Checklist of Pheidole species

The following species and subspecies belong to the genus Pheidole. Synonyms are listed under their senior names.

A
aana Wilson & Taylor, 1967 (Samoa)

aberrans Mayr, 1868 (Argentina)

• Pheidole aberrans diversiceps Santschi, 1916

• Pheidole aberrans fartilia Forel, 1913

• Pheidole aberrans mutica Emery, 1906

absurda Forel, 1886 (Guatemala)

• Pheidole ridicula Wheeler, W.M., 1916

acamata Wilson, 2003 (Nicaragua)

acantha Eguchi, 2001 (Borneo)

accinota Wheeler, W.M., 1925 (Peru)

aciculata Wilson, 2003 (Peru)

aculifera Wilson, 2003 (Panama)

acutidens (Santschi, 1922) (Argentina)

ademonia Wilson, 2003 (Peru)

adrianoi Naves, 1985 (United States)

aeberlii Forel, 1894 (Senegal)

aeberlii erythraea Emery, 1901 (Ethiopia)

aenescens Wilson, 2003 (Brazil)

aequiseta Santschi, 1923 (Bolivia)

aglae Forel, 1913 (Indonesia (Java))

• Pheidole exasperata concordia Santschi, 1916

• Pheidole exasperata fusiformis Viehmeyer, 1914

• Pheidole exasperata polita Viehmeyer, 1914

agricola Wilson, 2003 (Mexico)

ajax Forel, 1899 (Costa Rica)

akermani Arnold, 1920 (South Africa)

alacris Santschi, 1923 (Argentina)

• Pheidole flavens mediorubra Santschi, 1933

alayoi Wilson, 2003 (Cuba)

albidula Santschi, 1928 (Democratic Republic of Congo)

albipes Wilson, 2003 (Mexico)

alexeter Wilson, 2003 (Brazil)

alfaroi Emery, 1896 (Costa Rica)

alienata Borgmeier, 1929 (Brazil)

allani Bingham, 1903 (Myanmar)

allarmata Wilson, 2003 (Brazil)

alpestris Wilson, 2003 (Ecuador)

alpinensis Forel, 1912 (Brazil)

alticola Wilson, 2003 (Colombia)

amabilis Wilson, 2003 (Costa Rica)

amata Forel, 1901 (Colombia)

amazonica Wilson, 2003 (Peru)

amber Donisthorpe, 1941 (New Guinea)

ambigua Wilson, 2003 (Brazil)

ambonensis Karavaiev, 1930 (Indonesia (Ambon Island))

ampla Forel, 1893 (Australia)

ampla mackayensis Forel, 1902 (Australia)

ampla parviceps Forel, 1915 (Australia)

ampla perthensis Crawley, 1922 (Australia)

amplificata Viehmeyer, 1914 (New Guinea)

anastasii Emery, 1896 (Costa Rica)

andrieui Santschi, 1930 (Sudan)

androsana Wheeler, W.M., 1905 (Bahamas)

angulicollis Eguchi, 2001 (Borneo)

angulifera Wilson, 2003 (Costa Rica)

angusta Forel, 1908 (Brazil)

• Pheidole angusta idulis Forel, 1912

angusticeps Wilson, 2003 (Costa Rica)

anima Wilson, 2003 (Honduras)

annemariae Forel, 1918 (Madagascar)

annexa Eguchi, 2001 (Borneo)

anthracina Forel, 1902 (Australia)

anthracina grandii Emery, 1914 (Australia)

anthracina orba Forel, 1902 (Australia)

antillana Forel, 1893 (Antilles)

aper Forel, 1912 (Brazil)

aphrasta Zhou & Zheng, 1999 (China)

arachnion Wilson, 2003 (Costa Rica)

• Pheidole iracunda Wilson, 2003

araneoides Wilson, 2003 (Brazil)

arboricola Wilson, 2003 (Costa Rica)

arcifera Santschi, 1925 (Brazil)

arctos Wilson, 2003 (Mexico)

areniphila Forel, 1910

• Pheidole arenicola Forel, 1910

areniphila aurora Santschi, 1928 (Tanzania)

argentina (Bruch, 1932) (Argentina)

arhuaca Forel, 1901 (Colombia)

• Pheidole arhuaca bimons Forel, 1912

ariel Wilson, 2003 (Mexico)

aripoensis Wilson, 2003 (Trinidad)

aristotelis Forel, 1911 (Borneo)

arnoldi Forel, 1913 (Zimbabwe)

arnoldi ballaensis Arnold, 1920 (Zimbabwe)

arnoldi rufescens Arnold, 1920 (Zimbabwe)

aspera Mayr, 1862 (South Africa)

asperata Emery, 1895 (Myanmar)

asperithorax Emery, 1894 (Bolivia)

aspidata Eguchi & Bui, 2005 (Vietnam)

astur Wilson, 2003 (Peru)

athertonensis Forel, 1915 (Australia)

athertonensis cedarensis Forel, 1915 (Australia)

athertonensis tambourinensis Forel, 1915 (Australia)

atticola Forel, 1912 (Argentina)

atua Wilson & Taylor, 1967 (Samoa)

aurea Wilson, 2003 (United States)

aurivillii Mayr, 1896 (Cameroun)

aurivillii attenuata Santschi, 1910 (Congo)

aurivillii kasaiensis Forel, 1911 (Democratic Republic of Congo)

aurivillii rubricalva Forel, 1915 (Democratic Republic of Congo)

auropilosa Mayr, 1887 (Brazil)

avia Forel, 1908 (Brazil)

azteca Wilson, 2003 (Mexico)

B
bahai (Forel, 1922) (United States)

bajaensis Wilson, 2003 (Mexico)

bakeri Forel, 1912 (Cuba)

balzani Emery, 1894 (Bolivia)

bambusarum Forel, 1908 (Brazil)

• Pheidole ogloblini Kusnezov, 1952

bandata Bharti, 2004 (India)

barbata Wheeler, W.M., 1908 (United States)

barreleti Forel, 1903 (Sri Lanka)

barumtaun Donisthorpe, 1938 (New Guinea)

batrachorum Wheeler, W.M., 1922 (Democratic Republic of Congo)

beauforti Emery, 1911 (New Guinea)

bellatrix Wilson, 2003 (Colombia)

belli Mann, 1919 (Solomon Is)

beloceps Wilson, 2003 (Costa Rica)

bequaerti Forel, 1913 (Democratic Republic of Congo)

bergi Mayr, 1887 (Argentina)

• Pheidole bergi pulliventris Santschi, 1929

• Pheidole bergi subparallela Emery, 1906

• Pheidole goetschi Santschi, 1939

bessonii Forel, 1891 (Madagascar)

bicarinata Mayr, 1870 (United States)

• Pheidole bicarinata vinelandica Forel, 1886

• Pheidole hayesi Smith, M.R., 1924

• Pheidole vinelandica buccalis Wheeler, W.M., 1908

• Pheidole vinelandica laeviuscula Emery, 1895

• Pheidole vinelandica longula Emery, 1895

biconstricta Mayr, 1870 (Colombia)

• Pheidole biconstricta bicolor Emery, 1890

• Pheidole biconstricta burtoni Mann, 1916

• Pheidole biconstricta hybrida Emery, 1894

• Pheidole biconstricta rubicunda Emery, 1890

• Pheidole biconstricta surda Forel, 1912

• Pheidole holmgreni Wheeler, W.M., 1925

• Pheidole holmgreni festata Wheeler, W.M., 1925

• Pheidole radoszkowskii lallemandi Forel, 1901

bicornis Forel, 1899 (Panama)

bidens Wilson, 2003 (Brazil)

bifurca Donisthorpe, 1941 (New Guinea)

bigote Longino, 2009 (Mexico)

bilimeki Mayr, 1870 (Mexico)

• Pheidole anastasii cellarum Forel, 1908

• Pheidole anastasii johnsoni Wheeler, W.M., 1907

• Pheidole anastasii venezuelana Forel, 1905

• Pheidole floridana antoniensis Forel, 1901

• Pheidole floridana ares Forel, 1908

• Pheidole floridana deplanata Pergande, 1896

• Pheidole punctatissima annectens Wheeler, W.M., 1905

• Pheidole punctatissima insulana Wheeler, W.M., 1905

• Pheidole rectiluma Wilson, 2003

biloba (Karavaiev, 1935) (Cambodia)

binasifera Wilson, 2003 (Colombia)

binghamii Forel, 1902 (India)

biolleyi Forel, 1908 (Costa Rica)

• Pheidole biolleyi tristani Forel, 1908

bison Wilson, 2003 (Bolivia)

blumenauensis Kempf, 1964 (Brazil)

• Pheidole diligens Santschi Santschi, 1923

bluntschlii Forel, 1911 (Indonesia (Sumatra))

boliviana Wilson, 2003 (Bolivia)

• Pheidole mincana Wilson, 2003

• Pheidole scitula Wilson, 2003

boltoni Wilson, 2003 (Costa Rica)

• Pheidole humida Wilson, 2003

borgmeieri Kempf, 1972 (Brazil)

boruca Wilson, 2003 (Costa Rica)

bos Forel, 1893 (Australia)

bos baucis Forel, 1910 (Australia)

bos eubos Forel, 1915 (Australia)

brachyops Wilson, 2003 (Costa Rica)

brandaoi Wilson, 2003 (Colombia)

branstetteri Longino, 2009 (Mexico)

braueri Forel, 1897 (Seychelles Is)

brevicona Mayr, 1887 (Brazil)

