Checklist of Aenictus species

The following species and subspecies belong to the genus Aenictus. Synonyms are listed under their senior names. For valid names only see Aenictus species and for distribution information see Aenictus species by Country.

A
abeillei (André, 1886) (Algeria)

acerbus Shattuck, 2008 (Australia)

aitkenii Forel, 1901 (India, Sri Lanka)

• Aenictus aratus asiatica Forel, 1911

alluaudi Santschi, 1910 (Democratic Republic of Congo, Kenya, Zimbabwe)

alluaudi falcifer Santschi, 1924 (Democratic Republic of Congo)

alticola Wheeler, W.M. & Chapman, 1930 (Philippines)

ambiguus Shuckard, 1840 (India, Nepal)

anceps Forel, 1910 (Eritrea, Sudan)

annae Forel, 1911 (Indonesia)

appressipilosus Jaitrong & Yamane, 2013 (Malaysia, Philippines)

arabicus Sharaf & Aldawood, 2012 (Saudi Arabia)

aratus Forel, 1900 (Australia, Borneo, China, India, Indonesia, Malaysia, New Guinea)

• Aenictus bengalensis impressus Forel, 1927

artipus Wilson, 1964 (China, Thailand, Vietnam)

arya Forel, 1901 (India)

asantei Campione, Novak & Gotwald, 1983 (Ghana, Nigeria)

asperivalvus Santschi, 1919 (Ivory Coast)

B
bakeri Menozzi, 1925 (Philippines)

baliensis Jaitrong & Yamane, 2013 (Indonesia)

bayoni Menozzi, 1933 (Uganda)

bidentatus Donisthorpe, 1942 (Cameroun, Ghana, Ivory Coast, Kenya, Nigeria, Uganda)

binghamii Forel, 1900 (Bangladesh, China, India, Laos, Myanmar, Thailand, Vietnam)

• Aenictus binghami gatesi Wheeler, W.M., 1927

biroi Forel, 1907 (Sri Lanka)

bobaiensis Zhou & Chen, 1999 (China)

bodongjaya Jaitrong & Yamane, 2011 (Indonesia)

boltoni Gomez, 2022 (Ghana, Ivory Coast, Nigeria)

bottegoi Emery, 1899 (Ethiopia, Zimbabwe)

bottegoi noctivagus Santschi, 1913 (Ethiopia)

breviceps Forel, 1912 (Indonesia)

brevicornis (Mayr, 1879) (Bangladesh, India, Vietnam)

brevinodus Jaitrong & Yamane, 2011 (Indonesia, Sulawesi)

brevipodus Jaitrong & Yamane, 2013 (Vietnam)

buttelreepeni Forel, 1913 (Indonesia)

buttgenbachi Forel, 1913 (Democratic Republic of Congo)

C
camposi Wheeler, W.M. & Chapman, 1925 (Borneo, China, Indonesia, Malaysia, Philippines, Singapore, Thailand)

carolianus Zettel & Sorger, 2010 (Philippines)

certus Westwood, 1842 (India)

ceylonicus (Mayr, 1866) (India, Japan, Sri Lanka)

• Aenictus ceylonicus latro Forel, 1901

• Aenictus papuanus Donisthorpe, 1941

• Aenictus similis Donisthorpe, 1948

changmaianus Terayama & Kubota, 1993 (Cambodia, Thailand, Vietnam)

chapmani Wilson, 1964 (Borneo, Indonesia, Malaysia, New Guinea)

clavatus Forel, 1901 (India)

clavatus atripennis Forel, 1913 (Indonesia)

clavatus kanariensis Forel, 1901 (India)

clavatus sundaicus Forel, 1909 (Indonesia)

clavitibia Forel, 1901 (Bangladesh, India)

clavitibia facetus Forel, 1911 (Indonesia)

concavus Jaitrong & Yamane, 2013 (Thailand, Vietnam)

congolensis Santschi, 1911 (Central African Republic, Congo, Gabon, Zambia)

cornutus Forel, 1900 (Borneo, Indonesia, Malaysia)

crucifer Santschi, 1914 (Kenya)

crucifer tuberculatus Arnold, 1915 (Zimbabwe)

currax Emery, 1900 (New Guinea)

cylindripetiolus Jaitrong & Yamane, 2013 (Singapore, Thailand)

D
decolor (Mayr, 1879) (Benin, Cameroun, Gabon, Ghana, Guinea, Ivory Coast, Kenya, Nigeria)

