Colombia

The following valid species and subspecies of ants are known to occur in Colombia. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Colombian ants.

Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Colombian Ants web page. Please advise if any corrections are warranted.

Acanthognathus

 * Acanthognathus brevicornis Smith, M.R., 1944
 * Acanthognathus laevigatus Galvis & Fernández, 2009
 * Acanthognathus ocellatus Mayr, 1887
 * Acanthognathus teledectus Brown & Kempf, 1969

Acanthoponera

 * Acanthoponera minor Forel, 1899
 * Acanthoponera mucronata Roger, 1860

Acanthostichus

 * Acanthostichus fuscipennis Emery, 1895
 * Acanthostichus sanchezorum Mackay, 1985
 * Acanthostichus truncatus Mackay, 1996

Acromyrmex

 * Acromyrmex aspersus (Smith, F., 1858)
 * Acromyrmex coronatus panamensis Forel, 1899
 * Acromyrmex landolti Forel, 1885
 * Acromyrmex rugosus Smith, F., 1858
 * Acromyrmex santschii Forel, 1912

Acropyga

 * Acropyga exsanguis Wheeler, W.M., 1909
 * Acropyga fuhrmanni Forel, 1914
 * Acropyga goeldii Forel, 1893
 * Acropyga guianensis Weber, 1944
 * Acropyga smithii Forel, 1893
 * Acropyga tricuspis LaPolla, 2004

Adelomyrmex

 * Adelomyrmex costatus Fernández, 2003
 * Adelomyrmex cristiani Fernández, 2003
 * Adelomyrmex grandis Fernández, 2003
 * Adelomyrmex myops (Wheeler, W.M., 1910)
 * Adelomyrmex striatus Fernández, 2003
 * Adelomyrmex tristani (Menozzi, 1931)
 * Adelomyrmex vaderi Fernández, 2003

Allomerus

 * Allomerus dentatus Fernández, 2007

Anochetus

 * Anochetus bispinosus Smith, F., 1858
 * Anochetus chocoensis Zabala, 2008
 * Anochetus diegensis Forel, 1912
 * Anochetus elegans Lattke, 1987
 * Anochetus emarginatus Fabricius, 1804
 * Anochetus inermis André, 1889
 * Anochetus neglectus Emery, 1894
 * Anochetus simoni Emery, 1890
 * Anochetus vallensis Lattke, 1987

Aphaenogaster

 * Aphaenogaster inermis Forel, 1899
 * Aphaenogaster phalangium Emery, 1890

Apterostigma

 * Apterostigma ancilonodum Lattke, 1997
 * Apterostigma auriculatum Wheeler, W.M., 1925
 * Apterostigma callipygium Lattke, 1997
 * Apterostigma carinatum Lattke, 1997
 * Apterostigma chocoense Lattke, 1997
 * Apterostigma collare Emery, 1896
 * Apterostigma dentigerum Wheeler, W.M., 1925
 * Apterostigma ierense Weber, 1937
 * Apterostigma manni Weber, 1938
 * Apterostigma megacephala Lattke, 1999
 * Apterostigma reburrum Lattke, 1997
 * Apterostigma turgidum Lattke, 1997

Atta

 * Atta cephalotes Linnaeus, 1758
 * Atta colombica Guérin-Méneville, 1844
 * Atta laevigata Smith, F., 1858
 * Atta sexdens Linnaeus, 1758

Azteca

 * Azteca alfari Emery, 1893
 * Azteca aurita Emery, 1893
 * Azteca chartifex multinida Forel, 1899
 * Azteca delpini antillana Forel, 1899
 * Azteca forelii Emery, 1893
 * Azteca hypophylla Forel, 1899
 * Azteca instabilis Smith, F., 1862
 * Azteca jelskii Emery, 1893
 * Azteca lallemandi Forel, 1899
 * Azteca linamariae Guerrero, et al., 2010
 * Azteca nigricans Forel, 1899
 * Azteca polymorpha Forel, 1899
 * Azteca salti Wheeler, W.M., 1930
 * Azteca schimperi Emery, 1893
 * Azteca schumannii Emery, 1893
 * Azteca theresiae Forel, 1899
 * Azteca trigona gaigei Forel, 1914
 * Azteca trigona Emery, 1893
 * Azteca velox Forel, 1899

Basiceros

 * Basiceros conjugans Brown, 1974
 * Basiceros disciger Mayr, 1887

Belonopelta

 * Belonopelta attenuata Mayr, 1870
 * Belonopelta deletrix Mann, 1922

Blepharidatta

 * Blepharidatta brasiliensis Wheeler, W.M., 1915
 * Blepharidatta fernandezi Brandao, Feitosa & Diniz, 2015

Brachymyrmex

 * Brachymyrmex antennatus Santschi, 1929
 * Brachymyrmex aphidicola Forel, 1909
 * Brachymyrmex australis Forel, 1901
 * Brachymyrmex bruchi Forel, 1912
 * Brachymyrmex cavernicola Wheeler, 1938
 * Brachymyrmex cordemoyi Forel, 1895
 * Brachymyrmex degener Emery, 1906
 * Brachymyrmex donisthorpei Santschi, 1939
 * Brachymyrmex flavidulus Roger, 1863
 * Brachymyrmex heeri Forel, 1874
 * Brachymyrmex tristis Mayr, 1870
 * Brachymyrmex minutus Forel, 1893
 * Brachymyrmex modestus Santschi, 1923
 * Brachymyrmex musculus Forel, 1899
 * Brachymyrmex myops Emery, 1906
 * Brachymyrmex obscurior Forel, 1893
 * Brachymyrmex patagonicus Mayr, 1868
 * Brachymyrmex pictus Mayr, 1887
 * Brachymyrmex termitophilus Forel, 1895b

