Camponotus blandus

Camponotus blandus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Formicinae
Tribe:	Camponotini
Genus:	Camponotus
Species:	C. blandus
Binomial name
Camponotus blandus (Smith, F., 1858) Specimen labels
Subspecies
Camponotus blandus aequinotus Forel, 1902 Camponotus blandus denudatus Emery, 1903 Camponotus blandus pronotalis Santschi, 1936 Camponotus blandus semiceps Emery, 1925

Koch et al. (2018) sampled this species in Caryocar barsiliense trees, in southeastern Brazil cerrado, as part of a study examining species interactions in ant-plants.

Distribution

Distribution based on Regional Taxon Lists

Neotropical Region: Argentina, Bolivia, Brazil (type locality), Colombia, Costa Rica, French Guiana, Guyana, Honduras, Mexico, Paraguay, Peru, Suriname, Uruguay, Venezuela.

Distribution based on AntMaps

Distribution based on AntWeb specimens

Check data from AntWeb

Biology

Passos and Leal (2019) - This species was found attending extrafloral nectaries of the plant Turnera subulata in northeastern Brazil (12°16′24″S 38°57′20″W) Caatinga vegetation. Camponotus blandus, classified as a subordinate species, was the most frequently observed ant at the extrafloral nectaries.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

• blandus. Formica blanda Smith, F. 1858b: 43 (w.) BRAZIL (Pará).
  • Combination in Camponotus: Roger, 1863b: 5;
  • combination in C. (Myrmocamelus): Forel, 1913l: 249;
  • combination in C. (Myrmaphaenus): Emery, 1920b: 256.
  • Status as species: Mayr, 1863: 412; Roger, 1863b: 5; Mayr, 1886c: 356; Emery, in Dalla Torre, 1893: 223; Forel, 1895b: 101; Emery, 1896d: 375 (in list); Emery, 1903: 67; Emery, 1906c: 192; Forel, 1907e: 10; Forel, 1908b: 72; Forel, 1908e: 62; Santschi, 1912e: 534; Forel, 1912i: 75; Forel, 1913l: 249; Bruch, 1914: 228; Forel, 1915c: 363; Luederwaldt, 1918: 50; Santschi, 1922c: 110; Emery, 1925b: 153; Borgmeier, 1927c: 151; Santschi, 1936b: 216; Kusnezov, 1952d: 198; Kusnezov, 1953b: 339; Kempf, 1960e: 399; Kempf, 1961b: 523; Kempf, 1970b: 340; Kempf, 1972a: 44; Zolessi, et al. 1988: 6; Brandão, 1991: 332; Bolton, 1995b: 89; Branstetter & Sáenz, 2012: 255; Guénard & Economo, 2015: 227; Mackay & Mackay, 2019: 753.
  • Current subspecies: nominal plus aequinotus, denudatus, pronotalis, semiceps.

References

• Adams, R.M.M., Wells, R.L., Yanoviak, S.P., Frost, C.J., Fox, E.G.P. 2020. Interspecific Eavesdropping on Ant Chemical Communication. Frontiers in Ecology and Evolution 8. (doi:10.3389/fevo.2020.00024).
• Bolton, B. 1995b. A new general catalogue of the ants of the world. Cambridge, Mass.: Harvard University Press, 504 pp. (page 89, catalogue)
• Emery, C. 1920b. Le genre Camponotus Mayr. Nouvel essai de la subdivision en sous-genres. Rev. Zool. Afr. (Bruss.) 8: 229-260 (page 256, combination in C. (Myrmaphaenus))
• Forel, A. 1913m. Fourmis d'Argentine, du Brésil, du Guatémala & de Cuba reçues de M. M. Bruch, Prof. v. Ihering, Mlle Baez, M. Peper et M. Rovereto. Bull. Soc. Vaudoise Sci. Nat. 49: 203-250 (page 249, combination in C. (Myrmocamelus))
• Gallegro-Ropero, M.C., Feitosa, R.M. 2014. Evidences of Batesian mimicry and parabiosis in ants of the Brazilian savanna. Sociobiology 61, 281-285.
• Koch, E. B. A., W. Dattilo, F. Camarota, and H. L. Vasconcelos. 2018. From species to individuals: does the variation in ant-plant networks scale result in structural and functional changes? Population Ecology. 60:309-318. doi:10.1007/s10144-018-0634-5
• Passos, F. C. S. and L. C. Leal. 2019. Protein matters: ants remove herbivores more frequently from extrafloral nectary-bearing plants when habitats are protein poor. Biological Journal of the Linnean Society. 127:407-416. doi:10.1093/biolinnean/blz033
• Roger, J. 1863b. Verzeichniss der Formiciden-Gattungen und Arten. Berl. Entomol. Z. 7(B Beilage: 1-65 (page 5, combination in Camponotus)
• Smith, F. 1858a. Catalogue of hymenopterous insects in the collection of the British Museum. Part VI. Formicidae. London: British Museum, 216 pp. (page 43, worker described)
• van der Kooi, C.J., Stavenga, D.G., Arikawa, K., Belušič, G., Kelber, A. 2020. Evolution of insect color vision: From spectral sensitivity to visual ecology. Annual Review of Entomology 66, annurev-ento-061720–071644. (doi:10.1146/annurev-ento-061720-071644).

