Leptogenys diminuta

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Leptogenys diminuta
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Ponerinae
Tribe: Ponerini
Genus: Leptogenys
Species: L. diminuta
Binomial name
Leptogenys diminuta
(Smith, F., 1857)

Leptogenys diminuta casent0106010 profile 1.jpg

Leptogenys diminuta casent0106010 dorsal 1.jpg

Specimen labels

Subspecies
Synonyms

Photo Gallery

  • Leptogenys diminuta with milliped prey, India. Photo by Manoj Vembayam.

Identification

A member of the Leptogenys diminuta species group.

Keys including this Species

Distribution

Distribution based on Regional Taxon Lists

Australasian Region: Australia (type locality).
Indo-Australian Region: Borneo, Indonesia, Malaysia (type locality), New Guinea, Philippines, Solomon Islands.
Oriental Region: Bangladesh, India (type locality), Laos, Myanmar (type locality), Nepal, Sri Lanka (type locality), Thailand, Vietnam.
Palaearctic Region: China.


Distribution based on AntMaps

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Distribution based on AntWeb specimens

Check data from AntWeb

Biology

Castes

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • diminuta. Ponera diminuta Smith, F. 1857a: 69 (w.) BORNEO. Forel, 1900d: 312 (q.m.); Wheeler, G.C. & Wheeler, J. 1976a: 49 (l.); Imai, Brown, et al. 1984: 67 (k.); Imai, Kubota, et al. 1985: 46 (k.). Combination in Lobopelta: Mayr, 1862: 734; in Leptogenys: Emery, 1895k: 461. Senior synonym of simillima: Mayr, 1863: 428; of bismarckensis, ferox, stitzi, santschii and material of the unavailable names amboinensis, huruensis referred here: Wilson, 1958a: 118; of yarrabahna: Taylor, 1988: 34. Current subspecies: nominal plus deceptrix, diminutolaeviceps, fruhstorferi, laeviceps, nongnongi, opacinodis, palliseri, sarasinorum, striatula, tjibodana, woodmasoni. See also: Bingham, 1903: 61.
  • simillima. Ponera simillima Smith, F. 1860b: 104 (w.) INDONESIA (Batjan I.). Combination in Lobopelta: Mayr, 1862: 734. Junior synonym of diminuta: Mayr, 1863: 428.
  • ferox. Ponera ferox Smith, F. 1865: 70 (w.) INDONESIA (Salawaty I.). Combination in Leptogenys: Donisthorpe, 1932c: 472. Junior synonym of diminuta: Wilson, 1958a: 118.
  • bismarckensis. Leptogenys (Lobopelta) diminuta var. bismarckensis Forel, 1901b: 7 (w.) NEW GUINEA (Bismarck Archipelago). Junior synonym of diminuta: Wilson, 1958a: 118.
  • papuana. Leptogenys (Lobopelta) diminuta var. papuana Stitz, 1912: 498, fig. 1 (w.) NEW GUINEA. [Junior primary homonym of papuana Emery, above.] Replacement name: stitzi Viehmeyer, 1924b: 310.
  • yarrabahna. Leptogenys (Lobopelta) diminuta var. yarrabahna Forel, 1915b: 29 (w.m.) AUSTRALIA. Junior synonym of diminuta: Taylor, 1988: 34.
  • santschii. Leptogenys (Lobopelta) diminuta subsp. santschii Mann, 1919: 299 (w.q.m.) SOLOMON IS. Junior synonym of diminuta: Wilson, 1958a: 118.
  • stitzi. Leptogenys (Lobopelta) diminuta var. stitzi Viehmeyer, 1924b: 310. Replacement name for papuana Stitz, 1912: 498. [Junior primary homonym of papuana Emery, 1897d: 556.] Junior synonym of diminuta: Wilson, 1958a: 118.
