Checklist of Tetramorium species

The following species and subspecies belong to the genus Tetramorium. Synonyms are listed under their senior names. For valid names only see Tetramorium species and for distribution information see Tetramorium species by Country.

A
aculeatum (Mayr, 1866) (Cameroun, Democratic Republic of Congo, Ghana, Guinea, Ivory Coast, Kenya, Liberia, Senegal, Sierra Leone, Sudan, Uganda, United Republic of Tanzania)

• Macromischa wasmanni Forel, 1901

• Macromischoides aculeatus inermis Bernard, 1953

• Macromischoides aculeatus melanogyne Santschi, 1923

• Macromischoides aculeatus militaris Santschi, 1924

• Macromischoides aculeatus pulchellus Santschi, 1924

• Macromischoides viridis Weber, 1943

• Macromischoides zumpti Santschi, 1937

• Tetramorium aculeatum andricum Emery, 1908

• Tetramorium aculeatum major Forel, 1915

• Tetramorium aculeatum rubroflava Forel, 1916

adamsi Hita Garcia & Fisher, 2012 (Madagascar)

adelphon Bolton, 1979 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

adpressum (Bolton, 1976) (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Sulawesi)

aegeum Radchenko, 1992 (Azerbaijan, Russian Federation)

• Tetramorium inerme laevigatum Arnoldi, 1948

africanum (Mayr, 1866) (Equatorial Guinea, Ghana, Guinea)

• Rhoptromyrmex tessmanni Forel, 1910

• Tetramorium lamottei Bernard, 1953

agile Arnold, 1960 (United Republic of Tanzania, Zimbabwe)

agna (Santschi, 1935) (Democratic Republic of Congo)

aherni Hita Garcia & Fisher, 2012 (Madagascar)

aisha Mbanyana, Robertson & Hita Garcia, 2018 (Namibia, South Africa)

akengense (Wheeler, W.M., 1922) (Democratic Republic of Congo)

akermani Arnold, 1926 (South Africa)

• Tetramorium akermani myersi Arnold, 1958

ala Hita Garcia & Fisher, 2012 (Madagascar)

alperti Hita Garcia & Fisher, 2014 (Madagascar)

alpestre Steiner, Schlick-Steiner & Seifert, 2010 (Austria, France, Italy, Switzerland)

alternans Santschi, 1929 (Algeria, Morocco, Tunisia)

• Tetramorium biskrensis kahenae Menozzi, 1934

altivagans Santschi, 1914 (Kenya, Sudan, United Republic of Tanzania)

• Tetramorium simillimum isis Weber, 1943

amalae Sharaf & Aldawood, 2012 (Saudi Arabia)

amatongae Bolton, 1980 (Mozambique, United Republic of Tanzania)

amaurum Bolton, 1980 (South Africa, Zimbabwe)

ambanizana Hita Garcia & Fisher, 2012 (Madagascar)

ambatovy Hita Garcia & Fisher, 2012 (Madagascar)

amentete Bolton, 1980 (Ghana, Ivory Coast)

amissum Bolton, 1980 (Democratic Republic of Congo)

anatolicum Csösz & Schulz, 2007 (Turkey)

andohahela Hita Garcia & Fisher, 2012 (Madagascar)

andrei Forel, 1892 (Madagascar)

• Tetramorium andrei robustior Forel, 1892

andrynicum Bolton, 1977 (Australia)

angulinode Santschi, 1910 (Botswana, Cameroun, Congo, Nigeria, Sudan, Zimbabwe)

• Tetramorium humerosum Bernard, 1953

• Tetramorium nullispinum Bolton, 1980

• Xiphomyrmex angulinode daphnis Santschi, 1920

• Xiphomyrmex papyri Weber, 1943

ankarana Hita Garcia & Fisher, 2012 (Madagascar)

annectens Pisarski, 1969 (China)

anodontion Bolton, 1979 (Madagascar)

antennatum (Mann, 1919) (Solomon Islands)

antipodum Wheeler, W.M., 1927 (Australia)

antrema (Bolton, 1976) (Cameroun)

anxium Santschi, 1914 (Benin, Cameroun, Gabon, Ghana, Guinea, Ivory Coast, Togo, United Republic of Tanzania)

aptum Bolton, 1977 (Borneo, Brunei Darussalam, China, Indonesia, Malaysia, Singapore, Thailand, Vietnam)

argenteopilosum Arnold, 1926 (Kenya, South Africa, United Republic of Tanzania, Zimbabwe)

argentirubrum Dietrich, 2004 (Jordan)

armatum Santschi, 1927 (Afghanistan, Iran, Kyrgyzstan, Mongolia, Turkmenistan)

• Tetramorium caespitum annauensis Karavaiev, 1931

• Tetramorium caespitum brevinodis Karavaiev, 1927

arnoldi (Forel, 1913) (South Africa, Zimbabwe)

artemis Hita Garcia & Fisher, 2012 (Madagascar)

arzi Tohmé, G., 1969 (Lebanon)

asetyum Bolton, 1980 (Nigeria)

aspersum (Smith, F., 1865) (Indonesia, New Guinea, Philippines, Solomon Islands)

• Tetramorium bismarckii Forel, 1901

• Xiphomyrmex costatus willowsi Wheeler, W.M., 1934

aspina Wagner, Karaman, Aksoy & Kiran, 2018 (Turkey)

aspis Hita Garcia & Fisher, 2014 (Madagascar)

ataxium Bolton, 1980 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Gabon, Ghana, Guinea, Ivory Coast, Nigeria, Senegal)

atlante Cagniant, 1970 (Morocco, Tunisia)

atratulum (Schenck, 1852) (Albania, Armenia, Austria, Belarus, Belgium, Bulgaria, Channel Islands, Czech Republic, Denmark, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Norway, Poland, Republic of Moldova, Romania, Russian Federation, Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland, United States)

• Anergates friedlandi Creighton, 1934

avaratra Hita Garcia & Fisher, 2012 (Madagascar)

avium Bolton, 1980 (South Africa)

azcatltontlium Marques, Vásquez-Bolaños & Quesada, 2011 (Mexico)

B
barbigerum Bolton, 1980 (Namibia)

barryi Mathew, 1981 (India)

basum Bolton, 1977 (New Guinea)

baufra (Bolton, 1976) (Zimbabwe)

beesoni (Mukerjee, 1934) (India)

belgaense Forel, 1902 (India, Nigeria)

bellerophoni Salata & Borowiec, 2017 (Turkey)

bellicosum Bolton, 1980 (Nigeria)

bendai Hita Garcia, Fischer & Peters, 2010 (Burundi)

bequaerti Forel, 1913 (Democratic Republic of Congo, United Republic of Tanzania)

• Tetramorium humile Santschi, 1914

berbiculum Bolton, 1980 (Zimbabwe)

bessonii Forel, 1891 (Madagascar)

bevisi Arnold, 1958 (Lesotho, South Africa, Zimbabwe)

bicarinatum (Nylander, 1846) (American Samoa, Aruba, Australia, Austria, Bangladesh, Barbados, Belgium, Bhutan, Borneo, Canary Islands, Cape Verde, China, Comoros, Costa Rica, Cuba, Denmark, Dominican Republic, Ecuador, Eritrea, Fiji, French Guiana, Galapagos Islands, Germany, Greater Antilles, Grenada, Guadeloupe, Guam, Guyana, Haiti, Hawaii, Honduras, Hungary, Iberian Peninsula, India, Indonesia, Israel, Japan, Kiribati, Krakatau Islands, Lord Howe Island, Madagascar, Malaysia, Malta, Marshall Islands, Mayotte, Mexico, Micronesia (Federated States of), Montenegro, Netherlands Antilles, New Caledonia, New Guinea, New Zealand, Nicobar Island, Niue, Norfolk Island, Panama, Peru, Philippines, Puerto Rico, Republic of Korea, Samoa, Seychelles, Singapore, Solomon Islands, Spain, Sri Lanka, Sweden, Thailand, Tokelau, Tonga, Trinidad and Tobago, United Arab Emirates, United States, Vanuatu, Vietnam, Yemen)

• Myrmica cariniceps Guérin-Méneville, 1852

• Myrmica kollari Mayr, 1853

• Myrmica modesta Smith, F. Smith, F., 1860

• Myrmica reticulata Smith, F., 1862

bicolor Viehmeyer, 1914 (New Guinea)

bicolorum Vásquez-Bolaños, 2007 (Mexico)

binodis (Linnaeus, 1763)

biskrense Forel, 1904 (Algeria, Canary Islands, Croatia, Greece, Morocco, Oman, Portugal, Republic of Macedonia, Saudi Arabia, Spain, Tunisia)

