Checklist of Crematogaster species

The following species and subspecies belong to the genus Crematogaster. Synonyms are listed under their senior names. For valid names only see Crematogaster species and for distribution information see Crematogaster species by Country.

A
abdominalis Motschoulsky, 1863 (India)

aberrans Forel, 1892 (China, India, Thailand)

• Crematogaster aberrans assmuthi Forel, 1913

• Crematogaster aberrans inglebyi Forel, 1902

• Crematogaster soror Forel, 1902

abrupta Mann, 1919 (Solomon Islands)

abstinens Forel, 1899 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guyana, Mexico, Panama, Venezuela)

• Crematogaster abstinens chacoana Santschi, 1933

• Crematogaster abstinens suturalis Forel, 1912

• Crematogaster ornatipilis Wheeler, W.M., 1918

• Crematogaster quadriformis erici Santschi, 1929

acaciae Forel, 1892 (Ethiopia, Namibia, Oman, Saudi Arabia, Sudan, United Republic of Tanzania, Yemen, Zimbabwe)

acaciae generosa Santschi, 1919 (South Africa)

acaciae gloriosa Santschi, 1914 (Democratic Republic of Congo)

acaciae victoriosa Santschi, 1916 (Zambia, Zimbabwe)

aculeata Donisthorpe, 1941 (New Guinea)

acuta (Fabricius, 1804) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Venezuela)

• Crematogaster acuta centralis Santschi, 1932

• Crematogaster quadriceps Smith, F., 1858

aegyptiaca Mayr, 1862 (Egypt, Eritrea, Oman, Saudi Arabia, United Arab Emirates)

aegyptiaca pharaonis Santschi, 1914 (Egypt)

aegyptiaca robusta Emery, 1877 (Eritrea, Ethiopia)

aegyptiaca turkanensis Santschi, 1935 (Kenya)

afghanica Pisarski, 1967 (Afghanistan, Iran)

africana Mayr, 1895 (Cameroun, Ivory Coast, Kenya)

africana alligatrix Forel, 1911 (Nigeria)

africana biemarginata Forel, 1910 (Cameroun)

africana camena Wheeler, W.M., 1922 (Democratic Republic of Congo)

africana polymorphica Weber, 1943 (Sudan)

africana schumanni Mayr, 1895 (Cameroun)

africana stanleyi Wheeler, W.M., 1922 (Congo)

africana stolonis Santschi, 1937 (Democratic Republic of Congo)

africana thoracica Santschi, 1921 (Democratic Republic of Congo)

africana tibialis Santschi, 1922 (Democratic Republic of Congo)

africana variegata Mayr, 1902 (Cameroun)

agnetis Forel, 1892 (Madagascar)

agniae Karavaiev, 1935 (Vietnam)

alafara Blaimer, 2013 (Madagascar)

algirica (Lucas, 1849) (Algeria)

alluaudi Emery, 1893 (Canary Islands)

alluaudi noualhieri Emery, 1893 (Canary Islands)

aloysiisabaudiae Menozzi, 1930 (Somalia)

alulai Emery, 1901 (Eritrea, Ethiopia)

alulai scrutans Forel, 1910 (Ethiopia)

amabilis Santschi, 1911 (Kenya, United Republic of Tanzania)

amabilis retiaria Santschi, 1933 (Kenya)

amapaensis Kempf, 1960 (Brazil)

ambigua Santschi, 1926 (Democratic Republic of Congo)

amita Forel, 1913 (Zimbabwe)

amita bushimana Santschi, 1926 (South Africa)

amita caffra Santschi, 1926 (South Africa)

amita makololo Santschi, 1926 (Zimbabwe)

amita matabele Arnold, 1920 (Zimbabwe)

ampla Forel, 1912 (Bolivia, Brazil, Colombia, Mexico, Panama, Paraguay)

ampullaris Smith, F., 1861 (Indonesia, Philippines, Sulawesi)

ancipitula Forel, 1917 (Angola)

angulosa André, 1896 (Borneo, Indonesia, Malaysia)

angusticeps Santschi, 1911 (Mali, Sudan)

antaris Forel, 1894 (Algeria, Iran, Israel, Oman, Qatar, Saudi Arabia, United Arab Emirates, Yemen)

• Crematogaster auberti sordida Forel, 1909

anthracina Smith, F., 1857 (Borneo, China, India, Indonesia, Malaysia, Singapore, Sri Lanka)

apicalis Motschoulsky, 1863 (Sri Lanka)

arata Emery, 1906 (Argentina, Brazil, Paraguay)

arcuata Forel, 1899 (Bolivia, Brazil, Colombia, Costa Rica, Panama, Panama, Peru, Venezuela)

• Crematogaster arcuata aruga Forel, 1913

arnoldi Forel, 1914 (South Africa)

arnoldi loveridgei Santschi, 1926 (United Republic of Tanzania)

aroensis Menozzi, 1935 (New Guinea)

arthurimuelleri Forel, 1894 (Mozambique)

ashmeadi Mayr, 1886 (Bahamas, United States)

• Crematogaster ashmeadi matura Wheeler, W.M., 1932

atra Mayr, 1870 (Argentina, Brazil, Mexico, Venezuela)

auberti Emery, 1869 (Algeria, Balearic Islands, Bulgaria, Egypt, France, Gibraltar, Greece, Iberian Peninsula, Iran, Iraq, Israel, Malta, Montenegro, Morocco, Portugal, Republic of Macedonia, Saudi Arabia, Spain, Turkey)

• Crematogaster auberti iberica Forel, 1909

auberti levithorax Forel, 1902 (Algeria, Canary Islands)

auberti nigripes Menozzi, 1940 (Israel, Libya)

auberti regilla Santschi, 1937 (Morocco)

auberti savinae Zimmermann, 1935 (Montenegro, Republic of Macedonia, Slovakia)

auberti vogti Forel, 1909 (Spain)

augusti Emery, 1895 (Indonesia)

aurita Karavaiev, 1935 (Cambodia, Laos, Thailand, Vietnam)

australis Mayr, 1876 (Australia)

australis chillagoensis Forel, 1915 (Australia)

australis sycites Forel, 1916 (Australia)

