Camponotus substitutus

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Camponotus substitutus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Camponotini
Genus: Camponotus
Species: C. substitutus
Binomial name
Camponotus substitutus
Emery, 1894

Camponotus substitutus casent0173454 profile 1.jpg

Camponotus substitutus casent0173454 dorsal 1.jpg

Specimen labels

Subspecies
Synonyms

Identification

Keys including this Species

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 22.29073333° to -37.533333°.

     
North
Temperate 		North
Subtropical 		Tropical South
Subtropical 		South
Temperate

Distribution based on Regional Taxon Lists

Neotropical Region: Argentina, Brazil (type locality), Chile, Colombia, French Guiana, Guatemala, Guyana, Mexico, Panama, Paraguay, Suriname, Uruguay, Venezuela.

Distribution based on AntMaps

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Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
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Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
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Biology

Flight Period

X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

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Castes

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • substitutus. Camponotus landolti r. substitutus Forel, 1899c: 136.
    • [First available use of Camponotus extensus subsp. melanoticus var. substitutus Emery, 1894f: 3 (s.w.) BRAZIL, BOLIVIA, PARAGUAY; unavailable (infrasubspecific) name.]
    • Type-material: syntype workers, syntype queens (numbers not stated).
    • [Note: this taxon is based on workers of sexguttatus (Fabricius, 1793: 354), sensu Smith, F. 1858b: 44, misidentification, sensu Mayr, 1862: 656, misidentification, and sensu Forel, 1879a: 71, misidentification.]
    • Type-localities: workers, queen Brazil: Pará, Santarem (H.W. Bates) (invalid restriction of type-locality by Kempf, 1972a: 72; no lectotype designated), queen Brazil: Rio de Janeiro (Novara Expd.), workers, queens Brazil: Rio de Janeiro (Naegeli), Brazil: Bahia, Caravellas (Joseph), Brazil: Pará (ex coll. Saussure), French Guiana: Cayenne (ex coll. Saussure), Mexico: Orizaba (ex coll. Saussure).
    • Type-depositories: BMNH, MHNG, NHMW.
    • Combination in C. (Myrmoturba): Bruch, 1914: 229;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 82.
    • As unavailable (infrasubspecific) name: Emery, 1896d: 371 (in list); Forel, 1908e: 62.
    • [Note: substitutus appears twice in Forel, 1908e: 62, first (entry no. 4) as a species, second (entry no.7) as an infrasubspecific name.]
    • Subspecies of landolti: Forel, 1899c: 136.
    • Subspecies of melanoticus: Emery, 1906c: 188; Forel, 1907e: 10; Forel, 1911c: 310; Forel, 1914e: 13; Bruch, 1914: 229; Mann, 1916: 476; Santschi, 1922d: 262.
    • Subspecies of maculatus: Santschi, 1916e: 396; Santschi, 1919f: 56.
    • Subspecies of fumidus: Santschi, 1922c: 101.
    • Status as species: Forel, 1902b: 171; Forel, 1908e: 62; Emery, 1925b: 82; Borgmeier, 1927c: 146; Santschi, 1929d: 313; Kusnezov, 1952d: 225; Kusnezov, 1953b: 339; Kempf, 1972a: 72; Zolessi, et al. 1988: 7; Brandão, 1991: 335; Bolton, 1995b: 125; Wild, 2007b: 29, 49; Mackay & Delsinne, 2009: 497 (in key); Branstetter & Sáenz, 2012: 256; Mackay & Mackay, 2019: 769.
    • Senior synonym of multipilis: Wild, 2007b: 49.
    • Distribution: Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Mexico, Panama, Paraguay, Uruguay, Venezuela.
    • Current subspecies: nominal plus clarus (unresolved junior homonym), pullulus.
  • multipilis. Camponotus melanoticus var. multipilis Forel, 1907e: 10 (w.q.m.) PARAGUAY.
    • Type-material: syntype major and minor workers, syntype queens, syntype males (numbers not stated).
    • Type-locality: Paraguay: Puerto Max, Estancia Postillon (L. Des Arts).
    • Type-depository: MHNG.
    • Combination in C. (Myrmoturba): Bruch, 1914: 229;
    • combination in C. (Tanaemyrmex): Emery, 1925b: 83.
    • Subspecies of melanoticus: Forel, 1912i: 70; Bruch, 1914: 229.
    • Subspecies of substitutus: Emery, 1925b: 83; Kusnezov, 1952d: 225; Kempf, 1972a: 72; Bolton, 1995b: 112.
    • Junior synonym of substitutus: Wild, 2007b: 49.

