Checklist of Carebara species

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Carebara
Myrmicinae
Carebara distincta
Distribution
Species Richness
Statistics
Valid Species	251
Invalid Species	47
Fossils	9
Countries Occupied	94
Year Described	1840
First Species Described	1840
Last Species Described	2024

The following species and subspecies belong to the genus Carebara. Synonyms are listed under their senior names. For valid names only see Carebara species and for distribution information see Carebara species by Country.

A

aberrans (Santschi, 1937) (Angola, Namibia)
aborensis (Wheeler, W.M., 1913) (India)
acuta (Weber, 1952) (Democratic Republic of Congo)
acutispina (Xu, Z., 2003) (China)
adisaki Wimolsuthikul & Jaitrong, 2024 (Thailand)
affinis (Jerdon, 1851) (Australia, Bangladesh, Borneo, Brunei Darussalam, China, India, Indonesia, Laos, Malaysia, Myanmar, Nicobar Island, Philippines, Singapore, Sri Lanka, Thailand)
   • Atta bellicosa Smith, F., 1858
   • Pheidologeton affinis australis Forel, 1915
   • Pheidologeton australis mjobergi Forel, 1918
   • Solenopsis calida Smith, F., 1863
   • Solenopsis laboriosa Smith, F., 1861

affinis javana (Emery, 1893) (Indonesia)
affinis minor (Emery, 1900) (New Guinea)
affinis spinosior (Forel, 1911) (Indonesia)
affinis sumatrensis (Forel, 1913) (Indonesia)
afghana (Pisarski, 1970) (Afghanistan)
africana (Forel, 1910) (Botswana)
alluaudi (Santschi, 1913) (Kenya)
   • Oligomyrmex kenyensis Ettershank, 1966

alluaudi cataractae (Santschi, 1919) (Zimbabwe)
alperti Fernández, 2010 (Malaysia, Philippines)
alpha (Forel, 1905) (Brunei Darussalam, Indonesia)
altinodus (Xu, Z., 2003) (China)
amia (Forel, 1913) (China, Taiwan)
ampla Santschi, 1912 (Democratic Republic of Congo)
ampla cincta Santschi, 1926 (Democratic Republic of Congo)
ampla obscurithorax Santschi, 1926 (Democratic Republic of Congo)
ampla rugosa Santschi, 1928 (Democratic Republic of Congo)
angolensis (Santschi, 1914) (Angola)
angolensis congolensis (Forel, 1916) (Democratic Republic of Congo)
angulata Fernández, 2004 (Colombia, Ecuador)
anophthalma (Emery, 1906) (Brazil, Ecuador, Guyana, Peru)
   • Carebara winifredae Wheeler, W.M., 1922

arabara Fernández, 2010 (Brazil, Colombia, Peru, Trinidad and Tobago)
   • Carebara striata Fernández Fernández, 2004

arabica (Collingwood & van Harten, 2001) (Oman, Saudi Arabia, Yemen)
   • Carebara abuhurayri Sharaf & Aldawood, 2011

armata (Donisthorpe, 1948) (New Guinea)
arnoldi (Forel, 1913) (Zimbabwe)
arnoldiella (Santschi, 1919) (South Africa)
asina (Forel, 1902) (Bangladesh, India)
atoma (Emery, 1900) (Australia, Brunei Darussalam, Fiji, Micronesia (Federated States of), New Guinea, Palau, Samoa, Solomon Islands, Tonga, Wallis and Futuna Islands)
audita Fernández, 2004 (Colombia)

B

bara Azorsa & Fisher, 2018 (Madagascar)
bartrumi Weber, 1943 (Sudan)
bengalensis (Forel, 1902) (Bangladesh, India, Nepal)
berivelo Azorsa & Fisher, 2018 (Madagascar)
beta (Forel, 1905) (Indonesia)
betsi Azorsa & Fisher, 2018 (Madagascar)
bicarinata Santschi, 1912 (French Guiana, Guyana)
bihornata (Xu, Z., 2003) (China)
bokorensis Matsuura & Hosoishi, 2024 (Cambodia, Thailand)
borealis (Terayama, 1996) (Japan)
bouvardi (Santschi, 1913) (Vietnam)
brasiliana Fernández, 2004 (Brazil)
brevipilosa Fernández, 2004 (Argentina, Brazil, Colombia, Costa Rica, El Salvador, Honduras, Mexico, Nicaragua, Panama, Venezuela)
bruchi (Santschi, 1933) (Argentina)
bruni (Forel, 1913) (Sri Lanka)
butteli (Forel, 1913) (Sri Lanka)