References based on Global Ant Biodiversity Informatics

• Andrade-Silva J., E. K. C. Pereira, O. Silva, C. L. C. Santos, J. H. C. Delabie, and J. M. M. Rebelo. 2015. Ants (Hymenoptera: Formicidae) associated with pig carcasses in an urban area. Sociobiology 62(4): 527-532.
• Antoniazzi R., R. N. S. L. Garoo, W. Dattilo, S. P. Ribeiro, and F. S. Neves. 2019. Ant species richness and interactions in canopies of two distinct successional stages in a tropical dry forest. The Science of Nature 106: 20
• Boscardin J., Correa Costa E., J. Garlet, and A. Fiorentini. 2013. Métodos de Captura para Formigas em Pré-plantio de Eucalyptus grandis. Floresta e Ambiente 20(3): 361-370.
• Brandao, C.R.F. 1991. Adendos ao catalogo abreviado das formigas da regiao neotropical (Hymenoptera: Formicidae). Rev. Bras. Entomol. 35: 319-412.
• Bruch C. 1914. Catálogo sistemático de los formícidos argentinos. Revista del Museo de La Plata 19: 211-234.
• Buren W. F. 1983. Artificial faunal replacement for imported fire ant control. Florida Entomologist 66: 93-100.
• Bächtold A., K. Del-Claro, L. A. Kaminski, A. V. L. Freitas, and P. S. Oliveira. 2012. Natural history of an ant-plant-butterfly interaction in a Neotropical savanna. Journal of Natural History 46: 943-954.
• Cardoso, D.C., T.G. Sobrinho and J.H. Schoereder. 2010. Ant community composition and its relationship with phytophysiognomies in a Brazilian Restinga. Insectes Sociaux 57:293-301
• Carvalho K. S., M. A. F. Carneiro, I. S. Nascimento, A. K. Saha, and E. M. Bruna. Lower ant diversity on earth mounds in a semi-arid ecosystem: natural variation or a sign of degradation? Sociobiology 63(4): 1022-1030.
• Clemes Cardoso D., and J. H. Schoereder. 2014. Biotic and abiotic factors shaping ant (Hymenoptera: Formicidae) assemblages in Brazilian coastal sand dunes: the case of restinga in Santa Catarina. Florida Entomologist 97(4): 1443-1450.
• Clemes Cardoso D., and M. Passos Cristiano. 2010. Myrmecofauna of the Southern Catarinense Restinga sandy coastal plain: new records of species occurrence for the state of Santa Catarina and Brazil. Sociobiology 55(1b): 229-239.
• Coelho M. S., G. W. Fernandes, J. C. Santos, and J. H. C. Delabie. 2009. Ants (Hymenoptera: Formicidae) as bioindicators of land restoration in a Brazilian Atlantic forest fragment. Sociobiology 54(1): 51-63.
• Costa-Milanez C. B., G. Lourenco-Silva, P. T. A. Castro, J. D. Majer, and S. P. Ribeiro. 2014. Are ant assemblages of Brazilian veredas characterised by location or habitat type? Braz. J. Biol. 74(1): 89-99.
• Cuezzo, F. 1998. Formicidae. Chapter 42 in Morrone J.J., and S. Coscaron (dirs) Biodiversidad de artropodos argentinos: una perspectiva biotaxonomica Ediciones Sur, La Plata. Pages 452-462.
• Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
• Diehl-Fleig E. 2014. Termites and Ants from Rio Grande do Sul, Brazil. Sociobiology (in Press).
• Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
• Fernandes I., and J. de Souza. 2018. Dataset of long-term monitoring of ground-dwelling ants (Hymenoptera: Formicidae) in the influence areas of a hydroelectric power plant on the Madeira River in the Amazon Basin. Biodiversity Data Journal 6: e24375.
• Fernandes T. T., R. R. Silva, D. Rodrigues de Souza-Campana, O. Guilherme Morais da Silva, and M. Santina de Castro Morini. 2019. Winged ants (Hymenoptera: Formicidae) presence in twigs on the leaf litter of Atlantic Forest. Biota Neotropica 19(3): http://dx.doi.org/10.1590/1676-0611-bn-2018-0694
• Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
• Forel A. 1908. Catálogo systemático da collecção de formigas do Ceará. Boletim do Museu Rocha 1(1): 62-69.
