Tetramorium sada

Tetramorium sada lives in tropical dry forests, sometimes on tsingy, and was mainly collected from the leaf litter or lower vegetation.

Identification
A member of the Tetramorium bonibony-species group

The following character set separates T. sada from all other species in the T. bonibony group: mesosoma with well developed anterior face of pronotum but without an anterodorsal median protuberance; petiolar node in profile triangular nodiform, strongly anteroposteriorly compressed dorsally, and highly transverse in dorsal view (LPeI 25 - 30; DPeI 400 - 463); postpetiole distinctly narrower than petiolar node (PPI 89 - 95); distinct bicolouration with dark brown head and mesosoma contrasting with yellow waist segments and gaster. (Hita Garcia and Fisher 2012)

Distribution
The distribution of T. sada is limited to western Madagascar. It seems to common from Tsingy de Bemaraha to Namoroka, but further north it is only known from Ankarafantsika.

Distribution based on Regional Taxon Lists
Malagasy Region: Madagascar.

Nomenclature

 *  sada. Tetramorium sada Hita Garcia & Fisher, 2012: 47, figs. 30, 37, 38, 60-62 (w.) MADAGASCAR.

Worker
HL 0.70 - 0.84 (0.79); HW 0.70 - 0.86 (0.80); SL 0.46 - 0.58 (0.54); EL 0.16 - 0.19 (0.18); PH 0.41 - 0.52 (0.48); PW 0.50 - 0.64 (0.59); WL 0.85 - 1.09 (1.01); PSL 0.30 - 0.37 (0.34); PTL 0.07 - 0.09 (0.08); PTH 0.26 - 0.34 (0.30); PTW 0.30 - 0.39 (0.36); PPL 0.21 - 0.25 (0.23); PPH 0.27 - 0.37 (0.33); PPW 0.27 - 0.37 (0.33); CI 100 - 103 (102); SI 65 - 70 (68); OI 21 - 22 (22); DMI 56 - 60 (58); LMI 46 - 49 (48); PSLI 42 - 45 (43); PeNI 59 - 63 (61); LPeI 25 - 30 (28); DPeI 400 - 463 (425); PpNI 54 - 60 (56); LPpI 67 - 76 (70); DPpI 132 - 152 (142); PPI 89 - 95 (92) (12 measured).

Head as wide as long to weakly wider (CI 100 - 103). Anterior clypeal margin with median impression. Frontal carinae moderately developed, ending between posterior eye margin and posterior head margin. Antennal scrobes absent. Antennal scapes short, not reaching posterior head margin (SI 65 - 70). Eyes comparatively small (OI 21 - 22). Mesosoma with well developed anterior face of pronotum but without an anterodorsal median protuberance, mesosoma weakly to moderately marginate from lateral to dorsal mesosoma; promesonotal suture and metanotal groove absent; mesosoma very high, compact, and stout (LMI 46 - 49). Propodeal spines very long, spinose, and acute (PSLI 42 - 45); propodeal lobes small and triangular. Petiolar node in profile triangular cuneiform and strongly anteroposteriorly compressed dorsally, approximately 3.3 to 3.9 times higher than long (LPeI 25 - 30), anterior and posterior faces not parallel, node in dorsal view highly transverse and very thin, between 4.0 to 4.6 times wider than long (DPeI 400 - 463). Postpetiole in profile approximately rounded, approximately 1.3 to 1.5 times higher than long (LPpI 67 - 76), in dorsal view approximately 1.3 to 1.5 times wider than long (DPpI 132 - 152). Postpetiole in profile more voluminous than petiolar node, in dorsal view distinctly less broad than petiolar node, only approximately 0.9 times width of petiolar node (PPI 89 - 95). Mandibles striate; clypeus with one distinct median longitudinal ruga and one or two often weaker rugae at each side; cephalic dorsum between frontal carinae anteriorly and centrally longitudinally rugose and posteriorly reticulate-rugose, dorsum with five to nine longitudinal rugae, rugae ending close to posterior head margin but often broken or with cross-meshes, always with one well-developed longitudinal median ruga, median ruga running from posterior head margin to posterior clypeal margin, approximately at eye level diverging into two rugae leading to posterior clypeal margin; lateral and ventral head mainly reticulate-rugose to longitudinally rugose. Ground sculpture on head weakly to moderately reticulate-punctate. Lateral mesosoma reticulate-rugose to irregularly longitudinally rugose; dorsal mesosoma anteriorly strongly reticulate-rugose, posteriorly reticulate-rugose to longitudinally rugose. Waist segments and gaster unsculptured, smooth, and shiny. All dorsal surfaces of body with abundant, moderately long, fine, and erect pilosity. Body distinctly bicoloured, head and mesosoma brown to dark brown, waist segments and gaster yellow, appendages of lighter brown than head and mesosoma.

Type Material
Holotype worker, MADAGASCAR, Mahajanga, Parc National Tsingy de Bemaraha, 3.4 km 93° E Bekopaka, Tombeau Vazimba, 19.14194 S, 44.828 E, 50 m, tropical dry forest, beating low vegetation, collection code BLF4234, 6.-10.XI.2001 (B.L. Fisher, C. Griswold et al.) (: CASENT0443274). Paratypes, 49 workers with same data as holotype (: CASENT0443394; CASC: CASENT0443271; CASENT0443273; CASENT0443277; CASENT0443280; CASENT0443330; CASENT0443304; CASENT0443311; CASENT0443312; CASENT0443313; CASENT0443342; CASENT0443345; CASENT0443362; CASENT0443363; CASENT0443377; CASENT0443382; CASENT0443383; CASENT0443384; CASENT0443385; CASENT0443398; CASENT0443399; CASENT0443422; CASENT0443433; CASENT0443438; : CASENT0443364; : CASENT0443439; : CASENT0443381).

Etymology
The name of the new species is Malagasy and means "mixture of colour". It refers to the distinctive bicolouration of the new species. The species name is a noun in apposition, and thus invariant.