• Pheidole kraepelini Forel, 1901

brevicornis Mayr, 1876 (Australia)

breviseta Santschi, 1919 (Argentina)

browni Wilson, 2003 (Costa Rica)

bruchella Forel, 1915 (Argentina)

bruchi Forel, 1914 (Argentina)

bruesi Wheeler, W.M., 1911 (Grenada Island)

brunnescens Santschi, 1929 (Brazil)

bucculenta Forel, 1908 (Brazil)

buchholzi Mayr, 1901 (Cameroun)

buckleyi Smith, M.R., 1951

• Atta pennsylvanica Buckley, 1867

bufo Wilson, 2003 (Brazil)

bula Sarnat, 2008 (Fiji Is)

bulliceps Wilson, 2003 (Ecuador)

bureni Wilson, 2003 (United States)

butteli Forel, 1913 (Indonesia (Sumatra))

C
caffra Emery, 1895 (South Africa)

caffra abyssinica Forel, 1910 (Ethiopia)

caffra amoena Forel, 1911 (Ethiopia)

caffra bayeri Forel, 1916 (Democratic Republic of Congo)

caffra montivaga Santschi, 1939 (Zimbabwe)

caffra senilifrons Wheeler, W.M., 1922 (Democratic Republic of Congo)

cairnsiana Forel, 1902 (Australia)

caldwelli Mann, 1921 (Fiji Is)

calens Forel, 1901 (Mexico)

californica Mayr, 1870 (United States)

• Pheidole californica hagermani Cole, 1936

• Pheidole californica incenata Wheeler, W.M., 1915

• Pheidole californica nevadensis Wheeler, W.M., 1915

• Pheidole californica pyramidensis Emery, 1922

• Pheidole californica satura Wheeler, W.M., 1915

• Pheidole oregonica Emery, 1895

calimana Wilson, 2003 (Colombia)

caltrop Wilson, 2003 (Panama)

cameroni Mayr, 1887 (Ecuador)

• Pheidole monticola Cameron, 1891

camilla Wilson, 2003 (Ecuador)

camptostela Kempf, 1972 (Brazil)

capellinii Emery, 1887 (Indonesia)

• Pheidole attila Forel, 1913

• Pheidole lighti Wheeler, W.M., 1927

capensis Mayr, 1862 (South Africa)

capensis dregei Emery, 1895 (South Africa)

capensis modestior Santschi, 1919 (South Africa)

capensis reddenburgensis Forel, 1913 (South Africa)

capillata Emery, 1906 (Brazil)

caracalla Wilson, 2003 (Peru)

carapuna Mann, 1916 (Brazil)

• Pheidole carapuna chaquimayensis Wheeler, W.M., 1925

carapunco Kusnezov, 1952 (Argentina)

cardiella Wilson, 2003 (Peru)

cardinalis Wilson, 2003 (Brazil)

caribbaea Wheeler, W.M., 1911 (Jamaica)

• Pheidole caribbaea sloanei Wheeler, W.M., 1917

carinata Wilson, 2003 (Costa Rica)

cariniceps Eguchi, 2001 (Borneo)

carinote Longino, 2009 (Costa Rica)

carrolli Naves, 1985 (United States)

casta Wheeler, W.M., 1908 (United States)

castanea (Smith, F., 1858) (Brazil)

cataphracta Wilson, 2003 (Colombia)

cataractae Wheeler, W.M., 1916 (Guyana)

caulicola Wilson, 2003 (Costa Rica)

cavifrons Emery, 1906 (Uruguay)

• Pheidole arciruga Forel, 1908

• Pheidole cavifrons fuscipunctis Santschi, 1916

cavigenis Wheeler, W.M., 1915 (United States)

ceibana Wilson, 2003 (Honduras)

celaena Wilson, 2003 (Costa Rica)

centeotl Wheeler, W.M., 1914 (Mexico)

cerebrosior Wheeler, W.M., 1915 (United States)

ceres Wheeler, W.M., 1904 (United States)

• Pheidole ceres tepaneca Wheeler, W.M., 1914

cerina Wilson, 2003 (Costa Rica)

cervicornis Emery, 1900 (New Guinea)

ceylonica (Motschoulsky, 1863) (Sri Lanka)

chalca Wheeler, W.M., 1914 (Mexico)

chalcoides Wilson, 2003 (Mexico)

charazana Wilson, 2003 (Colombia)

cheesmanae Donisthorpe, 1941 (Indonesia)

chilensis Mayr, 1862 (Chile)

chloe Forel, 1908 (Costa Rica)

chocoensis Wilson, 2003 (Colombia)

christinae Fischer, Hita Garcia & Peters, 2012 (Uganda)

christopherseni Forel, 1912 (Panama)

chrysops Wilson, 2003 (Brazil)

cielana Wilson, 2003 (Mexico)

cingulata (Smith, F., 1857) (Borneo)

citrina Wilson, 2003 (Costa Rica)

clavata (Emery, 1877) (Ethiopia)

claviscapa Santschi, 1925 (Brazil)

clementensis Gregg, 1969 (United States)

clydei Gregg, 1950 (United States)

clypeocornis Eguchi, 2001 (Borneo)

cocciphaga Borgmeier, 1934 (Suriname)

cockerelli Wheeler, W.M., 1908 (United States)

coffeicola Borgmeier, 1934 (Suriname)

colaensis Mann, 1921 (Fiji Is)

colobopsis Mann, 1916 (Brazil)

colpigaleata Eguchi, 2006 (Vietnam)

comata Smith, F., 1858 (Borneo)

• Pheidole multicoma Eguchi, 1999

concentrica Forel, 1902 (Australia)

concentrica recurva Forel, 1910 (Australia)

concinna Santschi, 1910 (Congo)

conficta Forel, 1902 (Australia)

confoedusta Wheeler, W.M., 1909 (Mexico)

constanciae Forel, 1902 (India)

constanciae nigra Forel, 1902 (India)

constipata Wheeler, W.M., 1908 (United States)

coonoorensis Forel, 1902 (India)

coracina Wilson, 2003 (Mexico)

cordiceps Mayr, 1868 (Argentina)

• Pheidole cordiceps lilianae Forel, 1912

cornicula Wilson, 2003 (Bolivia)

corticicola Santschi, 1910 (Congo)

coveri Wilson, 2003 (Trinidad)

cramptoni Wheeler, W.M., 1916 (Guyana)

• Pheidole cramptoni petiolicola Wheeler, W.M., 1921

crassicornis Emery, 1895 (United States)

crassinoda Emery, 1895 (South Africa)

crassinoda pluto Arnold, 1920 (Mozambique)

crassinoda ruspolii Emery, 1897 (Ethiopia)

crassinoda sordidula Santschi, 1937 (Tanzania)

creightoni Gregg, 1955 (United States)

crinita Wilson, 2003 (Ecuador)

crozieri Wilson, 2003 (Peru)

cryptocera Emery, 1900 (New Guinea)

cubaensis Mayr, 1862 (Cuba)

• Pheidole cubaensis grayi Mann, 1920

cuevasi Wilson, 2003 (Peru)

cuitensis Forel, 1910 (Angola)

cuprina Wilson, 2003 (Peru)

cursor Wilson, 2003 (Peru)

cyrtostela Wilson, 2003 (Brazil)

D
dammermani Wheeler, W.M., 1924 (Indonesia (Sebesi Island))

daphne Wilson, 2003 (Costa Rica)

darlingtoni Wheeler, W.M., 1936 (Haiti)

darwini Fischer, Hita Garcia & Peters, 2012 (Gabon)

dasypyx Wilson, 2003 (Panama)

davidsonae Wilson, 2003 (Peru)

davisi Wheeler, W.M., 1905 (United States)

dea Santschi, 1921 (Democratic Republic of Congo)

debilis Longino, 2009 (Costa Rica)

decarinata Santschi, 1929 (Sudan)

deceptrix Forel, 1899 (Guatemala)

• Pheidole chiapasana Wilson, 2003

• Pheidole variceps Wilson, 2003

defecta Santschi, 1923 (Guatemala)

deima Wilson, 2003 (Brazil)

delecta Forel, 1899 (Mexico)

delicata Wilson, 2003 (Bolivia)

deltea Eguchi, 2001 (Borneo)

demeter Wilson, 2003 (Peru)

dentata Mayr, 1886 (United States)

• Leptothorax tennesseensis Cole, 1938

• Pheidole commutata Mayr, 1886

• Pheidole dentata faisonsica Forel, 1901

dentigula Smith, M.R., 1927 (United States)

descolei Kusnezov, 1952 (Argentina)

deserticola Forel, 1910 (Australia)

deserticola foveifrons Viehmeyer, 1924 (Australia)

desertorum Wheeler, W.M., 1906 (United States)

• Pheidole desertorum comanche Wheeler, W.M., 1906

• Pheidole desertorum maricopa Wheeler, W.M., 1906

diabolus Wilson, 2003 (Colombia)

diana Forel, 1908 (Costa Rica)

diffidens (Walker, 1859) (Sri Lanka)

diffusa (Jerdon, 1851) (India)

diligens (Smith, F., 1858) (Brazil)

dinophila Wilson, 2003 (Brazil)

dione Forel, 1913 (Argentina)

distincta Donisthorpe, 1943 (New Guinea)

distorta Forel, 1899 (Colombia)

diversipilosa Wheeler, W.M., 1908 (United States)

dolon Wilson, 2003 (Peru)

dorsata Wilson, 2003 (Colombia)

dossena Wilson, 2003 (Costa Rica)

drepanon Wilson, 2003 (Dominican Republic)

dryas Wilson, 2003 (Costa Rica)

dugasi Forel, 1911 (Vietnam)

dumicola Wilson, 2003 (Mexico)

duneraensis Bharti, 2001 (India)

durionei Santschi, 1923 (Argentina)

dwyeri Gregg, 1969 (Mexico)

dyctiota Kempf, 1972 (Brazil)

