Checklist of Aphaenogaster species

The following species and subspecies belong to the genus Aphaenogaster. Synonyms are listed under their senior names. For valid names only see Aphaenogaster species and for distribution information see Aphaenogaster species by Country.

A
afra Santschi, 1933 (Algeria)

aktaci Kiran & Tezcan, 2008 (Greece, Turkey)

angulata Viehmeyer, 1922 (China)

annandalei Mukerjee, 1930 (India)

araneoides Emery, 1890 (Costa Rica, El Salvador, Guatemala, Honduras, Nicaragua, Panama)

ashmeadi (Emery, 1895) (United States)

• Aphaenogaster treatae harnedi Wheeler, W.M., 1919

asmaae Sharaf, 2018 (Oman)

atlantis Santschi, 1929 (Morocco)

B
balcanica (Emery, 1898) (Albania, Bosnia and Herzegovina, Croatia, Greece, Italy, Malta, Slovenia, Turkey)

• Aphaenogaster semipolita ionia Baroni Urbani Baroni Urbani, 1968

• Aphaenogaster semipolita ionia Santschi Santschi, 1933

balcanicoides Boer, 2013 (Greece)

baogong Terayama, 2009 (Taiwan)

barbara Shattuck, 2008 (Australia)

barbigula Wheeler, W.M., 1916 (Australia)

baronii Cagniant, 1988 (Morocco)

beccarii Emery, 1887 (China, India, Indonesia, Nicobar Island, Sri Lanka)

beesoni Donisthorpe, 1933 (India, Pakistan)

boulderensis Smith, M.R., 1941 (Mexico, United States)

bressleri Csősz & Fisher, 2021 (Madagascar)

brevicollis Forel, 1899 (Panama)

burri (Donisthorpe, 1950) (Turkey)

C
caeciliae Viehmeyer, 1922 (China)

campana Emery, 1878 (Albania, Italy, Malta)

carbonaria Pergande, 1894 (Mexico)

cardenai Espadaler, 1981 (Iberian Peninsula, Slovenia, Spain)

carolinensis Wheeler, W.M., 1915 (United States)

cavernicola Donisthorpe, 1938 (India)

cecconii Emery, 1894 (Greece)

charesi Salata & Borowiec, 2016 (Greece)

concolor Watanabe & Yamane, 1999 (China, Japan)

cristata (Forel, 1902) (India)

crocea André, 1881 (Algeria)

crocea croceoides Forel, 1890 (Algeria)

crocea lenis Santschi, 1911 (Tunisia)

crocea splendidoides Forel, 1890 (Algeria)

curiosa Santschi, 1933 (Morocco)

D
dejeani Cagniant, 1982 (Morocco)

depilis Santschi, 1911 (Algeria)

depilis numida Santschi, 1933 (Tunisia)

depressa Bolton, 1995 (Turkey)

• Ceratopheidole perganderi Donisthorpe, 1950

dlusskyi Radchenko & Arakelian, 1991 (Armenia)

donann Watanabe & Yamane, 1999 (Japan)

dromedaria (Emery, 1900) (New Guinea)

• Aphaenogaster dromedarius nigra Donisthorpe, 1938

• Ischnomyrmex dromedarius fusca Emery, 1900

dulciniae Emery, 1924 (Gibraltar, Iberian Peninsula, Spain)

E
edentula Watanabe & Yamane, 1999 (Japan)

epirotes (Emery, 1895) (Albania, Bulgaria, Croatia, Greece, Israel, Italy, Montenegro, Republic of Macedonia, Serbia, Slovenia, Turkey)

equestris Borowiec & Salata, 2014 (Turkey)

erabu Nishizono & Yamane, 1990 (Japan)

espadaleri Cagniant, 1984 (Morocco)

exasperata Wheeler, W.M., 1921 (China, Vietnam)

F
fabulosa Arnoldi, 1968 (Turkmenistan)

fallax Cagniant, 1992 (Algeria, Morocco)

famelica (Smith, F., 1874) (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Aphaenogaster famelica frontosa Wheeler, W.M., 1928

