Pheidole punctulata

Pheidole punctulata
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Myrmicinae
Tribe:	Attini
Genus:	Pheidole
Species:	P. punctulata
Binomial name
Pheidole punctulata Mayr, 1866 Specimen Label
Subspecies
Pheidole punctulata subatrox Santschi, 1937
Synonyms
Pheidole inquilina Forel Forel, 1914

Identification

A member of the megacephala species group

Distribution

Latitudinal Distribution Pattern

Latitudinal Range: 13.38675° to -33.3°.

North Temperate	North Subtropical	Tropical	South Subtropical	South Temperate

• Source: AntMaps

Distribution based on Regional Taxon Lists

Afrotropical Region: Comoros, Eritrea, Gambia, Guinea, Ivory Coast, Kenya, Mozambique, Namibia, South Africa (type locality), Zimbabwe, Zimbabwe.
Malagasy Region: Madagascar, Mayotte, Seychelles.

Distribution based on AntMaps

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.

Castes

.	Owned by Museum of Comparative Zoology.

Images from AntWeb

Worker (major/soldier). Specimen code casent0173311. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0173312. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Worker (major/soldier). Specimen code casent0173313. Photographer April Nobile, uploaded by California Academy of Sciences.	Owned by MHNG, Geneva, Switzerland.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

• punctulata. Pheidole punctulata Mayr, 1866b: 899 (s.) SOUTH AFRICA. Arnold, 1920a: 421 (w.q.m.). Subspecies of megacephala: Forel, 1894b: 94; Stitz, 1911b: 385; Emery, 1915j: 235; Emery, 1921f: 85; Wheeler, W.M. 1922a: 132; Santschi, 1930b: 67. Status as species: Forel, 1910e: 441; Santschi, 1914b: 76; Forel, 1914d: 240; Arnold, 1920a: 420; Santschi, 1935a: 256. Current subspecies: nominal plus atrox, spinosa, subatrox.

References

• Arnold, G. 1920a. A monograph of the Formicidae of South Africa. Part IV. Myrmicinae. Ann. S. Afr. Mus. 14: 403-578 (page 420, status as species)
• Arnold, G. 1920a. A monograph of the Formicidae of South Africa. Part IV. Myrmicinae. Ann. S. Afr. Mus. 14: 403-578 (page 421, worker, queen, male described)
• Borowiec, L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
• Borowiec, L., Salata, S. 2018. Notes on ants (Hymenoptera: Formicidae) from Gambia (Western Africa). Annals of the Upper Silesian Museum in Bytom, Entomology 26 (online 010), 1-13 (doi:10.5281/ZENODO.1243767).
• Cantone S. 2018. Winged Ants, The queen. Dichotomous key to genera of winged female ants in the World. The Wings of Ants: morphological and systematic relationships (self-published).
• Emery, C. 1915i. Les Pheidole du groupe megacephala (Formicidae). Rev. Zool. Afr. (Bruss.) 4: 223-250 (page 235, subspecies of megacephala)
• Emery, C. 1921c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174A:1-94 94: 1-94 + 7 (page 85, subspecies of megacephala)
• Forel, A. 1894b. Abessinische und andere afrikanische Ameisen, gesammelt von Herrn Ingenieur Alfred Ilg, von Herrn Dr. Liengme, von Herrn Pfarrer Missionar P. Berthoud, Herrn Dr. Arth. Müller etc. Mitt. Schweiz. Entomol. Ges. 9: 64-100 (page 94, subspecies of megacephala)
• Forel, A. 1910f. Note sur quelques fourmis d'Afrique. Ann. Soc. Entomol. Belg. 54: 421-458 (page 441, status as species)
• Forel, A. 1914d. Formicides d'Afrique et d'Amérique nouveaux ou peu connus. Bull. Soc. Vaudoise Sci. Nat. 50: 211-288 (page 240, status as species)
• Mayr, G. 1866b. Diagnosen neuer und wenig gekannter Formiciden. Verh. K-K. Zool.-Bot. Ges. Wien 16: 885-908 (page 899, soldier described)
• Salata, S., Fisher, B.L. 2022. Taxonomic revision of the Pheidole megacephala species-group (Hymenoptera, Formicidae) from the Malagasy Region. PeerJ 10: e13263 (doi:10.7717/peerj.13263).
• Santschi, F. 1914b. Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale, 1911-1912. Résultats scientifiques. Insectes Hyménoptères. II. Formicidae. Paris: Libr. A. Schulz, pp. 41-148. (page 76, status as species)
• Santschi, F. 1930b. Résultats de la Mission scientifique suisse en Angola, 1928-1929. Formicides de l'Angola. Rev. Suisse Zool. 37: 53-81 (page 67, subspecies of megacephala)
• Santschi, F. 1935b. Hymenoptera. I. Formicidae. Mission Sci. Omo 2: 255-277 (page 256, status as species)
• Stitz, H. 1911b. Formicidae. Wiss. Ergeb. Dtsch. Zent.-Afr.-Exped. 3: 375-392 (page 385, subspecies of megacephala)
• Taylor, B., Agoinon, N., Sinzogan, A., Adandonon, A., Kouaguou, Y. N., Bello, S., Wargui, R., Anato, F., Ouagoussounon, I., Houngbo, H., Tchibozo, S., Todjihounde, R., Vayssieres, J.F. 2018. Records of ants (Hymenoptera: Formicidae) from the Republic of Benin, with particular reference to the mango farm ecosystem. Journal of Insect Biodiversity 8(1): 6-29 (doi:10.12976/jib/2018.08.1.2).
• Wheeler, W. M. 1922b. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. II. The ants collected by the American Museum Congo Expedition. Bull. Am. Mus. Nat. Hist. 45: 39-269 (page 132, subspecies of megacephala)