C

capreola (Wheeler, W.M., 1927) (China, Vietnam)
capreola laeviceps (Wheeler, W.M., 1928) (China)
carinata Bharti & Kumar, 2013 (India)
castanea Smith, F., 1858 (China, Laos, Singapore, Thailand)
castanea augustata Santschi, 1920 (Laos)
ceylonensis (Forel, 1911) (Sri Lanka)
coeca Fernández, 2004 (Brazil, Colombia)
colobopsis Hosoishi & Yamane, 2022 (Cambodia)
concinna (Mayr, 1867) (Indonesia)
convexa (Weber, 1950) (Uganda)
coqueta Fernández, 2006 (Colombia)
cornigera (Forel, 1902) (Australia)
   • Oligomyrmex corniger parvicornis Forel, 1915

crassiuscula (Emery, 1900) (Australia, New Guinea)
creolei Azorsa & Fisher, 2018 (Mauritius, Seychelles)
cribriceps (Wheeler, W.M., 1927) (Vietnam)
crigensis (Belshaw & Bolton, 1994) (Ghana)
curvispina (Xu, Z., 2003) (China)

D

debilis (Santschi, 1913) (Guinea)
demeter Azorsa & Fisher, 2018 (Madagascar)
dentata Bharti & Kumar, 2013 (India)
deponens (Walker, 1859) (Sri Lanka)
diabola (Santschi, 1913) (Cameroun, Ivory Coast)
diabolica (Baroni Urbani, 1969) (Chad)
distincta (Bolton & Belshaw, 1993) (Ghana, Ivory Coast, Kenya, Nigeria)
diversa (Jerdon, 1851) (Bangladesh, Borneo, Cambodia, China, Guinea, India, Indonesia, Japan, Laos, Malaysia, Myanmar, Nepal, Philippines, Singapore, Sri Lanka, Thailand, Vietnam)
   • Myrmica polita Smith, F. 1860 Smith, F., 1860
   • Pheidole megacephala Smith, F. Smith, F., 1860
   • Pheidole megacephalotes Dalla Torre, 1892
   • Pheidole militaris Smith, F., 1860
   • Pheidole ocellifera Smith, F., 1858
   • Pheidole pabulator Smith, F., 1860
   • Pheidologeton diversus ficta (Forel, 1911)

diversa draco (Santschi, 1920) (China)
diversa laotina (Santschi, 1920) (China, Laos)
diversa macgregori (Wheeler, W.M., 1929) (Philippines)
diversa philippina (Wheeler, W.M., 1929) (Philippines)
diversa standfussi (Forel, 1911) (Guinea)
diversa taprobanae (Smith, F., 1858) (Sri Lanka)
diversa tenuirugosa (Wheeler, W.M., 1929) (Philippines)
diversa williamsi (Wheeler, W.M., 1929) (Philippines)
donisthorpei (Weber, 1950) (Central African Republic)
dota Azorsa & Fisher, 2018 (Madagascar)

E

elmenteitae (Patrizi, 1948) (Ivory Coast, Kenya)
elongata Fernández, 2004 (Colombia)
erythraea (Emery, 1915) (Eritrea, Ethiopia)
escherichi (Forel, 1911) (Sri Lanka)

F

fayrouzae Sharaf, 2013 (Saudi Arabia)
frontalis (Weber, 1950) (Democratic Republic of Congo)

G

globularia Fernández, 2004 (Colombia, Ecuador)
grandidieri (Forel, 1891) (Comoros, Madagascar, Mayotte)
   • Oligomyrmex voeltzkowi Forel, 1907

guineana Fernández, 2006 (Guinea)
   • Carebara silvestrii Santschi Santschi, 1914

gustavmayri Fischer, Azorsa & Fisher, 2014 (Namibia)
   • Pheidologeton mayri Santschi, 1928