• Forel A. 1908. Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles 44: 35-72.
• Forel A. 1912. Formicides néotropiques. Part VI. 5me sous-famille Camponotinae Forel. Mémoires de la Société Entomologique de Belgique. 20: 59-92.
• Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
• Gallego-Ropero M.C., R.M. Feitosa & J.R. Pujol-Luz, 2013. Formigas (Hymenoptera, Formicidae) Associadas a Ninhos de Cornitermes cumulans Kollar (Isoptera, Termitidae) no Cerrado do Planalto Central do Brasil. EntomoBrasilis, 6(1): 97-101.
• INBio Collection (via Gbif)
• Kempf W. W. 1961. A survey of the ants of the soil fauna in Surinam (Hymenoptera: Formicidae). Studia Entomologica 4: 481-524.
• Kempf W. W. 1978. A preliminary zoogeographical analysis of a regional ant fauna in Latin America. 114. Studia Entomologica 20: 43-62.
• Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
• Kusnezov N. 1952. El género Camponotus en la Argentina (Hymenoptera, Formicidae). Acta Zoologica Lilloana 12: 183-252.
• Kusnezov N. 1953. La fauna mirmecológica de Bolivia. Folia Universitaria. Cochabamba 6: 211-229.
• Kusnezov N. 1978. Hormigas argentinas: clave para su identificación. Miscelánea. Instituto Miguel Lillo 61:1-147 + 28 pl.
• LeBrun, E.G., C. V. Tillberg, A. V. Suarez, P. J. Folgarait, C. R. Smith and D. A. Holway. 2007. An Experimental Study of Competition between Fire Ants and Argentine Ants in Their Native Range. Ecology 88(1):63-75
• Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
• Lutinski J. A., B. C. Lopes, and A. B. B.de Morais. 2013. Diversidade de formigas urbanas (Hymenoptera: Formicidae) de dez cidades do sul do Brasil. Biota Neotrop. 13(3): 332-342.
• Marcus H. 1945. La diferencia en la fauna de las hormigas de Cochabamba y Liriuni. Revista de Agricultura 3: 28-29.
• Osorio Rosado J. L, M. G. de Goncalves, W. Drose, E. J. Ely e Silva, R. F. Kruger, and A. Enimar Loeck. 2013. Effect of climatic variables and vine crops on the epigeic ant fauna (Hymenoptera: Formicidae) in the Campanha region, state of Rio Grande do Sul, Brazil. J Insect Conserv 17: 1113-1123.
• Pignalberi C. T. 1961. Contribución al conocimiento de los formícidos de la provincia de Santa Fé. Pp. 165-173 in: Comisión Investigación Científica; Consejo Nacional de Investigaciones Científicas y Técnicas (Argentina) 1961. Actas y trabajos del primer Congreso Sudamericano de Zoología (La Plata, 12-24 octubre 1959). Tomo III. Buenos Aires: Librart, 276 pp.
• Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
• Queiroz Haddad G., F. Jorge Cividanes, I. C. Fernandes Martins, L. R. Batista Correa. 2012. Population Fluctuations of Formicidae (Hymenoptera) and Araneae (Arachnida) in Two Tillage Systems in the Region of Guaíra-Sp. Florida Entomologist 95(4): 1012-1018.
• Ribas C. R., J. H. Schoereder, M. Pic, and S. M. Soares. 2003. Tree heterogeneity, resource availability, and larger scale processes regulating arboreal ant species richness. Austral Ecology 28(3): 305-314.
• Ribeiro L. F., R. R. C. Solar, T. G. Sobrinho, D. C. Muscardi, J. H. Schoereder, and A. N. Andersen. 2019. Different trophic groups of arboreal ants show differential responses to resource supplementation in a neotropical savanna. Oecologia 190(2): 433-443.
• Santos M. P. C. J., A. F. Carrano-Moreira, and J. B. Torres. 2012. Diversity of soil ant (Hymenoptera: Formicidae) in dense Atlantic Forest and sugarcane plantationsin the County of Igarassu-PE. Revista Brasileira de Ciências Agrárias 7(4): 648-656.
• Santschi F. 1922. Camponotus néotropiques. Annales de la Société Entomologique de Belgique 62: 97-124.
