Stigmatomma pallipes

In California this species has been found in chaparral, riparian woodland, oak woodland and mixed forest (with oak, douglas fir and pine). Isolated workers or nest fragments have been collected in litter samples and under stones. Because of their subterranean habits the workers are infrequently encountered, but males are not uncommon at light in late summer in some foothill localities. Foraging behavior is probably similar to that of S. pallipes workers from eastern United States, which have been reported feeding on centipedes and other soil arthropods. (Ward 1988)

Identification
Worker: inner border of mandible and anterior clypeal margin broadly convex in outline; California specimens smaller in size than those of Stigmatomma oregonense (HW 0.99-1.15), with narrower heads (CI 0.86-0.90).

Stigmatomma pallipes in the eastern United States is not a common ant but it can be discovered by turning over stones and searching carefully through rotten wood on the ground. These small, dark reddish-brown ants are about 1/4 inch in length. The eyes are very small, with only one or two facets. There is a row of small teeth on the anterior margin of the clypeus and the mandibles are long and linear with a row of bi-dentate teeth on the inner margin. The petiole is broadly joined to the gaster.

There is only one other eastern species, the rare Stigmatomma trigonignathum, which lacks the sharp projections that flank the outer bases of the mandibles in S. pallipes, and has the two most basal large teeth mounted on an inward expansion of the mandible. In the West, Stigmatomma oregonense, which is similar to S. pallipes, has an overlapping distribution in part of northern California (Ward 1988).

Distribution
This species is widely distributed in the United States. It is known from Quebec, west to Wisconsin, south through Florida (Smith 1979) and then west and south, possibly in a series of disjunct populations, into the Coastal Range of California (Ward 1988). It found sporadically in the mountains of Arizona.

In the eastern part of its range the habitat of S. pallipes is usually moist woodland with a heavy canopy, but as Creighton pointed out (1950), the species may be subterranean, and therefore unobserved, in more open habitats. In Florida S. pallipes is usually found in sandy woodlands with thick leaf litter. It is seldom found in wet areas, such as the edges of marshes and swamp forests.

Distribution based on Regional Taxon Lists
Nearctic Region: Canada, United States.

Habitat
The small colonies are most common in wooded areas where they are found in rotten logs and under stones. In New England these ants are often found nesting under rock walls.

Abundance
This ant species is seldom if ever abundant. Due to their small colony size and nest location under rocks and in logs in forests, this ant species is difficult to find.

Biology
The workers are slow moving and not regularly encountered. The pupae are covered with a brown silk case that helps in identification in the field. The queen is worker-like and inconspicuous. The males and females mate during the late summer months. Females mate with males on rocks on the ground in forested areas during the first week of September in New England.

Nests are small, often far fewer than 100 workers, and diffuse. In the eastern United States workers have been found collecting centipedes and other soil arthropods. Prey are captured and held in the worker's heavily toothed mandibles, then paralyzed by a powerful sting. The crytpic foragers are not regularly seen above ground as they prefer to search for food in the same places where this species' forms their nests - in the soil, litter and decaying wood. Stigmatomma pallipes is a forest dweller throughout most of its range. In California this species has been found in chaparral, riparian woodland, oak woodland and mixed forest (oak, Douglas fir and pine) habitats. Isolated workers or nest fragments have been collected in litter samples and under stones.

Haskins (1924) provides this account of their nesting biology: ....ordinarily found in thick, damp woodlands, the typical localities in which most timid hypogeaic forms find refuge. Although the majority of these forms seem dependent for their continued existence upon these forests, with their concomitant opportunity of escape from more dominant types, it is certain that pallipes has not become entirely so, nor has it completely lost the ability to exist in proximity with glade or even field forms. In Petersham, Massachusetts, a colony taken under a broad stone in an open clump of young white pine, which was nesting beside two species of open-woods ants (Lasius americanus and Aphaenogaster picea).