  • hodgsoni. Leptogenys (Lobopelta) diminuta r. hodgsoni Forel, 1900d: 308 (diagnosis in key) (w.) MYANMAR. Raised to species: Bingham, 1903: 62. Junior synonym of diminuta: Xu, Z.-H., He, Q.-J. 2015: 144.
  • sarasinorum. Leptogenys (Lobopelta) diminuta var. sarasinorum Forel, 1900d: 307 (diagnosis in key) (w.) SRI LANKA.
    • As unavailable (infrasubspecific) name: Forel, 1912n: 52.
    • Subspecies of diminuta: Bingham, 1903: 62; Emery, 1911d: 103; Forel, 1913e: 661; Menozzi, 1939a: 329; Chapman & Capco, 1951: 35; Bolton, 1995b: 233 (error).
    • Status as species: Collingwood, 1970: 373.
    • Junior synonym of diminuta: Xu & He, 2015: 138.
  • deceptrix. Leptogenys (Lobopelta) diminuta r. deceptrix Forel, 1901h: 46 (w.) INDIA (no state data).
    • Type-material: syntype workers (number not stated).
    • [Note: the type-material consists of the Indian samples misidentified as diminuta by Forel, 1900d: 312.]
    • Type-localities: India: Coonoor (Daly), Coonoor, Darjeeling, Thana, and Nilgiris (Wroughton), Orissa (Taylor), and Assam (Smythies).
    • Type-depository: MHNG (possibly also in BMNH).
    • As unavailable (infrasubspecific) name: Emery, 1911d: 103.
    • Subspecies of diminuta: Chapman & Capco, 1951: 33; Bolton, 1995b: 230; Bharti & Wachkoo, 2013c: 17 (in key); Bharti, Guénard, et al. 2016: 51 (error).
    • Junior synonym of diminuta: Xu & He, 2015: 138.
  • palliseri. Leptogenys (Lobopelta) diminuta r. palliseri Forel, 1900d: 307 (diagnosis in key) (w.) INDIA (Karnataka).
    • Type-material: syntype workers (number not stated), syntype male.
    • Type-localities: India: Kanara (Palliser), and Kanara (Wroughton).
    • Type-depository: MHNG.
    • Forel, 1900d: 312 (m.).
    • Status as species: Bingham, 1903: 63.
    • Subspecies of diminuta: Emery, 1911d: 103; Forel, 1913k: 17; Karavaiev, 1925c: 279; Wheeler, W.M. 1927e: 1; Wheeler, W.M. 1929g: 59; Wheeler, W.M. 1930a: 95; Chapman & Capco, 1951: 35; Bolton, 1995b: 233; Tiwari, 1999: 23; Bharti & Wachkoo, 2013c: 17 (in key); Bharti, Guénard, et al. 2016: 52 (error).
    • Junior synonym of diminuta: Xu & He, 2015: 138.

Type Material

  • Leptogenys (Lobopelta) diminuta yarrabahna Forel, 1915: Syntype, 1 worker, Yarrabah, Queensland, Australia, Mjoberg, ANIC32-017977, Australian National Insect Collection.
  • Leptogenys (Lobopelta) diminuta yarrabahna Forel, 1915: Syntype, workers, Yarrabah and Bellenden Ker, Queensland, Australia.
  • Ponera diminuta Smith, 1857: Syntype, uncertain, Borneo (Sarawak), Malaysia, museum unknown (probably BMNH (London)).

The following notes on F. Smith type specimens have been provided by Barry Bolton (details):

Ponera diminuta

Two worker syntypes in Oxford University Museum of Natural History. Labelled “SAR. 31” (= Sarawak) and with Donisthorpe type-label.

Ponera ferox

Holotype worker in Oxford University Museum of Natural History. Labelled “S” (= Salawaty I.) and with a Donisthorpe type-label.

Ponera simillima

Two worker syntypes in Oxford University Museum of Natural History. Labelled “Bac” (= Batjan I.) and with a Donisthorpe type-label.

