Acropyga acutiventris

AntWiki: The Ants --- Online
Acropyga acutiventris
Acropyga acutiventris
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Plagiolepidini
Genus: Acropyga
Species group: acutiventris
Species: A. acutiventris
Binomial name
Acropyga acutiventris
Roger, 1862

Acropyga acutiventris side view

Acropyga acutiventris top view

Synonyms

A common wide-ranging species, A. acutiventris is known to occur in rainforest and, in Australia, savanna woodlands. Nests are located in rotten wood on the ground, under stones, and in the leaf litter. This species is a trophophoretic ant that has been found to be associated with two species of mealybugs (Xenococcus acropygae and X annandalei). Queens carry live mealbugs in their mandibles on their nuptual flights, with the mealybugs providing honeydew for her newly established nest.

Photo Gallery

  • A. acutiventris worker carrying a mealybug. From Sakaerat, Korat province, Thailand. (Image by Christian Peeters)
  • An Acropyga acutiventris worker ant pulls a cocoon to safety after the photographer disturbs her nest. Cape Tribulation, Queensland, Australia. (Image by Alex Wild.)
  • Acropyga acutiventris dealate queen with mealybug. Photo by Manoj Vembayam.
  • Acropyga acutiventris queen and male from a nest in captivity. Photo by Dulneth Wijewardana.

Identification

LaPolla (2004) - A member of the Acropyga acutiventris species group. Worker: 11 segmented antennae; mandible with 5 teeth and distinct ridges that run along dorsal surface; many erect hairs on head, mesosoma and gaster giving ant a "spiky appearance"; eyes prominent. Queen: As in worker with modifications expected for caste. Male: 12 segmented antennae; parameres with dense covering of erect hairs and with dorsolateral expansions medially. Compare with Acropyga rubescens.

This species is fairly easy to recognize, with the only morphologically similar species being its close relative, Acropyga rubescens. A combination of large eyes (more than 20 facets), unique mesosomal shape, and 5-toothed mandibles with distinct striate sculpture on the dorsal surface, enable identification of A. acutiventris. Separating workers of this species from A. rubescens can be more difficult, though in general it is not as hairy as in A. rubescens, is slightly smaller in total length, and the striate sculpture extends approximately across the entire dorsal surface of the mandible. Males of A. acutiventris are also less hairy than in A. rubescens, but the most reliable means to identify them is by dissecting out the penis valves, which are distinct for each species. Given the large range and morphological variability of this species, it is not surprising that so many synonyms have been created. In the past, species have been based on variation in color, overall size, head shape differences and antennal segment shape differences. Here, they are all treated as being the same of a wide ranging, variable species. Dissection of males from across the range (penis valve dissections from Australia, India, Papua New Guinea, Singapore, and the Solomon Islands) of A. acutiventris revealed no discernable genitalic differences.

Australia

Within Australia this species can be separated from other members of the genus by the presence of long erect hairs on the antennal scapes and posterior margin of the head.

Keys including this Species

Distribution

Acropyga acutiventris has a large range that occupies much of the Australasian, Indo-Australian, and Oriental regions. It can be found from India east to northern Australia. It is one of the most commonly encountered species in the genus.

Australia

Limited to the Top End and northern Queensland where it occurs primarily in rainforest with a few records from savanna woodlands.

Latitudinal Distribution Pattern

Latitudinal Range: 30.3416° to -21.08333015°.

   
North
Temperate 		North
Subtropical 		Tropical South
Subtropical 		South
Temperate

Distribution based on Regional Taxon Lists

Australasian Region: Australia (type locality).
Indo-Australian Region: Borneo, Indonesia, Krakatau Islands, Malaysia, New Guinea, Philippines, Singapore, Solomon Islands.
Oriental Region: India, Nepal, Nicobar Island, Sri Lanka (type locality), Thailand, Vietnam.
Palaearctic Region: China.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
pChart

Elevation Range

Occurrence at collecting sites during elevational surveys of rainforest in the Eungella region, Queensland, Australia (Burwell et al., 2020).
Species Elevation (m asl)
200 400 600 800 1000 1200
Acropyga acutiventris 30-40
Shading indicates the bands of elevation where species was recorded.
Numbers are the percentage of total samples containing this species.

