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Camponotus ethicus
| Camponotus ethicus | |
|---|---|
| |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Hymenoptera |
| Family: | Formicidae |
| Subfamily: | Formicinae |
| Tribe: | Camponotini |
| Genus: | Camponotus |
| Subgenus: | Mayria |
| Species group: | robustus |
| Species: | C. ethicus |
| Binomial name | |
| Camponotus ethicus Forel, 1897
| |
Known from the dry forests of the Parc National Sahamalaza and the Reserve Spéciale Ambre, and the transitional rainforest of the Galoko Chain in the northwestern part of Madagascar, C. ethicus is arboreal, forages most often on lower vegetation, and nests in dead branches above the ground. (Rakotonirina et al. 2016)
Identification
Rasoamanana and Fisher (2022) - A member of the Camponotus robustus species group.
Rakotonirina et al. (2016) - Larger species (CS: 1.92–2.58; ML: 3.49–4.18) with uniformly black to dark brown body color; in profile anterior margin of petiolar node convex and posterior margin straight; level of the propodeal dorsum abruptly lower than level of the promesonotal dorsum; pronotal dorsum without numerous erect hairs; humeral angle extended anteriorly into a narrow ridge.
The lower level of the propodeal dorsum relative to the promesonotum and the larger body size combined with the dark color of the legs make C. ethicus easy to separate from Camponotus robustus.
Keys including this Species
- Key to Malagasy Camponotus edmondi species group
- Key to Malagasy Camponotus subgenus Mayria minor workers
Distribution
Latitudinal Distribution Pattern
Latitudinal Range: -12.46889° to -14.30889°.
| North Temperate |
North Subtropical |
Tropical | South Subtropical |
South Temperate |
- Source: AntMaps
Distribution based on Regional Taxon Lists
Malagasy Region: Madagascar (type locality).
Distribution based on AntMaps
Distribution based on AntWeb specimens
Check data from AntWeb
Countries Occupied
| Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species. |
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Estimated Abundance
| Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species. |
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Biology
Castes
Worker
Images from AntWeb
   
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| Lectotype of Camponotus ethicus. Worker. Specimen code casent0101389. Photographer April Nobile, uploaded by California Academy of Sciences. | Owned by MHNG, Geneva, Switzerland. |
   
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| Paralectotype of Camponotus ethicus. Worker. Specimen code casent0101176. Photographer April Nobile, uploaded by California Academy of Sciences. | Owned by NHMB, Basel, Switzerland. |
Male
Images from AntWeb
   
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| Paralectotype of Camponotus ethicus. Male (alate). Specimen code casent0101387. Photographer April Nobile, uploaded by California Academy of Sciences. | Owned by MHNG, Geneva, Switzerland. |
  