Camponotus

 * Camponotus abscisus Roger, 1863
 * Camponotus ager Smith, F., 1858
 * Camponotus amoris Forel, 1904
 * Camponotus apicalis Mann, 1916
 * Camponotus arhuacus Forel, 1902
 * Camponotus atriceps Smith, F., 1858
 * Camponotus auricomus Roger, 1862
 * Camponotus bidens Mayr, 1870
 * Camponotus bispinosus Mayr, 1870
 * Camponotus blandus Smith, 1858
 * Camponotus brevis Forel, 1899
 * Camponotus bugnioni Forel, 1899
 * Camponotus canescens Mayr, 1870
 * Camponotus chartifex Smith, F., 1860
 * Camponotus cheesmanae Donisthorpe, 1932
 * Camponotus circularis Mayr, 1870
 * Camponotus coloratus Forel, 1904
 * Camponotus conulus Mayr, 1870
 * Camponotus coruscus Smith, F., 1862
 * Camponotus dalmasi Forel, 1899
 * Camponotus eurynotus Forel, 1907
 * Camponotus excisus Mayr, 1870
 * Camponotus falco Forel, 1902
 * Camponotus fasciatellus Dalla Torre, 1892
 * Camponotus fastigatus Roger, 1863
 * Camponotus improprius Forel, 1879
 * Camponotus indianus Forel, 1879
 * Camponotus integellus Forel, 1899
 * Camponotus landolti Forel, 1879
 * Camponotus lespesii Forel, 1886
 * Camponotus lindigi Mayr, 1870
 * Camponotus linnaei Forel, 1886
 * Camponotus mus Roger, 1863
 * Camponotus nidulans Smith, F., 1860
 * Camponotus nitens Mayr, 1870
 * Camponotus nitens fuhrmanni Forel, 1921
 * Camponotus nitidior Santschi, 1921
 * Camponotus novogranadensis Mayr, 1870
 * Camponotus orthocephalus Emery, 1894
 * Camponotus pachylepis Emery, 1920
 * Camponotus planatus Roger, 1863
 * Camponotus planatus colombicus Forel, 1899
 * Camponotus polymorphicus Mackay, López-Castro & Fernández, 2002
 * Camponotus propinquus Mayr, 1887
 * Camponotus raphaelis Forel, 1869
 * Camponotus rufipes Fabricius, 1775
 * Camponotus sanctaefidei Dalla Torre, 1892
 * Camponotus santschii Forel, 1899
 * Camponotus senex Smith, F., 1858
 * Camponotus sericeiventris Guérin-Méneville, 1838
 * Camponotus sericeiventris satrapus Wheeler, W.M. 1931
 * Camponotus sexguttatus Fabricius, 1793
 * Camponotus sexguttatus perturbans Kutter, 1931
 * Camponotus silvicola Forel, 1902
 * Camponotus simillimus Smith, F., 1862
 * Camponotus sphenoidalis Mayr, 1870
 * Camponotus striatus Smith, 1862
 * Camponotus substitutus Emery, 1894
 * Camponotus tonduzi Forel, 1899
 * Camponotus traili Mayr, 1878
 * Camponotus trapezoideus Mayr, 1870
 * Camponotus ustus Forel, 1879
 * Camponotus zoc Forel, 1879

Carebara

 * Carebara angulata Fernández, 2004
 * Carebara arabara Fernández, 2010
 * Carebara audita Fernández, 2004
 * Carebara brevipilosa Fernández, 2004
 * Carebara coeca Fernández, 2004
 * Carebara coqueta Fernández, 2006
 * Carebara elongata Fernández, 2004
 * Carebara globularia Fernández, 2004
 * Carebara kofana Fernández, 2004
 * Carebara paya Fernández, 2004
 * Carebara reina Fernández, 2004
 * Carebara reticulata Fernández, 2004
 * Carebara urichi Wheeler, W.M., 1922

Centromyrmex

 * Centromyrmex brachycola Roger, 1861

Cephalotes

 * Cephalotes argentatus (Smith, F., 1853)
 * Cephalotes atratus Linnaeus, 1758
 * Cephalotes basalis Smith, F., 1876
 * Cephalotes christopherseni Forel, 1912
 * Cephalotes clypeatus Fabricius, 1804
 * Cephalotes coffeae (Kempf, 1953)
 * Cephalotes columbicus (Forel, 1912)
 * Cephalotes complanatus Guérin-Méneville, 1844
 * Cephalotes conspersus (F. Smith, 1867)
 * Cephalotes cordatus Smith, F., 1853
 * Cephalotes crenaticeps (Mayr, 1866)
 * Cephalotes cristatus Emery, 1890
 * Cephalotes decolor De Andrade, 1999
 * Cephalotes depressus (Klug, 1824)
 * Cephalotes duckei Forel, 1906
 * Cephalotes femoralis (Smith, F., 1853)
 * Cephalotes foliaceus Emery, 1906
 * Cephalotes grandinosus Smith, F., 1860
 * Cephalotes inaequalis Mann, 1916
 * Cephalotes laminatus (F. Smith, 1860)
 * Cephalotes maculatus Smith, F., 1876
 * Cephalotes manni Kempf, 1951
 * Cephalotes marginatus Fabricius, 1804
 * Cephalotes minutus Fabricius, 1804
 * Cephalotes mompox De Andrade, 1999
 * Cephalotes opacus Santschi, 1920
 * Cephalotes pallens Klug, 1824
 * Cephalotes pallidoides de Andrade, 1999
 * Cephalotes pallidus de Andrade, 1999
 * Cephalotes palta De Andrade, 1999
 * Cephalotes palustris de Andrade, 1999
 * Cephalotes patei (Kempf, 1951)
 * Cephalotes pavonii Latreille, 1809
 * Cephalotes placidus Smith, F., 1860
 * Cephalotes porrasi Wheeler, W.M., 1942
 * Cephalotes pusillus Klug, 1824
 * Cephalotes ramiphilus Forel, 1904
 * Cephalotes scutulatus Smith, F., 1867
 * Cephalotes serraticeps Smith, F., 1858
 * Cephalotes setulifer Emery, 1894
 * Cephalotes simillimus Kempf, 1951
 * Cephalotes spinosus Mayr, 1862
 * Cephalotes umbraculatus Fabricius, 1804

Cheliomyrmex

 * Cheliomyrmex andicola Emery, 1894
 * Cheliomyrmex audax Santschi, 1921
 * Cheliomyrmex morosus (Smith, F. 1859)

