Camponotus maculatus

AntWiki: The Ants --- Online
Camponotus maculatus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Camponotini
Genus: Camponotus
Species: C. maculatus
Binomial name
Camponotus maculatus
(Fabricius, 1782)

Camponotus maculatus casent0104630 profile 1.jpg

Camponotus maculatus casent0104630 dorsal 1.jpg

Specimen labels

Subspecies
Synonyms

Well known, common and widely distributed throughout Africa. A common ground nesting and foraging ant which often ascends trees to tend aphids, but does not build tents (Taylor 1977). It was not among the commoner species in the 76-farm survey of Taylor & Adedoyin (1978). Known to be prey of Oecophylla longinoda (Taylor et al., 2018). While this species has been suggested as being invasive in may parts of the world, these reports appear to be largely the result of misidentifications or are presented with little or no justification; additional details are required to confirm its occurrence outside its native range.

Identification

Keys including this Species

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 49.78306° to -34.76667°.

       
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Afrotropical Region: Angola, Benin, Cameroun, Central African Republic, Chad, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Ivory Coast, Kenya, Lesotho, Liberia, Mali, Mozambique, Namibia, Nigeria, Saudi Arabia, Senegal, Sierra Leone, South Africa, Sudan, São Tomé & Principe, Uganda, United Republic of Tanzania, Zimbabwe.
Indo-Australian Region: Micronesia (Federated States of), Palau.
Malagasy Region: Madagascar, Mauritius, Mayotte, Réunion.
Palaearctic Region: Algeria, Iran, Oman, Türkiye.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Biology

Association with Other Organisms

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  • This species is a mutualist for the aphid Cinara apini (a trophobiont) (Jones, 1927; Saddiqui et al., 2019).
  • This species is a host for the eucharitid wasp Hydrorhoa striaticeps Kieffer complex (a parasitoid) (Quevillon, 2018) (multiple encounter modes; direct transmission; transmission outside nest).
  • This species is a host for the eucharitid wasp Stilbula cyniformis (a parasite) (Universal Chalcidoidea Database) (primary host).
  • This species is a host for the phorid fly Apocephalus coquilletti (a parasite) (phorid.net) (attacked).
  • This species is a host for the strepsipteran Pseudacteon simplex (a parasite) (Quevillon, 2018) (encounter mode secondary; indirect transmission; transmission outside nest).

Castes

Worker

Mcz ent camponotus maculatus hef.jpgMcz ent camponotus maculatus hal.jpgMcz ent camponotus maculatus had.jpgMcz ent camponotus maculatus lbs.jpg
. Owned by Museum of Comparative Zoology.