• Aphaenogaster famelica ruida Wheeler, W.M., 1928

• Aphaenogaster vapida Wheeler, W.M., 1928

faureli Cagniant, 1969 (Algeria)

feae Emery, 1889 (China, India, Myanmar, Philippines)

feae imbellis (Emery, 1900) (Indonesia)

feae nicobarensis (Forel, 1903) (India, Nicobar Island)

feae simulans Forel, 1915 (Indonesia)

fengbo Terayama, 2009 (Taiwan)

festae Emery, 1915 (Bulgaria, Greece, Iraq, Turkey)

finzii Müller, 1921 (Bosnia and Herzegovina, Croatia, Greece, Italy, Montenegro, Republic of Macedonia, Serbia)

fiorii Emery, 1915 (Italy)

flemingi Smith, M.R., 1928 (United States)

• Aphaenogaster texana macrospina Smith, M.R., 1934

• Aphaenogaster texana nana Wheeler, W.M., 1932

floridana Smith, M.R., 1941 (United States)

foreli Cagniant, 1995 (Morocco)

fulva Roger, 1863 (United States)

• Aphaenogaster fulva pusilla Enzmann, J., 1947

• Aphaenogaster fulva rubida Enzmann, J., 1947

• Myrmica aquia Buckley, 1867

G
gamagumayaa Naka & Maruyama, 2018 (Japan)

geei Wheeler, W.M., 1921 (China)

gemella (Roger, 1862) (Algeria, Balearic Islands)

gemella marocana (Forel, 1903) (Morocco)

• Aphaenogaster gemella tingitana Santschi, 1933

georgica Arnoldi, 1968 (Georgia)

gibbosa (Latreille, 1798) (Algeria, Andorra, Armenia, Bulgaria, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Iran, Israel, Montenegro, Portugal, Republic of Macedonia, Russian Federation, Saudi Arabia, Spain, Switzerland, Turkey, Turkmenistan)

• Aphaenogaster gibbosa barcinensis Menozzi, 1922

• Aphaenogaster gibbosa chorassanica Arnoldi, 1968

• Aphaenogaster striola laevior Forel, 1892

• Atta striola Roger, 1859

gibbosa homonyma Emery, 1921 (Tunisia)

gonacantha (Emery, 1899) (Madagascar)

gracillima Watanabe & Yamane, 1999 (Japan)

graeca Schulz, 1994 (Greece)

H
haarlovi Collingwood, 1961 (Afghanistan)

hamaensis Salata, Karaman, Kiran & Borowiec, 2021

hesperia Santschi, 1911 (Canary Islands)

holtzi (Emery, 1898) (Turkey)

honduriana Mann, 1922 (Honduras)

huachucana Creighton, 1934 (Mexico, United States)

hunanensis Wu, J. & Wang, C., 1992 (China)

I
iberica Emery, 1908 (Balearic Islands, Canary Islands, Gibraltar, Iberian Peninsula, Portugal, Spain)

• Aphaenogaster angusta Santschi, 1925

• Aphaenogaster conica Viehmeyer, 1914

• Aphaenogaster testaceopilosa vieirai Santschi, 1933

ichnusa Santschi, 1925 (Italy)

illyrica Bračko, Lapeva-Gjonova, Salata, Borowiec & Polak, 2019 (Bulgaria, Croatia, Greece, Slovenia)

incurviclypea Wang, W. & Zheng, 1997 (China)

inermis Forel, 1899 (Colombia, Costa Rica, Nicaragua, Panama)

• Aphaenogaster araneoides canalis Enzmann, J., 1947

• Aphaenogaster araneoides nitidiventris Forel, 1912

inermita Bolton, 1995 (Italy, Malta)

• Aphaenogaster semipolita inermis Santschi, 1933

iranica Kiran & Alipanah, 2013 (Iran)

irrigua Watanabe & Yamane, 1999 (Japan)

isekram Bernard, 1977 (Algeria)

italica Bondroit, 1918 (Italy, Switzerland)

izuensis Terayama & Kubota, 2013 (Japan)