Crematogaster

 * Crematogaster abstinens Forel, 1899
 * Crematogaster acuta Fabricius, 1804
 * Crematogaster ampla Forel, 1912
 * Crematogaster arcuata Forel, 1899
 * Crematogaster brasiliensis Mayr, 1878
 * Crematogaster bryophilia Longino, 2003
 * Crematogaster carinata Mayr, 1862
 * Crematogaster crinosa Mayr, 1862
 * Crematogaster crucis Forel, 1912
 * Crematogaster curvispinosa Mayr, 1862
 * Crematogaster distans Mayr, 1870
 * Crematogaster erecta Mayr, 1866
 * Crematogaster evallans Forel, 1907
 * Crematogaster flavomicrops Longino, 2003
 * Crematogaster flavosensitiva Longino, 2003
 * Crematogaster levior Longino, 2003
 * Crematogaster limata Smith, F., 1858
 * Crematogaster longispina Emery, 1890
 * Crematogaster montezumia Smith, F., 1858
 * Crematogaster nigropilosa Mayr, 1870
 * Crematogaster obscurata Emery, 1895
 * Crematogaster quadriformis Roger, 1863
 * Crematogaster rochai Forel, 1903
 * Crematogaster sotobosque Longino, 2003
 * Crematogaster stollii Forel, 1885
 * Crematogaster tenuicula Forel, 1904
 * Crematogaster torosa Mayr, 1870

Cryptopone

 * Cryptopone guianensis Weber, 1939

Cylindromyrmex

 * Cylindromyrmex boliviae Wheeler, W.M., 1924
 * Cylindromyrmex escobari De Andrade, 1998
 * Cylindromyrmex godmani Forel, 1899
 * Cylindromyrmex whymperi Cameron, 1891

Cyphomyrmex

 * Cyphomyrmex bicarinatus Snelling, R.R. & Longino, 1992
 * Cyphomyrmex bicornis Forel, 1895
 * Cyphomyrmex castagnei MacKay & Baena, 1993
 * Cyphomyrmex cornutus Kempf, 1968
 * Cyphomyrmex costatus Mann, 1922
 * Cyphomyrmex hamulatus Weber, 1938
 * Cyphomyrmex kirbyi Mayr, 1887
 * Cyphomyrmex longiscapus Weber, 1940
 * Cyphomyrmex minutus Mayr, 1862
 * Cyphomyrmex muelleri Schultz & Solomon, 2002
 * Cyphomyrmex rimosus Spinola, 1851
 * Cyphomyrmex salvini Forel, 1899

Dinoponera

 * Dinoponera longipes Emery, 1901

Discothyrea

 * Discothyrea sexarticulata (Borgmeier, 1954)

Dolichoderus

 * Dolichoderus abruptus Smith, F., 1858
 * Dolichoderus baenae Mackay, 1993
 * Dolichoderus bispinosus Olivier, 1792
 * Dolichoderus curvilobus Lattke, 1987
 * Dolichoderus debilis Emery, 1890
 * Dolichoderus decollatus Smith, F., 1858
 * Dolichoderus diversus Emery, 1894
 * Dolichoderus doloniger (Roger)
 * Dolichoderus fernandezi Mackay, 1993
 * Dolichoderus inpai (Harada)
 * Dolichoderus lamellosus Mayr, 1870
 * Dolichoderus laminatus Mayr, 1870
 * Dolichoderus laurae Mackay, 1993
 * Dolichoderus lobicornis (Kempf)
 * Dolichoderus longicollis Mackay, 1993
 * Dolichoderus lugens Emery
 * Dolichoderus lutosus Smith, F., 1858
 * Dolichoderus piceus Mackay, 1993
 * Dolichoderus schulzi Emery, 1894
 * Dolichoderus shattucki MacKay
 * Dolichoderus spinicollis (Latreille)
 * Dolichoderus superaculus Lattke, 1987
 * Dolichoderus utriensis Ortiz & Fernández, 2011
 * Dolichoderus varians Mann

Dorymyrmex

 * Dorymyrmex amazonicus Cuezzo & Guerrero, 2011
 * Dorymyrmex bicolor Wheeler, 1906
 * Dorymyrmex biconis Forel, 1912
 * Dorymyrmex brunneus Forel, 1908
 * Dorymyrmex goeldii Forel, 1904
 * Dorymyrmex insanus (Buckley, 1866)
 * Dorymyrmex pyramicus Roger, 1863
 * Dorymyrmex tuberosus Cuezzo & Guerrero, 2011
 * Dorymyrmex xerophylus Cuezzo & Guerrero, 2011

Eciton

 * Eciton burchellii Westwood, 1842
 * Eciton burchellii cupiens Santschi, 1923
 * Eciton burchellii foreli Mayr, 1886
 * Eciton hamatum Fabricius, 1782
 * Eciton jansoni Forel, 1912
 * Eciton lucanoides Emery, 1894
 * Eciton mexicanum Roger, 1863

Ectatomma

 * Ectatomma brunneum Smith, F., 1858
 * Ectatomma confine Mayr, 1870
 * Ectatomma edentatum Roger, 1863
 * Ectatomma gibbum Kugler, C. & Brown, 1982
 * Ectatomma goninion Kugler, C. & Brown, 1982
 * Ectatomma ruidum Roger, 1860
 * Ectatomma tuberculatum Olivier, 1792

Eurhopalothrix

 * Eurhopalothrix pilulifera Brown & Kempf, 1960

Forelius

 * Forelius damiani Guerro & Fernández, 2008

Fulakora

 * Fulakora lurilabes
 * Fulakora orizabana (Brown, 1960)