• Aenictus batesi Forel, 1911

dentatus Forel, 1911 (Borneo, China, India, Indonesia, Malaysia, Thailand)

diclops Shattuck, 2008 (Australia)

dlusskyi Arnoldi, 1968 (Armenia, Iran, Turkmenistan)

doryloides Wilson, 1964 (India)

doydeei Jaitrong & Yamane, 2011 (Laos, Thailand, Vietnam)

duengkaei Jaitrong & Yamane, 2012 (Thailand)

E
eguchii Jaitrong & Yamane, 2013 (Vietnam)

eugenii Emery, 1895 (Botswana, Burundi, Central African Republic, Congo, Democratic Republic of Congo, Eritrea, Eswatini, Ethiopia, Kenya, Mozambique, South Africa, Uganda, Zimbabwe)

• Aenictus eugeniae kenyensis Santschi, 1933

• Aenictus eugenii brazzai Santschi, 1910

• Aenictus eugenii caroli Forel, 1910

• Aenictus eugenii henrii Santschi, 1924

exilis Wilson, 1964 (New Guinea)

F
feae Emery, 1889 (Bangladesh, China, Myanmar)

fergusoni Forel, 1901 (Bangladesh, China, India, Myanmar, Nicobar Island, Sri Lanka, Thailand, Vietnam)

• Aenictus fergusoni karawaiewi Wheeler, W.M. & Chapman, 1930

• Aenictus fergusoni montanus Forel, 1901

• Aenictus fergusoni piltzi Forel, 1901

foreli Santschi, 1919 (Ivory Coast)

formosensis Forel, 1913 (China, Taiwan)

fuchuanensis Zhou, 2001 (Cambodia, China, Laos, Thailand, Vietnam)

fulvus Jaitrong & Yamane, 2011 (Borneo, Indonesia, Malaysia, Thailand)

furculatus Santschi, 1919 (Senegal)

furculatus andrieui Santschi, 1930 (Sudan)

fuscipennis Forel, 1913 (Indonesia)

fuscovarius Gerstäcker, 1859 (Eritrea, Kenya, Mozambique, Senegal)

fuscovarius laetior Forel, 1910 (Eritrea, Ethiopia)

fuscovarius magrettii Emery, 1892 (Kenya, Sudan)

fuscovarius sagittarius Santschi, 1938 (Egypt)

G
gibbosus Dalla Torre, 1893 (Indonesia)

gibbosus ashaverus Forel, 1913 (Indonesia)

glabratus Jaitrong & Nur-Zati, 2010 (Malaysia)

glabrinotum Jaitrong & Yamane, 2011 (Borneo, Indonesia, Malaysia)

gleadowii Forel, 1901 (India)

gonioccipus Jaitrong & Yamane, 2013 (Indonesia)

gracilis Emery, 1893 (Bangladesh, Borneo, India, Indonesia, Malaysia, Myanmar, Philippines, Singapore, Sri Lanka, Vietnam)

• Aenictus martini Forel, 1901

• Aenictus martini boelianensis Forel, 1913

• Eciton fergusoni elongatus Karavaiev, 1926

grandis Bingham, 1903 (China, Myanmar)

guineensis Santschi, 1924 (Benin, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Nigeria, Senegal)

gutianshanensis Staab, 2014 (China)

H
hamifer Emery, 1896 (Eritrea, Kenya)

hamifer spinosior Stitz, 1917 (Algeria)

henanensis Li & Wang, Y.-L., 2005 (China)

hilli Clark, 1928 (Australia)

hitai Gomez, 2022 (Kenya, Mozambique)

hodgsoni Forel, 1901 (Cambodia, China, India, Indonesia, Laos, Myanmar, Thailand, Vietnam)

hoelldobleri Stabb, 2015 (China)

hottai Terayama & Yamane, 1989 (Indonesia, Malaysia, Thailand)

humeralis Santschi, 1910 (Senegal, Sudan)

humeralis chevalieri Santschi, 1910 (Senegal)

humeralis viridans Santschi, 1915 (Benin)

huonicus Wilson, 1964 (New Guinea)

I
icarus Forel, 1911 (Indonesia)

icarus incautus Forel, 1911 (Indonesia)

idoneus Menozzi, 1928 (Indonesia)

inconspicuus Westwood, 1845 (South Africa)

indicus Bharti, Wachkoo & Kumar, 2012 (India)

inflatus Yamane & Hashimoto, 1999 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

itoi Jaitrong & Yamane, 2013 (Indonesia)

J
jacki Gomez, 2022 (Mozambique)

jacobsoni Forel, 1909 (Indonesia)

jarujini Jaitrong & Yamane, 2010 (Thailand)

javanus Emery, 1896 (Borneo, Brunei Darussalam, China, Indonesia, Malaysia, Thailand, Vietnam)

jawadwipa Jaitrong & Yamane, 2013 (Indonesia, Malaysia)

