Crematogaster sordidula

This is a common species in Greece, known from all provinces. It prefers warm mediterranean habitats such as pastures, mediterranean shrubs and roadsides. Nests located under stones or directly in ground (Borowiec & Salata, 2021). This ant was observed on larvae of that were feeding on Erophaca baetica in Sierra Morena, southern Spain. It has also been associated with the butterflies and  (Obregon et al. 2015).

Distribution
This is a common species, known from all Greek provinces (Borowiec et al., 2022).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Azerbaijan, Balearic Islands, Bulgaria, Croatia, France, Georgia, Gibraltar, Greece, Hungary, Iberian Peninsula, Israel, Italy, Montenegro, Portugal, Republic of Macedonia, Romania, Russian Federation, Slovenia, Spain, Switzerland, Turkey.

Nomenclature

 * . Myrmica sordidula Nylander, 1849: 44 (w.) ITALY (Sicily).
 * Type-material: 3 syntype workers (Radchenko, 2007: 33).
 * Type-locality: Italy: Sicily, Messina (C. Zeller).
 * Type-depository: ZMHF.
 * Mayr, 1855: 472 (q.); Forel, 1870: 308 (m.).
 * Combination in Crematogaster: Mayr, 1855: 472;
 * combination in C. (Orthocrema): Santschi, 1918d: 182.
 * Status as species: Mayr, 1855: 472 (redescription); Nylander, 1856b: 101; Smith, F. 1858b: 135; Mayr, 1861: 75 (in key); Mayr, 1862: 768; Mayr, 1863: 405; Roger, 1863b: 36; Mayr, 1865: 108; André, 1874: 201 (in key); Mayr, 1877: 19; Emery, 1878a: x (in list); Emery, 1878b: 59; Emery & Forel, 1879: 464; Mayr, 1880: 39; Emery, 1881b: 535; André, 1881b: 77; André, 1883b: 391 (in key); Forel, 1886e: clxvi; Forel, 1889: 257; Saunders, E. 1890: 205; Emery, 1891b: 15; Dalla Torre, 1893: 86; Medina, 1893: 105; Forel, 1894d: 24; Mayr, 1895: 135 (in key); Ruzsky, 1902d: 31; Ruzsky, 1905b: 502; Forel, 1906c: 189; Karavaiev, 1910b: 48; Emery, 1912e: 669; Karavaiev, 1912a: 11; Karavaiev, 1912b: 583; Emery, 1914d: 157; Forel, 1915d: 32 (in key); Emery, 1916b: 156; Bondroit, 1918: 113; Santschi, 1919e: 244; Arnold, 1920a: 537; Menozzi, 1921: 27; Emery, 1922e: 130; Menozzi, 1922b: 328; Wheeler, W.M. 1922a: 850, 1024; Kuznetsov-Ugamsky, 1923: 252; Müller, 1923a: 69; Müller, 1923b: 74; Finzi, 1924a: 13; Emery, 1924c: 164; Santschi, 1925g: 347; Emery, 1926: 5; Donisthorpe, 1926a: 8; Karavaiev, 1926b: 109; Kutter, 1927: 99; Wheeler, W.M. 1928d: 111; Kuznetsov-Ugamsky, 1929b: 43; Santschi, 1929e: 147; Finzi, 1930d: 313; Santschi, 1931a: 5; Grandi, 1935: 100; Santschi, 1936c: 201; Santschi, 1937e: 311; Weber, 1943c: 357; Donisthorpe, 1950e: 1061; Schmitz, 1950: 15; Ceballos, 1956: 304; Bernard, 1959: 346; Fromantin & Soulié, 1961: 93; Dlussky, 1962: 179; Baroni Urbani, 1964b: 43; Baroni Urbani, 1964c: 152; Cagniant, 1964: 88; Cagniant, 1966b: 279; Bernard, 1967: 163 (redescription); Pisarski, 1967: 388; Cagniant, 1968a: 143; Baroni Urbani, 1969a: 331; Collingwood & Yarrow, 1969: 66; Cagniant, 1970a: 417; Pisarski, 1970: 308; Baroni Urbani, 1971c: 82; Tarbinsky, 1976: 76 (redescription); Kutter, 1977c: 89; Arnol'di & Dlussky, 1978: 538 (in key); Collingwood, 1978: 81 (in key); Agosti & Collingwood, 1987a: 54; Agosti & Collingwood, 1987b: 272 (in key); Kugler, J. 1988: 258; Atanassov & Dlussky, 1992: 182; Collingwood, 1993b: 194; Bolton, 1995b: 163; Poldi, et al. 1995: 4; Espadaler, 1997b: 29; Collingwood & Prince, 1998: 14 (in key); Markó & Csösz, 2002: 116; Cagniant, 2005: 10; Bračko, 2006: 136; Cagniant, 2006a: 198; Markó, Sipos, et al. 2006: 69; Petrov, 2006: 93 (in key); Bračko, 2007: 17; Seifert, 2007: 216; Radchenko, 2007: 33; Gratiashvili & Barjadze, 2008: 138; Casevitz-Weulersse & Galkowski, 2009: 487; Vonshak, et al. 2009: 42; Lapeva-Gjonova, et al. 2010: 18; Karaman, M.G. 2011a: 10 (redescription); Csösz, et al. 2011: 57; Karaman, M.G. 2011b: 25; Legakis, 2011: 14; Borowiec, L. & Salata, 2012: 491; Kiran & Karaman, 2012: 18; Borowiec, L. 2014: 69 (see note in bibliography); Bračko, et al. 2014: 13; Lebas, et al. 2016: 276; Borowiec, L. & Salata, 2018: 5; Salata & Borowiec, 2018c: 44; Seifert, 2018: 180.
 * Senior synonym of caspica: Atanassov & Dlussky, 1992: 182; Bolton, 1995b: 163.
