Camponotus atriceps

This ant is known to be a synanthropic species. De Oliveira et al. (2015) found two colonies of C. atriceps opportunistically nesting in Cecropia pachystachya trees (southwest Bahia, Brazil).

Distribution based on Regional Taxon Lists
Nearctic Region: United States. Neotropical Region: Argentina, Barbados, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Uruguay, Venezuela.

Biology
De la Mora et al. (2016) found this species nesting in logs found in coffee farms in the Soconusco region of Chiapas, Mexico. A small percentage of pupae were found to be parasitized by eucharitid wasps.

Koch et al. (2018) sampled this species in Caryocar barsiliense trees, in southeastern Brazil cerrado, as part of a study examining species interactions in ant-plants.

Leite et al. (2013) studied the foraging behavior of Celeus obrieni (Kaempfer's Woodpecker), a bamboo specialists that feeds on ants. This field study was conducted in central-western Brazil on the plains of the Araguiai River from 2008-2010. The research focused on foraging by C. obrieni on Guadua paniculata, a large bamboo species, in cerradão habitat. Celeus obrieni are known to forage and breed in patches of G. paniculata. Ants use the hollow stems of the bamboo as a nesting resource.

Leite et al. found the woodpeckers, in comparison with standardized foraging observations on other plants around these same sites, concentrated their foraging on bamboo. A sample of stems (n = 900) that were not visited by the woodpecker showed ~40% contained ant nests. Thirteen ant species were found, with three being common (76% of all nests): Azteca fasciata, Camponotus atriceps, and Camponotus depressus. Most nests for these were located 2-4 m from the ground. Dead stems (59%) contained more nests than live stems. A second set of bamboo stem samples, those foraged on and drilled into by C. obrieni, were also inspected. Drilling was concentrated at the same height where the majority of ant nests were found in the unforaged-stem samples. Despite the much larger proportion of live stems within each stand, more dead (62%) than live stems were drilled. Fifty-nine percent of the drilled stems contained ant colonies. Nests of Camponotus depressus were the most common, Azteca fasciata less frequent, and Camponotus atriceps found in only four of the drilled stems.

Vanoye-Eligio et al. (2020) provide the following notes based on collections using Multilure-type traps from northeastern Mexico: Four worker minors and 1 major of Camponotus atriceps were detected. This is a synanthropic species often observed nesting on trunks in coffee farms (De la Mora et al. 2015).

While this species is known to nest in ant gardens it does not seem to be a true ant-garden taxon (i.e., one that is able to initiate ant gardens or is restricted to ant gardens) and is more likely to be a secondary resident or opportunistic ant-garden nester (Campbell et al., 2022; Marini-Filho, 1999).

Fungi

 * This taxon is a host for the fungi (Araujo et al., 2018; Shrestha et al., 2017; Sobczak et al., 2017),,  (Araujo et al., 2018; Shrestha et al., 2017), ,  and  (Quevillon, 2018; encounter mode primary; direct transmission; transmission outside nest).

Strepsiptera


Sobczak et al. (2017) - Studied C. atriceps infection by Ophiocordyceps camponoti-atricipis (an Entomopathogenic fungi) in an Atlantic forest biome area in Brazil. Describing the fungi: "All parasitized ants presented an abundant and visible external mycelium covering its thorax and mouthparts. The fruiting bodies of the fungi are initially white, turning light-brown over time. Single stroma produced from dorsal pronotum, averaging 8.6–16.9 mm, cylindrical, velvety and ginger brown at the base, becoming cram-pinkish towards apex. Fertile region of lateral cushions chocolate brown, darkening with age, variable in size. Perithecia immersed in the stroma, partially erumpent, flask-shaped, averaging 2–2.3 mm. Region near the stroma basis with short, exposed neck or ostiole, averaging 0.8–1.4 mm above the ant’s dorsal pronotum." The majority of the ants were found attached to leaves, biting onto the edge with closed mandibles, of coffee plants (Coffea arabica) at a height of 100-250 cm. It is unclear how typical such a response, i.e., height, plant leaf, plant species, etc., specificity holds for these two species but there does appear to be some behavioral modification produced by the fungus.

