Myrmica aloba
Myrmica aloba | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | Myrmicinae |
Tribe: | Myrmicini |
Genus: | Myrmica |
Species group: | scabrinodis |
Species: | M. aloba |
Binomial name | |
Myrmica aloba Forel, 1909 | |
Synonyms | |
|
M. aloba is almost exclusively an Iberian form that is associated with hot, but moist grassland conditions, where it nests in very similar places to Myrmica scabrinodis living in cooler climates. In the extreme south-west of Europe near Cape St. Vincent in Portugal, we observed many populous nests living in the tops of large grass tussocks in river flood meadows that are grazed by cattle. They have been found on marshy grassland in north Portugal, where they were hosts to the Large Blue Butterfly Phengaris alcon (Denis et Schiffermiiller) (I. Wynhoff, pers. comm.). In Spain in summer they are often found foraging on rocks near dried-up water-courses, lakes and springs. In the mountains of Soria we have found nests living in soil on cattle meadows near a spring. Here M. aloba lived sympatrically with M. scabrinodis and both species appeared to be hosts to Ph. alcon. The ecological separation of these species on such sites has not yet been studied in detail, but it seemed that M. aloba avoided the very boggy conditions close to the spring, where some M. scabrinodis nests were observed. Sexuals were found in nests from the end of July until mid-September, which indicates that M. aloba swarms in late summer. (Radchenko and Elmes 2010)
Photo Gallery
Identification
Radchenko and Elmes (2010) – A member of the scabrinodis complex of the scabrinodis species group. It is most similar to Myrmica scabrinodis: the female castes of the two species have a strongly curved (though not angularly bent) scape, but M. aloba differs from M. scabrinodis by the absence of a lobe on the bend of the scape having at most only a small longitudinal ridge. Males were described relatively recently (Tinaut and Espadaler 1987) based on nest samples containing both workers and males; the males of M. aloba are very similar to those of M. scabrinodis by the length of scape and character of pilosity on the scape and legs. Garcia et al. (2024), Table 1. Biometric measurements and indexes following Radchenko and Elmes (2010) of M. babiensis, M. aloba (own data, in microns, mean ± standard deviation (minimum; maximum)) and other big sized parasitic Myrmica gynes (minimum–maximum, data taken from Radchenko and Elmes, 2003).
Species | Myrmica babiensis n=19 |
Myrmica aloba n=14 |
Myrmica laurae n=14 |
Myrmica bibikoffi n=2 |
Myrmica hirsuta n=36 |
Myrmica myrmicoxena n=6 |
---|---|---|---|---|---|---|
HL | 1155.5 ± 34.9 (1078; 1212) |
1321.8 ± 28 (1278; 1361) |
950–1090 | 1240–1400 | 1000–1180 | 1020–1050 |
HW | 1070.6 ± 37.3 (979; 1137) |
1185.1 ± 32.9 (1145; 1228) |
870–1050 | 1220–1340 | 880–1100 | 940–970 |
SL | 753.1 ± 25 (712; 792) |
942.8 ± 26.1 (888; 988) |
680–790 | 960–1000 | 720–880 | 670–700 |
AL | 1802.6 ± 70.1 (1650; 1925) |
2015.4 ± 67.8 (1925; 2131) |
1500–1850 | 2140 | 1620–2000 | 1520–1580 |
HL/HW | 1.077 ± 0.034 (1.033; 1.186) |
0.993 ± 0.021 (0.962; 1.025) |
- | - | - | - |
FW/HW | 0.457 ± 0.022 (0.429; 0.516) |
0.349 ± 0.015 (0.326; 0.376) |
0.41–0.49 | 0.35–0.39 | 0.39–0.46 | 0.45–0.46 |
