Checklist of Dolichoderus species

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Dolichoderus
Dolichoderinae
Dolichoderus attelaboides casent0106108 head 1.jpg
Dolichoderus attelaboides
Dolichoderus Distribution.png
Distribution
Dolichoderus Species Richness.png
Species Richness
Statistics
Valid Species 150
Invalid Species 77
Fossils 51
Countries Occupied 85
Year Described 1831
First Species Described 1758
Last Species Described 2022

The following species and subspecies belong to the genus Dolichoderus. Synonyms are listed under their senior names. For valid names only see Dolichoderus species and for distribution information see Dolichoderus species by Country.

A

abruptus (Smith, F., 1858) (Bolivia, Brazil, Colombia, Ecuador, Peru)
   • Hypoclinea ursus Mayr, 1866

affinis Emery, 1889 (Borneo, Brunei Darussalam, China, India, Indonesia, Laos, Malaysia, Myanmar, Nepal, Philippines, Singapore, Vietnam)
   • Dolichoderus affinis nigricans Emery, 1895

affinis glabripes Forel, 1895 (India)
affinis mus Santschi, 1920 (Vietnam)
albamaculus Shattuck & Marsden, 2013 (Australia)
andinus (Kempf, 1962) (Peru)
angusticornis Clark, 1930 (Australia)
attelaboides (Fabricius, 1775) (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)
   • Dolichoderus attelaboides pulla Santschi, 1923
   • Dolichoderus imbecillus Mann, 1916
   • Dolichoderus imbecillus heterogaster Santschi, 1923

australis André, 1896 (Australia)

B

baenae Mackay, W.P., 1993 (Colombia, Ecuador)
bakhtiari Barabag & Jaitrong, 2022 (Thailand)
beccarii Emery, 1887 (Borneo, Indonesia, Malaysia)
bidens (Linnaeus, 1758) (Brazil, Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)
   • Dolichoderus auromaculatus Forel, 1885
   • Dolichoderus bidens attenuatus Forel, 1903
   • Dolichoderus bidens bahiana Santschi, 1921
   • Formica perditor Fabricius, 1804

bispinosus (Olivier, 1792) (Argentina, Belize, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Formica fungosa Fabricius, 1798
   • Hypoclinea vestita Mayr, 1862
   • Polyrhachis arboricola Norton, 1868

brevis Santschi, 1920 (Laos)
brevithorax Menozzi, 1928 (Indonesia)
burmanicus Bingham, 1903 (Myanmar)
butteli Forel, 1913 (Indonesia)

C

canopus Shattuck & Marsden, 2013 (Australia)
carbonarius Emery, 1895 (Malaysia)
carbonarius latisquamis Emery, 1900 (Indonesia)
clarki Wheeler, W.M., 1935 (Australia)
   • Dolichoderus tristis Clark Clark, 1930

clusor Forel, 1907 (Australia)
cogitans Forel, 1912 (Bolivia, Brazil)
coniger (Mayr, 1870) (Borneo, Indonesia, Malaysia)
crawleyi Donisthorpe, 1917 (Singapore)
curvilobus (Lattke, 1987) (Colombia, Costa Rica)
cuspidatus (Smith, F., 1857) (Borneo, Brunei Darussalam, Indonesia, Malaysia, Malaysia, Thailand)
   • Dolichoderus atra Menozzi, 1930
   • Dolichoderus bifurcatus Karavaiev, 1926
   • Dolichoderus cuspidatus surbecki Santschi, 1925
   • Dolichoderus cuspidatus tenuispina Viehmeyer, 1922