Nomenclature

 * . Formica atriceps Smith, F. 1858b: 44 (s.w.) BRAZIL (Pará, Rio de Janeiro).
 * Replacement name for Formica abdominalis Fabricius, 1804: 409. [Junior primary homonym of Formica abdominalis Latreille, 1802c: 175 (Bolton, 1995b: 86).]
 * [Note: atriceps oldest junior synonym of abdominalis Fabricius (synonymy by Roger, 1862c: 285, confirmed by Hashmi, 1973: 83); hence atriceps first available replacement name (Bolton, 1995b: 83).]
 * Mayr, 1862: 660 (q.); Forel, 1878: 76 (m.); Mariano, et al. 2001: 269 (k.).
 * Combination in Camponotus: Mayr, 1862: 660; Roger, 1862c: 285.
 * Junior synonym of abdominalis: Roger, 1862c: 285; Mayr, 1863: 398; Roger, 1863b: 4; Dalla Torre, 1893: 220; Forel, 1895b: 101; Emery, 1896d: 372; Forel, 1899c: 131; Santschi, 1913h: 42; Emery, 1925b: 107; Borgmeier, 1927c: 148; Kusnezov, 1952d: 197; Hashmi, 1973: 83; Brandão, 1991: 334.
 * Synonym of abdominalis: Forel, 1879a: 76.
 * [Note: Forel, 1879a: 76 gives atriceps as senior synonym, but abdominalis has priority.]
 * Subspecies of abdominalis: Emery, 1906c: 187; Forel, 1911c: 310; Santschi, 1912e: 534; Forel, 1912i: 70; Bruch, 1914: 228; Donisthorpe, 1915d: 349; Mann, 1916: 476; Luederwaldt, 1918: 50; Donisthorpe, 1927b: 404; Santschi, 1936b: 208; Kusnezov, 1952d: 197; Kempf, 1972a: 59.
 * Status as species: Mayr, 1862: 660 (redescription); Mayr, 1870a: 377; Forel, 1879a: 76; Mayr, 1884: 30; Forel, 1885a: 339 (redescription); Mayr, 1886d: 423; Cresson, 1887: 255; Emery, 1888c: 364; Emery, 1890a: 70; Emery, 1890b: 56; Cameron, 1891: 89; von Jhering, 1894: 372; Emery, 1894k: 61; Bolton, 1995b: 86; Wild, 2007b: 27; Branstetter & Sáenz, 2012: 255; Bezděčková, et al. 2015: 111; Wetterer, et al. 2016: 9; Mackay & Mackay, 2019: 752.
 * Senior synonym of atricipitoesuriens: Bolton, 1995b: 86.
 * Senior synonym of costaricensis: Bolton, 1995b: 86.
 * Senior synonym of cupiens: Bolton, 1995b: 86.
 * Senior synonym of depressidens: Bolton, 1995b: 86.
 * Senior synonym of epistomatus: Bolton, 1995b: 86.
 * Senior synonym of esuriens: Bolton, 1995b: 86.
 * Senior synonym of fuchsae: Bolton, 1995b: 86.
 * Senior synonym of fulvacea: Mayr, 1877a: 377; Bolton, 1995b: 86.
 * Senior synonym of fumata: Bolton, 1995b: 86.
 * Senior synonym of laevilata: Bolton, 1995b: 86.
 * Senior synonym of mediopallidus: Bolton, 1995b: 86.
 * Senior synonym of rabidus: Bolton, 1995b: 86.
 * Senior synonym of romani: Bolton, 1995b: 86.
 * Senior synonym of stercorarius: Bolton, 1995b: 86.
 * Senior synonym of taeniatus: Mayr, 1870a: 378 (in text); Forel, 1879a: 76; Forel, 1895b: 101; Bolton, 1995b: 86.
 * Senior synonym of transvectus: Bolton, 1995b: 86.
 * Senior synonym of ustulatus: Bolton, 1995b: 86.
 * Senior synonym of vulpinus: Bolton, 1995b: 86.
 * [Note: all the junior synonyms in Bolton, 1995b: 86, were initially synonymised with the junior primary homonym abdominalis Fabricius, by Hashmi, 1973: 82–83.]
 * Material of the unavailable names defrictus, flavobasalis, luisae, misionensis, panamanus referred to abdominalis by Hashmi, 1973: 82-83, Brandão, 1991: 334; referred to atriceps by Bolton, 1995b: 87.
 * Current subspecies: nominal plus nocens (status uncertain): Hashmi, 1973: 120; Bolton, 1995b: 114.
 * abdominalis. Formica abdominalis Fabricius, 1804: 409 (w.) (no state data, “Habitat in America meridionali”).
 * [Junior primary homonym of Formica abdominalis Latreille, 1802c: 175 (Emery, 1921a: 25).]
 * Wheeler, G.C. & Wheeler, J. 1953e: 196 (l.).
 * Combination in Camponotus: Roger, 1862c: 285;
 * combination in C. (Myrmothrix): Forel, 1912i: 91.
 * Status as species: Dalla Torre, 1893: 219; Forel, 1895b: 101; Emery, 1896d: 372 (in list); Forel, 1899c: 131; Forel, 1901c: 136; Wheeler, W.M. 1905b: 133; Wheeler, W.M. 1907a: 276; Forel, 1907e: 10; Forel, 1908b: 71; Forel, 1908e: 62; Santschi, 1913h: 42; Donisthorpe, 1915d: 349; Wheeler, W.M. 1916c: 14; Wheeler, W.M. 1916d: 330; Mann, 1916: 476; Crawley, 1916b: 376; Wheeler, W.M. 1922c: 15; Mann, 1922: 53; Santschi, 1922c: 107; Wheeler, W.M. 1923a: 5; Emery, 1925b: 107; Borgmeier, 1927c: 148; Donisthorpe, 1927b: 403; Borgmeier, 1934: 111; Menozzi, 1935b: 202; Santschi, 1936b: 207; Wheeler, W.M. 1942: 256; Creighton, 1950a: 396; Kusnezov, 1952d: 197; Kempf, 1961b: 522; Kempf, 1970b: 341; Kempf, 1972a: 59; Hashmi, 1973: 82 (redescription); Zolessi, et al. 1988: 6; Brandão, 1991: 333.
 * Senior synonym of atriceps: Roger, 1862c: 285; Mayr, 1863: 398; Roger, 1863b: 4; Dalla Torre, 1893: 220; Forel, 1895b: 101; Emery, 1896d: 372; Forel, 1899c: 131; Santschi, 1913h: 42; Emery, 1925b: 107; Borgmeier, 1927c: 148; Kusnezov, 1952d: 197; Hashmi, 1973: 83; Brandão, 1991: 334.
 * Replacement name: Formica atriceps Smith, F. 1858b: 44.
 * [Note: atriceps oldest junior synonym of abdominalis Fabricius (synonymy by Roger, 1862c: 285, confirmed by Hashmi, 1973: 83); hence atriceps first available replacement name (Bolton, 1995b: 83).]
 * atricipitoesuriens. Camponotus abdominalis var. atricipitoesuriens Dalla Torre, 1893: 220.
 * [First available use of Camponotus atriceps r. esuriens var. atricipitoesuriens Forel, 1879a: 77 (w.) MEXICO (Veracruz), HAITI; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Emery, 1896d: 372 (in list); Forel, 1899c: 132; Emery, 1925b: 108; Kempf, 1972a: 61.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 87.
 * costaricensis. Camponotus abdominalis var. costaricensis Forel, 1899c: 132 (w.) COSTA RICA.
 * Menozzi, 1927d: 339 (q.).
 * Combination in C. (Myrmothrix): Emery, 1925b: 107.
 * As unavailable (infrasubspecific) name: Santschi, 1936b: 212; Kempf, 1972a: 61.
 * Subspecies of abdominalis: Forel, 1902b: 170; Forel, 1914e: 13; Emery, 1925b: 107; Menozzi, 1927c: 268; Menozzi, 1927d: 339; Wheeler, W.M. 1942: 256.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 94.
 * cupiens. Camponotus abdominalis subsp. cupiens Forel, 1908c: 410 (s.w.q.) BRAZIL (São Paulo).
 * Subspecies of abdominalis: Forel, 1911e: 290; Luederwaldt, 1918: 50; Emery, 1925b: 107; Borgmeier, 1927c: 149; Eidmann, 1936b: 94; Santschi, 1936b: 210 Santschi, 1939e: 328; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 95.
 * depressidens. Camponotus (Myrmothrix) abdominalis st. depressidens Santschi, 1922c: 107 (s.w.) FRENCH GUIANA.