Images from AntWeb

Camponotus maculatus casent0055753 head 1.jpgCamponotus maculatus casent0055753 profile 1.jpgCamponotus maculatus casent0055753 dorsal 1.jpgCamponotus maculatus casent0055753 label 1.jpg
Worker. Specimen code casent0055753. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0063127 head 1.jpgCamponotus maculatus casent0063127 profile 1.jpgCamponotus maculatus casent0063127 dorsal 1.jpgCamponotus maculatus casent0063127 label 1.jpg
Worker. Specimen code casent0063127. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0101095 head 1.jpgCamponotus maculatus casent0101095 profile 1.jpgCamponotus maculatus casent0101095 dorsal 1.jpgCamponotus maculatus casent0101095 label 1.jpg
Lectotype of Camponotus hova maculatoidesWorker. Specimen code casent0101095. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101096 head 1.jpgCamponotus maculatus casent0101096 profile 1.jpgCamponotus maculatus casent0101096 dorsal 1.jpgCamponotus maculatus casent0101096 label 1.jpg
Worker (major/soldier). Specimen code casent0101096. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101109 head 1.jpgCamponotus maculatus casent0101109 profile 1.jpgCamponotus maculatus casent0101109 dorsal 1.jpgCamponotus maculatus casent0101109 label 1.jpg
Paralectotype of Camponotus hova obscuratusWorker (major/soldier). Specimen code casent0101109. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101110 head 1.jpgCamponotus maculatus casent0101110 profile 1.jpgCamponotus maculatus casent0101110 dorsal 1.jpgCamponotus maculatus casent0101110 label 1.jpg
Lectotype of Camponotus hova obscuratusWorker. Specimen code casent0101110. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101111 head 1.jpgCamponotus maculatus casent0101111 profile 1.jpgCamponotus maculatus casent0101111 dorsal 1.jpgCamponotus maculatus casent0101111 label 1.jpg
Worker. Specimen code casent0101111. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101182 head 1.jpgCamponotus maculatus casent0101182 profile 1.jpgCamponotus maculatus casent0101182 dorsal 1.jpgCamponotus maculatus casent0101182 label 1.jpg
Worker. Specimen code casent0101182. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101183 head 1.jpgCamponotus maculatus casent0101183 profile 1.jpgCamponotus maculatus casent0101183 dorsal 1.jpgCamponotus maculatus casent0101183 label 1.jpg
Worker. Specimen code casent0101183. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101338 head 1.jpgCamponotus maculatus casent0101338 profile 1.jpgCamponotus maculatus casent0101338 dorsal 1.jpgCamponotus maculatus casent0101338 label 1.jpg
Paralectotype of Camponotus hova maculatoidesWorker (major/soldier). Specimen code casent0101338. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101339 head 1.jpgCamponotus maculatus casent0101339 profile 1.jpgCamponotus maculatus casent0101339 dorsal 1.jpgCamponotus maculatus casent0101339 label 1.jpg
Paralectotype of Camponotus hova maculatoidesWorker. Specimen code casent0101339. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101340 head 1.jpgCamponotus maculatus casent0101340 profile 1.jpgCamponotus maculatus casent0101340 dorsal 1.jpgCamponotus maculatus casent0101340 label 1.jpg
Worker. Specimen code casent0101340. Photographer Michele Esposito, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101341 head 1.jpgCamponotus maculatus casent0101341 profile 1.jpgCamponotus maculatus casent0101341 dorsal 1.jpgCamponotus maculatus casent0101341 label 1.jpg
Worker (major/soldier). Specimen code casent0101341. Photographer Michele Esposito, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101352 head 1.jpgCamponotus maculatus casent0101352 profile 1.jpgCamponotus maculatus casent0101352 dorsal 1.jpgCamponotus maculatus casent0101352 label 1.jpg
Worker (major/soldier). Specimen code casent0101352. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101353 head 1.jpgCamponotus maculatus casent0101353 profile 1.jpgCamponotus maculatus casent0101353 dorsal 1.jpgCamponotus maculatus casent0101353 label 1.jpg
Worker. Specimen code casent0101353. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101354 head 1.jpgCamponotus maculatus casent0101354 profile 1.jpgCamponotus maculatus casent0101354 dorsal 1.jpgCamponotus maculatus casent0101354 label 1.jpg
Worker (major/soldier). Specimen code casent0101354. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101356 head 1.jpgCamponotus maculatus casent0101356 profile 1.jpgCamponotus maculatus casent0101356 dorsal 1.jpgCamponotus maculatus casent0101356 label 1.jpg
Worker. Specimen code casent0101356. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101747 head 1.jpgCamponotus maculatus casent0101747 profile 1.jpgCamponotus maculatus casent0101747 dorsal 1.jpgCamponotus maculatus casent0101747 label 1.jpg
Paralectotype of Camponotus hova maculatoidesWorker (major/soldier). Specimen code casent0101747. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0101929 head 1.jpgCamponotus maculatus casent0101929 profile 1.jpgCamponotus maculatus casent0101929 dorsal 1.jpgCamponotus maculatus casent0101929 label 1.jpg
Worker (major/soldier). Specimen code casent0101929. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0103332 head 1.jpgCamponotus maculatus casent0103332 profile 1.jpgCamponotus maculatus casent0103332 dorsal 1.jpgCamponotus maculatus casent0103332 label 1.jpg
Worker (major/soldier). Specimen code casent0103332. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0055750 head 1.jpgCamponotus maculatus casent0055750 profile 1.jpgCamponotus maculatus casent0055750 dorsal 1.jpgCamponotus maculatus casent0055750 label 1.jpg
Worker. Specimen code casent0055750. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0104631 head 1.jpgCamponotus maculatus casent0104631 profile 1.jpgCamponotus maculatus casent0104631 dorsal 1.jpgCamponotus maculatus casent0104631 label 1.jpg
Syntype of Camponotus grandidieri comorensisWorker. Specimen code casent0104631. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by ZMHB, Berlin, Germany.
Camponotus maculatus casent0104633 head 1.jpgCamponotus maculatus casent0104633 profile 1.jpgCamponotus maculatus casent0104633 dorsal 1.jpgCamponotus maculatus casent0104633 label 1.jpg
Paratype of Camponotus rubripes hagensiiWorker. Specimen code casent0104633. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by ZMHB, Berlin, Germany.
Camponotus maculatus casent0104638 head 1.jpgCamponotus maculatus casent0104638 profile 1.jpgCamponotus maculatus casent0104638 dorsal 1.jpgCamponotus maculatus casent0104638 label 1.jpg
Type of unavailable quadrinomial: Camponotus maculatus hova laticollusWorker (major/soldier). Specimen code casent0104638. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by ZMHB, Berlin, Germany.
Camponotus maculatus casent0135006 head 1.jpgCamponotus maculatus casent0135006 profile 1.jpgCamponotus maculatus casent0135006 dorsal 1.jpgCamponotus maculatus casent0135006 label 1.jpg
Worker. Specimen code casent0135006. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0146442 head 1.jpgCamponotus maculatus casent0146442 profile 1.jpgCamponotus maculatus casent0146442 dorsal 1.jpgCamponotus maculatus casent0146442 label 1.jpg
Worker (major/soldier). Specimen code casent0146442. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0179509 head 1.jpgCamponotus maculatus casent0179509 profile 1.jpgCamponotus maculatus casent0179509 dorsal 1.jpgCamponotus maculatus casent0179509 label 1.jpg
Worker. Specimen code casent0179509. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Queen

