Probolomyrmex dammermani

Nothing is known about the biology of .

Identification
Taylor (1965) - P. dammermani is closely related to the other Indo-Australian species, Probolomyrmex salomonis and Probolomyrmex greavesi. It is distinguished from them by the proportions of the node and the shape of its subpetiolar process. P. salomonis has a proportionately narrow head, and the antennal scapes of greavesi are somewhat shorter.

Eguchi et al. (2006) - In the worker this species is most similar to P. longinodus, but well distinguished from the latter by the relatively shorter petiole (LPtI < 135) and poorly developed propodeal teeth in the worker (see also Terayama & Ogata, 1988). In the male, P. dammermani is similar to Probolomyrmex longiscapus and Probolomyrmex longinodus, but the former can be separated from the latter two by the following characters: 1) thinner head in lateral view, 2) lower frontal carina, 3) narrower eye, and 4) narrower abdominal sternum IX and genitalia.

The nontype workers agree with the syntype worker. However, a certain variation is seen in the following characters: punctation on the body surface; convexity of lateral outlines of head in full-face view; concavity of occipital border (also mentioned by Taylor, 1965); development of posterior portion of subpetiolar process (also mentioned by Taylor, 1965); and development of posterolateral lamella of propodeum (sometimes forming a triangular propodeal spine). The flagellum is more slender in the worker and queen from Bali than in those from the other localities. Workers from colony Eg01iv06-08 are smaller, antennal scape shorter, and petiole in profile shorter (N=2; WL 0.69–0.73 mm, SI 89, LPtI 110) than in the material from the other localities (N=5; WL 0.88–0.96 mm, SI 103–117, LPtI 116–131).

Distribution based on Regional Taxon Lists
Indo-Australian Region: Indonesia, Philippines. Oriental Region: Thailand.

Nomenclature

 *  dammermani. Probolomyrmex dammermani Wheeler, W.M. 1928e: 7, fig. 1 (w.) INDONESIA (Java). Eguchi, Yoshimura & Yamane, 2006: 7 (q.m.). See also: Taylor, 1965d: 356.

Worker
Eguchi et al. (2006) - HL, 0.54–0.66 mm; HW, 0.35–0.42 mm; SL, 0.31–0.49 mm; CI, 63–67; SI, 89–117; WL, 0.69–0.96 mm; PW, 0.26–0.34 mm; DPtW, 0.15–0.19 mm; DPtI, 55–59; PtH, 0.21–0.26 mm; PtNL, 0.23–0.34 mm; LPtI, 110–131 (N=7).

Taylor (1965) - The following notes are based on a single syntype worker (labelled “cotype”) in the Museum of Comparative Zoology (type No. 26427). The general accuracy of Wheeler's original description has been confirmed by study of this specimen, but a few additions and corrections are indicated.

(1) HL, 0.60 mm.; HW, 0.41 mm.; SL, 0.42 mm.; Cl, 68; SI, 103; WL, 0.81 mm.; PW, 0.33 mm.; dorsal petiole width, 0.19 mm.; petiolar node index, 58; petiolar height, 0.25 mm.; petiolar node length, 0•30 nun.; lateral petiolar index, 120.

(2) First funicular segment of antenna about one-fifth longer than broad, and terminal segment one-quarter longer than the three preceding segments together.

(3) The two sculptural components normal for the genus present. Fine shagreening everywhere well developed, and overlying puncturation very distinct. Punctures of frons and dorsa of mesosoma, petiole and gaster about 0.01 mm. in diameter and spaced at about the same distance. Postgenal punctures, and those of sides of mesosoma and ventral and lateral aspects of petiole and gaster, larger, averaging about 0.02 mm. in diameter, and variously spaced, from intervals about equal to their maximum diameter to near contiguity.

(4) Mandibular dentition not visible, but apical two maxillary and terminal labial palpomeres with proportions similar to those of P. angusticeps.

Two additional worker specimens in the Museum of Comparative Zoology are apparently referable to P. dammermani and allow extension of its range to the Philippine Island of Negros. One of these is teneral, and was not measured; the other has the following dimensions: HL, 0.60 mm.; HW, 0.40 mm.; CI, 67; WL, 0.88 mm.; PW, 0.31 mm.; dorsal petiole width, 0.17 mm.; petiolar node index, 55; petiole height, 0.26 mm.; petiolar node length, 0.30 mm.; lateral petiolar index, 115. This specimen has lost its antennae, but the other has a scape index of about 100. The mandibular dentition consists of a single large apical tooth followed by a smaller preapical one and five small denticles. The palpal formula is maxillary 4: labial 2 (teneral dissected), with the segmental proportions as in P. angusticeps.

These specimens are very similar to the Javanese syntype in size, general form, sculpturation, pilosity and pubescence. The following slight differences from the syntype are noted: (1) the occipital border is somewhat less concave; (2) the posterior part of the subpetiolar process is more expanded (c.f. figs. 18 and 19); (3) the colour, even that of the less teneral specimen, is a much lighter yellowish-brown; this is almost - certainly due, however, to that specimen being partly callow.