• Tetramorium biskrense jarbas Cagniant, 1970

• Tetramorium caespitum oxyomma Karavaiev, 1912

boehmei Hita Garcia & Fisher, 2010 (Kenya, Uganda)

boltoni Hita Garcia, Fischer & Peters, 2010 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Gabon, Kenya, Uganda, United Republic of Tanzania)

bonibony Hita Garcia & Fisher, 2012 (Madagascar)

bothae Forel, 1910 (Lesotho, South Africa)

• Tetramorium guillarmodi Arnold, 1960

bressleri Hita Garcia & Fisher, 2014 (Madagascar)

brevicorne Bondroit, 1918 (France, Italy, Turkey)

brevidentatum (Kutter, 1932) (Indonesia)

breviscapus Wagner, Arthofer, Seifert, Muster, Steiner & Schlick-Steiner, 2017 (Bosnia and Herzegovina, Croatia, Greece)

brevispinosum (Stitz, 1910) (Cameroun, Central African Republic, Guinea, Ivory Coast, Kenya)

• Triglyphothrix nion Bernard, 1953

brigitteae Mbanyana, Robertson & Hita Garcia, 2018 (South Africa)

browni Bolton, 1980 (Ghana)

bulawayense Forel, 1913 (Kenya, South Africa, Zimbabwe)

• Tetramorium bequaerti bruni Santschi, 1917

buschingeri (Lapeva-Gjonova, 2017) (Bulgaria)

buthrum Bolton, 1980 (Central African Republic, Democratic Republic of Congo, Mozambique)

C
caespitum (Linnaeus, 1758) (Afghanistan, Albania, Armenia, Austria, Azerbaijan, Balearic Islands, Belarus, Belgium, Bulgaria, Canary Islands, Channel Islands, China, Croatia, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, France, Georgia, Germany, Gibraltar, Hungary, Iberian Peninsula, Iran, Israel, Italy, Jersey, Kazakhstan, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Malta, Monaco, Montenegro, Netherlands, Norway, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Spain, Sweden, Switzerland, Turkey, Turkmenistan, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Formica fusca Leach Leach, 1825

• Myrmica brevinodis transversinodis Enzmann, J., 1946

• Myrmica fuscula Nylander, 1846

• Myrmica modesta Foerster Foerster, 1850

• Tetramorium caespitum hammi Donisthorpe, 1915

• Tetramorium caespitum himalayanum Viehmeyer, 1914

• Tetramorium jiangxiense Wang, M. & Xiao, 1988

• Tetramorium semilaeve transbaicalense Ruzsky, 1936

caespitum barabense Ruzsky, 1925 (Russian Federation)

caespitum caespitomoravicum Kratochvíl, 1941 (Czech Republic)

caespitum japonicum Röszler, 1936 (Japan)

caespitum pallidum Stitz, 1934 (China)

caespitum typicum Ruzsky, 1902 (Kazakhstan)

caldarium (Roger, 1857) (Balearic Islands, Barbados, Bermuda, Brazil, Canary Islands, Colombia, Dominican Republic, Ecuador, Egypt, Galapagos Islands, Germany, Greater Antilles, Guadeloupe, Haiti, Iberian Peninsula, India, Ivory Coast, Kenya, Madagascar, Mauritius, Mexico, Monaco, Norfolk Island, Oman, Peru, Poland, Puerto Rico, Saint Helena, Saudi Arabia, Socotra Archipelago, Spain, United Kingdom of Great Britain and Northern Ireland, United Republic of Tanzania, United States, Yemen)

• Tetramorium minutum Donisthorpe, 1942

• Tetramorium pusillum hemisi Wheeler, W.M., 1922

calidum Forel, 1907 (Oman, Saudi Arabia, United Arab Emirates)

calinum Bolton, 1980 (Ghana, Ivory Coast)

camelliae Hita Garcia & Fisher, 2014 (Madagascar)

camerunense Mayr, 1895 (Cameroun, Gabon, Ghana, Guinea, Ivory Coast, Kenya)

candidum Bolton, 1980 (Democratic Republic of Congo, Kenya, United Republic of Tanzania)

capense Mayr, 1865 (South Africa, Zimbabwe)

• Tetramorium braunsi Forel, 1913

• Tetramorium popovici Forel, 1914

capillosum Bolton, 1980 (Cameroun, Central African Republic, Gabon, Uganda)

capitale (McAreavey, 1949) (Australia)

cardiocarenum Xu, Z. & Zheng, 1994 (China)

carinatum (Smith, F., 1859) (Indonesia, New Guinea, Philippines, Solomon Islands)

• Xiphomyrmex aruensis Karavaiev, 1935

• Xiphomyrmex costatus Emery, 1897

• Xiphomyrmex costatus deficiens Emery, 1897

• Xiphomyrmex costatus flavescens Emery, 1897

caritum (Bolton, 1986) (Uganda)

caucasicum Wagner, Arthofer, Seifert, Muster, Steiner & Schlick-Steiner, 2017 (Armenia, Azerbaijan, Georgia, Russian Federation)

cavernicola Hita Garcia & Fisher, 2015 (Madagascar)

centum Bolton, 1977 (New Guinea)

chapmani Bolton, 1977 (Philippines)

chefketi Forel, 1911 (Armenia, Azerbaijan, Bulgaria, China, Georgia, Greece, Iran, Kyrgyzstan, Republic of Macedonia, Russian Federation, Saudi Arabia, Turkey, Ukraine)

• Tetramorium caespitum sarkissiani Forel, 1911

• Tetramorium caespitum turcomanicum Santschi, 1921

• Tetramorium taurocaucasicum Arnoldi, 1968

chepocha (Bolton, 1976) (Borneo, Indonesia, Malaysia)

chloe (Santschi, 1920) (Mozambique, United Republic of Tanzania, Zimbabwe)

christiei Forel, 1902 (India)

ciliatum Bolton, 1977 (China, Thailand)

clunum Forel, 1913 (Namibia, South Africa)

cognatum Bolton, 1979 (Comoros, Madagascar)

coillum Bolton, 1979 (Madagascar)

coloreum Mayr, 1901 (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Guinea, Uganda)

• Tetramorium humerosum muscicola Bernard, 1953

concaviceps Bursakov, 1984 (Kazakhstan, Mongolia)

confusum Bolton, 1977 (Australia)

constanciae Arnold, 1917 (Central African Republic, Mozambique, United Republic of Tanzania, Zimbabwe)

• Triglyphothrix longispinosa Arnold, 1956

convexum Bolton, 1980 (Ghana)

coonoorense Forel, 1902 (India)

cordatum Sheela & Narendran, 1998 (India)

crepum Wang, M. & Wu, J., 1988 (China)

cristatum Stitz, 1910 (Angola, Democratic Republic of Congo, Ghana, Guinea, Ivory Coast, Kenya, Sudan, Togo, Uganda, United Republic of Tanzania, Zimbabwe)

• Tetramorium guineense medje Wheeler, W.M., 1922

critchleyi (Bolton, 1976) (Nigeria)

crypticum (Bolton, 1976) (Ghana, Uganda)

cuneinode Bolton, 1977 (China, Thailand)

curtulum Emery, 1895 (Borneo, China, Indonesia, Malaysia, Myanmar, Singapore)

curvispinosum Mayr, 1897 (Sri Lanka)

cyclolobium Xu, Z. & Zheng, 1994 (China)

cynicum Bolton, 1977 (New Guinea, Philippines)

D
dalek Hita Garcia & Fisher, 2014 (Madagascar)

davidi Forel, 1911 (Greece, Iran, Israel, Malta, Turkey)

decamerum (Forel, 1902) (India)

decem Forel, 1913 (Ivory Coast, Kenya, United Republic of Tanzania, Zimbabwe)

deceptum Bolton, 1977 (Australia)

dedefra (Bolton, 1976) (Angola)

degener Santschi, 1911 (Madagascar)

delagoense Forel, 1894 (Angola, Comoros, Israel, Kenya, Madagascar, Mayotte, Mozambique, South Africa, Sudan, United Republic of Tanzania)

• Tetramorium simillimum madecassum Forel, 1895

densopilosum Radchenko & Arakelian, 1990 (Armenia, Georgia, Russian Federation, Turkey)

depressiceps Menozzi, 1933 (Egypt, Iraq, Israel, Saudi Arabia)

depressum Forel, 1892 (Canary Islands)