Biology

Association with Other Organisms

Explore-icon.png Explore: Show all Associate data or Search these data. See also a list of all data tables or learn how data is managed.
  • This species is a prey for the phorid fly Puliciphora boltoni (a predator) (Quevillon, 2018).
  • This species is a prey for the phorid fly Puliciphora boltoni (a predator) (Quevillon, 2018).

Castes

Worker

A. acutiventris hd.jpgA. acutiventris sd.jpgA. acutiventris th.jpgA. acutiventris lb.jpg
. Owned by Museum of Comparative Zoology.

Phylogeny

Acropyga

Acropyga ayanganna

Acropyga donisthorpei

Acropyga panamensis

Acropyga guianensis

Acropyga stenotes

Acropyga fuhrmanni

Acropyga smithii

Acropyga romeo

Acropyga hirsutula

Acropyga dubitata

Acropyga decedens, Acropyga goeldii

Acropyga epedana

Acropyga arnoldi

Acropyga silvestrii

Acropyga kinomurai

Acropyga lauta

Acropyga sauteri

Acropyga acutiventris

Acropyga myops

Acropyga butteli

Acropyga ambigua

Acropyga pallida

Based on Blaimer et al., 2016. Note only selected Acropyga species are included, and undescribed species are excluded.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • acutiventris. Acropyga acutiventris Roger, 1862a: 243 (w.q.) SRI LANKA.
    • Type-material: syntype workers, syntype queens (numbers not stated).
    • Type-locality: “probably Sri Lanka” (“Ceylon”): (no further data).
    • Type-depository: unknown (LaPolla, 2004a: 31).
    • [Note: formicid type-material from Roger’s collections is present in DEIB, MNHN, MNHU, ZSBS, plus isolated examples in other museums.]
    • Imai, et al. 1984: 68 (k.).
    • Combination in Plagiolepis: Mayr, 1863: 442,
    • combination in Acropyga: Roger, 1863b: 11.
    • Status as species: Mayr, 1862: 769; Roger, 1863b: 11; Mayr, 1863: 442; Smith, F. 1871a: 319; Emery, 1888a: 530; Dalla Torre, 1893: 174; Forel, 1894c: 418; Mayr, 1897: 432; Emery, 1900d: 697; Emery, 1901f: 121; Emery, 1901g: 567; Bingham, 1903: 333; Forel, 1903d: 407; Forel, 1909e: 395; Forel, 1911e: 286; Forel, 1913e: 663; Emery, 1925b: 28; Wheeler, W.M. 1929g: 57; Karavaiev, 1933a: 308; Chapman & Capco, 1951: 210; Taylor, 1992a: 58; Bolton, 1995b: 57; Terayama, Fellowes & Zhou, 2002: 23 (redescription); LaPolla, 2004a: 31 (redescription); Jaitrong & Nabhitabhata, 2005: 10; Mohanraj, et al. 2010: 6; Pfeiffer, et al. 2011: 36; Ran & Zhou, 2011: 65; Guénard & Dunn, 2012: 27; Sarnat, et al. 2013: 69; Bharti, Guénard, et al. 2016: 23; Dias, R.K.S. et al. 2020: 28; Khachonpisitsak, et al. 2020: 38; Wang, W.Y., Soh, et al. 2022: 35.
    • Senior synonym of australis: Taylor, 1992a: 58; Bolton, 1995b: 57; Terayama, Fellowes & Zhou, 2002: 23; LaPolla, 2004a: 31.
    • Senior synonym of carinata: LaPolla, 2004a: 31.
    • Senior synonym of crassicornis: LaPolla, 2004a: 31.
    • Senior synonym of flava: Forel, 1894c: 418; Bolton, 1995b: 57; Terayama, Fellowes & Zhou, 2002: 23; LaPolla, 2004a: 31.
    • Senior synonym of indosinensis: LaPolla, 2004a: 31.
    • Senior synonym of javana: LaPolla, 2004a: 31.
    • Senior synonym of moluccana: LaPolla, 2004a: 31.
    • Senior synonym of mysolensis: LaPolla, 2004a: 31.
    • Senior synonym of occipitalis: LaPolla, 2004a: 31.
    • Senior synonym of opaca: LaPolla, 2004a: 31.
    • Senior synonym of papuana: LaPolla, 2004a: 31.
    • Senior synonym of undecemus: LaPolla, 2004a: 31.
    • Distribution
    • Austral: Australia.
    • Malesian: Indonesia (Flores, Irian Jaya, Java, Kalimantan, Misool, Seram, Sulawesi, Sumatra), Malaysia (Peninsula, Sabah, Sarawak), Papua New Guinea, Singapore, Solomon Is.