 * Subspecies of abdominalis: Emery, 1925b: 107; Santschi, 1936b: 211; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82.
 * Junior synonym of atriceps: Bolton, 1995b: 96.
 * epistomatus. Camponotus (Myrmothrix) abdominalis st. epistomatus Santschi, 1936b: 208 (w.) COLOMBIA.
 * Subspecies of abdominalis: Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 97.
 * esuriens. Formica esuriens Smith, F. 1858b: 54 (s.w.) MEXICO (no state data).
 * Norton, 1868c: 1 (q.m.).
 * Combination in Camponotus: Roger, 1863b: 5;
 * combination in C. (Myrmothrix): Emery, 1925b: 107.
 * Status as species: Roger, 1863b: 5; Mayr, 1863: 398; Norton, 1868a: 59; Norton, 1868c: 1; Emery, 1878a: ix (in list); Cresson, 1887: 255; Dalla Torre, 1893: 229; Pergande, 1896: 862.
 * Subspecies of atriceps: Forel, 1879a: 76; Forel, 1885a: 340; Dalla Torre, 1893: 230.
 * Subspecies of abdominalis: Emery, in Dalla Torre, 1893: 219 (footnote); Emery, 1896d: 372 (in list); Forel, 1899c: 133; Forel, 1901h: 70; Forel, 1909a: 252; Emery, 1925b: 107; Kutter, 1931a: 62; Wheeler, W.M. 1934f: 138; Wheeler, W.M. 1935g: 40; Santschi, 1936b: 213; Stitz, 1937: 135; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 83; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 98.
 * fuchsae. Camponotus abdominalis subsp. fuchsae Forel, 1908c: 409 (s.w.q.m.) BRAZIL (São Paulo).
 * Combination in C. (Myrmothrix): Bruch, 1914: 228.
 * Subspecies of abdominalis: Bruch, 1914: 228; Luederwaldt, 1918: 50; Emery, 1925b: 108; Borgmeier, 1927c: 149; Santschi, 1936b: 213; Kusnezov, 1952d: 198; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * fulvacea. Formica fulvacea Norton, 1868a: 60, pl. 2, fig. 1 (s.w.) MEXICO (Veracruz).
 * Combination in Camponotus: Norton, 1868c: 2.
 * As unavailable (infrasubspecific) name: Emery, 1896d: 372 (in list) Forel, 1899c: 133; Kempf, 1972a: 60.
 * Status as species: Norton, 1868c: 2.
 * Subspecies of esuriens: Dalla Torre, 1893: 230.
 * Junior synonym of esuriens: Forel, 1879a: 76; Forel, 1885a: 340.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Mayr, 1870a: 377; Bolton, 1995b: 86.
 * fumata. Camponotus (Tanaemyrmex) abdominalis var. fumata Stitz, 1933: 71 (w.) VENEZUELA.
 * Subspecies of abdominalis: Santschi, 1936b: 209; Kempf, 1972a: 59.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * laevilata. Camponotus (Myrmothrix) abdominalis var. laevilata Santschi, 1922c: 108 (s.q.) BRAZIL (São Paulo).
 * As unavailable (infrasubspecific) name: Santschi, 1936b: 209; Kempf, 1972a: 60.
 * Subspecies of abdominalis: Emery, 1925b: 107.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * mediopallidus. Camponotus abdominalis r. mediopallidus Forel, 1901h: 70.
 * [First available use of Camponotus abdominalis r. ustulatus var. mediopallidus Forel, 1899c: 132 (w.q.) COLOMBIA; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Wheeler, W.M. 1918b: 26.
 * Subspecies of abdominalis: Forel, 1902b: 170; Forel, 1907a: 34; Crawley, 1916b: 376; Santschi, 1922c: 107; Emery, 1925b: 108; Borgmeier, 1934: 111; Santschi, 1936b: 209; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * rabidus. Camponotus (Myrmothrix) abdominalis st. rabidus Santschi, 1922c: 107 (s.w.q.) PERU.
 * Subspecies of abdominalis: Emery, 1925b: 108; Santschi, 1936b: 214; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * romani. Camponotus (Myrmothrix) abdominalis subsp. romani Wheeler, W.M. 1923a: 5 (s.w.) BRAZIL (Amazonas).
 * Subspecies of abdominalis: Emery, 1925b: 108; Borgmeier, 1927c: 149; Santschi, 1936b: 215; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * stercorarius. Camponotus atriceps r. stercorarius Forel, 1885a: 340 (w.q.m.) GUATEMALA.
 * Wheeler, G.C. & Wheeler, J. 1953e: 196 (l.).
 * Combination in C. (Myrmothrix): Crawley, 1916b: 376.
 * Status as species: Dalla Torre, 1893: 253.
 * Subspecies of atriceps: Emery, 1890b: 56; Emery, 1894k: 61.
 * Subspecies of abdominalis: Emery, in Dalla Torre, 1893: 219 (footnote); Emery, 1896d: 372 (in list); Forel, 1899c: 133; Forel, 1901h: 70; Wheeler, W.M. 1907a: 276; Forel, 1908b: 71; Forel, 1908e: 62; Donisthorpe, 1915d: 349; Crawley, 1916b: 376; Forel, 1922: 99; Mann, 1922: 53; Wheeler, W.M. 1923a: 5; Emery, 1925b: 108; Donisthorpe, 1927b: 404; Wheeler, W.M. 1933b: 28; Santschi, 1936b: 213; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * taeniatus. Camponotus taeniatus Roger, 1863a: 149 (q.) MEXICO (no state data).
 * Status as species: Roger, 1863b: 5; Mayr, 1863: 459.
 * Junior synonym of abdominalis: Dalla Torre, 1893: 220; Forel, 1899c: 131; Santschi, 1913h: 42; Emery, 1925b: 107; Kempf, 1972a: 59; Hashmi, 1973: 82.
 * Junior synonym of atriceps: Mayr, 1870a: 378 (in text); Forel, 1879a: 76; Forel, 1895b: 101; Bolton, 1995b: 86.
 * transvectus. Camponotus abdominalis subsp. transvectus Wheeler, W.M. 1910d: 326 (s.w.q.m.) U.S.A. (Texas).
 * Combination in C. (Myrmothrix): Emery, 1925b: 108.
 * Subspecies of abdominalis: Emery, 1925b: 108; Santschi, 1936b: 213; Creighton, 1950a: 396; Smith, M.R. 1951a: 844; Smith, D.R. 1979: 1431.
 * Junior synonym of abdominalis: Hashmi, 1973: 82.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * ustulatus. Camponotus atriceps r. ustulatus Forel, 1885a: 339 (s.w.) GUATEMALA.
 * Santschi, 1936b: 211 (q.m.).
 * Combination in C. (Myrmothrix): Emery, 1925b: 108.
 * As unavailable (infrasubspecific) name: Forel, 1914e: 13 (in text).
 * Status as species: Dalla Torre, 1893: 255.
 * Subspecies of abdominalis: Emery, in Dalla Torre, 1893: 219 (footnote); Emery, 1896d: 372 (in list); Forel, 1899c: 132; Wheeler, W.M. 1907a: 276; Wheeler, W.M. 1918b: 26; Emery, 1925b: 108; Santschi, 1936b: 211; Kempf, 1972a: 61.
 * Junior synonym of abdominalis: Hashmi, 1973: 82; Brandão, 1991: 334.
 * Junior synonym of atriceps: Bolton, 1995b: 86.
 * vulpinus. Camponotus vulpinus Mayr, 1862: 658, 770 (w.q.) VENEZUELA, MEXICO (no state data).
 * Junior synonym of esuriens: Roger, 1863b: 5; Mayr, 1863: 398; Mayr, 1870a: 377 (in text); Forel, 1879a: 76; Forel, 1885a: 340; Dalla Torre, 1893: 229; Emery, 1896d: 372; Pergande, 1896: 862; Emery, 1925b: 108; Kempf, 1972a: 60.
 * Junior synonym of abdominalis: Hashmi, 1973: 83.
 * Junior synonym of atriceps: Bolton, 1995b: 86.