• Tetramorium semilaeve guancha Santschi, 1929

desertorum (Forel, 1910) (Botswana, Namibia)

dichroum Santschi, 1932 (South Africa, Zambia)

difficile Bolton, 1977 (China, Nepal)

diligens (Smith, F., 1865) (New Guinea)

• Pheidole diligentissima Dalla Torre, 1892

diomandei Bolton, 1980 (Cameroun, Ivory Coast)

diomedeum Emery, 1908 (Bulgaria, China, Croatia, Greece, Italy, Malta, Montenegro, Turkey)

• Tetramorium caespitum bariensis Forel, 1911

• Tetramorium caespitum laevior Menozzi, 1936

distinctum (Bolton, 1976) (Ghana, Ivory Coast)

do Forel, 1914 (Mozambique, South Africa, United Republic of Tanzania, Zimbabwe)

dogieli Karavaiev, 1931 (Kenya, United Republic of Tanzania)

dolichosum Bolton, 1980 (Democratic Republic of Congo)

dominum Bolton, 1980 (South Africa)

doriae Emery, 1881 (Eritrea, Ethiopia, Oman, Saudi Arabia, United Arab Emirates, Yemen)

dumezi Menozzi, 1942 (Cameroun, Democratic Republic of Congo, Ghana, Kenya, Nigeria, Uganda)

• Tetramorium dumezi Bolton Bolton, 1980

duncani Mbanyana, Robertson & Hita Garcia, 2018 (Namibia, South Africa)

dunhuangense Chang & He, 2001 (China)

dysalum Bolton, 1979 (Madagascar)

dysderke Bolton, 1980 (Nigeria)

E
edouardi Forel, 1894 (Ethiopia, Ivory Coast, Kenya)

• Tetramorium kivuense Stitz, 1911

• Tetramorium tersum Santschi, 1911

• Xiphomyrmex kivuense atrinodis Santschi, 1928

eleates Forel, 1913 (Indonesia, Philippines, Singapore, Thailand)

electrum Bolton, 1979 (Madagascar)

elf Hita Garcia & Fisher, 2012 (Madagascar)

elidisum Bolton, 1980 (Ghana)

elisabethae Forel, 1904 (India)

emeryi Mayr, 1901 (South Africa, Zimbabwe)

• Tetramorium emeryi cristulatum Forel, 1913

eminii (Forel, 1894) (Angola, Ethiopia, Kenya, South Africa, Sudan, United Republic of Tanzania)

• Tetramorium marleyi akermani Arnold, 1926

• Triglyphothrix cinereus Weber, 1943

• Triglyphothrix guillodi Santschi, 1937

• Triglyphothrix guillodi mus Santschi, 1937

• Triglyphothrix marleyi Forel, 1914

• Triglyphothrix marthae Forel, 1911

enkidu Hita Garcia & Fisher, 2014 (Madagascar)

erectum Emery, 1895 (Botswana, Democratic Republic of Congo, Namibia, South Africa, Uganda)

• Tetramorium bacchus Forel, 1910

ericae Arnold, 1917 (Zimbabwe)

• Triglyphothrix pauper Santschi, 1917

etiolatum Bolton, 1977 (New Guinea)

exasperatum Emery, 1891 (Gibraltar, Morocco, Spain, Tunisia)

• Tetramorium exasperatum acutiseta Santschi, 1921

• Tetramorium parvioculum Guillem & Bensusan, 2009

• Tetramorium zahrae Santschi, 1923

exile Csösz & Radchenko, 2007 (Afghanistan)

F
fergusoni Forel, 1902 (India)

ferox Ruzsky, 1903 (Armenia, Azerbaijan, Bulgaria, China, Croatia, Czech Republic, Georgia, Greece, Hungary, Iran, Kyrgyzstan, Malta, Montenegro, Republic of Macedonia, Romania, Russian Federation, Turkmenistan)

• Tetramorium confinis Radchenko & Arakelian, 1990

• Tetramorium ferox perspicax Radchenko, 1992

• Tetramorium ferox silhavyi Kratochvíl, 1941

feroxoides Dlussky & Zabelin, 1985 (Kazakhstan, Kyrgyzstan, Russian Federation, Turkey, Turkmenistan)

• Tetramorium bursakovi Radchenko, 1992

fezzanense Bernard, 1948 (Algeria)

flabellum Bolton, 1980 (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Ghana, Ivory Coast)

flagellatum Bolton, 1977 (Borneo, Indonesia, Malaysia)

flaviceps Arnold, 1960 (Mozambique, South Africa, Zimbabwe)

• Tetramorium do mus Arnold, 1960

flavidulum Emery, 1924 (Turkey)

flavipes Emery, 1893 (China, Laos, Thailand, Vietnam)

flavithorax (Santschi, 1914) (Cameroun, Ghana, Ivory Coast)

flavum Chang & He, 2001 (China)

forte Forel, 1904 (Algeria, Balearic Islands, France, Gibraltar, Iberian Peninsula, Morocco, Portugal, Spain)

• Tetramorium caespitum hispanicum Bondroit, 1918

• Tetramorium caespitum ruginode Stitz, 1917

• Tetramorium ferox marocanum De Haro & Collingwood, 1994

• Tetramorium maura tingitanum Santschi, 1929

• Tetramorium ruginode marocana Cagniant, 1997

frenchi Forel, 1914 (South Africa, United Republic of Tanzania, Zimbabwe)

freya Hita Garcia & Fisher, 2014 (Madagascar)

frigidum Arnold, 1926 (Lesotho, South Africa)

• Tetramorium akermani drakensbergensis Arnold, 1926

fulviceps (Emery, 1897) (New Guinea)

furtivum (Arnold, 1956) (Uganda, United Republic of Tanzania, Zimbabwe)

fusciclava Consani & Zangheri, 1952 (Italy)

fuscipes (Viehmeyer, 1925) (Australia)

G
gabonense (André, 1892) (Cameroun, Central African Republic, Gabon, Uganda)

• Triglyphothrix areolata burgeoni Santschi, 1935

• Triglyphothrix areolatus Stitz, 1910

• Triglyphothrix gabonensis boulognei Forel, 1916

• Triglyphothrix gabonensis kamerunensis Santschi, 1937

• Triglyphothrix gabonensis soyauxi Forel, 1901

• Triglyphothrix mucidus Forel, 1909

galaticum Menozzi, 1936 (Turkey)

galoasanum Santschi, 1910 (Congo)

gambogecum (Donisthorpe, 1941) (China, New Guinea)

• Xiphomyrmex gambogecus flavus Donisthorpe, 1941

gazense Arnold, 1958 (Democratic Republic of Congo, Kenya, United Republic of Tanzania, Zimbabwe)

gegaimi Forel, 1916 (Cameroun, Democratic Republic of Congo, Gabon, Kenya, Uganda)

geminatum Bolton, 1980 (Cameroun, Gabon)

gestroi (Menozzi, 1933) (Uganda)

ghindanum Forel, 1910 (Eritrea, Ethiopia, Sudan)

gilgamesh Hita Garcia & Fisher, 2014 (Madagascar)

glabratum Stitz, 1923 (Namibia, South Africa)

• Tetramorium rutilum Prins, 1973

gladius Hita Garcia & Fisher, 2014 (Madagascar)

gladstonei Forel, 1913 (Benin, Mozambique, South Africa, Zimbabwe)

globulinode (Mayr, 1901) (China, Democratic Republic of Congo, South Africa, Zimbabwe)

• Rhoptromyrmex globulinodis alberti Forel, 1916

• Rhoptromyrmex globulinodis obscurus Santschi, 1932

gollum Hita Garcia & Fisher, 2014 (Madagascar)

goniommoide Poldi, 1979 (Turkey)

gracile Forel, 1894 (Ethiopia)

grandinode Santschi, 1913 (Namibia, South Africa)

• Tetramorium grandinode hopensis Forel, 1914

granulatum Bolton, 1980 (Nigeria)

grassii Emery, 1895 (Eswatini, New Zealand, South Africa)

• Tetramorium grassii laevigatum Mayr, 1901

• Tetramorium grassii simulans Santschi, 1914

• Tetramorium grazsii mayri Emery, 1924

• Tetramorium joffrei Forel, 1914

• Tetramorium joffrei algoa Arnold, 1917

guangxiense Zhou & Zheng, 1997 (China)

guineense (Bernard, 1953) (Cameroun, Canada, Central African Republic, China, Cook Islands, Cuba, Democratic Republic of Congo, Gabon, Ghana, Guinea, Ivory Coast, Liberia, New Guinea, Nigeria, Pitcairn, Senegal, Wallis and Futuna Islands)