    • Oriental: China, India (+ Andaman Is, Nicobar Is), Myanmar, Sri Lanka, Thailand, Vietnam.
  • australis. Acropyga moluccana var. australis Forel, 1902h: 477 (w.) AUSTRALIA (Queensland).
    • Type-material: syntype workers (number not stated).
    • [Note: LaPolla, 2004a: 31, cites 7w syntypes BMNH.]
    • Type-locality: Australia: Queensland, Mackay (G. Turner).
    • Type-depositories: ANIC, BMNH, MHNG.
    • Wheeler, G.C. & Wheeler, J. 1968: 209 (l.).
    • Subspecies of acutiventris: Forel, 1911e: 286; Emery, 1925b: 28.
    • Subspecies of moluccana: Forel, 1910b: 58; Taylor & Brown, 1985: 108; Taylor, 1987a: 5.
    • Junior synonym of acutiventris: Taylor, 1992a: 58; Bolton, 1995b: 57; Terayama, Fellowes & Zhou, 2002: 23; LaPolla, 2004a: 31.
  • carinata. Acropyga acutiventris var. carinata Karavaiev, 1933a: 310 (w.) INDONESIA (Sumatra).
    • Type-material: syntype workers (number not stated, “a few”).
    • Type-locality: Indonesia: Sumatra, Payacombo, 25.iii.1913, no. 5360 (O. John).
    • Type-depository: unknown (not in SIZK (Radchenko, Fisher, et al. 2023: 10)).
    • Subspecies of acutiventris: Chapman & Capco, 1951: 210; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 32.
  • crassicornis. Acropyga crassicornis Emery, 1900d: 698 (footnote), fig. 13 (w.) NEW GUINEA (no state data).
    • Type-material: syntype workers (number not stated).
    • [Note: LaPolla, 2004a: 31, cites 1w NHMB as the holotype, but this is unlikely as the material was collected by Biró and described by Emery, which implies syntypes in HNHM, MSNG, or both.]
    • Type-locality: New Guinea (probably Papua New Guinea): (no further data) (L. Biró).
    • Type-depository: NHMB (perhaps also HNHM, MSNG).
    • Status as species: Forel, 1911e: 286; Viehmeyer, 1914c: 530; Emery, 1925b: 28; Chapman & Capco, 1951: 211; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • flava. Plagiolepis flava Mayr, 1862: 699 (w.m.) INDIA (Nicobar Is).
    • Type-material: syntype workers (number not stated), 1 syntype male.
    • Type-locality: India: Nicobar Is, Milu I., (Novara Expd.) (no collector’s name) (sent by R. von Frauenfeld).
    • Type-depository: NHMW.
    • Combination in Acropyga: Mayr, 1862: 769.
    • Status as species: Roger, 1863b: 11; Mayr, 1863: 442; Mayr, 1865: 53 (redescription); Emery, 1889b: 510; Dalla Torre, 1893: 174; Emery, 1895k: 476.
    • Subspecies of acutiventris: Forel, 1913k: 100; Emery, 1925b: 28; Wheeler, W.M. 1935g: 38; Chapman & Capco, 1951: 210.
    • Junior synonym of acutiventris: Forel, 1894c: 418; Bolton, 1995b: 57; Terayama, Fellowes & Zhou, 2002: 23; LaPolla, 2004a: 31.
  • indosinensis. Acropyga (Rhizomyrma) indosinensis Wheeler, W.M. 1935c: 72.
    • Replacement name for Acropyga (Rhizomyrma) silvestrii Wheeler, W.M. 1927h: 100. [Junior primary homonym of Acropyga silvestrii Emery, 1915g: 21.]
    • Status as species: Chapman & Capco, 1951: 212; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 32.
  • javana. Acropyga (Acropyga) acutiventris var. javana Karavaiev, 1933a: 308 (w.m.) INDONESIA (Java).
    • Type-material: syntype workers, syntype males (numbers not stated, “numerous”).
    • [Note: Martynov & Radchenko, 2016: 247, cite 5w syntypes NMAS (plus 211w in alcohol).]
    • Type-localities: Indonesia: Java, Depok, Djungeln, 22.xii.1912, no. 2344 (W. Karawajew), Java, Buitenzorg (= Bogor), Botanical Garden, no. 2370 (W. Karawajew).
    • Type-depository: NMAS (perhaps also SIZK).
