana Wilson Mexico
Pheidole obrima Myrmicinae Attini Pheidole obrima Wilson Venezuela
Pheidole oliveirai Myrmicinae Attini Pheidole oliveirai Wilson Colombia
Pheidole olsoni Myrmicinae Attini Pheidole olsoni Wilson Costa Rica
Pheidole onyx Myrmicinae Attini Pheidole onyx Wilson Costa Rica
Pheidole otisi Myrmicinae Attini Pheidole otisi Wilson Costa Rica
Pheidole palenquensis Myrmicinae Attini Pheidole palenquensis Wilson Ecuador
Pheidole paraensis Myrmicinae Attini Pheidole paraensis Wilson Brazil
Pheidole pariana Myrmicinae Attini Pheidole pariana Wilson Venezuela
Pheidole peckorum Myrmicinae Attini Pheidole peckorum Wilson Ecuador
Pheidole pedana Myrmicinae Attini Pheidole pedana Wilson Brazil
Pheidole pelor Myrmicinae Attini Pheidole pelor Wilson United States
Pheidole peltastes Myrmicinae Attini Pheidole peltastes Wilson Brazil
Pheidole pepo Myrmicinae Attini Pheidole pepo Wilson Colombia
Pheidole perdiligens Myrmicinae Attini Pheidole perdiligens Wilson Synonym, see Pheidole plebecula
Pheidole perkinsi Myrmicinae Attini Pheidole perkinsi Wilson Dominica
Pheidole perpilosa Myrmicinae Attini Pheidole perpilosa Wilson United States
Pheidole perryorum Myrmicinae Attini Pheidole perryorum Wilson Brazil
Pheidole peruviana Myrmicinae Attini Pheidole peruviana Wilson Peru
Pheidole petersoni Myrmicinae Attini Pheidole petersoni Wilson Synonym, see Pheidole erratilis
Pheidole petrensis Myrmicinae Attini Pheidole petrensis Wilson Synonym, see Pheidole nubicola
Pheidole pholeops Myrmicinae Attini Pheidole pholeops Wilson Peru
Pheidole pidax Myrmicinae Attini Pheidole pidax Wilson Colombia
Pheidole pilispina Myrmicinae Attini Pheidole pilispina Wilson Panama
Pheidole pilosior Myrmicinae Attini Pheidole pilosior Wilson United States
Pheidole pinicola Myrmicinae Attini Pheidole pinicola Wilson Mexico
Pheidole plato Myrmicinae Attini Pheidole plato Wilson Ecuador
Pheidole polymorpha Myrmicinae Attini Pheidole polymorpha Wilson Mexico
Pheidole portalensis Myrmicinae Attini Pheidole portalensis Wilson United States
Pheidole potosiana Myrmicinae Attini Pheidole potosiana Wilson Mexico
Pheidole praeses Myrmicinae Attini Pheidole praeses Wilson Brazil
Pheidole prattorum Myrmicinae Attini Pheidole prattorum Wilson Costa Rica
Pheidole prolixa Myrmicinae Attini Pheidole prolixa Wilson Synonym, see Pheidole harrisonfordi
Pheidole prostrata Myrmicinae Attini Pheidole prostrata Wilson Costa Rica
Pheidole protensa Myrmicinae Attini Pheidole protensa Wilson Mexico
Pheidole psilogaster Myrmicinae Attini Pheidole psilogaster Wilson Mexico
Pheidole pygmaea Myrmicinae Attini Pheidole pygmaea Wilson Colombia
Pheidole quadriceps Myrmicinae Attini Pheidole quadriceps Wilson Colombia
Pheidole quercicola Myrmicinae Attini Pheidole quercicola Wilson Mexico
Pheidole rectiluma Myrmicinae Attini Pheidole rectiluma Wilson Synonym, see Pheidole bilimeki
Pheidole rectisentis Myrmicinae Attini Pheidole rectisentis Wilson Nicaragua
Pheidole rectispina Myrmicinae Attini Pheidole rectispina Wilson Costa Rica
Pheidole rectitrudis Myrmicinae Attini Pheidole rectitrudis Wilson Costa Rica
Pheidole renae Myrmicinae Attini Pheidole renae Wilson Costa Rica
Pheidole rhytifera Myrmicinae Attini Pheidole rhytifera Wilson Venezuela
