Checklist of Pachycondyla species

The following species and subspecies belong to the genus Pachycondyla. Synonyms are listed under their senior names.

A
aciculata Emery, 1901 (New Guinea)

acuta Emery, 1900 (New Guinea)

• Ponera dahlii Forel, 1901

aenescens Mayr, 1870 (Colombia)

aenigmatica (Arnold, 1949) (South Africa)

aequalis (Mann, 1919) (Solomon Is)

agilis (Forel, 1901) (United States)

• Neoponera golbachi Kusnezov, 1969

• Neoponera goyana Borgmeier, 1937

ambigua André, 1890 (Sierra Leone)

amblyops (Emery, 1887) (Indonesia (Sumatra))

amblyops oculatior (Forel, 1909) (Indonesia (Java))

analis (Latreille, 1802) (Guinea)

• Formica foetens Fabricius, 1793

• Megaponera dohrni Emery, 1902

• Ponera abyssinica Guérin-Méneville, 1849

• Ponera laeviuscula Gerstäcker, 1859

analis amazon (Santschi, 1935) (Ethiopia)

analis crassicornis (Gerstäcker, 1859) (Mozambique)

analis rapax (Santschi, 1914) (Tanzania)

analis subpilosa (Santschi, 1937) (Angola)

analis termitivora (Santschi, 1930) (Democratic Republic of Congo)

annamita (André, 1892) (Viet Nam)

annamita arcuata (Forel, 1900) (India)

antecurvata Mackay & Mackay, 2010 (Costa Rica)

apicalis (Latreille, 1802) (South America)

• Neoponera latreillei Forel, 1905

• Neoponera obscuricornis latocciput Forel, 1921

arabica (Collingwood & Agosti, 1996) (Yemen)

arhuaca (Forel, 1901) (Colombia)

astuta Smith, F., 1858 (Australia)

astuta obscura Karavaiev, 1935 (Vietnam)

atrata (Karavaiev, 1925) (Indonesia (Ambon Island))

australis (Forel, 1900) (Australia)

B
barbata Stitz, 1911 (Australia)

batak Yamane, 2007 (Indonesia (Sumatra))

becculata Mackay & Mackay, 2010 (Ecuador)

berthoudi (Forel, 1890) (South Africa)

berthoudi pubescens (Weber, 1942) (Sudan)

bispinosa Smith, F., 1858 (India (not Australia))

brevidorsa (Xu, 1994) (China)

breviscapa Mackay & Mackay, 2010 (Bolivia)

brunoi Forel, 1913 (Zimbabwe)

• Ectomomyrmex nigeriensis katangana Santschi, 1933

• Euponera bayoni Menozzi, 1933

• Euponera dentis Weber, 1942

• Euponera lamottei Bernard, 1953

• Euponera nigeriensis Santschi, 1914

bucki (Borgmeier, 1927) (Brazil)

bugabensis Forel, 1899 (Panama)

butteli (Forel, 1913) (Indonesia (Java))

butteli (Forel, 1913) (Indonesia (Sumatra))

C
caffraria (Smith, F., 1858) (South Africa)

• Ponera guineensis André, 1890

caffraria affinis (Santschi, 1935) (Congo)

caffraria caffra (Santschi, 1935) (Guinea)

cambouei (Forel, 1891) (Madagascar)

carbonaria (Smith, F., 1858) (Ecuador)

• Euponera atrovirens splendida Forel, 1901

• Pachycondyla atrovirens Mayr, 1866

carinulata (Roger, 1861) (Guyana)

• Neoponera carinulata gibbinota Forel, 1909

• Pachycondyla carinulata azteca Forel, 1899

cariosa (Emery, 1895) (Mozambique)

castanea (Mayr, 1865) (New Zealand)

• Euponera castanea striata Stitz, 1911

castaneicolor (Dalla Torre, 1893)

• Ponera castanea Smith, 1876

cavernosa (Roger, 1860) (South Africa)

cavernosa montivaga (Arnold, 1947) (South Africa)

cavimaculata Wang & Zhao, 2009 (China)

cavinodis (Mann, 1916) (Brazil)

cernua Mackay & Mackay, 2010 (Ecuador)

chinensis (Emery, 1895)