D

dajiensis Wang, W. & Zheng, 2005 (China)
debilis Emery, 1890 (Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Panama, Suriname, Trinidad and Tobago, Venezuela)
   • Dolichoderus debilis parabiotica Forel, 1912
   • Dolichoderus debilis sieversi Forel, 1901

decollatus Smith, F., 1858 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)
   • Dolichoderus capitatus Santschi, 1921
   • Dolichoderus decollatus neglectus Menozzi, 1935

dentatus Forel, 1902 (Australia)
diversus Emery, 1894 (Brazil, Colombia, Costa Rica, Ecuador, Guyana, Mexico, Panama, Trinidad and Tobago, Venezuela)
   • Dolichoderus championi Forel, 1899
   • Dolichoderus championi ornatus Mann, 1916
   • Dolichoderus championi taeniatus Wheeler, W.M., 1916
   • Dolichoderus championi trinidadensis Forel, 1899
   • Dolichoderus germaini garbei Forel, 1911
   • Dolichoderus germaini leviusculus Emery, 1906
   • Iridomyrmex mazaruni Donisthorpe, 1939

doloniger (Roger, 1862) (Bolivia, Colombia, Venezuela)
doriae Emery, 1887 (Australia)

E

epetreius (Lattke, 1987) (Venezuela)
erectilobus Santschi, 1920 (Thailand, Vietnam)
etus Shattuck & Marsden, 2013 (Australia)
extensispinus Forel, 1915 (Australia)

F

feae Emery, 1889 (China, India, Myanmar, Thailand)
   • Dolichoderus feae caligatus Wheeler, W.M., 1927
   • Dolichoderus feae fuscus Emery, 1889

fernandezi Mackay, W.P., 1993 (Colombia)
ferrugineus Forel, 1903 (Bolivia, Brazil)
   • Dolichoderus bidens inferior Mann, 1916

flatidorsus Zhou & Zheng, 1997 (China)
formosus Clark, 1930 (Australia)
   • Dolichoderus occidentalis Clark, 1930

furcifer Emery, 1887 (Indonesia)

G

gagates Emery, 1890 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Suriname, Venezuela)
germaini Emery, 1894 (Argentina, Brazil, Ecuador, Paraguay, Peru)
ghilianii Emery, 1894 (Brazil, Peru)
gibbifer Emery, 1887 (Indonesia)
   • Dolichoderus gibbifer gibbosior Forel, 1912

gibbus (Smith, F., 1861) (Indonesia, Sulawesi)
gordoni Shattuck & Marsden, 2013 (Australia)
goudiei Clark, 1930 (Australia)

H

haradae Mackay, W.P., 1993 (Brazil)

I

imitator Emery, 1894 (Bolivia, Brazil, Ecuador)
incisus Xu, Z., 1995 (China)
indrapurensis Forel, 1912 (Borneo, Indonesia, Malaysia, Singapore)
indrapurensis nigrogaster Viehmeyer, 1922 (Borneo, Indonesia, Malaysia)
inermis Mackay, W.P., 1993 (Costa Rica)
inferus Shattuck & Marsden, 2013 (Australia)
inpai (Harada, 1987) (Brazil, Colombia, Ecuador)

J

jacobsoni Forel, 1909 (Indonesia)

K

kathae Shattuck & Marsden, 2013 (Australia)
kinabaluensis Dill, 2002 (Borneo, Indonesia, Malaysia)

L

lactarius (Smith, F., 1860) (Indonesia)
lamellosus (Mayr, 1870) (Argentina, Bolivia, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Paraguay, Suriname, Venezuela)
   • Dolichoderus lamellosa missionensis Santschi, 1916

laminatus (Mayr, 1870) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Suriname, Venezuela)
   • Dolichoderus laminatus luteiventris Emery, 1894

laotius Santschi, 1920 (China, Laos)
laurae Mackay, W.P., 1993 (Colombia)
lobicornis (Kempf, 1959) (Brazil, Colombia, Ecuador)
longicollis Mackay, W.P., 1993 (Colombia)
luederwaldti Santschi, 1921 (Brazil)
lugens Emery, 1894 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Suriname)
lujae Santschi, 1923 (Brazil)
lutosus (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Suriname, Trinidad and Tobago, Venezuela)
   • Dolichoderus lutosus nigriventris Forel, 1893
   • Hypoclinea cingulata Mayr, 1862