Queen
Eguchi et al. (2006) - (alate and dealate queens). HL, 0.61–0.66 mm; HW, 0.42–0.44 mm; SL, 0.40–0.49 mm; EL, 0.11–0.13 mm; CI, 67–70; SI, 93–111; EI, 26–30; WL, 0.91–1.02 mm; PW, 0.37–0.38 mm; DPtW, 0.20 mm; DPtI, 53–54; PtH, 0.26–0.28 mm; PtNL, 0.30–0.33 mm; LPtI, 115–127 (N=3, but N=2 for DPtW and DPtI). Body ferruginous brown. Head in full-face view elongate, with very weakly convex sides and almost straight occipital border. Eye a little shorter than the width of apical antennal segment. Antennal scape when laid backward reaching or a little extending beyond the level of posterior margin of median ocellus; relative lengths of antennal segments II–XII as in the worker (see Fig. 10A); segment III almost as long as IV. Pronotum large; mesoscutum ca. 1.1–1.2 times as long as broad, in profile weakly convex; notauli absent; parapsidal lines very fine; scutoscutellar suture fine, very weakly and roundly curved posteriad; scutellum in profile with relatively steep posterior slope; axilla poorly separated from scutellum by an obscure impression but not by a suture; mesopleuron (except in its posteriormost part) well divided by a suture into anepisternum and katepisternum; raised median area of metanotum with gentle lateral slopes; suture between metepisternum and propodeum absent; a weak depression present dividing metepisternum into anepisternum and katepisternum; orifice of metapleural gland small, opening posterolaterad; posterior margin of propodeal dorsum in dorsal view moderately and broadly concave; posterior face of propodeum margined laterally with a well-developed translucent lamella which in profile forms an angle in upper portion. Petiole including subpetiolar process longer than high, in profile with very weakly concave posterior outline (above the articulation with gaster), in dorsal view gradually widened toward the midlength and parallel-sided in posterior half; posterodorsal margin of petiolar node in dorsal view almost straight or very weakly convex; subpetiolar process low, with an acute anterior projection and posteriorly with a low and sometimes obtusely angulate prominence. Abdominal segment III (gastral segment I) in profile relatively short, gently narrowed anteriad in the anterior 3/4 (in the queen from Bali the segment in profile a little slenderer). Wing structure and venation as in the male. Hind wing with three hamuli.

Male
Eguchi et al. (2006) - HL, 0.46–0.47 mm; HW, 0.46–0.47 mm; CI, 98–102; HD, 0.38–0.41 mm; HDI, 83–87; EL, 0.23–0.24 mm; SL, 0.30–0.32 mm; SI, 64–68; MstlL, 0.20–0.22 mm; MstlW, 0.20–0.23 mm; MstlI, 95–110; WL, 0.95 mm; PtNL, 0.28–0.32 mm; PtH, 0.16–0.20 mm; LPtI, 155–175 (N=3 but N=1 for WL). Head in lateral view relatively thin (HDI<90); protrusion of the frontoclypeal region relatively long, so that antennal insertion situated in the middle of its dorsal surface. Frontal carina low, not or slightly exceeding posterior margin of antennal insertions in full-face view. Eye relatively narrow, weakly widened ventrally. Antennal flagellum relatively long, filiform, with antennal segment III distinctly longer than segment II; segment XI (the third from apex) distinctly longer than broad; ventrolateral surface of the apical segment not concave. Mandible elongated triangular, with only a single strong apical tooth on its masticatory margin; basal angle distinct. Palpal formula: maxillary 4, labial 2; maxillary palp moderately long; the third palpomere distinctly longer than the second, and the apical distinctly longer than the third. With the mesosoma in lateral view, pronotum distinctly higher than mesoscutum; metanotum strongly produced posteriorly, and a suture separating it from mesoscutellum strongly notched; anepisternum of metapleuron separated from metakatepisternum and propodeum by a deep furrow; a suture between metakatepisternum and propodeum weak, but separation between the plates recognizable; dorsal margin of propodeum forming an angle with posterior slope. With the mesosoma in dorsal view, mesonotum lacking notauli; parapsidal lines distinct; axillae distinct; mesoscutellum slightly broader than long; declivitous face of the propodeum very slightly concave with weak, obtuse lateral lamellae. Petiole in lateral view without a distinct peduncle; node long with gentle anterior slope and steep posterior slope; posterodorsal margin angular; subpetiolar process narrowly developed with its apex sharp. Abdominal sternum IX long, with its apical margin gently pointed medially. Genitalia retractile. With the phallus in lateral view, the basal ring moderately long and its dorsal margin gently declined; dorsal margin of the basiparamere gently raised in its basal portion; posterodorsal slope gentle; digitus volsellaris simply curved ventrally in its apical portion; penis valve relatively thick and nearly straight, not strongly narrowed apically, and its apical portion blunt and slightly curved ventrally. Paramere thin; its expanded inner faces directed ventrally, of which apical portion is curved dorsally. Aedeagal apodeme relatively narrow. On fore wing, costa and radius apical to stigma vestigial; Rsf2 and Rsf3 completely absent; radial sector never reaching costal margin; Mf1, Rs+M and media apical to Rs+M present; cu-a cross vein present. On hind wing, Rsf4+5 present; jugal lobe absent.

Type Material
Eguchi et al. (2006) - Syntypes: 4 workers, Buitenzorg [Bogor], Java, Indonesia, 18/xii/1922, K. Dammerman [one worker examined].