• Ponera solitaria Smith, F., 1874

christmasi (Donisthorpe, 1935) (Christmas Island)

chyzeri (Forel, 1907) (Colombia)

claudata (Menozzi, 1926) (Philippines)

cognata (Emery, 1896) (Costa Rica)

commutata (Roger, 1860) (South America)

comorensis (André, 1887) (Madagascar)

concava Mackay & Mackay, 2010 (Brazil)

conicula Mackay & Mackay, 2010 (Venezuela)

constricta (Mayr, 1884) (French Guiana)

• Ponera josephi Forel, 1886

constricticeps Mackay & Mackay, 2010 (Argentina)

cooki Mackay & Mackay, 2010 (Brazil)

coveri Mackay & Mackay, 2010 (Peru)

crassa (Emery, 1877) (Ethiopia)

crassa crassior (Santschi, 1930) (Kenya)

crassa gamzea Özdikmen, 2010

• Pachycondyla crassa ilgii Forel, 1910

crassicornis (Emery, 1897) (New Guinea)

crassinoda (Latreille, 1802) (Surinam)

crenata (Roger, 1861)

• Ponera pallipes Smith, F., 1858

cribrata Santschi, 1910 (Congo)

croceicornis (Emery, 1900)

• Euponera luteipes inops Forel, 1910

• Ponera bicolor Donisthorpe, 1949

curiosa Mackay & Mackay, 2010 (Brazil)

curvinodis Forel, 1899 (Guatemala)

D
darwinii (Forel, 1893) (Australia)

• Belonopelta darwinii indica Forel, 1900

• Cryptopone rufotestaceus Donisthorpe, 1943

• Euponera lamarki Santschi, 1913

darwinii africana (Forel, 1909) (Democratic Republic of Congo)

darwinii indica (Emery, 1899) (Myanmar)

• Belonopelta darwinii indica Forel

darwinii madecassa (Emery, 1899) (Madagascar)

denticulata (Kirby, W.F., 1896) (Australia)

depilis (Emery, 1902) (São Tomé & Principe (São Tomé Island))

dismarginata Mackay & Mackay, 2010 (Costa Rica)

donosoi Mackay & Mackay, 2010 (Ecuador)

dubitata (Forel, 1900) (Australia)

E
eleonorae (Forel, 1921) (Ecuador)

elisae (Forel, 1891) (Madagascar)

elisae divaricata (Emery, 1915) (Ethiopia)

elisae redbankensis (Forel, 1913) (Zimbabwe)

elisae rotundata (Emery, 1895) (South Africa)

• Ponera arnoldi Forel, 1913

emiliae (Forel, 1901) (Venezuela)

escherichi Forel, 1910 (Ethiopia)

exarata Emery, 1901 (New Guinea)

excavata (Emery, 1893) (Australia)

excavata acuticostata (Forel, 1900) (Australia)

F
fauveli Emery, 1896 (Bolivia)

ferruginea (Smith, F., 1858) (Mexico)

• Ponera ferruginea panamensis Forel, 1899

fiebrigi (Forel, 1912) (Panama)

fisheri Mackay & Mackay, 2010 (Panama)

flavipes Yamane, 2007 (Myanmar)

flavopilosa (Weber, 1942) (Sudan)

foetida (Linnaeus, 1758) ("America meridionali")

• Formica lobata De Geer, 1773

fossigera (Mayr, 1901) (South Africa)

fugax Forel, 1907 (Tanzania)

fusca Mackay & Mackay, 2010 (Colombia)

fusciceps (Emery, 1900) (New Guinea)

• Cryptopone mayri fuscior Mann, 1919

• Cryptopone mayri minor Mann, 1919

fuscoatra (Roger, 1861) (Colombia)

G
gigas (Wu & Wang, 1995) (China)

gilberti (Kempf, 1960) (Brazil)

gilloglyi Mackay & Mackay, 2010 (Mexico)

gilva (Roger, 1863) (United States)

• Euponera gilva harnedi Smith, M.R., 1929

• Euponera obsoleta Menozzi, 1931

• Ponera ochracea guatemalensis Forel, 1899

glabripes (Emery, 1893) (Philippines)

globularia Mackay & Mackay, 2010 (Bolivia)

goeldii (Forel, 1912) (Brazil)