H

hainteny Azorsa & Fisher, 2018 (Madagascar)
hammoniae (Stitz, 1923) (Namibia)
hannya (Terayama, 1996) (Japan)
hiragasy Azorsa & Fisher, 2018 (Madagascar)
hornata Bharti & Kumar, 2013 (India)
hostilis (Smith, F., 1858) (South Africa)
hunanensis (Wu, J. & Wang, C., 1995) (China)

I

inca Fernández, 2004 (Costa Rica, Honduras, Peru)
incerta (Santschi, 1919) (Brazil, Zimbabwe)
incierta Fernández, 2004 (Brazil)
   • Carebara incerta Santschi Santschi, 1923

infima (Santschi, 1913) (Guinea)
intermedia Fernández, 2004 (Belize, Guatemala, Honduras, Mexico)

J

jacobsoni (Forel, 1911) (Indonesia)
jajoby Azorsa & Fisher, 2018 (Madagascar)
jeanneli (Santschi, 1913) (Eritrea, Kenya)
jiangxiensis (Wu, J. & Wang, C., 1995) (China)
junodi Forel, 1904 (South Africa)

K

kabosy Azorsa & Fisher, 2018 (Madagascar)
khamiensis (Arnold, 1952) (Zimbabwe)
kofana Fernández, 2004 (Colombia)
kunensis (Ettershank, 1966) (Namibia)
   • Pheidologeton arnoldi Santschi, 1928

L

laeviceps Liu & Zhong, 2024 (China)
lamellifrons (Forel, 1902) (India)
langi Wheeler, W.M., 1922 (Democratic Republic of Congo)
latinoda (Zhou & Zheng, 1997) (China)
latro (Santschi, 1937) (Democratic Republic of Congo)
leei (Forel, 1902) (India)
lignata Westwood, 1840 (Bangladesh, China, India, Indonesia, Nepal)
lilith Fischer, Azorsa & Hita Garcia, 2015 (Ivory Coast)
longiceps (Santschi, 1929) (Argentina)
longii (Wheeler, W.M., 1903) (United States)
lova Azorsa & Fisher, 2018 (Madagascar)
lucida (Santschi, 1917) (Zimbabwe)
lusciosa (Wheeler, W.M., 1928) (China)
luzonensis Fischer, Azorsa & Fisher, 2014 (Philippines)
   • Pheidologeton silvestrii Wheeler, W.M., 1929

M

maccus (Wheeler, W.M., 1929) (Philippines)
madibai Fischer & Azorsa, 2014 (Central African Republic, Democratic Republic of Congo, Gabon, Uganda)
mahafaly Azorsa & Fisher, 2018 (Madagascar)
majeri Fernández, 2004 (Brazil)
malagasy Azorsa & Fisher, 2018 (Madagascar)
manni (Donisthorpe, 1941) (New Guinea)
mayri (Forel, 1901) (Argentina, Brazil, French Guiana, Guyana, Paraguay, Suriname)
melanocephala (Donisthorpe, 1948) (New Guinea)
melasolena (Zhou & Zheng, 1997) (China)
menozzii (Ettershank, 1966) (Equatorial Guinea)
   • Aneleus eidmanni Menozzi, 1942

minima (Emery, 1900) (New Guinea)
minuta Fernández, 2004 (Brazil)
mjobergi (Forel, 1915) (Australia)
mukkaliensis Bharti, Akbar & Aldawood, 2014 (India)

N

nana (Roger, 1863) (India, Sri Lanka)
nanningensis (Li & Tang, 1986) (China)
nayana (Sheela & Narendran, 1997) (India)
nicotianae (Arnold, 1948) (Zambia, Zimbabwe)
norfolkensis (Donisthorpe, 1941) (Australia, Norfolk Island)
nosindambo (Forel, 1891) (Madagascar)
nuda Fernández, 2004 (Brazil)

O

obscura (Viehmeyer, 1914) (Indonesia)
obtusidenta (Xu, Z., 2003) (China, India)
octata (Bolton & Belshaw, 1993) (Kenya)
oertzeni (Forel, 1886) (Albania, Greece, Türkiye)
oertzeni aeolia (Forel, 1911) (Türkiye)
omasi Azorsa & Fisher, 2018 (Madagascar)
oni (Terayama, 1996) (China, Japan)
osborni Wheeler, W.M., 1922 (Democratic Republic of Congo)
overbecki (Viehmeyer, 1916) (Singapore)