Pheidole rosula Myrmicinae Attini Pheidole rosula Wilson Argentina
Pheidole rotundiceps Myrmicinae Attini Pheidole rotundiceps Wilson Ecuador
Pheidole roushae Myrmicinae Attini Pheidole roushae Wilson Mexico
Pheidole rubiceps Myrmicinae Attini Pheidole rubiceps Wilson Venezuela
Pheidole rugiceps Myrmicinae Attini Pheidole rugiceps Wilson Costa Rica
Pheidole ruida Myrmicinae Attini Pheidole ruida Wilson Synonym, see Pheidole harrisonfordi
Pheidole rutilana Myrmicinae Attini Pheidole rutilana Wilson Brazil
Pheidole sabella Myrmicinae Attini Pheidole sabella Wilson Peru
Pheidole sabina Myrmicinae Attini Pheidole sabina Wilson Mexico
Pheidole sagax Myrmicinae Attini Pheidole sagax Wilson Colombia
Pheidole sagittaria Myrmicinae Attini Pheidole sagittaria Wilson Costa Rica
Pheidole sarpedon Myrmicinae Attini Pheidole sarpedon Wilson Ecuador
Pheidole scabriventris Myrmicinae Attini Pheidole scabriventris Wilson Synonym, see Pheidole nebulosa
Pheidole scalaris Myrmicinae Attini Pheidole scalaris Wilson Synonym, see Pheidole transversostriata
Pheidole scimitara Myrmicinae Attini Pheidole scimitara Wilson Peru
Pheidole scitula Myrmicinae Attini Pheidole scitula Wilson Synonym, see Pheidole boliviana
Pheidole scolioceps Myrmicinae Attini Pheidole scolioceps Wilson Peru
Pheidole securigera Myrmicinae Attini Pheidole securigera Wilson Colombia
Pheidole seligmanni Myrmicinae Attini Pheidole seligmanni Wilson Colombia
Pheidole servilia Myrmicinae Attini Pheidole servilia Wilson Colombia
Pheidole setsukoae Myrmicinae Attini Pheidole setsukoae Wilson Mexico
Pheidole sicaria Myrmicinae Attini Pheidole sicaria Wilson Costa Rica
Pheidole sigillata Myrmicinae Attini Pheidole sigillata Wilson Brazil
Pheidole simonsi Myrmicinae Attini Pheidole simonsi Wilson Panama
Pheidole sitiens Myrmicinae Attini Pheidole sitiens Wilson United States
Pheidole spathicornis Myrmicinae Attini Pheidole spathicornis Wilson Mexico
Pheidole spathipilosa Myrmicinae Attini Pheidole spathipilosa Wilson Costa Rica
Pheidole specularis Myrmicinae Attini Pheidole specularis Wilson Costa Rica
Pheidole sphaerica Myrmicinae Attini Pheidole sphaerica Wilson Colombia
Pheidole spilota Myrmicinae Attini Pheidole spilota Wilson Colombia
Pheidole stigma Myrmicinae Attini Pheidole stigma Wilson Ecuador
Pheidole strigosa Myrmicinae Attini Pheidole strigosa Wilson Brazil
Pheidole styrax Myrmicinae Attini Pheidole styrax Wilson Venezuela
Pheidole subnuda Myrmicinae Attini Pheidole subnuda Wilson Colombia
Pheidole subsphaerica Myrmicinae Attini Pheidole subsphaerica Wilson Colombia
Pheidole superba Myrmicinae Attini Pheidole superba Wilson Colombia
Pheidole synarmata Myrmicinae Attini Pheidole synarmata Wilson Guyana
Pheidole tachirana Myrmicinae Attini Pheidole tachirana Wilson Venezuela
Pheidole tambopatae Myrmicinae Attini Pheidole tambopatae Wilson Peru
Pheidole tanyscapa Myrmicinae Attini Pheidole tanyscapa Wilson Costa Rica
Pheidole tarchon Myrmicinae Attini Pheidole tarchon Wilson Argentina
Pheidole tayrona Myrmicinae Attini Pheidole tayrona Wilson Synonym, see Pheidole sculptior
Pheidole tenebra Myrmicinae Attini Pheidole tenebra Wilson Synonym, see Pheidole harrisonfordi
Pheidole tennantae Myrmicinae Attini Pheidole tennantae Wilson Costa Rica