FLW/FW | 1.059 ± 0.015 (1.033; 1.091) |
1.216 ± 0.042 (1.144; 1.274) |
1.05–1.17 | 1.26–1.37 | 1.10–1.30 | 1.16–1.21 |
SL/HL | 0.652 ± 0.025 (0.605; 0.690) |
0.713 ± 0.021 (0.677; 0.754) |
0.70–0.75 | 0.71–0.77 | 0.68–0.81 | 0.65–0.66 |
SL/HW | 0.703 ± 0.024 (0.660; 0.746) |
0.708 ± 0.022 (0.669; 0.747) |
0.74–0.80 | 0.79–0.80 | 0.73–0.84 | 0.71–0.72 |
PL/PH | 1.006 ± 0.066 (0.870; 1.110) |
1.294 ± 0.069 (1.208; 1.421) |
- | - | - | - |
PPW/HW | 0.759 ± 0.043 (0.700; 0.888) |
0.462 ± 0.019 (0.434; 0.497) |
0.62–0.69 | 0.60–0.61 | 0.57–0.72 | 0.56–0.57 |
ESL/HW | 0.321 ± 0.0367 (0.258; 0.382) |
0.286 ± 0.012 (0.259; 0.301) |
0.35–0.41 | 0.30–0.35 | 0.24–0.36 | 0.18–0.23 |
ESD/ESL | 1.977 ± 0.203 (1.594; 2.314) |
1.471 ± 0.073 (1.341; 1.559) |
- | - | - | - |
Keys including this Species
Distribution
Latitudinal Distribution Pattern
Latitudinal Range: 43.686667° to 36.087222°.
North Temperate |
North Subtropical |
Tropical | South Subtropical |
South Temperate |
- Source: AntMaps
Distribution based on Regional Taxon Lists
Palaearctic Region: Balearic Islands, China, France, Iberian Peninsula, Portugal, Spain (type locality).
Distribution based on AntMaps
Distribution based on AntWeb specimens
Check data from AntWeb
Countries Occupied
Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species. |
Estimated Abundance
Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species. |
Biology
Association with Other Organisms
- Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.
- This species is a host for the ant Myrmica babiensis (a workerless inquiline) in Spain (Garcia et al. (2024)).
Flight Period
X | X | X | |||||||||
Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
Source: Radchenko & Elmes, 2010.
- Check details at Worldwide Ant Nuptial Flights Data, AntNupTracker and AntKeeping.
- Explore: Show all Flight Month data or Search these data. See also a list of all data tables or learn how data is managed.
Castes
Nomenclature
The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.
- aloba. Myrmica scabrinodis var. aloba Forel, 1909c: 103 (w.) SPAIN. Santschi, 1925g: 340 (q.); Finzi, 1926: 95 (w.); Tinaut & Espadaler, 1987: 61 (m.). Subspecies of rugulosoides: Finzi, 1926: 95. Raised to species: Santschi, 1931b: 340; Santschi, 1936c: 199. Senior synonym of albuferensis: Santschi, 1931b: 340; of rolandi: Seifert, 1988b: 29. See also: Collingwood & Yarrow, 1969: 56; Radchenko & Elmes, 2010: 84.
- rolandi. Myrmica rolandi Bondroit, 1918: 101 (w.q.) FRANCE. Subspecies of sulcinodis: Santschi, 1919e: 244; Menozzi, 1922b: 325; of scabrinodis: Emery, 1921f: 40; Santschi, 1925g: 340; Santschi, 1931b: 344. Revived status as species: Finzi, 1926: 89; Bernard, 1967: 115. Junior synonym of aloba: Seifert, 1988b: 29.
- albuferensis. Myrmica albuferensis Lomnicki, 1925b: 2 (w.) SPAIN (Balearic Is.). Junior synonym of rolandi: Finzi, 1926: 89; Bernard, 1956b: 257; of aloba: Santschi, 1931b: 340; Seifert, 1988b: 29.
Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Description
Worker
Etymology
M. aloba: from a combination of loba derived from the Greek = lobe [of ear] and the prefix α (a) = not or without, to indicate the mostly absent (or extremely reduced) lobe on the bend of the scape compared to its close relative M. scabrinodis.