More than one individual may assist in subduing prey, but only one transports the prey. Back at the nest, the larvae move onto the prey or are carried to it. The prey is not dismembered. Workers do not regurgitate food for larvae or for each other. The queen supplements her diet by periodically squeezing a larva until a drop of clear liquid is regurgitated. In most ways, queens and workers show few behavioral differences, except that functional queens do not forage. Workers do not change tasks as they mature. The organization of the colony is not completely understood, but circumstantial evidence suggests that groups of 9-16 workers form small colonies with one or more queens, and these colonies in turn fission when the group increases after the emergence of reproductives and workers in late summer: Within an area, the workers from apparently isolated colonies are compatible. Alates leave the nest to mate, but females usually return to the nest after mating. Alate females probably occasionally disperse and found new colonies removed from the parental population.

New Mexico
This species nests under stones in moist areas. Colonies apparently consist of only a few workers. They prey primarily on chilopods. These secretive ants are rarely collected, but well worth the search (Mackay and Mackay 2002)

Foraging/Diet
The highly modified mandibles appear to be specialized for hunting centipedes. Predators of centipedes and beetle larvae in forest litter. James Trager observed workers and larvae of this species under a stone in Missouri USA, feeding on an Elateridae (beetle) larva, which appeared to have been killed by them (fresh, no external damage evident on the larva, except where the larvae were chewing into the intersegmental membranes).

Associations with other Organisms
It is possible that Strumigenys species are most often found under the same stone.

Castes
Peeters & Molet (2010) - Unlike in Amblyopone australis, queens and workers of S. pallipes are similar in body size, and workers show no variation in size.