H
hapale Bolton, 1980 (Ivory Coast)

hasinae Yamane & Jaitrong, 2011 (Thailand)

hecate Hita Garcia & Fisher, 2013 (Gabon)

hector Hita Garcia & Fisher, 2012 (Madagascar)

hippocratis Agosti & Collingwood, 1987 (Greece, Turkey)

hirsutum Collingwood & van Harten, 2005 (Saudi Arabia, Yemen)

hispidum (Wheeler, W.M., 1915) (Mexico, United States)

hobbit Hita Garcia & Fisher, 2014 (Madagascar)

hortorum Arnold, 1958 (South Africa, Zambia, Zimbabwe)

humbloti Forel, 1891 (Comoros, Kenya, Madagascar, Mayotte, Mozambique, Namibia, South Africa, United Republic of Tanzania, Zambia, Zimbabwe)

• Tetramorium humbloti pembensis Forel, 1907

• Tetramorium humblotii victoriensis Forel, 1913

hungaricum Röszler, 1935 (Austria, Bulgaria, Czech Republic, Greece, Hungary, Romania, Russian Federation, Slovenia, Turkey)

I
ibycterum Bolton, 1979 (Madagascar)

ictidum Bolton, 1980 (Cameroun, Gabon, Ghana, Uganda)

imbelle (Emery, 1915) (Eritrea, Ethiopia)

immigrans Santschi, 1927 (Argentina, Azerbaijan, Bulgaria, Chile, Turkey, United States)

impressum (Viehmeyer, 1925) (Australia)

• Xiphomyrmex viehmeyeri venustus Wheeler, W.M., 1934

impurum (Foerster, 1850) (Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Germany, Hungary, Iberian Peninsula, Italy, Montenegro, Netherlands, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, Switzerland, Turkey)

• Tetramorium caespitum penninum Santschi, 1927

• Tetramorium staerckei gregori Kratochvíl, 1944

incruentatum Arnold, 1926 (Zambia, Zimbabwe)

• Rhoptromyrmex arnoldi Santschi, 1916

indicum Forel, 1913 (Borneo, India, Indonesia, Malaysia, Sri Lanka)

indocile Santschi, 1927 (Armenia, Azerbaijan, Bulgaria, France, Hungary, Iran, Kyrgyzstan, Russian Federation, Spain, Switzerland, Ukraine)

• Tetramorium semilaeve kutteri Santschi, 1927

indosinense Wheeler, W.M., 1927 (China, Vietnam)

inerme Mayr, 1877 (Armenia, Azerbaijan, China, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Russian Federation, Turkey, Turkmenistan)

• Tetramorium caespitum picta Karavaiev, 1912

• Tetramorium caespitum plana Karavaiev, 1927

• Tetramorium caespitum reticuliventre Ruzsky, 1902

inezulae (Forel, 1914) (South Africa, United Republic of Tanzania, Zambia, Zimbabwe)

• Tetramorium hepburni Arnold, 1917

• Triglyphothrix hepburni mashonana Arnold, 1949

infraspinosum Karavaiev, 1935 (Vietnam)

infraspinum Forel, 1905 (Indonesia)

inglebyi Forel, 1902 (India)

inquilinum Ward, Brady, Fisher & Schultz, 2014 (France, Iberian Peninsula, Russian Federation, Spain, Switzerland, Turkmenistan)

• Teleutomyrmex schneideri Kutter, 1950

insolens (Smith, F., 1861) (American Samoa, Borneo, China, Fiji, Hungary, Indonesia, Laos, Malaysia, Mexico, New Caledonia, New Guinea, Niue, Philippines, Poland, Réunion, Samoa, Singapore, Solomon Islands, Sri Lanka, Sulawesi, Thailand, Tonga, United States, Vanuatu)

• Tetramorium guineense macra Emery, 1914

• Tetramorium melanogyna pallidiventre Wheeler, W.M., 1934

• Tetramorium pacificum wilsoni Mann, 1921

intermedium Hita Garcia, Fischer & Peters, 2010 (Ghana)

intextum Santschi, 1914 (Kenya, Zambia, Zimbabwe)

• Tetramorium intextum cataractae Santschi, 1916

intonsum Bolton, 1980 (Ghana, Ivory Coast, Nigeria, United Republic of Tanzania)

invictum Bolton, 1980 (Ghana, Ivory Coast)

isectum Bolton, 1979 (Madagascar)

isipingense Forel, 1914 (South Africa)

isoelectrum Hita Garcia & Fisher, 2012 (Madagascar)

J
jarawa Agavekar, Hita Garcia & Economo, 2017 (India)

jauresi Forel, 1914 (South Africa, United Republic of Tanzania, Zimbabwe)

• Tetramorium latens Arnold, 1948

jedi Hita Garcia & Fisher, 2012 (Madagascar)

jejunum Arnold, 1926 (Zimbabwe)

jizani Collingwood, 1985 (Saudi Arabia)

jordani Santschi, 1937 (Namibia, South Africa)

• Tetramorium aspinatum Prins, 1973

juba Collingwood, 1985 (Algeria, Saudi Arabia, United Arab Emirates)

judas Wheeler, W.M. & Mann, 1916 (Israel)

jugatum Bolton, 1980 (Angola, Cameroun, Ghana, Ivory Coast, Nigeria)

K
kabulistanicum Pisarski, 1967 (Afghanistan, Turkmenistan)

kakamega Hita Garcia, Fischer & Peters, 2010 (Kenya)

kali Hita Garcia & Fisher, 2012 (Madagascar)

karthala Hita Garcia & Fisher, 2014 (Madagascar)

katypum (Bolton, 1976) (Philippines)

kelleri Forel, 1887 (Madagascar, Mayotte)

kephalosi Salata & Borowiec, 2017 (Croatia, Greece)

keralense Sheela & Narendran, 1998 (India)

kestrum Bolton, 1980 (Angola, Democratic Republic of Congo, Kenya, Sudan, Uganda)

kheperra (Bolton, 1976) (Borneo, Brunei Darussalam, Cambodia, China, India, Indonesia, Malaysia, Philippines, Singapore, Vietnam)

khnum Bolton, 1977 (China, Philippines)

khyarum Bolton, 1980 (Botswana, Democratic Republic of Congo, Ivory Coast, Namibia, Nigeria, Saudi Arabia, South Africa, Yemen)

kieti Roncin, 2002 (Vietnam)

kisilkumense Dlussky, 1990 (Turkmenistan)

kraepelini Forel, 1905 (Brunei Darussalam, China, Indonesia, Japan, Philippines, Taiwan, Vietnam)

• Tetramorium mixtum amia Forel, 1912

• Tetramorium yanoi Santschi, 1937

krishnani Agavekar, Hita Garcia & Economo, 2017 (India)

krynitum Bolton, 1980 (Namibia)

kutteri (Tinaut, 1990) (Iberian Peninsula, Spain)

kydelphon Bolton, 1979 (New Guinea)

L
laevithorax Emery, 1895 (Kenya, Namibia, South Africa, Sudan, Uganda)

• Tetramorium jeanae Weber, 1943

lanuginosum Mayr, 1870 (American Samoa, Aruba, Australia, Bangladesh, Barbados, Borneo, Cambodia, China, Comoros, Dominican Republic, Ecuador, Fiji, Galapagos Islands, Greater Antilles, Guam, Iberian Peninsula, India, Indonesia, Israel, Japan, Kiribati, Krakatau Islands, Laos, Madagascar, Malaysia, Malta, Marshall Islands, Mauritius, Mayotte, Mexico, Micronesia (Federated States of), Myanmar, Netherlands Antilles, New Guinea, Nicobar Island, Oman, Philippines, Puerto Rico, Réunion, Samoa, Saudi Arabia, Seychelles, Singapore, Socotra Archipelago, Solomon Islands, Spain, Sri Lanka, Thailand, Tokelau, Tonga, United States, Vietnam, Wallis and Futuna Islands, Yemen)