References
- Blatrix, R., Aubert, C., Decaëns, T., Berquier, C., Andrei-Ruiz, M.-C., Galkowski, C. 2020. Contribution of a DNA barcode to an assessment of the specificity of ant taxa (Hymenoptera: Formicidae) on Corsica. European Journal of Entomology 117: 420-429 (doi:10.14411/eje.2020.046).
- Borowiec, L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
- Catarineu, C., Barberá, G.G., Reyes-López, J.L. 2018. Zoogeography of the ants (Hymenoptera: Formicidae) of southeastern Iberian Peninsula. Sociobiology 65, 383-396 (doi:10.13102/sociobiology.v65i3.2822).
- Collingwood, C. A.; Yarrow, I. H. H. 1969. A survey of Iberian Formicidae (Hymenoptera). EOS. Rev. Esp. Entomol. 44: 53-101 (page 56, see also)
- Collingwood, C.A., Prince, A. 1998. A guide to ants of Continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Supl nº5, pp 49.
- Ebsen, J.R., Boomsma, J.J. & Nash, D.R. 2019. Phylogeography and cryptic speciation in the Myrmica scabrinodis Nylander, 1846 species complex (Hymenoptera: Formicidae), and their conservation implications. Insect Conservation and Diversity 12: 467-480 (doi:10.1111/icad.12366).
- Finzi, B. 1926. Le forme europee del genere Myrmica Latr. Primo contributo. Boll. Soc. Adriat. Sci. Nat. Trieste 29: 71-119 (page 95, male described, Variety of rugulosoides)
- Forel, A. 1909c. Fourmis d'Espagne récoltées par M. O. Vogt et Mme Cécile Vogt, Docteurs en médecine. Ann. Soc. Entomol. Belg. 53: 103-106 (page 103, worker described)
- García, F., Arnal, J.M., Espadaler, X. 2008. Primeros registros de Myrmica bibikoffi Kutter, 1963 (Hymenoptera: Formicidae) en la Península Ibérica. Heteropterus Rev. Entomol. 8, 211–215.
- García, F., Cuesta-Segura, A.D., Espadaler, X. 2024. Myrmica babiensis sp. nov. (Hymenoptera: Formicidae), a new social parasite from the NW Iberian Peninsula. Annales Zoologici 74(1), 113-127 (doi:10.3161/00034541anz2024.74.1.006).
- Radchenko, A.G. & Elmes, G.W. 2010. Myrmica ants of the Old World. Fauna Mundi 3: 1-789.
- Santschi, F. 1925g. Fourmis d'Espagne et autres espèces paléarctiques (Hymenopt.). EOS. Rev. Esp. Entomol. 1: 339-360 (page 340, queen, male described)
- Santschi, F. 1931c. Notes sur le genre Myrmica (Latreille). Rev. Suisse Zool. 38: 335-355 (page 340, Raised to species, Senior synonym of albuferensis)
- Santschi, F. 1936c. Liste et descriptions de fourmis du Maroc. Bull. Soc. Sci. Nat. Maroc 16: 198-210 (page 199, Raised to species)
- Seifert, B. 1988b. A taxonomic revision of the Myrmica species of Europe, Asia Minor, and Caucasia (Hymenoptera, Formicidae). Abh. Ber. Naturkundemus. Görlitz 62(3): 1-75 (page 29, Senior synonym of rolandi)
- Seifert, B. 2024. Pleistocene population differentiation in the ant Myrmica scabrinodis (Hymenoptera: Formicidae) – a taxonomic borderline case. Soil Organisms 96(1): 11-21 (doi:10.25674/357).
