Checklist of Cephalotes species

The following species and subspecies belong to the genus Cephalotes. Synonyms are listed under their senior names. For valid names only see Cephalotes species and for distribution information see Cephalotes species by Country.

A
adolphi (Emery, 1906) (Brazil)

alfaroi (Emery, 1890) (Costa Rica, Panama)

angustus (Mayr, 1862) (Argentina, Brazil)

• Cryptocerus angustus clarior Forel, 1912

• Cryptocerus cristatus ruficeps Santschi, 1929

argentatus (Smith, F., 1853) (Colombia)

argentiventris De Andrade, 1999 (Dominican Republic, Greater Antilles)

atratus (Linnaeus, 1758) (Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Uruguay, Venezuela)

• Cephalotes atratus crassispina Santschi, 1920

• Cephalotes atratus erectus Kempf, 1951

• Cephalotes atratus nitidiventris Santschi, 1920

• Cephalotes decemspinosus Santschi, 1920

• Cephalotes decemspinosus spissus Santschi, 1929

• Cryptocerus atratus rufiventris Emery, 1894

• Cryptocerus dubitatus Smith, F., 1853

• Cryptocerus marginatus Fabricius, 1804

• Formica quadridens De Geer De Geer, 1773

auricomus (Wheeler, W.M., 1936) (Dominican Republic, Greater Antilles, Haiti)

auriger De Andrade, 1999 (Mexico)

B
basalis (Smith, F., 1876) (Brazil, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Cryptocerus cordatus multispinus Emery, 1890

betoi De Andrade & Baroni Urbani, 1999 (Brazil, Ecuador)

biguttatus (Emery, 1890) (Belize, Costa Rica, Guatemala, Honduras, Mexico)

bimaculatus (Smith, F., 1860) (Mexico)

• Cryptocerus aztecus Forel, 1901

bivestitus (Santschi, 1922) (Argentina)

• Cryptocerus guttatus trivialis Santschi, 1929

bohlsi (Emery, 1896) (Argentina, Brazil, Ecuador, Paraguay)

• Cryptocerus bohlsi medusa Santschi, 1919

borgmeieri (Kempf, 1951) (Argentina, Brazil, Ecuador, Paraguay)

bruchi (Forel, 1912) (Argentina, Brazil, Ecuador)

• Cryptocerus iheringi pampaensis Santschi, 1931

• Cryptocerus ridiculus Santschi, 1915

C
chacmul Snelling, R.R., 1999 (Mexico)

christopherseni (Forel, 1912) (Colombia, Panama, Venezuela)

clypeatus (Fabricius, 1804) (Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)

coffeae (Kempf, 1953) (Colombia)

columbicus (Forel, 1912) (Colombia)

complanatus (Guérin-Méneville, 1844) (Bolivia, Brazil, Colombia, French Guiana, Peru, Suriname)

• Cryptocerus angulatus Smith, F., 1858

• Cryptocerus multispinus amazonensis Forel, 1911

conspersus (Smith, F., 1867) (Brazil, Colombia, Ecuador, Trinidad and Tobago)

• Cryptocerus denticulatus variegata Forel, 1911

cordatus (Smith, F., 1853) (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Peru)

• Cryptocerus cordatus boliviensis Santschi, 1921

cordiae (Stitz, 1913) (Bolivia, Brazil, Peru)

cordiventris (Santschi, 1931) (Costa Rica, Panama)

crenaticeps (Mayr, 1866) (Colombia, Venezuela)

cristatus (Emery, 1890) (Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico)

curvistriatus (Forel, 1899) (Costa Rica, Mexico)

D
decolor De Andrade, 1999 (Colombia, Venezuela)

decoloratus De Andrade, 1999 (Dominican Republic, Greater Antilles, Haiti)

• Cryptocerus varians marginata Wheeler, W.M. & Mann, 1914

dentidorsum De Andrade, 1999 (Ecuador, Peru)

depressus (Klug, 1824) (Argentina, Bolivia, Brazil, Colombia, Ecuador, Guyana, Paraguay, Venezuela)

• Cryptocerus depressus sorocabensis Forel, 1912

dorbignyanus (Smith, F., 1853)

duckei (Forel, 1906) (Brazil, Colombia)

E
ecuadorialis De Andrade, 1999 (Ecuador)

eduarduli (Forel, 1921) (Argentina, Bolivia, Brazil, Paraguay)

emeryi (Forel, 1912) (Lesser Antilles, Mexico)

F
femoralis (Smith, F., 1853) (Colombia, Ecuador, Venezuela)

• Cryptocerus silvae Forel, 1906

fiebrigi (Forel, 1906) (Argentina, Brazil, Greater Antilles, Paraguay)