• Tetramorium obesum striatidens Emery, 1889

• Triglyphothrix ceramensis Stitz, 1912

• Triglyphothrix mauricei Donisthorpe, 1946

• Triglyphothrix striatidens australis Forel, 1902

• Triglyphothrix striatidens felix Forel, 1912

• Triglyphothrix striatidens flavescens Wheeler, W.M., 1929

• Triglyphothrix striatidens laevidens Forel, 1900

• Triglyphothrix striatidens orissana Forel, 1902

• Triglyphothrix tricolor Donisthorpe, 1948

laparum Bolton, 1977 (Borneo, China, Indonesia, Malaysia, Philippines)

laticephalum Bolton, 1977 (Australia)

latinode Collingwood & Agosti, 1996 (Saudi Arabia, Saudi Arabia, Yemen)

latreillei Forel, 1895 (Madagascar)

legone Bolton, 1980 (Benin, Ghana, Nigeria)

lerouxi Mbanyana, Robertson & Hita Garcia, 2018 (South Africa)

lobulicorne Santschi, 1916 (South Africa, Zimbabwe)

longicorne Forel, 1907 (Burkina Faso, Cameroun, Kenya, Mozambique, Nigeria, South Africa, United Republic of Tanzania, Zimbabwe)

longoi Forel, 1915 (South Africa)

lucayanum Wheeler, W.M., 1905 (Bahamas, Brazil, Cameroun, Cayman Islands, Cuba, Democratic Republic of Congo, Dominican Republic, Ecuador, Equatorial Guinea, Gabon, Gambia, Ghana, Greater Antilles, Guinea, Ireland, Italy, Ivory Coast, Kenya, Liberia, Mali, Mexico, Netherlands Antilles, Nigeria, Puerto Rico, Sierra Leone, Uganda)

• Tetramorium camerunense waelbroeki Forel, 1909

• Tetramorium lucayanum sexdens Forel, 1915

• Tetramorium rectinodis Menozzi, 1942

lucidulum Menozzi, 1933 (Israel, Montenegro, Republic of Macedonia, Syria, Turkey)

lucyae Sorger, 2011 (Borneo, Indonesia, Malaysia)

luteipes Santschi, 1910 (Congo)

luteolum Arnold, 1926 (Namibia, South Africa, Zimbabwe)

M
mackae Media:Hita Garcia & Fisher 2012a.pdf|Hita Garcia & Fisher, 2012 (Madagascar)

magnificum Bolton, 1980 (Cameroun, Ivory Coast)

mahafaly Hita Garcia & Fisher, 2011 (Madagascar)

mai Wang, M., 1993 (China)

malabarense Sheela & Narendran, 1998 (India)

malagasy Hita Garcia & Fisher, 2012 (Madagascar)

mallenseana Hita Garcia & Fisher, 2012 (Madagascar)

manni Bolton, 1985 (Fiji)

• Tetramorium manni Dlussky Dlussky, 1994

• Triglyphothrix pacifica Mann, 1921

manobo (Calilung, 2000) (Philippines)

marginatum Forel, 1895 (Madagascar)

margueriteae Mbanyana, Robertson & Hita Garcia, 2018 (South Africa)

marojejy Hita Garcia & Fisher, 2012 (Madagascar)

mars Hita Garcia & Fisher, 2014 (Madagascar)

matopoense Arnold, 1926 (South Africa, Zimbabwe)

maurum Santschi, 1918 (Tunisia)

mayri (Forel, 1912) (India, Sri Lanka)

megalops Bolton, 1977 (Australia)

meghalayense Bharti, 2011 (India)

• Tetramorium browni Tiwari Tiwari, 2000

melanogyna Mann, 1919 (Solomon Islands)

melleum Emery, 1897 (Australia, New Guinea)

• Ireneella papuensis Donisthorpe, 1941

menkaura (Bolton, 1976) (Ghana)

meressei Forel, 1916 (Democratic Republic of Congo, Ghana, Sudan)

meridionale Emery, 1870 (Balearic Islands, Greece, Iberian Peninsula, Israel, Italy, Malta, Spain)

merina Hita Garcia & Fisher, 2014 (Madagascar)

meshena (Bolton, 1976) (Borneo, Indonesia, Malaysia, Singapore)

metactum Bolton, 1980 (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Kenya, Uganda)

mexicanum Bolton, 1979 (Mexico)

microgyna Santschi, 1918 (Angola, South Africa, Zimbabwe)

microps (Mayr, 1901) (South Africa, Zimbabwe)

• Tetramorium auropunctatus bulawayensis Arnold, 1917

• Triglyphothrix auropunctata rhodesiana Forel, 1913

• Triglyphothrix auropunctatus Forel, 1910

• Triglyphothrix auropunctatus fusciventris Forel, 1913

• Triglyphothrix auropunctatus pallens Forel, 1910

minimum (Bolton, 1976) (Ghana, Ivory Coast)

minusculum (Santschi, 1914) (Cameroun, Ghana, Ivory Coast, Sudan)

• Xiphomyrmex minusculus amen Weber, 1943

miserabile Santschi, 1918 (Kenya, United Republic of Tanzania)

mixtum Forel, 1902 (India)

mkomazi Hita Garcia, Fischer & Peters, 2010 (United Republic of Tanzania)

monardi (Santschi, 1937) (Angola)

monticola Hita Garcia & Fisher, 2014 (Madagascar)

moravicum Kratochvíl, 1941 (Austria, Azerbaijan, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Iran, Italy, Montenegro, Poland, Romania, Russian Federation, Serbia, Slovenia, Turkey, Ukraine)

• Tetramorium rhenanum Schulz, 1996

mossamedense Forel, 1901 (Angola, Botswana, Namibia, South Africa, United Republic of Tanzania, Zimbabwe)

• Tetramorium caespitum schultzei Forel, 1910

mpala Hita Garcia & Fischer, 2014 (Kenya)

muralti Forel, 1910 (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Ghana, Ivory Coast, Uganda)

• Xiphomyrmex muralti trilineata Santschi, 1919

muscorum Arnold, 1926 (Ivory Coast, Mozambique, Uganda, United Republic of Tanzania)

mutatum Bolton, 1985 (Solomon Islands)

• Triglyphothrix pulchella Mann, 1919

myops Bolton, 1977 (India)

myrmidon Hita Garcia & Fisher, 2014 (Madagascar)

N
nacta (Bolton, 1976) (Thailand)

naganum Bolton, 1979 (Comoros, Madagascar)

nama Hawkes, 2020 (South Africa)

nassonowii Forel, 1892 (Madagascar)

nautarum Santschi, 1918 (Equatorial Guinea)

navum Bolton, 1977 (New Guinea)

nazgul Hita Garcia & Fisher, 2012 (Madagascar)

nefassitense Forel, 1910 (Eritrea, Ethiopia)

nify Hita Garcia & Fisher, 2012 (Madagascar)

nigrum Forel, 1907 (Kenya, South Africa, Sudan, United Republic of Tanzania)

• Tetramorium brevis Weber, 1943

nipponense Wheeler, W.M., 1928 (China, Japan, Thailand, Vietnam)

nitidissimum Pisarski, 1967 (China, Turkmenistan)

nodiferum (Emery, 1901) (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Ghana, Guinea, Kenya, Uganda)

noeli Hita Garcia & Fisher, 2012 (Madagascar)

noratum Bolton, 1977 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

norvigi Hita Garcia & Fisher, 2012 (Madagascar)

nosybe Hita Garcia & Fisher, 2012 (Madagascar)

notiale Bolton, 1980 (Botswana, Democratic Republic of Congo, Eswatini, Malawi, Mozambique, Namibia, South Africa, Zambia, Zimbabwe)

• Tetramorium guineense striatum Arnold, 1917

notomelanum Vásquez-Bolaños, Castaño-Meneses & Guzmán-Mendoza, 2011 (Mexico)

nube Weber, 1943 (Ghana, Sudan)

nursei Bingham, 1903 (China, India, Iran, Pakistan)

O
obesum André, 1887 (Bangladesh, India, Sri Lanka)

obiwan Hita Garcia & Fisher, 2014 (Madagascar)

obtusidens Viehmeyer, 1916 (Borneo, Brunei Darussalam, China, Indonesia, Malaysia, Philippines, Singapore)

occidentale (Santschi, 1916) (Benin, Cameroun, Central African Republic, Ghana, Ivory Coast, Kenya, Togo, Uganda)

• Xiphomyrmex insularis Menozzi, 1924

ochrothorax Chang & He, 2001 (China)

ocothrum Bolton, 1979 (Borneo, Indonesia, Malaysia)

oculatum Forel, 1913 (Angola, Botswana, Mozambique, South Africa, Zimbabwe)

olana Hita Garcia & Fisher, 2012 (Madagascar)

opacum (Emery, 1909) (Cameroun, Guinea, Kenya)