- Tinaut, A.; Espadaler, X. 1987. Description [sic] del macho de Myrmica aloba Forel, 1909 (Hymenoptera, Formicidae). Nouv. Rev. Entomol. (n.s.) 4: 61-69 (page 61, male described)
References based on Global Ant Biodiversity Informatics
- Abril S., C. Gómez. 2012. Lista actualizada de las especies de hormigas de Menorca (Islas Baleares, España) y primera cita de Monomorium andrei Saunders, 1890 (Hymenoptera, Formicidae) de la isla Boletín de la Sociedad Entomológica Aragonesa 50: 403-407.
- Acosta, F. J., M. D. Martínez Ibáñez, and M. A. Morales. "Contribución al conocimiento de la mirmecofauna del encinar peninsular. I." Boletín de la Asociación Española de Entomología 6 (2) (1983): 379-391.
- Alvarez-Blanco P., S. Caut, X. Cerda, and E. Angulo. 2017. Native predators living in invaded areas: responses of terrestrial amphibian species to an Argentine ant invasion. Oecologia 185: 95-106.
- AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
- Antarea (at www.antarea.fr on June 11th 2017)
- Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
- Bernard, F. "Remarques sur le peuplement des Baléares en fourmis." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 47 (1956): 254-266.
- Bondroit, J. "Les fourmis de France et de Belgique." Annales de la Société Entomologique de France 87 (1918): 1-174.
- Bondroit, J. "Notes diverses sur des fourmis d'Europe." Annales de la Société Entomologique de Belgique 59 (1920): 143-158.
- Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
- Cagniant, H. "Deuxième liste de fourmis d'Algérie récoltées principalement en forêt. (1re partie)." Bulletin de la Société d'Histoire Naturelle de Toulouse 105 (1970): 405-430.
- Cagniant, H. "Liste actualisee des fourmis du Maroc (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 193-200.
- Cagniant, H. "Liste préliminaire de fourmis forestières d'Algérie. Résultats obtenus de 1963 à 1966." Bulletin de la Societe d'Histoire Naturelle de Toulouse 104 (1968): 138-147.
- Cagniant, H. "Une nouvelle fourmi parasite d'Algérie: Sifolinia kabylica (nov. sp.), Hyménoptères. Formicidae. Myrmicinae." Insectes Sociaux 17 (1970): 39-47.
- Cagniant, H. "Étude de quelques fourmis marocaines. Statistique provisoire des Formicidae du Maroc." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 53 (1964): 83-118.
- Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
- Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
- Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
- Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
- Collingwood, C. A. "A provisional list of Iberian Formicidae with a key to the worker caste." EOS (Revista española de entomología) Nº LVII (1978): 65-95.
- Comín del Río, P., and Andrés de Haro. "Datos iniciales para un estudio ecológico de las hormigas de Menorca." Bolletí de la Societat d´Historia Natural de les Balears 24 (1980): 23-48.
- Comín del Río, P. "Los Formícidos de Menorca. Contribución al estudio taxonómico, geográfico y biológico." Tesina de licenciatura Universida (1977): 135 pp.
- Comín del Río, P. Estudio de los formícidos de Baleares: Contribución al estudio taxonómico, geográfìco y biológico. Palma de Mallorca: Universidad de las Islas Baleares, 1988.
- Ebsen J. R., J. J. Boomsma, and D. R. Nash. 2019. Phylogeography and cryptic speciation in the Myrmica scabrinodis Nylander, 1846 species complex (Hymenoptera: Formicidae), and their conservation implications. Insect Conservation and Diversity doi: 10.1111/icad.12366
- Eidmann, H. "Die Ameisenfauna der Balearen." Zeitschrift für Morphologie und Ökologie der Tiere 6 (1926): 694-742.
- Eidmann, H. "Zur Kenntnis der Insektekfauna der Balearischen Inseln." Entomologische Mitteilungen 16 (1927): 24-37.
- Emery, C. "Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]." en Wytsman, P. "Genera Insectorum" 174B (1922): 95-206.