J
januschevi Arnoldi, 1976 (Tajikistan)

japonica Forel, 1911 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Aphaenogaster smythiesi sinensis Wheeler, W.M., 1928

• Aphaenogaster verecunda Wheeler, W.M., 1928

jolantae Borowiec & Salata, 2014 (Greece)

K
karpathica Boer, 2013 (Greece)

kervillei Forel, 1910 (Israel, Lebanon, Turkey)

kimberleyensis Shattuck, 2008 (Australia)

koniari Cagniant & Galkowski, 2013 (Morocco)

kumejimana Watanabe & Yamane, 1999 (Japan)

kurdica Ruzsky, 1905 (Armenia, Georgia, Iran, Russian Federation)

L
laevior Emery, 1887 (Indonesia, Sulawesi)

lamellidens Mayr, 1886 (United States)

• Aphaenogaster lamellidens nigripes Smith, M.R., 1923

ledouxi Tohmé, G., 1969 (Greece, Lebanon)

lepida Wheeler, W.M., 1930 (China, Republic of Korea, Taiwan)

• Aphaenogaster funkikoensis Creighton, 1950

• Aphaenogaster phillipi Menozzi, 1932

• Aphaenogaster silvestrii Wheeler, W.M. Wheeler, W.M., 1929

lesbica Forel, 1913 (Greece, Israel, Montenegro)

leveillei Emery, 1881 (Algeria, Morocco)

leveillei laurenti Santschi, 1939 (Algeria)

longiceps (Smith, F., 1858) (Australia, China, India, Nicobar Island)

• Aphaenogaster longiceps flava Emery, 1921

• Stenamma longiceps ruginota Forel, 1902

loriai (Emery, 1897) (New Guinea)

• Aphaenogaster loriai atra Stitz, 1938

lustrans Smith, M.R., 1961 (New Guinea)

luteipes Watanabe & Yamane, 1999 (Japan)

lykiaensis Borowiec & Salata, 2014 (Turkey)

M
maculifrons Kiran & Aktaç, 2008 (Turkey)

makay Csősz & Fisher, 2021 (Madagascar)

mariae Forel, 1886 (United States)

mauritanica Dalla Torre, 1893 (Algeria, Tunisia)

mediterrae Shattuck, 2008 (Australia)

megommata Smith, M.R., 1963 (Mexico, United States)

melitensis Santschi, 1933 (Malta)

messoroides Dlussky, 1990 (Turkmenistan)

mexicana (Pergande, 1896) (Mexico)

miamiana Wheeler, W.M., 1932 (Mexico, United States)

• Aphaenogaster fulva azteca Enzmann, J., 1947

miniata Cagniant, 1990 (Morocco)

minutula Watanabe & Yamane, 1999 (Japan)

montana Mackay, 2017 (Mexico)

muschtaidica Emery, 1908 (Georgia)

mutica Pergande, 1896 (Mexico)

N
nadigi Santschi, 1923 (Morocco)

O
obsidiana (Mayr, 1861) (Georgia, Iran, Israel, Montenegro, Republic of Macedonia, Russian Federation, Turkey)

occidentalis (Emery, 1895) (Canada, United States)

• Aphaenogaster subterranea borealis Wheeler, W.M., 1915

• Aphaenogaster subterranea valida Wheeler, W.M., 1915

olympica Borowiec & Salata, 2014 (Greece)

omotoensis Terayama & Kubota, 2013 (Japan)

opposita (Say, 1836)

osimenseis Teranishi, 1940 (Japan, Micronesia (Federated States of))

ovaticeps (Emery, 1898) (Albania, Croatia, Greece, Italy, Montenegro, Republic of Macedonia, Slovenia)

• Aphaenogaster ovaticeps muelleriana Wolf, 1915

P
pachei (Forel, 1906) (India, Nepal)

pallescens Walker, 1871

pallida (Nylander, 1849) (Algeria, Bulgaria, Canary Islands, Italy, Republic of Macedonia)

patruelis Forel, 1886 (Mexico, United States)

• Aphaenogaster patruelis willowsi Wheeler, W.M., 1933

• Stenamma patruele bakeri Wheeler, W.M., 1904

peloponnesiaca Salata, Karaman, Kiran & Borowiec, 2021

perplexa Smith, M.R., 1961 (New Guinea)