Pheidole tenuis Myrmicinae Attini Pheidole tenuis Wilson Colombia
Pheidole tepuicola Myrmicinae Attini Pheidole tepuicola Wilson Venezuela
Pheidole terribilis Myrmicinae Attini Pheidole terribilis Wilson Suriname
Pheidole tetroides Myrmicinae Attini Pheidole tetroides Wilson Mexico
Pheidole texticeps Myrmicinae Attini Pheidole texticeps Wilson Synonym, see Pheidole plebecula
Pheidole thrasys Myrmicinae Attini Pheidole thrasys Wilson Synonym, see Pheidole simonsi
Pheidole tigris Myrmicinae Attini Pheidole tigris Wilson Colombia
Pheidole tobini Myrmicinae Attini Pheidole tobini Wilson Peru
Pheidole torosa Myrmicinae Attini Pheidole torosa Wilson Mexico
Pheidole trageri Myrmicinae Attini Pheidole trageri Wilson Brazil
Pheidole traini Myrmicinae Attini Pheidole traini Wilson Panama
Pheidole trinitatis Myrmicinae Attini Pheidole trinitatis Wilson Trinidad and Tobago
Pheidole triplex Myrmicinae Attini Pheidole triplex Wilson Trinidad and Tobago
Pheidole tristicula Myrmicinae Attini Pheidole tristicula Wilson Synonym, see Pheidole carapuna
Pheidole tristops Myrmicinae Attini Pheidole tristops Wilson Colombia
Pheidole triumbonata Myrmicinae Attini Pheidole triumbonata Wilson Synonym, see Pheidole walkeri
Pheidole truncula Myrmicinae Attini Pheidole truncula Wilson Costa Rica
Pheidole tschinkeli Myrmicinae Attini Pheidole tschinkeli Wilson Guatemala
Pheidole tuxtlasana Myrmicinae Attini Pheidole tuxtlasana Wilson Mexico
Pheidole ulothrix Myrmicinae Attini Pheidole ulothrix Wilson Costa Rica
Pheidole umphreyi Myrmicinae Attini Pheidole umphreyi Wilson Panama
Pheidole unicornis Myrmicinae Attini Pheidole unicornis Wilson Colombia
Pheidole valens Myrmicinae Attini Pheidole valens Wilson Brazil
Pheidole variceps Myrmicinae Attini Pheidole variceps Wilson Synonym, see Pheidole deceptrix
Pheidole veletis Myrmicinae Attini Pheidole veletis Wilson Bolivia
Pheidole venatrix Myrmicinae Attini Pheidole venatrix Wilson Colombia
Pheidole verricula Myrmicinae Attini Pheidole verricula Wilson Costa Rica
Pheidole vestita Myrmicinae Attini Pheidole vestita Wilson Costa Rica
Pheidole violacea Myrmicinae Attini Pheidole violacea Wilson Costa Rica
Pheidole viriosa Myrmicinae Attini Pheidole viriosa Wilson Ecuador
Pheidole vomer Myrmicinae Attini Pheidole vomer Wilson Colombia
Pheidole wardi Myrmicinae Attini Pheidole wardi Wilson Honduras
Pheidole xanthogaster Myrmicinae Attini Pheidole xanthogaster Wilson Peru
Pheidole xyston Myrmicinae Attini Pheidole xyston Wilson Mexico
Pheidole yucatana Myrmicinae Attini Pheidole yucatana Wilson Mexico
Pheidole zelata Myrmicinae Attini Pheidole zelata Wilson Bolivia
Pheidole zoster Myrmicinae Attini Pheidole zoster Wilson Peru
Platythyrea dentata Ponerinae Platythyreini Platythyrea dentata Lattke Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Platythyrea procera Ponerinae Platythyreini Platythyrea procera Lattke Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Platythyrea scalpra Ponerinae Platythyreini Platythyrea scalpra Lattke Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Polyrhachis punjabi Formicinae Camponotini Polyrhachis punjabi Bharti India
Pristomyrmex acerosus Myrmicinae Crematogastrini Pristomyrmex acerosus Wang, M. Vanuatu