- Espadaler, X. "Formicidos de las sierras de Cazorla, del Pozo y Segura (Jaén, España)." Ecología 11 (1997): 489-499.
- Finzi B. 1926. Le forme europee del genere Myrmica Latr. Primo contributo. Bollettino della Società Adriatica di Scienze Naturali in Trieste. 29: 71-119.
- Finzi, B. "Le forme europee del genere Myrmica Latr. Primo contributo." Bollettino della Società Adriatica di Scienze Naturali in Trieste 29 (1926): 71-119.
- Garcia F, Arnal J, Espadaler X. 2008. Primeros registros de Myrmica bibikoffi Kutter, 1963 (Hymenoptera: Formicidae) en la Península Ibérica. Heteropterus Revista de Entomologia 8: 211-215.
- Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
- Goetsch, W. "Beiträge zur Biologie spanischer Ameisen." EOS (Revista española de entomología) 18 (1942): 175-241.
- Gomez K., D. Martinez, and X. Espadaler. 2018. Phylogeny of the ant genus Aphaenogaster (Hymenoptera: Formicidae) in the Iberian Peninsula, with the description of a new species. Sociobiology 65: 215-224.
- Guillem, R. "A survey of the ants of Minorca (Hymenoptera: Formicidae) with two new species for the island: Hypoponera punctatissima (Roger, 1859) and Temnothorax algiricus (Forel, 1894)." Boletín de la Asociación Española de Entomología 33, no. 3-4 (2010): 447-460.
- Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
- Jansen G., R. Savolainen, K. Vespalainen. 2010. Phylogeny, divergence-time estimation, biogeography and social parasite–host relationships of the Holarctic ant genusMyrmica(Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution 56: 294-304.
- Lomnicki, J. "Une contribution à la connaissance de la faune des fourmis des îles Baléares." Polskie Pismo Entomologiczne - Bulletin Entomologique de Pologne 4 (1925): 1-3.
- Martínez Ibáñez, M. D., and X. Espadaler. "Revisión de las hormigas ibéricas de la colección M. Medina y nuevos datos de distribución." Actas de las VIII Jornadas de la Asociación Española de Entomología (1986): 1022-1034.
- Menozzi, C. "Contribution à la faune myrmécologique de l'Espagne." Boletín de la Real Sociedad española de Historia natural (Madrid) 22 (1922): 324-332.
- Menozzi, C. "Zur Kenntnis der Ameisenfauna der Balearen." Zoologischer Anzeiger 66 (1926): 180-182.
- Ordóñez-Urbano C., J. Reyes-López, and S. Carpintero-Ortega. 2007. Estudio faunísidos de los formícidos (Hymenoptera: Formicidae) asociados a los bosques de ribera en la Provincia de Córdoba (España). Primeras Aportaciones. Boletin Sociedad Entomológica Aragonesa 40: 367-375.
- Ortiz, F. J. "Formícidos del litoral granadino." Memoria de Licenciatura Universida (1985): 206 pp.
- Pascual, M. R. "Estudio taxonómico y ecológico de los Formícidos de las Sierras de Alfacar, La Yedra, Huétor y Harana." Tesis Doctoral Universida (1986): 264 pp.
- Radchenko A. G., and G. W. Elmes. 2010. Myrmica ants (Hymenoptera: Formicidae) of the Old World. Fauna Mundi 3. Warsaw: Natura Optima Dux Foundation, 790 pp.
- Reyes-Lopez J., S. Carpintero. 2014. Comparison of the exotic and native ant communities (Hymenoptera: Formicidae) in urban green areas at inland, coastal and insular sites in Spain. Eur. J. Entomol. 111(3): 421428.
- Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
- Santschi F. 1925. Fourmis d'Espagne et autres espéces paléartiques EOS (Revista española de entomología) 1: 339-360.
- Santschi F. 1931. Notes sur le genre Myrmica (Latreille). Revue Suisse de Zoologie 38: 335-355.