Description

Karyotype

  • 2n = 38 (India; Malaysia) (Goni et al., 1982; Imai et al., 1983; Imai et al., 1984) (in Gona et al. 1982 as Leptogenys sp.1).
  • 2n = 32 (Indonesia) (Imai et al., 1985).

References

  • Arimoto, K., Yamane, S. 2018. Taxonomy of the Leptogenys chalybaea species group (Hymenoptera, Formicidae, Ponerinae) from Southeast Asia. Asian Myrmecology 10, e010008 (DOI 10.20362/am.010008).
  • Bingham, C. T. 1903. The fauna of British India, including Ceylon and Burma. Hymenoptera, Vol. II. Ants and Cuckoo-wasps. London: Taylor and Francis, 506 pp. (page 61, see also)
  • Emery, C. 1895m. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXIII. Formiche di Birmania del Tenasserim e dei Monti Carin raccolte da L. Fea. Parte II. Ann. Mus. Civ. Stor. Nat. 34[=(2(14): 450-483 (page 461, Combination in Leptogenys)
  • Forel, A. 1900f. Les Formicides de l'Empire des Indes et de Ceylan. Part VII. J. Bombay Nat. Hist. Soc. 13: 303-332 (page 312, queen, male described)
  • General, D.E.M., Buenavente, P.A.C., Rodriguez, L.J.V. 2020. A preliminary survey of nocturnal ants, with novel modifications for collecting nocturnal arboreal ants. Halteres 11: 1-12 (doi:10.5281/ZENODO.3707151).
  • Imai, H. T.; Brown, W. L., Jr.; Kubota, M.; Yong, H.-S.; Tho, Y. P. 1984. Chromosome observations on tropical ants from western Malaysia. II. Annu. Rep. Natl. Inst. Genet. Jpn. 34: 66-69 (page 67, karyotype described)
  • Imai, H. T.; Kubota, M.; Brown, W. L., Jr.; Ihara, M.; Tohari, M.; Pranata, R. I. 1985. Chromosome observations on tropical ants from Indonesia. Annu. Rep. Natl. Inst. Genet. Jpn. 35: 46-48 (page 46, karyotype described)
  • Mayr, G. 1862. Myrmecologische Studien. Verh. K-K. Zool.-Bot. Ges. Wien 12: 649-776 (page 734, Combination in Lobopelta)
  • Mayr, G. 1863a. Formicidarum index synonymicus. Verh. K-K. Zool.-Bot. Ges. Wien 13: 385-460 (page 428, Senior synonym of simillima)
  • Smith, F. 1857a. Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace. [part]. J. Proc. Linn. Soc. Lond. Zool. 2: 42-88 (page 69, worker described)
  • Taylor, R. W. 1988. The nomenclature and distribution of some Australian and New Caledonian ants of the genus Leptogenys Roger (=Prionogenys Emery, n. syn.) (Hymenoptera: Formicidae: Ponerinae). Gen. Appl. Entomol. 20: 33-37 (page 34, Senior synonym of yarrabahana)
  • Wheeler, G. C.; Wheeler, J. 1976a. Supplementary studies on ant larvae: Ponerinae. Trans. Am. Entomol. Soc. 102: 41-64 (page 49, larva described)
  • Wilson, E. O. 1958c. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bull. Mus. Comp. Zool. 118: 101-153 (page 118, Senior synonym of bismarchensis, ferox, stitzi and santschii, and material of the unavailable names amboinensis and buruensis referred heere.)
  • Xu, Z. 1996b. A taxonomic study of the ant genus Leptogenys (Hymenoptera: Formicidae) in China. J. Yunnan Agric. Univ. 11: 222-227 (page 224, see also)
  • Xu, Z.-H., He, Q.-J. 2015. Taxonomic review of the ponerine ant genus Leptogenys ROGER, 1861 (Hymenoptera: Formicidae) with a key to the Oriental species. Myrmecological News, 21, 137-161.