Common Name
Pale Footed Primitive Ant

Nomenclature

 * . Typhlopone pallipes Haldeman, 1844: 54 (w.) U.S.A. (no state data).
 * Type-material: syntype (?) workers.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: U.S.A.: (no further data; the only information supplied is “found in old stumps in June”).
 * Type-depository: unknown (no type-material known to exist).
 * [Note: Creighton, 1940b: 4, comments that he examined two ‘autotypes’ that had been sent by Haldeman to Harris, which were deposited in the Boston Society of Natural History (now MSBM). While not syntypes, these were probably specimens collected and identified by Haldeman himself.]
 * [Unjustified emendation of spelling to pallidipes: Dalla Torre, 1893: 14; Santschi, 1914a: 429.]
 * Emery, 1895c: 261 (q.m.); Wheeler, W.M. 1900c: 61 (l.); Wheeler, G.C. & Wheeler, J. 1952a: 114 (l.).
 * Combination in Amblyopone: Haldeman, 1849a: 201 (footnote); Cresson, 1887: 259; Brown, 1960a: 169;
 * combination in Stigmatomma (Amblyopone): Wheeler, W.M. 1900c: 56;
 * combination in Stigmatomma: Roger, 1863b: 20; Emery, in Dalla Torre, 1893: 14; Yoshimura & Fisher, 2012a: 19.
 * Status as species: Roger, 1863b: 20; Mayr, 1863: 457; Mayr, 1886d: 439; Cresson, 1887: 259; Dalla Torre, 1893: 14; Emery, 1895c: 261; Wheeler, W.M. 1900c: 56; Wheeler, W.M. 1904e: 299; Wheeler, W.M. 1905f: 374; Wheeler, W.M. 1906b: 1; Wheeler, W.M. 1908e: 399; Wheeler, W.M. 1910g: 561; Emery, 1911d: 24; Wheeler, W.M. 1913c: 112; Wheeler, W.M. 1917i: 460; Smith, M.R. 1928b: 243; Smith, M.R. 1930a: 2; Wheeler, W.M. 1932a: 2; Dennis, 1938: 275; Wesson, L.G. & Wesson, R.G. 1940: 90; Creighton, 1940b: 3 (redescription); Brown, 1949c: 84; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Cole, 1953c: 84; Smith, M.R. 1958c: 110; Whelden, 1958: 1; Brown, 1960a: 169, 183, 192 (in key); Carter, 1962a: 6 (in list); Francoeur & Béique, 1966: 141; Smith, M.R. 1967: 346; Hunt & Snelling, 1975: 20; Francoeur, 1977b: 206; Francoeur, 1979: 34 (redescription); Smith, D.R. 1979: 1335; Traniello, 1982: 65; Wheeler, G.C. & Wheeler, J. 1986g: 17; DuBois & LaBerge, 1988: 135; Ward, 1988: 105; Deyrup, et al. 1989: 92; Lattke, 1991c: 6 (in key); Wheeler, G.C., et al. 1994: 302; Bolton, 1995b: 62; Lacau & Delabie, 2002: 40 (in key); Mackay & Mackay, 2002: 31; Deyrup, 2003: 44; Coovert, 2005: 23; MacGown & Forster, 2005: 67; Ward, 2005: 25; Ellison, et al. 2012: 89; Deyrup, 2017: 13.
 * Senior synonym of arizonense: Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Senior synonym of binodosus: Emery, 1895c: 261; Wheeler, W.M. 1908e: 400; Emery, 1911d: 24; Wheeler, W.M. 1911f: 159; Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Francoeur & Béique, 1966: 141; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Senior synonym of montigena: Brown, 1949c: 84; Brown, 1960a: 169; Smith, M.R. 1967: 346; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Senior synonym of serratum: Mayr, 1886d: 439; Emery, 1895c: 261; Emery, 1911d: 24; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Senior synonym of subterranea: Brown, 1960a: 169; Smith, M.R. 1967: 346; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Senior synonym of wheeleri: Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * Distribution: Canada, U.S.A.
 * arizonense. Stigmatomma pallipes subsp. arizonense Wheeler, W.M. 1915b: 389 (w.) U.S.A. (Arizona).
 * Type-material: holotype worker.
 * Type-locality: U.S.A.: Arizona, Huachuca Mts, Ramsay Cañon, 5800 ft (W.M. Mann).
 * Type-depository: MCZC.
 * Subspecies of pallipes: Cole, 1937a: 98.
 * Junior synonym of pallipes: Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 61.
 * binodosus. Arotropus binodosus Provancher, 1881a: 206, figs. 32, 33 (q.) CANADA (no state data, probably Québec).
 * Type-material: holotype male.
 * [Note: the holotype was misprinted with a queen symbol in the original description, but was actually a male (Provancher, 1887: 240, 241). Lectotype designation of a worker by Gahan & Rohwer, 1917: 306, is set aside as no worker was represented in the original description. Lectotype male designation by Francoeur & Béique, 1966: 141, appears unnecessary as Provancher described only a single specimen, automatically the holotype.]
 * Type-locality: Canada (no further data) (L. Provancher).
 * [Note: type-locality was probably Québec, Cap Rouge (Provancher, 1887: 241).]
 * Type-depository: QMNH.
 * [Misspelled as binodus by Creighton, 1940b: 3.]
 * Provancher, 1887: 240 (w.).
 * Combination in Amblyopone: Provancher, 1887: 240;
 * combination in Stigmatomma: Emery, in Dalla Torre, 1893: 14.
 * Status as species: Provancher, 1887: 240; Cresson, 1887: 258; Dalla Torre, 1893: 14.
 * Junior synonym of pallipes: Emery, 1895c: 261; Wheeler, W.M. 1908e: 400; Emery, 1911d: 24; Wheeler, W.M. 1911f: 159; Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Francoeur & Béique, 1966: 141; Smith, D.R. 1979: 1335; Bolton, 1995b: 61.
 * montigena. Stigmatomma pallipes subsp. montigena Creighton, 1940b: 7, figs. 6, 8 (w.q.) U.S.A. (North Carolina).
 * Type-material: holotype ergatoid queen, paratype workers (number not stated).
 * Type-locality: holotype U.S.A.: North Carolina, Little Switzerland, nr Spruce Pines (W.S. Creighton); paratypes with same data.
 * Type-depositories: AMNH (holotype); AMNH, LACM, MCZC (paratypes).
 * Subspecies of pallipes: Creighton, 1950a: 33; Smith, M.R. 1951a: 782.
 * Junior synonym of pallipes: Brown, 1949c: 84; Brown, 1960a: 169; Smith, M.R. 1967: 346; Smith, D.R. 1979: 1335; Bolton, 1995b: 62.
 * serratum. Stigmatomma serratum Roger, 1859: 251 (w.) U.S.A. (Maryland).
 * Type-material: holotype worker.
 * Type-locality: U.S.A.: Maryland, Baltimore (no collector’s name).
 * Type-depository: MNHU.
 * Status as species: Roger, 1861a: 53; Roger, 1863b: 20; Mayr, 1863: 454.
 * Junior synonym of pallipes: Mayr, 1886d: 439; Emery, 1895c: 261; Emery, 1911d: 24; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 63.
 * subterranea. Stigmatomma pallipes subsp. subterranea Creighton, 1940b: 8, fig. 4 (w.) U.S.A. (Kansas).
 * Type-material: holotype worker, 14 paratype workers.
 * Type-locality: holotype U.S.A.: Kansas, Elmo, 6.ix.1927 (W.S. Creighton); paratypes with same data.
 * Type-depositories: AMNH (holotype); AMNH, LACM, MCZC (paratypes).
 * Status as species: Brown, 1949c: 85.
 * Subspecies of pallipes: Buren, 1944a: 279; Creighton, 1950a: 34; Smith, M.R. 1951a: 783; Smith, M.R. 1958c: 110.
 * Junior synonym of pallipes: Brown, 1960a: 169; Smith, M.R. 1967: 346; Smith, D.R. 1979: 1335; Bolton, 1995b: 63.
 * wheeleri. Stigmatomma pallidipes [sic] var. wheeleri Santschi, 1914a: 429 (w.q.m.) U.S.A. (Connecticut).
 * Type-material: syntype workers, syntype queens, syntype males (numbers not stated).
 * Type-locality: U.S.A.: Connecticut, Colebrook (W.M. Wheeler).
 * Type-depository: MCZC, NHMB.
 * Subspecies of pallipes: Wheeler, W.M. 1916m: 581.
 * Junior synonym of pallipes: Creighton, 1940b: 3; Creighton, 1950a: 33; Smith, M.R. 1951a: 783; Brown, 1960a: 169; Smith, D.R. 1979: 1335; Bolton, 1995b: 63.