M

magnipastor Dill, 2002 (Borneo, Indonesia, Malaysia)
mariae Forel, 1885 (Mexico, United States)
   • Dolichoderus mariae blatchleyi Wheeler, W.M., 1917
   • Dolichoderus mariae davisi Wheeler, W.M., 1905

maschwitzi Dill, 2002 (Borneo, Indonesia, Malaysia)
mesonotalis Forel, 1907 (Brazil, Peru)
   • Dolichoderus grandii Menozzi, 1924
   • Dolichoderus simplex Forel, 1912

modiglianii Emery, 1900 (Indonesia)
   • Dolichoderus modiglianii rubescens Emery, 1900

moggridgei Forel, 1886 (Afghanistan, China, India)
moggridgei bicolor Santschi, 1920 (India)
moggridgei lugubris Santschi, 1920 (India)
monoceros Emery, 1897 (New Guinea)
mucronifer (Roger, 1862) (French Guiana, Suriname)
   • Dolichoderus spinicollis ensiger Forel, 1910

N

niger Crawley, 1922 (Australia)
nigricornis Clark, 1930 (Australia)

O

omacanthus (Kempf, 1972) (Brazil)
omicron Shattuck & Marsden, 2013 (Australia)

P

parvus Clark, 1930 (Australia)
   • Dolichoderus glauerti Wheeler, W.M., 1934

pastorulus Dill, 2002 (Borneo, Indonesia, Malaysia)
patens (Mayr, 1870) (Borneo, Indonesia, Malaysia)
patens pubiventris Emery, 1900 (Indonesia, Philippines)
piceus Mackay, W.P., 1993 (Colombia)
pilinomas Dill, 2002 (Borneo, Indonesia, Malaysia)
pilosus Zhou & Zheng, 1997 (China)
plagiatus (Mayr, 1870) (Canada, Mexico, United States)
   • Dolichoderus borealis Provancher, 1888
   • Dolichoderus plagiatus inornatus Wheeler, W.M., 1905

pustulatus Mayr, 1886 (United States)
   • Dolichoderus plagiatus beutenmuelleri Wheeler, W.M., 1904

Q

quadridenticulatus (Roger, 1862) (Bolivia, Brazil, Ecuador, Peru, Venezuela)
   • Dolichoderus analis Emery, 1894
   • Dolichoderus gibbosus gibbosoanalis Forel, 1922
   • Dolichoderus gibbosus integra Forel, 1911
   • Dolichoderus gibbosus nitidior Emery, 1894
   • Formica gibbosa Smith, F. Smith, F., 1858

quadripunctatus (Linnaeus, 1771) (Albania, Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Croatia, Czechia, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Liechtenstein, Lithuania, Luxembourg, Monaco, Montenegro, North Macedonia, Norway, Poland, Portugal, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Türkiye, Ukraine)
   • Dolichoderus quadripunctatus kratochvili Novak, 1941
   • Dolichoderus quadripunctatus unicolor Ruzsky, 1905

R

reflexus Clark, 1930 (Australia)
   • Dolichoderus armstrongi McAreavey, 1949

rosenbergi Forel, 1911 (Ecuador)
rufescens Mann, 1912 (Brazil)
rufotibialis Clark, 1930 (Australia)
rugocapitus Zhou, 2001 (China)
rugosus (Smith, F., 1858) (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname)
rutilus Shattuck & Marsden, 2013 (Australia)