Images from AntWeb

Camponotus maculatus casent0147206 head 1.jpgCamponotus maculatus casent0147206 profile 1.jpgCamponotus maculatus casent0147206 profile 2.jpgCamponotus maculatus casent0147206 dorsal 1.jpgCamponotus maculatus casent0147206 label 1.jpg
Queen (alate/dealate). Specimen code casent0147206. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0125148 head 1.jpgCamponotus maculatus casent0125148 profile 1.jpgCamponotus maculatus casent0125148 dorsal 1.jpgCamponotus maculatus casent0125148 label 1.jpg
Queen (alate/dealate). Specimen code casent0125148. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0101184 head 1.jpgCamponotus maculatus casent0101184 profile 1.jpgCamponotus maculatus casent0101184 dorsal 1.jpgCamponotus maculatus casent0101184 dorsal 2.jpgCamponotus maculatus casent0101184 label 1.jpg
Queen (alate/dealate). Specimen code casent0101184. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0055751 head 1.jpgCamponotus maculatus casent0055751 profile 1.jpgCamponotus maculatus casent0055751 dorsal 1.jpgCamponotus maculatus casent0055751 label 1.jpg
Queen (alate/dealate). Specimen code casent0055751. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0101879 head 1.jpgCamponotus maculatus casent0101879 profile 1.jpgCamponotus maculatus casent0101879 profile 2.jpgCamponotus maculatus casent0101879 dorsal 1.jpgCamponotus maculatus casent0101879 label 1.jpg
Queen (alate/dealate). Specimen code casent0101879. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.