• Cryptocerus guttifer Santschi, 1919

flavigaster De Andrade, 1999 (Dominican Republic, Greater Antilles)

foliaceus (Emery, 1906) (Bolivia, Colombia, Panama, Peru)

fossithorax (Santschi, 1921) (Argentina)

frigidus (Kempf, 1960) (Brazil)

G
gabicamacho Oliveira, Powell & Feitosa, 2021 (Brazil)

goeldii (Forel, 1912) (Brazil)

goniodontus De Andrade, 1999 (Mexico)

grandinosus (Smith, F., 1860) (Brazil, Colombia, Costa Rica, Ecuador, Guyana, Panama, Trinidad and Tobago)

• Cryptocerus grandinosus nevadensis Forel, 1912

guayaki De Andrade, 1999 (Argentina, Brazil, Paraguay)

H
haemorrhoidalis (Latreille, 1802) (Haiti)

hamulus (Roger, 1863) (Dominican Republic, Greater Antilles, Haiti)

• Cryptocerus hamulus haytianus Forel, 1901

• Hypocryptocerus haemorrhoidalis affinis Wheeler, W.M., 1936

hirsutus De Andrade, 1999 (Mexico)

I
inaequalis (Mann, 1916) (Brazil, Colombia)

inca (Santschi, 1911) (Ecuador, Peru)

incertus (Emery, 1906) (Argentina, Brazil, Paraguay)

insularis (Wheeler, W.M., 1934) (Mexico)

J
jamaicensis (Forel, 1922) (Greater Antilles, Jamaica)

jheringi (Emery, 1894) (Argentina, Brazil, Paraguay)

• Cryptocerus peltatus Emery, 1896

• Cryptocerus peltatus ellenriederi Forel, 1911

K
klugi (Emery, 1894) (Brazil)

kukulcan Snelling, R.R., 1999 (Belize, Costa Rica, Honduras, Mexico)

L
laminatus (Smith, F., 1860) (Brazil, Colombia, Ecuador, Peru)

lanuginosus (Santschi, 1919) (Argentina, Paraguay)

• Cryptocerus lobigaster Santschi, 1919

lenca De Andrade, 1999 (Honduras, Nicaragua)

liepini De Andrade & Baroni Urbani, 1999 (Brazil)

liogaster (Santschi, 1916) (Argentina)

• Cryptocerus lobigaster fides Santschi, 1925

liviaprado Oliveira, Powell & Feitosa, 2021 (Brazil)

M
maculatus (Smith, F., 1876) (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Trinidad and Tobago, Venezuela)

• Cryptocerus grandinosus magdalenensis Forel, 1899

• Cryptocerus maculatus nanus Forel, 1912

manni (Kempf, 1951) (Brazil, Colombia, Ecuador, Guyana, Peru, Venezuela)

mariadeandrade Oliveira, Powell & Feitosa, 2021 (Brazil)

marycorn Oliveira, Powell & Feitosa, 2021 (Brazil)

membranaceus (Klug, 1824) (Brazil)

• Cryptocerus fervidus Smith, F., 1876

minutus (Fabricius, 1804) (Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Guyana, Honduras, Lesser Antilles, Mexico, Nicaragua, Panama, Paraguay, Peru, Trinidad and Tobago, Venezuela)

• Cryptocerus cognatus Smith, F., 1862

• Cryptocerus exiguus Smith, F., 1867

• Cryptocerus quadrimaculatus Klug, 1824

• Cryptocerus volxemi Emery, 1878

mompox De Andrade, 1999 (Colombia, Venezuela)

monicaulyssea Oliveira, Powell & Feitosa, 2021 (Brazil)

multispinosus (Norton, 1868) (Costa Rica, Honduras, Mexico, Nicaragua, Panama)

• Cryptocerus gibbosus Smith, F., 1876

N
nilpiei De Andrade, 1999 (Brazil)

notatus (Mayr, 1866) (Brazil)

• Paracryptocerus fleddermanni Kempf, 1958

O
oculatus (Spinola, 1851) (Brazil, Peru)

• Cryptocerus aethiops Smith, F., 1853

opacus Santschi, 1920 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela)

• Cephalotes abdominalis Santschi, 1929

P
pallens (Klug, 1824) (Argentina, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Grenada, Guatemala, Guyana, Honduras, Lesser Antilles, Mexico, Nicaragua, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Cryptocerus araneolus Smith, F., 1853

• Cryptocerus discocephalus Smith, F., 1853

pallidicephalus (Smith, F., 1876) (Brazil, Mexico)