References based on Global Ant Biodiversity Informatics

• André E. 1895. Formicides de l'Ogooué (Congo français). Rev. Entomol. (Caen) 14: 1-5.
• Arnold G. 1914. Nest-changing migrations of two species of ants. Proceedings and Transactions of the Rhodesia Scientific Association 13: 25-32.
• Arnold G. 1917. A monograph of the Formicidae of South Africa. Part III. Myrmicinae. Annals of the South African Museum. 14: 271-402.
• Arnold G. 1920. A monograph of the Formicidae of South Africa. Part IV. Myrmicinae. Annals of the South African Museum. 14: 403-578.
• Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) from Gambia (Western Africa). Annals of the Upper Silesian Museum in Bytom Entomology 26: 1-13.
• Crawley W. C. 1920. Ants from Mesopotamia and north-west Persia (concluded). Entomol. Rec. J. Var. 32: 177-179.
• Dorow, Wolfgang H. O. 1995. Review and Bibliography of the ants of the Seychelles (Hymenoptera: Formicidae). J. Afr. Zool. 110:73-96
• Dorow, Wolfgang H.O. 1996. Review and bibliography of the ants of the Seychelles. Journal of African Zoology 110(2): 73-95.
• Emery C. 1881. Spedizione italiana nell'Africa equatoriale. Risultati zoologici. Formiche. Annali del Museo Civico di Storia Naturale 16: 270-276.
• Emery C. 1892. Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et août 1886. Formicides. Annales de la Société Entomologique de France 60: 553-574.
• Finzi B. 1939. Materiali zoologici dell'Eritrea raccolti da G. Müller durante la spedizione dell'Istituto Sieroterapico Milanese e conservati al Museo di Trieste. Parte III. Hymenoptera: Formicidae. Atti del Museo Civico di Storia Naturale di Trieste 14: 153-168.
• Fisher B. L. 1997. Biogeography and ecology of the ant fauna of Madagascar (Hymenoptera: Formicidae). Journal of Natural History 31: 269-302.
• Fisher B. L. 2003. Formicidae, ants. Pp. 811-819 in: Goodman, S. M.; Benstead, J. P. (eds.) 2003. The natural history of Madagascar. Chicago: University of Chicago Press, xxi + 1709 pp.
• Forel A. 1897. Ameisen aus Nossi-Bé, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar, gesammelt von Herrn Dr. A. Voeltzkow aus Berlin. Mit einem Anhang über die von Herrn Privatdocenten Dr. A. Brauer in Marburg auf den Seychellen und von Herrn Perrot auf Ste. Marie (Madagaskar) gesammelten Ameisen. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 21: 185-208.
• Forel A. 1901. Einige neue Ameisen aus Südbrasilien, Java, Natal und Mossamedes. Mitteilungen der Schweizerischen Entomologischen Gesellschaft. 10: 297-311.
• Forel A. 1903. Einige neue Ameisen aus Sud-Angola. Pp 559-564, in: Baum, H. Kunene-Sambesi-Expedition, 1903. Berlin: Verlag des Kolonial-Wirtschaftlichen Komitees, 593pp.