E
ectatommoides Wilson, 2003 (Costa Rica)

ecuadorana Wilson, 2003 (Ecuador)

eidmanni Menozzi, 1926 (Brazil)

elecebra (Wheeler, W.M., 1904) (United States)

elegans Donisthorpe, 1938 (New Guinea)

elisae Emery, 1900 (Indonesia (Engano Island))

• Pheidole elisae nenia Forel, 1913

elongicephala Eguchi, 2008 (Vietnam)

embolopyx Brown, 1968 (Brazil)

emmae Forel, 1905 (Indonesia (Java))

ensifera Forel, 1897 (Madagascar)

eowilsoni Longino, 2009 (Costa Rica)

eparmata Wilson, 2003 (Venezuela)

epetrion Wilson, 2003 (Ecuador)

epiphyta Longino, 2009 (Costa Rica)

erato Mann, 1919 (Solomon Islands)

erethizon Wilson, 2003 (Mexico)

eriophora Wilson, 2003 (Colombia)

ernsti Forel, 1912 (Taiwan)

erratilis Wilson, 2003 (Costa Rica)

• Pheidole petersoni Wilson, 2003

escherichii Forel, 1910 (Ethiopia)

euryscopa Wilson, 2003 (Colombia)

exarata Emery, 1896 (Costa Rica)

• Pheidole crassipes grantae Forel, 1908

exasperata (Mayr, 1866) ("ohne Vaterlandsangabe")

excellens Mayr, 1862 (Ghana)

excellens fulvobasalis Santschi, 1921 (Benin)

excellens weissi Santschi, 1910 (Congo)

excubitor Wilson, 2003 (Costa Rica)

exigua Mayr, 1884 (French Guiana)

exquisita Wilson, 2003 (Ecuador)

F
fabricator (Smith, F., 1858) (Brazil)

• Atta nigriventris Smith, F., 1858

fadli Sharaf, 2007 (Egypt)

fallax Mayr, 1870 (Cuba)

• Pheidole fallax britoi Forel, 1912

• Pheidole fallax columbica Forel, 1886

• Pheidole fallax ovalis Forel, 1912

• Pheidole fallax rubens Forel, 1899

• Pheidole jelskii fallacior Forel, 1901

fantasia Chapman, 1963 (Philippines)

fatigata Bolton, 1995

• Pheidole granulatus Donisthorpe, 1949

feae Emery, 1895 (Myanmar)

fera Santschi, 1925 (Brazil)

fergusoni Forel, 1902 (India)

fervens Smith, F., 1858 (Singapore)

• Pheidole amia Forel, 1912

• Pheidole cavannae Emery, 1887

• Pheidole javana Mayr, 1867

• Pheidole javana desucta Wheeler, W.M., 1929

• Pheidole javana dharmsalana Forel, 1902

• Pheidole javana dolenda Forel, 1912

• Pheidole javana soror Santschi, 1937

• Pheidole nodus azumai Santschi, 1941

• Pheidole oceanica nigriscapa Santschi, 1928

• Solenopsis pungens Smith, F., 1861

fervens pectinata Stitz, 1912 (Indonesia (Seram Island))

fervida Smith, F., 1874 (Japan)

fervida kwazana Teranishi, 1940 (Korea)

fimbriata Roger, 1863 ("Rio Paraguay")

• Pheidole diversa Smith, F., 1860

• Pheidole fimbriata tucumana Forel, 1913

• Pheidole smithii Dalla Torre, 1892

fiorii Emery, 1890 (Costa Rica)

fissiceps Wilson, 2003 (Peru)

flavens Roger, 1863 (Cuba)

• Pheidole exigua tuberculata Mayr, 1887

• Pheidole flavens gracilior Forel, 1901

• Pheidole flavens haytiana Forel, 1907

• Pheidole flavens navigans Forel, 1901

• Pheidole flavens spei Santschi, 1930

• Pheidole flavens vincentensis Forel, 1893

• Pheidole floridana aechmeae Wheeler, W.M., 1934

• Pheidole greggi Naves, 1985

flavens farquharensis Forel, 1907 (Madagascar)

flavens thomensis Emery, 1894 (Antilles Is)

flaveria Zhou & Zheng, 1999 (China)

flavida Mayr, 1887 (Brazil)

flavifrons Wilson, 2003 (Peru)

flavothoracica Viehmeyer, 1914 (New Guinea)

floricola Wilson, 2003 (Honduras)

floridana Emery, 1895 (United States)

• Pheidole lauta Wheeler, W.M., 1908

foederalis Borgmeier, 1928 (Brazil)

foreli Mayr, 1901 (South Africa)

foreli pubens Forel, 1910 (South Africa)

fortis Eguchi, 2006 (Vietnam)

fossimandibula Longino, 2009 (Costa Rica)

foveolata Eguchi, 2006 (Vietnam)

fowleri Wilson, 2003 (Brazil)

fracticeps Wilson, 2003 (Trinidad)

fullerae Wilson, 2003 (Ecuador)

funki LaPolla, 2005 (Guyana)

funkikoensis Wheeler, W.M., 1929 (Taiwan)

furcata Sarnat, 2008 (Fiji Is)

furtiva Wilson, 2003 (United States)

fuscula Emery, 1900 (New Guinea)

G
gagates Wilson, 2003 (Peru)

gaigei Forel, 1914 (Colombia)

galba Wilson, 2003 (Peru)

gambogia Donisthorpe, 1948 (New Guinea)

gangamon Wilson, 2003 (Mexico)

gatesi (Wheeler, W.M., 1927) (Myanmar)

gauthieri Forel, 1901 (Colombia)

• Pheidole gauthieri oxymora Forel, 1912

gellibrandi Clark, 1934 (Australia)

geminata Wilson, 2003 (Ecuador)

gemmula Wilson, 2003

geraesensis Santschi, 1929 (Brazil)

germaini Emery, 1896 (Brazil)

gertrudae Forel, 1886 (Brazil)

• Megalomyrmex humilis Borgmeier, 1930

• Pheidole gertrudae leonhardi Forel, 1901

• Pheidole gertrudae loretensis Santschi, 1933

ghatica Forel, 1902 (India)

ghigii Emery, 1900 (Indonesia (Sumatra))

gibba Mayr, 1887 (Brazil)

• Pheidole gibba lopesi Borgmeier, 1937

gibbata Borgmeier, 1934 (Surinam)

gigaflavens Wilson, 2003 (Argentina)

gigas Wilson, 2003 (Brazil)

gigliolii Menozzi, 1935 (Guyana)

gilva Wilson, 2003 (Peru)

gilvescens Creighton & Gregg, 1955 (United States)

glabrella Fischer, Hita Garcia & Peters, 2012 (Cameroon)

globularia Wilson, 2003 (Ecuador)

glomericeps Wilson, 2003 (Mexico)

gnomus Wilson, 2003 (Ecuador)

godmani Forel, 1893 (Antilles)

goeldii Forel, 1895 (Brazil)

gombakensis Eguchi, 2001 (West Malaysia)

gouldi Forel, 1886 (Guatemala)

gracilescens (Smith, F., 1860) (Indonesia (Sulawesi))

gracilipes (Motschoulsky, 1863) (Sri Lanka)

gradifer Wilson, 2003 (Costa Rica)

grallatrix Emery, 1899 (Madagascar)

grandinodus Wilson, 2003 (Brazil)

granulata Pergande, 1896 (Mexico)

gravida Wilson, 2003 (Venezuela)

grayi Forel, 1902 (India)

grex Wilson, 2003 (Colombia)

grundmanni Smith, M.R., 1953 (United States)

guajirana Wilson, 2003 (Colombia)

guayasana Wilson, 2003 (Ecuador)

guerrerana Wilson, 2003 (Mexico)

guilelmimuelleri Forel, 1886 (Brazil)

• Pheidole guilelmimuelleri gustavi Forel, 1911

guineensis (Fabricius, 1793) (Guinea)

gulo Wilson, 2003 (Nicaragua)

gymnoceras Longino, 2009 (Costa Rica)

H
hainanensis Chen, Ye, Lu & Zhou, 2011 (China)

hamtoni Wilson, 2003 (Mexico)

harlequina Wilson, 2003 (Dominican Republic)

harrisonfordi Wilson, 2003 (Honduras)

• Pheidole prolixa Wilson, 2003

• Pheidole ruida Wilson, 2003

• Pheidole tenebra Wilson, 2003

hartmeyeri Forel, 1907 (Australia)

haskinsorum Wilson, 2003 (Bolivia)

hasticeps Wilson, 2003 (Costa Rica)

havilandi Forel, 1911 (Borneo)

• Pheidole havilandi sapuana Forel, 1911

• Pheidole havilandi selangorensis Forel, 1913

haywardi Kusnezov, 1952 (Argentina)

hazenae Wilson, 2003 (Ecuador)

hecate Wheeler, W.M., 1911 (Jamaica)

• Pheidole hecate bruesi Wheeler, W.M., 1917

• Pheidole hecate malevola Wheeler, W.M., 1911

hector Wilson, 2003 (Costa Rica)

hedlundorum Wilson, 2003 (Venezuela)

heliosa Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

hercules Donisthorpe, 1941 (New Guinea)

heterothrix Santschi, 1923 (Argentina)

• Pheidole gavrilovi Kusnezov, 1952

hetschkoi Emery, 1896 (Brazil)

• Pheidole hetschkoi frivola Santschi, 1923

hewitti Santschi, 1932 (South Africa)

• Pheidole capensis hewitti Arnold, 1944

heyeri Forel, 1899 (Brazil)

• Pheidole guilelmimuelleri ultrix Forel, 1912

hierax Wilson, 2003 (Panama)

hirsuta Emery, 1896 (Costa Rica)

hirtula Forel, 1899 (Mexico)