    • Subspecies of acutiventris: Chapman & Capco, 1951: 211; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • moluccana. Acropyga moluccana Mayr, 1879: 658 (w.) INDONESIA (Seram).
    • Type-material: holotype (?) worker.
    • [Note: no indication of number of specimens is given.]
    • Type-locality: Indonesia: Ceram (no collector’s name) (received from F. Smith).
    • Type-depository: NHMW.
    • Karavaiev, 1933a: 310 (m.); Wheeler, G.C. & Wheeler, J. 1953c: 137 (l.).
    • Status as species: Emery, 1887a: 247; Dalla Torre, 1893: 174; Emery, 1900d: 697 (footnote); Forel, 1901b: 24; Dahl, 1901: 12; Forel, 1905c: 24; Emery, 1925b: 29; Karavaiev, 1933a: 310; Chapman & Capco, 1951: 211; Taylor & Brown, 1985: 107; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • mysolensis. Acropyga moluccana subsp. mysolensis Forel, 1911e: 285 (w.) INDONESIA (Misool).
    • Type-material: syntype workers (number not stated).
    • Type-locality: Indonesia: Mysol (H.W. Bates).
    • Type-depositories: MHNG, ZSBS.
    • Subspecies of moluccana: Chapman & Capco, 1951: 211; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • occipitalis. Acropyga moluccana var. occipitalis Stitz, 1912: 510, fig. 12 (w.) NEW GUINEA (no state data).
    • Type-material: holotype worker.
    • Type-locality: New Guinea: (no further data) (Kaiserin Augustafluss Expd.) (Bürgers).
    • Type-depository: MNHU.
    • Subspecies of moluccana: Chapman & Capco, 1951: 211; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • opaca. Acropyga moluccana var. opaca Stitz, 1911a: 370 (w.) NEW GUINEA (Papua New Guinea).
    • Type-material: 5 syntype workers.
    • Type-locality: Papua New Guinea (“Neu-Guinea (Germaniahuk)”): (no further data) (Schultze).
    • Type-depository: MNHU.
    • Donisthorpe, 1948g: 139 (m.).
    • Subspecies of moluccana: Mann, 1919: 365; Wheeler, W.M. 1935g: 38; Donisthorpe, 1948g: 139; Chapman & Capco, 1951: 211; Bolton, 1995b: 57.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • papuana. Acropyga (Acropyga) moluccana subsp. papuana Mann, 1919: 364 (w.) SOLOMON IS (Ugi).
    • Type-material: syntype workers (number not stated, “very common”).
    • [Note: LaPolla, 2004a: 31, cites 7w syntypes MCZC, USNM, plus 3 other specimens BMNH that are not from the type-series.]
    • Type-locality: Solomon Is: Ugi I., Pawa, 19.v.-24.xi.1916 (W.M. Mann).
    • Type-depositories: MCZC, USNM.
    • Wheeler, G.C. & Wheeler, J. 1953c: 136 (l.).
    • Subspecies of moluccana: Wheeler, W.M. 1935g: 38; Bolton, 1995b: 58.
    • Junior synonym of acutiventris: LaPolla, 2004a: 31.
  • silvestrii. Acropyga (Rhizomyrma) silvestrii Wheeler, W.M. 1927h: 100, fig. 7 (w.) VIETNAM.
    • Type-material: 3 syntype workers.
    • Type-locality: Vietnam (“Indochina”): Coxan (F. Silvestri).
    • Type-depository: MCZC.
    • [Junior primary homonym of Acropyga silvestrii Emery, 1915g: 21.]
    • Replacement name: Acropyga (Rhizomyrma) indosinensis Wheeler, W.M. 1935c: 72.
  • undecemus. Pseudolasius undecemus Donisthorpe, 1949c: 756 (w.) NEW GUINEA (Indonesia, Papua New Guinea).
    • Type-material: 5 syntype workers.
    • Type-localities: 1 worker Indonesia: Irian Jaya (“Dutch New Guinea”): Maffin Bay, 10.vi.1944 (E.S. Ross), 4 workers Papua New Guinea (“New Guinea”): Finschhafen, 27.iv.1944 (E.S. Ross).
    • Type-depositories: BMNH, CASC.
    • Combination in Acropyga: Bolton, 1995b: 58.
    • Status as species: Chapman & Capco, 1951: 207; Bolton, 1995b: 58.
    • Junior synonym of acutiventris: LaPolla, 2004a: 32.