Gigantiops

 * Gigantiops destructor Fabricius, 1804

Gnamptogenys

 * Gnamptogenys acuminata Emery, 1896
 * Gnamptogenys acuta Brown, 1956
 * Gnamptogenys alfaroi Emery, 1894
 * Gnamptogenys andina Lattke, 1995
 * Gnamptogenys annulata Mayr, 1887
 * Gnamptogenys bispinosa Emery, 1890
 * Gnamptogenys bisulca Kempf & Brown, 1968
 * Gnamptogenys brunnea Lattke, 1995
 * Gnamptogenys caelata Kempf, 1967
 * Gnamptogenys concinna Smith, F., 1858
 * Gnamptogenys continua Mayr, 1887
 * Gnamptogenys curvoclypeata Lattke, 1990
 * Gnamptogenys dichotoma MacKay & Mackay, 2008
 * Gnamptogenys ejuncida Lattke, 1995
 * Gnamptogenys enodis Lattke, Fernández & Palacio, 2004
 * Gnamptogenys ericae Forel, 1912
 * Gnamptogenys extra Lattke, 1995
 * Gnamptogenys fernandezi Lattke, 1990
 * Gnamptogenys gentryi Lattke, 1995
 * Gnamptogenys haenschi Emery, 1902
 * Gnamptogenys hartmani Wheeler, W.M., 1915
 * Gnamptogenys horni Santschi, 1929
 * Gnamptogenys ingeborgae Brown, 1993
 * Gnamptogenys interrupta Mayr, 1887
 * Gnamptogenys kempfi Lenko, 1964
 * Gnamptogenys lanei Kempf, 1960
 * Gnamptogenys mecotyle Brown, 1958
 * Gnamptogenys mina Brown, 1956
 * Gnamptogenys minuta Emery, 1896
 * Gnamptogenys moelleri Forel, 1912
 * Gnamptogenys mordax Smith, F., 1858
 * Gnamptogenys nigrivitrea Lattke, 1995
 * Gnamptogenys perspicax Kempf & Brown, 1970
 * Gnamptogenys pilosa Lattke, 1995
 * Gnamptogenys pleurodon Emery, 1896
 * Gnamptogenys porcata Emery, 1896
 * Gnamptogenys regularis Mayr, 1870
 * Gnamptogenys relicta Mann, 1916
 * Gnamptogenys stellae Lattke, 1995
 * Gnamptogenys striatula Mayr, 1884
 * Gnamptogenys strigata Norton, 1868
 * Gnamptogenys sulcata Smith, F., 1858
 * Gnamptogenys tortuolosa Smith, F., 1858
 * Gnamptogenys triangularis Mayr, 1887

Heteroponera

 * Heteroponera inca Brown, 1958
 * Heteroponera microps Borgmeier, 1957
 * Heteroponera monticola Kempf & Brown, 1970

Hylomyrma

 * Hylomyrma blandiens Kempf, 1961
 * Hylomyrma columbica Forel, 1912
 * Hylomyrma dolichops Kempf, 1973
 * Hylomyrma immanis Kempf, 1973
 * Hylomyrma praepotens Kempf, 1973
 * Hylomyrma sagax Kempf, 1973
 * Hylomyrma reitteri (Mayr, 1887)
 * Hylomyrma transversa Kempf, 1973

Hypoponera

 * Hypoponera distinguenda Emery, 1890
 * Hypoponera fiebrigi antoniensis Forel, 1912
 * Hypoponera fiebrigi famini Forel, 1912
 * Hypoponera opaciceps gaigei Forel, 1914

Kalathomyrmex

 * Kalathomyrmex emeryi (Forel, 1907)

Kempfidris

 * Kempfidris inusualis (Fernández, 2007)

Labidus

 * Labidus coecus Latreille, 1802
 * Labidus praedator Smith, F., 1858
 * Labidus praedator sedulus Menozzi, 1926
 * Labidus spininodis Emery, 1890

Lachnomyrmex

 * Lachnomyrmex fernandezi Feitosa & Brandao, 2008
 * Lachnomyrmex grandis Fernández & Baena, 1997
 * Lachnomyrmex longinodus Fernández & Baena, 1997
 * Lachnomyrmex pilosus Weber, 1950

Lenomyrmex

 * Lenomyrmex foveolatus Fernández & Palacio, 1999
 * Lenomyrmex inusitatus Fernández, 2001
 * Lenomyrmex mandibularis Fernández & Palacio, 1999
 * Lenomyrmex wardi Fernández & Palacio, 1999

Leptanilloides

 * Leptanilloides legionarius Brandão, Diniz, Agosti & Delabie, 1999
 * Leptanilloides sculpturatus Brandão, Diniz, Agosti & Delabie, 1999

Leptogenys

 * Leptogenys amu Lattke, 2011
 * Leptogenys carbonaria Lattke, 2011
 * Leptogenys erugata Lattke, 2011
 * Leptogenys foraminosa Lattke, 2011
 * Leptogenys gaigei 	Wheeler, W.M., 1923
 * Leptogenys gatu Lattke, 2011
 * Leptogenys glabra Lattke, 2011
 * Leptogenys gorgona Lattke, 2011
 * Leptogenys ingens Mayr, 1866
 * Leptogenys langi Wheeler, W.M., 1923
 * Leptogenys linda Lattke, 2011
 * Leptogenys mucronata columbica Forel, 1901
 * Leptogenys nigricans Lattke, 2011
 * Leptogenys phylloba Lattke, 2011
 * Leptogenys pubiceps Emery, 1890
 * Leptogenys pusilla Emery, 1890
 * Leptogenys quadrata Lattke, 2011
 * Leptogenys ritae Forel, 1899
 * Leptogenys serrata Lattke, 2011
 * Leptogenys socorda Lattke, 2011
 * Leptogenys tama Lattke, 2011
 * Leptogenys unistimulosa Santschi, 1929
 * Leptogenys volcanica Lattke, 2011

Linepithema

 * Linepithema angulatum (Emery, 1894)
 * Linepithema dispertitum Forel, 1885
 * Linepithema gallardoi (Brèthes, 1914)
 * Linepithema hirsutum Escarraga & Guerrero, 2016
 * Linepithema iniquum Mayr, 1870
 * Linepithema neotropicum Wild, 2007
 * Linepithema piliferum (Mayr, 1870)
 * Linepithema tsachila Wild, 2007

Mayaponera

 * Mayaponera arhuaca (Forel, 1901)
 * Mayaponera becculata MacKay & Mackay, 2010
 * Mayaponera conicula MacKay & Mackay, 2010
 * Mayaponera constricta Mayr, 1884
 * Mayaponera longidentata MacKay & Mackay, 2010
 * Mayaponera pergandei (Forel, 1909)

Megalomyrmex

 * Megalomyrmex balzani Emery, 1894
 * Megalomyrmex bidentatus Fernández & Baena, 1997
 * Megalomyrmex cuatiara Brandão, 1990
 * Megalomyrmex cupecuara Brandão, 1990
 * Megalomyrmex cyendyra Brandão, 1990
 * Megalomyrmex drifti Kempf, 1961
 * Megalomyrmex emeryi Forel, 1904
 * Megalomyrmex foreli Emery, 1890
 * Megalomyrmex incisus Smith, M.R., 1947
 * Megalomyrmex leoninus Forel, 1885
 * Megalomyrmex megadrifti Boudinot, Sumnicht & Adams, 2013 41
 * Megalomyrmex modestus Emery, 1896
 * Megalomyrmex pacova Brandão, 1990
 * Megalomyrmex peetersi Prado & Adams, 2020
 * Megalomyrmex poatan Brandão, 1990
 * Megalomyrmex silvestrii Wheeler, W.M., 1909
 * Megalomyrmex staudingeri Emery, 1890
 * Megalomyrmex wallacei Mann, 1916 66