• Forel A. 1907. Ameisen von Madagaskar, den Comoren und Ostafrika. Wissenschaftliche Ergebnisse. Reise in Ostafrika 2: 75-92.
• Forel A. 1909. Fourmis du Musée de Bruxelles. Fourmis de Benguela récoltées par M. Creighton Wellman, et fourmis du Congo récoltées par MM. Luja, Kohl et Laurent. Annales de la Société Entomologique de Belgique 53: 51-73.
• Forel A. 1910. Formicides australiens reçus de MM. Froggatt et Rowland Turner. Rev. Suisse Zool. 18: 1-94
• Forel A. 1912. The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, M.A. Volume 4. No. XI. Fourmis des Seychelles et des Aldabras, reçues de M. Hugh Scott. Trans. Linn. Soc. Lond. Zool. (2) 15: 159-167.
• Forel A. 1914. Formicides d'Afrique et d'Amérique nouveaux ou peu connus. Bulletin de la Société Vaudoise des Sciences Naturelles 50: 211-288.
• Garcia F.H., Wiesel E. and Fischer G. 2013.The Ants of Kenya (Hymenoptera: Formicidae)—Faunal Overview, First Species Checklist, Bibliography, Accounts for All Genera, and Discussion on Taxonomy and Zoogeography. Journal of East African Natural History, 101(2): 127-222
• IZIKO South Africa Museum Collection
• Kutter H. 1932. Ameisen aus dem Museum zu Dresden. Mitt. Schweiz. Entomol. Ges. 15: 207-210.
• Madl M. 2019. Notes on the ant fauna of Eritrea (Insecta: Hymenoptera: Formicidae): type specimens deposited in the Natural History Museum Vienna (Austria) and a preliminary checklist. Ann. Naturhist. Mus. Wien, B 121: 9-18.
• Prins A. J. 1963. A list of the ants collected in the Kruger National Park with notes on their distribution. Koedoe 6: 91-108.
• Prins A. J. 1964. Revised list of the ants collected in the Kruger National Park. Koedoe 7: 77-93.
• Prins A. J., and J. J. Cillie. 1968. The ants collected in the Hluhluwe and Umfolozi game reserves. The Lammergeyer 8: 40-47.
• Santschi F. 1914. Formicides de l'Afrique occidentale et australe du voyage de Mr. le Professeur F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 8: 309-385.
• Santschi, F. "Résultats de la Mission scientifique suisse en Angola, 1928-1929. Formicides de l'Angola." Revue Suisse de Zoologie 37 (1930): 53-81.
• Weber N. A. 1964. Termite prey of some African ants. Entomological News 75: 197-204.
• Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. II. The ants collected by the American Museum Congo Expedition. Bulletin of the American Museum of Natural History 45: 39-269.
• Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bulletin of the American Museum of Natural History 45: 1005-1055
• Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bulletin of the American Museum of Natural History 45: 711-1004