• Pheidole vasliti acolhua Wheeler, W.M., 1914

hispaniolae Wilson, 2003 (Dominican Republic)

hizemops Wilson, 2003 (Costa Rica)

hoelldobleri Wilson, 2003 (Costa Rica)

hongkongensis Wheeler, W.M., 1928 (China (Hong Kong))

hoplitica Wilson, 2003 (United States)

horni Emery, 1901 (Sri Lanka)

horribilis Wilson, 2003 (Ecuador)

hortensis Forel, 1913 (Indonesia (Java))

hortonae Wilson, 2003 (Venezuela)

hospes Smith, F., 1865 (New Guinea)

hospita Bingham, 1903 (India)

huacana Wilson, 2003 (Peru)

huberi Forel, 1911 (Indonesia (Sumatra))

• Pheidole huberi perakensis Forel, 1911

huilana Wilson, 2003 (Colombia)

humeralis Wheeler, W.M., 1908 (United States)

humeridens Wilson, 2003 (Argentina)

hyatti Emery, 1895 (United States)

• Pheidole hyatti ecitonodora Wheeler, W.M., 1908

• Pheidole hyatti solitanea Wheeler, W.M., 1915

• Pheidole vaslitii Pergande, 1896

I
impressa Mayr, 1870 (Brazil)

impressa detrita Emery, 1890 (Venezuela)

impressiceps Mayr, 1876 (Australia)

impressiceps commista Forel, 1901 (New Guinea (Bismarck Archipelago))

inca Wilson, 2003 (Peru)

incerta (Smith, F., 1863) (Indonesia (Misool Island))

incisa Mayr, 1870 (Colombia)

incurvata Viehmeyer, 1924 (Australia)

indagatrix Wilson, 2003 (Costa Rica)

indica Mayr, 1879 (India)

• Pheidole indica himalayana Forel, 1902

• Pheidole indica rotschana Forel, 1902

• Pheidole striativentris Mayr, 1879

indosinensis Wheeler, W.M., 1928 (China)

industa Santschi, 1939 (Uruguay)

inermis Mayr, 1870 (Mexico)

infernalis Wilson, 2003 (Bolivia)

inflexa Santschi, 1923 (Ecuador)

innotata Mayr, 1866 ("ohne Vaterlandsangabe")

innupta Menozzi, 1931 (Costa Rica)

inornata Eguchi, 2001 (Borneo)

inquilina (Wheeler, W.M., 1903) (United States)

inscrobiculata Viehmeyer, 1916 (West Malaysia)

insipida Forel, 1899 (Mexico)

inversa Forel, 1901 (Colombia)

irritans (Smith, F., 1858) (South Africa)

isis Mann, 1919 (Solomon Islands)

isis taki Mann, 1919 (Solomon Islands)

J
jacobsoni Forel, 1911 (Indonesia (Java))

• Pheidole orophila Eguchi, 2001

jaculifera Wilson, 2003 (Brazil)

jamaicensis Wheeler, W.M., 1908 (Jamaica)

janzeni Longino, 2009 (Costa Rica)

jeannei Wilson, 2003 (Brazil)

jelskii Mayr, 1884 (French Guiana)

• Pheidole fallax emiliae Forel, 1901

• Pheidole jelskii antillensis Forel, 1901

• Pheidole jelskii arenicola Emery, 1894

jivaro Wilson, 2003 (Ecuador)

jonas Forel, 1907 (Comoro Is)

jordanica Saulcy, 1874 (Israel)

• Pheidole megacephala obtusa Stitz, 1917

• Pheidole schmitzi Forel, 1911

• Pheidole sinaitica laticeps Mayr, 1904

jubilans Forel, 1911 (Indonesia (Java))

jucunda Forel, 1885 (India)

jucunda fossulata Forel, 1902 (India)

jujuyensis Forel, 1913 (Argentina)

juniperae Wilson, 2003 (United States)

K
karolmorae Longino, 2009 (Costa Rica)

karolsetosa Longino, 2009 (Costa Rica)

katonae Forel, 1907 (Tanzania)

kikutai Eguchi, 2001 (Borneo)

kitschneri Forel, 1910 (South Africa)

knowlesi Mann, 1921 (Fiji Is)

• Pheidole knowlesi extensa Mann, 1921

kochi (Emery, 1911) (New Guinea)

kohli Mayr, 1901 (Kenya)

kugleri Wilson, 2003 (Peru)

kukrana Wilson, 2003 (Nicaragua)

kuna Wilson, 2003 (Panama)

kusnezovi Wilson, 2003 (Argentina)

• Eriopheidole symbiotica Kusnezov, 1952

L
laelaps Wilson, 2003 (Colombia)

laevicolor Eguchi, 2006 (Vietnam)

laevifrons Mayr, 1887 (Brazil)

laevinota Forel, 1908 (Brazil)

laevithorax Eguchi, 2008 (Vietnam)

laeviventris Mayr, 1870 (Colombia)

laevivertex Forel, 1901 (Mexico)

lagunculinoda Longino, 2009 (Mexico)

laidlowi Mann, 1916 (Brazil)

lamellinoda Forel, 1902 (India)

lamia Wheeler, W.M., 1901 (United States)

laminata Emery, 1900 (New Guinea)

lancifera Wilson, 2003 (Brazil)

lanigera Wilson, 2003 (Peru)

lanuginosa Wilson, 1984 (India)

laselva Wilson, 2003 (Costa Rica)

• Pheidole ebenina Wilson, 2003

laticornis Wilson, 2003 (Costa Rica)

laticrista Santschi, 1916 (Argentina)

latinoda Roger, 1863 (Sri Lanka)

latinoda angustior Forel, 1902 (Sri Lanka)

latinoda major Forel, 1885 (India)

latinoda peradeniyae Forel, 1911 (Sri Lanka)

lattkei Wilson, 2003 (Venezuela)

laudatana Wilson, 2003 (Dominica)

lemnisca Wilson, 2003 (Colombia)

lemur Forel, 1912 (Brazil)

leoncortesi Longino, 2009 (Mexico)

leonina Wilson, 2003 (Peru)

leptina Wilson, 2003 (Peru)

liengmei Forel, 1894 (Mozambique)

liengmei malindana Forel, 1913 (Zimbabwe)

liengmei micrartifex Forel, 1913 (Zimbabwe)

liengmei shinsendensis Forel, 1913 (Democratic Republic of Congo)

lignicola Mayr, 1887 (Brazil)

• Pheidole lignicola levocciput Forel, 1908

lilloi (Kusnezov, 1952) (Argentina)

liteae Forel, 1910 (Australia)

littoralis Cole, 1952 (United States)

lobulata Emery, 1900 (New Guinea)

lokitae Forel, 1913 (Indonesia (Sumatra))

longiceps Mayr, 1876 (Australia)

longiceps doddi Forel, 1910 (Australia)

longiceps frontalis Forel, 1902 (Australia)

longicornis Emery, 1888 (Indonesia (Nias Island))

longinoi Wilson, 2003 (Costa Rica)

longior Santschi, 1933 (Argentina)

longipes (Latreille, 1802) (French Guiana)

longiscapa Forel, 1901 (Venezuela)

• Pheidole longiscapa martensis Forel, 1914

longiseta Wilson, 2003 (Peru)

longispinosa Forel, 1891 (Madagascar)

longispinosa scabrata Forel, 1895 (Madagascar)

lourothi Wilson, 2003 (Costa Rica)

lovejoyi Wilson, 2003 (Brazil)

lucaris Wilson, 2003 (Brazil)

lucida Forel, 1895 (Madagascar)

lucioccipitalis Eguchi, 2001 (Borneo)

lucretii Santschi, 1923 (Brazil)

lupus Wilson, 2003 (Peru)

lustrata Wilson, 2003 (Costa Rica)

luteipes Emery, 1914 (New Caledonia)

luteipes obvia Emery, 1914 (New Caledonia)

lutzi Forel, 1905 (Brazil)

• Pheidole lutzi heinzi Forel, 1913

M
macclendoni Wheeler, W.M., 1908 (United States)

machetula Wilson, 2003 (Peru)

mackayi Wilson, 2003 (Mexico)

macracantha Wilson, 2003 (Panama)

macromischoides Wilson, 2003 (Cuba)

macrops Wilson, 2003 (Mexico)

maculifrons Wheeler, W.M., 1929 (Philippines)

madecassa Forel, 1892 (Madagascar)

madrensis Wilson, 2003 (Mexico)

magna Eguchi, 2006 (Vietnam)

magrettii Emery, 1887 (Indonesia (Java))

• Pheidole nodgii maxwellensis Forel, 1913

maja Forel, 1886 (Guatemala)

makilingi Viehmeyer, 1916 (Philippines)

malabarica (Jerdon, 1851) (India)

malinsii Forel, 1902 (Sri Lanka)

mallota Wilson, 2003 (Panama)

mamore Mann, 1916 (Brazil)

manteroi Emery, 1897 (New Guinea)

mantilla Wilson, 2003 (Costa Rica)

manuana Wilson, 2003 (Peru)

manukana Eguchi, 2001 (Borneo)

marcidula Wheeler, W.M., 1908 (United States)

maufei Arnold, 1920 (Zimbabwe)

mayri Forel, 1894 (Senegal)

medioflava Donisthorpe, 1941 (New Guinea)

megacephala (Fabricius, 1793) (Mauritius)