B
baduvi Forel, 1912 (Brunei Darussalam, Indonesia, Krakatau Islands, Malaysia)

bakeri Menozzi, 1925 (Philippines)

bandarensis Forel, 1913 (Brunei Darussalam, Indonesia, Malaysia, Thailand)

• Crematogaster biroi andelis Santschi, 1928

bara Blaimer, 2013 (Madagascar)

batesi Forel, 1911 (Nigeria)

bequaerti Forel, 1913 (Democratic Republic of Congo, Guinea)

bequaerti atraplex Santschi, 1922 (Democratic Republic of Congo)

bequaerti gerardi Santschi, 1915 (Democratic Republic of Congo)

bequaerti ludia Forel, 1913 (Democratic Republic of Congo)

bequaerti modica Santschi, 1926 (Kenya, Uganda)

• Crematogaster bequaerti saga Santschi, 1937

bequaerti mutabilis Santschi, 1914 (Kenya)

bequaerti semiclara Santschi, 1928 (Zimbabwe)

betapicalis Bolton, 1995 (India)

• Crematogaster apicalis Smith, F. Smith, F., 1878

bicolor Smith, F., 1860 (Indonesia)

bicolor imbellis Emery, 1893 (Philippines)

biformis André, 1892 (Borneo, Indonesia, Malaysia)

binghamii Forel, 1904 (Bangladesh, China, India, Nepal, Thailand, Vietnam)

bingo Forel, 1908 (Argentina, Brazil)

biroi Mayr, 1897 (China, India, Pakistan, Sri Lanka, Taiwan)

• Crematogaster aitkenii Forel, 1902

• Crematogaster biroi smythiesii Forel, 1902

• Crematogaster urvijae Bharti, 2003

bison Forel, 1913 (Taiwan)

boera Santschi, 1926 (Zimbabwe)

bogojawlenskii Ruzsky, 1905 (Armenia, Georgia, Iran, Kyrgyzstan, Turkmenistan, Uzbekistan)

• Crematogaster sordidula kosti Ruzsky, 1905

boliviana (Wheeler, W.M., 1922) (Bolivia)

borneensis André, 1896 (Borneo, Indonesia, Malaysia, Singapore)

• Crematogaster borneensis capax Forel, 1911

• Crematogaster borneensis hosei Forel, 1911

• Crematogaster borneensis macarangae Viehmeyer, 1916

• Crematogaster borneensis sembilana Forel, 1911

bouvardi Santschi, 1920 (Laos, Vietnam)

brasiliensis Mayr, 1878 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Trinidad and Tobago)

• Crematogaster armandi Forel, 1921

• Crematogaster brasiliensis cocciphila Borgmeier, 1934

• Crematogaster inca Wheeler, W.M., 1925

• Crematogaster limata ludio Forel, 1912

brasiliensis arawak Weber, 1938 (Guyana)

brevidentata Forel, 1912 (Barbados, Brazil)

brevimandibularis Donisthorpe, 1943 (New Guinea)

brevis Emery, 1887 (Indonesia)

brevispina Hosoishi & Ogata, 2016 (Philippines)

breviventris Santschi, 1920 (Cameroun)

bruchi Forel, 1912 (Argentina, Paraguay)

brunensis Hosoishi & Ogata, 2016 (Brunei Darussalam)

brunnea Smith, F., 1857 (Borneo, China, India, Indonesia, Malaysia, Sri Lanka)

brunnea contemta Mayr, 1879 (Bangladesh, China, India, Sri Lanka)

brunnea latipetiolata Karavaiev, 1935 (Vietnam)

brunnea nicevillei Emery, 1922 (China, India, Sri Lanka)

brunnea nilgirica Emery, 1922 (India)

brunnea rabula Forel, 1902 (India, Sri Lanka)

brunnea ruginota Santschi, 1928 (China, India, Philippines)

brunnea sundaica Forel, 1909 (Indonesia)

brunneipennis André, 1890 (Sierra Leone)

brunneipennis omniparens Forel, 1914 (South Africa)

brunneipennis yorubosa Santschi, 1933 (Nigeria)

brunnescens Motschoulsky, 1863 (Sri Lanka)

bryophilia Longino, 2003 (Colombia, Costa Rica, Ecuador, Panama, Venezuela)

buchneri Forel, 1894 (Angola, Ghana)

• Crematogaster halli Donisthorpe, 1945

buchneri composita Santschi, 1933 (Democratic Republic of Congo)

buchneri graeteri Forel, 1916 (Democratic Republic of Congo)

buchneri uasina Santschi, 1935 (Kenya)

buddhae Forel, 1902 (India)

butteli Forel, 1913 (Indonesia)

C
capensis Mayr, 1862 (South Africa)

capensis calens Forel, 1910 (Mozambique)

capensis tropicorum Forel, 1910 (Mozambique)

captiosa Forel, 1911 (Borneo, Indonesia, Malaysia)

• Crematogaster borneensis harpyia Forel, 1911

• Crematogaster borneensis insulsa Forel, 1911

• Crematogaster borneensis novem Forel, 1911

• Crematogaster borneensis symbia Forel, 1911

carinata Mayr, 1862 (Barbados, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Lesser Antilles, Mexico, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Crematogaster limata parabiotica Forel, 1904

castanea Smith, F., 1858 (Comoros, Eritrea, Kenya, Madagascar, Mauritius, Mayotte, Namibia, South Africa, Uganda, Zimbabwe)