References based on Global Ant Biodiversity Informatics

  • Andersen A. N., M. Houadria, M. Berman, and M. van der Geest. Rainforest ants of the Tiwi Islands: a remarkable centr of endemism in Australia's monsoonal tropics. Insectes Sociaux 59: 433-441.
  • Bakhtiar E.Y., and S.L. Chiang. 2010. Leptogenys ants (Hymenoptera: Formicidae: Ponerinae) of Sabah. Serangga 15(1-2): 37-55.
  • Bharti H., Y. P. Sharma, M. Bharti, and M. Pfeiffer. 2013. Ant species richness, endemicity and functional groups, along an elevational gradient in the Himalayas. Asian Myrmecology 5: 79-101.
  • Bui T.V. 2002. Result of ant survey in Tam Dao National Park. Hoi Nghi Con Trung Hoc Toan Quoc, Ha Noi 495-498.
  • CSIRO Collection
  • Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
  • Chapman, J.W. and S.R. Capco. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monographs of the Institute of Science and Technology (Manila) 1: 1- 327
  • Chavhan A., and S.S. Pawar. 2011. Distribution and diversity of ant species (Hymenoptera: Formicidae) in and around Amravati City of Maharashtra, India. World Journal of Zoology 6(4): 395-400.
  • Cheng D., Z. Chen, and S. Zhou. 2015. An analysis on the ant fauna of Jinzhongshan Nature Reserve in Gunagxi, China. Journal of Guangxi Normal University: Natural Science Edition 33(3): 129.137.
  • Chhotani O. B., and P. K. Maiti. 1977. Contribution to the knowledge of Formicidae of the Andaman Islands. Zoological Survey of India 3(1): 17-20.
  • Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
  • Dahl F. 1901. Das Leben der Ameisen im Bismarck-Archipel, nach eigenen Beobachtungen vergleichend dargestellt. Mitt. Zool. Mus. Berl. 2: 1-70.
  • Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
  • Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
  • Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
  • Donisthorpe, Horace. 1943. The Ants of Waigeu Island, North Dutch New Guinea. The Annals and Magazine of Natural History 11 (10): 433-475.
  • Eguchi K., B. T. Viet, and S. Yamane. 2014. Generic Synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part II—Cerapachyinae, Aenictinae, Dorylinae, Leptanillinae, Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae. Zootaxa 3860: 001-046.
  • Eguchi K., T. V. Bui, S. Yamane, H. Okido, and K. Ogata. 2004. Ant faunas of Ba Vi and Tam Dao, North Vietnam (Insecta: Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 27: 77-98.
  • Eguchi K., and S. Yamane. 2003. Species diversity of ants (Hymenoptera, Formicidae) in a lowland rainforest, northwestern Borneo. New Entomol. 52(1,2): 49-59.
  • Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). [concl.]. Ann. Mus. Civ. Stor. Nat. 25(5): 427-473.
  • Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 14:565-580.
  • Emery C. 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1-125.
  • Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
  • Emery, C.. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine)." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 5, no. 25 (1887): 427-473.
  • Emery, C.. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
  • Fayle T. M., D. P. Edwards, E. C. Turner, A. J. Dumbrell, P. Eggleton, and W. A. Foster. 2012. Public goods, public services and by-product mutualism in an ant–fern symbiosis. Oikos 121(8): 1279-1286.
  • Forel A. 1900. Les Formicides de l'Empire des Indes et de Ceylan. Part VII. J. Bombay Nat. Hist. Soc. 13: 303-332.
  • Forel A. 1901. Formiciden aus dem Bismarck-Archipel, auf Grundlage des von Prof. Dr. F. Dahl gesammelten Materials. Mitt. Zool. Mus. Berl. 2: 4-37.