• Atta testacea Smith, F., 1858

• Formica edax Forskål, 1775

• Myrmica agilis Smith, F., 1857

• Myrmica suspiciosa Smith, F., 1859

• Myrmica trinodis Losana, 1834

• Oecophthora perniciosa Gerstäcker, 1859

• Oecophthora pusilla Heer, 1852

• Pheidole janus Smith, F., 1858

megacephala costauriensis Santschi, 1914 (Ghana)

megacephala duplex Santschi, 1937 (Angola)

megacephala ilgi Forel, 1907 (Ethiopia)

megacephala impressifrons Wasmann, 1905

• Pheidole megacephala impressiceps Wasmann, 1904

megacephala melancholica Santschi, 1912 (Ivory Coast)

megacephala nkomoana Forel, 1916 (Democratic Republic of Congo)

megacephala rotundata Forel, 1894 (Mozambique)

megacephala scabrior Forel, 1891 (Madagascar)

megacephala speculifrons Stitz, 1911 (Tanzania)

megacephala talpa Gerstäcker, 1871 (Kenya)

meihuashanensis Li & Chen, 1992 (China)

meinerti Forel, 1905 (Venezuela)

• Pheidole mimula Wheeler, W.M., 1916

meinertopsis Wilson, 2003 (Peru)

melanogaster Donisthorpe, 1943 (New Guinea)

melastomae Wilson, 2003 (Panama)

mendanai Mann, 1919 (Solomon Islands)

mendicula Wheeler, W.M., 1925 (Panama)

mentita Santschi, 1914 (Guinea)

mentita pullata Santschi, 1914 (Guinea)

mera Wilson, 2003 (United States)

merimbun Eguchi, 2001 (Brunei)

mesomontana Longino, 2009 (Costa Rica)

metallescens Emery, 1895 (United States)

• Pheidole metallescens splendidula Wheeler, W.M., 1908

metana Wilson, 2003 (Colombia)

micon Wilson, 2003 (Colombia)

micridris Wilson, 2003 (Peru)

microgyna Wheeler, W.M., 1928 (Guyana)

microps Wilson, 2003 (Peru)

micula Wheeler, W.M., 1915 (United States)

midas Wilson, 2003 (Trinidad)

militicida Wheeler, W.M., 1915 (United States)

minax Wilson, 2003 (Peru)

minensis Santschi, 1923 (Brazil)

minima Mayr, 1901 (Cameroun)

minima catella Santschi, 1914 (Nigeria)

minima faurei Santschi, 1920 (Gabon)

minima malelana Wheeler, W.M., 1922 (Democratic Republic of Congo)

minor (Jerdon, 1851) (India)

minuscula Bernard, 1953 (Guinea)

minutula Mayr, 1878 (Brazil)

• Pheidole minutula folicola Forel, 1904

• Pheidole minutula personata Santschi, 1929

mirabilis Wilson, 2003 (Colombia)

miseranda Wheeler, W.M., 1924 (Indonesia (Krakatau Island))

mittermeieri Wilson, 2003 (Bolivia)

mixteca Wilson, 2003 (Mexico)

mjobergi Forel, 1915 (Australia)

modiglianii Emery, 1900 (Indonesia (Mentawei Island))

moerens Wheeler, W.M., 1908 (Puerto Rico)

• Pheidole moerens creola Wheeler, W.M. & Mann, 1914

• Pheidole moerens dominicensis Wheeler, W.M., 1913

moffetti Wilson, 2003 (French Guiana)

monstrosa Wilson, 2003 (Brazil)

montana Eguchi, 1999 (Borneo)

monteverdensis Wilson, 2003 (Costa Rica)

mooreorum Wilson, 2003 (Mexico)

• Pheidole fariasana Wilson, 2003

morelosana Wilson, 2003 (Mexico)

morrisii Forel, 1886 (United States)

• Pheidole morrisi impexa Wheeler, W.M., 1908

• Pheidole morrisii vanceae Forel, 1901

moseni Wheeler, W.M., 1925 (Brazil)

mosenopsis Wilson, 2003 (Bolivia)

multidens Forel, 1902 (India)

multispina Wilson, 2003 (Panama)

mus Forel, 1902 (India)

mutisi Fernández & Wilson, 2008 (Colombia)

mylognatha Wheeler, W.M., 1922 (Democratic Republic of Congo)

N
nana Emery, 1894 (Brazil)

naoroji Forel, 1902 (India)

napoensis Wilson, 2003 (Ecuador)

nasifera Wilson, 2003 (Peru)

nasutoides Hölldobler & Wilson, 1992 (Costa Rica)

naylae Wilson, 2003 (Cuba)

nebulosa Wilson, 2003 (Costa Rica)

• Pheidole scabriventris Wilson, 2003

nemoralis Forel, 1892 (Madagascar)

nemoralis petax Forel, 1895 (Madagascar)

neokohli Wilson, 1984

• Anergatides kohli Wasmann, 1915

neolongiceps Brown, 1950 (Cuba)

• Pheidole longiceps Aguayo, 1932

neolongiscapa Özdikmen, 2010

• Pheidole longiscapa Zhou & Zheng, 1999

neoschultzi LaPolla, 2006

• Pheidole schultzi LaPolla, 2005

nesiota Wilson, 2003 (Brazil)

nietneri Emery, 1901 (Sri Lanka)

nigella Emery, 1894 (Bolivia)

• Pheidole dimidiata Emery, 1894

nigeriensis Santschi, 1914 (Nigeria)

nigricula Wilson, 2003 (Costa Rica)

nigritella Bernard, 1953 (Guinea)

nimba Bernard, 1953 (Guinea)

nindi Mann, 1919 (Solomon Islands)

nitella Wilson, 2003 (Costa Rica)

nitidicollis Emery, 1896 (Costa Rica)

• Pheidole sagana Wheeler, W.M., 1934

nitidula Emery, 1888 (Argentina)

• Pheidole nitidula daguerrei Santschi, 1931

• Pheidole perversa Forel, 1908

• Pheidole perversa richteri Forel, 1909

• Pheidole strobeli Emery, 1906

• Pheidole strobeli misera Santschi, 1916

• Pheidole strobeli silvicola Borgmeier, 1927

njassae Viehmeyer, 1914 (Tanzania)

njassae legitima Santschi, 1916

• Pheidole njassae sculptior Viehmeyer, 1914

noar Wilson, 2003 (Dominican Republic)

noda Smith, F., 1874 (Japan)

• Pheidole nodus flebilis Santschi, 1937

• Pheidole nodus praevexata Wheeler, W.M., 1929

• Pheidole rhombinoda Mayr, 1879

• Pheidole rhombinoda formosensis Forel, 1913

• Pheidole rhombinoda micantiventris Mayr, 1897

• Pheidole rhombinoda stella Forel, 1911

• Pheidole rhombinoda taprobanae Forel, 1902

• Pheidole treubi Forel, 1905

nodgii Forel, 1905 (Indonesia (Java))

nodgii verlatenensis Wheeler, W.M., 1937 (Indonesia (Verlaten Island))

nodifera (Smith, F., 1858) (China)

nubicola Wilson, 2003 (Mexico)

nubila Emery, 1906 (Argentina)

nuculiceps Wheeler, W.M., 1908 (United States)

O
oaxacana Wilson, 2003 (Mexico)

obnixa Forel, 1912 (Argentina)

obrima Wilson, 2003 (Venezuela)

obscurifrons Santschi, 1925 (Argentina)

obscurithorax Naves, 1985 (Argentina)

obtusopilosa Mayr, 1887 (Uruguay)

obtusospinosa Pergande, 1896 (Mexico)

• Pheidole subdentata Pergande, 1896

• Pheidole arizonica Santschi, 1911

occipitalis André, 1890 (Sierra Leone)

occipitalis adami Santschi, 1939 (Guinea)

occipitalis neutralis Santschi, 1914 (Guinea)

oceanica Mayr, 1866 (Fiji)

• Pheidole oceanica boraborensis Wheeler, W.M., 1932

• Pheidole oceanica pattesoni Mann, 1919

• Pheidole oceanica upoluana Santschi, 1928

ocellata Zhou, 2001 (China)

ochracea Eguchi, 2008 (Vietnam)

oculata (Emery, 1899) (Madagascar)

oliveirai Wilson, 2003 (Colombia)

olsoni Wilson, 2003 (Costa Rica)

onifera Mann, 1921 (Fiji Is)

onyx Wilson, 2003 (Costa Rica)

opaciventris Mayr, 1876 (Australia)

optiva Forel, 1901 (Mexico)

orbica Forel, 1893 (Antilles)

• Pheidole californica shoshoni Cole, 1933

oswaldi Forel, 1891 (Madagascar)

oswaldi decollata Forel, 1892 (Madagascar)

otisi Wilson, 2003 (Costa Rica)

oxyops Forel, 1908 (Paraguay)

• Pheidole genalis Borgmeier, 1929

• Pheidole oxyops regia Forel, 1908

P
paiute Gregg, 1959 (United States)

palenquensis Wilson, 2003 (Ecuador)

pallidula (Nylander, 1849) (Italy (Sicily))

• Oecophthora subdentata Mayr, 1853

• Pheidole pallidula arenarum Ruzsky, 1905

• Pheidole pallidula cicatricosa Stitz, 1917

• Pheidole pallidula orientalis Müller, 1923

• Xenoaphaenogaster inquilina Baroni Urbani, 1964

pallidula koshewnikovi Ruzsky, 1905 (Russia)

pallidula obscura Santschi, 1936

• Pheidole pallidula emeryi Krausse, 1912

pallidula recticeps Menozzi, 1932 (Egypt)

pallidula selenia Özdikmen, 2010

• Pheidole pallidula inermis Stitz, 1917

pallidula tasdelenia Özdikmen, 2010

• Pheidole pallidula tristis Forel, 1907

pampana Santschi, 1929 (Argentina)

paraensis Wilson, 2003 (Brazil)

pararugiceps Longino, 2009 (Costa Rica)

parasitica Wilson, 1984 (India)

pariana Wilson, 2003 (Venezuela)

parva Mayr, 1865 (Sri Lanka)