Male

Images from AntWeb

Camponotus maculatus casent0101185 head 1.jpgCamponotus maculatus casent0101185 profile 1.jpgCamponotus maculatus casent0101185 dorsal 1.jpgCamponotus maculatus casent0101185 label 1.jpg
Male (alate). Specimen code casent0101185. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Camponotus maculatus casent0101569 head 1.jpgCamponotus maculatus casent0101569 profile 1.jpgCamponotus maculatus casent0101569 profile 2.jpgCamponotus maculatus casent0101569 dorsal 1.jpgCamponotus maculatus casent0101569 label 1.jpg
Male (alate). Specimen code casent0101569. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Camponotus maculatus casent0146792 head 1.jpgCamponotus maculatus casent0146792 profile 1.jpgCamponotus maculatus casent0146792 profile 2.jpgCamponotus maculatus casent0146792 profile 3.jpgCamponotus maculatus casent0146792 dorsal 1.jpgCamponotus maculatus casent0146792 label 1.jpg
Male (alate). Specimen code casent0146792. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Camponotus maculatus casent0101355 head 1.jpgCamponotus maculatus casent0101355 profile 1.jpgCamponotus maculatus casent0101355 dorsal 1.jpgCamponotus maculatus casent0101355 label 1.jpg
Male (alate). Specimen code casent0101355. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • maculatus. Formica maculata Fabricius, 1782: 491 (w.) (no state data,”Habitat in Africa aequinoctiali”).
    • Smith, F. 1858b: 29 (s.w.); Mayr, 1862: 654 (q.m.); Forel, 1891b: 29 (s.w.q.m.).
    • Combination in Camponotus: Mayr, 1862: 654;
    • combination in C. (Myrmoturba): Forel, 1912i: 91;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 86.
    • Subspecies of sylvaticus: Forel, 1879a: 64; Emery & Forel, 1879: 448; Mayr, 1880: 21; Emery, 1881b: 526; André, 1882a: 145; André, 1884b: 536; Emery, 1884a: 375; Emery, 1887a: 212 (footnote).
    • Subspecies of rubripes: Forel, 1886f: 143, 149; Emery, 1887a: 212 (footnote).
    • Status as species: Fabricius, 1787: 308; Gmelin, 1790: 2800; Olivier, 1792: 495; Fabricius, 1793: 356; Latreille, 1802c: 283; Fabricius, 1804: 403; Lepeletier de Saint-Fargeau, 1835: 214; Haliday, 1836: 329; Smith, F. 1858b: 28 (redescription); Gerstäcker, 1859: 262; Mayr, 1862: 654 (redescription); Gerstäcker, 1862: 509; Mayr, 1863: 400; Roger, 1863b: 2; Mayr, 1865: 27; Gerstäcker, 1871: 355; Mayr, 1877: 20 (in list); Emery, 1877b: 364; Emery, 1878b: 44; Forel, 1891b: 23, 29 (redescription); Emery, 1891b: 17; Forel, 1892e: 349; Emery, 1892a: 119; Emery, 1892d: 566; Dalla Torre, 1893: 241; Emery, 1893i: 671; Mayr, 1893: 195; Mayr, 1895: 150; Emery, 1895a: 180; Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1896i: 158; Mayr, 1896: 249; Mayr, 1897: 432; Emery, 1897e: 604; Emery, 1898a: 225; Forel, 1899c: 134; Emery, 1899a: 501; Mayr, 1901b: 28; Mayr, 1904b: 11; Emery, 1905d: 37; Ruzsky, 1905b: 191; Mayr, 1907a: 389; Mayr, 1907b: 19; Forel, 1907e: 16; Forel, 1907g: 88; Forel, 1908a: 16; Emery, 1908a: 188 (infraspecific taxa key); Santschi, 1908: 518; Bondroit, 1910: 487; Forel, 1910c: 266; Forel, 1910f: 26; Karavaiev, 1910b: 9; Stitz, 1910: 148; Wheeler, W.M. 1910d: 301; Stitz, 1911b: 392; Forel, 1911d: 371; Viehmeyer, 1912: 8; Forel, 1913a: 145; Forel, 1913b: 340; Santschi, 1913h: 42; Santschi, 1914b: 129; Santschi, 1914c: 288; Santschi, 1914d: 379; Forel, 1914d: 249; Wheeler, W.M. & Mann, 1914: 47; Donisthorpe, 1915a: 221 (redescription of holotype); Wheeler, W.M. 1915g: 814; Forel, 1915d: 70 (in key); Emery, 1915f: 79; Emery, 1916b: 224; Stitz, 1916: 403; Mann, 1916: 474; Stitz, 1917: 351; Luederwaldt, 1918: 49; Mann, 1920: 435; Emery, 1920c: 5, 12; Arnold, 1922: 621; Wheeler, W.M. 1922a: 234, 949, 1040; Soudek, 1922: 91; Viehmeyer, 1923: 93; Stitz, 1923: 164; Wheeler, W.M. 1924b: 253; Wheeler, W.M. 1925b: 3; Emery, 1925b: 86; Santschi, 1925h: 166; Crawley, 1926: 393 (redescription); Menozzi, 1926a: 41; Karavaiev, 1926e: 189; Arnold, 1926: 292; Karavaiev, 1927a: 294; Santschi, 1932f: 519; Finzi, 1936: 189; Santschi, 1936e: 81; Finzi, 1939a: 159; Finzi, 1940: 163; Weber, 1943c: 385; Donisthorpe, 1946e: 33; Chapman & Capco, 1951: 248; Bernard, 1953b: 260; Prins, 1963: 106; Baroni Urbani, 1972: 125; Hamann & Klemm, 1976: 676; Collingwood, 1985: 282; Taylor & Brown, 1985: 116; Marsh, 1986: 342; Bolton, 1995b: 109; Collingwood & Agosti, 1996: 374; Blard, et al. 2003: 129; Paknia, et al. 2008: 153; Mackay & Delsinne, 2009: 496 (in key); Legakis, 2011: 31; Borowiec, L. & Salata, 2012: 480; Hita Garcia, et al. 2013: 205; Borowiec, L. 2014: 36 (see note in bibliography); Madl, 2019: 14.