  • Forel A. 1901. Formiciden des Naturhistorischen Museums zu Hamburg. Neue Calyptomyrmex-, Dacryon-, Podomyrma- und Echinopla-Arten. Mitt. Naturhist. Mus. Hambg. 18: 43-82.
  • Forel A. 1901. Nouvelles espèces de Ponerinae. (Avec un nouveau sous-genre et une espèce nouvelle d'Eciton). Revue Suisse de Zoologie 9: 325-353.
  • Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
  • Forel A. 1906. Les fourmis de l'Himalaya. Bulletin de la Société Vaudoise des Sciences Naturelles 42: 79-94.
  • Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
  • Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
  • Forel, A. 1908. Fourmis de Ceylan et d'Égypte récoltées par le Prof. E. Bugnion. Lasius carniolicus. Fourmis de Kerguelen. Pseudandrie? Strongylognathus testaceus. Bull. Soc. Vaudoise Sci. Nat. 44: 1-22
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Hashimoto Y., S. Yamane, and T. Itioka. 1997. A preliminary study on dietary habits of ants in a Bornean rain forest. Japanese Journal of Entomology 65(4): 688-695.
  • Hashimoto Y., Y. Morimoto, and M. Mohamed. 2003. Species List of Ground and Leaf Litter Ants Collected in Lower Kinabatangan. Pp 13-18. In Lower Kinabatangan Scientific Expedition 2002, 176 pp. ISBN-13: 983-2369-11-8
  • Hu C.-H. 2006. Indigenized conservation and biodiversity maintenance on Orchid Island. PhD Thesis, graduate school of the University of Minnesota. 150 pages.
  • Hua Li-zhong. 2006. List of Chinese insects Vol. IV. Pages 262-273. Sun Yat-sen university Press, Guangzhou. 539 pages.
  • Huong N. T. T., P. V. Sang, and B. T. Viet. 2015. A preliminary study on diversity of ants (Hymenoptera: Formicidae) at Hon Ba Nature Reserve. Environmental Scientific Conference 7: 614-620.
  • IZIKO South Africa Museum Collection
  • Imai H. T., M. Kubota, W. L. Brown, Jr., M. Ihara, M. Tohari, and R. I. Pranata. 1985. Chromosome observations on tropical ants from Indonesia. Annu. Rep. Natl. Inst. Genet. Jpn. 35: 46-48.
  • Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
  • Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
  • Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
  • Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
  • Janda M., G. D. Alpert, M. L. Borowiec, E. P. Economo, P. Klimes, E. Sarnat, and S. O. Shattuck. 2011. Cheklist of ants described and recorded from New Guinea and associated islands. Available on http://www.newguineants.org/. Accessed on 24th Feb. 2011.
  • Kishimoto-Yamata K., F. Hyodo, M. Matsuoka, Y. Hashimoto, M. Kon, T. Ochi, S. Yamane, R. Ishii, and T. Itioka. 2012. Effects of remnant primary forests on ant and dung beetle species diversity in a secondary forest in Sarawak, Malaysia. Journal of Insect Conservation DOI 10.1007/s10841-012-9544-6
  • Li Q., Y. Chen, S. Wang, Y. Zheng, Y. Zhu, and S. Wang. 2009. Diversity of ants in subtropical evergreen broadleaved forest in Pu'er City, Yunnan. Biodiversity Science 17(3): 233-239.
  • Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
  • Lu Z., B. D. Hoffmann, and Y. Chen. 2016. Can reforested and plantation habitats effectively conserve SW China’s ant biodiversity? Biodivers. Conserv. DOI 10.1007/s10531-016-1090-1
  • Lucky A., E. Sarnat, and L. Alonso. 2011. Ants of the Muller Range, Papua New Guinea, Chapter 10. In Richards, S. J. and Gamui, B. G. (editors). 2013. Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: surveying the biodiversity of Papua New Guinea’s sublime karst environments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA.
  • Mann W. M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology 63:273-391.