Description

References

References based on Global Ant Biodiversity Informatics

  • Boer P. 2019. Ants of Curacao, species list. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/SPECIES%20LIST%20CURACAO.html
  • Brandao, C.R.F. 1991. Adendos ao catalogo abreviado das formigas da regiao neotropical (Hymenoptera: Formicidae). Rev. Bras. Entomol. 35: 319-412.
  • Bruch C. 1914. Catálogo sistemático de los formícidos argentinos. Revista del Museo de La Plata 19: 211-234.
  • Calcaterra L. A., F. Cuezzo, S. M. Cabrera, and J. A. Briano. 2010. Ground ant diversity (Hymenoptera: Formicidae) in the Ibera nature reserve, the largest wetland of Argentina. Ann. Entomol. Soc. Am. 103(1): 71-83.
  • Cuezzo, F. 1998. Formicidae. Chapter 42 in Morrone J.J., and S. Coscaron (dirs) Biodiversidad de artropodos argentinos: una perspectiva biotaxonomica Ediciones Sur, La Plata. Pages 452-462.
  • Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
  • Emery C. 1886. Saggio di un catalogo sistematico dei generi Camponotus, Polyrhachis e affini. Memorie della Reale Accademia delle Scienze dell'Istituto di Bologna 5: 363-382
  • Emery C. 1894. Camponotus sexguttatus Fab. e C. sexguttatus Sm. et auct. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 9(187): 1-4.
  • Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
  • Fernández F., E. E. Palacio, W. P. Mackay, and E. S. MacKay. 1996. Introducción al estudio de las hormigas (Hymenoptera: Formicidae) de Colombia. Pp. 349-412 in: Andrade M. G., G. Amat García, and F. Fernández. (eds.) 1996. Insectos de Colombia. Estudios escogidos. Bogotá: Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 541 pp
  • Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
  • Forel A. 1902. Quatre notices myrmécologiques. Annales de la Société Entomologique de Belgique 46: 170-182.
  • Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
  • Forel A. 1908. Catálogo systemático da collecção de formigas do Ceará. Boletim do Museu Rocha 1(1): 62-69.
  • Forel A. 1912. Formicides néotropiques. Part VI. 5me sous-famille Camponotinae Forel. Mémoires de la Société Entomologique de Belgique. 20: 59-92.
  • Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
  • INBio Collection (via Gbif)
  • Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
  • Kusnezov N. 1952. El género Camponotus en la Argentina (Hymenoptera, Formicidae). Acta Zoologica Lilloana 12: 183-252.
  • Kusnezov N. 1953. La fauna mirmecológica de Bolivia. Folia Universitaria. Cochabamba 6: 211-229.
  • Kusnezov N. 1978. Hormigas argentinas: clave para su identificación. Miscelánea. Instituto Miguel Lillo 61:1-147 + 28 pl.
  • Maes, J.-M. and W.P. MacKay. 1993. Catalogo de las hormigas (Hymenoptera: Formicidae) de Nicaragua. Revista Nicaraguense de Entomologia 23.
  • Mann W. M. 1916. The Stanford Expedition to Brazil, 1911, John C. Branner, Director. The ants of Brazil. Bulletin of the Museum of Comparative Zoology 60: 399-490
  • Murua A. F., F. Cuezzo, and J. C. Acosta. 1999. La fauna de hormigas del Gran Bajo Oriental del departamento Valle Fertíl (San Juan, Argentina). Revista de la Sociedad Entomológica Argentina 58(3/4): 135-138.
  • Nunes F. A., G. B. Martins Segundo, Y. B. Vasconcelos, R. Azevedo, and Y. Quinet. 2011. Ground-foraging ants (Hymenoptera: Formicidae) and rainfall effect on pitfall trapping in a deciduous thorn woodland (Caatinga), Northeastern Brazil. Rev. Biol. Trop. 59 (4): 1637-1650.
  • Pignalberi C. T. 1961. Contribución al conocimiento de los formícidos de la provincia de Santa Fé. Pp. 165-173 in: Comisión Investigación Científica; Consejo Nacional de Investigaciones Científicas y Técnicas (Argentina) 1961. Actas y trabajos del primer Congreso Sudamericano de Zoología (La Plata, 12-24 octubre 1959). Tomo III. Buenos Aires: Librart, 276 pp.
  • Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
  • Santschi F. 1916. Formicides sudaméricains nouveaux ou peu connus. Physis (Buenos Aires). 2: 365-399.
  • Santschi F. 1925. Fourmis des provinces argentines de Santa Fe, Catamarca, Santa Cruz, Córdoba et Los Andes. Comunicaciones del Museo Nacional de Historia Natural "Bernardino Rivadavia" 2: 149-168.
  • Santschi F. 1929. Nouvelles fourmis de la République Argentine et du Brésil. Anales de la Sociedad Cientifica Argentina. 107: 273-316.
  • Ulyssea M. A., and C. R. F. Brandao. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217–224.
  • Ulysséa M. A., C. R. F. Brandão. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217-224.
  • Vittar, F. 2008. Hormigas (Hymenoptera: Formicidae) de la Mesopotamia Argentina. INSUGEO Miscelania 17(2):447-466
  • Vittar, F., and F. Cuezzo. "Hormigas (Hymenoptera: Formicidae) de la provincia de Santa Fe, Argentina." Revista de la Sociedad Entomológica Argentina (versión On-line ISSN 1851-7471) 67, no. 1-2 (2008).
  • Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
  • Zolessi L. C. de, Y. P. Abenante, and M. E. de Philippi. 1988. Lista sistematica de las especies de Formicidos del Uruguay. Comun. Zool. Mus. Hist. Nat. Montev. 11: 1-9.
  • Zolessi L. C. de; Y. P. de Abenante, and M. E. Philippi. 1989. Catálogo sistemático de las especies de Formícidos del Uruguay (Hymenoptera: Formicidae). Montevideo: ORCYT Unesco, 40 + ix pp.
  • de Zolessi, L.C., Y.P. de Abenante and M.E. Philippi. 1987. Lista sistemática de las especies de formícidos del Uruguay. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo 11(165):1-9
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