Monomorium

 * Monomorium floricola Jerdon, 1851
 * Monomorium pharaonis Linnaeus, 1758

Mycetarotes

 * Mycetarotes parallelus (Emery, 1906)

Mycetomoellerius

 * Mycetomoellerius gaigei (Forel, 1914)
 * Mycetomoellerius urichii (Forel, 1893)
 * Mycetomoellerius zeteki (Weber, 1940)

Mycocepurus

 * Mycocepurus smithii Forel, 1893

Myrmelachista

 * Myrmelachista dalmasi Forel, 1912
 * Myrmelachista reclusi Forel, 1903
 * Myrmelachista schumanni Emery, 1890
 * Myrmelachista zeledoni Emery, 1896

Myrmicocrypta

 * Myrmicocrypta boliviana Weber, 1938
 * Myrmicocrypta ednaella Mann, 1922
 * Myrmicocrypta foreli Mann, 1916
 * Myrmicocrypta occipitalis Weber, 1938
 * Myrmicocrypta spinosa Weber, 1937
 * Myrmicocrypta unidentata Weber, 1937
 * Myrmicocrypta urichi Weber, 1937

Neivamyrmex

 * Neivamyrmex curvinotus 	Watkins, 1994
 * Neivamyrmex emersoni Wheeler, W.M., 1921
 * Neivamyrmex gibbatus Borgmeier, 1953
 * Neivamyrmex guyanensis Santschi, 1916
 * Neivamyrmex halidaii Shuckard, 1840
 * Neivamyrmex punctaticeps Emery, 1894
 * Neivamyrmex rosenbergi Forel, 1911
 * Neivamyrmex swainsonii Shuckard, 1840

Neoponera

 * Neoponera aenescens Mayr, 1870
 * Neoponera agilis (Forel, 1901)
 * Neoponera apicalis Latreille, 1802
 * Neoponera bugabensis Forel, 1899
 * Neoponera carbonaria (Smith, F., 1858)
 * Neoponera carinulata (Roger, 1861)
 * Neoponera chyzeri Forel, 1907
 * Neoponera commutata Roger, 1860
 * Neoponera crenata Roger, 1861
 * Neoponera curvinodis Forel, 1899
 * Neoponera fauveli Emery, 1896
 * Neoponera foetida Linnaeus, 1758
 * Neoponera fusca MacKay & Mackay, 2010
 * Neoponera hispida MacKay & Mackay, 2010
 * Neoponera holcotyle MacKay & Mackay, 2010
 * Neoponera inversa Smith, F., 1858
 * Neoponera laevigata Smith, F., 1858
 * Neoponera magnifica Borgmeier, 1929
 * Neoponera moesta Mayr, 1870
 * Neoponera recava MacKay & Mackay, 2010
 * Neoponera striatinodis Emery, 1890
 * Neoponera unidentata Mayr, 1862
 * Neoponera villosa Fabricius, 1804

Nesomyrmex

 * Nesomyrmex anduzei Weber, 1943
 * Nesomyrmex antoniensis (Forel, 1912)
 * Nesomyrmex echinatinodis Forel, 1886
 * Nesomyrmex pleuriticus Kempf, 1959
 * Nesomyrmex spininodis Mayr, 1887
 * Nesomyrmex tonsuratus Kempf, 1959

Nomamyrmex

 * Nomamyrmex esenbeckii (Westwood, 1842)
 * Nomamyrmex hartigii Westwood, 1842

Nylanderia

 * Nylanderia fulva Mayr, 1862
 * Nylanderia johannae Forel, 1912
 * Nylanderia johannae dibullana (Forel, 1912)
 * Nylanderia nodifera Mayr, 1870
 * Nylanderia steinheili Forel, 1893

Ochetomyrmex

 * Ochetomyrmex neopolitus Fernández, 2003
 * Ochetomyrmex semipolitus Mayr, 1878

Octostruma

 * Octostruma amrishi (Makhan, 2007)
 * Octostruma balzani (Emery, 1894)
 * Octostruma convallisur Longino, 2013
 * Octostruma iheringi (Emery, 1888)
 * Octostruma impressa Palacio, 1997
 * Octostruma inca Brown & Kempf, 1960
 * Octostruma obtusidens Longino, 2013
 * Octostruma pexidorsum Longino, 2013
 * Octostruma rugifera (Mayr, 1887)
 * Octostruma stenoscapa Palacio, 1997

Odontomachus

 * Odontomachus affinis Guérin-Méneville, 1844
 * Odontomachus allolabis Kempf, 1974
 * Odontomachus bauri Emery, 1892
 * Odontomachus chelifer Latreille, 1802
 * Odontomachus erythrocephalus Emery, 1890
 * Odontomachus haematodus Linnaeus, 1758
 * Odontomachus hastatus Fabricius, 1804
 * Odontomachus meinerti Forel, 1905
 * Odontomachus mormo Brown, 1976
 * Odontomachus opaciventris Forel, 1899
 * Odontomachus panamensis Forel, 1899

Oxyepoecus

 * Oxyepoecus inquilinus Kusnezov, 1952

Pachycondyla

 * Pachycondyla crassinoda Latreille, 1802
 * Pachycondyla fuscoatra Roger, 1861
 * Pachycondyla harpax Fabricius, 1804
 * Pachycondyla impressa Roger, 1861

Paraponera

 * Paraponera clavata Fabricius, 1775

Paratrachymyrmex

 * Paratrachymyrmex bugnioni (Forel, 1912)
 * Paratrachymyrmex cornetzi (Forel, 1912)
 * Paratrachymyrmex irmgardae (Forel, 1912)