• Crematogaster tricolor decolor Forel, 1891

castanea adusta Santschi, 1935 (Namibia)

castanea analis Santschi, 1910 (Congo)

castanea aquila Forel, 1907 (Kenya)

castanea arborea Smith, F., 1858 (South Africa)

castanea bruta Santschi, 1926 (Kenya)

castanea busschodtsi Emery, 1899 (Democratic Republic of Congo, Kenya)

castanea durbanensis Forel, 1914 (South Africa)

castanea ferruginea Forel, 1892 (Eritrea, Kenya, Somalia)

castanea hararica Forel, 1894 (Eritrea, Ethiopia)

castanea insidiosa Santschi, 1920 (Namibia)

castanea inversa Forel, 1907 (Kenya)

castanea mediorufa Forel, 1907 (Kenya)

castanea museisapientiae Forel, 1911

castanea rufimembrum Santschi, 1914 (United Republic of Tanzania)

castanea rufonigra Emery, 1895 (South Africa)

castanea simia Forel, 1910 (Botswana)

castanea ulugurensis Forel, 1911 (Kenya)

castanea yambatensis Forel, 1913 (Democratic Republic of Congo)

celebensis Hosoishi & Ogata, 2016 (Indonesia)

censor Forel, 1910 (Senegal)

censor junodi Forel, 1916 (South Africa)

cephalotes Smith, F., 1857 (Borneo, Indonesia, Malaysia)

cerasi (Fitch, 1855) (Canada, Mexico, United States)

• Crematogaster browni Buren, 1968

• Crematogaster kennedyi Wheeler, W.M., 1930

chhangi Hosoishi & Ogata, 2014 (Cambodia)

chiarinii Emery, 1881 (Eritrea, Ethiopia, Gambia, Kenya, Oman, Saudi Arabia, Somalia, Yemen)

• Crematogaster chiarinii affabilis Forel, 1907

chiarinii aethiops Forel, 1907 (Somalia)

chiarinii bayeri Santschi, 1926 (Kenya)

chiarinii cincta Emery, 1896 (Somalia)

chiarinii nigra Forel, 1910 (Congo)

chiarinii sellula Santschi, 1914 (United Republic of Tanzania)

chiarinii subsulcata Santschi, 1914 (Sudan)

chiarinii taediosa Forel, 1894 (Eritrea, Ethiopia)

chiarinii v-nigra Forel, 1910 (Democratic Republic of Congo)

chlorotica Emery, 1899 (Benin, Cameroun, Mali, Sierra Leone)

chodati Forel, 1921 (Paraguay)

chopardi Bernard, 1950 (Niger)

chungi Brown, 1949 (China)

cicatriculosa Roger, 1863 (Senegal, Sierra Leone, South Africa)

cisplatinalis Mayr, 1887 (Argentina, Brazil, Paraguay, Uruguay)

clariventris Mayr, 1895 (Cameroun, Congo, Guinea, Kenya, Uganda)

clariventris biimpressa Mayr, 1895 (Congo)

claudiae Feldhaar, Maschwitz & Fiala, 2016 (Borneo, Malaysia)

clydia Forel, 1912 (Brazil)

coarctata Mayr, 1870 (Mexico, United States)

• Crematogaster coarctata mormonum Wheeler, W.M., 1919

• Crematogaster laeviuscula californica Wheeler, W.M., 1919

coelestis Santschi, 1911 (Guinea, Senegal)

coelestis mercatori Santschi, 1939 (Guinea)

concava Emery, 1899 (Cameroun, Kenya, Sierra Leone)

consternens (Walker, 1859) (Sri Lanka)

constructor Emery, 1895 (South Africa, Zimbabwe)

constructor kirbyella Emery, 1922 (South Africa)

• Crematogaster constructor kirbyi Mayr, 1895

coriaria Mayr, 1872 (Borneo, Indonesia, Malaysia, Thailand)

• Crematogaster coriaria fraudatrix Forel, 1911

• Crematogaster spengeli Forel, 1912

• Crematogaster spengeli taipingensis Forel, 1913

coriaria compacta Crawley, 1924 (Indonesia)

coriaria picata Emery, 1922 (Malaysia)

cornigera Forel, 1902 (Australia)

cornuta Crawley, 1924 (Indonesia)

corporaali Santschi, 1928 (Indonesia)

corticicola Mayr, 1887 (Argentina, Bolivia, Brazil, Paraguay)

• Crematogaster distans rugiceps Forel, 1911

corvina Mayr, 1870 (Honduras, Mexico)

crassicornis Emery, 1893 (Philippines)

crinosa Mayr, 1862 (Argentina, Barbados, Belize, Bolivia, Brazil, Chile, Colombia, Costa Rica, Ecuador, French Guiana, Greater Antilles, Grenada, Guatemala, Guyana, Honduras, Lesser Antilles, Martinique, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States, Uruguay, Venezuela)

• Crematogaster brevispinosa Mayr, 1870

• Crematogaster brevispinosa chathamensis Wheeler, W.M., 1933

• Crematogaster brevispinosa minutior Forel, 1893

• Crematogaster brevispinosa recurvispina Forel, 1912

• Crematogaster brevispinosa sampaioi Forel, 1912

• Crematogaster brevispinosa schuppi Forel, 1901

• Crematogaster brevispinosa striatinota Forel, 1912

• Crematogaster brevispinosa townsendi Wheeler, W.M., 1925

cristata Santschi, 1929 (Brazil)

• Crematogaster terricola Borgmeier, 1929

crucis Forel, 1912 (Colombia, Costa Rica, Ecuador)

cubaensis Mann, 1920 (Cuba, Greater Antilles)

• Crematogaster barbouri Weber, 1934

curvispinosa Mayr, 1862 (Argentina, Barbados, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Galapagos Islands, Grenada, Guadeloupe, Guatemala, Guyana, Honduras, Lesser Antilles, Martinique, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Uruguay, Venezuela)