• Rhoptromyrmex opacus esta Forel, 1909

• Rhoptromyrmex opacus laeviceps Santschi, 1916

• Rhoptromyrmex opacus monodi Bernard, 1953

orc Hita Garcia & Fisher, 2012 (Madagascar)

orientale Forel, 1895 (Madagascar)

ornatum Emery, 1897 (Australia, New Guinea)

• Tetramorium ornatum obscurius Forel, 1901

osiris (Bolton, 1976) (Central African Republic, Ghana)

P
pacificum Mayr, 1870 (Australia, Borneo, Brunei Darussalam, China, Cook Islands, Fiji, French Polynesia, India, Indonesia, Krakatau Islands, Malaysia, Micronesia (Federated States of), New Caledonia, New Guinea, Niue, Palau, Philippines, Samoa, Singapore, Solomon Islands, Sri Lanka, Thailand, Tonga, United States, Vanuatu, Wallis and Futuna Islands)

• Tetramorium pacificum subscabrum Emery, 1893

palaense Bolton, 1979 (Borneo, Indonesia, Malaysia, Thailand)

parasiticum Bolton, 1980 (South Africa)

parvispinum (Emery, 1893) (Borneo, China, Indonesia, Malaysia, Philippines, Taiwan, Thailand)

• Triglyphothrix parvispina formosae Forel, 1912

parvum Bolton, 1977 (Borneo, Indonesia, Malaysia, Thailand)

pauper Forel, 1907 (Kenya, Socotra Archipelago, South Africa, United Republic of Tanzania, Zimbabwe)

pelagium Poldi, 1995 (Italy)

peringueyi Arnold, 1926 (Botswana, Namibia, South Africa)

perlongum Santschi, 1923 (Angola)

persignatum Bolton, 1995 (Israel)

• Tetramorium signatum Menozzi Menozzi, 1933

petersi Forel, 1910 (Namibia)

petiolatum Sheela & Narendran, 1998 (India)

peutli Forel, 1916 (Angola, Cameroun, Democratic Republic of Congo, Gabon, Ghana, Ivory Coast, Uganda)

phasias Forel, 1914 (Angola, Democratic Republic of Congo, Malawi, Mozambique, South Africa, United Republic of Tanzania, Zambia, Zimbabwe)

• Tetramorium guineense hertigi Santschi, 1937

philippwagneri Hita Garcia, Fischer & Peters, 2010 (Cameroun, Democratic Republic of Congo, Gabon, United Republic of Tanzania, Zambia)

pialtum Bolton, 1980 (Central African Republic, Ghana, Kenya)

pilosum Emery, 1893 (India, Sri Lanka)

pinnipilum Bolton, 1980 (Angola, Kenya)

pisarskii Radchenko & Scupola, 2015 (Afghanistan)

placidum Bolton, 1979 (Mexico)

platynode Bolton, 1980 (South Africa)

pleganon Bolton, 1979 (Madagascar)

plesiarum Bolton, 1979 (Madagascar)

plumosum Bolton, 1980 (Eswatini)

pnyxis (Bolton, 1976) (Malaysia)

pogonion Bolton, 1980 (Namibia, South Africa)

politum Emery, 1897 (New Guinea)

polymorphum Yamane & Jaitrong, 2011 (China, Laos, Thailand)

popell Hita Garcia & Fisher, 2012 (Madagascar)

postpetiolatum Santschi, 1919 (Democratic Republic of Congo, Gabon, Uganda)

poweri Forel, 1914 (South Africa)

praetextum Bolton, 1980 (Namibia)

proximum Bolton, 1979 (Madagascar)

pseudogladius Hita Garcia & Fisher, 2014 (Madagascar)

psymanum Bolton, 1980 (Ghana)

pulchellum Emery, 1897 (New Guinea, Singapore)

pulcherrimum (Donisthorpe, 1945) (Central African Republic, Uganda)

pullulum Santschi, 1924 (Angola, Cameroun, Central African Republic, Democratic Republic of Congo, Kenya, Sudan, Uganda, Zambia)

• Tetramorium fernandensis Menozzi, 1942

• Xiphomyrmex uelensis Santschi, 1935

punctatum Santschi, 1927 (Greece, Iran, Italy)

• Tetramorium semilaeve liparaeum Santschi, 1927

punctiventre Emery, 1887 (New Guinea)

punicum (Smith, F., 1861) (Azerbaijan, Canary Islands, Georgia, Greece, Iran, Israel, Turkey)

punicum cyrenaicum Menozzi, 1927 (Libya)

pusillum Emery, 1895 (South Africa)

• Tetramorium caespitum ladismithensis Forel, 1913

• Tetramorium pusillum tablensis Forel, 1914

pylacum Bolton, 1980 (Cameroun, Central African Republic, Ghana, Ivory Coast)

pyrenaeicum Röszler, 1937 (Andorra, Spain)

Q
quadridentatum Stitz, 1910 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Equatorial Guinea, Gabon, Ghana, Kenya, Nigeria, Uganda)

• Tetramorium commodum Santschi, 1924

qualarum Bolton, 1980 (Gabon, Ghana)

quasirum Bolton, 1979 (Madagascar)

R
rala Hita Garcia & Fisher, 2014 (Madagascar)

ranarum Forel, 1895 (Madagascar)

raptor Hita Garcia, 2014 (Cameroun, Nigeria)

rawlinsoni (Taylor, 1992) (Indonesia, Krakatau Islands)

regulare Bolton, 1980 (South Africa)

rekhefe Bolton, 1979 (Indonesia, Timor-Leste)

renae Hita Garcia, Fischer & Peters, 2010 (São Tomé & Principe)

repentinum Arnold, 1926 (South Africa, Zambia, Zimbabwe)

repletum Wang, M. & Xiao, 1988 (China)

reptana (Bolton, 1976) (Uganda)

reticuligerum Bursakov, 1984 (Kazakhstan, Turkmenistan)

rhetidum Bolton, 1980 (Cameroun, Democratic Republic of Congo, Gabon, Ghana, Ivory Coast, South Africa, United Republic of Tanzania)

rhodium Emery, 1924 (Cyprus, Greece, Turkey)

rigidum Bolton, 1977 (New Guinea)

rimytyum Bolton, 1980 (Democratic Republic of Congo, Ghana, Senegal)

rinatum Bolton, 1977 (Philippines)

robertsoni Hita Garcia, Fischer & Peters, 2010 (United Republic of Tanzania)

robitika Hita Garcia & Fisher, 2012 (Madagascar)

rogatum Bolton, 1980 (Angola)

rossi (Bolton, 1976) (India)

rothschildi (Forel, 1907) (Ethiopia, Kenya)

rotundatum (Santschi, 1924) (Democratic Republic of Congo, Gabon)

rubrum Hita Garcia, Fischer & Peters, 2010 (Cameroun)

rufescens Stitz, 1923 (Mozambique, Namibia, South Africa)

rugigaster Bolton, 1977 (India)

rumo Hita Garcia & Fisher, 2014 (Madagascar)

ryanphelanae Hita Garcia & Fisher, 2012 (Madagascar)

S
sabatinellii Radchenko & Scupola, 2015 (Jordan)

sabatra Hita Garcia & Fisher, 2012 (Madagascar)

sada Hita Garcia & Fisher, 2012 (Madagascar)

saginatum Bolton, 1980 (Angola, Ivory Coast, Uganda)

sahlbergi Finzi, 1936 (China, Egypt, Greece, Israel)

salomo Mann, 1919 (Solomon Islands)

salvatum Forel, 1902 (India, Pakistan)

salwae Mohamed, Zalat, Fadl, Gadalla & Sharaf, 2001 (Egypt)

sanetrai Schulz & Csösz, 2007 (Italy)

sargina Hita Garcia & Fisher, 2012 (Madagascar)

saudiae (Collingwood & Agosti, 1996) (Saudi Arabia)

saudicum Sharaf, 2013 (Saudi Arabia)

scabrosum (Smith, F., 1859) (Indonesia, New Guinea)

• Tetramorium papuanum Emery, 1887

scabrum Mayr, 1879 (Borneo, Brunei Darussalam, China, India, Indonesia, Malaysia, Singapore)

schaufussii Forel, 1891 (Madagascar)

schmidti Forel, 1904 (Greece, Israel, Republic of Macedonia, Turkey)

schmitzi (Forel, 1910) (Israel)

schneideri Emery, 1898 (Afghanistan, China, Iran, Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan)

schoutedeni Santschi, 1924 (Democratic Republic of Congo)

sculptatum Bolton, 1977 (New Guinea)

scutum Hita Garcia & Fisher, 2014 (Madagascar)

scytalum Bolton, 1979 (Comoros, Madagascar, Mayotte)

securis Roncin, 2002 (Vietnam)

seiferti (Kiran & Karaman, 2017) (Turkey)

semilaeve André, 1883 (Azerbaijan, France, Iberian Peninsula, Italy, Malta, Portugal, Spain)