    • Senior synonym of atramentarius: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of ballioni: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of boera: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of cavallus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of cluisoides: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of cognata: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
    • Senior synonym of conakryensis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of erythraea: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of flavifemur: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of flavominor: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of hannae: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of hieroglyphicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of intonsus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of lacteipennis: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
    • Senior synonym of liengmei: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of liocnemis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of lividior: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of lohieri: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of manzer: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of mathildae: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of melanocnemis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of miserabilis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of nubis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of proletaria: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of radamoides: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of sarmentus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of schereri: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of schultzei: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of semispicatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • Senior synonym of sudanicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
    • Senior synonym of thomensis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of tuckeri: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Senior synonym of zumpti: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
    • Material of the unavailable names calceatus, cataractae, citinus, cluis, contaminatus, diffusus, hansingi, importunoides, madecassa, pessimus referred here by Baroni Urbani, 1972: 124-127; Bolton, 1995b: 110.
    • Current subspecies: nominal plus foveolatus (unresolved junior homonym), miserabilis (unresolved junior homonym), obfuscatus, subnudus, sylvaticomaculatus, ugandensis.
  • atramentarius. Camponotus maculatus var. atramentarius Forel, 1904b: 379 (w.) ETHIOPIA.
    • Forel, 1910f: 26 (q.).
    • Combination in C. (Myrmoturba): Forel, 1913b: 340;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Forel, 1905b: 180; Forel, 1907e: 14; Forel, 1907g: 88; Forel, 1909b: 54, 67; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1911f: 280; Forel, 1913b: 340; Santschi, 1914b: 131; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 952; Stitz, 1923: 164; Emery, 1925b: 88; Santschi, 1930b: 75; Santschi, 1937d: 240; Bernard, 1953b: 261.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 86.
  • ballioni. Camponotus maculatus r. ballioni Forel, 1904d: 176 (w.q.) SOUTH AFRICA.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
    • As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 954.
    • Status as species: Santschi, 1925h: 166.
    • Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 87.
  • boera. Camponotus (Myrmoturba) ballioni var. boera Santschi, 1925h: 166.
    • [First available use of Camponotus maculatus subsp. cognatus var. boera Forel, 1910f: 27 (s.w.q.) SOUTH AFRICA; unavailable (infrasubspecific) name.]
    • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Arnold, 1922: 632; Wheeler, W.M. 1922a: 954; Emery, 1925b: 87.