• Crematogaster curvispinosa antillana Forel, 1893

• Crematogaster curvispinosa kemali Santschi, 1923

• Crematogaster curvispinosa obscura Santschi, 1929

• Crematogaster curvispinosa panamana Wheeler, W.M., 1942

• Crematogaster fuliginea Santschi, 1925

• Crematogaster sculpturata Pergande, 1896

• Crematogaster sculpturata accola Wheeler, W.M., 1934

• Crematogaster sculpturata phytoeca Wheeler, W.M., 1934

cuvierae Donisthorpe, 1945 (Ghana)

cylindriceps Wheeler, W.M., 1927 (Cambodia, Philippines)

cypria Santschi, 1930 (Cyprus)

D
dahlii Forel, 1901 (New Guinea)

daisyi Forel, 1901 (Borneo, Indonesia, Malaysia)

dalyi Forel, 1902 (India)

decamera Forel, 1910 (Borneo, Indonesia, Malaysia)

degeeri Forel, 1886 (Madagascar)

• Crematogaster degeeri lunaris Santschi, 1928

delagoensis Forel, 1894 (Mozambique, Saudi Arabia)

delagoensis acutidens Arnold, 1944 (South Africa)

delagoensis merwei Santschi, 1932 (South Africa)

delagoensis rhodesiana Arnold, 1920 (Zimbabwe)

delitescens Wheeler, W.M., 1921 (French Guiana, Guyana, Lesser Antilles, Suriname)

dentata Dalla Torre, 1893 (Madagascar, Mauritius, Mayotte)

• Crematogaster sewellii improba Forel, 1907

dentinodis Forel, 1901 (Mexico, United States)

depilis Wheeler, W.M., 1919 (Mexico, United States)

• Crematogaster larreae Buren, 1968

depressa (Latreille, 1802) (Guinea, Ivory Coast, Sierra Leone)

• Crematogaster buchneri foreli Mayr, 1895

• Crematogaster mandibularis André, 1889

• Formica platygnatha Roger, 1863

depressa adultera Santschi, 1915 (Congo)

depressa fuscipennis Emery, 1899 (Cameroun)

desecta Forel, 1911 (Sri Lanka)

desperans Forel, 1914 (South Africa)

difformis Smith, F., 1857 (Borneo, Indonesia, Malaysia, Philippines, Singapore, Thailand)

• Crematogaster edentata Mayr, 1867

diffusa (Jerdon, 1851) (India)

dispar Forel, 1902 (Australia)

dispar bipartita Emery, 1922 (Australia)

distans Mayr, 1870 (Argentina, Brazil, Colombia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama, Peru, United States, Venezuela)

• Crematogaster descolei Kusnezov, 1949

• Crematogaster distans cordinoda Forel, 1914

• Crematogaster distans paraensis Forel, 1904

• Crematogaster distans parviceps Forel, 1908

• Crematogaster distans pevsnerae Forel, 1912

• Crematogaster distans vanda Borgmeier, 1929

dohrni Mayr, 1879 (China, India, Sri Lanka)

dohrni artifex Mayr, 1879 (China, India, Singapore, Thailand)

dohrni fabricans Forel, 1911 (Indonesia)

dohrni gigas Forel, 1913 (Sri Lanka)

dohrni kerri Forel, 1911 (Thailand)

dohrni kiangsiensis Forel, 1903 (China)

dolens Forel, 1910 (United Republic of Tanzania)

donisthorpei Santschi, 1934 (Zimbabwe)

dorsidens Santschi, 1925 (Brazil)

dubia Karavaiev, 1935 (Cambodia)

E
ebenina Forel, 1902 (China, India, Myanmar)

• Crematogaster ebenina corax Forel, 1902

edentula Santschi, 1914 (Benin, Guinea, Mali)

egidyi Forel, 1903 (Borneo, China, Indonesia, Malaysia)

egidyi ingvei Forel, 1911 (Indonesia)

egidyi spinozai Forel, 1911 (Borneo, Indonesia, Malaysia)

egidyi szaboi Forel, 1913 (Singapore)

egregior Forel, 1912 (Bolivia, Brazil, Ecuador, Peru)

elegans Smith, F., 1859 (Indonesia, New Guinea)

elysii Mann, 1919 (New Guinea, Solomon Islands)

emeryana Creighton, 1950 (Mexico, United States)

• Crematogaster marioni Buren, 1968

emeryi Forel, 1907 (New Guinea)

• Crematogaster biroi Emery Emery, 1900

enneamera Emery, 1900 (New Guinea)

ensifera Forel, 1910 (Madagascar)

erecta Mayr, 1866 (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Mexico, Panama, Peru, Suriname)

• Crematogaster brevispinosa projecta Santschi, 1925

• Crematogaster virgula Forel, 1899

erectepilosa Salata & Borowiec, 2015 (Greece)

eurydice Forel, 1915 (Australia)

euterpe Santschi, 1922 (Argentina)

evallans Forel, 1907 (Brazil, Colombia, Costa Rica, Guatemala)

• Crematogaster evallans carbonescens Forel, 1913

excisa Mayr, 1895 (Congo, Sierra Leone, Zimbabwe)

excisa bomaella Santschi, 1935 (Democratic Republic of Congo)

excisa cavinota Stitz, 1916 (Congo)

excisa lacustris Santschi, 1914 (Kenya)

F
ferrarii Emery, 1888 (Borneo, Cambodia, China, Indonesia, Krakatau Islands, Laos, Malaysia)

ferrarii lumpurensis Forel, 1907 (Malaysia)

flava Forel, 1886 (Bangladesh, India)

flavicornis Emery, 1897 (New Guinea)

flavitarsis Emery, 1900 (New Guinea)

flaviventris Santschi, 1910 (Democratic Republic of Congo, Yemen)

flavomicrops Longino, 2003 (Brazil, Colombia, Costa Rica, Ecuador, Honduras, Panama, Venezuela)

flavosensitiva Longino, 2003 (Brazil, Colombia, Panama, Venezuela)

foliocrypta Longino, 2003 (Costa Rica, Ecuador)

foraminiceps Santschi, 1913 (Kenya)

foraminiceps mirmillo Santschi, 1935 (Kenya)

foraminiceps staitchi Forel, 1915 (Democratic Republic of Congo)

formosa Mayr, 1870 (Guatemala, Mexico)

