Polyrhachis illaudata

Polyrhachis illaudata
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Formicinae
Tribe:	Camponotini
Genus:	Polyrhachis
Subgenus:	Myrma
Species:	P. illaudata
Binomial name
Polyrhachis illaudata Walker, 1859 Specimen Labels
Subspecies
Polyrhachis illaudata intermedia Forel, 1886 Polyrhachis illaudata pauperata Emery, 1889 Polyrhachis illaudata proximomayri Emery, 1896
Synonyms
Polyrhachis duodentata Donisthorpe, 1942 Polyrhachis latispinosa Donisthorpe, 1942 Polyrhachis mayri Roger, 1863

Photo Gallery

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  Polyrhachis illaudata worker from Malabar WLS, India. Photo by Kalesh Sadasivan.
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  Polyrhachis illaudata worker from Ponmudi, India. Photo by Kalesh Sadasivan.
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  Polyrhachis illaudata worker returning to its nest in rotten wood with a prey item. Western Ghats, Kerala, India, 1000m (photo by Dr Kalesh Sadasivan).
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  Polyrhachis illaudata alate queen. Dehradun, Uttarakhand, India. Photo by Yathumon M A.

Identification

Keys including this Species

• Key to Polyrhachis of India

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 32.812778° to -6°.

North Temperate	North Subtropical	Tropical	South Subtropical	South Temperate

• Source: AntMaps

Distribution based on Regional Taxon Lists

Indo-Australian Region: Borneo, Indonesia, Krakatau Islands, Malaysia, Philippines, Singapore.
Oriental Region: Bangladesh, India, Laos, Nepal, Sri Lanka (type locality), Thailand.
Palaearctic Region: China.

Distribution based on AntMaps

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.

Biology

Association with Other Organisms

Diptera

• This species is a host for the phorid fly Megaselia holosericei (a parasite) (Brown et al., 2015) (injured).
• This species is a host for the milichiid fly Milichia sp. (a myrmecophile) in Malaysia (Yusah & Fayle, 2014; Milichiidae online).

Fungi

• This species is a host for the fungus Ophiocordyceps formicarum (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
• This species is a host for the fungus Ophiocordyceps irangiensis (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
• This species is a host for the fungus Ophiocordyceps myrmecophila (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
• This species is a host for the fungus Ophiocordyceps unilateralis (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
• This species is a host for the fungus Ophiocordyceps unilateralis (a pathogen) in Taiwan (Lin et al., 2020).

Castes

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  Liu, C. et al. 2020. Ants of the Hengduan Mountains, Figure 47, Polyrhachis illaudata.

.	Owned by Museum of Comparative Zoology.

Images from AntWeb

Holotype of Polyrhachis illaudata. Worker. Specimen code casent0903443. Photographer Z. Lieberman, uploaded by California Academy of Sciences.	Owned by NHMUK, London, UK.

Holotype of Polyrhachis duodentata. Queen (alate/dealate). Specimen code casent0903444. Photographer Z. Lieberman, uploaded by California Academy of Sciences.	Owned by NHMUK, London, UK.

Holotype of Polyrhachis latispinosa. Queen (alate/dealate). Specimen code casent0903445. Photographer Z. Lieberman, uploaded by California Academy of Sciences.	Owned by NHMUK, London, UK.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

• illaudata. Polyrhachis illaudatus Walker, 1859: 373 (w., not m.) SRI LANKA. Imai, Brown, et al. 1984: 68 (k.). Combination in P. (Myrma): Donisthorpe, 1932b: 576. Senior synonym of mayri: Donisthorpe, 1932b: 576; of duodentata, latispinosa: Bolton, 1974b: 176. Current subspecies: nominal plus intermedia, pauperata, proximomayri.
• mayri. Polyrhachis mayri Roger, 1863b: 7 (w.) SRI LANKA. Bingham, 1903: 405 (q.); Karavaiev, 1927e: 46 (m.). Combination in P. (Myrma): Viehmeyer, 1916a: 165. Junior synonym of illaudata: Donisthorpe, 1932b: 576.
• duodentata. Polyrhachis (Myrma) duodentata Donisthorpe, 1942d: 461, pl. 2, fig. 2 (q.) INDIA. Junior synonym of illaudata: Bolton, 1974b: 176.
• latispinosa. Polyrhachis (Myrma) latispinosa Donisthorpe, 1942d: 460, pl. 2, fig. 1 (q.) INDIA. Junior synonym of illaudata: Bolton, 1974b: 176.