  • Mann William. 1916. The Ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology at Harvard College 63(7): 273-391
  • Mann, W.M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology of Harvard College 63: 273-391
  • Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
  • Menozzi C. 1930. Formiche di Sumatra raccolte dal Prof. J. C. van der Meer Mohr. Miscellanea Zoologica Sumatrana 47: 1-5.
  • Mohamed M. 1995. A preliminary list of ants (Hymenoptera: Formicidae) of the Tawau Hills Park, Sabah. In: Ghazally Ismail et al. (eds.), Tawau Hills Park, Sabah, Pelanduk Pub. Pp. 205-213.
  • Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
  • Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
  • Narendra A., H. Gibb, and T. M. Ali. 2011. Structure of ant assemblages in Western Ghats, India: role of habitat, disturbance and introduced species. Insect Conservation and diversity 4(2): 132-141.
  • Ogata K. 2005. Asian ant inventory and international networks. Report on Insect inventory Project in Tropic Asia TAIIV: 145-170.
  • Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
  • Radchenko A. G. 1993. Ants from Vietnam in the collection of the Institute of Zoology, PAS, Warsaw. I. Pseudomyrmicinae, Dorylinae, Ponerinae. Annales Zoologici (Warsaw) 44: 75-82.
  • Sakchoowong W., W. Jaitrong, and K. Ogata. 2009. Comparison of ground-ant diversity between natural forests and disturbed forests along a natural gas pipeline transect in Thong Pha Phum National Park, Kanchanaburi province. Kasetsart J. (Nat. Sci) 43: 64-73.
  • Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
  • Sitthicharoenchai D., and N. Chantarasawat. 2006. Ant species diversity in the establishing area for Advanced Technology Institute at Lai-Nan Sub-district, Wiang Sa district, Nan Province, Thailand. The Natural History Journal of Chulalongkorn University 6(2): 67-74
  • Smith F. 1865. Descriptions of new species of hymenopterous insects from the islands of Sumatra, Sula, Gilolo, Salwatty, and New Guinea, collected by Mr. A. R. Wallace. Journal and Proceedings of the Linnean Society of London. Zoology 8: 61-94.
  • Smith, F.. "Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace." Journal of the Proceedings of the Linnean Society of London, Zoology 2 (1857): 42-88.
  • Smith, Fr.. "Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the Islands of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea." Journal of the Proceedings of the Linnean Society of London, Zoology 5 (1860): 93-143.
  • Snelling R. R. 1998. Insect Part 1: The social Hymenoptera. In Mack A. L. (Ed.) A Biological Assessment of the Lakekamu Basin, Papua New Guinea, RAP 9. 189 ppages
  • Sonune B. V., and R. J. Chavan. 2016. Distribution and diversity of ants (Hymenoptera: Formicidae) around Gautala Autramghat Sanctuary, Aurangabad Maharashtra, India. Journal of Entomology and Zoology Studies 4(2): 361-364.
  • Stitz H. 1912. Ameisen aus Ceram und Neu-Guinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1912: 498-514.
  • Suriyapong Y. 2003. Study of ground dwelling ant populations and relationship to some ecological factors in Sakaerat environmental research station, Nakhon Ratchasima. PhD thesis Suranaree University of Technology 203 pages.
  • Taylor R. W., and D. R. Brown. 1985. Formicoidea. Zoological Catalogue of Australia 2: 1-149. 
  • Terayama M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta: Hymenoptera). Research Bulletin of Kanto Gakuen University. Liberal Arts 17:81-266.
  • Terayama, M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta; Hymenoptera). The Research Bulletin of Kanto Gakuen University 17: 81-266.