    • Subspecies of ballioni: Santschi, 1930b: 57.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 89.
  • cavallus. Camponotus maculatus st. cavallus Santschi, 1911g: 211 (s.w.q.) ANGOLA.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Stitz, 1916: 403; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Emery, 1925b: 88; Santschi, 1930b: 57; Santschi, 1937d: 241.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 91.
  • cluisoides. Camponotus (Myrmoturba) maculatus var. cluisoides Forel, 1913h: 354 (s.w.) KENYA.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
    • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Wheeler, W.M. 1922a: 952.
    • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1927f: 361.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
  • cognata. Formica cognata Smith, F. 1858b: 35 (s.w.) SOUTH AFRICA.
    • Mayr, 1862: 655 (q.m.).
    • Combination in Camponotus: Mayr, 1862: 655;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • As unavailable (infrasubspecific) name: Emery, 1894i: 46; Emery, 1895h: 46.
    • Status as species: Mayr, 1877: 20 (in list); Dalla Torre, 1893: 225.
    • Subspecies of sylvaticus: Forel, 1879a: 65; Emery & Forel, 1879: 448; Mayr, 1880: 22; Emery, 1881b: 526; André, 1882a: 146; Emery, 1882: 448; Emery, 1884a: 375; André, 1884b: 536; Forel, 1885b: 176; Forel, 1886d: 239; Forel, 1886e: clxvii; Medina, 1893: 104.
    • Subspecies of rubripes: Forel, 1886f: 143, 151; Emery, 1887a: 212 (footnote); Mayr, 1889: 278; Forel, 1889: 255; Forel, 1890a: lxv; Saunders, E. 1890: 202.
    • Subspecies of maculatus: Mayr, 1862: 655; Emery, 1891b: 19; Emery, 1892d: 566; Emery, in Dalla Torre, 1893: 225, 241 (footnote); Mayr, 1893: 195; André, 1895a: 1; Mayr, 1896: 249; Forel, 1904b: 378; Forel, 1904c: 12; Ruzsky, 1905b: 205; Stitz, 1910: 148; Forel, 1910f: 25; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Arnold, 1922: 631; Emery, 1925b: 88; Menozzi, 1942: 180; Eidmann, 1944: 469.
    • Junior synonym of maculatus: Roger, 1863b: 2; Mayr, 1863: 400; Mayr, 1865: 27; Gerstäcker, 1871: 355; Baroni Urbani, 1972: 127; Bolton, 1995b: 93.
  • conakryensis. Camponotus (Myrmoturba) maculatus var. conakryensis Emery, 1920c: 13 (s.w.) GUINEA.
    • Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1930a: 73.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
  • erythraea. Camponotus (Myrmoturba) maculatus var. erythraea Emery, 1920c: 14 (s.) ERITREA.
    • Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1935b: 275.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 98.
  • flavifemur. Camponotus (Myrmoturba) maculatus var. flavifemur Santschi, 1937g: 84 (w.q.) TANZANIA.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 99.
  • flavominor. Camponotus (Tanaemyrmex) maculatus var. flavominor Emery, 1925b: 87.
    • [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. flavominor Santschi, 1920i: 4 (s.w.) BENIN (footnote); unavailable (infrasubspecific) name.]
    • As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 956.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 100.
  • hannae. Camponotus (Myrmoturba) maculatus st. hannae Santschi, 1919a: 349, fig. 4 (s.w.) ZIMBABWE.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88; Arnold, 1926: 294.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 102.
  • hieroglyphicus. Camponotus maculatus st. hieroglyphicus Santschi, 1917b: 290 (s.w.q.) ANGOLA.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 103.
  • intonsus. Camponotus maculatus subsp. intonsus Emery, 1905d: 29 (footnote) (s.w.) SOUTH AFRICA.
    • As unavailable (infrasubspecific) name: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
    • Subspecies of maculatus: Arnold, 1922: 632; Wheeler, W.M. 1922a: 955.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 105.
  • lacteipennis. Formica lacteipennis Smith, F. 1858b: 34 (s.w.q.m.) SOUTH AFRICA.
    • Combination in Camponotus: Roger, 1863b: 2;
    • combination in C. (Myrmoturba): Forel, 1914a: 267;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 90.
    • Status as species: Mayr, 1863: 416; Roger, 1863b: 2; Dalla Torre, 1893: 237.
    • Subspecies of sylvaticus: Mayr, 1886c: 355.