• Crematogaster formosa aterrima Wheeler, W.M., 1909

foxi Mann, 1919 (Solomon Islands)

fraxatrix Forel, 1911 (Borneo, Indonesia, Malaysia, Thailand)

fritzi Emery, 1901 (Brunei Darussalam, Indonesia, Malaysia, Micronesia (Federated States of), New Guinea, Sulawesi)

frivola Forel, 1902 (Australia)

• Crematogaster perthensis Crawley, 1922

frivola sculpticeps Forel, 1907 (Australia)

fruhstorferi Emery, 1901 (Indonesia, Sulawesi)

fuentei Menozzi, 1922 (Iberian Peninsula, Spain)

fumikoae Hosoishi & Ogata, 2015 (Thailand, Vietnam)

fusca Mayr, 1876 (Australia)

G
gabonensis Emery, 1899 (Gabon, Ivory Coast, Kenya)

gabonensis fuscitatis Forel, 1913 (Democratic Republic of Congo)

gallicola Forel, 1894 (Mozambique, Zimbabwe)

gallicola latro Forel, 1910 (Botswana)

gambiensis André, 1889 (Gambia, Ghana, Kenya, Sierra Leone)

• Crematogaster fulva Donisthorpe, 1945

gambiensis krantziana Forel, 1914 (South Africa)

gambiensis longiruga Forel, 1907 (Kenya)

gambiensis sejuncta Stitz, 1916 (Democratic Republic of Congo)

gambiensis transversiruga Santschi, 1914 (Sudan)

gavapiga Menozzi, 1935 (Indonesia)

gerstaeckeri Dalla Torre, 1892 (Kenya)

• Crematogaster cephalotes Gerstäcker Gerstäcker, 1871

gerstaeckeri godefreyi Forel, 1914 (South Africa, Zimbabwe)

• Crematogaster neuvillei carininotum Santschi, 1917

gerstaeckeri infaceta Santschi, 1916 (Zimbabwe)

gerstaeckeri inquieta Santschi, 1928 (Zimbabwe)

gerstaeckeri kohliella Santschi, 1918 (Democratic Republic of Congo)

gerstaeckeri maledicta Forel, 1914 (Zimbabwe)

gerstaeckeri oraclum Forel, 1913 (Democratic Republic of Congo)

• Crematogaster bulawayensis Santschi, 1916

• Crematogaster godfreyi foraminicipoides Arnold, 1920

gerstaeckeri pudica Santschi, 1930 (Zimbabwe)

gerstaeckeri pulla Wheeler, W.M., 1922 (Kenya)

gerstaeckeri rufescens Wheeler, W.M., 1922 (South Africa)

gerstaeckeri zulu Wheeler, W.M., 1922 (South Africa)

goeldii Forel, 1903 (Argentina, Bolivia, Brazil)

gordani Karaman, 2008 (Bulgaria, Montenegro)

gratiosa Santschi, 1926 (Democratic Republic of Congo)

grevei Forel, 1891 (Madagascar)

gryllsi Sharaf & Hita Garcia, 2019 (Saudi Arabia)

gullukdagensis Salata & Borowiec, 2015 (Greece)

gutenbergi Santschi, 1914 (Namibia)

H
hafahafa Blaimer, 2013 (Madagascar)

hashimi Hosoishi, 2015 (Malaysia, Thailand)

hazolava Blaimer, 2013 (Madagascar)

heathi Mann, 1916 (Brazil)

hemiceros Santschi, 1926 (Zimbabwe)

hezaradjatica Pisarski, 1967 (Afghanistan, Iran)

himalayana Forel, 1902 (India, Pakistan)

hodgsoni Forel, 1902 (India, Myanmar)

homeri Forel, 1913 (Cameroun)

hottentota Emery, 1899 (Lesotho, South Africa, Zimbabwe)

hottentota bassuto Santschi, 1925 (Lesotho)

hova Forel, 1887 (Madagascar)

• Crematogaster hova latinoda Forel, 1892

huberi (Forel, 1907) (Guyana, Suriname)

hullettii Feldhaar, Maschwitz & Fiala, 2016 (Borneo, Malaysia)

I
iheringi Forel, 1908 (Argentina, Brazil)

iheringi bosqi Santschi, 1934 (Argentina)

iheringi nemesis Santschi, 1934 (Argentina)

ilgii Forel, 1910 (Ethiopia)

imperfecta Hosoishi, 2015 (Malaysia)

impressa Emery, 1899 (Cameroun, Gambia, Guinea, Ivory Coast)

• Crematogaster excisa andrei Forel, 1911

impressa brazzai Santschi, 1910 (Congo)

impressa maynei Forel, 1913 (Democratic Republic of Congo)

impressa sapora Forel, 1916 (Democratic Republic of Congo)

impressiceps Mayr, 1902 (Equatorial Guinea)

impressiceps frontalis Santschi, 1922 (Democratic Republic of Congo)

impressiceps longiscapa Stitz, 1916 (Democratic Republic of Congo)

impressiceps lujana Forel, 1915 (Democratic Republic of Congo)

inconspicua Mayr, 1896 (Cameroun)

incorrecta Santschi, 1917 (Gabon)

indefensa Kempf, 1968 (Bolivia)

indosinensis Hosoishi & Ogata, 2016 (Cambodia)

inermis Mayr, 1862 (Algeria, Egypt, Iran, Israel, Spain)

inermis aphrodite Santschi, 1937 (Cyprus)

inermis armatula Emery, 1926 (Israel, Syrian Arab Republic)

inermis lucida Forel, 1890 (Tunisia)

inflata Smith, F., 1857 (Borneo, India, Indonesia, Malaysia, Philippines, Singapore, Thailand)