References based on Global Ant Biodiversity Informatics

 * Adams B. J., S. A. Schnitzer, and S. P. Yanoviak. 2019. Connectivity explains local ant community structure in a Neotropical forest canopy: a large-scale experimental approach. Ecology 100(6): e02673.
 * Andrade T., G. D. V. Marques, K. Del-Claro. 2007. Diversity of ground dwelling ants in Cerrado: an analysis of temporal variations and distinctive physiognomies of vegetation (Hymenoptera: Formicidae). Sociobiology 50(1): 121-134.
 * Antoniazzi R., R. N. S. L. Garoo, W. Dattilo, S. P. Ribeiro, and F. S. Neves. 2019. Ant species richness and interactions in canopies of two distinct successional stages in a tropical dry forest. The Science of Nature 106: 20
 * Armbrecht I., I. Tischer, and P. Chacon. 2001. Nested subsets and partition patterns in ant assemblages (Hymenoptera, Formicidae) of Colombian dry forest fragments. Pan-Pacific Entomologist 77(3): 196-209.
 * Arruda F. V., M. A. Pesquero, D. G. Marcelino, G. A. Leiter, J. H. C. Delabie, and R. Fagundes. 2015. Size and condition of bamboo as structural factors behind the vertical stratification of the bamboo-nesting ant community. Insectes Sociaux DOI 10.1007/s00040-015-0440-4
 * Boer P. 2019. Ants of Curacao, species list. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/SPECIES%20LIST%20CURACAO.html
 * Borgmeier T. 1934. Contribuição para o conhecimento da fauna mirmecológica dos cafezais de Paramaribo, Guiana Holandesa (Hym. Formicidae). Archivos do Instituto de Biologia Vegetal (Rio de Janeiro) 1: 93-111.
 * Bouwma, A.M., K.J. Howard and R.L. Jeanne. 2007. Rates of Predation by Scouting-and-Recruiting Ants on the Brood of a Swarm-Founding Wasp in Costa Rica. Biotropica 39(6):719-724
 * Brandao, C.R.F. 1991. Adendos ao catalogo abreviado das formigas da regiao neotropical (Hymenoptera: Formicidae). Rev. Bras. Entomol. 35: 319-412.
 * Bruch C. 1914. Catálogo sistemático de los formícidos argentinos. Revista del Museo de La Plata 19: 211-234.
 * Byk J., K. Del-Claro. 2010. Nectar- and pollen-gathering Cephalotes ants provide no protection against herbivory: a new manipulative experiment to test ant protective capabilities. Acta Ethol. 13: 33-38.
 * Bächtold A., K. Del-Claro, L. A. Kaminski, A. V. L. Freitas, and P. S. Oliveira. 2012. Natural history of an ant-plant-butterfly interaction in a Neotropical savanna. Journal of Natural History 46: 943-954.
 * Cameron P. 1891. Appendix. Hym. Formicidae. Pp. 89-95 in: Whymper, E. Travels amongst the Great Andes. London: J. Murray, xxii + 147 pp.
 * Campos R. B. F., J. H. Schoereder, and C. F. Sperber. 2007. Small-scale patch dynamics after disturbance in litter ant communities. Basic and Applied Ecology 8: 3643.
 * Castano-Meneses, G., M. Vasquez-Bolanos, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha, and I. Alcala-Martinez. 2015. Avances de Formicidae de Mexico. Universidad Nacional Autonoma de Mexico.
 * Chacon de Ulloa P., A. M. Osorio-Garica, R. Achury, and C. Bermudez-Rivas. 2012. Hormigas (Hymenoptera: Formicidae) del Bosque seco tropical (Bs-T) de la cuenca alta del rio Cauca, Colombia. Biota Colombiana 13(2): 165-181.
 * Chacon de Ulloa P., S. Valdes-Rodriguez, A. Hurtado-Giraldo, and M. Cleopatra Pimienta. 2014. Arboreal ants of Gorgona National Park (Pacific of Colombia). Rev. Biol. Trop. 62(1): 277-287.
 * Coelho M. S., G. W. Fernandes, J. C. Santos, and J. H. C. Delabie. 2009. Ants (Hymenoptera: Formicidae) as bioindicators of land restoration in a Brazilian Atlantic forest fragment. Sociobiology 54(1): 51-63.
 * Cokendolpher J. C., and O. F. Francke. 1990. The ants (Hymenoptera, Formicidae) of western Texas. Part II. Subfamilies Ecitoninae, Ponerinae, Pseudomyrmecinae, Dolichoderinae, and Formicinae. Special Publications, the Museum. Texas Tech University 30:1-76.
 * Costa-Milanez C. B., G. Lourenco-Silva, P. T. A. Castro, J. D. Majer, and S. P. Ribeiro. 2014. Are ant assemblages of Brazilian veredas characterised by location or habitat type? Braz. J. Biol. 74(1): 89-99.
 * Crawley W. C. 1916. Ants from British Guiana. Ann. Mag. Nat. Hist. 8(17): 366-378.
 * Cuautle, M. and V. Rico-Gray. 2003. The Effect of Wasps and Ants on the Reproductive Success of the Extrafloral Nectaried Plant Turnera ulmifolia (Turneraceae). Functional Ecology 17(3):417-423
 * Cupul-Magana F. G., and M. Vasquez-Bolanos. 2019. New records of ants (Hymenoptera: Formicidae) from María Cleofas Island, Gulf of California, Mexico. Dugesiana 26(1): 27-29.
 * Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
 * Dattilo W., N. Sibinel, J. C. F. Falcao, and R. V. Nunes. 2011. Ant fauna in a urban remnant of Atlantic forest in the municipuality of Marilia, state of Sao Paulo, Brazil. Bioscience Journal 27(3): 494-504.
 * Davidson, D.W. 2005. Ecological stoichiometry of ants in a New World rain forest. Oecologia 142:221-231
 * De la Mora, A., C. J. Murnen, and S. M. Philpott. 2013. Local and landscape drivers of biodiversity of four groups of ants in Neotropical coffee landscapes. Biodiversity and Conservation 22: 871-888.
 * De la Mora, A., G. Perez-Lachaud, J. P.Lachaud, and S. M. Philpott. 2015. Local and landscape drivers of ant parasitism in a coffee landscape. Environmental Entomology 44: 939-950.
 * Dejean, A., S. Durou, I. Olmsted, R.R. Snelling and J. Orivel. 2003. Nest Site Selection by Ants in a Flooded Mexican Mangrove, with Special Reference to the Epiphytic Orchid Myrmecophila christinae. Journal of Tropical Ecology 19(3) :325-331
 * Dejean, A., S. Durou, I. Olmsted, R.R. Snelling and J. Orivel. 2003. Nest Site Selection by Ants in a Flooded Mexican Mangrove, with Special Reference to the Epiphytic Orchid Myrmecophila christinae. Journal of Tropical Ecology 19(3):325-331