P

paeta (Santschi, 1937) (Angola)
panamensis (Wheeler, W.M., 1925) (Ecuador, Panama)
panhai Jaitrong, Pitaktunsakul & Jantarit, 2021 (Thailand)
parva Wimolsuthikul & Jaitrong, 2024 (Thailand)
patrizii Menozzi, 1927 (Somalia)
paya Fernández, 2004 (Colombia, Ecuador)
periyarensis Dhadwal & Bharti, 2022 (India)
perpusilla (Emery, 1895) (Kenya, Rwanda, South Africa, Uganda, United Republic of Tanzania, Zambia, Zimbabwe)
   • Aneleus perpusillus concedens Santschi, 1914
   • Oligomyrmex perpusillus arnoldianus Ettershank, 1966
   • Pheidologeton perpusillum arnoldi Forel, 1914
   • Pheidologeton perpusillus spinosus Forel, 1907

peruviana (Emery, 1906) (Peru)
petulca (Wheeler, W.M., 1922) (Democratic Republic of Congo)
petulens (Santschi, 1920) (Laos)
phragmotica Fischer, Azorsa & Hita Garcia, 2015 (Kenya)
pilosa Fernández, 2004 (Brazil)
pisinna (Bolton & Belshaw, 1993) (Kenya)
placida Azorsa & Fisher, 2018 (Madagascar)
polita (Santschi, 1914) (Cameroun, Central African Republic, Gabon, Guinea, Kenya, Rwanda, Uganda, United Republic of Tanzania)
polyphemus (Wheeler, W.M., 1928) (China)
propomegata Bharti & Kumar, 2013 (India)
pseudolusciosa (Wu, J. & Wang, C., 1995) (China)
pullata (Santschi, 1920) (Indonesia)
pumilia Fischer, Azorsa & Fisher, 2014 (Mozambique)
   • Oligomyrmex nanus Santschi, 1919

pygmaea (Emery, 1887) (Borneo, Brunei Darussalam, Indonesia, Laos, Malaysia, Philippines, Sri Lanka, Thailand)
   • Pheidologeton pygmaeus albipes Emery, 1893
   • Pheidologeton pygmaeus bugnioni Forel, 1915
   • Pheidologeton pygmaeus simalurensis Forel, 1915

pygmaea densistriata (Stitz, 1925)

Q

qianliyan Terayama, 2009 (China, Taiwan)
quratulain Akbar & Bharti, 2017 (India)

R

raberi Azorsa & Fisher, 2018 (Madagascar)
raja (Forel, 1902) (India, Pakistan)
rara (Bolton & Belshaw, 1993) (Kenya)
rectangulata Bharti & Kumar, 2013 (India)
rectidorsa (Xu, Z., 2003) (China, India)
reina Fernández, 2004 (Colombia, Costa Rica, Honduras, Nicaragua, Panama)
   • Carebara semistriata Fernández, 2004

reticapita (Xu, Z., 2003) (China)
reticulata Fernández, 2004 (Colombia)
robertsoni (Bolton & Belshaw, 1993) (South Africa)
rothneyi (Forel, 1902) (India)
rubra (Smith, F., 1860) (Indonesia)
   • Pheidole rubrobrunea Dalla Torre, 1892

rugata (Forel, 1913) (Indonesia)
rugoflabella Fischer, Azorsa & Fisher, 2014 (Indonesia)
   • Pheidologeton rugosus Karavaiev, 1935

S

sakamotoi Terayama, Lin & Eguchi, 2012 (China, Taiwan)
salegi Azorsa & Fisher, 2018 (Madagascar)
sampi Azorsa & Fisher, 2018 (Madagascar)
sangi (Eguchi & Bui, 2007) (Vietnam)
santschii (Weber, 1943) (Sudan)
sarasinorum (Emery, 1901) (Indonesia, Sulawesi)
sarita (Bolton & Belshaw, 1993) (Ghana, Ivory Coast)
satana (Karavaiev, 1935) (Indonesia)
sauteri (Forel, 1912) (China, Japan, Taiwan)
semilaevis (Mayr, 1901) (South Africa)
   • Oligomyrmex hewitti Santschi, 1919

sicheli Mayr, 1862 (Benin, Senegal)
silenus (Smith, F., 1858) (Borneo, Indonesia, Malaysia, Singapore, Sri Lanka, Thailand)
   • Amauromyrmex speculifrons Wheeler, W.M., 1929