    • Subspecies of maculatus: Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1920c: 5; Arnold, 1922: 626; Wheeler, W.M. 1922a: 955; Emery, 1925b: 89.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 127; Bolton, 1995b: 107.
  • liengmei. Camponotus sexpunctatus var. liengmei Forel, 1894b: 67 (s.w.) MOZAMBIQUE.
    • Forel, 1907g: 88 (m.).
    • Combination in C. (Myrmoturba): Forel, 1914a: 267;
    • combination in C. (Dinomyrmex): Santschi, 1921f: 312;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 87.
    • As unavailable (infrasubspecific) name: Emery, 1895h: 46; Emery, 1896d: 371 (in list.
    • Subspecies of maculatus: Forel, 1901h: 69; Forel, 1907g: 88; Forel, 1910e: 452; Forel, 1911e: 288; Forel, 1914d: 249; Emery, 1920c: 5; Arnold, 1922: 636; Wheeler, W.M. 1922a: 955; Emery, 1925b: 87; Prins, 1963: 107.
    • Status as species: Santschi, 1923e: 292.
    • Junior synonym of maculatus Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
  • liocnemis. Camponotus maculatus subsp. liocnemis Emery, 1905d: 30 (footnote) (s.w.) SOUTH AFRICA.
    • [Misspelled as lioculmis by Forel, 1905b: 180 (in text); misspelled as liochnemis by Santschi, 1914e: 38.]
    • Santschi, 1914e: 38 (q.m.).
    • Combination in C. (Myrmoturba): Santschi, 1914d: 379.
    • As unavailable (infrasubspecific) name: Forel, 1907e: 16; Forel, 1909b: 54; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1913b: 340; Santschi, 1917b: 293; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 958; Emery, 1925b: 88; Santschi, 1925h: 166; Santschi, 1930a: 73; Santschi, 1930b: 76.
    • Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1914e: 38.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 108.
  • lividior. Camponotus maculatus var. lividior Santschi, 1911e: 128 (s.w.q.m.) COMORO IS.
    • Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 88.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
  • lohieri. Camponotus (Myrmoturba) maculatus var. lohieri Emery, 1915g: 22.
    • [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. lohieri Santschi, 1913c: 313 (w.) IVORY COAST; unavailable (infrasubspecific) name.]
    • As unavailable (infrasubspecific) name: Santschi, 1914d: 379; Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
    • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1942: 180; Eidmann, 1944: 463.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 109.
  • manzer. Camponotus maculatus subsp. manzer Forel, 1910e: 452 (s.) ZIMBABWE.
    • Forel, 1911f: 280 (w.); Santschi, 1916b: 509 (m.).
    • Combination in C. (Myrmoturba): Santschi, 1916b: 509;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Forel, 1911f: 280; Santschi, 1916b: 509; Santschi, 1917b: 291; Emery, 1920c: 5; Arnold, 1922: 624; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88; Santschi, 1930b: 76.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • mathildae. Camponotus maculatus subsp. mathildae Forel, 1910c: 266 (s.w.q.) SOUTH AFRICA.
    • Combination in C. (Myrmoturba): Santschi, 1914b: 130;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • As unavailable (infrasubspecific) name: Arnold, 1922: 649.
    • Subspecies of maculatus: Santschi, 1914b: 130; Emery, 1920c: 5; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
  • melanocnemis. Camponotus maculatus var. melanocnemis Santschi, 1911c: 368 (s.) DEMOCRATIC REPUBLIC OF CONGO.
    • Santschi, 1915c: 278 (w.).
    • As unavailable (infrasubspecific) name: Emery, 1920c: 5.
    • Subspecies of maculatus: Forel, 1913h: 355; Santschi, 1913c: 313; Forel, 1914d: 250; Santschi, 1914d: 379; Santschi, 1915c: 278; Santschi, 1920i: 4 (footnote); Wheeler, W.M. 1922a: 234, 956; Emery, 1925b: 87; Weber, 1943c: 333; Bernard, 1953b: 260.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 111.
  • miserabilis. Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379 (s.w.) GUINEA.
    • [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
    • Subspecies of maculatus: Santschi, 1915c: 281; Emery, 1920c: 5; Wheeler, W.M. 1922a: 219, 957; Emery, 1925b: 88.
    • Replacement name: Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126.
  • nubis. Camponotus (Myrmoturba) maculatus subsp. nubis Weber, 1943c: 385, pl. 16, fig. 30 (s.w.) SOUTH SUDAN.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 114.