Description

Karyotype

• n = 14, 2n = 28 (Indonesia) (Imai et al., 1985).
• n = 18 (Malaysia) (Imai et al., 1983).

References

• Baidya, P., Bagchi, S. 2021. Influence of human land use and invasive species on beta diversity of tropical ant assemblages. Insect Conservation and Diversity, icad.12536 (doi:10.1111/icad.12536).
• Baltazar, C.R. 1966. A catalogue of Philippine Hymenoptera (with a bibliography, 1758-1963). Pacific Insects Monographs 8: 1-488. (page 276, listed)
• Bolton, B. 1974b [1973]. New synonymy and a new name in the ant genus Polyrhachis F. Smith (Hym., Formicidae). Entomol. Mon. Mag. 109: 172-180 (page 176, Senior synonym of duodentata and latispinosa)
• Brassard, F., Leong, C.-M., Chan, H.-H., Guénard, B. 2021. High diversity in urban areas: How comprehensive sampling reveals high ant species richness within one of the most urbanized regions of the world. Diversity 13, 358 (doi:10.3390/d13080358).
• Dendup, K.C., Dorji, C., Dhadwal, T., Bharti, H., Pfeiffer, M. 2021. A preliminary checklist of ants from Bhutan. Asian Myrmecology 14, e014005 (doi:10.20362/am.014005).
• Donisthorpe, H. 1932b. On the identity of some ants from Ceylon described by F. Walker. Ann. Mag. Nat. Hist. 10(9): 574-576 (page 576, Combination in P. (Myrma))
• Donisthorpe, H. 1932b. On the identity of some ants from Ceylon described by F. Walker. Ann. Mag. Nat. Hist. 10(9): 574-576 (page 576, Senior synonym of mayri)
• General, D.E.M., Buenavente, P.A.C., Rodriguez, L.J.V. 2020. A preliminary survey of nocturnal ants, with novel modifications for collecting nocturnal arboreal ants. Halteres 11: 1-12 (doi:10.5281/ZENODO.3707151).
• Imai, H. T.; Brown, W. L., Jr.; Kubota, M.; Yong, H.-S.; Tho, Y. P. 1984. Chromosome observations on tropical ants from western Malaysia. II. Annu. Rep. Natl. Inst. Genet. Jpn. 34: 66-69 (page 68, karyotype described)
• Khachonpisitsak, S., Yamane, S., Sriwichai, P., Jaitrong, W. 2020. An updated checklist of the ants of Thailand (Hymenoptera, Formicidae). ZooKeys 998, 1–182 (doi:10.3897/zookeys.998.54902).
• Lee, C.-C., Weng, Y.-M., Lai, L.-C., Suarez, A.V., Wu, W.-J., Lin, C.-C., Yang, C.-C.S. 2020. Analysis of recent interception records reveals frequent transport of arboreal ants and potential predictors for ant invasion in Taiwan. Insects 11, 356 (doi:10.3390/INSECTS11060356).
• Lin, W.-J., Lee, Y.-I., Liu, S.-L., Lin, C.-C., Chung, T.-Y., Chou, J.-Y. 2020. Evaluating the tradeoffs of a generalist parasitoid fungus, Ophiocordyceps unilateralis, on different sympatric ant hosts. Scientific Reports 10:6428 (doi:10.1038/s41598-020-63400-1).
• Liu, C., Fischer, G., Hita Garcia, F., Yamane, S., Liu, Q., Peng, Y.Q., Economo, E.P., Guénard, B., Pierce, N.E. 2020. Ants of the Hengduan Mountains: a new altitudinal survey and updated checklist for Yunnan Province highlight an understudied insect biodiversity hotspot. ZooKeys 978, 1–171 (doi:10.3897/zookeys.978.55767).
• Rafiqi, A.M., Rajakumar, A., Abouheif, E. 2020. Origin and elaboration of a major evolutionary transition in individuality. Nature 585, 239–244. (doi:10.1038/s41586-020-2653-6).
• Walker, F. 1859. Characters of some apparently undescribed Ceylon insects. [part]. Ann. Mag. Nat. Hist. 3(4): 370-376 (page 373, worker described)
• Wong, T.L., Guénard, B. 2020. Review of ants from the genus Polyrhachis Smith (Hymenoptera: Formicidae: Formicinae) in Hong Kong and Macau, with notes on their natural history. Asian Myrmecology 13: e013001 (doi:10.20362/am.013001).
• Wu, J. & Wang, C. 1992. Formicidae (pp. 1301-1320). In Peng, J. et al. Iconography of Forest Insects in Hunan, China. Forest Bureau of Hunan Province: 1473 pp. Hunan Scientific and Technical Publishing House.
• Yamane, S., Tanaka, H.O., Hasimoto, Y., Ohashi, M., Meleng, P., Itioka, T. 2021. A list of ants from Lambir Hills National Park and its vicinity, with their biological information: Part II. Subfamilies Leptanillinae, Proceratiinae, Amblyoponinae, Ponerinae, Dorylinae, Dolichoderinae, Ectatomminae and Formicinae. Contributions from the Biological Laboratory, Kyoto University 31, 87–157.
• Yano, M. 1911a. The genus Polyrhachis of Japan. [In Japanese.] Dobutsugaku Zasshi (Zool. Mag.) 23:249-256.
• Yusah, K., Fayle, T. 2014. The first record of a fly of the family Milichiidae (Diptera) interacting with an ant of the genus Polyrhachis Smith, 1857 (Hymenoptera: Formicidae). Biodiversity Data Journal 2, e4168 (doi:10.3897/bdj.2.e4168).

