Crematogaster bandarensis

Inhabits developed forests, and nests in soil or in leaf litter.

Identification
Hosoishi and Ogata (2016) - A member of the Crematogaster quadriruga group. In the worker this species can be distinguished from all other members of the Crematogaster quadriruga group by the distinct compound eyes, V-shaped metanotal groove in lateral view, large propodeal spiracles touching metapleural gland bulla, and short propodeal spines. This species is similar to Crematogaster philippinensis and Crematogaster sundalandensis, but can be distinguished from them by the short and stout propodeal spines (PSL 0.05–0.08 vs. 0.08–0.11 and 0.08–0.11 in the latters).

Distribution
S. Thailand, Malaysia (Peninsula and Borneo), Brunei, Indonesia (Bali, Sumatra, Krakatau).

Distribution based on Regional Taxon Lists
Indo-Australian Region: Brunei Darussalam, Indonesia, Malaysia, Singapore. Oriental Region: Thailand.

Nomenclature

 * . Crematogaster biroi var. bandarensis Forel, 1913k: 76 (w.q.) INDONESIA (Sumatra).
 * Type-material: lectotype worker (by designation of Hosoishi & Ogata, 2016a: 575), paralectotype workers (number not stated), 1 paralectotype queen.
 * [Note: the syntype queen in the original description is not mentioned by Hosoishi & Ogata.]
 * Type-locality: lectotype Indonesia: Sumatra, Bandar Baroe (von Buttel-Reepen); paralectotypes with same data.
 * Type-depository: MHNG.
 * Combination in C. (Orthocrema): Emery, 1922e: 132.
 * Subspecies of biroi: Emery, 1922e: 132; Menozzi, 1935a: 113 (in key); Chapman & Capco, 1951: 97; Bolton, 1995b: 148.
 * Status as species: Hosoishi & Ogata, 2016a: 575 (redescription); Khachonpisitsak, et al. 2020: 86.
 * Senior synonym of andelis: Hosoishi & Ogata, 2016a: 575.
 * Distribution: Brunei, Malaysia (Peninsula, Sabah, Sarawak), Indonesia (Bali, Krakatau, Sumatra), Thailand.
 * andelis. Crematogaster biroi var. andelis Santschi, 1928h: 129 (w.) INDONESIA (Sumatra).
 * Type-material: 9 syntype workers.
 * Type-locality: Indonesia: Sumatra, Ile Varela (Pulu Berhala), 2.ix.1919 (J.B. Corporaal).
 * Type-depository: NHMB.
 * Stärcke, 1930: 373 (m.).
 * Subspecies of biroi: Stärcke, 1930: 373; Menozzi, 1935a: 113 (in key); Chapman & Capco, 1951: 97; Bolton, 1995b: 147.
 * Junior synonym of bandarensis: Hosoishi & Ogata, 2016a: 575.

Worker
Hosoishi and Ogata (2016) - (n = 10): HW 0.40–0.47; HL 0.42–0.50; CI 92–98; SL 0.34–0.39; SI 81–86; EL 0.09–0.11; PW 0.26–0.29; WL 0.48–0.54; PSL 0.05–0.08; PtL 0.14–0.17; PtW 0.12–0.15; PtH 0.10–0.12; PpL 0.09–0.11; PpW 0.13–0.16; PtHI 67–82; PtWI 80–88; PpWI 127–150; WI 100–117.

Workers monomorphic. Head subquadratic in full-face view. Mandibles with four teeth arranged at an equal distance, apical and subapical teeth large, basal two teeth smaller. Anterior clypeal margin convex in medial portion. Compound eyes slightly projecting beyond lateral margins of head in full-face view. Scapes reaching posterolateral corners of head.

Pronotal collar with weakly concave anterior margin in dorsal view, distinctly lower than pronotum in lateral view. Pronotal dorsum without distinct ridges laterally. Mesonotal dorsum with lateral ridges that irregularly extend posteriad to tips of propodeal spines. Pronotum and mesonotum in lateral view not clearly forming continuous dorsal outline; mesonotal dorsum flat. Metanotal groove in dorsal view transverse, almost straight in median portion, forming deep concavity that is laterally margined by lamellate ridges. Propodeal spiracles elliptical, situated at posterolateral corners of propodeum, touching to metapleural gland bullae. Propodeal spines developed, as long as diameter of propodeal spiracles, in dorsal view directed posteriad.

Petiole in dorsal view with subparallel sides, longer than wide. Posterior portion of petiole with short process that is slightly higher than posterior margin of petiole disc in lateral view. Subpetiolar process weakly developed as angulate tubercle. Postpetiole in lateral view with weakly convex dorsum, as high as petiole, in dorsal view as wide as petiole, weakly bilobed posteriorly but without longitudinal sulcus. Subpostpetiolar process undeveloped, but venter of postpetiole convex.

Integument essentially smooth and shining. Dorsal surface of head smooth and shining. Mandibles with feeble rugulae and smooth interspaces. Clypeus generally smooth and shining, but with one pair of longitudinal rugulae; rugulae not extending to posterior clypeal margin. Dorsal and lateral surfaces of pronotum smooth and shining; anterolateral shoulders of pronotum with rugulae. Mesopleura smooth and shining. Dorsal surface of propodeum smooth and shining, but one pair of rugulae running from metanotal groove to tips of propodeal spines. Dorsal surface pf petiole smooth. Lateral surface of petiole generally smooth, but with one longitudinal rugula. Dorsal and lateral surfaces of postpetiole smooth and shining.

Standing pilosity sparse. Dorsal face of head with suberect setae sparsely. Clypeus with two pairs of long setae in anterior portion, one directed upward and the other downward. Anterior clypeal margin with one pair of long setae medially and short setae laterally. Scapes with suberect setae. Mesosoma with five pairs of long erect and stout setae (ps1PN, ps2PN, psaMN, pspMN, and ps1PS) that are much longer than other erect setae. Posterolateral tubercles of petiole posteriorly with two pairs of long setae. Postpetiole with three pairs of long setae on disc anteriorly, posteriorly and laterally. Fourth abdominal tergite with erect to suberect setae abundantly, and short decumbent setae sparsely.

Body yellow. All flagellar segments yellow.

Type Material
Hosoishi and Ogata (2016) - Syntype workers, Bandar Baroe, Sumatra, Indonesia (v. Buttel-Reepen) (, examined). One syntype worker (middle specimen of three on one pin) in MHNG here designated Lectotype.

Determination Clarifications
This species corresponds to sp. 52 of SKY (Ito et al., 2001; Eguchi & Yamane, 2003).

References based on Global Ant Biodiversity Informatics

 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
 * Hosoichi S., and K. Ogata. 2016. Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae). Zoological Journal of the Linnean Society 176: 547–606.
 * Hosoishi S. and K. Ogata. 2009. A check list of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 32: 43-83.
 * Itino T., and S. Yamane. The vertical distribution of ants on canopy trees in a Bornean lowland rainf forest. Tropics 4(2/3): 277-281.
 * Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.