• Crematogaster moorei Donisthorpe, 1941

innocens Forel, 1911 (Borneo, Indonesia, Malaysia)

insularis Smith, F., 1859 (New Guinea)

ionia Forel, 1911 (Croatia, Greece, Israel, Montenegro, Republic of Macedonia, Turkey)

iridipennis Smith, F., 1865 (New Guinea)

irritabilis Smith, F., 1860 (New Guinea)

irritabilis leguilloui Emery, 1900 (New Guinea)

irritabilis subtilis Viehmeyer, 1914 (New Guinea)

isolata Buren, 1968 (Mexico, United States)

izanami Terayama, 2013 (Japan)

J
jacindae Sharaf & Hita Garcia, 2019 (Oman)

jacobsoni Forel, 1911 (Indonesia)

jardinero Longino, 2003 (Costa Rica, Ecuador)

javanica Menozzi, 1935 (Indonesia)

jeanneli Santschi, 1914 (Kenya)

jehovae Forel, 1907 (China, Greece, Israel, Montenegro, Saudi Arabia, United Arab Emirates, Yemen)

jehovae crawleyi Emery, 1926 (Iraq)

jullieni Santschi, 1910 (Gabon, Guinea)

juventa Santschi, 1926 (Democratic Republic of Congo)

K
kachelibae Arnold, 1954 (Kenya)

karawaiewi Menozzi, 1935 (New Guinea)

• Crematogaster wladimiri Consani, 1953

kasaiensis Forel, 1913 (Democratic Republic of Congo)

kelleri Forel, 1891 (Madagascar, Seychelles)

• Crematogaster adrepens Forel, 1897

• Crematogaster gibba Emery, 1894

keris Hosoishi, 2020 (Malaysia)

khmerensis Hosoishi & Ogata, 2017 (Cambodia)

kirbii (Sykes, 1835) (India)

kneri Mayr, 1862 (Benin, Ghana, Senegal)

kneri behanzini Santschi, 1915 (Benin)

kneri dakarensis Santschi, 1914 (Senegal)

kneri funerea Santschi, 1915 (Cameroun)

kneri pronotalis Santschi, 1914 (Nigeria)

kohli Forel, 1909 (Democratic Republic of Congo, Zimbabwe)

kohli winkleri Forel, 1909 (Cameroun)

kojimai Hosoishi & Ogata, 2012 (Malaysia)

kutteri Viehmeyer, 1924 (Australia)

L
laestrygon Emery, 1869 (Balearic Islands, Canary Islands, Georgia, Iberian Peninsula, Malta, Morocco, Saudi Arabia, Spain, Turkey, Yemen)

• Crematogaster laestrygon vaucheri Emery, 1926

laestrygon airensis Santschi, 1937 (Niger)

laestrygon atlantis Santschi, 1937 (Algeria)

laestrygon canariensis Barquin, 1981 (Canary Islands)

laestrygon diminuta Santschi, 1910 (Tunisia)

laestrygon granulata Santschi, 1910 (Tunisia)

laestrygon maura Forel, 1909 (Algeria)

laestrygon submaura Lomnicki, 1925 (Spain)

laestrygon surcoufi Santschi, 1937 (Algeria)

laestrygon theryi Santschi, 1921 (Morocco)

laestrygon vivax Santschi, 1937 (Morocco)

laeviceps Smith, F., 1858 (Australia)

laeviceps broomensis Forel, 1915 (Australia)

laeviceps chasei Forel, 1902 (Australia)

laeviceps clarior Forel, 1902 (Australia)

laevis Mayr, 1878 (Bolivia, Brazil, Peru)

laevis bruesi Wheeler, W.M., 1911 (Grenada, Lesser Antilles)

laevissima Smith, F., 1860 (Indonesia, Sulawesi)

laeviuscula Mayr, 1870 (Mexico, United States)

• Crematogaster atkinsoni Wheeler, W.M., 1919

• Crematogaster atkinsoni helveola Wheeler, W.M., 1919

• Crematogaster castanea decolorata Santschi, 1926

• Crematogaster clara Mayr, 1870

• Crematogaster hespera Buren, 1968

• Crematogaster opaca cedrosensis Wheeler, W.M., 1934

• Oecodoma arborea Buckley, 1867

• Oecodoma bicolor Buckley, 1867

lamottei Bernard, 1953 (Guinea)

lango Weber, 1943 (Central African Republic, Sudan)

latuka Weber, 1943 (Sudan)

laurenti Forel, 1909 (Democratic Republic of Congo)

laurenti zeta Emery, 1922 (Democratic Republic of Congo)

ledouxi Soulié, 1961 (Cambodia)

levior Longino, 2003 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela)

libengensis Stitz, 1916 (Democratic Republic of Congo)

libengensis rufula Santschi, 1926 (Democratic Republic of Congo)

liengmei Forel, 1894 (Mozambique)

liengmei caculata Forel, 1915 (South Africa)

liengmei weitzeckeri Emery, 1895 (South Africa)

• Crematogaster coelestis kloofensis Forel, 1914

limata Smith, F., 1858 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Trinidad and Tobago, Venezuela)

• Crematogaster inca ascendens Wheeler, W.M., 1925

• Crematogaster limata dextella Santschi, 1929

• Crematogaster limata palans Forel, 1912

lineolata (Say, 1836) (Mexico, United States)

• Crematogaster lineolata subopaca Emery, 1895

• Crematogaster opaca texana Santschi, 1929

• Myrmica columbiana Buckley, 1867

• Myrmica marylandica Buckley, 1867

• Myrmica novaeboracensis Buckley, 1867

linsenmairi Feldhaar, Maschwitz & Fiala, 2016 (Singapore)

litoralis Arnold, 1955 (Kenya)

lobata Emery, 1895 (Madagascar)

longicephala Özdikmen, 2010 (Australia)