• Cryptocerus striativentris Emery, 1894

pallidoides De Andrade, 1999 (Argentina, Bolivia, Brazil, Colombia, Ecuador, Guyana, Paraguay, Trinidad and Tobago, Venezuela)

pallidus De Andrade, 1999 (Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, Suriname)

palta De Andrade, 1999 (Colombia)

palustris De Andrade, 1999 (Brazil, Colombia, Guyana)

patei (Kempf, 1951) (Colombia)

patellaris (Mayr, 1866) (Brazil)

pavonii (Latreille, 1809) (Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname)

pellans De Andrade, 1999 (Argentina, Bolivia, Brazil, Paraguay)

persimilis De Andrade, 1999 (Argentina, Brazil, Paraguay)

persimplex De Andrade, 1999 (Bolivia, Brazil)

peruviensis De Andrade, 1999 (Costa Rica, Ecuador, Peru)

pileini De Andrade, 1999 (Argentina)

pilosus (Emery, 1896) (Argentina, Brazil, Paraguay)

pinelii (Guérin-Méneville, 1844) (Argentina, Brazil, Paraguay)

placidus (Smith, F., 1860) (Brazil, Colombia, Guyana, Peru)

• Cryptocerus fenestralis Smith, F., 1876

porrasi (Wheeler, W.M., 1942) (Belize, Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Nicaragua, Panama)

prodigiosus (Santschi, 1921) (Argentina)

pusillus (Klug, 1824) (Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Cryptocerus elongatus Klug, 1824

• Cryptocerus obtusus Smith, F., 1858

• Cryptocerus pusillus brevispinosa Santschi, 1921

• Formica caustica Kollar, 1832

Q
quadratus (Mayr, 1868) (Argentina, Paraguay)

• Cryptocerus convexus Santschi, 1916

R
ramiphilus (Forel, 1904) (Bolivia, Brazil, Colombia, Ecuador, Peru)

resinae De Andrade, 1999 (Dominican Republic)

rohweri (Wheeler, W.M., 1916) (Mexico, United States)

S
scutulatus (Smith, F., 1867) (Belize, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Panama, Venezuela)

• Cryptocerus angulosus Smith, F., 1867

• Cryptocerus jucundus Smith, F., 1876

serraticeps (Smith, F., 1858) (Brazil, Colombia, Ecuador, Peru)

setulifer (Emery, 1894) (Colombia, Costa Rica, Honduras, Panama)

• Cryptocerus setulifer orbis Forel, 1906

simillimus (Kempf, 1951) (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname)

sobrius (Kempf, 1958) (Guatemala)

solidus (Kempf, 1974) (Brazil, Ecuador)

specularis Brandão, Feitosa, Powell & Del-Claro, 2014 (Brazil)

spinosus (Mayr, 1862) (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Cryptocerus laminatus peruvianus Forel, 1911

• Cryptocerus punctatus Mayr, 1862

supercilii De Andrade, 1999 (Argentina, Ecuador)

T
taino De Andrade, 1999 (Greater Antilles)

targionii (Emery, 1894) (Argentina, Bolivia, Brazil, Guyana, Paraguay, Trinidad and Tobago, Venezuela)

• Cryptocerus denticulatus Emery, 1894

texanus (Santschi, 1915) (Mexico, United States)

toltecus De Andrade, 1999 (Mexico)

trichophorus De Andrade, 1999 (Brazil, Ecuador, Peru)

U
umbraculatus (Fabricius, 1804) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Cryptocerus elegans Smith, F., 1853

• Cryptocerus flavomaculatus Mayr, 1862

• Cryptocerus quadriguttatus Guérin-Méneville, 1844

unimaculatus (Smith, F., 1853) (Dominican Republic, Greater Antilles, Haiti)

• Cryptocerus hamulus steinheili Forel, 1908

• Hypocryptocerus haemorrhoidalis signatus Wheeler, W.M., 1936

ustus (Kempf, 1973) (Brazil)

V
varians (Smith, F., 1876) (Bahamas, Cuba, Greater Antilles, United States)

vinosus (Wheeler, W.M., 1936) (Greater Antilles, Haiti)

W
wheeleri (Forel, 1901) (Mexico)

=Fossil Species=

A
alveolatus (Vierbergen & Scheven, 1995)

B
bloosi Baroni Urbani & De Andrade, 1999

brevispineus De Andrade, 1999

C
caribicus De Andrade & Baroni Urbani, 1999

D
dieteri Baroni Urbani, 1999

H
hispaniolicus De Andrade, 1999

I
integerrimus (Vierbergen & Scheven, 1995)

J
jansei (Vierbergen & Scheven, 1995)

M
maya De Andrade, 1999

O
obscurus (Vierbergen & Scheven, 1995)

olmecus De Andrade, 1999

P
poinari Baroni Urbani, 1999

S
serratus (Vierbergen & Scheven, 1995)

squamosus (Vierbergen & Scheven, 1995)

sucinus De Andrade, 1999

V
ventriosus De Andrade, 1999