Camponotus herculeanus sachalinensis

Camponotus herculeanus sachalinensis
Scientific classification (junior synonym of Camponotus herculeanus)
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Formicinae
Tribe:	Camponotini
Genus:	Camponotus
Species:	Camponotus herculeanus sachalinensis Forel, 1904 Specimen Labels

This taxon is not in use as it is currently considered to be a junior synonym of Camponotus herculeanus.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

• sachalinensis. Camponotus herculeanus var. sachalinensis Forel, 1904b: 38 (q.) MONGOLIA, RUSSIA.
  • Karavaiev, 1912b: 592 (m.).
  • Combination in C. (Camponotus): Emery, 1925b: 72.
  • As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 221.
  • Junior synonym of saxatilis: Kuznetsov-Ugamsky, 1928b: 4; Kuznetsov-Ugamsky, 1929a: 18.
  • Status as species: Collingwood, 1981: 29; Bolton, 1995b: 121; Collingwood & Heatwole, 2000: 12; Imai, et al. 2003: 39.
  • Subspecies of herculeanus: Ruzsky, 1905b: 222; Wheeler, W.M. 1906c: 325; Yano, 1910: 422; Karavaiev, 1912b: 592; Ruzsky, 1926: 108; Arnol'di, 1967: 1821; Pisarski, 1969a: 230; Pisarski, 1969b: 304; Dlussky & Pisarski, 1970: 86; Collingwood, 1976: 306; Pisarski & Krzysztofiak, 1981: 159; Kupyanskaya, 1986b: 96; Kupyanskaya, 1990: 166; Morisita, et al. 1991: 41; Radchenko, 1994b: 116 (in key); Radchenko, 1996b: 1203 (in key); Radchenko, 1997a: 555; Radchenko, 2005b: 158.
  • Senior synonym of altaica: Arnol'di, 1967: 1821.
  • Senior synonym of jacuticus: Arnol'di, 1967: 1821.
  • Junior synonym of herculeanus: Yasumatsu & Brown, 1957: 49; Schar et al., 2018: 6.

Taxonomic Notes

Aibek & Yamane (2009) - Various authors have treated this form as a subspecies of C. herculeanus or as a distinct species. Collingwood (1981) first raised this form to distinct species rank, but he did not mention any difference between the two. Radchenko (2005) argued that it is a subspecies of C. herculeanus, since sachalinensis and the typical form differ only in coloration, and intermediate specimens can be collected in Altai and Tuva. He mentioned that the two forms are indistinguishable also in the queen and male. The typical C. herculeanus is distinctly bicolored; especially in the queen caste the lateral part of mesonotum, whole propodeum, petiole and legs are reddish brown, while the queen of sachalinensis is almost wholly black except for legs. In the typical form, even the major worker has reddish brown alitrunk, petiole and legs, while the major worker of sachalinensis has a wholly black alitrunk; only propodeal declivity, petiole and legs are tinged with brown or red. Furthermore, in the worker of the typical form from Finland the pubescence on gastral tergites is longer and denser than in sachalinensis. Although the difference is minor, we tentatively follow Collingwood (1981) and Bolton (1995a) for the above-mentioned reason (see also Japanese Ant Database Group, 2003 for the Japanese population).

References

• Aibek, U., Yamane, S. 2009. Taxonomic review of the genus Camponotus (Hymenoptera, Formicidae, Formicinae) from Mongolia. Biogeography 11, 97-108.
• Arnol'di, K. V. 1967. New data on the ant genus Camponotus (Hymenoptera, Formicidae) from the USSR fauna. 1. Camponotus (s. str.). Zool. Zh. 4 46: 1815-1830 (page 1821, Revived from synonymy as subspecies of herculeanus)
• Arnol'di, K. V. 1967. New data on the ant genus Camponotus (Hymenoptera, Formicidae) from the USSR fauna. 1. Camponotus (s. str.). Zool. Zh. 4 46: 1815-1830 (page 1821, Senior synonym of altaica, jacuticus)
• Collingwood, C. A. 1976a. Ants (Hymenoptera: Formicidae) from North Korea. Ann. Hist.-Nat. Mus. Natl. Hung. 68: 295-309 (page 306, Subspecies of herculeanus)
• Collingwood, C. A. 1981. Ants (Hymenoptera: Formicidae) from Korea, 2. Folia Entomol. Hung. 42[3 34: 25-30 (page 29, Raised to species)
• Emery, C. 1925d. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183: 1-302 (page 72, Combination in C. (Camponotus))
• Forel, A. 1904d. Miscellanea myrmécologiques. Rev. Suisse Zool. 12: 1-52 (page 38, queen described)
• Karavaiev, V. 1912b. Ameisen aus dem paläarktischen Faunengebiete. Rus. Entomol. Obozr. 12: 581-596 (page 592, male described)
• Kupyanskaya, A. N. 1990a. Ants of the Far Eastern USSR. Vladivostok: Akademiya Nauk SSSR, 258 pp. (page 166, Subspecies of herculeanus)
• Kuznetsov-Ugamsky, N. N. 1928b. Ants of the South Ussuri Region. Zap. Vladivost. Otd. Gos. Rus. Geogr. Obshch. 1(1 18: 1-47 (page 4, Provisional junior synonym of saxatilis)
• Kuznetsov-Ugamsky, N. N. 1929a. Die Ameisen des Süd-Ussuri-Gebietes. Zool. Anz. 83: 16-34 (page 18, Provisional junior synonym of saxatilis)
• Radchenko, A. G. 1996e. A key to the ant genus Camponotus (Hymenoptera, Formicidae) in Palearctic Asia. Zool. Zh. 7 75: 1195-1203 (page 1203, Subspecies of herculeanus)
• Ruzsky, M. 1926. A systematic list of the ants found in Siberia. I. Review of the species of the genera Camponotus (s. ext.) and Formica (s. str.). Izv. Tomsk. Gos. Univ. 77: 107-111 (page 108, Subspecies of herculeanus)
• Schar, S., Talavera, G., Espadaler, X., Rana, J.D., Andersen, A.A., Cover, S.P., Vila, R. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 2018:1–12 (doi:10.1111/jbi.13380).
• Yasumatsu, K.; Brown, W. L., Jr. 1957. A second look at the ants of the Camponotus herculeanus group in eastern Asia. J. Fac. Agric. Kyushu Univ. 11: 45-51 (page 49, Junior synonym of herculeanus)