 * Del Toro, I., M. Vázquez, W.P. Mackay, P. Rojas and R. Zapata-Mata. Hormigas (Hymenoptera: Formicidae) de Tabasco: explorando la diversidad de la mirmecofauna en las selvas tropicales de baja altitud. Dugesiana 16(1):1-14.
 * Delabie J. H. C., R. Céréghino, S. Groc, A. Dejean, M. Gibernau, B. Corbara, and A. Dejean. 2009. Ants as biological indicators of Wayana Amerindian land use in French Guiana. Comptes Rendus Biologies 332(7): 673-684.
 * Díaz-Castelazo C., V. Rico-Gray, P. S. Oliveira, and M. Cuautle. 2004. Extrafloral nectary-mediated ant-plant interactions in the coastal vegetation of Veracruz, Mexico: Richness, occurrence, seasonality and ant foraging patterns. Ecoscience 11: 472-481.
 * Emery C. 1890. Studii sulle formiche della fauna neotropica. Bull. Soc. Entomol. Ital. 22: 38-8
 * Emery C. 1890. Voyage de M. E. Simon au Venezuela (Décembre 1887 - Avril 1888). Formicides. Annales de la Société Entomologique de France (6)10: 55-76.
 * Emery C. 1894. Estudios sobre las hormigas de Costa Rica. Anales del Museo Nacional de Costa Rica 1888-1889: 45-64.
 * Emery C. 1896. Formiche raccolte dal dott. E. Festa nei pressi del golfo di Darien. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 11(229): 1-4.
 * Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
 * Escalante Gutiérrez J. A. 1993. Especies de hormigas conocidas del Perú (Hymenoptera: Formicidae). Revista Peruana de Entomología 34:1-13.
 * Fernandes I., and J. de Souza. 2018. Dataset of long-term monitoring of ground-dwelling ants (Hymenoptera: Formicidae) in the influence areas of a hydroelectric power plant on the Madeira River in the Amazon Basin. Biodiversity Data Journal 6: e24375.
 * Fernandes, P.R. XXXX. Los hormigas del suelo en Mexico: Diversidad, distribucion e importancia (Hymenoptera: Formicidae).
 * Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
 * Flores-Flores, R. V., A. Aguirre, D. V. Anjos, F.R. Neves, R. I. Campos, and W. Dattilo. 2018. Food source quality and ant dominance hierarchy influence the outcomes of ant-plant interactions in an arid environment. Acta Oecologica 87: 13-19.
 * Flores-Maldonado K. Y., S. A. Phillips, and G. Sanchez-Ramos. 1999. The myrmecofauna (Hymenoptera: Formicidae) along an altitudinal gradient in the Sierra Madre Oriental of Northeastern Mexico. The Southwestern Naturalist 44(4): 457-461.
 * Flores-Maldonado, K. Y., S. A. Phillips-Jr, and G. Sanchez-Ramos. 1999. The myrmecofauna (Hymenoptera: Formicidae) along an altitudinal gradient in the Sierra Madre Oriental of Northeastern Mexico. The Southwestern Naturalist 44: 457-461.
 * Forel A. 1897. Quelques Formicides de l'Antille de Grenada récoltés par M. H. H. Smith. Transactions of the Entomological Society of London. 1897: 297-300.
 * Forel A. 1901. Formiciden des Naturhistorischen Museums zu Hamburg. Neue Calyptomyrmex-, Dacryon-, Podomyrma- und Echinopla-Arten. Mitt. Naturhist. Mus. Hambg. 18: 43-82.
 * Forel A. 1902. Quatre notices myrmécologiques. Annales de la Société Entomologique de Belgique 46: 170-182.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1907. Formicides du Musée National Hongrois. Ann. Hist.-Nat. Mus. Natl. Hung. 5: 1-42.
 * Forel A. 1908. Ameisen aus Sao Paulo (Brasilien), Paraguay etc. gesammelt von Prof. Herm. v. Ihering, Dr. Lutz, Dr. Fiebrig, etc. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 58: 340-418.
 * Forel A. 1908. Catálogo systemático da collecção de formigas do Ceará. Boletim do Museu Rocha 1(1): 62-69.
 * Forel A. 1908. Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles 44: 35-72.
 * Forel A. 1911. Ameisen des Herrn Prof. v. Ihering aus Brasilien (Sao Paulo usw.) nebst einigen anderen aus Südamerika und Afrika (Hym.). Deutsche Entomologische Zeitschrift 1911: 285-312.
 * Forel A. 1911. Die Ameisen des K. Zoologischen Museums in München. Sitzungsber. Math.-Phys. Kl. K. Bayer. Akad. Wiss. Münch. 11: 249-303.
 * Forel A. 1922. Glanures myrmécologiques en 1922. Revue Suisse de Zoologie. 30: 87-102.
 * Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
 * Gove, A. D., J. D. Majer, and V. Rico-Gray. 2009. Ant assemblages in isolated trees are more sensitive to species loss and replacement than their woodland counterparts. Basic and Applied Ecology 10: 187-195.
 * Hashmi A. A. 1973. A Revision of the Neotropical Ant Subgenus Myrmothrix of Genus Camponotus (Hymenoptera: Formicidae). Studia Ent. 16(1-4): 1-140.
 * Hernandez, F. Varela and G. Castano-Meneses. 2010. Checklist, Biological Notes and Distribution of Ants (Hymenoptera: Formicidae) from Barranca de Metztitlán Biosphere Reserve, Hidalgo, Mexico. Sociobiology 56(2):397-434
 * Hernandez-Ruiz P., G. Castano-Meneses, and Z. Cano-Santana. 2009. Composition and functional groups of epiedaphic ants (Hymenoptera: Formicidae) in irrigated agroecosystem and in nonagricultural areas. Pesq. agropec. bras., Brasília, 44(8): 904-910.
 * INBio Collection (via Gbif)
 * Jaffe, Klaus and Lattke, John. 1994. Ant Fauna of the French and Venezuelan Islands in the Caribbean in Exotic Ants, editor D.F. Williams. 182-190.
 * Kempf W. W. 1961. A survey of the ants of the soil fauna in Surinam (Hymenoptera: Formicidae). Studia Entomologica 4: 481-524.
 * Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
 * Kusnezov N. 1952. El género Camponotus en la Argentina (Hymenoptera, Formicidae). Acta Zoologica Lilloana 12: 183-252.
 * Kusnezov N. 1953. La fauna mirmecológica de Bolivia. Folia Universitaria. Cochabamba 6: 211-229.
 * Kusnezov N. 1978. Hormigas argentinas: clave para su identificación. Miscelánea. Instituto Miguel Lillo 61:1-147 + 28 pl.