• Neoponera lydiae Santschi, 1920

grandis (Donisthorpe, 1947) (Vietnam)

granosa (Roger, 1860) (South Africa)

guianensis (Weber, 1939) (Guyana)

• Wadeura haskinsi Weber, 1939

gulera Özdikmen, 2010 (Sudan)

• Ponera ambigua Weber, 1942

H
harpax (Fabricius, 1804) (South America)

• Pachycondyla harpax concinna Wheeler, W.M., 1925

• Pachycondyla harpax dibullana Forel, 1901

• Pachycondyla harpax irina Wheeler, W.M., 1925

• Pachycondyla montezumia Smith, F., 1858

• Ponera amplinoda Buckley, 1866

hartwigi (Arnold, 1948) (South Africa)

havilandi (Forel, 1901) (South Africa)

• Euponera sulcigera Mayr, 1904

havilandi fochi (Forel, 1918) (South Africa)

havilandi godfreyi (Arnold, 1926) (South Africa)

havilandi marleyi (Arnold, 1926) (South Africa)

havilandi Forel, 1901 (Singapore)

henryi Donisthorpe, 1942 (India)

hispida Mackay & Mackay, 2010 (Colombia)

holcotyle Mackay & Mackay, 2010 (Colombia)

holmgreni (Wheeler, W.M., 1925) (Peru)

hottentota Emery, 1886 (South Africa)

• Ophthalmopone lanceolata Mayr, 1895

I
ilgii (Forel, 1894) (Ethiopia)

impressa (Roger, 1861) (Colombia)

• Pachycondyla fuscoatra andicola Santschi, 1913

• Pachycondyla fuscoatra montana Forel, 1912

• Pachycondyla fuscoatra transversa Emery, 1890

inca Emery, 1901 (Peru)

• Pachycondyla fuscoatra cearensis Forel, 1901

incisa Emery, 1911 (New Guinea)

ingesta (Wheeler, W.M., 1922) (Democratic Republic of Congo)

insignis Mackay & Mackay, 2010 (Costa Rica)

insulana (Mayr, 1876) (Samoa)

insularis (Emery, 1889) (Indonesia (Java))

insularis brevior (Forel, 1901) (Borneo)

inversa (Smith, F., 1858) (South America)

J
javana (Mayr, 1867) (Indonesia (Java))

• Ectomomyrmex denticeps Wheeler, W.M., 1929

• Ectomomyrmex javanus maternus Forel, 1900

• Ectomomyrmex sundaicus Mayr, 1867

• Pachycondyla astuta cambodjana Forel, 1911

• Pachycondyla horni Forel, 1913

• Pachycondyla japonica Emery, 1902

jerdonii (Forel, 1900) (India)

jerdonii glabricollis (Stärcke, 1930) (Indonesia (Berhala Island))

jinxiuensis (Zhou, 2001) (China)

jonesii (Forel, 1891) (Madagascar)

K
karawaiewi Özdikmen, 2010 (Indonesia)

• Euponera luteipes arcuata Karavaiev, 1925

kenyensis (Santschi, 1937) (Kenya)

• Paltothyreus tarsatus striatus Santschi, 1930

kruegeri Forel, 1910 (South Africa)

kruegeri asina Santschi, 1912 (Kenya)

kruegeri rhodesiana Forel, 1913 (Zimbabwe)

L
laevigata (Smith, F., 1858) (Brazil)

• Euponera laevigata whelpleyi Wheeler, W.M., 1922

• Pachycondyla gagatina Emery, 1890

laevissima Arnold, 1915 (South Africa)

laevissima aspera (Arnold, 1962) (South Africa)

lamottei (Bernard, 1953) (Guinea)

latinoda Mackay & Mackay, 2010 (Brazil)

lattkei Mackay & Mackay, 2010 (Venezuela)

leeuwenhoeki (Forel, 1886) (India)

leeuwenhoeki jacobsoni Forel, 1915 (Indonesia (Simalur Island))

leeuwenhoeki sumatrensis Forel, 1901 (Indonesia (Sumatra))

lenis Kempf, 1961 (Brazil)

lenkoi Kempf, 1962 (Brazil)

leveillei (Emery, 1890) (Venezuela)

lineaticeps Mayr, 1866 (Mexico)

lobocarena Xu, 1995 (China)

longidentata Mackay & Mackay, 2010 (Colombia)

lunaris (Emery, 1896) (Paraguay)

lutea (Mayr, 1862) (Australia)