References based on Global Ant Biodiversity Informatics

• Abe T., S. Yamane, and K. Onoyama. Ants collected on the Krakatau Islands 100 years after the great eruptions. Biogeography 14: 65-75.
• Andersen A. N., R. J. Kohout, and C. R. Trainor. 2013. Biogeography of Timor and Surrounding Wallacean Islands: Endemism in Ants of the Genus Polyrhachis Fr. Smith. Diversity 5: 139-148.
• André E. 1892. Voyage de M. Chaper à Bornéo. Catalogue des fourmis et description des espèces nouvelles. Mém. Soc. Zool. Fr. 5: 46-55.
• Ashmead W. H. 1904. A list of the Hymenoptera of the Philippine Islands, with descriptions of new species. J. N. Y. Entomol. Soc. 12:1-22.
• Bharti H., Y. P. Sharma, M. Bharti, and M. Pfeiffer. 2013. Ant species richness, endemicity and functional groups, along an elevational gradient in the Himalayas. Asian Myrmecology 5: 79-101.
• Bolton B. 1974. New synonymy and a new name in the ant genus Polyrhachis F. Smith (Hym., Formicidae). Entomol. Mon. Mag. 109: 172-180
• Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
• Chen Y. Q., Q. Li, Y. L. Chen, Z. X. Lu, X. Y. Zhou. 2011. Ant diversity and bio-indicators in land management of lac insect agroecosystem in Southwestern China. Biodivers. Conserv. 20: 3017-3038.
• Chen Y., C. W. Luo, H. W. Li, Y. J. Liu, H. F. Zheng, and F. C. Yang. 2013. Investigation of ant species and distribution on Wuliang Mountain. Journal of Henan Agricultural Sciences 42(5): 118-122.
• Chen Z. L., S. Y. Zhou, D. D. Ye, Y. Chen, and C. W. Lu. 2013. Moleular phylogeny of the ant subfamily Formicinae (Hymenoptera, Formicidae) from China based on Mitochondrial genes. Sociobiology 60(2): 135-144.
• Chhotani O. B., and P. K. Maiti. 1977. Contribution to the knowledge of Formicidae of the Andaman Islands. Zoological Survey of India 3(1): 17-20.
• Collingwood C.A. 1970. Formicidae (Hymenopter: Aculeata) of Nepal. Himalaya Khumbu Himal, 3: 371-388.
• Crawley W.C. 1924. Ants from Sumatra, with biological notes by Edward Jacobson. Annals and Magazine of Natural History (9)13: 380-409
• Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
• Davidson D. W., S. C. Cook, R. R. Snelling and T. H. Chua. 2003. Explaining the Abundance of Ants in Lowland Tropical Rainforest Canopies. Science 300: 969-972.
• Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
• Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
• Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
• Dias R. K. S., and H. P. G. R. C. Ruchirani. 2014. Nest density of Aneuretus simoni Emery (Sri Lankan Relict Ant) and Stereomyrmex horni Emery in three forest regions in western and southern Sri Lanka. Asian Myrmecology 6: 83-90.
• Donisthorpe H. 1942. Ants from the Colombo Museum Expedition to Southern India, September-October 1938. Annals and Magazine of Natural History (11)9: 449-461.
• Donisthorpe, Horace. 1943. The Ants of Waigeu Island, North Dutch New Guinea. The Annals and Magazine of Natural History 11 (10): 433-475.
• Emery C. 1886. Saggio di un catalogo sistematico dei generi Camponotus, Polyrhachis e affini. Memorie della Reale Accademia delle Scienze dell'Istituto di Bologna 5: 363-382
• Emery C. 1888. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza . Formiche raccolte dal sig. Elio Modigliani in Sumatra e nell'isola Nias. Annali del Museo Civico di Storia Naturale 25: 528-534.