Type Material

The following notes on V.A. Karavaiev type specimens have been provided by Martynov & Radchenko (2016) based on material held in the Institute of Zoology, Ukrainian Academy of Science:

Acropyga acutiventris var. javana

Type locality(-ies): “Depok, W. Java, Djungeln, 22.XII.1912 (Nr. 2344), zahlreiche ww von 2,5-3 mm Länge. Kolonie im Boden zwischen den Wurzeln eines großen Baumes, in der Nachbarschaft einer Termitenkolonie.— Bujtenzorg, Java, Botan. Garten (Nr. 2370), zahlreiche ww von 3-3,5 mm Länge und mm. Koloniein der Erde”.

Original Karawajew label(-s): “Acrotyga (Acropyga) / acutiventris v. javana Kar. / Buitenzorg, 2370.”.

Material: SYNTYPES, 5 w (newly mounted, IN 314/6/1), 211 w (in alcohol, IN 314/BAMS1/Karaw61).

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Worker

LaPolla (2004) - (n=l0): TL: 2.67-3.53; HW: 0.8-0.912; HL: 0.803-0.896; SL: 0.628-0.827; ML: 0.836-1.03; GL: 0.933-1.74; CI: 91.07-104.25; SI: 78.7-97.97.

Head: yellow to light brownish-yellow; head about as broad as long; posterior margin concave; covered in a thick layer of appressed hairs, with many scattered erect hairs, especially along posterior margin; eyes relatively large, placed at lower 1/4 of head; 11 segmented, incrassate antennae; scape reaches or surpasses posterior margin by about length of pedicel; clypeus broad, slightly convex medially with many erect hairs, the longest ones along anterior clypeal margin; mandible with 5 uneven teeth; 3rd and 5th tooth (from apical to basal) smaller than others; dorsal surface of mandible with distinct ridges across surface from about base of teeth toward insertion of mandibles; slight gap present between inner mandibular surface and anterior clypeal margin. Mesosoma: yellow to light brownish-yellow; pronotum in lateral view typically with short shelf before rising sharply toward mesonotum; mesonotum rounded with many erect and appressed hairs at apex; metanotal area distinct; propodeum below level of mesonotum, dorsum flat before steep declivity; propodeum with short erect and appressed hairs. Gaster: yellow to light brownish-yellow to reddish-yellow; petiole thick and erect reaching height of propodeum; yellow to light brownish-yellow; gaster covered in thick layer of appressed hairs with many scattered erect hairs throughout giving a "spiky" appearance.

Queen

LaPolla (2004) - (n=3): TL: 4.56-5.09; HW: 1.02-1.06; HL: 0.967-1.01; SL: 0.902-0.971; ML: 1.4-1.63; GL: 2.19-2.63; CI: 102.97-106; SI: 85.09-95.2. As in worker with modifications expected for caste.

Male

LaPolla (2004) - (n=3): TL: 2.89-3.33; HW: 0.604-0.644; HL: 0.588-0.626; SL: 0.529-0.617; ML: 1.03-l. l6; GL: 1.27-1.56; CI: 102.72-103.09; SI: 87.58-95.81.

Head: yellow to light brownish-yellow, darker around three prominent ocelli; head about as broad as long; covered in layer of appressed hairs with scattered suberect to erect hairs; 12 segmented, slightly incrassate antennae; scape surpasses posterior margin by about length of first two funicular segments; clypeus broad, relatively flat, with scattered short erect hairs; mandible with 3-4 teeth; gap present between inner mandibular margin and anterior clypeal margin. Mesosoma: yellow to light brownish-yellow; pronotum short and collar-like; mesonotum large, rounded anteriorly; mesonotum flat dorsally, with layer of shorter appressed hairs and scattered longer erect hairs throughout; propodeum lower than mesonotum and scutellum; declivity not distinct from propodeum. Gaster: petiole thick and erect; gaster yellow to light brownish-yellow to reddish-yellow, darker dorsally; covered in dense layer of appressed hairs with scattered erect hairs throughout. Genitalia: in lateral view parameres thick, tapering to rounded apices; parameres with medial dorsolateral extensions; cuspi short, bent toward approximately middle of digiti, where cuspi meet digiti with short, peg-like teeth at apices; digiti long and erect; apex rounded; digiti with short-peg-like teeth where cuspi meet.