Pristomyrmex boltoni Myrmicinae Crematogastrini Pristomyrmex boltoni Wang, M. New Guinea
Pristomyrmex browni Myrmicinae Crematogastrini Pristomyrmex browni Wang, M. Mauritius
Pristomyrmex collinus Myrmicinae Crematogastrini Pristomyrmex collinus Wang, M. Philippines
Pristomyrmex costatus Myrmicinae Crematogastrini Pristomyrmex costatus Wang, M. Malaysia
Pristomyrmex curvulus Myrmicinae Crematogastrini Pristomyrmex curvulus Wang, M. Philippines
Pristomyrmex divisus Myrmicinae Crematogastrini Pristomyrmex divisus Wang, M. Philippines
Pristomyrmex flatus Myrmicinae Crematogastrini Pristomyrmex flatus Wang, M. Philippines
Pristomyrmex hirsutus Myrmicinae Crematogastrini Pristomyrmex hirsutus Wang, M. Philippines
Pristomyrmex inermis Myrmicinae Crematogastrini Pristomyrmex inermis Wang, M. Papua New Guinea
Pristomyrmex largus Myrmicinae Crematogastrini Pristomyrmex largus Wang, M. Micronesia (Federated States of)
Pristomyrmex longispinus Myrmicinae Crematogastrini Pristomyrmex longispinus Wang, M. Philippines
Pristomyrmex longus Myrmicinae Crematogastrini Pristomyrmex longus Wang, M. New Guinea
Pristomyrmex minusculus Myrmicinae Crematogastrini Pristomyrmex minusculus Wang, M. Tonga
Pristomyrmex modestus Myrmicinae Crematogastrini Pristomyrmex modestus Wang, M. Borneo
Pristomyrmex occultus Myrmicinae Crematogastrini Pristomyrmex occultus Wang, M. Borneo
Pristomyrmex profundus Myrmicinae Crematogastrini Pristomyrmex profundus Wang, M. Malaysia
Pristomyrmex pulcher Myrmicinae Crematogastrini Pristomyrmex pulcher Wang, M. Malaysia
Pristomyrmex quindentatus Myrmicinae Crematogastrini Pristomyrmex quindentatus Wang, M. Indonesia
Pristomyrmex rigidus Myrmicinae Crematogastrini Pristomyrmex rigidus Wang, M. Thailand
Pristomyrmex simplex Myrmicinae Crematogastrini Pristomyrmex simplex Wang, M. Papua New Guinea
Proceratium angulinode Proceratiinae Proceratiini Proceratium angulinode De Andrade Borneo
Proceratium australe Proceratiinae Proceratiini Proceratium australe De Andrade Australia
Proceratium austronesicum Proceratiinae Proceratiini Proceratium austronesicum De Andrade New Guinea
Proceratium avioide Proceratiinae Proceratiini Proceratium avioide De Andrade Synonym, see Proceratium avium
Proceratium banjaranense Proceratiinae Proceratiini Proceratium banjaranense De Andrade Borneo
Proceratium bhutanense Proceratiinae Proceratiini Proceratium bhutanense De Andrade Synonym, see Proceratium williamsi
Proceratium burundense Proceratiinae Proceratiini Proceratium burundense De Andrade Burundi
Proceratium caledonicum Proceratiinae Proceratiini Proceratium caledonicum De Andrade New Caledonia
Proceratium catio Proceratiinae Proceratiini Proceratium catio De Andrade Colombia
Proceratium cavinodus Proceratiinae Proceratiini Proceratium cavinodus De Andrade Australia
Proceratium chickasaw Proceratiinae Proceratiini Proceratium chickasaw De Andrade United States
Proceratium colombicum Proceratiinae Proceratiini Proceratium colombicum De Andrade Colombia
Proceratium confinium Proceratiinae Proceratiini Proceratium confinium De Andrade Pakistan
Proceratium creek Proceratiinae Proceratiini Proceratium creek De Andrade United States
Proceratium cubanum Proceratiinae Proceratiini Proceratium cubanum De Andrade Cuba