Worker
Minutely punctured, dull reddish-brown, terminal segments of the abdomen, antennae, and mandibles, paler; legs testaceous. 6 mill. long. Head subquadrate, narrowing posteriorly; mandibles widest in the middle, direct, with the tip incurved, inner margin coarsely serrate-dentate: pronotum inflated, with an impressed mesial line : abdomen divided into three principal segments, of which the first is subglobular, the second considerably larger, conical, largest posteriorly, terminal segment of equal diameter with the preceding, but twice as long, with a dark indistinct transverse band about the middle: provided with a sting.

Queen
Emery (1895) - Das ♀ ist nicht grosser als die ♀♀, ja sogar kleiner als mein grösster ♀ sonst, abgesehen von den grösseren Augen sowie der Anwesenheit der Punktaugen und der Flügel, vom ♀ nicht verschieden

Male
Emery (1895) - Das ♂ ist dem von Forel beschriebenen S. gheorgieffi sehr ähnlich und nur in folgenden Punkten von der Beschreibung abweichend: der Clypeus hat eine grössere Zahl sehr kleiner Zähne; die Fühler sind weniger schlank, nur das 2. Geisselglied ist mehr als doppelt so lang wie dick, die übrigen weniger als zweimal so lang wie dick; das Mesonotum ist durchaus matt, die Seiten des Thorax wenig glänzend, das Scutellum glänzend, die seitlich gerandete, flache, abschüssige Fläche des Metanotums kaum glänzend. Das Stielchen des Abdomens ziemlich grob runzlig punktirt, daher minder glänzend als diefolgenden Segmente. Pechschwarz. Mundtheile, Fühler, Beine und Genitalien gelbbraun, die Schenkel etwas dunkler.