  • Tiwari R. N., B. G. Kundu, S. Roy Chowdhury, and S. N. Ghosh. 2003. Insecta: Hymenoptera: Formicidae. Fauna of Sikkim. Part 4. State Fauna Series. 9.Zool.Surv.India. i-iii, 1-512. Chapter pagination: 467-506.
  • Tiwari R.N., B.G. Kundu, S. Roychowdhury, S.N. Ghosh. 1999. Insecta: Hymenoptera: Formicidae. Pp. 211-294 in: Director; Zoological Survey of India (ed.) 1999. Fauna of West Bengal. Part 8. Insecta (Trichoptera, Thysanoptera, Neuroptera, Hymenoptera and Anoplura). Calcutta: Zoological Survey of India, iv + 442 pp.
  • Tiwari, R.N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4):1-96
  • Trainor C.R. and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
  • Trainor C.R., and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
  • Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
  • Viehmeyer H. 1914. Ameisen aus Perak, Bali und Ceram (Hym.) (Freiburger Molukken-Expedition), gesammelt von E. Streesemann. Entomologische Mitteilungen. Berlin-Dahlem 3: 112-116.
  • Wang C. and Wu J.. 1992. Ants of the Jianfengling forest region in Hainan Province (Hymenoptera: Formicidae). Scientia Silvae Sinicae 28: 561-564.
  • Wheeler W. M. 1909. Ants of Formosa and the Philippines. Bulletin of the American Museum of Natural History 26: 333-345.
  • Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
  • Wheeler W. M. 1921. Chinese ants collected by Prof. C. W. Howard. Psyche (Cambridge) 28: 110-115.
  • Wheeler W. M. 1927. A few ants from China and Formosa. American Museum Novitates 259: 1-4.
  • Wheeler W. M. 1930. A list of the known Chinese ants. Peking Natural History Bulletin 5: 53-81.
  • Wheeler W. M., and J. W. Chapman. 1925. The ants of the Philippine Islands. Part I, Dorylinae and Ponerinae. Philipp. J. Sci. 28: 47-73.
  • Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
  • Wheeler, William Morton.1935.Checklist of the Ants of Oceania.Occasional Papers 11(11): 3-56
  • Wilson E. O. 1958. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bulletin of the Museum of Comparative Zoology 118: 101-153.
  • Wilson E.O. 1959. Adaptive shift and dispersal in a tropical ant fauna. Evolution 13(1): 122-144.
  • Wilson Edward O. 1959. Adaptive Shift and Dispersal in a Tropical Ant Fauna. Evolution 13(1): 122-144
  • Xu Z. H., and Q. J. He. 2015. Taxonomic review of the ponerine ant genus Leptogenys ROGER, 1861 (Hymenoptera: Formicidae) with a key to the Oriental species. Myrmecological News 21: 137-161.
  • Xu Z. 2000. Five new species and one new record species of the ant genus Leptogenys Roger (Hymenoptera: Formicidae) from Yunnan province, China. Entomologia Sinica 7(2): 117-126
  • Yamane S.; Bui T. V.; Ogata K.; Okido H.; Eguchi K. 2002. Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture Kyushu University 25: 51-62.
  • Yamane S.; Nona, A. R. 1994. Ants from Lambir Hills National Park, Sarawak. Pp. 222-226 in: Inoue, T.; Hamid, A. A. (eds.) 1994. Plant reproductive systems and animal seasonal dynamics. Long-term study of dipterocarp forests in Sarawak. Kyoto: Center for Ecological Research, Kyoto University, vii + 255 pp. 
  • Zhao S., F. L. Jia, G. Q. Liang, Y. L. Ke, W. J. Tian. 2009. Ants and their distribution in Guangdong Province, China. Journal of Environmental Entomology 31(2): 156-161.
  • Zhou S., Y. Chen, Z. Chen, P. Zhou, D. Ban, and M. Huang. 2012. Two new species of the genus Leptogenys from Guangxi, China (Hymenoptera: Formicidae). Sociobiology 59: 885-892.
  • Zhou S.-Y. 2001. Ants of Guangxi. Guangxi Normal University Press, Guilin, China, Guilin, China. 255 pp.
  • Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.