• Emery C. 1893. Voyage de M. E. Simon à l'île de Ceylan (janvier-février 1892). Formicides. Annales de la Société Entomologique de France 62: 239-258.
• Emery C. 1901. Ameisen gesammelt in Ceylon von Dr. W. Horn 1899. Deutsche Entomologische Zeitschrift 1901: 113-122.
• Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
• Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
• Figueras G. S., and O. M. Nuneza. 2013. Species diversity of ants in karst limestone habitats in Bukidnon and Davao Oriental, Mindanao, Philippines. Advances in Environmental Sciences - International Journal of the Bioflux Society 5(3): 306-315.
• Fontanilla A. M., A. Nakamura, Z. Xu, M. Cao, R. L. Kitching, Y. Tang, and C. J. Burwell. 2019. Taxonomic and functional ant diversity along tropical, subtropical, and subalpine elevational transects in southwest China. Insects 10, 128; doi:10.3390/insects10050128
• Forel A. 1893. Les Formicides de l'Empire des Indes et de Ceylan. Part III. J. Bombay Nat. Hist. Soc. 8: 17-36.
• Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
• Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
• Forel A. 1909. Ameisen aus Java und Krakatau beobachtet und gesammelt von Herrn Edward Jacobson. Notes Leyden Mus. 31: 221-232.
• Forel A. 1911. Ameisen aus Ceylon, gesammelt von Prof. K. Escherich (einige von Prof. E. Bugnion). Pp. 215-228 in: Escherich, K. Termitenleben auf Ceylon. Jena: Gustav Fischer, xxxii + 262 pp.
• Forel A. 1911. Ameisen aus Java beobachtet und gesammelt von Herrn Edward Jacobson. II. Theil. Notes Leyden Mus. 33: 193-218.
• Forel A. 1912. Ameisen aus Java beobachtet und gesammelt von Edward Jacobson. III. Theil. Notes Leyden Mus. 34: 97-112
• Forel A. 1912. Einige neue und interessante Ameisenformen aus Sumatra etc. Zool. Jahrb. Suppl. 15: 51-78.
• Forel A. 1913. H. Sauter's Formosa-Ausbeute: Formicidae II. Arch. Naturgesch. (A)79(6): 183-202
• Forel A. 1913k. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
• Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
• Forel, A. 1908. Fourmis de Ceylan et d'Égypte récoltées par le Prof. E. Bugnion. Lasius carniolicus. Fourmis de Kerguelen. Pseudandrie? Strongylognathus testaceus. Bull. Soc. Vaudoise Sci. Nat. 44: 1-22
• Ghosh S. N., S. Sheela, B. G. Kundu, S. Roychowdhury, and R. N. Tiwari. 2006. Insecta: Hymenoptera: Formicidae. Pp. 369-398 in: Alfred, J. R. B. (ed.) 2006. Fauna of Arunachal Pradesh. (Part -2). [State Fauna Series 13.]. New Delhi: Zoological Survey of India, iv + 518 pp.
• Gumawardene, N.R., J.D. Majer and J.P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2:71-83
• Gunawardene N. R., J. D. Majer, and J. P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2: 71-83.
• Guo Zhenchao. 2006. Taxonomic Study on the ant fauna from Jigongshan and Funiushan in Henan Province. Guangxi Normal University. Master's thesis 91 pages.
• Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
• Hashimoto Y., Y. Morimoto, E. S. Widodo, and M. Mohamed. 2006. Vertical distribution pattern of ants in a Bornean tropical rainforest (Hymenoptera: Formicidae). Sociobiology 47(3): 697- 710.
• Imai H. T., M. Kubota, W. L. Brown, Jr., M. Ihara, M. Tohari, and R. I. Pranata. 1985. Chromosome observations on tropical ants from Indonesia. Annu. Rep. Natl. Inst. Genet. Jpn. 35: 46-48.
• Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
• Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
• Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
• Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
• Kaleeswaran B., S. Ezil, B. P. Ganesh, and S. Bhavatarini. 2008. Biodiversity and niches of Ants in Alagar hills, Tamil nadu Wildlife Biodiversity Conservation Published by Day publishing house pp 188-208.
• Karavaiev V. 1930. Ameisen von den Molukken und Neuguinea. (Ergebnisse der Sunda-Expedition der Notgemeinschaft der deutschen Wissenschaft 1929/30.). Zool. Anz. 92: 206-214.
• Karmaly K. A.; S. Sumesh, T. P. Rabeesh, and L. Kishore. 2010. A checklist of ants of Thirunelli in Wayanad, Kerala. J. of the Bombay Natural History Society 107(1): 64-67.
• Katayama M., K. Kishimoto-Yamada, H. O. Tanaka, T. Endo, Y. Hashimoto, Sk. Yamane, and T. Itioka. 2015. Negative correlation between ant and spider abundances in the canopy of a Bornean tropical rain forest. Biotropica (in press).
• Kishimoto-Yamata K., F. Hyodo, M. Matsuoka, Y. Hashimoto, M. Kon, T. Ochi, S. Yamane, R. Ishii, and T. Itioka. 2012. Effects of remnant primary forests on ant and dung beetle species diversity in a secondary forest in Sarawak, Malaysia. Journal of Insect Conservation DOI 10.1007/s10841-012-9544-6
• Kohout R.J., and M. Mohamed. 2008. A preliminary list of the Polyrhachis ants of the Maliau Basin Conservation area in Sabah, Borneo (Hymenoptera: Formicidae: Formicinae). Asian Myrmecology 2: 63-70.
• Leong C. M., S. F. Shiao, and B. Guenard. 2017. Ants in the city, a preliminary checklist of Formicidae (Hymenoptera) in Macau, one of the most heavily urbanized regions of the world. Asian Myrmecology 9: e009014.
• Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
• Liu X. 2012. Taxonomy, diversity and spatial distribution characters of the ant family Formicidae (Insecta: Hymenoptera) in southeastern Tibet. PhD Thesis 139 pages
• Liu X., Z. Xu, N. Yu, and C. Zhang. 2016. Distribution patterns of ant species ( Hymenoptera: Formicidae) in Galongla Mountains and Medog Valley of Southeastern Tibet. Scientia Silvae Sinicae 52(11): 88-95.
• Malsch A. K. F., K. Rosciszewski, and U. Maschwitz. 2003. The ant species richness and diversity of a primary lowland rain forest, the Pasoh Forest reserve, West Malaysia. in T. Okuda, N. Manokaran, Y. Matsumoto, K. Niiyama, S. C. Thomas, and P. S. Ashton, eds. Pasoh: Ecology and Natural History of a Southeast Asin Lowland Tropical Rain Forest, pp 347-374.
• Mathew R. 2003. On Formicidae (Insecta: Hymenoptera) of Nongkhyllem Wild Life Sanctuary, Ri-Bhoi District, Meghalaya. Records of the Zoological Survey of India 101:195-207.
• Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
• Mathew, R. 2000. Insecta : Hymenoptera : Formicidae. State Fauna Series. , Fauna of Tripura , Zool. Surv. India. 7 (3) : 347-354.
• Menozzi C. 1930. Formiche di Sumatra raccolte dal Prof. J. C. van der Meer Mohr. Miscellanea Zoologica Sumatrana 47: 1-5.
• Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
• Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
• Musthak Ali T. M. 1992. Ant Fauna of Karnataka-2. Newsletter of IUSSI Indian Chapter 6(1-2): 1-9.