• Ectatomma socialis MacLeay, 1873

• Euponera lutea clara Crawley, 1915

luteipes (Mayr, 1862) (India (Nicobar Is))

luteipes continentalis (Karavaiev, 1925) (India)

luteola (Roger, 1861) (Brazil)

M
magnifica Borgmeier, 1929 (Brazil)

malayana (Wheeler, W.M., 1929) (Malaysia)

manni (Viehmeyer, 1924) (Solomon Is)

• Euponera robiginosa Donisthorpe, 1941

marginata (Roger, 1861) (Brazil)

mayri (Emery, 1887) (Australia)

melanaria (Emery, 1893) (Sri Lanka)

melanaria macra (Emery, 1894) (Seychelles Is)

melancholica Smith, F., 1865 (Indonesia (Morotai Island))

mesoponeroides (Radchenko, 1993) (Vietnam)

metanotalis Luederwaldt, 1918 (Brazil)

minuta Mackay & Mackay, 2010 (Mexico)

• Euponera rubra javana Forel, 1905

mirabilis Mackay & Mackay, 2010 (Bolivia)

mlanjiensis (Arnold, 1946) (Malawi)

mocquerysi (Emery, 1902) (São Tomé & Principe (São Tomé Island))

modiglianii Emery, 1900 (Indonesia (Sumatra))

moesta Mayr, 1870 (Colombia)

• Neoponera crenata confusa Santschi, 1921

• Neoponera mesonotalis Santschi, 1923

• Neoponera stipitum Forel, 1901

• Neoponera unidentata sulcatula Santschi, 1919

• Pachycondyla orizabana Norton, 1868

motschulskyi (Donisthorpe, 1943) (New Guinea)

myropola (Menozzi, 1925) (Philippines)

N
nakasujii Yashiro, Matsuura, Guénard, Terayama & Dunn, 2010 (Japan)

neonimba Özdikmen, 2010

• Euponera nimba Bernard, 1953

nicobarensis (Forel, 1905) (India (Nicobar Is))

nigrita (Emery, 1895) (Myanmar)

nigrita nigritella (Stärcke, 1930) (Indonesia (Sumatra))

novemdentata (Bernard, 1953) (Guinea)

O
oberthueri Emery, 1890 (Brazil)

obesa (Emery, 1897) (New Guinea)

obscurans (Walker, 1859) (Sri Lanka)

obscuricornis Emery, 1890 (Brazil)

obtusa Emery, 1900 (Borneo)

ochracea (Mayr, 1855) (Italy)

ochracea sicula (Emery, 1909) (Italy (Sicily))

oculata (Smith, F., 1858) (Australia)

overbecki Viehmeyer, 1916 (Singapore)

P
pachyderma Emery, 1901 (Cameroon)

pachyderma attenata Santschi, 1920 (Democratic Republic of Congo)

pachyderma funerea (Wheeler, W.M., 1922) (Democratic Republic of Congo)

pachyderma postsquamosa Santschi, 1920 (Congo)

pachynoda (Clark, 1930) (Australia)

pallidipennis (Smith, F., 1860) (Indonesia (Sulawesi))

papuana (Viehmeyer, 1914) (New Guinea)

• Euponera pulchella Donisthorpe, 1941

• Euponera viehmeyeri Donisthorpe, 1948

pergandei (Forel, 1909) (Guatemala)

peringueyi (Emery, 1899) (South Africa)

peringueyi saldanhae (Arnold, 1951) (South Africa)

perroti (Forel, 1891) (Madagascar)

perroti admista (Forel, 1892) (Madagascar)

picardi Forel, 1901 (Angola)

picea (Bernard, 1953) (Guinea)

pilidorsalis Yamane, 2007 (West Malaysia)

piliventris Smith, F., 1858 (Australia)

piliventris intermedia (Forel, 1900) (Australia)

piliventris regularis Forel, 1907 (Australia)

pilosior (Wheeler, W.M., 1928) (Japan)