Etymology
Pale-footed

References based on Global Ant Biodiversity Informatics

 * Annotated Ant Species List Ordway-Swisher Biological Station. Downloaded at http://ordway-swisher.ufl.edu/species/os-hymenoptera.htm on 5th Oct 2010.
 * Banschbach V. S., and E. Ogilvy. 2014. Long-term Impacts of Controlled Burns on the Ant Community (Hymenoptera: Formicidae) of a Sandplain Forest in Vermont. Northeastern Naturalist 21(1): 1-12.
 * Belcher A. K., M. R. Berenbaum, and A. V. Suarez. 2016. Urbana House Ants 2.0.: revisiting M. R. Smith's 1926 survey of house-infesting ants in central Illinois after 87 years. American Entomologist 62(3): 182-193.
 * Brown W. L., Jr. 1949. A new American Amblyopone, with notes on the genus (Hymenoptera: Formicidae). Psyche (Cambridge). 56: 81-88.
 * Brown W. L., Jr. 1960. Contributions toward a reclassification of the Formicidae. III. Tribe Amblyoponini (Hymenoptera). Bull. Mus. Comp. Zool. 122: 143-230.
 * Canadensys Database. Dowloaded on 5th February 2014 at http://www.canadensys.net/
 * Carroll T. M. 2011. The ants of Indiana (Hymenoptera: Formicidae). Master's Thesis Purdue university, 385 pages.
 * Choate B., and F. A. Drummond. 2012. Ant Diversity and Distribution (Hymenoptera: Formicidae) Throughout Maine Lowbush Blueberry Fields in Hancock and Washington Counties. Environ. Entomol. 41(2): 222-232.
 * Choate B., and F. A. Drummond. 2013. The influence of insecticides and vegetation in structuring Formica Mound ant communities (Hymenoptera: Formicidae) in Maine lowbush blueberry. Environ. Entomol. 41(2): 222-232.
 * Clark A. T., J. J. Rykken, and B. D. Farrell. 2011. The Effects of Biogeography on Ant Diversity and Activity on the Boston Harbor Islands, Massachusetts, U.S.A. PloS One 6(11): 1-13.
 * Cokendolpher J. C., and O. F. Francke. 1990. The ants (Hymenoptera, Formicidae) of western Texas. Part II. Subfamilies Ecitoninae, Ponerinae, Pseudomyrmecinae, Dolichoderinae, and Formicinae. Special Publications, the Museum. Texas Tech University 30:1-76.
 * Cole A. C., Jr. 1949. The ants of Mountain Lake, Virginia. Journal of the Tennessee Academy of Science 24: 155-156.
 * Coovert G. A. 2005. The Ants of Ohio (Hymenoptera: Formicidae). Ohio Biological Survey, Inc. 15(2): 1-207.
 * Coovert, G.A. 2005. The Ants of Ohio (Hymenoptera: Formicidae) Ohio Biological Survey Bulletin New Series Volume 15(2):1-196
 * Cover S. P., and R. A. Johnson. 20011. Checklist of Arizona Ants. Downloaded on January 7th at http://www.asu.edu/clas/sirgtools/AZants-2011%20updatev2.pdf
 * Creighton W. S. 1940. A revision of the forms of Stigmatomma pallipes. American Museum Novitates 1079: 1-8.
 * Dash S. T. and L. M. Hooper-Bui. 2008. Species diversity of ants (Hymenoptera: Formicidae) in Louisiana. Conservation Biology and Biodiversity. 101: 1056-1066
 * Davis W. T., and J. Bequaert. 1922. An annoted list of the ants of Staten Island and Long Island, N. Y. Bulletin of the Brooklyn Entomological Society 17(1): 1-25.
 * Del Toro I., K. Towle, D. N. Morrison, and S. L. Pelini. 2013. Community Structure, Ecological and Behavioral Traits of Ants (Hymenoptera: Formicidae) in Massachusetts Open and Forested Habitats. Northeastern Naturalist 20: 1-12.
 * Del Toro, I. 2010. PERSONAL COMMUNICATION. MUSEUM RECORDS COLLATED BY ISRAEL DEL TORO
 * Deyrup, M. and J. Trager. 1986. Ants of the Archbold Biological Station, Highlands County, Florida (Hymenoptera: Formicidae). Florida Entomologist 69(1):206-228
 * Drummond F. A., A. M. llison, E. Groden, and G. D. Ouellette. 2012. The ants (Formicidae). In Biodiversity of the Schoodic Peninsula: Results of the Insect and Arachnid Bioblitzes at the Schoodic District of Acadia National Park, Maine. Maine Agricultural and forest experiment station, The University of Maine, Technical Bulletin 206. 217 pages
 * DuBois M. B. 1981. New records of ants in Kansas, III. State Biological Survey of Kansas. Technical Publications 10: 32-44
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-187