• Tetramorium banyulense Bernard, 1983

• Tetramorium caespitum tenuicornis Emery, 1925

• Tetramorium semilaeve siciliense Santschi, 1927

semilaeve fortunatarum Barquin Diez, 1981 (Canary Islands)

semilaeve hoggarense Santschi, 1929 (Algeria)

semilaeve italicum Menozzi, 1932 (Italy)

semilaeve jugurtha Menozzi, 1934 (Morocco, Tunisia)

• Tetramorium semilaeve gaetulum Santschi, 1936

semireticulatum Arnold, 1917 (Zimbabwe)

• Tetramorium semireticulatum politum Arnold, 1948

seneb Bolton, 1977 (Borneo, Indonesia, Malaysia)

sentosum Sheela & Narendran, 1998 (India)

sepositum Santschi, 1918 (Mozambique, Zimbabwe)

sepultum Bolton, 1980 (Eswatini, Mozambique)

sericeiventre Emery, 1877 (Algeria, Angola, Botswana, Burkina Faso, Cameroun, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Ivory Coast, Kenya, Lesotho, Liberia, Libya, Madagascar, Malawi, Mali, Mozambique, Namibia, Oman, Saudi Arabia, Senegal, Seychelles, South Africa, Sudan, Tunisia, United Arab Emirates, United Republic of Tanzania, Zimbabwe)

• Atopula hortensis Bernard, 1948

• Tetramorium blochmanni calvum Stitz, 1923

• Tetramorium blochmanni continentis Forel, 1910

• Tetramorium blochmanni nigriventre Stitz, 1910

• Tetramorium blochmannii Forel, 1887

• Tetramorium blochmannii montanum Forel, 1891

• Tetramorium neuvillei Forel, 1907

• Tetramorium quadrispinosum Emery, 1886

• Tetramorium quadrispinosum angolense Santschi, 1930

• Tetramorium quadrispinosum beirae Arnold, 1926

• Tetramorium quadrispinosum elegans Santschi, 1918

• Tetramorium quadrispinosum eudoxia Santschi, 1918

• Tetramorium quadrispinosum otaviensis Arnold, 1926

• Tetramorium sericeiventre arenarium Santschi, 1918

• Tetramorium sericeiventre bipartita Santschi, 1918

• Tetramorium sericeiventre cinnamomeum Santschi, 1918

• Tetramorium sericeiventre debile Forel, 1894

• Tetramorium sericeiventre femoratum Emery, 1895

• Tetramorium sericeiventre gamaii Santschi, 1918

• Tetramorium sericeiventre hori Santschi, 1918

• Tetramorium sericeiventre inversa Santschi, 1910

• Tetramorium sericeiventre jasonis Santschi, 1918

• Tetramorium sericeiventre munda Santschi, 1918

• Tetramorium sericeiventre repertum Santschi, 1926

• Tetramorium sericeiventre vascoi Santschi, 1918

• Tetramorium sericeiventre vividum Santschi, 1926

sericeum Arnold, 1926 (Zimbabwe)

setigerum Mayr, 1901 (Democratic Republic of Congo, Gabon, Kenya, Mozambique, South Africa, Sudan, Uganda, United Republic of Tanzania, Zimbabwe)

• Tetramorium setigerum anteversa Santschi, 1921

• Tetramorium setigerum quaerens Forel, 1914

setuliferum Emery, 1895 (Angola, Botswana, Democratic Republic of Congo, Eswatini, Lesotho, Malawi, Mozambique, Namibia, South Africa, United Republic of Tanzania, Zambia, Zimbabwe)

• Tetramorium setuliferum cucalense Santschi, 1911

• Tetramorium setuliferum triptolemus Arnold, 1917

severini (Emery, 1895) (Madagascar)

shamshir Hita Garcia & Fisher, 2012 (Madagascar)

shensiense Bolton, 1977 (China)

shilohense Forel, 1913 (Kenya, Malawi, Zambia, Zimbabwe)

shirlae Sharaf, 2007 (Egypt, Saudi Arabia)

shivalikense Bharti & Kumar, 2012 (India)

sibiricum Seifert, 2021 (Russian Federation)

sigillum Bolton, 1980 (Ivory Coast)

signatum Emery, 1895 (Angola, Israel, Namibia, South Africa)

• Tetramorium solidum grootensis Forel, 1913

• Tetramorium solidum lugubre Forel, 1910

• Tetramorium solidum tuckeri Arnold, 1926

sikorae Forel, 1892 (Madagascar, Réunion)

• Xiphomyrmex latior Santschi, 1926

silvicola Hita Garcia & Fisher, 2012 (Madagascar)

simillimum (Smith, F., 1851) (Australia, Bangladesh, Barbados, Bermuda, Borneo, Brazil, Cameroun, Canada, China, Comoros, Cook Islands, Costa Rica, Cuba, Denmark, Dominican Republic, Ecuador, Eritrea, Estonia, Ethiopia, Fiji, Galapagos Islands, Greater Antilles, Grenada, Guadeloupe, Guam, Guinea, Haiti, Hawaii, India, Indonesia, Israel, Ivory Coast, Japan, Kenya, Kiribati, Krakatau Islands, Madagascar, Malawi, Malaysia, Marshall Islands, Mauritius, Mayotte, Mexico, Micronesia (Federated States of), Mozambique, New Caledonia, New Guinea, Niue, Norfolk Island, Northern Mariana Islands, Palau, Paraguay, Philippines, Puerto Rico, Réunion, Samoa, Seychelles, Socotra Archipelago, Solomon Islands, Sri Lanka, Thailand, Tokelau, Tonga, Uganda, United Kingdom of Great Britain and Northern Ireland, United Republic of Tanzania, United States, Vanuatu, Wallis and Futuna Islands, Yemen, Zambia)

• Myrmica parallela Smith, F., 1859

• Tetramorium pusillum bantouana Santschi, 1910

• Tetramorium pusillum exoleta Santschi, 1914

• Tetramorium pygmaeum Emery, 1877

• Tetramorium simillimum denticulatum Forel, 1902

• Tetramorium simillimum insulare Santschi, 1928

• Tetramorium simillimum opacior Forel, 1913

• Wasmannia auropunctata brevispinosa Borgmeier, 1928

simulator Arnold, 1917 (South Africa, Zimbabwe)

singletonae Hita Garcia & Fisher, 2012 (Madagascar)

sitefrum Bolton, 1980 (Ghana)

sjostedti Forel, 1915 (Australia)

smaug Hita Garcia & Fisher, 2012 (Madagascar)

smithi Mayr, 1879 (Bangladesh, Borneo, China, Guam, India, Indonesia, Japan, Laos, Malaysia, Marshall Islands, Niue, Pakistan, Philippines, Singapore, Sri Lanka, Thailand, Vietnam)

• Tetramorium simillimum laevinode Forel, 1902

• Tetramorium smithi kanariense Forel, 1902

snellingi Hita Garcia, Fischer & Peters, 2010 (Central African Republic, Kenya, Uganda)

solidum Emery, 1886 (Botswana, Namibia, South Africa)

solomonense Ward, Brady, Fisher & Schultz, 2014 (Solomon Islands)

• Triglyphothrix mayri Mann, 1919

somniculosum Arnold, 1926 (Mozambique)

spininode Bolton, 1977 (Australia)

spinosum (Pergande, 1896) (Mexico, United States)

• Tetramorium wheeleri Forel, 1901

• Xiphomyrmex spinosus insons Wheeler, W.M., 1915

splendens Ruzsky, 1902 (Croatia, Greece, Republic of Macedonia, Russian Federation, Turkey)

splendidior (Viehmeyer, 1925) (Australia)

squaminode Santschi, 1911 (Kenya, Saudi Arabia, South Africa, United Republic of Tanzania, Zimbabwe)

staerckei Kratochvíl, 1944 (Austria, Bulgaria, Czech Republic, Hungary, Kyrgyzstan, Republic of Macedonia, Romania, Russian Federation, Serbia, Slovakia, Turkey)

steinheili Forel, 1892 (Madagascar)

striabdomen Chang & He, 2001 (China)

striativentre Mayr, 1877 (Afghanistan, China, Iran, Israel, Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan)

strictum Bolton, 1977 (Australia)

striolatum Viehmeyer, 1914 (Australia)

subcoecum Forel, 1907 (Botswana, Eritrea, Ethiopia, Kenya, Namibia, South Africa, United Republic of Tanzania, Zimbabwe)

