Leptanilla copiosa

Leptanilla copiosa
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hymenoptera
Family:	Formicidae
Subfamily:	Leptanillinae
Tribe:	Leptanillini
Genus:	Leptanilla
Species:	L. copiosa
Binomial name
	Leptanilla copiosa; (Petersen, 1968)

Described from two males capture in a malaise trap. Nothing is known about this species.

Identification

Griebenow (2024), Figure 57. Position of the male anteromedian ocellus relative to the compound eye in Leptanilla, diagrammatic, after Griebenow (2020: fig. 12B). A, Leptanilla copiosa. B, Leptanilla zhg-my10.

Keys including this Species

Keys to Leptanillinae

Distribution

Distribution based on Regional Taxon Lists

Indo-Australian Region: Philippines (type locality).

Distribution based on AntMaps

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.

Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.

Biology

Castes

Known only from males.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

copiosa. Noonilla copiosa Petersen, 1968: 585, figs. 6-10 (m.) PHILIPPINES.
- In Leptanillinae: Boudinot, 2015: 32.
- Combination in Leptanilla: Griebenow, 2024: 129.

Type Material

Holotype, male. Philippines, PALAWAN: Mantalingajan Range, Pinigisan, 600 m., 13 September, 1961. Caught in a Malaise trap outside primary forest.
Paratype, male, same locality, but 6 September and caught in a trap inside primary forest. Types in the Zoological Museum, Copenhagen.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Male

Petersen 1968. Figures 6.

Petersen 1968. Figures 7-10.

Length of holotype without antennae 3.6 mm, without protruding portion of genitalia 2.9 mm; antennae 2.6 mm; fore wing 2.3 mm.

Colour of the alcohol preserved specimens mainly various shades from light to medium brown; yellowish are antennae especially pedicellus, mandibles, fore tarsi in strong contrast to other parts of fore legs, mid and hind legs apart from coxae, and genitalia apart from volsellar digiti.

Sculpture of any conspicous kind lacking.

Pilosity also inconspicuous. Semi-erect setae of moderate length and density present all over, longest at apices of terga, increasing in length caudally. Antennal setae almost as long as diameter of joints; setae at outline of head and on mandibles are shown in fig. 7 and pilosity scheme of genitalia is sketched in figs. 9 and 10.

Head seen from in front almost circular (fig. 7), measured over the eyes somewhat broader than high (ratio 8:7), behind the eyes only slightly narrower than high. Eyes almost hemispheric, strongly bulging, hind margin straightened; ratio of longest and shortest diameter 6:5; distance between eyes about half as long as width of head. Ocelli large and prominent, distance from front ocellus to lateral ocelli hardly as long as ocellar diameter, distance between lateral ocelli 2.5 times as long as ocellar diameter or about as long as their distance to occiput. Rim of antennal sockets prominent. Antennae almost as long as head, thorax and abdomen (without genitalia) combined; relative lengths of scape, pedicel, 1st., 2nd., 10th and 11th flagellar joints 32:12:37:31:25:36, relative widths 9:7:6:6:5:5; scape thus slightly longer than 2nd flagellar joint, flagellar joints decrease in size distally, apical joint, however, considerably longer than preceding ones and about as long as joint 1. Frontoclypeal area not differentiated either by sutures or carinae; shape of clypeal margin shown in fig. 7. Mandibles finger-shaped, twice as long as malar space or half as long as 1st flagellar joint, densely setaceous at apices. Labial and maxillary palpi almost equal in size, ovoid.

Thorax twice as long as broad and 1.5 times as long as high; relative lengths of pronotum, mesonotum, scutellum and propodeum measured at mid-line 8:38:18:18. Lateral view of thorax shown in fig. 6; ventral view of propleura and prosternum sketched in fig. 8.

Legs very long and slender, e. g. hind legs as long as combined length of head, thorax and abdomen without genitalia. Approximate lengths of various parts of legs are measurable from fig. 6. Fore coxae in ventral view a little more than twice as long as wide at apex (fig. 8); fore femora curved and strong, length slightly morc than four times greatest width.

Fore wing 2.5 times longer than wide; proportions of venation may be measured from fig. 6. Hind wing about 7 times longer than wide, less than half as long as fore wing; 3 long hamuli present .distally to mid-length of wing.

Abdomen strongly curved as shown in fig. 6, probably also in life. Segment 2 (petiole) almost twice as long as greatest diameter of posterior cylindrical portion; spiracles placed in anterior one third of petiole, interdistance as long as petiolar diameter. Lengths of terga 3-7 at mid-line almost equal, slightly more than half length of attenuated, apically rounded tergum 8. Sternum 7 half as long as preceding sterna; sternum 8 and segment 9 described in the diagnosis of the genus. Genitalia as long as thorax, measurements may be taken from figs. 9 and 10.

References

Boudinot, B.E. 2015. Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): a new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages. European Journal of Taxonomy 120, 1-62 (http://dx.doi.org/10.5852/ejt.2015.120).
Griebenow, Z. 2020. Delimitation of tribes in the subfamily Leptanillinae (Hymenoptera: Formicidae), with a description of the male of Protanilla lini Terayama, 2009. Myrmecological News 30: 229-250. (doi:10.25849/MYRMECOL.NEWS_030:229).
Griebenow, Z.H. 2021. Synonymisation of the male-based ant genus Phaulomyrma (Hymenoptera:Formicidae) with Leptanilla based upon Bayesian total-evidence phylogenetic inference. Invertebrate Systematics 35, 603–636 (doi:10.1071/is20059).
Griebenow, Z. 2024. Systematic revision of the ant subfamily Leptanillinae (Hymenoptera, Formicidae). ZooKeys 1189, 83–184 (doi:10.3897/zookeys.1189.107506).
Petersen, B. 1968. Some novelties in presumed males of Leptanillinae (Hym., Formicidae). Entomol. Medd. 36: 577-598 (page 585, figs. 6-10 male described)

References based on Global Ant Biodiversity Informatics

Baroni Urbani C. 1977. Materiali per una revisione della sottofamiglia Leptanillinae Emery (Hymenoptera: Formicidae). Entomologica Basiliensia 2: 427-488.
General D. M., and G. D. Alpert. 2012. A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. Zookeys 200: 1-111.
Petersen B. 1968. Some novelties in presumed males of Leptanillinae (Hym., Formicidae). Entomologiske Meddelelser 36: 577-598.