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-188
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-189
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-190
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-191
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-192
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-193
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-194
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-195
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-196
 * Dubois, M.B. and W.E. Laberge. 1988. An Annotated list of the ants of Illionois. pages 133-156 in Advances in Myrmecology, J. Trager
 * Ellison A. M., S. Record, A. Arguello, and N. J. Gotelli. 2007. Rapid Inventory of the Ant Assemblage in a Temperate Hardwood Forest: Species Composition and Assessment of Sampling Methods. Environ. Entomol. 36(4): 766-775.
 * Ellison A. M., and E. J. Farnsworth. 2014. Targeted sampling increases knowledge and improves estimates of ant species richness in Rhode Island. Northeastern Naturalist 21(1): NENHC-13NENHC-24.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Forster J.A. 2005. The Ants (hymenoptera: Formicidae) of Alabama. Master of Science, Auburn University. 242 pages.
 * Francoeur, A. 1979. Le fourmis du Quebec. Annales de la Societe Entomologique du Quebec 24(1):48-64.
 * Frye J. A., T. Frye, and T. W. Suman. 2014. The ant fauna of inland sand dune communities in Worcester County, Maryland. Northeastern Naturalist, 21(3): 446-471.
 * General D. M., and L. C. Thompson. 2011. New Distributional Records of Ants in Arkansas for 2009 and 2010 with Comments on Previous Records. Journal of the Arkansas Academy of Science 65: 166-168.
 * General D., and L. Thompson. 2008. New distributional records of ants in Arkansas. Journal of the Arkansas Academy of Science 62: 148-150.
 * Gibbs M. M., P. L. Lambdin, J. F. Grant, and A. M. Saxton. 2003. Ground-inhabiting ants collected in a mixed hardwood southern Appalachian forest in Eastern Tennessee. Journal of the Tennessee Academy of Science 78(2): 45-49.
 * Gregg, R.T. 1963. The Ants of Colorado.
 * Guénard B., K. A. Mccaffrey, A. Lucky, and R. R. Dunn. 2012. Ants of North Carolina: an updated list (Hymenoptera: Formicidae). Zootaxa 3552: 1-36.
 * Headley A. E. 1943. The ants of Ashtabula County, Ohio (Hymenoptera, Formicidae). The Ohio Journal of Science 43(1): 22-31.
 * Herbers J. M. 2011. Nineteen years of field data on ant communities (Hymenoptera: Formicidae): what can we learn. Myrmecological News 15: 43-52.
 * Hill, J.G. 2006. Ants collected at Okatibbee Lake, Lauderdale County, Mississippi
 * Ipser R. M. 2004. Native and exotic ants (Hymenoptera: Formicidae) of Georgia: Ecological Relationships with implications for development of biologically-based management strategies. Doctor of Philosophy thesis, University of Georgia. 165 pages.
 * Ipser, R.M., M.A. Brinkman, W.A. Gardner and H.B. Peeler. 2004. A Survey of Ground-Dwelling Ants (Hymenoptera: Formicidae) in Georgia. The Florida Entomologist 87(3) 253-260.
 * Ivanov, K. 2019. The ants of Ohio (Hymenoptera, Formicidae): an updated checklist. Journal of Hymenoptera Research 70: 65–87.
 * Ivanov K., L. Hightower, S. T. Dash, and J. B. Keiper. 2019. 150 years in the making: first comprehensive list of the ants (Hymenoptera: Formicidae) of Virginia, USA. Zootaxa 4554 (2): 532–560.
 * Johnson C. 1986. A north Florida ant fauna (Hymenoptera: Formicidae). Insecta Mundi 1: 243-246
 * Johnson R. Personnal Database. Accessed on February 5th 2014 at http://www.asu.edu/clas/sirgtools/resources.htm
 * Kansas State Entomology Collection. Dowloaded the 30th of May 2011 at http://biodis.k-state.edu/collections/entomology/
 * Kjar D. 2009. The ant community of a riparian forest in the Dyke Marsh Preserve, Fairfax County, Virginiam and a checklist of Mid-Atlantic Formicidae. Banisteria 33: 3-17.
 * Kjar D., and E. M. Barrows. 2004. Arthropod community heterogeneity in a mid-Atlantic forest highly invaded by alien organisms. Banisteria 23: 26-37.