• Crematogaster longiceps Forel Forel, 1910

longicephala curticeps Wheeler, W.M., 1915 (Australia)

longiceps Santschi, 1910 (Mali, Sudan)

longiclava Emery, 1893 (Philippines)

longiclava placens Forel, 1911 (Malaysia)

longipilosa Forel, 1907 (Borneo, Indonesia, Malaysia, Thailand)

longispina Emery, 1890 (Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Nicaragua, Peru, Suriname)

longispina naumannae Forel, 1921 (Ecuador)

longispina nura Özdikmen, 2010 (Bolivia)

• Crematogaster longispina boliviana Wheeler, W.M., 1925

lorteti Forel, 1910 (Greece, Israel, Montenegro, Syrian Arab Republic, Turkey)

lorteti hellenica Forel, 1911 (Greece)

lotti Weber, 1943 (Sudan)

lucayana Wheeler, W.M., 1905 (Bahamas)

lucayana etiolata Wheeler, W.M., 1905 (Bahamas)

luctans Forel, 1907 (Eritrea, Iraq, Kenya, Sierra Leone)

luctans liebknechti Forel, 1915 (Democratic Republic of Congo)

luctans sordidoides Santschi, 1930 (Sierra Leone)

lutzi Forel, 1905 (Brazil)

lutzi florida Forel, 1908 (Brazil)

luzonensis Hosoishi & Ogata, 2016 (Philippines)

M
macaoensis Wu, J. & Wang, C., 1995 (China)

macracantha Creighton, 1945 (Borneo, Indonesia, Malaysia)

madagascariensis André, 1887 (Madagascar)

madecassa Emery, 1895 (Madagascar)

mafybe Blaimer, 2013 (Madagascar)

magitae Forel, 1910 (Sierra Leone)

magnifica Santschi, 1925 (Brazil)

• Crematogaster magnifica nociva Borgmeier, 1937

mahery Blaimer, 2010 (Madagascar)

maina Blaimer, 2013 (Comoros, Madagascar, Mayotte)

major Donisthorpe, 1941 (New Guinea)

major flavior Donisthorpe, 1941 (New Guinea)

malahelo Blaimer, 2013 (Madagascar)

malala Blaimer, 2010 (Madagascar)

mancocapaci Santschi, 1911 (Ecuador)

manni Buren, 1968 (Cuba, Greater Antilles)

margaritae Emery, 1895 (Democratic Republic of Congo)

margaritae brevarmata Forel, 1915 (Democratic Republic of Congo)

margaritae cupida Santschi, 1935 (Democratic Republic of Congo)

margaritae lujae Forel, 1913 (Democratic Republic of Congo)

marthae Forel, 1892 (Madagascar)

maryatii Feldhaar, Maschwitz & Fiala, 2016 (Borneo, Indonesia, Malaysia)

masokely Blaimer, 2013 (Madagascar)

masukoi Hosoishi, Yamane & Ogata, 2010 (Borneo, Indonesia, Malaysia)

matsumurai Forel, 1901 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea)

• Crematogaster matsumurai iwatensis Santschi, 1937

meijerei Emery, 1911 (New Guinea)

melanogaster Emery, 1895 (Oman)

melanogaster homonyma Emery, 1922 (South Africa)

menilekii Forel, 1894 (Benin, Ethiopia, Guinea)

menilekii completa Santschi, 1928 (Zimbabwe)

menilekii occidentalis Mayr, 1902 (Equatorial Guinea)

menilekii proserpina Santschi, 1919 (Democratic Republic of Congo)

menilekii satan Forel, 1916 (Democratic Republic of Congo)

menilekii spuria Forel, 1913 (Democratic Republic of Congo)

menilekii suddensis Weber, 1943 (Sudan)

menilekii viehmeyeri Santschi, 1921 (United Republic of Tanzania)

mesonotalis Emery, 1911 (New Guinea)

microspina Menozzi, 1942 (Equatorial Guinea)

mimicans Donisthorpe, 1932 (New Guinea)

mimosae Santschi, 1914 (Kenya, Oman, Saudi Arabia, United Arab Emirates, Yemen)

minutissima Mayr, 1870 (Costa Rica, Mexico, United States)

misella Arnold, 1920 (Zimbabwe)

missouriensis Emery, 1895 (United States)

• Crematogaster minutissima smithi Creighton, 1950

• Crematogaster minutissima thoracica Creighton, 1939

mjobergi Forel, 1915 (Australia)

moatensis Hosoishi & Ogata, 2016 (Indonesia)

modiglianii Emery, 1900 (Borneo, Indonesia, Laos, Malaysia, Myanmar, Philippines, Thailand)

• Crematogaster modiglianii anamita Santschi, 1925

• Crematogaster modiglianii anoemica Santschi, 1925

• Crematogaster modiglianii clemensae Forel, 1910

• Crematogaster modiglianii sarawakana Forel, 1911

• Crematogaster modiglianii surbeki Santschi, 1925

moelleri Forel, 1912 (Argentina, Brazil, Costa Rica)

monocula Hosoishi, 2019 (Cambodia)

montana Borgmeier, 1939 (Brazil)

montenigrina Karaman, 2008 (Montenegro)

monteverdensis Longino, 2003 (Costa Rica)

montezumia Smith, F., 1858 (Argentina, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Paraguay, Peru)

• Crematogaster montezumia cristulata Santschi, 1925

• Crematogaster montezumia functa Forel, 1911

• Crematogaster montezumia proletaria Santschi, 1933

• Crematogaster sulcata Mayr, 1870

• Crematogaster sulcata ramulinida Forel, 1899

monticola Arnold, 1920 (Lesotho, South Africa)

moqorensis Pisarski, 1967 (Afghanistan)

mosis Forel, 1909 (Israel, Lebanon, United Arab Emirates, Yemen)

mottazi Santschi, 1928 (Ivory Coast)

mpanjono Blaimer, 2012 (Madagascar)

mucronata Emery, 1900 (Indonesia)

• Crematogaster fulmeki Forel, 1922

muralti Forel, 1910 (Cameroun)

muralti livingstonei Santschi, 1919 (Zambia)

muralti ugandensis Santschi, 1914 (Uganda)

mutans Buren, 1968 (United States)

myops Forel, 1911 (Borneo, Indonesia, Malaysia, Singapore)