• Norval G., W. F. Hsiao, C. C. Lin, and S. C. Huang. 2011. Ambushing the supply line: a report on Anolis sagrei predation on ants in Chiayi County, Taiwan. Russian Journal of Herpetology 18(1): 39-46.
• Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
• Qin F., S. Wei, Z. Zeng, S. Zhou, and X. Qin. 2008. Molecular phylogenetic research on five genera of Formicidae. Agricultural Science and Technology 9(5): 66-69.
• Ran H., and S. Y. Zhou. 2013. Checklist of Chinese Ants: Formicomorph Subfamilies ( Hymenoptera: Formicidae) (III). Journal of Guangxi Normal University : Natural Science Edition 31(1): 104-111.
• Robson Simon Ant Collection, 05-Sept-2014
• Robson Simon Database Polyrhachis -05 Sept 2014
• Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
• Sheela S. 2008. Handbook of Hymenoptera, Formicidae. Zoological Survey of India, 56 pages
• Sitthicharoenchai D., and N. Chantarasawat. 2006. Ant species diversity in the establishing area for Advanced Technology Institute at Lai-Nan Sub-district, Wiang Sa district, Nan Province, Thailand. The Natural History Journal of Chulalongkorn University 6(2): 67-74
• Skarbek C. J., M. Noack, H. Bruelheide, W. Hardtle, G. von Oheimb, T. Scholten, S. Seitz, M. Staab. 2019. A tale of scale: plot but not neighbourhood tree diversity increases leaf litter ant diversity. Journal of Animal Ecology DOI: 10.1111/1365-2656.13115
• Song Y., Z. Xu, C. Li, N. Zhang, L. Zhang, H. Jiang, and F. Mo. 2013. An Analysis on the Ant Fauna of the Nangun river Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
• Staab M., A. Schuldt, T. Assmann, H. Bruelheide, and A.M. Klein. 2014. Ant community structure during forest succession in a subtropical forest in South-East China. Acta Oecologia 61: 32-40.
• Stitz H. 1925. Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1923: 110-136.
• Sureh P. V., V. V. Sudheendrakumar, C. F. Binoy, G. Mathew, and T. C. Narendran. 1999. The macro Hymenopteran fauna of Parambikulam wildlife Sanctuary. Zoos' Print Journal 14(4): 1-2.
• Tang Jue, Li Shen, Huang Enyou, Zhang Benyue. 1985. Notes on ants from Zhoushan Islands Zhejiang (Hymenoptera: Formicidae).  Journal of Zhejiang University (Agric.& Life Sci.) 3.
• Terayama M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta: Hymenoptera). Research Bulletin of Kanto Gakuen University. Liberal Arts 17:81-266.
• Terayama, M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta; Hymenoptera). The Research Bulletin of Kanto Gakuen University 17: 81-266.
• Thapa V. K. 2000. An Inventory of Nepal's Insects, Vol. III. IUCN Nepal, Kathmandu, xi + 475 pp.
• Tiwari R. N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4): 1-96.
• Tiwari R. N., B. G. Kundu, S. Roy Chowdhury, and S. N. Ghosh. 2003. Insecta: Hymenoptera: Formicidae. Fauna of Sikkim. Part 4. State Fauna Series. 9.Zool.Surv.India. i-iii, 1-512. Chapter pagination: 467-506.
• Tiwari R.N., B.G. Kundu, S. Roychowdhury, S.N. Ghosh. 1999. Insecta: Hymenoptera: Formicidae. Pp. 211-294 in: Director; Zoological Survey of India (ed.) 1999. Fauna of West Bengal. Part 8. Insecta (Trichoptera, Thysanoptera, Neuroptera, Hymenoptera and Anoplura). Calcutta: Zoological Survey of India, iv + 442 pp.
• Tiwari, R.N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4):1-96
• Trainor C.R., and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