Paratrechina

 * Paratrechina longicornis Latreille, 1802

Pheidole

 * Pheidole ajax	Forel, 1899
 * Pheidole alfaroi Emery, 1896
 * Pheidole allarmata Wilson, 2003
 * Pheidole alticola Wilson, 2003
 * Pheidole amata Forel, 1901
 * Pheidole angulifera Wilson, 2003
 * Pheidole arhuaca Forel, 1901
 * Pheidole astur Wilson, 2003
 * Pheidole bellatrix Wilson, 2003
 * Pheidole biconstricta Mayr, 1870
 * Pheidole bilimeki Mayr, 1870
 * Pheidole binasifera Wilson, 2003
 * Pheidole boliviana Wilson, 2003
 * Pheidole boruca Wilson, 2003
 * Pheidole brandaoi Wilson, 2003
 * Pheidole browni Wilson, 2003
 * Pheidole bufo Wilson, 2003
 * Pheidole bulliceps Wilson, 2003
 * Pheidole calens Forel, 1901
 * Pheidole calimana Wilson, 2003
 * Pheidole carapuna Mann, 1916
 * Pheidole cataphracta Wilson, 2003
 * Pheidole charazana Wilson, 2003
 * Pheidole chocoensis Wilson, 2003
 * Pheidole christopherseni Forel, 1912
 * Pheidole colobopsis Mann, 1916
 * Pheidole cramptoni Wheeler, W.M., 1916
 * Pheidole diabolus Wilson, 2003
 * Pheidole distorta Forel, 1899
 * Pheidole dorsata Wilson, 2003
 * Pheidole ectatommoides Wilson, 2003
 * Pheidole eriophora Wilson, 2003
 * Pheidole euryscopa Wilson, 2003
 * Pheidole exarata Emery, 1896
 * Pheidole exigua Mayr, 1884
 * Pheidole fabricator Smith, F., 1858
 * Pheidole fallax Mayr, 1870
 * Pheidole fimbriata Roger, 1863
 * Pheidole flavens Roger, 1863
 * Pheidole gaigei Forel, 1914
 * Pheidole gauthieri Forel, 1901
 * Pheidole gertrudae Forel, 1886
 * Pheidole gibbata Borgmeier, 1934
 * Pheidole grex Wilson, 2003
 * Pheidole guajirana Wilson, 2003
 * Pheidole guilelmimuelleri Forel, 1866
 * Pheidole harrisonfordi Wilson, 2003
 * Pheidole hasticeps Wilson, 2003
 * Pheidole hazenae Wilson, 2003
 * Pheidole huilana Wilson, 2003
 * Pheidole iceni Fernandez, 2011
 * Pheidole incisa Mayr, 1870
 * Pheidole indica Mayr, 1879
 * Pheidole inversa Forel, 1901
 * Pheidole laelaps Wilson, 2003
 * Pheidole laeviventris Mayr, 1870
 * Pheidole laselva Wilson, 2003
 * Pheidole lemnisca Wilson, 2003
 * Pheidole longiscapa Forel, 1901
 * Pheidole megacephala Fabricius, 1793
 * Pheidole melastomae Wilson, 2003
 * Pheidole mendicula Wheeler, W.M., 1925
 * Pheidole metana Wilson, 2003
 * Pheidole micon Wilson, 2003
 * Pheidole mirabilis Wilson, 2003
 * Pheidole mutisi Fernández & Wilson, 2008
 * Pheidole nitella Wilson, 2003
 * Pheidole oliveirai Wilson, 2003
 * Pheidole pepo Wilson, 2003
 * Pheidole perpusilla Emery, 1894
 * Pheidole pidax Wilson, 2003
 * Pheidole praeusta Roger, 1863
 * Pheidole pubiventris Mayr, 1887
 * Pheidole punctatissima Mayr, 1870
 * Pheidole pygmaea Wilson, 2003
 * Pheidole quadriceps Wilson, 2003
 * Pheidole radoszkowskii Mayr, 1884
 * Pheidole reclusi Forel, 1899
 * Pheidole reichenspergeri Santschi, 1923
 * Pheidole rhinoceros Forel, 1899
 * Pheidole riveti Santschi, 1911
 * Pheidole sagax Wilson, 2003
 * Pheidole scolioceps Wilson, 2003
 * Pheidole sculptior Forel, 1893
 * Pheidole securigera Wilson, 2003
 * Pheidole seeldrayersi Forel, 1910
 * Pheidole seligmanni Wilson, 2003
 * Pheidole sensitiva Borgmeier, 1959
 * Pheidole servilia Wilson, 2003
 * Pheidole sicaria Wilson, 2003
 * Pheidole sospes Forel, 1908
 * Pheidole sphaerica Wilson, 2003
 * Pheidole spilota Wilson, 2003
 * Pheidole steinheili Forel, 1901
 * Pheidole subarmata Mayr, 1884
 * Pheidole subnuda Wilson, 2003
 * Pheidole subsphaerica Wilson, 2003
 * Pheidole superba Wilson, 2003
 * Pheidole susannae Forel, 1886
 * Pheidole tambopatae Wilson, 2003
 * Pheidole tennantae Wilson, 2003
 * Pheidole tenuis Wilson, 2003
 * Pheidole tigris Wilson, 2003
 * Pheidole transversostriata Mayr, 1887
 * Pheidole tristops Wilson, 2003
 * Pheidole unicornis Wilson, 2003
 * Pheidole vafra Santschi, 1923
 * Pheidole vallifica Forel, 1901
 * Pheidole venatrix Wilson, 2003
 * Pheidole vomer Wilson, 2003
 * Pheidole vorax
 * Pheidole xanthogaster Wilson, 2003

Platythyrea

 * Platythyrea pilosula Smith, F., 1858
 * Platythyrea prizo Kugler, C., 1977
 * Platythyrea punctata Smith, F., 1858

Pogonomyrmex

 * Pogonomyrmex mayri Forel, 1899
 * Pogonomyrmex naegelii Emery 1878
 * Pogonomyrmex striatinodus Fernández & Palacio, 1998

Prionopelta

 * Prionopelta amabilis Borgmeier, 1949
 * Prionopelta antillana Forel, 1909
 * Prionopelta dubia Ladino & Feitosa, 2020
 * Prionopelta minuta Ladino & Feitosa, 2020
 * Prionopelta modesta Forel, 1909

Probolomyrmex

 * Probolomyrmex boliviensis Mann, 1923
 * Probolomyrmex petiolatus Weber, 1940

Proceratium

 * Proceratium catio De Andrade, 2003
 * Proceratium colombicum De Andrade, 2003
 * Proceratium mancum Mann, 1922
 * Proceratium micrommatum Roger, 1863
 * Proceratium transitionis De Andrade, 2003