 * Senior synonym of esterelana: Casevitz-Weulersse & Galkowski, 2009: 487.
 * Senior synonym of flachii: Finzi, 1930d: 313; Agosti & Collingwood, 1987a: 54; Bolton, 1995b: 163; Legakis, 2011: 13; Kiran & Karaman, 2012: 18.
 * Senior synonym of mayri: Mayr, 1855: 472; Nylander, 1856b: 101; Smith, F. 1858b: 136; Mayr, 1863: 405; Roger, 1863b: 36; André, 1874: 204 (in list); Emery & Forel, 1879: 464; Dalla Torre, 1893: 86; Ruzsky, 1905b: 502; Emery, 1922e: 130; Fromantin & Soulié, 1961: 94; Agosti & Collingwood, 1987a: 54; Bolton, 1995b: 163; Karaman, M.G. 2011a: 14 (in text); Legakis, 2011: 13; Kiran & Karaman, 2012: 18.
 * Distribution: Albania, Algeria, Azerbaijan, Bosnia, Bulgaria, Croatia, Cyprus, France, Georgia, Germany, Gibraltar, Greece, Hungary, Israel, Italy (+ Sicily), Macedonia, Montenegro, Morocco, Portugal, Romania, Russia, Serbia, Slovenia, Spain (+ Balearics), Switzerland, Turkey.
 * [Note: distribution records after Borowiec, L. 2014: 69.]
 * Current subspecies: nominal plus aeolia, marocana, molongori, osmanica.
 * caspica. Crematogaster sordidula var. caspica Emery, 1922e: 130.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Russia: Petrovsk (Korb).
 * Type-depository: MSNG.
 * [First available use of Crematogaster sordidula subsp. sordidula var. caspica Emery, 1912e: 670 (w.) RUSSIA; unavailable (infrasubspecific) name.]
 * Junior synonym of sordidula: Atanassov & Dlussky, 1992: 182; Bolton, 1995b: 150.
 * esterelana. Orthocrema esterelana Bernard, 1978b: 43, fig. 1 (w.q.m.) FRANCE.
 * Type-material: 30 syntype workers, 3 syntype queens, 2 syntype males.
 * Type-locality: France: Dept. Var, Fréjus, 45 m., on Valescure rhyolites (F. Bernard).
 * Type-depository: MNHN.
 * Combination in Crematogaster: Bolton, 1995b: 152.
 * Status as species: Bolton, 1995b: 152.
 * Junior synonym of sordidula: Casevitz-Weulersse & Galkowski, 2009: 487.
 * flachii. Crematogaster sordidula var. flachii Forel, 1895d: 229 (w.) SERBIA.
 * Type-material: syntype workers (number not stated).
 * [Note: Forel says the taxon had already been found on Crete by v. Oertzen, and by himself, but it is uncertain whether this material constitutes part of the type-series.]
 * Type-locality: Serbia: Belgrade (Flache).
 * Type-depository: MHNG.
 * Combination in C. (Orthocrema): Emery, 1922e: 130.
 * As unavailable (infrasubspecific) name: Emery, 1912e: 670.
 * Subspecies of sordidula: Forel, 1901h: 64; Forel, 1904b: 372; Ruzsky, 1905b: 505; Forel, 1911d: 343; Forel, 1913d: 430; Emery, 1914d: 157; Santschi, 1921a: 114; Emery, 1922e: 130; Finzi, 1923: 4; Karavaiev, 1927b: 106; Finzi, 1928c: 789; Menozzi, 1933b: 61 (in key); Menozzi, 1936d: 283.
 * Status as species: Tohmé, G. & Tohmé, 2014: 135 (error).
 * Junior synonym of mayri: Müller, 1923b: 74; Emery, 1926: 5; Finzi, 1927a: 10; Zimmermann, 1935: 26.
 * Junior synonym of sordidula: Finzi, 1930d: 313; Agosti & Collingwood, 1987a: 54; Bolton, 1995b: 153; Legakis, 2011: 13; Kiran & Karaman, 2012: 18.
 * mayri. Acrocoelia mayri Mayr, 1853b: 114 (w.) CROATIA (attributed to Schmidt).
 * Type-material: syntype workers (number not stated).
 * Type-locality: Croatia (“Dalmatia”): Zara (v. Manderstjerna).
 * Type-depository: NHMW.
 * Müller, 1923b: 76 (q.); Emery, 1926: 6 (m.).
 * Combination in Crematogaster: Mayr, 1855: 472.
 * Subspecies of sordidula: Müller, 1923b: 74; Emery, 1926: 6; Finzi, 1927a: 10; Cori & Finzi, 1931: 238; Zimmermann, 1935: 26; Finzi, 1939c: 156; Baroni Urbani, 1971c: 84; Aktaç, 1977: 121; Poldi, et al. 1995: 4.
 * Junior synonym of sordidula: Mayr, 1855: 472; Nylander, 1856b: 101; Smith, F. 1858b: 136; Mayr, 1863: 405; Roger, 1863b: 36; André, 1874: 204 (in list); Emery & Forel, 1879: 464; Dalla Torre, 1893: 86; Ruzsky, 1905b: 502; Emery, 1922e: 130; Fromantin & Soulié, 1961: 94; Agosti & Collingwood, 1987a: 54; Bolton, 1995b: 157; Karaman, M.G. 2011a: 14 (in text); Legakis, 2011: 13; Kiran & Karaman, 2012: 18.

References based on Global Ant Biodiversity Informatics

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