• Euponera chosonensis Teranishi, 1940

porcata (Emery, 1897) (Australia)

procidua Emery, 1890 (Surinam)

pseudogigas (Zhou & Zheng, 1997) (China)

pumicosa (Roger, 1860) (South Africa)

pumicosa sculpturata Santschi, 1912 (Zimbabwe)

punctata Karavaiev, 1935 (Vietnam)

purpurascens Forel, 1899 (Costa Rica)

R
recava Mackay & Mackay, 2010 (Colombia)

recticlypea (Xu, 1998) (China)

rostrata Emery, 1890 (Venezuela)

rotundiceps (Emery, 1914) (New Caledonia)

• Ponera mjobergi Forel, 1915

rubescens (Santschi, 1937) (Democratic Republic of Congo)

rubiginosa (Emery, 1889) (Myanmar)

rubra (Smith, F., 1857) (Singapore)

rubra minirubra Özdikmen, 2010

• Euponera rubra javana Forel, 1905

ruficornis (Clark, 1934) (Australia)

rufipes (Jerdon, 1851) (India)

rufipes ceylonensis Forel, 1911 (Sri Lanka)

rufonigra (Clark, 1934) (Australia)

• Euponera clarki Wheeler, W.M., 1934

rugosula (Emery, 1902) (Peru)

• Neoponera laevinodis André, 1902

rupinicola Mackay & Mackay, 2010 (Colombia)

S
sakishimensis Terayama, 1999 (Japan)

sandakana (Wheeler, W.M., 1919) (Borneo)

sanguinea Santschi, 1920 (Democratic Republic of Congo)

sauteri Forel, 1912 (Taiwan)

sauteri Wheeler, W.M., 1906 (Japan)

schoedli Mackay & Mackay, 2006 (Ecuador)

schultzi Mackay & Mackay, 2010 (Brazil)

scobina (Wilson, 1958)

• Bothroponera striata Donisthorpe, 1949

scolopax (Emery, 1899) (Cameroun)

senegalensis (Santschi, 1914) (Senegal)

sennaarensis (Mayr, 1862) (Sudan)

• Ponera sorghi Roger, 1863

sennaarensis decolor (Santschi, 1921) (Congo)

sennaarensis ruginota (Stitz, 1916) (Cameroun)

sharpi (Forel, 1901) (Singapore)

sheldoni (Mann, 1919) (Solomon Is)

sikorae (Forel, 1891) (Madagascar)

silvestrii Santschi, 1914 (Ghana)

silvestrii nimba (Bernard, 1953) (Guinea)

simillima (Donisthorpe, 1949) (New Guinea)

sinensis (Wang, M., 1992) (China)

sjostedti (Mayr, 1896) (Cameroun)

solisi Mackay & Mackay, 2010 (Costa Rica)

solitaria (Smith, F., 1860) (Indonesia (Batjan Island))

soror (Emery, 1899) (Cameroun)

soror ancilla (Emery, 1899) (Congo)

soror suturalis Forel, 1907 (Ethiopia)

stigma (Fabricius, 1804)

• Euponera brunneus Donisthorpe, 1948

• Euponera nixoni Donisthorpe, 1943

• Euponera sexdentatus Donisthorpe, 1949

• Euponera stigma rufescens Wheeler, W.M. & Mann, 1914

• Ponera americana Mayr, 1862

• Ponera myopa Roger, 1860

• Ponera quadridentata Smith, F., 1859

• Ponera stigma attrita Forel, 1893

striata Smith, F., 1858 (Brazil)

• Pachycondyla striata nitidiventris Santschi, 1921

striatinodis Emery, 1890 (Costa Rica)

• Neoponera bakeri Mann, 1916

• Neoponera rugosinodis André, 1902

striatula Karavaiev, 1935

• Pachycondyla striata Karavaiev, 1927

strigulosa (Emery, 1895) (South Africa)

• Pachycondyla berthoudi Forel, 1901

striolata Donisthorpe, 1933 (India)

subiridescens (Wheeler, W.M., 1922) (Democratic Republic of Congo)

sublaevis (Emery, 1887) (Australia)

sublaevis kurandensis Forel, 1910 (Australia)

sublaevis murina Forel, 1910 (Australia)

sublaevis reticulata (Forel, 1900) (Australia)

sublaevis rubicunda (Emery, 1893) (Australia)

succedanea (Roger, 1863) (Cuba)