Proceratium dayak Proceratiinae Proceratiini Proceratium dayak De Andrade Borneo
Proceratium dominicanum Proceratiinae Proceratiini Proceratium dominicanum De Andrade Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Proceratium dusun Proceratiinae Proceratiini Proceratium dusun De Andrade Borneo
Proceratium ecuadoriense Proceratiinae Proceratiini Proceratium ecuadoriense De Andrade Ecuador
Proceratium foveolatum Proceratiinae Proceratiini Proceratium foveolatum De Andrade Borneo
Proceratium galilaeum Proceratiinae Proceratiini Proceratium galilaeum De Andrade Israel
Proceratium gibberum Proceratiinae Proceratiini Proceratium gibberum De Andrade Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Proceratium gigas Proceratiinae Proceratiini Proceratium gigas De Andrade Australia
Proceratium gracile Proceratiinae Proceratiini Proceratium gracile De Andrade Australia
Proceratium hirsutum Proceratiinae Proceratiini Proceratium hirsutum De Andrade Australia
Proceratium ivimka Proceratiinae Proceratiini Proceratium ivimka De Andrade New Guinea
Proceratium lattkei Proceratiinae Proceratiini Proceratium lattkei De Andrade Venezuela
Proceratium longiscapus Proceratiinae Proceratiini Proceratium longiscapus De Andrade Dominican Republic
Proceratium malesianum Proceratiinae Proceratiini Proceratium malesianum De Andrade Malaysia
Proceratium melitense Proceratiinae Proceratiini Proceratium melitense De Andrade Malta
Proceratium mexicanum Proceratiinae Proceratiini Proceratium mexicanum De Andrade Mexico
Proceratium microsculptum Proceratiinae Proceratiini Proceratium microsculptum De Andrade Borneo
Proceratium oceanicum Proceratiinae Proceratiini Proceratium oceanicum De Andrade Fiji
Proceratium panamense Proceratiinae Proceratiini Proceratium panamense De Andrade Panama
Proceratium poinari Proceratiinae Proceratiini Proceratium poinari Baroni Urbani & De Andrade Yes Burdigalian, Early Miocene Dominican amber, Dominican Republic
Proceratium politum Proceratiinae Proceratiini Proceratium politum De Andrade New Caledonia
Proceratium pumilio Proceratiinae Proceratiini Proceratium pumilio De Andrade Australia
Proceratium robustum Proceratiinae Proceratiini Proceratium robustum De Andrade Australia
Proceratium siamense Proceratiinae Proceratiini Proceratium siamense De Andrade Thailand
Proceratium snellingi Proceratiinae Proceratiini Proceratium snellingi Baroni Urbani & De Andrade New Guinea
Proceratium striativenter Proceratiinae Proceratiini Proceratium striativenter De Andrade Malaysia
Proceratium sulawense Proceratiinae Proceratiini Proceratium sulawense De Andrade Indonesia
Proceratium taino Proceratiinae Proceratiini Proceratium taino De Andrade Dominican Republic
Proceratium terraealtae Proceratiinae Proceratiini Proceratium terraealtae De Andrade Malaysia
Proceratium transitionis Proceratiinae Proceratiini Proceratium transitionis De Andrade Colombia
Strumigenys jiangxiensis Myrmicinae Attini Strumigenys jiangxiensis Zhou & Xu, Z. China
Thaumatomyrmex mandibularis Ponerinae Thaumatomyrmecini Thaumatomyrmex mandibularis Baroni Urbani & De Andrade Cuba
Thaumatomyrmex nageli Ponerinae Ponerini Thaumatomyrmex nageli Baroni Urbani & De Andrade Cuba
Tyrannomyrmex rex Myrmicinae Solenopsidini Tyrannomyrmex rex Fernández Malaysia