silvestrii (Santschi, 1914) (Cameroun, Central African Republic, Equatorial Guinea, Gabon, Ghana, Guinea, Ivory Coast, Kenya, Nigeria, Senegal, Uganda, Zimbabwe)
   • Aneleus punctatus Karavaiev, 1931

similis (Mayr, 1862) (India, Nicobar Island)
sinhala Fischer, Azorsa & Fisher, 2014 (Sri Lanka)
   • Oligomyrmex taprobanae Forel, 1911

sodalis (Emery, 1914) (New Caledonia)
solitaria (Stitz, 1910) (Togo)
spinata Bharti & Kumar, 2013 (India)
stenoptera (Kusnezov, 1952) (Argentina)
striata (Xu, Z., 2003) (China)
sublatro (Forel, 1913) (Indonesia)
subreptor (Emery, 1900) (New Guinea)
sudanensis (Weber, 1943) (Sudan)
sudanica Santschi, 1933 (Sudan)
sundaica (Forel, 1913) (Indonesia)

T

tahitiensis (Wheeler, W.M., 1936) (French Polynesia)
taiponica (Wheeler, W.M., 1928) (China)
tana Azorsa & Fisher, 2018 (Madagascar)
tanana Azorsa & Fisher, 2018 (Madagascar)
tenua Fernández, 2004 (Brazil)
terayamai Bharti, Akbar & Aldawood, 2014 (India)
termitolestes (Wheeler, W.M., 1918) (Democratic Republic of Congo, Nigeria)
thoracica (Weber, 1950) (Ivory Coast, Kenya, Uganda)
traegaordhi (Santschi, 1914) (South Africa)
transversalis (Smith, F., 1860) (Indonesia, Sulawesi)
trechideros (Zhou & Zheng, 1997) (China, Thailand, Vietnam)

U

ugandana (Santschi, 1923) (Uganda)
urichi (Wheeler, W.M., 1922) (Argentina, Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Erebomyrma eidmanni Eidmann, 1936
   • Erebomyrma morai Menozzi, 1931
   • Erebomyrma nevermanni Mann, 1926

V

varia (Santschi, 1920) (Vietnam)
vazimba Azorsa & Fisher, 2018 (Madagascar)
vespillo (Wheeler, W.M., 1921) (China, Vietnam)
vidua Smith, F., 1858 (Eritrea, Kenya, Mozambique, Namibia, Rwanda, South Africa, Zimbabwe)
   • Carebara colossus Gerstäcker, 1859
   • Carebara dux Smith, F., 1858
   • Carebara vidua abdominalis Santschi, 1912

vidua fur Santschi, 1928 (Congo)
viehmeyeri (Mann, 1919) (Solomon Islands)
villiersi (Bernard, 1953) (Cameroun, Central African Republic, Gabon, Guinea)
volsellata (Santschi, 1937) (Cameroun)
vorax (Santschi, 1914) (Ghana)

W

weyeri (Karavaiev, 1930) (Indonesia)
wheeleri (Ettershank, 1966) (China)
   • Oligomyrmex silvestrii Wheeler, W.M., 1928

wroughtonii (Forel, 1902) (Bangladesh, India)

Y

yamatonis (Terayama, 1996) (China, Japan)
yanoi (Forel, 1912) (China, Taiwan)
   • Pheidologeton dentiviris Forel, 1913

Z

zengchengensis (Zhou, Zhao & Jia, 2006) (China)

Fossil Species

A

antiqua (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

G

groehni Radchenko, Dlussky & Gröhn, 2019 (Baltic amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

K

kutscheri Radchenko, Dlussky & Gröhn, 2019 (Bitterfeld amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

N

nitida (Dlussky, 2002) (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

R

rugiceps (Heer, 1849) (Oeningen, Switzerland (Messinian, Late Miocene))

S

schossnicensis (Assmann, 1870) (Schossnitz (= Sosnica?), Silesia, Poland (Late Miocene))
sophiae (Emery, 1891) (Sicilian amber, Italy (Late/Upper Miocene))

T

thorali (Théobald, 1937) (Aix-en-Provence, France (Late Oligocene))

U

ucrainica (Dlussky, 2002) (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))