 * Landero Torres I., M. A. Garcia Martinez, H. Oliva Rivera, M. E. Galindo Tovar, H. Lee Espinosa, and J. Murguia Gonzales. 2010. Comparación de dos muestreos de hormigas del suelo en la barranca de Metlác, Fortín de las Flores, Veracruz, México. Revista Científica UDO Agrícola 10 (1): 173-178.
 * Lattke J. E., and M. Velez, and N. Aguirre. 2016. Survey of ants in dry forests of Southwestern Ecuador (Hymenoptera: Formicidae). Sociobiology 63(3): 909-918.
 * Longino J. T. 2013. Ants of Honduras. Consulted on 18 Jan 2013. https://sites.google.com/site/longinollama/reports/ants-of-honduras
 * Longino J. T. 2013. Ants of Nicargua. Consulted on 18 Jan 2013. https://sites.google.com/site/longinollama/reports/ants-of-nicaragua
 * Longino J. et al. ADMAC project. Accessed on March 24th 2017 at https://sites.google.com/site/admacsite/
 * Longino, J.T. 2010. Personal Communication. Longino Collection Database
 * Lopez-Riquelme, G. O., and M. L. Fanjul Moles. 2006. The ant Camponotus atriceps shows electroretinogram circadian rhythms. Biological Rhythm Research 37: 111-115.
 * Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
 * Luna Bello G. O. 2005. Composicion y estructura de la comunidad de hormigas a lo largo de un gradiente de intensificacion agricola en zonas de bosque tropical humedo en la region autonoma Atlantico Sur, Nicaragua. PhD Universidad Centroamericana, Facultad de Ciencia y Tecnologia del Ambiente carrera de Ecologi y Desarrollo. 127 pages
 * Luna, P., Y. Penaloza-Arellanes, A. L. Castillo-Meza, J. H. Garcia-Chavez, and W. Dattilo. 2018. Beta diversity of ant-plant interactions over day-night periods and plant physiognomies in a semiarid environment. Journal of Arid Environments 156: 69-76.
 * Maes, J.-M. and W.P. MacKay. 1993. Catalogo de las hormigas (Hymenoptera: Formicidae) de Nicaragua. Revista Nicaraguense de Entomologia 23.
 * Majer J. D., and J. H. C. Delabie. 1994. Comparison of the ant communities of annually inundated and terra firme forests at Trombetas in the Brazilian Amazon. Insectes Sociaux 41: 343-359.
 * Mann W. M. 1922. Ants from Honduras and Guatemala. Proceedings of the United States National Museum 61: 1-54.
 * Marques G. D. V., and K. Del-Claro. 2006. The Ant Fauna in a Cerrado area: The Influence of Vegetation Structure and Seasonality (Hymenoptera: Formicidae). Sociobiology 47(1): 1-18.
 * Menozzi C. 1927. Formiche raccolte dal Sig. H. Schmidt nei dintorni di San José di Costa Rica. Entomologische Mitteilungen. Berlin-Dahlem. 16: 266-277.
 * Menozzi C. 1935. Spedizione del Prof. Nello Beccari nella Guiana Britannica. Hymenoptera-Formicidae. Redia. 21: 189-203.
 * Mertl A. L., J. F. A. Traniello, K. Ryder Wilkie, and R. Constantino. 2012. Associations of two ecologically significant social insect taxa in the litter of an amazonian rainforest: is there a relationship between ant and termite species richness? Psyche doi:10.1155/2012/312054
 * Mirmecofauna de la reserva ecologica de San Felipe Bacalar
 * Munhae C. B., Z. A. F. N. Bueno, M. S. C. Morini, and R. R. Silva. 2009. Composition of the Ant Fauna (Hymenoptera: Formicidae) in Public Squares in Southern Brazil. Sociobiology 53(2B): 455-472.
 * Nascimento Santos M., J. H. C. Delabie, and J. M. Queiroz. 2019. Biodiversity conservation in urban parks: a study of ground-dwelling ants (Hymenoptera: Formicidae) in Rio de Janeiro City. Urban Ecosystems https://doi.org/10.1007/s11252-019-00872-8
 * Navarro, E.V. Vergara, H. Echavarria Sanchez, F.J. Serna Cardona. 2007. Hormigas (Hymenoptera: Formicidae) asociadas al arboretum de la Universidad Nacional de Colombia, sede Medellin. Boletín Sociedad Entomológica Aragonesa 40:497-505.
 * Neves F. S., K. S. Queiroz-Dantas, W. D. da Rocha, and J. H. C. Delabie. 2013. Ants of Three Adjacent Habitats of a Transition Region Between the Cerrado and Caatinga Biomes: The Effects of Heterogeneity and Variation in Canopy Cover. Neotrop Entomol 42: 258268.
 * Neves F. S., R. F. Braga, M. M. do Espirito-Santo, J. H. C. Delabie, G. Wilson Fernandes, and G. A. Sanchez-Azofeifa. 2010. Diversity of Arboreal Ants In a Brazilian Tropical Dry Forest: Effects Of Seasonality and Successional Stage. Sociobiology 56(1): 1-18.
 * Nogueira Rossi M., and H. G. Fowler. 2004. Predaceous Ant Fauna in New Sugarcane Fields in the State of São Paulo, Brazil. Brazilian Archives of Biology and Technology 47(5): 805-811.
 * Nunes F. A., G. B. Martins Segundo, Y. B. Vasconcelos, R. Azevedo, and Y. Quinet. 2011. Ground-foraging ants (Hymenoptera: Formicidae) and rainfall effect on pitfall trapping in a deciduous thorn  woodland (Caatinga), Northeastern Brazil. Rev. Biol. Trop. 59 (4): 1637-1650.
 * O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
 * Pacheco R., H. L. Vasconcelos, S. Groc, G. P. Camacho, and T. L. M. Frizzo. 2013. The importance of remnants of natural vegetation for maintaining ant diversity in Brazilian agricultural landscapes. Biodivers. Conserv. DOI 10.1007/s10531-013-0463-y
 * Pergande, T. 1895. Mexican Formicidae. Proceedings of the California Academy of Sciences Ser. 2 :850-896
 * Philpott, S.M., P. Bichier, R. Rice, and R. Greenberg. 2007. Field testing ecological and economic benefits of coffee certification programs. Conservation Biology 21: 975-985.
 * Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
 * Quiroz-Robledo L., and J. Valenzuela-Gonzalez. 1995. A comparison of ground ant communities in a tropical rainforest and adjacent grassland in Los Tuxtlas, Veracruz, Mexico. Southwestern Entomologist 20(2): 203-213.
 * Radoszkowsky O. 1884. Fourmis de Cayenne Française. Trudy Russkago Entomologicheskago Obshchestva 18: 30-39.