K
khaoyaiensis Jaitrong & Yamane, 2013 (Thailand)

kodungallurensis Antony & Prasad, 2022 (India)

koloi Gomez, 2022 (Ghana, Ivory Coast)

kutai Jaitrong & Wiwatwitaya, 2013 (Indonesia)

L
laeviceps (Smith, F., 1857) (Bangladesh, Borneo, China, India, Indonesia, Malaysia, Myanmar, Philippines, Thailand, Vietnam)

• Aenictus laeviceps smythiesii Forel, 1901

• Eciton fergusoni sundaica Karavaiev, 1927

latifemoratus Terayama & Yamane, 1989 (Borneo, Indonesia, Malaysia)

latiscapus Forel, 1901 (China, India)

latiscapus fumatus Wheeler, W.M., 1927 (China)

latiscapus sauteri Forel, 1913 (Taiwan)

leliepvrei Bernard, 1953 (Algeria)

leptotyphlatta Jaitrong & Eguchi, 2010 (Thailand)

levior (Karavaiev, 1926) (Borneo, Brunei Darussalam, Indonesia, Malaysia)

lifuiae Terayama, 1984 (China, Japan, Taiwan)

longi Forel, 1901 (Bangladesh, India)

longi taivanae Forel, 1913 (Taiwan)

longicephalus Jaitrong & Yamane, 2013 (Indonesia)

longinodus Jaitrong & Yamane, 2012 (Thailand)

luteus Emery, 1892 (Ivory Coast, Sierra Leone)

luteus moestus Santschi, 1930 (Sudan)

luzoni Wheeler, W.M. & Chapman, 1925 (Philippines)

M
malakkaparensis Antony & Prasad, 2022 (India)

maneerati Jaitrong & Yamane, 2013 (China, Thailand, Vietnam)

mariae Emery, 1895 (South Africa, Zimbabwe)

• Aenictus mariae natalensis Forel, 1901

mauritanicus Santschi, 1910 (Morocco)

mentu Weber, 1942 (Sudan)

minimus Jaitrong & Hashimoto, 2012 (Vietnam)

minipetiolus Jaitrong & Yamane, 2013 (Indonesia)

minutulus Terayama & Yamane, 1989 (Borneo, Indonesia, Malaysia)

mocsaryi Emery, 1901 (New Guinea)

moebii Emery, 1895 (Ivory Coast, Togo)

moebii pudicus Santschi, 1919 (Ivory Coast)

moebii sankisianus Forel, 1913 (Democratic Republic of Congo)

montivagus Jaitrong & Yamane, 2011 (Borneo, Indonesia, Malaysia)

mutatus Santschi, 1913 (Benin, Ivory Coast)

mvuvii Gomez, 2022 (Central African Republic, Gabon, Ghana, Ivory Coast, Uganda)

N
nesiotis Wheeler, W.M. & Chapman, 1930 (Australia, New Guinea, Philippines)

nganduensis Wilson, 1964 (New Guinea)

nishimurai Terayama & Kubota, 1993 (Laos, Thailand, Vietnam)

nuchiti Jaitrong & Ruangsittichai, 2018 (Thailand)

nyuyi Gomez, 2022 (Senegal)

O
obscurus Smith, F., 1865 (New Guinea)

orientalis (Karavaiev, 1926) (Indonesia, New Guinea)

P
pachycerus (Smith, F., 1858) (India, Sri Lanka)

• Aenictus bengalensis continuus Forel, 1901

• Typhlatta bengalensis Mayr, 1879

pangantihoni Zettel & Sorger, 2010 (Philippines)

paradentatus Jaitrong & Yamane, 2012 (China, Laos, Thailand, Vietnam)

parahuonicus Jaitrong & Yamane, 2011 (Laos, Thailand, Vietnam)

peguensis Emery, 1895 (India, Myanmar, Vietnam)

pfeifferi Zettel & Sorger, 2010 (Borneo, Indonesia, Malaysia)

pharao Santschi, 1924 (Sudan)

philiporum Wilson, 1964 (Australia, New Guinea)

philippinensis Chapman, 1963 (Philippines)

piercei Wheeler, W.M. & Chapman, 1930 (China, India, Philippines)

pilosus Jaitrong & Yamane, 2013 (Philippines)

pinkaewi Jaitrong & Yamane, 2013 (Thailand)

popeyei Gómez, 2022 (Cameroun, Nigeria)

porizonoides Walker, 1860 (India, Sri Lanka)