S

sagmanotus Xu, Z., 2001 (China)
satanus Bolton, 1995 (Brazil)
   • Formica obscura Smith Smith, F., 1858

scabridus Roger, 1862 (Australia)
   • Dolichoderus scabridus ruficornis Santschi, 1916
   • Polyrhachis foveolatus Lowne, 1865

schulzi Emery, 1894 (Brazil, Colombia, Costa Rica)
   • Dolichoderus biolleyi Forel, 1908
   • Dolichoderus schulzi columbica Forel, 1914

scrobiculatus (Mayr, 1876) (Australia)
semiorbis Shattuck & Marsden, 2013 (Australia)
semirugosus (Mayr, 1870) (Borneo, Indonesia, Malaysia, Thailand)
septemspinosus Emery, 1894 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela)
setosus (Kempf, 1959) (Brazil)
shattucki Mackay, W.P., 1993 (Colombia, Ecuador, Panama)
sibiricus Emery, 1889 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation)
   • Dolichoderus abietis Kono & Sugihara, 1939
   • Dolichoderus quadripunctatus japonicus Yoshioka, 1939
   • Dolichoderus quadripunctatus yoshiokae Wheeler, W.M., 1933
   • Dolichoderus sinensis Wheeler, W.M., 1921
   • Dolichoderus sinensis atriceps Wheeler, W.M., 1928

siggii Forel, 1895 (Thailand)
smithi Mackay, W.P., 1993 (Bolivia)
spinicollis (Latreille, 1817) (Brazil, Colombia, Venezuela)
spurius Forel, 1903 (French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)
   • Dolichoderus bidens albatus Viehmeyer, 1922

squamanodus Xu, Z., 2001 (China)
sulcaticeps (Mayr, 1870) (Borneo, Brunei Darussalam, Indonesia, Malaysia, Singapore, Thailand, Vietnam)
   • Dolichoderus sulcaticeps nigriventris Santschi, 1920

sundari Tiwari, 2000 (India)
superaculus (Lattke, 1987) (Colombia)

T

taprobanae (Smith, F., 1858) (Borneo, China, India, Indonesia, Laos, Malaysia, Nepal, Sri Lanka, Vietnam)
   • Dolichoderus semirufus André, 1887
   • Dolichoderus taprobanae obscuripes Santschi, 1920
   • Dolichoderus taprobanae tonkina Santschi, 1920
   • Formica ingruens Walker, 1859
   • Hypoclinea gracilis Motschoulsky, 1863

taprobanae borneensis Forel, 1911 (Borneo, Indonesia, Malaysia)
taprobanae ceramensis Stitz, 1912 (Indonesia)
taprobanae friedrichsi Forel, 1914 (Vietnam)
taprobanae gracilipes (Mayr, 1879) (India, Sri Lanka)
taprobanae siamensis Forel, 1911 (Thailand)
taschenbergi (Mayr, 1866) (Canada, United States)
   • Dolichoderus taschenbergi aterrimus Wheeler, W.M., 1915
   • Dolichoderus taschenbergi gagates Wheeler, W.M., 1905
   • Dolichoderus taschenbergi wheeleriella Forel, 1916

thoracicus (Smith, F., 1860) (Borneo, Cambodia, China, India, Indonesia, Laos, Malaysia, Philippines, Singapore, Sri Lanka, Sulawesi, Thailand, Vietnam)
   • Hypoclinea bituberculata Mayr, 1862
   • Hypoclinea sellaris Roger, 1863

thoracicus bilikanus Santschi, 1925 (Indonesia)
thoracicus borneonensis (Roger, 1863) (Borneo, Indonesia, Malaysia)
thoracicus emarginatus Santschi, 1920 (Laos, Vietnam)
thoracicus lacciperdus Santschi, 1925 (Cambodia)
thoracicus levior Karavaiev, 1926 (Indonesia)
thoracicus nasutus Karavaiev, 1935 (Cambodia)
thoracicus rufescens Stitz, 1925 (Borneo, Indonesia, Malaysia)
tricolor Emery, 1914 (New Caledonia)
tricornis Emery, 1897 (New Guinea)
tristis Mann, 1916 (Brazil, Peru)
tuberifer Emery, 1887 (Indonesia, Thailand)
turneri Forel, 1902 (Australia)

U

utriensis Ortiz & Fernández, 2011 (Colombia, Ecuador)

V

validus (Kempf, 1959) (Costa Rica, Ecuador, Panama)
varians Mann, 1916 (Brazil, Colombia, Ecuador)
voraginosus Mackay, W.P., 1993 (Brazil)