 * Ramirez M., and M. L. Enriquez. 2003. Importance and diversity of ants in silvopastoral systems in the Cauca Valley, Colombia. Livestock Research for Rural Development 15(1): http://www.lrrd.org/lrrd15/1/rami151.htm
 * Ramos-Lacau L. S., P. S. D. Silva, S. Lacau , J. H. C. Delabie, and O. C. Bueno. 2012. Nesting architecture and population structure of the fungus-growing ant Cyphomyrmex transversus (Formicidae: Myrmicinae: Attini) in the Brazilian coastal zone of Ilhéus, Bahia, Annales de la Société entomologique de France (N.S.): International Journal of Entomology. 48(3-4): 439-445.
 * Reynoso-Campos J. J., J. A. Rodriguez-Garza, and M. Vasquez-Bolanos. 2015. Hormigas (Hymenoptera: Formicidae) de la Isla Cozumel, Quintana Roo, Mexico (pp. 27-39). En: Castaño Meneses G., M. Vásquez-Bolaños, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha e I. Alcalá-Martínez (Coords.). Avances de Formicidae de  México.  UNAM,  Universiad  de  Guadalajara, Guadalajara, Jalisco.
 * Ribas C. R., J. H. Schoereder, M. Pic, and S. M. Soares. 2003. Tree heterogeneity, resource availability, and larger scale processes regulating arboreal ant species richness. Austral Ecology 28(3): 305-314.
 * Ribas C. R., R. R. C. Solar, R. B. F. Campos, F. A. Schmidt, C. L. Valentim, and J. H. Schoereder. 2012. Can ants be used as indicators of environmental impacts caused by arsenic? Insect Conserv 16: 413421.
 * Rico-Gray, V., and L. B. Thien. 1989. Ant-mealybug interaction decreases reproductive fitness of Schomburgkia tibicinis (Orchidaceae) in Mexico. Journal of Tropical Ecology 5: 109-112.
 * Rico-Gray, V., and L. B. Thien. 1989. Effect of different ant species on reproductive fitness of Schomburgkia tibicinis (Orchidaceae). Oecologia 81: 487-489.
 * Rico-Gray,V., J.G. Garcia-Franco, M. Palacios-Rios, C. Diaz-Castelazo, V. Parra-Tabla and J.A. Navarro. 1998. Geographical and Seasonal Variation in the Richness of Ant-Plant Interactions in Mexico. Biotropica 30(2):190-200.
 * Rios-Casanova, L., A. Valiente-Banuet, and V. Rico-Gray. (2004). Las hormigas del Valle de Tehuacan (Hymenoptera: Formicidae): una comparacion con otras zonas aridas de Mexico. Acta Zoologica Mexicana 20: 37-54.
 * Rivas-Arancibia, S. P., H. Carrillo-Ruiz, A. Bonilla-Arce, D. M. Figueroa-Castro, and A. R. Andres-Hernandez. 2014. Effect of disturbance on the ant community in a semiarid region of central Mexico. Applied Ecology and Environmental Research 12: 703-716.
 * Rodriguez E. R., and J. E. Lattke. 2012. Diversidad de hormigas en un gradiente altitudinal de la cordillera de la Costa, Venezuela. Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 50: 295?304.
 * Romero H., and K. Jaffe. 1989. A comparison of methods for sampling ants (Hymenoptera, Formicidae) in savannas. Biotropica 21(4): 348-352.
 * Rosa da Silva R. 1999. Formigas (Hymenoptera: Formicidae) do oeste de Santa Catarina: historico das coletas e lista atualizada das especies do Estado de Santa Catarina. Biotemas 12(2): 75-100.
 * Ríos-Casanova, L., A. Valiente-Banuet and V. Rico-Gray. 2004. Las hormigas del Valle de Tehuacan (Hymenoptera: Formicidae): Una comparacion con otras zonas aridas de Mexico. Acta Zoologica Mexicana 20(1):37-54
 * Salinas P. J. 2010. Catalogue of the ants of the Táchira State, Venezuela, with notes on their biodiversity, biogeography and ecology (Hymenoptera: Formicidae: Amblyioponinae, Ponerinae, Proceratiinae, Myrmicinae, Ecitoninae, Formicinae, Pseudomyrmecinae, Dolichoderinae). Boletín de la SEA 47: 315-328.
 * Santos J. C., and Del-Claro K. 2001. Interactions between ants, herbivores and extrafloral nectaries in Tocoyena formosa (Cham & Schlechtd.) K. Schum. (Rubiaceae) in cerrado vegetation. Rev. Bras. Zoociencias Juiz de Fora 3(1): 77-92.
 * Santschi F. 1922. Camponotus néotropiques. Annales de la Société Entomologique de Belgique 62: 97-124.
 * Santschi F. 1936. Contribution à l'étude des fourmis de l'Amérique du Sud. Revista de Entomologia (Rio de Janeiro). 6: 196-218.
 * Santschi F. 1939. Études et descriptions de fourmis néotropiques. Revista de Entomologia (Rio de Janeiro). 10: 312-330.
 * Schoereder J. H., T. G. Sobrinho, M. S. Madureira, C. R. Ribas, and P. S. Oliveira. 2010. The arboreal ant community visiting extrafloral nectaries in the Neotropical cerrado savanna. Terrestrial Arthropod Reviews 3: 3-27.
 * Silva T. S. R., and R. M. Feitosa. 2019. Using controlled vocabularies in anatomical terminology: A case study with Strumigenys (Hymenoptera: Formicidae). Arthropod Structure and Development 52: 1-26.
 * Silvestre R., C. R. F. Brandão, and R. R. Silva da 2003. Grupos funcionales de hormigas: el caso de los gremios del cerrado. Pp. 113-148 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp.
 * Siqueira de Castro F., A. B. Gontijo, W. Duarte da Rocha, and S. Pontes Ribeiro. 2011. As comunidades de formigas de serapilheira nas florestas semidecíduas do Parque Estadual do Rio Doce, Minas Gerais. MG.BIOTA, Belo Horizonte 3(5): 5-24.
 * Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
 * Soares N. S., L. de O. Almeida, C. A. Goncalves, M. T. Marcolino, and A. M. Bonetii. 2006. Survey of Ants (Hymenoptera: Formicidae) in the Urban Area of Uberlândia, MG, Brazil. Neotropical Entomology 35(3): 324-328.
 * Solomon, S.E. and A.S. Mikheyev. 2005. The ant (Hymenoptera: Formicidae) fauna of Cocos Island, Costa Rica. Florida Entomologist 88(4):415-423
 * Stitz H. 1933. Neue Ameisen des Hamburger Museums (Hym. Form.). Mitt. Dtsch. Entomol. Ges. 4: 67-75.
 * Suguituru S. S., M. Santina de Castro Morini, R. M. Feitosa, and R. Rosa da Silva. 2015. Formigas do Alto Tiete. Canal 6 Editora 458 pages