 * Kjar D., and Z. Park. 2016. Increased ant (Hymenoptera: Formicidae) incidence and richness are associated with alien plant cover in a small mid-Atlantic riparian forest. Myrmecological News 22: 109-117.
 * Lessard J. P., R. R. Dunn, C. R. Parker, and N. J. Sanders. 2007. Rarity and Diversity in Forest Ant Assemblages of Great Smoky Mountains National Park. Southeastern Naturalist 1: 215-228.
 * Lynch J. F. 1981. Seasonal, successional, and vertical segregation in a Maryland ant community. Oikos 37: 183-198.
 * Lynch J. F. 1988. An annotated checklist and key to the species of ants (Hymenoptera: Formicidae) of the Chesapeake Bay region. The Maryland Naturalist 31: 61-106
 * Lynch J. F., and A. K. Johnson. 1988. Spatial and temporal variation in the abundance and diversity of ants (Hymenoptera: Formicidae) in the soild and litter layers of a Maryland forest. American Midland Naturalist 119(1): 31-44.
 * MacGown J. A., J. G. Hill, R. L. Brown, T. L. Schiefer, J. G. Lewis. 2012. Ant diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi. Mississippi Agricultural and Forestry Experiment Station Bulletin 1197: 1-30
 * MacGown J. A., J. G. Hill, and M. Deyrup. 2009. Ants (Hymenoptera: Formicidae) of the Little Ohoopee River Dunes, Emanuel County, Georgia. J. Entomol. Sci. 44(3): 193-197.
 * MacGown J. A., J. G. Hill, and R. L. Brown. 2010.  Native and exotic ant in Mississippi state parks.  Proceedings:  Imported Fire Ant Conference, Charleston, South Carolina, March 24-26, 2008: 74-80.
 * MacGown J. A., and R. L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
 * MacGown, J.A. and JV.G. Hill. Ants of the Great Smoky Mountains National Park (Tennessee and North Carolina).
 * MacGown, J.A. and R.L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A., J.G. Hill, R.L. Brown and T.L. 2009. Ant Diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi Report #2009-01. Schiefer. 2009.
 * MacGown. J. 2011. Ants collected during the 25th Annual Cross Expedition at Tims Ford State Park, Franklin County, Tennessee
 * Mackay W. P., and E. E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
 * Mackay, W.P. and E. Mackay. XXXX. The Ants of New Mexico
 * Mahon M. B., K. U. Campbell, and T. O. Crist. 2017. Effectiveness of Winkler litter extraction and pitfall traps in sampling ant communities and functional groups in a temperate forest. Environmental Entomology 46(3): 470–479.
 * Mann H. R., E. Rowe, J. Selfridge, and D. L. Price. 2018. Leaf litter and arboreal ants (Hymenoptera: Formicidae) in a Mid-Atlantic Forest. Northeastern Naturalist 25(2): 341-354.
 * Martelli, M.G., M.M. Ward and Ann M. Fraser. 2004. Ant Diversity Sampling on the Southern Cumberland Plateau: A Comparison of Litter Sifting and Pitfall Trapping. Southeastern Naturalist 3(1): 113-126
 * Menke S. B., and N. Vachter. 2014. A comparison of the effectiveness of pitfall traps and winkler litter samples for characterization of terrestrial ant (Formicidae) communities in temperate savannas. The Great Lakes Entomologist 47(3-4): 149-165.
 * Moreau C. S., M. A. Deyrup, and L. R. David Jr. 2014. Ants of the Florida Keys: Species Accounts, Biogeography, and Conservation (Hymenoptera: Formicidae). J. Insect Sci. 14(295): DOI: 10.1093/jisesa/ieu157
 * Munsee J. R., W. B. Jansma, and J. R. Schrock. 1986. Revision of the checklist of Indiana ants with the addition of five new species (Hymenoptera: Formicidae). Indiana Academy of Science 95: 265-274.
 * O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
 * O'Neill J.C. and Dowling A.P.G. 2011. A Survey of the Ants (hymenoptera: Formicidae) of Arkansas and the Ozark Mountains. An Undergraduate Honors, University of Arkansas. 18pages.
 * Ouellette G. D. and A. Francoeur. 2012. Formicidae [Hymenoptera] diversity from the Lower Kennebec Valley Region of Maine. Journal of the Acadian Entomological Society 8: 48-51
 * Ouellette G. D., F. A. Drummond, B. Choate and E. Groden. 2010. Ant diversity and distribution in Acadia National Park, Maine. Environmental Entomology 39: 1447-1556