• Santschi F. 1934. Fourmis de Misiones et du Chaco argentin. Revista de la Sociedad Entomológica Argentina. 6: 23-34.
• Schoereder J. H., T. G. Sobrinho, M. S. Madureira, C. R. Ribas, and P. S. Oliveira. 2010. The arboreal ant community visiting extrafloral nectaries in the Neotropical cerrado savanna. Terrestrial Arthropod Reviews 3: 3-27.
• Sendoya S. F., A. V. L. Freitas, and P. S. Oliveira. 2009. Egg-laying butterflies distinguish predaceous ants by sight. American Naturalist 174: 134-140.
• Silvestre R., C. R. F. Brandão, and R. R. Silva da 2003. Grupos funcionales de hormigas: el caso de los gremios del cerrado. Pp. 113-148 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp.
• Solomon, S.E. and A.S. Mikheyev. 2005. The ant (Hymenoptera: Formicidae) fauna of Cocos Island, Costa Rica. Florida Entomologist 88(4):415-423
• Suguituru S. S., M. Santina de Castro Morini, R. M. Feitosa, and R. Rosa da Silva. 2015. Formigas do Alto Tiete. Canal 6 Editora 458 pages
• Ulyssea M. A., and C. R. F. Brandao. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217–224.
• Ulyssea M.A., C. E. Cereto, F. B. Rosumek, R. R. Silva, and B. C. Lopes. 2011. Updated list of ant species (Hymenoptera, Formicidae) recorded in Santa Catarina State, southern Brazil, with a discussion of research advances and priorities. Revista Brasileira de Entomologia 55(4): 603-–611.
• Ulysséa M. A., C. R. F. Brandão. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217-224.
• Vasconcelos, H.L., J.M.S. Vilhena, W.E. Magnusson and A.L.K.M. Albernaz. 2006. Long-term effects of forest fragmentation on Amazonian ant communities. Journal of Biogeography 33:1348-1356
• Vittar, F. 2008. Hormigas (Hymenoptera: Formicidae) de la Mesopotamia Argentina. INSUGEO Miscelania 17(2):447-466
• Vittar, F., and F. Cuezzo. "Hormigas (Hymenoptera: Formicidae) de la provincia de Santa Fe, Argentina." Revista de la Sociedad Entomológica Argentina (versión On-line ISSN 1851-7471) 67, no. 1-2 (2008).
• Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
• Zolessi L. C. de, Y. P. Abenante, and M. E. de Philippi. 1988. Lista sistematica de las especies de Formicidos del Uruguay. Comun. Zool. Mus. Hist. Nat. Montev. 11: 1-9.
• Zolessi L. C. de; Y. P. de Abenante, and M. E. Philippi. 1989. Catálogo sistemático de las especies de Formícidos del Uruguay (Hymenoptera: Formicidae). Montevideo: ORCYT Unesco, 40 + ix pp.
• da Silva de Oliveira A. B., and F. A. Schmidt. 2019. Ant assemblages of Brazil nut trees Bertholletia excelsa in forest and pasture habitats in the Southwestern Brazilian Amazon. Biodiversity and Conservation 28(2): 329-344.
• da Silva, R.R., C.R.F. Brandao, and R. Silvestre. 2004. Similarity Between Cerrado Localities in Central and Southeastern Brazil Based on the Dry Season Bait Visitors Ant Fauna. Studies on Neotropical Fauna and Environment 39(3):191-199.
• de Almeida Soares S., Y. R. Suarez, W. D. Fernandes, P. M. Soares Tenorio, J. H. C. Delabie, and W. F. Antonialli-Junior. 2013. Temporal variation in the composition of ant assemblages (Hymenoptera, Formicidae) on trees in the Pantanal floodplain, Mato Grosso do Sul, Brazil. Rev. Bras. entomol. 57: 84-90
• de Zolessi, L.C., Y.P. de Abenante and M.E. Philippi. 1987. Lista sistemática de las especies de formícidos del Uruguay. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo 11(165):1-9
• de Zolessi, L.C., Y.P. de Abenante and M.E. Phillipi. 1989. Catalago Systematico de las Especies de Formicidos del Uruguay (Hymenoptera: Formicidae). Oficina Regional de Ciencia y Technologia de la Unesco para America Latina y el Caribe- ORCYT. Montevideo, Uruguay