 * Tavares A. A., P. C. Bispo, and A. C. Zanzini. 2008. Effect of Collect Time on Communities of Epigaeic Ants (Hymenoptera: Formicidae) in Areas of Eucalyptus cloeziana and Cerrado. Neotropical Entomology 37(2): 126-130.
 * Torres, I. L., M. A. G. Martinez, H. O. Rivera, M. E. G. Tovar, H. E. L. Espinosa, and J. M. Gonzalez. 2018. Comparacion de dos muestreos de hormigas del suelo en la barranca de Metlac, Fortin de las Flores, Veracruz, Mexico. Revista Cientifica UDO Agricola 10: 173-178.
 * Ulyssea M. A., and C. R. F. Brandao. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217224.
 * Ulyssea M.A., C. E. Cereto, F. B. Rosumek, R. R. Silva, and B. C. Lopes. 2011. Updated list of ant species (Hymenoptera, Formicidae) recorded in Santa Catarina State, southern Brazil, with a discussion of research advances and priorities. Revista Brasileira de Entomologia 55(4): 603-611.
 * Ulysséa M. A., C. R. F. Brandão. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217-224.
 * Valenzuela-Gonzalez J., A. V. Parra-Cabanas, L. Quiroz-Robledo, D. L. Martinez-Tlapa, and E. D. Montes-de-Oca-Torres. 2013. Variación de la mirmecofauna en un gradiente altitudinal en la región central de Veracruz, México. In Formicidae de Mexico (eds. M. Vasquez-Bolanos, G. Castano-Meneses, A. Cisneros-Caballero, G. A. Quiroz-Rocha, and J. L. Navarrete-Heredia) p 75-82.
 * Varela-Hernandez, F., M. Rocha-Ortega, R. W. Jones, and W. P. Mackay. 2016. Insectos: Hormigas (Formicidae) del estado de Queretaro, Mexico. Pages 397-404 in W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
 * Varela-Hernandez, F., M. Rocha-Ortega, W. P. Mackay, and R. W. Jones. 2016. Lista preliminar de las hormigas (Hymenoptera: Formicidae) del estado de Queretaro, Mexico. Pages 429-435 in . W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
 * Vasconcelos, H.L., J.M.S. Vilhena, W.E. Magnusson and A.L.K.M. Albernaz. 2006. Long-term effects of forest fragmentation on Amazonian ant communities. Journal of Biogeography 33:1348-1356
 * Vasquez Bolanos M., and J. Escoto Rocha. 2018. Ants (Hymenoptera: Formicidae) of Aguascalientes. Investigacion y Ciencia 24(68): 36-40.
 * Vasquez-Bolanos M. 2011. Checklist of the ants (Hymenoptera: Formicidae) from Mexico. Dugesiana 18(1): 95-133.
 * Vergara-Torres C. A., M. Vasquez-Bolanos, A. M. Corona-Lopez, V. H. Toledo-Hernandez, and A. Flores-Palacios. 2016. Ant (Hymenoptera: Formicidae) Diversity in the Canopy of a Tropical Dry Forest in Tepoztl an, Central Mexico. Annals of the Entomological Society of America 2016: 1–7.
 * Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
 * Weber N. A. 1938. The food of the giant toad, Bufo marinus (L.), in Trinidad and British Guiana with special reference to the ants. Annals of the Entomological Society of America 31: 499-503.
 * Wetterer J. K., M. J. Liles, J. M. Sermeno, L. Serrano Cervantes, E. E. Echeverria, R. M. Estrada Hernandez, A. Henriquez, D. Perez, D. A. Sanchez Garcia, C. E. Gomez Peralta, R. Lopez Sorto, and G. Melendez. 2016. Predaceous fie ants (Hymenoptera: Formicidae) at sea turtle (Testudines: Cheloniidae) nestig beaches and hatcheries in El Salvador. Florida Entomologist 99(1): 106-109.
 * Wheeler W. M. 1905. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21: 79-135.
 * Wheeler W. M. 1907. A collection of ants from British Honduras. Bulletin of the American Museum of Natural History 23: 271-277.
 * Wheeler W. M. 1910. The North American ants of the genus Camponotus Mayr. Annals of the New York Academy of Sciences 20: 295-354.
 * Wheeler W. M. 1916. Ants collected in British Guiana by the expedition of the American Museum of Natural History during 1911. Bulletin of the American Museum of Natural History 35: 1-14.
 * Wheeler W. M. 1918. Ants collected in British Guiana by Mr. C. William Beebe. Journal of the New York Entomological Society 26: 23-28.
 * Wheeler W. M. 1919. The ants of Tobago Island. Psyche (Cambridge) 26: 113.
 * Wheeler W. M. 1922. The ants of Trinidad. American Museum Novitates 45: 1-16.
 * Wheeler W. M. 1942. Studies of Neotropical ant-plants and their ants. Bulletin of the Museum of Comparative Zoology 90: 1-262.
 * Wheeler W. M., and I. W. Bailey. 1920. The feeding habits of pseudomyrmine and other ants. Transactions of the American Philosophical Society (2)22: 235-279.
 * Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
 * Wheeler, G.C. and J. Wheeler. 1985. A checklist of Texas ants. Prairie Naturalist 17:49-64.
 * Wheeler, William Morton. 1911. Ants Collected in Grenada, W.I. by Mr. C. T. Brues. Bulletin of the Museum of Comparitive Zoology at Harvard College. 54(5):166-172.
 * Wheeler, William Morton. 1934. Ants From The Islands Off The West Coast Of Lower California and Mexico. The Pan-Pacific Entomologist. 10(3):131-144.
 * Wilson, E.O. 1987. The Arboreal Ant Fauna of Peruvian Amazon Forests: A First Assessment. Biotropica 19(3):245-251.
 * Yanoviak S. P., S, M. Berghoff, K. E. Linsenmair, and G. Zotz. 2011. Effects of an Epiphytic Orchid on Arboreal Ant Community Structure in Panama. Biotropica 43(6): 731-737.
 * Zolessi L. C. de; Y. P. de Abenante, and M. E. Philippi. 1989. Catálogo sistemático de las especies de Formícidos del Uruguay (Hymenoptera: Formicidae). Montevideo: ORCYT Unesco, 40 + ix pp.
 * de Zolessi, L.C., Y.P. de Abenante and M.E. Philippi. 1987. Lista sistemática de las especies de formícidos del Uruguay. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo 11(165):1-9
 * de Zolessi, L.C., Y.P. de Abenante and M.E. Phillipi. 1989. Catalago Systematico de las Especies de Formicidos del Uruguay (Hymenoptera: Formicidae). Oficina Regional de Ciencia y Technologia de la Unesco para America Latina y el Caribe- ORCYT. Montevideo, Uruguay