Karyotype

Explore-icon.png Explore: Show all Karyotype data or Search these data. See also a list of all data tables or learn how data is managed.
  • 2n = 28; 29 (Malaysia) (Imai et al., 1983) (suggested Robertsonian polymorphism).

References

References based on Global Ant Biodiversity Informatics

  • Abe T., S. Yamane, and K. Onoyama. Ants collected on the Krakatau Islands 100 years after the great eruptions. Biogeography 14: 65-75.
  • Andersen A. N., J. C. Z. Woinarski, and B. Hoffman. 2004. Biogeography of the ant fauna of the Tiwi Islands, in northern Australia's moonsoonal tropics. Australian Journal of Zoology 52: 97-110.
  • Andersen A. N., M. Houadria, M. Berman, and M. van der Geest. Rainforest ants of the Tiwi Islands: a remarkable centr of endemism in Australia's monsoonal tropics. Insectes Sociaux 59: 433-441.
  • Andersen, Alan N., John C.Z. Woinarski and Ben D. Hoffman. 2004. Biogeography of the ant fauna of the Tiwi Islands, in northern Australia's monsoonal tropics. Australian Journal of Zoology 52: 97-110.
  • Asfiya W., L. Lach, J. D. Majer, B. Heterick, and R. K. Didham. 2015. Intensive agroforestry practices negatively affect ant (Hymenoptera: Formicidae) diversity and composition in southeast Sulawesi, Indonesia. Asian Myrmecology 7: 87-104.
  • Blaimer B. B., J. S. LaPolla, M. G. Branstetter, M. W. Lloyd, and S. G. Brady. 2016. Phylogenomics, biogeography and diversification of obligate mealybug-tending ants in the genus Acropyga. Molecular Phylogenetics and Evolution 102: 20-29.
  • CSIRO Collection
  • Chapman J. W. 1965. Studies on the ecology of the army ants of the Philippines genus Aenictus Schuckard (Hymenoptera: Formicidae). Philippine Journal of Science. 93: 551-595.
  • Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
  • Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
  • Dahl F. 1901. Das Leben der Ameisen im Bismarck-Archipel, nach eigenen Beobachtungen vergleichend dargestellt. Mitt. Zool. Mus. Berl. 2: 1-70.
  • Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
  • Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
  • Dias R. K. S. 2013. Diversity and importance of soil-dweeling ants. Proceedings of the National Symposium on Soil Biodiversity, chapt 4, pp 19-22.
  • Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
  • Dias R. K. S., and H. A. W. S. Peiris. 2015. Ground-dwelling ant assemblages (Family: Formicidae) in six coconut (Cocos nucifera L. 1753) plantations in Sri Lanka. Journal ofInsect Biodiversity 3(14): 1–10. http://dx.doi.org/10.12976/jib/2015.3.14
  • Dias R. K. S., and K. R. K. Anuradha Kosgamage. 2012. Occurrence and species diversity of ground-dwelling worker ants (Family: Formicidae) in selected lands in the dry zone of Sri Lanka. J. Sci. Univ. Kelaniya 7: 55-72.
  • Dias R. K. S., and R. P. K. C. Rajapaksa. 2016. Geographic records of subfamilies, genera and species of ants (Hymenoptera: Formicidae) in the four climatic zones of Sri Lanka: a review. J. Sci. Univ. Kelaniya 11(2): 23-45.
  • Dias R. K. S.; Perera K. A. M. 2011. Worker ant community observed by repeated sampling and information on endemic Aneuretus simoni Emery in the Gilimale Forest Reserve in Sri Lanka. Asian Myrmecology 4: 69-78.
  • Dias, R.K.S. 2006. Overview of ant research in Sri Lanka: 2000-2004. ANeT Newsletter 8:7-10
  • Donisthorpe H. 1948. A fourth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)1: 131-143.
  • Donisthorpe H. 1949. A sixth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)1: 744-759.
  • Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia. [part]. Ann. Mus. Civ. Stor. Nat. 24(4): 209-258.
  • Emery C. 1888. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza . Formiche raccolte dal sig. Elio Modigliani in Sumatra e nell'isola Nias. Annali del Museo Civico di Storia Naturale 25: 528-534.
  • Emery C. 1901. Ameisen gesammelt in Ceylon von Dr. W. Horn 1899. Deutsche Entomologische Zeitschrift 1901: 113-122.
  • Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 14:565-580.
  • Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
  • Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 4, no. 24 (1887): 209-258.
  • Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
  • Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. J. Bombay Nat. Hist. Soc. 8: 396-420.
  • Forel A. 1901. Formiciden aus dem Bismarck-Archipel, auf Grundlage des von Prof. Dr. F. Dahl gesammelten Materials. Mitt. Zool. Mus. Berl. 2: 4-37.
  • Forel A. 1903. Les fourmis des îles Andamans et Nicobares. Rapports de cette faune avec ses voisines. Rev. Suisse Zool. 11: 399-411.
  • Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
  • Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
  • Forel A. 1911. Die Ameisen des K. Zoologischen Museums in München. Sitzungsber. Math.-Phys. Kl. K. Bayer. Akad. Wiss. Münch. 11: 249-303.
  • Forel A. 1913. Quelques fourmis des Indes, du Japon et d'Afrique. Rev. Suisse Zool. 21: 659-673
  • Forel A. 1913k. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
  • Gumawardene, N.R., J.D. Majer and J.P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2:71-83
  • Gunawardene N. R., J. D. Majer, and J. P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2: 71-83.
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
  • Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
  • Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
  • Janda M., G. D. Alpert, M. L. Borowiec, E. P. Economo, P. Klimes, E. Sarnat, and S. O. Shattuck. 2011. Cheklist of ants described and recorded from New Guinea and associated islands. Available on http://www.newguineants.org/. Accessed on 24th Feb. 2011.
  • Janda M., P. Matos-Maravi, M. Borovanska, J. Zima Jr., E. Youngerman, and N. E. Pierce. 2016. Phylogen y and population genetic structure of the ant genus Acropy ga (Hymenoptera : Formicidae) in Papua New Guinea. Invertebrate Systematics 30: 28-40.
  • Karavaiev V. 1933. Ameisen aus dem Indo-Australischen Gebiet, VII. Konowia 11: 305-320.
  • LaPolla J.S. 2004. Acropyga (Hymenoptera: Formicidae) of the world. Contributions of the American Entomological Institute 33(3): 1-130.
  • LaPolla, J. S. 2004. Acropyga (Hymenoptera: Formicidae) of the world. Contributions of the American Entomological Institute 33(3): 1-130.
  • Latumahina F., M. Borovanska, N. S. Putra, and M. Janda. 2015. Ants of Ambon Island – diversity survey and checklist. ZooKeys 472: 43–57.
  • Leong C. M., S. F. Shiao, and B. Guenard. 2017. Ants in the city, a preliminary checklist of Formicidae (Hymenoptera) in Macau, one of the most heavily urbanized regions of the world. Asian Myrmecology 9: e009014.
  • Lucky A., E. Sarnat, and L. Alonso. 2011. Ants of the Muller Range, Papua New Guinea, Chapter 10. In Richards, S. J. and Gamui, B. G. (editors). 2013. Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: surveying the biodiversity of Papua New Guinea’s sublime karst environments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA.
  • Lucky A., K. Sagata, and E. Sarnat. 2011. Ants of the Nakanai Mountains, East New Britain Province, Papua New Guinea, Chapter 1. In Richards, S. J. and Gamui, B. G. (editors). 2013. Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: surveying the biodiversity of Papua New Guinea’s sublime karst environments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA.
  • Lucky A., L. E. Alonso, E. Sarnat, and J. Hulr. 2015. Ants and scolytine beetles. In: Richards, S.J. and N. Whitmore (editors) 2015. A rapid biodiversity assessment of Papua New Guinea's Hindenburg Wall region. Wildlife Conservation Society Papua New Guinea Program. Goroka, PNG.
  • Mann W. M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology 63:273-391.
  • Mann William. 1916. The Ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology at Harvard College 63(7): 273-391