 * Peeters, C. and M. Molet. 2010. Evolution of advanced social traits in phylogenetically basal ants: striking worker polymorphism and large queens in Amblyopone australis. Insectes Sociaux
 * Ross H. H., G. L. Rotramel, and W. E. LeBerge. 1971. A synopsis of common and economic Illinois antsm with keys to the genera (Hymenoptera, Formicidae). Illinois Nat. history Surv. Biol. Notes 71: 1-22.
 * Shik, J., A. Francoeur and C. Buddle. 2005. The effect of human activity on ant species (Hymenoptera: Formicidae) richness at the Mont St. Hilaire Biosphere Reserve, Quebec. Canadian Field-Naturalist 119(1): 38-42.
 * Smith M. R. 1928. An additional annotated list of the ants of Mississippi. With a description of a new species of Aphaenogaster (Hym.: Formicidae). Entomological News 39: 242-246.
 * Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
 * Smith M. R. 1962. A new species of exotic Ponera from North Carolina (Hymenoptera, Formicidae). Acta Hymenopterologica 1: 377-382.
 * Sturtevant A. H. 1931. Ants collected on Cape Cod, Massachusetts. Psyche (Cambridge) 38: 73-79
 * Talbot M. 1957. Populations of ants in a Missouri woodland. Insectes Sociaux 4(4): 375-384.
 * Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
 * Van Pelt A. F. 1966. Activity and density of old-field ants of the Savannah River Plant, South Carolina. Journal of the Elisha Mitchell Scientific Society 82: 35-43.
 * Van Pelt A., and J. B. Gentry. 1985. The ants (Hymenoptera: Formicidae) of the Savannah River Plant, South Carolina. Dept. Energy, Savannah River Ecology Lab., Aiken, SC., Report SRO-NERP-14, 56 p.
 * Van Pelt, A. 1983. Ants of the Chisos Mountains, Texas (Hymenoptera: Formicidae) . Southwestern Naturalist 28:137-142.
 * Wang C., J. Strazanac and L. Butler. 2000. Abundance, diversity and activity of ants (Hymenoptera: Formicidae) in oak-dominated mixed Appalachian forests treated with microbial pesticides. Environmental Entomology. 29: 579-586
 * Ward P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55: 1-16
 * Ward P. S. 2007. The ant genus Leptanilloides: discovery of the male and evaluation of phylogenetic relationships based on DNA sequence data. Memoirs of the American Entomological Institute 80: 637-649.
 * Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
 * Wheeler G. C., J. N. Wheeler, and P. B. Kannowski. 1994. Checklist of the ants of Michigan (Hymenoptera: Formicidae). The Great Lakes Entomologist 26(4): 297-310
 * Wheeler G. C., and J. Wheeler J. 1989. A checklist of the ants of Oklahoma. Prairie Naturalist 21: 203-210.
 * Wheeler G. C., and J. Wheeler. 1986. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp.
 * Wheeler W. M. 1900. The habits of Ponera and Stigmatomma. Biological Bulletin (Woods Hole). 2: 43-69.
 * Wheeler W. M. 1905. An annotated list of the ants of New Jersey. Bulletin of the American Museum of Natural History. 21: 371-403.
 * Wheeler W. M. 1906. Fauna of New England. 7. List of the Formicidae. Occasional Papers of the Boston Society of Natural History 7: 1-24.
 * Wheeler W. M. 1913. Ants collected in Georgia by Dr. J. C. Bradley and Mr. W. T. Davis. Psyche (Cambridge) 20: 112-117.
 * Wheeler W. M. 1916. Formicoidea. Formicidae. Pp. 577-601 in: Viereck, H. L. 1916. Guide to the insects of Connecticut. Part III. The Hymenoptera, or wasp-like insects, of Connecticut. Connecticut State Geological and Natural History Survey. Bulletin 22: 1-824.
 * Wheeler, G.C. and J. Wheeler. 1985. A checklist of Texas ants. Prairie Naturalist 17:49-64.
 * Wheeler, G.C., J. Wheeler and P.B. Kannowski. 1994. CHECKLIST OF THE ANTS OF MICHIGAN (HYMENOPTERA: FORMICIDAE). Great Lakes Entomologist 26:1:297-310
 * Whelden R. M. 1958. Notes on the anatomy of the Formicidae. 1. Stigmatomma pallipes (Haldeman). Journal of the New York Entomological Society 65: 1-21.
 * Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
 * Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71