  • proletaria. Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
    • Replacement name for Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379. [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 118.
  • radamoides Camponotus maculatus var. radamoides Forel, 1891b: 213 (s.w.) MADAGASCAR.
    • Combination in C. (Myrmoturba): Santschi, 1917b: 292;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • As unavailable (infrasubspecific) name: Santschi, 1911e: 131 (caption of fig. 3d).
    • Subspecies of maculatus: Santschi, 1917b: 292; Santschi, 1919b: 239; Emery, 1920c: 5; Santschi, 1920g: 180; Arnold, 1922: 626; Wheeler, W.M. 1922a: 958, 1042; Emery, 1925b: 88; Santschi, 1925h: 166; Arnold, 1926: 292; Santschi, 1930a: 73; Santschi, 1930b: 76; Santschi, 1935a: 279; Santschi, 1937d: 240.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 119.
  • sarmentus. Camponotus (Myrmoturba) maculatus subsp. sarmentus Emery, 1920c: 14, fig. 2 (s.w.) TANZANIA.
    • Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Emery, 1925b: 88.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
  • schereri. Camponotus maculatus subsp. schereri Forel, 1911e: 289 (w.q.) LIBERIA.
    • Combination in C. (Myrmoturba): Santschi, 1915c: 280;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 88.
    • Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1915c: 280; Emery, 1920c: 5; Wheeler, W.M. 1922a: 959; Emery, 1925b: 88.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
  • schultzei. Camponotus maculatus var. schultzei Forel, 1912j: 179 (s.w.m.) UGANDA.
    • Menozzi, 1933a: 110 (q.).
    • As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
    • Subspecies of maculatus: Santschi, 1914b: 130; Menozzi, 1933a: 110; Menozzi, 1939c: 106.
    • Junior synonym of melanocnemis: Forel, 1914d: 250; Emery, 1925b: 87.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
    • schulzi. Camponotus schulzi Salata, Loss, et al. 2019: 95, figs. 1-10, 35 (s.w.) TURKEY.
    • [Note: subgenus C. (Myrmentoma).]
  • semispicatus. Camponotus (Tanaemyrmex) maculatus var. semispicatus Emery, 1925b: 87.
    • [First available use of Camponotus (Myrmoturba) maculatus st. mathildae var. semispicata Santschi, 1914b: 130 (w.) KENYA; unavailable (infrasubspecific) name.]
    • Menozzi, 1930b: 116 (q.).
    • As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
    • Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1930b: 115; Menozzi, 1939c: 107; Menozzi & Consani, 1952: 69; Bernard, 1953b: 261.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 123.
  • sudanicus. Camponotus (Myrmoturba) maculatus subsp. sudanicus Weber, 1943c: 385 (s.w.) SOUTH SUDAN.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 125.
  • thomensis. Camponotus (Myrmoturba) maculatus var. thomensis Santschi, 1920i: 3 (s.w.m.) SÃO TOMÉ & PRÍNCIPE (São Tomé I.).
    • [Misspelled as thomeensis by Santschi, 1939f: 167.]
    • Subspecies of maculatus: Wheeler, W.M. 1922a: 951; Emery, 1925b: 87; Santschi, 1939f: 167.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 127.
  • tuckeri. Camponotus (Myrmoturba) maculatus var. tuckeri Santschi, 1932a: 391 (s.w.m.) NAMIBIA.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 128.
  • zumpti. Camponotus (Myrmoturba) maculatus var. zumpti Santschi, 1937b: 103 (w.) CAMEROON.
    • Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 130.

Taxonomic Notes

Taylor et al. (2018): Camponotus lohieri Santschi, 1913, new status (here, originally described as Camponotus (Myrmoturba) maculatus Fab. stirps melanocnemis Sants. v. lohieri n. var. Santschi, 1913; raised status to Camponotus maculatus F., var. lohieri Sant., Emery, 1915g: 22), CASENT0905268), Haut-Dahomey, DESANTI.

This treatment of C. lohieri as a valid species by Taylor et al. (2018) was proposed without comment or justification, has has not been accepted by subsequent authors.

Taylor et al. (2018) treat Camponotus proletaria as a full, distinct species but provide no justification for the change. Subsequent authors have not followed this treatment.

Description

References

References based on Global Ant Biodiversity Informatics

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