  • Mann, W.M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology of Harvard College 63: 273-391
  • Martynov A. V., and A. G. Radchenko. 2016. Karawajew’s ant type specimens (Hymenoptera, Formicidae) in the National Museum of Natural History of the National Academy of Sciences of Ukraine. Zootaxa 4097 (2): 244–254.
  • Mohamed M. 1995. A preliminary list of ants (Hymenoptera: Formicidae) of the Tawau Hills Park, Sabah. In: Ghazally Ismail et al. (eds.), Tawau Hills Park, Sabah, Pelanduk Pub. Pp. 205-213.
  • Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
  • Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
  • Ogata K. 2005. Asian ant inventory and international networks. Report on Insect inventory Project in Tropic Asia TAIIV: 145-170.
  • Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
  • Ran H., and S. Y. Zhou. 2011. Checklist of Chinese Ants: the Formicomorph Subfamilies (Hymenoptera: Formicidae) (I). Journal of Guangxi Normal University: Natural Science Edition. 29(3): 65-73.
  • Rizali A., M. M. Bos, D. Buchori, Sk. Yamane, and C. H. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 15(2): 77-84.
  • Rizali A., M.M. Bos, D. Buchori, Sk. Yamane, C. Hans, and J. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 77-84.
  • Snelling R. R. 1998. Insect Part 1: The social Hymenoptera. In Mack A. L. (Ed.) A Biological Assessment of the Lakekamu Basin, Papua New Guinea, RAP 9. 189 ppages
  • Stitz H. 1911. Australische Ameisen. (Neu-Guinea und Salomons-Inseln, Festland, Neu-Seeland). Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1911: 351-381.
  • Stitz H. 1912. Ameisen aus Ceram und Neu-Guinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1912: 498-514.
  • Taylor R. W. 1987. A checklist of the ants of Australia, New Caledonia and New Zealand (Hymenoptera: Formicidae). CSIRO (Commonwealth Scientific and Industrial Research Organization) Division of Entomology Report 41: 1-92.
  • Taylor R. W. 1992. Nomenclature and distribution of some Australian and New Guinean ants of the subfamily Formicinae (Hymenoptera: Formicidae). Journal of the Australian Entomological Society 31: 57-69.
  • Terayama M.; Fellowes, J. R.; Zhou, S. 2002. The East Asian species of the ant genus Acropyga Roger, 1862 (Hymenoptera: Formicidae: Formicinae). Edaphologia 70:21-32. 
  • Varghese T. 2004. Taxonomic studies on ant genera of the Indian Institute of Science campus with notes on their nesting habits. Pp. 485-502 in : Rajmohana, K.; Sudheer, K.; Girish Kumar, P.; Santhosh, S. (eds.) 2004. Perspectives on biosystematics and biodiversity. Prof. T.C. Narendran commemoration volume. Kerala: Systematic Entomology Research Scholars Association, xxii + 666 pp.
  • Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
  • Viehmeyer H. 1914. Papuanische Ameisen. Deutsche Entomologische Zeitschrift 1914: 515-535.
  • Watanasit S., and N. Noon-anant. 2005. Ants at Ton Nga Chang Wildlife Sanctuary, Songkhla. Sonklanakarin J. Sci. Technolo. 27(2): 267-280.
  • Wheeler W. M. 1927. Ants collected by Professor F. Silvestri in Indochina. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 20: 83-106.
  • Wheeler W. M. 1929. Ants collected by Professor F. Silvestri in Formosa, the Malay Peninsula and the Philippines. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 24: 27-64.
  • Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
  • Wheeler, William Morton.1935.Checklist of the Ants of Oceania.Occasional Papers 11(11): 3-56
  • Wilson E. O. 1959. Some ecological characteristics of ants in New Guinea rain forests. Ecology 40: 437-447.
  • Woodcock P., D. P. Edwards, T. M. Fayle, R. J. Newton, C. Vun Khen, S. H. Bottrell, and K. C. Hamer. 2011. The conservation value of South East Asia's highly degraded forests: evidence from leaf-litter ants. Phil. Trans. R. Soc. B. 366: 3256-3264.
  • Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
Retrieved from "https://antwiki.org/wiki/index.php?title=Acropyga_acutiventris&oldid=720026"