• Pheidole bugi Wheeler, W.M., 1919

• Pheidole parva decanica Forel, 1902

• Pheidole rinae mala Forel, 1911

• Pheidole rinae tipuna Forel, 1912

• Pheidole sauteri Wheeler, W.M., 1909

parvicorpus Eguchi, 2001 (Borneo)

peckorum Wilson, 2003 (Ecuador)

pedana Wilson, 2003 (Brazil)

pegasus Sarnat, 2008 (Fiji Is)

peguensis Emery, 1895 (Myanmar)

pelor Wilson, 2003 (United States)

peltastes Wilson, 2003 (Brazil)

penetralis Smith, F., 1863 (Indonesia (Misool Island))

pepo Wilson, 2003 (Colombia)

peregrina Wheeler, W.M., 1916 (Brazil)

• Pheidole impariceps Santschi, 1923

• Pheidole impariceps fusciclava Santschi, 1923

perkinsi Wilson, 2003 (Dominica)

perpilosa Wilson, 2003 (United States)

perpusilla Emery, 1894 (Brazil)

• Pheidole decem Forel, 1901

• Pheidole emersoni Wheeler, W.M., 1922

• Pheidole flavens breviscapa Forel, 1899

perryorum Wilson, 2003 (Brazil)

peruviana Wilson, 2003 (Peru)

petrensis Wilson, 2003 (Mexico)

phanigaster Longino, 2009 (Mexico)

philemon Forel, 1910 (Solomon Is)

philippi Emery, 1915 (Ethiopia)

phipsoni Forel, 1902 (India)

pholeops Wilson, 2003 (Peru)

picata Forel, 1891 (Madagascar)

picata bernhardae Emery, 1915 (Madagascar)

picata gietleni Forel, 1905 (Madagascar)

picea (Buckley, 1867) (United States)

piceonigra Emery, 1922

• Pheidole picea Mayr Mayr, 1870

picobarva Longino, 2009 (Costa Rica)

pidax Wilson, 2003 (Colombia)

pieli Santschi, 1925 (China)

• Pheidole rinae incensa Wheeler, W.M., 1928

pilifera (Roger, 1863) (United States)

• Pheidole pennsylvanica Roger, 1863

• Pheidole pilifera artemisia Cole, 1933

• Pheidole pilifera coloradensis Emery, 1895

• Pheidole pilifera septentrionalis Wheeler, W.M., 1908

• Pheidole xerophila pacifica Wheeler, W.M., 1915

pilispina Wilson, 2003 (Panama)

piliventris (Smith, F., 1858) (Brazil)

pilosior Wilson, 2003 (United States)

pinealis Wheeler, W.M., 1908 (United States)

pinicola Wilson, 2003 (Mexico)

plagiaria Smith, F., 1860 (Indonesia (Batjan Island))

• Pheidole divergens Mayr, 1867

• Pheidole peguensis yomensis Forel, 1903

plagiaria moica Forel, 1911 (Vietnam)

plagiaria palawanica Stitz, 1925 (Philippines)

plagiaria rectilineata Viehmeyer, 1916 (Indonesia (Sulawesi) (in Copal))

planidorsum Eguchi, 2001 (Borneo)

planifrons Santschi, 1920 (Vietnam)

plato Wilson, 2003 (Ecuador)

platypus Crawley, 1915 (Australia)

plebecula Forel, 1899 (Costa Rica)

plinii Forel, 1911 (Borneo)

polita Emery, 1894 (Bolivia)

• Pheidole wheeleri Mann, 1916

polymorpha Wilson, 2003 (Mexico)

porcula Wheeler, W.M., 1908 (United States)

poringensis Eguchi, 2001 (Borneo)

portalensis Wilson, 2003 (United States)

potosiana Wilson, 2003 (Mexico)

praeses Wilson, 2003 (Brazil)

praeusta Roger, 1863 (Colombia)

prattorum Wilson, 2003 (Costa Rica)

prelli Forel, 1911 (Kenya)

prelli ingenita Santschi, 1928 (Ivory Coast)

prelli redbankensis Forel, 1913 (Zimbabwe)

pronotalis Forel, 1902 (Sri Lanka)

prostrata Wilson, 2003 (Costa Rica)

protea Forel, 1912 (Indonesia (Sumatra))

protensa Wilson, 2003 (Mexico)

providens (Sykes, 1835) (India)

proxima Mayr, 1876 (Australia)

proxima bombalensis Forel, 1910 (Australia)

proxima transversa Forel, 1902 (Australia)

psammophila Creighton & Gregg, 1955 (United States)

psilogaster Wilson, 2003 (Mexico)

pubiventris Mayr, 1887 (Brazil)

• Pheidole indistincta Forel, 1899

• Pheidole pubiventris cearensis Forel, 1901

• Pheidole pubiventris nevadensis Forel, 1901

• Pheidole pubiventris timmii Forel, 1901

• Pheidole pubiventris variegata Emery, 1896

pugnax Dalla Torre, 1892

• Pheidole radowszkowskii militaris Emery, 1890

pulchella Santschi, 1910 (Congo)

• Pheidole niapuana Wheeler, W.M., 1922

• Pheidole pulchella achantella Santschi, 1939

pullula Santschi, 1911 (Argentina)

punctatissima Mayr, 1870 (Mexico)

• Pheidole punctatissima napaea Wheeler, W.M., 1934

punctithorax Borgmeier, 1929 (Brazil)

punctulata Mayr, 1866 (South Africa)

• Pheidole inquilina Forel Forel, 1914

punctulata atrox Forel, 1913 (Democratic Republic of Congo)

• Pheidole inquilina Forel Forel, 1914

punctulata spinosa Forel, 1891 (Madagascar)

punctulata subatrox Santschi, 1937 (Democratic Republic of Congo)

purpurascens Emery, 1897 (New Guinea)

purpurea Longino, 2009 (Mexico)

• Myrmica laevigata Smith, F., 1855

• Pheidole laevigata Mayr, 1862

puttemansi Forel, 1911 (Brazil)

• Pheidole fallax janeirensis Forel, 1912

pygmaea Wilson, 2003 (Colombia)

pyriformis Clark, 1938 (Australia)

Q
quadrensis Forel, 1900 (Indonesia (Sumatra))

quadriceps Wilson, 2003 (Colombia)

quadricuspis Emery, 1900 (Indonesia (Sumatra))

quadriprojecta Smith, M.R., 1947 (New Guinea)

quadrispinosa (Smith, F., 1865) (Indonesia (Salwatty Island))

• Pheidolacanthinus armatus Smith, F., 1865

• Pheidole smithi Emery, 1887

quercicola Wilson, 2003 (Mexico)

quiaccana Wheeler, W.M., 1925 (Bolivia)

quinata Eguchi, 2000 (Borneo)

R
rabo Forel, 1913 (Indonesia (Sumatra))

• Pheidole concinna Wheeler, W.M., 1928

• Pheidole tsailuni Wheeler, W.M., 1929

radoszkowskii Mayr, 1884 (French Guiana)

• Pheidole medialis Wilson, 2003

• Pheidole radoszkowskii luteola Forel, 1893

• Pheidole radoszkowskii opacissima Forel, 1901

• Pheidole radoszkowskii parvinoda Forel, 1912

• Pheidole radowszkowskii acuta Emery, 1894

• Pheidole radowszkowskii australis Emery, 1890

• Pheidole triconstricta paranana Santschi, 1925

rebeccae Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

reclusi Forel, 1899 (Colombia)

recondita Clouse, 2007 (Guam Island)

rectisentis Wilson, 2003 (Nicaragua)

rectispina Wilson, 2003 (Costa Rica)

rectitrudis Wilson, 2003 (Costa Rica)

reflexans Santschi, 1933 (Argentina)

reichenspergeri Santschi, 1923 (Brazil)

renae Wilson, 2003 (Costa Rica)

retivertex Eguchi, 2001 (Borneo)

retronitens Santschi, 1930 (Democratic Republic of Congo)

rhea Wheeler, W.M., 1908 (United States)

rhinoceros Forel, 1899 (Panama)

rhinomontana Longino, 2009 (Costa Rica)

rhytifera Wilson, 2003 (Venezuela)

rinae Emery, 1900 (Indonesia (Sumatra))

risii Forel, 1892 (Argentina)

riveti Santschi, 1911 (Ecuador)

roberti Forel, 1902 (India)

rochai Forel, 1912 (Brazil)

rogeri Emery, 1896 (Costa Rica)

rogersi Forel, 1902 (India)

rogersi taylori Forel, 1902 (India)

rohani Santschi, 1925 (Angola)

rohani monardi Santschi, 1930 (Angola)

rohani pellax Santschi, 1930 (Angola)

roosevelti Mann, 1921 (Fiji Is)

rosae Forel, 1901 (Brazil)

• Pheidole silvestrii Emery, 1906

rosula Wilson, 2003 (Argentina)

rotundiceps Wilson, 2003 (Ecuador)

roushae Wilson, 2003 (Mexico)

rubiceps Wilson, 2003 (Venezuela)

rudigenis Emery, 1906 (Argentina)

• Pheidole bucolica Santschi, 1929

ruficeps (Smith, F., 1861) (Indonesia (Sulawesi))

• Pheidole mordax Smith, F., 1863

rufipilis Forel, 1908 (Brazil)

• Pheidole pubiventris impia Santschi, 1923

• Pheidole rufipilis divexa Forel, 1908

rugaticeps Emery, 1877 (Ethiopia)

rugaticeps arabs Emery, 1881 (Yemen)

rugatula Santschi, 1933 (Argentina)

rugiceps Wilson, 2003 (Costa Rica)

rugifera Eguchi, 2001 (Borneo)

rugithorax Eguchi, 2008 (Vietnam)

rugosa Smith, F., 1858 (Sri Lanka)

rugulosa Gregg, 1959 (United States)

rutilana Wilson, 2003 (Brazil)

ryukyuensis Ogata, 1982 (Japan)