• Aenictus greeni Bingham, 1903

powersi Wheeler, W.M. & Chapman, 1930 (Philippines)

prolixus Shattuck, 2008 (Australia)

pubescens Smith, F., 1859 (India)

punctatus Jaitrong & Yamane, 2012 (Brunei Darussalam, Indonesia, Malaysia)

punctiventris Emery, 1901 (Borneo, Indonesia, Malaysia)

punctiventris scutellaris Forel, 1912 (Indonesia)

punensis Forel, 1901 (China, India)

R
rabori Chapman, 1963 (Philippines)

raptor Forel, 1913 (Democratic Republic of Congo)

reyesi Chapman, 1963 (Philippines)

rhodiensis Menozzi, 1936 (Greece, Iran, Israel, Turkey)

rixator Forel, 1901 (South Africa, South Africa)

rotundatus Mayr, 1901 (Kenya, Lesotho, Namibia, South Africa, Sudan, Zimbabwe)

• Aenictus furibundus Arnold, 1959

• Aenictus rotundatus merwei Santschi, 1932

rotundicollis Jaitrong & Yamane, 2011 (Borneo, Indonesia, Malaysia)

rougieri André, 1893 (Tunisia)

S
sagei Forel, 1901 (Afghanistan, India, Nepal, Pakistan)

samungi Jaitrong & Ruangsittichai, 2018 (Thailand)

schneirlai Wilson, 1964 (New Guinea)

seletarius Wong & Guénard, 2016 (Singapore)

shilintongae Jaitrong & Schultz, 2016 (China)

shillongensis Tiwari, 2000 (India)

shuckardi Forel, 1901 (Bangladesh, China, India, Singapore)

siamensis Jaitrong & Yamane, 2011 (Thailand)

silvestrii Wheeler, W.M., 1929 (Borneo, Indonesia, Malaysia)

sirenicus Yamane & Wang, W., 2015 (Borneo, Malaysia)

sonchaengi Jaitrong & Yamane, 2011 (Borneo, Indonesia, Malaysia, Thailand)

soudanicus Santschi, 1910 (Senegal, Sudan)

soudanicus brunneus Forel, 1913 (Democratic Republic of Congo)

spathifer Santschi, 1928 (Indonesia)

steindachneri Mayr, 1901 (South Africa, Zimbabwe)

stenocephalus Jaitrong & Yamane, 2010 (Thailand)

subterraneus Jaitrong & Hashimoto, 2012 (Malaysia)

sulawesiensis Jaitrong & Wiwatwitaya, 2013 (Indonesia)

sumatrensis Forel, 1913 (Indonesia)

sumatrensis maxillosus Forel, 1913 (Indonesia)

sundalandensis Jaitrong & Yamane, 2013 (Indonesia, Malaysia)

susanae Gomez, 2022 (Cameroun, Central African Republic)

T
thailandianus Terayama & Kubota, 1993 (China, Thailand, Vietnam)

togoensis Santschi, 1915 (Ivory Coast, Togo)

trigonus Forel, 1911 (Indonesia)

turneri Forel, 1900 (Australia)

• Aenictus deuqueti Crawley, 1923

• Aenictus exiguus Clark, 1934

U
ugaduwensis Gomez, 2022 (Burkina Faso, Senegal)

V
vagans Santschi, 1924 (Niger)

vaucheri Emery, 1915 (Algeria, Morocco)

vieti Jaitrong & Yamane, 2010 (Vietnam)

villiersi Bernard, 1953 (Guinea)

W
watanasiti Jaitrong & Yamane, 2013 (China, Thailand, Vietnam)

wayani Jaitrong & Yamane, 2011 (Indonesia, Philippines, Sulawesi)

weissi Santschi, 1910 (Cameroun, Congo, Gabon, Ghana, Guinea, Ivory Coast, Kenya, Nigeria, Rwanda, Uganda)

westwoodi Forel, 1901 (China, India)

wilaiae Jaitrong & Yamane, 2013 (Thailand)

wilsoni Bharti, Wachkoo & Kumar, 2012 (India)

wiwatwitayai Jaitrong & Yamane, 2013 (Thailand, Vietnam)

wroughtonii Forel, 1890 (India)

wudangshanensis Wang, W., 2006 (China)

X
xegi Gomez, 2022 (Congo, Ghana)

Y
yamanei Wiwatwitaya & Jaitrong, 2011 (Borneo, Indonesia, Malaysia, Singapore, Thailand)

yangi Liu, Hita Garcia, Peng & Economo, 2015 (China)

Z
zhengi Zhang, W., 1994 (China)