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| Paralectotype of Camponotus ethicus. Male (alate). Specimen code casent0101177. Photographer April Nobile, uploaded by California Academy of Sciences. | Owned by NHMB, Basel, Switzerland. |
Nomenclature
The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.
- ethicus. Camponotus ethicus Forel, 1897c: 200 (w.m.) MADAGASCAR.
- Type-material: lectotype minor worker (by designation of Rakotonirina, et al. 2016: 122), 2 paralectotype minor workers, 2 paralectotype males.
- Type-locality: lectotype Madagascar: Nossi-Bé, Sakatia (A. Voeltzkow); paralectotypes with same data.
- Type-depositories: MHNG (lectotype); MHNG, NHMB (paralectotypes).
- Combination in C. (Myrmentoma): Forel, 1912i: 92;
- combination in C. (Orthonotomyrmex): Forel, 1914a: 273;
- combination in C. (Myrmisolepis): Santschi, 1921f: 310;
- combination in C. (Myrmepinotus): Emery, 1925b: 127;
- combination in C. (Mayria): Rasoamanana & Fisher, 2022: 147.
- Rakotonirina, et al. 2016: 124 (s.).
- Status as species: Emery, 1898a: 227; Wheeler, W.M. 1922a: 1049; Emery, 1925b: 127; Bolton, 1995b: 98; Rakotonirina, et al. 2016: 122 (redescription).
- Distribution: Madagascar.
Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Description
Worker
Rasoamanana and Fisher (2022) - Morphological measurements: see Appendix 1 and Ratios of morphometric data for majors and minors
Rakotonirina et al. (2016) - Minor. In full-face view head subquadrate (CWb/CL: 0.87–0.89), slightly diverging posteriorly; posterior margin more or less straight. Eyes not breaking lateral outline of head, their posterior level located at posterior fifth portion of head (PoOc/CL: 0.21–0.25). Anterior clypeal margin straight; posterior margin weakly notched medially. Mandible triangular, apical margin armed with six sharp teeth. Antennal scape long, apical half surpassing posterior cephalic margin. Pronotal dorsum flat, anterodorsal corner projecting anteriorly into narrow ridge; anterior margination present; pronotal dorsum and lateral portion anteriorly separated by sharp margination. In dorsal view, mesonotum as long as broad; in profile, mesonotal dorsum inclined posteriorly and propodeal dorsum nearly horizontal and distinctly situated at lower level than promesonotum; mesopleuron and propodeal surface together distinctly longer than lateral portion of pronotum; propodeal dorsum almost horizontal and declivitous surface nearly vertical; propodeal spiracle located anterior to posterolateral margin of propodeum. Width of procoxa larger than width of mesopleuron. In profile anterior margin of petiolar node convex and posterior margin more or less straight. Constriction between abdominal segments III and IV lacking.
Dorsum of head and mesosoma with fine and dense imbrication. Imbricate sculpture much finer and denser on gastral tergites. Mandible imbricate and superimposed with sparse large punctures. Erect hairs lacking on pronotum; one pair present on mesonotum, propodeum near junction of dorsum and declivity, and upper level of lateral margin of petiole. Few pairs of erect hairs on head dorsum from clypeus and edge of frontal lobe to posterior portion of head; several scattered pairs organized transversely on anterior and posterior portions of each gastral tergite; pubescence short and reduced in number. Body coloration black; appendages dark reddish black.
Major. With characteristics of minor worker, except: head much more square (CL/CWb: 1.008–1.08); lateral margins slightly convex. Eyes located more anteriorly, their posterior level on posterior fourth of head (PoOc/CL: 0.227–0.252). One third of apical portion of scape extending beyond posterior cephalic margin. Scattered piligerous punctures present laterally on head from the level of anterior margin of eyes to near base of mandible.
Type Material
- Lectotype (designated by Rakotonirina, Csősz & Fisher, 2016: 122), minor worker, Antsiranana Province, Sakatia bay, Nosy Be, Madagascar, Voeltzkow, CASENT0101389, Musee d'Histoire Naturelle Genève.
- Paralectotype (designated by Rakotonirina, Csősz & Fisher, 2016: 122), worker, Antsiranana Province, Sakatia bay, Nosy Be, Madagascar, Voeltzkow, CASENT0101388, Musee d'Histoire Naturelle Genève.
- Paralectotype (designated by Rakotonirina, Csősz & Fisher, 2016: 122), worker, Antsiranana Province, Sakatia bay, Nosy Be, Madagascar, Voeltzkow, CASENT0101176, Naturhistorisches Museum, Basel.
- Paralectotype (designated by Rakotonirina, Csősz & Fisher, 2016: 122), male, Antsiranana Province, Sakatia bay, Nosy Be, Madagascar, Voeltzkow, CASENT0101387, Musee d'Histoire Naturelle Genève.
- Paralectotype (designated by Rakotonirina, Csősz & Fisher, 2016: 122), male, Antsiranana Province, Sakatia bay, Nosy Be, Madagascar, Voeltzkow, CASENT0101177, Naturhistorisches Museum, Basel.
References
- Emery, C. 1925d. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183: 1-302 (page 127, combination in C. (Myrmepinotus))
- Forel, A. 1897d. Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen un (page 200, worker, male described)
- Forel, A. 1912j. Formicides néotropiques. Part VI. 5me sous-famille Camponotinae Forel. Mém. Soc. Entomol. Belg. 20: 59-92 (page 92, combination in C. (Myrmentoma))
- Forel, A. 1914a. Le genre Camponotus Mayr et les genres voisins. Rev. Suisse Zool. 22: 257-276 (page 273, combination in C. (Orthonotomyrmex))
- Rakotonirina J.C., Csosz S., and B.L. Fisher. 2016. Revision of the Malagasy Camponotus edmondi species group (Hymenoptera, Formicidae, Formicinae): integrating qualitative morphology and multivariate morphometric analysis. ZooKeys. 572:81-154. doi:10.3897/zookeys.572.7177
- Rasoamanana, N., Fisher, B.L. 2022. A taxonomic revision of the Malagasy endemic subgenus Mayria of the genus Camponotus (Hymenoptera, Formicidae) based on qualitative morphology and quantitative morphometric analyses. ZooKeys 1081: 137–231 (doi:10.3897/zookeys.1081.71872).
- Santschi, F. 1921f. Retouches aux sous-genres de Camponotus. Ann. Soc. Entomol. Belg. 61: 310-312 (page 310, combination in C. (Myrmisolepis))
- Wheeler, W. M. 1922k. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bull. Am. Mus. Nat. Hist. 4 45: 1005-1055 (see also)
References based on Global Ant Biodiversity Informatics
- Fisher B. L. 1997. Biogeography and ecology of the ant fauna of Madagascar (Hymenoptera: Formicidae). Journal of Natural History 31: 269-302.
- Fisher B. L. 2003. Formicidae, ants. Pp. 811-819 in: Goodman, S. M.; Benstead, J. P. (eds.) 2003. The natural history of Madagascar. Chicago: University of Chicago Press, xxi + 1709 pp.
- Forel A. 1897. Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn Perrot auf Ste. Marie (Madagaskar) gesammelten Ameisen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 21: 185-208.