Y

ypsilon Forel, 1902 (Australia)

Fossil Species

A

affectus Théobald, 1937 (Kleinkems, Germany (Early Oligocene))
antiquus Carpenter, 1930 (Florissant, Colorado, United States (Late Eocene))

B

balticus (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
brevicornis Dlussky, 2002 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
brevipalpis Dlussky, 2008 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
brevipennis Dlussky, 2008 (Bitterfeld amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
bruneti Théobald, 1937 (Kleinkems, Germany (Early Oligocene))

C

caribbaeus (Wilson, 1985) (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))
coquandi Théobald, 1937 (Aix-en-Provence, France (Late Oligocene))
cornutus (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

D

dibolius Wilson, 1985 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))
dlusskyi LaPolla & Greenwlalt, 2015 (Kishenehn Formation shale, Montana, United States (Lutetian, Middle Eocene))

E

elegans Wheeler, W.M., 1915 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
evolans Zhang, J., 1989 (Shanwang, China (Early Miocene))
explicans (Förster, 1891) (Kleinkems, Germany (Early Oligocene))

G

granulinotus Dlussky, 2008 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

H

heeri Dlussky & Putyatina, 2014 (Radoboj, Croatia (Burdigalian, Early Miocene))

I

intermedius Mackay, W.P., 1993 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

J

jiaoyanshanensis (Hong, 1985) (Shanwang, China (Early Miocene))
jonasi Dubovikoff & Zharkov, 2022 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

K

kohlsi Dlussky & Rasnitsyn, 2002 (Green River Formation, Colorado, United States (Lutetian, Middle Eocene))
kutscheri Dlussky, 2008 (Bitterfeld amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
kutschlinicus (Deichmüller, 1881) (Kuclín, near Bílina, Czechia (Late Eocene))

L

lacinius Zhang, J., 1989 (Shanwang, China (Early Miocene))
longipennis (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
longipilosus Dlussky, 2002 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
lucidus Dlussky, 2008 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))
luridivenosus Zhang, J., Sun & Zhang, X., 1994 (Shanwang, China (Early Miocene))

M

mesosternalis Wheeler, W.M., 1915 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

N

nanus Dlussky, 2002 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

O

obliteratus (Scudder, 1877) (Quesnel, British Columbia, Canada (Early Miocene?))
oviformis Théobald, 1937 (Célas, Gard, France (Late Eocene))

P

passalomma Wheeler, W.M., 1915 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
perkovskyi Dlussky, 2008 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))
pilipes Dlussky, 2008 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))
pinguis Dlussky, Rasnitsyn & Perfilieva, 2015 (Bolshaya Svetlovodnaya, Sikhote-Alin, Russia (Priabonian, Late Eocene))
polessus Dlussky, 2002 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))
polonicus Dlussky, 2002 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
primitivus (Wilson, 1985) (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))
prolaminatus (Wilson, 1985) (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))
punctatus Dlussky, 2008 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

R

robustus Dlussky, 2002 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))
rohweri Carpenter, 1930 (Florissant, Colorado, United States (Late Eocene))

S

sculpturatus (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))

T

tavridus Perfilieva, Dubovikoff & Dlussky, 2017 (Malyi Kamyshlak, Kerch, Crimea, Russian Federation (Middle Miocene))
tertiarius (Mayr, 1868) (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
transversipetiolaris Zhang, J., Sun & Zhang, X., 1994 (Shanwang, China (Early Miocene))

V

vectensis Donisthorpe, 1920 (Bembridge Marls, Isle of Wight, UK (Priabonian, Late Eocene))
vexillarius Wheeler, W.M., 1915 (Baltic amber, Baltic Sea region, Europe (Bartonian, Middle to Late Eocene))
vlaskini Dlussky, 2008 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))

Z

zherichini Dlussky, 2002 (Rovno amber, Baltic Sea region, Europe (Priabonian, Late Eocene))