• Aphaenogaster quadrispina wheeleri Donisthorpe, 1938

phalangium Emery, 1890 (Colombia, Costa Rica, Honduras, Panama)

phillipsi Wheeler, W.M. & Mann, 1916 (Israel, Jordan, Occupied Palestinian Territory)

picea (Wheeler, W.M., 1908) (United States)

• Aphaenogaster texana punctithorax Cole, 1938

picena Baroni Urbani, 1971 (Albania, Croatia, Italy, Slovenia)

polyodonta Zhou, 2001 (China)

poultoni Crawley, 1922 (Australia)

praedo Emery, 1908 (Algeria, Morocco)

praedo ellipsoida Santschi, 1933 (Morocco)

praenoda Santschi, 1933 (Morocco)

praenoda confinis Santschi, 1933 (Morocco)

projectens Donisthorpe, 1947 (New Guinea)

prudens (Forel, 1902) (India, Nepal)

pumilopuncta Zhou, 2001 (China)

punctaticeps Mackay, W.P., 1989 (Mexico, United States)

punctatissima Mackay, 2017 (Mexico)

pythia Forel, 1915 (Australia, New Guinea)

Q
quadrispina Emery, 1911 (New Guinea)

R
radchenkoi Kiran, Aktaç & Tezcan, 2008 (Bulgaria, Turkey)

reichelae Shattuck, 2008 (Australia)

relicta Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

• Aphaenogaster relicta epinotalis Wheeler, W.M. & Mann, 1914

reticulaticeps Mackay, 2017 (Mexico)

rhaphidiiceps (Mayr, 1877) (Afghanistan, Iran, Kazakhstan, Turkmenistan)

rifensis Cagniant, 1994 (Morocco)

rothneyi (Forel, 1902) (China, India)

rudis Enzmann, J., 1947 (United States)

rugosoferruginea Forel, 1889 (Greece)

rugulosa Watanabe & Yamane, 1999 (Japan)

rupestris Forel, 1909 (Algeria)

S
sagei (Forel, 1902) (China, India, Pakistan)

sahafina Csősz & Fisher, 2021 (Madagascar)

saharensis Bernard, 1953 (Algeria)

sangiorgii (Emery, 1901) (Greece, Republic of Macedonia)

sarae Sharaf, 2018 (Oman)

sardoa Mayr, 1853 (Algeria, Italy, Morocco)

• Aphaenogaster sardoa longispina Wolf, 1915

sardoa anoemica Santschi, 1910 (Morocco)

schmidti Karavaiev, 1912 (Republic of Korea)

schmitzi Forel, 1910 (Israel, Jordan)

• Aphaenogaster schmitzi syria Forel, 1910

schurri (Forel, 1902) (China, India)

semipolita (Nylander, 1856) (Georgia, Italy, Turkey)

senilis Mayr, 1853 (Balearic Islands, Canary Islands, France, Gibraltar, Iberian Peninsula, Italy, Portugal, Slovakia, Spain)

• Aphaenogaster senilis acoreensis Santschi, 1933

• Aphaenogaster senilis grata Santschi, 1933

• Aphaenogaster senilis occidua Santschi, 1933

• Aphaenogaster testaceopilosa fuentei Medina, 1893

senilis disjuncta Santschi, 1933 (Morocco)

sicardi Cagniant, 1990 (Morocco)

sicula Emery, 1908 (Italy, Malta, Montenegro)

simonellii Emery, 1894 (Greece, Turkey)

smithi Gregg, 1949 (United States)

smythiesii (Forel, 1902) (Afghanistan, China, India, Nepal, Pakistan, Republic of Korea)

spinosa Emery, 1878 (France, Italy, Switzerland)