Procryptocerus

 * Procryptocerus attenuatus Smith F., 1876
 * Procryptocerus balzani Emery, 1894
 * Procryptocerus batesi Forel, 1899
 * Procryptocerus belti Forel, 1899
 * Procryptocerus carbonarius Mayr, 1870
 * Procryptocerus convexus Forel, 1904
 * Procryptocerus coriarius Mayr, 1870
 * Procryptocerus ferreri Forel, 1912
 * Procryptocerus hylaeus Kempf, 1951
 * Procryptocerus impressus Forel, 1899
 * Procryptocerus kempfi Longino & Snelling, 2002
 * Procryptocerus mayri Forel, 1899
 * Procryptocerus pictipes Emery, 1896
 * Procryptocerus rudis Mayr, 1870
 * Procryptocerus scabriusculus Forel, 1899
 * Procryptocerus schmitti Forel, 1901
 * Procryptocerus spiniperdus Forel, 1899
 * Procryptocerus subpilosus (Smith, 1860)
 * Procryptocerus tortuguero Longino and Snelling, 2002

Protalaridris

 * Protalaridris arhuaca Guerrero, Lattke & Alpert, 2018
 * Protalaridris armata Brown, 1980

Pseudomyrmex

 * Pseudomyrmex atripes Smith, F., 1860
 * Pseudomyrmex boopis Roger, 1863
 * Pseudomyrmex browni Kempf, 1967
 * Pseudomyrmex cubaensis Forel, 1901
 * Pseudomyrmex curacaensis Forel, 1912
 * Pseudomyrmex dendroicus Forel, 1904
 * Pseudomyrmex depressus Forel, 1906
 * Pseudomyrmex eduardi Forel, 1912
 * Pseudomyrmex elongatulus Dalla Torre, 1892
 * Pseudomyrmex elongatus Mayr, 1870
 * Pseudomyrmex euryblemma Forel, 1899
 * Pseudomyrmex excisus Mayr, 1870
 * Pseudomyrmex faber Smith, 1858
 * Pseudomyrmex ferox Ward, 1999
 * Pseudomyrmex filiformis Fabricius, 1804
 * Pseudomyrmex flavidulus Smith, F., 1858
 * Pseudomyrmex fortis (Forel, 1899)
 * Pseudomyrmex gebellii Forel, 1899
 * Pseudomyrmex godmani Forel, 1899
 * Pseudomyrmex gracilis Fabricius, 1804
 * Pseudomyrmex insuavis Ward, 1999
 * Pseudomyrmex ita (Forel, 1906)
 * Pseudomyrmex kuenckeli Emery, 1890
 * Pseudomyrmex laevigatus Smith, F., 1877
 * Pseudomyrmex laevivertex Forel, 1906
 * Pseudomyrmex lisus Enzmann, 1944
 * Pseudomyrmex mordax Warming, 1894
 * Pseudomyrmex oculatus Smith, F., 1855
 * Pseudomyrmex oki Forel, 1906
 * Pseudomyrmex pallens landolti Forel, 1912
 * Pseudomyrmex pallens Mayr, 1870
 * Pseudomyrmex pallidus Smith, F., 1855
 * Pseudomyrmex pictus (Stitz, 1913)
 * Pseudomyrmex rochai Forel, 1912
 * Pseudomyrmex rubiginosus (Stitz, 1913)
 * Pseudomyrmex sericeus Mayr, 1870
 * Pseudomyrmex simplex Smith, F., 1877
 * Pseudomyrmex spiculus Ward, 1989
 * Pseudomyrmex spinicola (Emery, 1890)
 * Pseudomyrmex subtilissimus Emery, 1890
 * Pseudomyrmex tenuis Fabricius, 1804
 * Pseudomyrmex tenuissimus Emery, 1906
 * Pseudomyrmex termitarius Smith, F., 1855
 * Pseudomyrmex triplaridis Forel, 1904
 * Pseudomyrmex triplarinus (Weddell, 1850)
 * Pseudomyrmex urbanus Smith, F., 1877
 * Pseudomyrmex venustus Smith, F., 1858
 * Pseudomyrmex viduus Smith, 1858
 * Pseudomyrmex vinneni (Forel, 1906)
 * Pseudomyrmex vitabilis Ward, 1999

Pseudoponera

 * Pseudoponera stigma Fabricius, 1804

Rasopone

 * Rasopone ferruginea (Smith, F. 1858)
 * Rasopone lunaris (Emery, 1896)
 * Rasopone rupinicola MacKay & Mackay, 2010

Rhopalothrix

 * Rhopalothrix ciliata Mayr, 1870

Rogeria

 * Rogeria alzatei Kugler, C., 1994
 * Rogeria belti Mann, 1922
 * Rogeria blanda (F. Smith, 1858)
 * Rogeria curvipubens Emery, 1894
 * Rogeria foreli Emery, 1894
 * Rogeria gibba Kugler, C., 1994
 * Rogeria innotabilis Kugler, C., 1994
 * Rogeria leptonana Kugler, C., 1994
 * Rogeria lirata Kugler, C., 1994
 * Rogeria merenbergiana Kugler, C., 1994
 * Rogeria micromma Kempf, 1961
 * Rogeria nevadensis Kugler, C., 1994
 * Rogeria scobinata Kugler, C., 1994
 * Rogeria tribrocca Kugler, C., 1994

Sericomyrmex

 * Sericomyrmex amabilis Wheeler, W.M., 1925
 * Sericomyrmex bondari Borgmeier, 1937
 * Sericomyrmex mayri Forel, 1912
 * Sericomyrmex opacus Mayr, 1865

Simopelta

 * Simopelta fernandezi MacKay & Mackay, 2008
 * Simopelta laevigata MacKay & Mackay, 2008
 * Simopelta laticeps 	Gotwald & Brown, 1967
 * Simopelta longirostris MacKay & Mackay, 2008
 * Simopelta mayri MacKay & Mackay, 2008
 * Simopelta paeminosa Snelling, R.R. 1971
 * Simopelta pergandei 	Forel, 1909
 * Simopelta transversa MacKay & Mackay, 2008
 * Simopelta williamsi Wheeler, W.M., 1935