S
sabahna Eguchi, 2000 (Borneo)

sabella Wilson, 2003 (Peru)

sabina Wilson, 2003 (Mexico)

sagax Wilson, 2003 (Colombia)

sagei Forel, 1902 (India)

sagittaria Wilson, 2003 (Costa Rica)

sarawakana Forel, 1911 (Borneo)

sarcina Forel, 1912 (Brazil)

sarpedon Wilson, 2003 (Ecuador)

sauberi Forel, 1905 (Indonesia (Java))

saxicola Wheeler, W.M., 1922 (Democratic Republic of Congo)

sayapensis Eguchi, 2001 (Borneo)

scabriuscula Gerstäcker, 1871 (Kenya)

scalaris Wilson, 2003 (Costa Rica)

scapulata Santschi, 1923 (Argentina)

schmalzi Emery, 1894 (Brazil)

• Pheidole rata Borgmeier, 1929

• Pheidole rehi Forel, 1900

schoedli Eguchi, Hashimoto & Malsch, 2006 (Borneo)

schoutedeni Forel, 1913 (Democratic Republic of Congo)

schoutedeni platycephala Stitz, 1916 (Democratic Republic of Congo)

schultzei Forel, 1910 (Botswana)

schultzei ebangana Santschi, 1937 (Angola)

schultzei gwaaiensis Forel, 1913 (Zimbabwe)

schultzei woodvalensis Arnold, 1960 (Zimbabwe)

schwarzmaieri Borgmeier, 1939 (Brazil)

sciara Cole, 1955 (United States)

scimitara Wilson, 2003 (Peru)

sciophila Wheeler, W.M., 1908 (United States)

• Pheidole proserpina Wheeler, W.M., 1908

• Pheidole sciophila semilaevicephala Smith, M.R., 1934

scolioceps Wilson, 2003 (Peru)

scrobifera Emery, 1896 (Costa Rica)

sculptior Forel, 1893 (Antilles)

• Pheidole tayrona Wilson, 2003

sculpturata Mayr, 1866 (South Africa)

sculpturata areolata Forel, 1911 (South Africa)

sculpturata berthoudi Forel, 1894 (South Africa)

sculpturata dignata Santschi, 1915 (Congo)

sculpturata particeps Santschi, 1921 (Tanzania)

sculpturata rhodesiana Forel, 1913 (Zimbabwe)

sculpturata welgelegenensis Forel, 1913 (Democratic Republic of Congo)

sculpturata zambesiana Forel, 1913 (Zimbabwe)

sebofila Longino, 2009 (Costa Rica)

securigera Wilson, 2003 (Colombia)

seeldrayersi Forel, 1910 (Colombia)

selathorax Zhou, 2001 (China)

seligmanni Wilson, 2003 (Colombia)

semidea Fischer, Hita Garcia & Peters, 2012 (Nigeria)

semilaevis Forel, 1901 (Brazil)

senex Gregg, 1952 (United States)

• Pheidole pilifera anfracta Cole, 1952

senilis Santschi, 1929 (Brazil)

sensitiva Borgmeier, 1959 (Brazil)

sepulchralis Bingham, 1903 (India)

sericella Viehmeyer, 1914 (New Guinea)

servilia Wilson, 2003 (Colombia)

setosa Fischer, Hita Garcia & Peters, 2012 (Democratic Republic of Congo)

setsukoae Wilson, 2003 (Mexico)

severini Forel, 1904 (Suriname)

sexdentata Donisthorpe, 1948 (New Guinea)

sexspinosa Mayr, 1870 (Tuvalu)

• Pheidole sexspinosa adamsoni Wheeler, W.M., 1932

sexspinosa biroi Emery, 1900 (New Guinea)

sexspinosa fuscescens Emery, 1900 (New Guinea)

sharpi Forel, 1902 (India)

sharpi hoogwerfi Forel, 1902 (Sri Lanka)

sicaria Wilson, 2003 (Costa Rica)

sigillata Wilson, 2003 (Brazil)

sikorae Forel, 1891 (Madagascar)

sikorae litigiosa Forel, 1892 (Madagascar)

similigena Wheeler, W.M., 1937 (Cuba)

simoni Emery, 1893 (Philippines)

simonsi Wilson, 2003 (Costa Rica)

simplex Wheeler, W.M., 1925 (Peru)

simplispinosa Sarnat, 2008 (Fiji Is)

sinaitica Mayr, 1862 (Egypt)

sinaitica santschii Forel, 1907 (Tunisia)

singaporensis Özdikmen, 2010

• Myrmica longipes Smith, F., 1857

singaporensis conicollis (Emery, 1900) (Indonesia (Sumatra))

singaporensis continentis (Forel, 1911) (Myanmar)

singaporensis pseudola Forel, 1915 (Indonesia (Sumatra))

singularis Smith, F., 1863 (Indonesia (Misool Island))

sinica (Wu & Wang, 1992) (China)

sitiens Wilson, 2003 (United States)

skwarrae Wheeler, W.M., 1934 (Mexico)

smythiesii Forel, 1902 (India)

• Pheidole bhavanae Bingham, 1903

• Pheidole smythiesii bengalensis Forel, 1902

socrates Forel, 1912 (Brazil)

soesilae Makhan, 2007 (Surinam)

soritis Wheeler, W.M., 1908 (United States)

• Pheidole sitarches Wheeler, W.M., 1908

• Pheidole sitarches campestris Creighton, 1950

• Pheidole sitarches rufescens Wheeler, W.M., 1908

• Pheidole sitarches transvarians Wheeler, W.M., 1908

sospes Forel, 1908 (Brazil)

• Pheidole minutissima Kusnezov, 1952

spadonia Wheeler, W.M., 1915 (United States)

sparsisculpta Longino, 2009 (Costa Rica)

spathicornis Wilson, 2003 (Mexico)

spathifera Forel, 1902 (India)

spathifera aspatha Forel, 1902 (India)

spathifera yerburyi Forel, 1902 (Sri Lanka)

spathipilosa Wilson, 2003 (Costa Rica)

specularis Wilson, 2003 (Costa Rica)

speculifera Emery, 1877 (Ethiopia)

speculifera ascara Emery, 1901 (Ethiopia)

speculifera bispecula Santschi, 1930 (Democratic Republic of Congo)

speculifera cubangensis Forel, 1901 (Angola)

sperata Forel, 1915 (Indonesia (Sumatra))

sphaerica Wilson, 2003 (Colombia)

spilota Wilson, 2003 (Colombia)

spinicornis Eguchi, 2001 (Borneo)

spininodis Mayr, 1887 (Argentina)

• Pheidole hohenlohei Emery, 1888

• Pheidole spielbergi Emery, 1888

• Pheidole spininodis bruta Santschi, 1934

• Pheidole spininodis lucifuga Santschi, 1923

• Pheidole spininodis pencosensis Forel, 1914

• Pheidole spininodis solaris Santschi, 1929

spinoda (Smith, F., 1858) (Australia)

spinulosa Forel, 1910 (Lesotho)

spinulosa conigera Forel, 1910 (South Africa)

spinulosa messalina Forel, 1910 (Lesotho)

squalida Santschi, 1910 (Congo)

steinheili Forel, 1901 (Colombia)

stigma Wilson, 2003 (Ecuador)

stomachosa Wheeler, W.M., 1917 (Jamaica)

strator Forel, 1910 (Ethiopia)

strator fugax Arnold, 1920 (Mozambique)

strator tabida Menozzi, 1933 (Uganda)

striata Donisthorpe, 1947 (New Guinea)

striaticeps Mayr, 1870 (Mexico)

strigosa Wilson, 2003 (Brazil)

stulta Forel, 1886 (Brazil)

• Pheidole championi Forel, 1899

• Pheidole championii sima Forel, 1912

styrax Wilson, 2003 (Venezuela)

subaberrans (Kusnezov, 1952) (Argentina)

subarmata Mayr, 1884 (French Guiana)

• Pheidole cornutula Emery, 1890

• Pheidole cornutula dentimentum Santschi, 1929

• Pheidole cornutula imbecillis Emery, 1906

• Pheidole hondurensis Mann, 1922

• Pheidole subarmata borinquenensis Wheeler, W.M., 1908

• Pheidole subarmata elongatula Forel, 1893

• Pheidole subarmata nassavensis Wheeler, W.M., 1905

• Pheidole subarmata nefasta Santschi, 1929

submonticola Eguchi, 2001 (Borneo)

subnuda Wilson, 2003 (Colombia)

subreticulata Emery, 1894 (Brazil)

subsphaerica Wilson, 2003 (Colombia)

sulcaticeps Roger, 1863 (Sri Lanka)

sulcaticeps punensis Forel, 1902 (India)

sulcaticeps vellicans Forel, 1911 (Sri Lanka)

superba Wilson, 2003 (Colombia)

susannae Forel, 1886 (Guatemala)

• Pheidole incisa evoluta Borgmeier, 1929

• Pheidole partita Mayr, 1887

• Pheidole susannae atricolor Forel, 1901

• Pheidole susannae obscurior Forel, 1886

susanowo Onoyama & Terayama, 1999 (Japan)

sykesii Forel, 1902 (India)

symbiotica Wasmann, 1909 (Portugal)

synanthropica Longino, 2009 (Costa Rica)

synarmata Wilson, 2003 (Guyana)