• Tetramorium subcoecum inscia Forel, 1913

sudanense (Weber, 1943) (Sudan)

sulcinode Santschi, 1927 (Afghanistan, Azerbaijan, Kazakhstan, Kyrgyzstan, Pakistan, Russian Federation, Turkey, Turkmenistan, Turkmenistan)

• Tetramorium karakalense Dlussky & Zabelin, 1985

surrogatum Bolton, 1985 (Eritrea, Ethiopia)

• Triglyphothrix silvestrii Emery, 1915

susannae Hita Garcia, Fischer & Peters, 2010 (Central African Republic, Ghana)

syriacum Emery, 1924 (Georgia, Israel, Saudi Arabia, Syria, Turkey)

T
tabarum Bolton, 1980 (Central African Republic, Democratic Republic of Congo)

talpa (Bolton, 1976) (Democratic Republic of Congo)

tanaense Hita Garcia, Fischer & Peters, 2010 (Kenya, United Republic of Tanzania)

tanakai Bolton, 1977 (Japan)

tantillum Bolton, 1979 (Madagascar)

taueret Bolton, 1995 (Egypt, Israel)

• Tetramorium caespitum debilis Emery, 1924

taylori Bolton, 1985 (Australia)

• Tetramorium australe Bolton, 1977

tenebrosum Arnold, 1926 (Kenya, Mozambique)

tenuicrine (Emery, 1914) (Fiji, New Caledonia)

tenuinode Hita Garcia & Fisher, 2014 (Madagascar)

termitobium Emery, 1908 (Democratic Republic of Congo, Gabon, Ivory Coast, Uganda)

thalidum Bolton, 1977 (Australia)

thoth (Bolton, 1976) (South Africa)

titus Forel, 1910 (South Africa)

tonganum Mayr, 1870 (Borneo, Brunei Darussalam, China, Cook Islands, Fiji, French Polynesia, Guam, Indonesia, Japan, Malaysia, Marshall Islands, Micronesia (Federated States of), New Caledonia, New Guinea, Niue, Palau, Philippines, Samoa, Singapore, Solomon Islands, Sri Lanka, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Tetramorium magitae Forel, 1911

tortuosum Roger, 1863 (India, Philippines, Sri Lanka)

• Tetramorium tortuosum bellii Forel, 1902

• Tetramorium tortuosum ethica Forel, 1911

tosii Emery, 1899 (Madagascar)

traegaordhi Santschi, 1914 (South Africa)

trafo Hita Garcia & Fisher, 2012 (Madagascar)

transformans Santschi, 1914 (Kenya, Socotra Archipelago, Yemen)

transversarium Roger, 1863 (Sri Lanka)

transversinode (Mayr, 1901) (Cameroun, Kenya, South Africa)

• Rhoptromyrmex steini Forel, 1913

• Rhoptromyrmex transversinodis pretoriae Arnold, 1926

triangulatum Bharti & Kumar, 2012 (India)

tricarinatum Viehmeyer, 1914 (New Guinea)

• Xiphomyrmex tricolor Donisthorpe, 1949

trimeni (Emery, 1895) (South Africa, United Republic of Tanzania)

trirugosum Hita Garcia, Fischer & Peters, 2010 (Cameroun)

tsingy Hita Garcia & Fisher, 2012 (Madagascar)

tsushimae Emery, 1925 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation, United States)

• Tetramorium caespitum jacoti Wheeler, W.M., 1927

turneri Forel, 1902 (Australia)

tychadion Bolton, 1980 (United Republic of Tanzania)

tylinum Bolton, 1977 (Borneo, Indonesia, Malaysia)

typhlops Bolton, 1980 (Ivory Coast)

tyrion Hita Garcia & Fisher, 2012 (Madagascar)

U
ubangense Santschi, 1937 (Democratic Republic of Congo, Uganda)

uelense Santschi, 1923 (Congo, Democratic Republic of Congo, Ghana, Guinea, Nigeria)

• Decamorium decem nimba Bernard, 1953

ultor Forel, 1913 (Kenya, Mozambique, United Republic of Tanzania, Zambia, Zimbabwe)

umtaliense Arnold, 1926 (South Africa, Zimbabwe)

undatium Chang & He, 2001 (China)

unicum Bolton, 1980 (Cameroun, Democratic Republic of Congo, Uganda)

urbanii Bolton, 1977 (Bhutan, India)

V
validiusculum Emery, 1897 (Australia, New Guinea)

• Tetramorium longicarinum Donisthorpe, 1941

valky Hita Garcia & Fisher, 2012 (Madagascar)

vandalum Bolton, 1977 (New Guinea)

venator Hita Garcia, 2014 (Cameroun, Central African Republic, Democratic Republic of Congo, Gabon, Ghana, Kenya, Liberia, Uganda, United Republic of Tanzania)

vernicosum Radchenko, 1992 (Tajikistan)

versiculum Bolton, 1980 (Benin, Ghana, Ivory Coast)

vertigum Bolton, 1977 (Indonesia, Philippines, Sulawesi)

vexator Arnold, 1926 (South Africa)

viehmeyeri Forel, 1907 (Australia)

viticola Weber, 1943 (Central African Republic, Gabon, Kenya, Sudan, Uganda)

voasary Hita Garcia & Fisher, 2012 (Madagascar)

vohitra Hita Garcia & Fisher, 2012 (Madagascar)

vombis (Bolton, 1976) (Solomon Islands)

vony Hita Garcia & Fisher, 2012 (Madagascar)

W
wadje Bolton, 1980 (Ghana, Nigeria)

wagneri Viehmeyer, 1914 (New Guinea)

walshi (Forel, 1890) (Bangladesh, China, India, Philippines, Singapore, Sri Lanka, Thailand)

• Triglyphothrix musculus Forel, 1902

• Triglyphothrix walshi spuria Forel, 1912

wardi Hita Garcia & Fisher, 2012 (Madagascar)

warreni Arnold, 1926 (South Africa)

weitzeckeri Emery, 1895 (Eritrea, Eswatini, Ethiopia, Kenya, Mozambique, Namibia, South Africa, Sudan, United Republic of Tanzania, Zambia, Zimbabwe)

• Tetramorium ebeninum Arnold, 1926

• Tetramorium escherichi Forel, 1910

• Xiphomyrmex weitzeckeri edithae Weber, 1943

• Xiphomyrmex weitzeckeri nigellus Santschi, 1932

wroughtonii (Forel, 1902) (Australia, Borneo, Brunei Darussalam, China, India, Indonesia, Krakatau Islands, Malaysia, New Guinea, Philippines, Taiwan, Thailand, Vietnam)

• Rhoptromyrmex rothneyi intermedia Forel, 1913

• Rhoptromyrmex rothneyi leno Viehmeyer, 1914

• Rhoptromyrmex rothneyi sumatrensis Forel, 1913

• Rhoptromyrmex rothneyi taivanensis Wheeler, W.M., 1930

• Rhoptromyrmex wroughtonii rothneyi Forel, 1902

X
xanthogaster Santschi, 1911 (Madagascar)

xuthum Bolton, 1980 (Ghana)

Y
yammer Hita Garcia & Fisher, 2012 (Madagascar)

yarthiellum (Bolton, 1976) (Zambia)

yemene Collingwood & Agosti, 1996 (United Arab Emirates, Yemen)

yerburyi Forel, 1902 (India, Sri Lanka)

youngi Bolton, 1980 (Angola, Ghana)

yulongense Xu, Z. & Zheng, 1994 (China)

Z
zambezium Santschi, 1939 (Ivory Coast, Kenya, Mozambique, Zambia, Zimbabwe)

zapyrum Bolton, 1980 (Burkina Faso, Cameroun, Gabon, Ghana, Ivory Coast, Kenya, Liberia, Nigeria)

zenatum Bolton, 1979 (Madagascar)

zonacaciae (Weber, 1943) (Kenya, Sudan)

zypidum Bolton, 1977 (Philippines)

=Fossil Species=

K
kulickae Radchenko & Dlussky, 2015

P
paraarmatum Radchenko & Dlussky, 2015