Solenopsis

 * Solenopsis altinodis Forel, 1912
 * Solenopsis azteca Forel, 1893
 * Solenopsis basalis Forel, 1895
 * Solenopsis bicolor (Emery, 1906)
 * Solenopsis brevicornis Emery, 1888
 * Solenopsis corticalis Forel, 1881
 * Solenopsis decipiens Emery, 1906
 * Solenopsis franki Forel, 1908
 * Solenopsis gayi Spinola, 1851
 * Solenopsis geminata Fabricius, 1804
 * Solenopsis globularia Smith, F., 1858
 * Solenopsis hayemi Forel, 1908
 * Solenopsis laeviceps Mayr, 1870
 * Solenopsis picea Emery, 1896
 * Solenopsis pollux Forel, 1893
 * Solenopsis rugiceps Mayr, 1870
 * Solenopsis stricta Emery, 1896
 * Solenopsis subterranea Mackay, W.P. & Vinson, 1989
 * Solenopsis tenuis Mayr, 1878
 * Solenopsis texana Emery, 1895
 * Solenopsis vinsoni Pacheco & Mackay, 2013
 * Solenopsis virulens Smith, F., 1858
 * Solenopsis weyrauchi 	Trager, 1991
 * Solenopsis zeteki Wheeler, W.M., 1942

Stegomyrmex

 * Stegomyrmex manni Smith, M.R., 1946

Stenamma

 * Stenamma felixi Mann, 1922
 * Stenamma schmidti Menozzi, 1931

Strumigenys

 * Strumigenys alberti Forel, 1893
 * Strumigenys beebei Wheeler, W.M., 1915
 * Strumigenys biolleyi Forel, 1908
 * Strumigenys borgmeieri Brown, 1954
 * Strumigenys carinithorax Borgmeier, 1934
 * Strumigenys cassicuspis (Bolton, 2000)
 * Strumigenys castanea Brown, 1953
 * Strumigenys connectens Kempf, 1958
 * Strumigenys cordovensis Mayr, 1887
 * Strumigenys cosmostela Kempf, 1975
 * Strumigenys crassicornis Mayr, 1887
 * Strumigenys cultrigera Mayr, 1887
 * Strumigenys decipula (Bolton, 2000)
 * Strumigenys deltisquama Brown, 1957
 * Strumigenys denticulata Mayr, 1887
 * Strumigenys depressiceps Weber, 1934
 * Strumigenys dolichognatha Weber, 1934
 * Strumigenys doryceps (Bolton, 2000)
 * Strumigenys dyseides Bolton, 2000
 * Strumigenys eggersi Emery, 1890
 * Strumigenys elongata Roger, 1863
 * Strumigenys emmae (Emery, 1890)
 * Strumigenys enopla Bolton, 2000
 * Strumigenys fridericimuelleri Forel, 1886
 * Strumigenys gemella Kempf, 1975
 * Strumigenys godmani Forel, 1899
 * Strumigenys grytava Bolton, 2000
 * Strumigenys gundlachi Roger, 1862
 * Strumigenys hemidisca Brown, 1953
 * Strumigenys hyphata (Brown, 1953)
 * Strumigenys incuba Bolton, 2000
 * Strumigenys interfectiva Lattke & Goitia, 1997
 * Strumigenys kompsomala Bolton, 2000
 * Strumigenys laevipleura Kempf, 1958
 * Strumigenys lalassa Bolton, 2000
 * Strumigenys lanuginosa Wheeler, W.M., 1905
 * Strumigenys louisianae Roger, 1863
 * Strumigenys manis Bolton, 2000
 * Strumigenys margaritae Forel, 1893
 * Strumigenys marginiventris Santschi, 1931
 * Strumigenys membranifera Emery, 1869
 * Strumigenys metopia Brown, 1959
 * Strumigenys microthrix Kempf, 1975
 * Strumigenys monstra Bolton, 2000
 * Strumigenys nubila Lattke & Goitia, 1997
 * Strumigenys orchibia Brown, 1953
 * Strumigenys pariensis Lattke & Goitia, 1997
 * Strumigenys perissognatha (Bolton, 2000)
 * Strumigenys perparva Brown, 1958
 * Strumigenys precava Brown, 1954
 * Strumigenys raptans Bolton, 2000
 * Strumigenys rehi Forel, 1907
 * Strumigenys ruta Bolton, 2000
 * Strumigenys skia Bolton, 2000
 * Strumigenys smithii Forel, 1886
 * Strumigenys spathula Lattke & Goitia, 1997
 * Strumigenys subedentata Mayr, 1887
 * Strumigenys teratrix Bolton, 2000
 * Strumigenys trinidadensis Wheeler, W.M., 1922
 * Strumigenys trudifera Kempf & Brown, 1969
 * Strumigenys urrhobia Bolton, 2000
 * Strumigenys vartana Bolton, 2000
 * Strumigenys villiersi Perrault, 1986
 * Strumigenys wheeleriana Baroni Urbani, 2007
 * Strumigenys xenognatha Kempf, 1958
 * Strumigenys zeteki (Brown, 1959)

Syscia

 * Syscia tolteca Forel, 1909

Talaridris

 * Talaridris mandibularis Weber, 1941

Tapinoma

 * Tapinoma litorale Wheeler, W.M., 1905
 * Tapinoma melanocephalum Fabricius, 1793
 * Tapinoma ramulorum Emery, 1896

Tatuidris

 * Tatuidris tatusia Brown & Kempf, 1968

Technomyrmex

 * Technomyrmex fulvus (Wheeler, W.M., 1934)
 * Technomyrmex gorgona Fernández & Guerreo, 2008

Tetramorium

 * Tetramorium caldarium Roger, 1857

Thaumatomyrmex

 * Thaumatomyrmex atrox Weber, 1939
 * Thaumatomyrmex ferox Mann, 1922
 * Thaumatomyrmex manni Weber, 1939
 * Thaumatomyrmex paludis Weber, 1942
 * Thaumatomyrmex zeteki Smith, M.R. 1944

Tranopelta

 * Tranopelta gilva Mayr, 1866
 * Tranopelta subterranea (Mann, 1916)

Trichomyrmex

 * Trichomyrmex destructor (Jerdon, 1851)

Typhlomyrmex

 * Typhlomyrmex clavicornis Emery, 1906
 * Typhlomyrmex meire Lacau, Villemant & Delabie, 2004
 * Typhlomyrmex pusillus (Emery, 1894)
 * Typhlomyrmex rogenhoferi Mayr, 1862

Wasmannia

 * Wasmannia auropunctata

Uncertain species
The following species was described from Colombia but its true identity is currently unknown.
 * Cephalotes argentatus (Smith, F., 1853)