• Euponera cauta Mann, 1922

• Euponera stigma compressinodis Borgmeier, 1928

sulcata (Mayr, 1867) (("Vaterland Unbekannt") Emery, 1911: 78 gives India)

sulcata fossulata (Forel, 1900) (India)

sulcata sulcatotesserinoda (Forel, 1900) (India)

sumatrana Özdikmen, 2010

• Pachycondyla sculpturata Karavaiev, 1925

suspecta (Santschi, 1914) (Tanzania)

T
taivanae (Forel, 1913) (Taiwan)

• Euponera takahashii Wheeler, W.M., 1930

talpa (André, 1890) (Sierra Leone)

• Psalidomyrmex clavicornis Bernard, 1953

talpa variolata Santschi, 1912 (Congo)

tarsata (Fabricius, 1798) (Senegal)

• Pachycondyla simillima Smith, F., 1858

• Ponera gagates Guérin-Méneville, 1844

• Ponera pestilentia Smith, F., 1858

• Ponera spiniventris Smith, F., 1858

tarsata delagoensis (Emery, 1899) (Mozambique)

tarsata kaya Özdikmen, 2010

• Paltothyreus tarsatus striatus Santschi, 1930

tarsata mediana (Santschi, 1919) (Congo)

tarsata robusta (Santschi, 1919) (Somalia)

tarsata striatidens (Santschi, 1919) (Kenya)

tarsata subopaca (Santschi, 1919) (Gabon)

tengu (Terayama, 1999) (Japan)

terminalis Özdikmen, 2010 (Borneo)

• Ponera apicalis Smith, F., 1857

tesseronoda (Emery, 1877) (India)

testacea (Bernard, 1953) (Guinea)

testacea (Emery, 1893) (Sri Lanka)

• Cryptopone emeryi Donisthorpe, 1943

• Cryptopone mayri Mann, 1919

• Cryptopone sarawakana Wheeler, W.M., 1933

• Ponera anommata Donisthorpe, 1948

theresiae Forel, 1899 (Panama)

tianzun Terayama, 2009 (Taiwan)

tonkina Santschi, 1920 (Vietnam)

tridentata Smith, F., 1858 (Borneo)

tridentata debilior (Forel, 1901) (Borneo)

tridentata exasperans Forel, 1911 (West Malaysia)

typhlos (Karavaiev, 1935) (Vietnam)

U
unicolor (Smith, F., 1860) (Indonesia (Sulawesi))

unidentata Mayr, 1862 (Brazil)

• Neoponera unidentata eburneipes Wheeler, W.M. & Bequaert, 1929

• Neoponera unidentata maya Wheeler, W.M. & Bequaert, 1929

• Neoponera unidentata trinidadensis Wheeler, W.M. & Bequaert, 1929

V
variolosa (Arnold, 1947) (South Africa)

venusta (Forel, 1912) (Brazil)

verecundae (Donisthorpe, 1943) (New Guinea)

verenae (Forel, 1922) (Panama)

vermiculata (Emery, 1897) (Indonesia)

vidua (Smith, F., 1857) (Borneo)

vieirai Mackay & Mackay, 2010 (Ecuador)

villiersi (Bernard, 1953) (Guinea)

villosa (Fabricius, 1804) (Central America)

• Ponera bicolor Guérin-Méneville, 1844

• Ponera pedunculata Smith, F., 1858

• Ponera pilosa Smith, F., 1858

W
wallacei Yamane, 2007 (Indonesia (Lombok Island))

wasmannii (Forel, 1887) (Madagascar)

weberi (Bernard, 1953) (Guinea)

williamsi (Wheeler, W.M. & Chapman, 1925) (Philippines)

wroughtonii (Forel, 1901) (South Africa)

wroughtonii crudelis (Forel, 1901) (South Africa)

Z
zhengi Xu, 1995 (China)

zumpti (Santschi, 1937) (Cameroun)

zuparkoi Mackay & Mackay, 2010 (Peru)