T
tachigaliae Wheeler, W.M., 1921 (Guyana)

tachirana Wilson, 2003 (Venezuela)

taipoana Wheeler, W.M., 1928 (China)

taivanensis Forel, 1912 (Taiwan)

tambopatae Wilson, 2003 (Peru)

tandjongensis Forel, 1913 (Indonesia (Sumatra))

tanyscapa Wilson, 2003 (Costa Rica)

tarchon Wilson, 2003 (Argentina)

tarda Donisthorpe, 1947 (Mauritius)

tasmaniensis Mayr, 1866 (Australia)

tasmaniensis continentis Forel, 1902 (Australia)

taurus Emery, 1906 (Argentina)

tawauensis Eguchi, 2001 (Borneo)

templaria Forel, 1902 (India)

templaria euscrobata Forel, 1913 (Sri Lanka)

tenebricosa Eguchi, 2001 (Borneo)

tenerescens Wheeler, W.M., 1922 (Trinidad)

teneriffana Forel, 1893 (Spain (Canary Is))

• Pheidole teneriffana taina Aguayo, 1932

tennantae Wilson, 2003 (Costa Rica)

tenuicephala Longino, 2009 (Costa Rica)

tenuiclavata Donisthorpe, 1943 (New Guinea)

tenuinodis Mayr, 1901 (South Africa)

tenuinodis bothae Forel, 1901 (South Africa)

tenuinodis robusta Stitz, 1923 (Namibia)

tenuinodis sipapomae Arnold, 1920 (Zimbabwe)

tenuis Wilson, 2003 (Colombia)

tepicana Pergande, 1896 (Mexico)

• Pheidole carbonaria Pergande, 1896

• Pheidole kingi André, 1898

• Pheidole kingi instabilis Emery, 1901

• Pheidole kingi torpescens Wheeler, W.M., 1915

• Pheidole rugifrons Pergande, 1896

• Pheidole townsendi André, 1898

tepuicola Wilson, 2003 (Venezuela)

termitobia Forel, 1901 (Brazil)

termitophila Forel, 1904 (Sudan)

termitophila liberiensis Forel, 1911 (Liberia)

terraceensis Bharti, 2001 (India)

terresi Wheeler, W.M. & Mann, 1914 (Haiti)

• Pheidole terresi illota Wheeler, W.M. & Mann, 1914

terribilis Wilson, 2003 (Surinam)

tetra Creighton, 1950 (United States)

tetracantha Emery, 1897 (New Guinea)

tetrica Forel, 1913 (Brazil)

tetroides Wilson, 2003 (Mexico)

texana Wheeler, W.M., 1903 (United States)

texticeps Wilson, 2003 (Costa Rica)

• Pheidole perdiligens Wilson, 2003

thrasys Wilson, 2003 (Panama)

tigris Wilson, 2003 (Colombia)

tijucana Borgmeier, 1927 (Brazil)

tillandsiarum Wheeler, W.M., 1934 (Mexico)

tisiphone Wheeler, W.M., 1911 (Mexico)

titanis Wheeler, W.M., 1903 (United States)

tjibodana Forel, 1905 (Indonesia (Java))

tobini Wilson, 2003 (Peru)

tolteca Forel, 1901 (Mexico)

torosa Wilson, 2003 (Mexico)

trachyderma Emery, 1906 (Argentina)

trageri Wilson, 2003 (Brazil)

tragica Wheeler, W.M., 1934 (Mexico)

traini Wilson, 2003 (Panama)

transfigens Forel, 1911 (New Guinea)

transversostriata Mayr, 1887 (Guyana)

• Pheidole lacerta Wheeler, W.M., 1922

• Pheidole transversostriata nigridens Forel, 1901

trapezoidea Viehmeyer, 1914 (Australia)

tricarinata Santschi, 1914 (Guinea)

tricolor Donisthorpe, 1949 (New Guinea)

triconstricta Forel, 1886 (Argentina)

• Pheidole radoszkowskii discursans Forel, 1912

• Pheidole radoszkowskii saviozae Forel, 1911

• Pheidole triconstricta ambulans Emery, 1906

• Pheidole triconstricta hebe Santschi, 1923

• Pheidole triconstricta rosariensis Forel, 1913

trinitatis Wilson, 2003 (Trinidad)

triplex Wilson, 2003 (Trinidad)

tristicula Wilson, 2003 (Peru)

tristis (Smith, F., 1858) (Brazil)

• Atta fumipennis Smith, F., 1858

• Atta rubra Smith, F., 1858

• Pheidole crassipes Mayr, 1887

• Pheidole emeryi Mayr, 1887

• Pheidole emeryi alsia Santschi, 1923

tristops Wilson, 2003 (Colombia)

truncula Wilson, 2003 (Costa Rica)

tschinkeli Wilson, 2003 (Guatemala)

tumida Eguchi, 2008 (Vietnam)

turneri Forel, 1902 (Australia)

tuxtlasana Wilson, 2003 (Mexico)

tysoni Forel, 1901

U
ulothrix Wilson, 2003 (Costa Rica)

umbonata Mayr, 1870 (Tonga)

• Pheidole umbonata zimmermani Wheeler, W.M., 1936

umbonata fusciventris Emery, 1900 (New Guinea)

umphreyi Wilson, 2003 (Panama)

uncagena Sarnat, 2008 (Fiji Is)

unicornis Wilson, 2003 (Colombia)

upeneci Forel, 1913 (Indonesia (Java))

ursus Mayr, 1870 (Mexico)

• Macromischa cressoni André, 1887

• Pheidole ursus gracilinoda Forel, 1904

V
vafella Wheeler, W.M., 1925 (Bolivia)

vafra Santschi, 1923 (Brazil)

• Pheidole idiota Santschi, 1923

valens Wilson, 2003 (Brazil)

vallicola Wheeler, W.M., 1915 (United States)

vallifica Forel, 1901 (Colombia)

vanderveldi Forel, 1913 (Democratic Republic of Congo)

variabilis Mayr, 1876 (Australia)

variabilis aliena Emery, 1914 (New Caledonia)

variabilis croceithorax Emery, 1914 (New Caledonia)

variabilis latigena Forel, 1907 (Australia)

variabilis mediofusca Forel, 1902 (Australia)

variabilis ocior Forel, 1915 (Australia)

variabilis ocyma Forel, 1915 (Australia)

variabilis parvispina Forel, 1902 (Australia)

variabilis praedo Forel, 1902 (Australia)

variabilis redunca Crawley, 1915 (Australia)

variabilis rugocciput Forel, 1902 (Australia)

variabilis rugosula Forel, 1902 (Australia)

variolosa Emery, 1892 (Somalia)

vatu Mann, 1921 (Fiji Is)

veletis Wilson, 2003 (Bolivia)

velox Emery, 1887 (Indonesia (Ternate Island))

venatrix Wilson, 2003 (Colombia)

verricula Wilson, 2003 (Costa Rica)

vestita Wilson, 2003 (Costa Rica)

veteratrix Forel, 1891 (Madagascar)

veteratrix angustinoda Forel, 1892 (Madagascar)

victima Santschi, 1929 (Brazil)

victoris Forel, 1913 (Zimbabwe)

vieti Eguchi, 2008 (Vietnam)

vigilans (Smith, F., 1858) (Australia)

• Pheidole ampla norfolkensis Wheeler, W.M., 1927

• Pheidole ampla parallela Forel, 1902

• Pheidole ampla yarrensis Forel, 1902

• Pheidole dolichocephala André, 1896

• Pheidole vinelandica nebrascensis Forel, 1922

violacea Wilson, 2003 (Costa Rica)

virago Wheeler, W.M., 1915 (United States)

viriosa Wilson, 2003 (Ecuador)

vistana Forel, 1914 (United States)

• Pheidole grallipes Wheeler, W.M., 1916

• Pheidole susannae longipes Pergande, 1896

voeltzkowii Forel, 1894 (Madagascar)

vomer Wilson, 2003 (Colombia)

vorax (Fabricius, 1804) (Central America)

• Pheidole cephalica Smith, F., 1858

• Pheidole opaca Mayr, 1862

• Pheidole opaca apterostigmoides Weber, 1943

• Pheidole opaca incrustata Forel, 1908

• Pheidole opaca sarrita Forel, 1908

vulgaris Eguchi, 2006 (Vietnam)

W
walkeri Mann, 1922 (Honduras)

• Pheidole arietans Wilson, 2003

• Pheidole glyphoderma Wilson, 2003

• Pheidole triumbonata Wilson, 2003

wallacei Mann, 1916 (Brazil)

wardi Wilson, 2003 (Honduras)

watsoni Forel, 1902 (Myanmar)

weiseri Santschi, 1923 (Argentina)

wheelerorum Mackay, 1988 (United States)

wiesei Forel, 1910 (Australia)

williamsi Wheeler, W.M., 1919 (Ecuador (Galapagos Is))

• Pheidole williamsi seymourensis Wheeler, W.M., 1924

wilsoni Mann, 1921 (Fiji Is)

wolfringi Forel, 1908 (Brazil)

woodmasoni Forel, 1885 (India)

wroughtonii Forel, 1902 (India)

X
xanthocnemis Emery, 1914 (New Caledonia)

xanthogaster Wilson, 2003 (Peru)

xerophila Wheeler, W.M., 1908 (United States)

• Pheidole xerophila tucsonica Wheeler, W.M., 1908

xocensis Forel, 1913 (Zimbabwe)

xocensis bulawayensis Forel, 1913 (Zimbabwe)

xyston Wilson, 2003 (Mexico)

Y
yaqui Creighton & Gregg, 1955 (United States)

yeensis Forel, 1902 (Myanmar)

yucatana Wilson, 2003 (Mexico)

Z
zelata Wilson, 2003 (Bolivia)

zeteki Smith, M.R., 1947 (Panama)

zhoushanensis Li & Chen, 1992 (China)

zoceana Santschi, 1925 (China)

zoster Wilson, 2003 (Peru)