• Aphaenogaster corsica Casevitz-Weulersse, 2010

• Aphaenogaster spinosa etrusca Baroni Urbani, 1969

• Aphaenogaster testaceopilosum nitida Krausse, 1912

• Aphaenogaster testaceopilosum romana Santschi, 1933

splendida (Roger, 1859) (Algeria, Armenia, Croatia, Greece, Iberian Peninsula, Iran, Israel, Malta, Occupied Palestinian Territory, Republic of Macedonia, Russian Federation, Spain, Turkey)

sporadis Santschi, 1933 (Greece)

striativentris Forel, 1895 (Iberian Peninsula, Spain, Spain)

strioloides Forel, 1890 (Algeria, Tunisia)

subcostata Viehmeyer, 1922 (Greece, Turkey)

subexaperata Zhou, 2001 (China)

subterranea (Latreille, 1798) (Albania, Algeria, Andorra, Armenia, Austria, Balearic Islands, Belgium, Bulgaria, Channel Islands, China, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Malta, Montenegro, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Aphaenogaster friederichsi Forel, 1918

subterraneoides Emery, 1881 (Bulgaria, Croatia, Greece, Israel, Montenegro, Russian Federation, Ukraine)

• Aphaenogaster pallida cypriotes Emery, 1908

subterraneoides armeniaca Arnoldi, 1968 (Armenia)

swammerdami Forel, 1886 (Madagascar)

• Aphaenogaster swammerdami clara (Santschi, 1915)

• Aphaenogaster swammerdami curta Forel, 1891

• Aphaenogaster swammerdami spinipes Santschi, 1911

syriaca Emery, 1908 (Iran, Israel, Lebanon)

T
takahashii Wheeler, W.M., 1930 (China, Taiwan)

tennesseensis (Mayr, 1862) (United States)

• Atta laevis Mayr, 1862

• Myrmica subrubra Buckley, 1867

• Stenamma tennesseense ecalcaratum Emery, 1895

testaceopilosa (Lucas, 1849) (Algeria, Turkey)

testaceopilosa cabylica Stitz, 1917 (Algeria)

testaceopilosa canescens (Emery, 1895) (Algeria)

texana Wheeler, W.M., 1915 (Mexico, United States)

• Aphaenogaster huachucana crinimera Cole, 1953

• Aphaenogaster silvestrii Menozzi Menozzi, 1929

• Aphaenogaster texana furvescens Wheeler, W.M., 1915

theryi Santschi, 1923 (Morocco)

tibetana Donisthorpe, 1929 (China, Tibet)

tinauti Cagniant, 1992 (Morocco)

tipuna Forel, 1913 (Japan, Taiwan)

tokarainsulana Watanabe & Yamane, 1999 (China, Japan)

torossiani Cagniant, 1988 (Morocco)

transcaucasica Karavaiev, 1926 (Georgia)

treatae Forel, 1886 (United States)

• Aphaenogaster treatae pluteicornis Wheeler, G.C. & Wheeler, E.W., 1934

• Aphaenogaster treatae wheeleri Mann, 1915

trinacriae Alicata & Schifani, 2019 (Italy)

turkestanica Arnoldi, 1976 (Kazakhstan)

U
uinta Wheeler, W.M., 1917 (United States)

ujhelyii Szabó, 1910 (Tunisia)

ulibeli Gomez & Espadaler, 2018 (Spain)

umphreyi Deyrup & Davis, 1998 (United States)

V
vohraliki Salata, Karaman, Kiran & Borowiec, 2021

W
wangtian Terayama, 2009 (Taiwan)

wangye Terayama, 2009 (Taiwan)

weigoldi Viehmeyer, 1922 (China)

weulersseae Cagniant, 1989 (Morocco)

wilsoni Cagniant, 1988 (Morocco)

X
xuantian Terayama, 2009 (Taiwan)

=Fossil Species=

A
amphioceanica De Andrade, 1995

antiqua Dlussky, 2002

archaica (Meunier, 1915)

D
dlusskyana Radchenko & Perkovsky, 2016

donisthorpei Carpenter, 1930

E
enspelensis Jessen, 2020

L
longaeva Scudder, 1877

M
mayri Carpenter, 1930

mersa Wheeler, W.M., 1915

O
oligocenica Wheeler, W.M., 1915

P
praerelicta De Andrade, 1995

S
sommerfeldti Mayr, 1868