- Santschi, F. "Fourmis d'Espagne et autres espéces paléartiques." EOS (Revista española de entomología) 1 (1925): 339-360.
- Santschi, F. "Inventa entomologica itineris Hispanici et Maroccani, quod a. 1926 fecerunt Harald et Håkan Lindberg. Fourmis du Bassin Méditerranéen occidental et du Maroc récoltées par MM. Lindberg." Societas Scientiarum Fennica (Helsingfors) 3 (14) (1931): 1-13.
- Schmitz H. 1950. Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit. Brotéria Ser. Cienc. Nat. 19: 12-16.
- Schmitz, H. "Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit." Brotéria, Ciências Naturais 19 (1950): 12-16.
- Seifert B. 1988. A taxonomic revision of the Myrmica species of Europe, Asia Minor, and Caucasia (Hymenoptera, Formicidae). Abhandlungen und Berichte des Naturkundemuseums Görlitz 62(3): 1-75.
- Seifert, B. "A taxonomic revision of the Myrmica species of Europe, Asia Minor, and Caucasia." Abhandlungen und Berichte des Naturkundemuseums Goerlitz 62(3) (1988): 1-75.
- Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
- Tie Ru, and Xu Shengquan. 2004. Variety and distribution of ants in Northwest China. Journal of Ningxia Agricultural College 25(3): 4-9.
- Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
- Tinaut A., and F. J. Ortiz. 1988. Introduccion al conocimiento de las hormigas de la provincia de Almeria (Hymenoptera: Formicidae). Publicaciones del Instituto de Estudios Almerienses. Boletín (Ciencias) 8: 223-231.
- Tinaut, A., and F. Ruano. "Contribución al conocimiento de los formícidos de la Sierra de la Estrella (Portugal)." Boletín de la Asociación Española de Entomología 18 (3-4) (1994): 97-99.
- Tinaut, A., and X. Espadaler. "Description [sic] del macho de Myrmica aloba Forel, 1909." Nouvelle Revue d'Entomologie (n.s.) 4 ( (1987): 61-69.
- Tinaut, A. "Contribución al conocimiento de los formícidos del Parque Nacional de Doñana." Boletín de la Asociación Española de Entomología 15 (1991): 57-63.
- Trigos Peral G., and J. L. Reyes-Lopez. 2015. New record of Temnothorax algiricus (Hymenoptera, Formicidae) from the Iberian Peninsula and ten additional new records for the ant fauna from Málaga, with preliminary list of species from Natural Park Sierras de Tejeda, Almijara y Alhama (Andalusia, Spain). Boln. Asoc. esp. Ent. 39 (3-4): 411-416.
- Weber N. A. 1950. A revision of the North American ants of the genus Myrmica Latreille with a synopsis of the Palearctic species. III. Annals of the Entomological Society of America 43: 189-226.
- Weber, N. A. "A revision of the North American ants of the genus Myrmica Latreille with a synopsis of the Palearctic species. I." Ann. Entomol. Soc. Am. 40 (1947): 437-474.
- Weber, N. A. "A revision of the North American ants of the genus Myrmica Latreille with a synopsis of the Palearctic species. III." Annals of the Entomological Society of America 43 (1950): 189-226.
- de Haro, Andrés, C. A. Collingwood, and P. Comín del Río. "Prospección mirmecológica por Ibiza y Formentera (Baleares)." Orsis 2 (1986): 115-120.
- de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Extremadura (España) y Sao Brás-Almodovar, Alcácer do Sal, Serra da Estrela (Portugal)." Boletim da Sociedade Portuguesa de Entomologia Suplemento 3(1) (1992): 95-104.
- del Campo Gracia P., M. D. Martinez Ibanez, A. Tinaut, and S. Montagud Alario. 2014. Faunistic study of the Formicidae (Hymenoptera, Formicidae) from the Comunitat Valenciana (Spain). Boln. Asoc. esp. Ent. 38 (1-2): 33-65.