• Wang C. and Wu J. 1991. Taxonomic studies on the genus Polyrhachis of China (Hymenoptera: Formicidae). Forest Research 4: 596-601.
• Wang W. 2006. A taxonomic study of Formicidae ants from Hujiaping forest farm in Laifeng County. Journal of Hubei Institute for Nationalities (Natural Science Edition). 24(3): 298-300.
• Watanasit S., A. Nhusidum, and T. Takodee. 2007. Diversity of canopy ants at a reserve area of Prince of Songkla University, Songkhla. Songklanakarin J. Sci. Technol. 29(2): 307-320.
• Watanasit S., S. Sonthichai, and N. Noon-anant. 2003. Preliminary survey of ants at Tarutao National Park, Southern Thailand. Songklanakarin J. Sci. Technol. 25(1) : 115-122
• Wheeler W. M. 1909. Ants of Formosa and the Philippines. Bulletin of the American Museum of Natural History 26: 333-345.
• Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
• Wheeler W. M. 1921. Chinese ants collected by Prof. C. W. Howard. Psyche (Cambridge) 28: 110-115.
• Wheeler W. M. 1927. Chinese ants collected by Professor S. F. Light and Professor N. Gist Gee. American Museum Novitates 255: 1-12.
• Wheeler W. M. 1928. Ants collected by Professor F. Silvestri in China. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 22: 3-38.
• Wheeler W. M. 1929. Ants collected by Professor F. Silvestri in Formosa, the Malay Peninsula and the Philippines. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 24: 27-64.
• Wheeler W. M. 1930. A list of the known Chinese ants. Peking Natural History Bulletin 5: 53-81.
• Wheeler W. M. 1937. Additions to the ant-fauna of Krakatau and Verlaten Island. Treubia 16: 21-24.
• Wheeler, William Morton. 1924. Ants of Krakatau and Other Islands in the Sunda Strait. Treubia. 5(1-3):1-20.
• Widodo E. S., T. Naito, M. Mohamed, and Y. Hashimoto. 2004. Effects of selective logging on the arboreal ants of a Bornean rainforest. Entomological Science 7: 341-349.
• Widodo E.S., M. Mohamed, and Y. Hashimoto. 2001. Canopy ant diversity assessment in the fragmented rainforest of Sabah, East Malaysia. Nature and Human activities 6: 13-23.
• Xu F. F., and J. Chen. 2009. Comparison of the Differences in Response to the Change of the Extrafloral Nectar-ant-herbivore Interaction System Between a Native and an Introduced Passiflora Species. Acta Botanica Yunnanica 31(6): 543-550.
• Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
• Yamane S.; Nona, A. R. 1994. Ants from Lambir Hills National Park, Sarawak. Pp. 222-226 in: Inoue, T.; Hamid, A. A. (eds.) 1994. Plant reproductive systems and animal seasonal dynamics. Long-term study of dipterocarp forests in Sarawak. Kyoto: Center for Ecological Research, Kyoto University, vii + 255 pp. 
• Zhang R. J., L. W. Liang, and S. Y. Zhou. 2014. An analysis on the ant fauna of Nonggang Nature Reserve in Guangxi, China. Journal of Guangxi Normal university: Natural Science Edition 32(3): 86-93.
• Zhang W., G. Liu, P. Zhong, and S. Zhang. 2014. Investigation of Formicidae in Luofushan Mountain. Journal of Huizhou University 34(3): 46-50.
• Zhang W., and S. Zhou. 2016. An investigation on Formicidae species of Nanling National Park. Journal of Huizhou University 36(3): 27-30.
• Zhou S.-Y. 2001. Ants of Guangxi. Guangxi Normal University Press, Guilin, China, Guilin, China. 255 pp.
• Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.