N
natalensis Forel, 1910 (Lesotho, South Africa)

natalensis braunsi Emery, 1922 (South Africa)

natalensis dulcis Santschi, 1926 (South Africa)

navajoa Buren, 1968 (United States)

nawai Ito, 1914 (China, Japan, Republic of Korea)

• Crematogaster laboriosa Smith, F., 1874

nesiotis Mann, 1919 (New Guinea, Solomon Islands)

neuvillei Forel, 1907 (Ethiopia)

neuvillei cooperi Forel, 1914 (South Africa)

nigeriensis Santschi, 1914 (Nigeria)

nigeriensis wilnigra Forel, 1916 (Democratic Republic of Congo)

nigrans Forel, 1915 (Democratic Republic of Congo)

nigriceps Emery, 1897 (Israel, Kenya, Somalia)

nigriceps prelli Forel, 1911 (United Republic of Tanzania)

nigriceps saganensis Consani, 1952 (Ethiopia)

nigronitens Santschi, 1917 (Zimbabwe)

nigropilosa Mayr, 1870 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Venezuela)

nitidiceps Emery, 1895 (Argentina, Brazil, Paraguay, Peru, Uruguay)

nocturna Buren, 1968 (United States)

nosibeensis Forel, 1891 (Madagascar)

O
oasium Santschi, 1911 (Kuwait, Oman, Saudi Arabia, Tunisia, United Arab Emirates)

oasium saharensis Santschi, 1921 (Algeria)

obnigra Mann, 1919 (New Guinea, Solomon Islands)

obscura Smith, F., 1857 (Borneo, Indonesia, Malaysia)

obscurata Emery, 1895 (Belize, Colombia, Costa Rica, Guatemala, Mexico, Panama, United States, Venezuela)

• Crematogaster agnita Wheeler, W.M., 1934

obscurior Dalla Torre, 1892 (Indonesia)

• Crematogaster obscurus Smith, F., 1860

ocellata Hosoishi & Ogata, 2016 (Vietnam)

ochracea Mayr, 1862 (Philippines)

ochraceiventris Stitz, 1916 (Congo)

onusta Stitz, 1925 (Borneo, Indonesia, Malaysia, Philippines)

opaca Mayr, 1870 (Honduras, Mexico, United States)

opaciceps Mayr, 1901 (South Africa)

opaciceps cacodaemon Forel, 1914 (South Africa)

opaciceps clepens Forel, 1913 (Democratic Republic of Congo)

opaciceps defleta Forel, 1910 (South Africa)

orobia Santschi, 1919 (South Africa)

osakensis Forel, 1900 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea)

• Crematogaster sordidula japonica Forel, 1912

oscaris Forel, 1910 (Namibia, South Africa)

overbecki Viehmeyer, 1916 (Singapore)

oxygynoides Santschi, 1934 (Argentina, Brazil)

P
painei Donisthorpe, 1945 (Ghana)

pallida Lowne, 1865 (Australia)

pallipes Mayr, 1862 (Australia)

• Crematogaster piceus Lowne, 1865

paolii Menozzi, 1930 (Benin, Somalia)

paradoxa Emery, 1894 (New Guinea)

parallela Santschi, 1925 (Brazil)

patei Buren, 1968 (Mexico)

pauciseta Emery, 1899 (Cameroun)

pauciseta grossulior Forel, 1916 (Democratic Republic of Congo)

pauli Emery, 1901 (Indonesia, Sulawesi)

pellens Walker, 1859 (Sri Lanka)

perelegans Forel, 1902 (India)

peringueyi Emery, 1895 (Namibia, South Africa)

peringueyi angustior Arnold, 1920 (South Africa)

peringueyi cacochyma Forel, 1914 (South Africa)

peringueyi dentulata Stitz, 1923 (South Africa)

peringueyi gedeon Arnold, 1920 (South Africa)

peristerica Menozzi, 1925 (Brazil)

peruviana (Wheeler, W.M., 1922) (Peru)

petiolidens Forel, 1916 (Democratic Republic of Congo)

pfeifferi Hosoishi & Ogata, 2017 (Borneo, Malaysia)

philippinensis Hosoishi & Ogata, 2016 (Philippines)

phoenica Santschi, 1915 (Syrian Arab Republic)

phoenica pygmalion Santschi, 1934 (Syrian Arab Republic)

phoenix Santschi, 1921 (Kenya, United Republic of Tanzania)

physothorax Emery, 1889 (Borneo, Indonesia, Malaysia, Myanmar)

pia Forel, 1911 (Malaysia, Thailand)

pia soengeiensis Forel, 1913 (Indonesia)

pia taivanae Forel, 1913 (Taiwan)

pilosa Emery, 1895 (United States)

• Crematogaster creightoni Wheeler, W.M., 1933

pinicola Deyrup & Cover, 2007 (United States)

polita Smith, F., 1865 (New Guinea)

politula Forel, 1902 (Bangladesh, China, India)

polymnia Santschi, 1922 (Argentina)

polymnia oeagria Santschi, 1933 (Argentina)

popohana Forel, 1912 (Taiwan)

popohana amia Forel, 1913 (Taiwan)

pradipi Tiwari, 1999 (India)

pseudinermis Viehmeyer, 1923 (Kenya, Sudan)

pseudinermis muellerianus Finzi, 1939 (Eritrea, Ethiopia)

pulchella Bernard, 1953 (Cameroun, Guinea)

punctulata Emery, 1895 (Mexico, United States)

pygmaea Forel, 1904 (Brazil)

pythia Forel, 1915 (Australia)

Q
quadriformis Roger, 1863 (Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Peru, Suriname, Uruguay)

• Crematogaster quadriformis gracilior Forel, 1901

• Crematogaster quadriformis vezenyii Forel, 1907

quadriformis roveretoi Forel, 1913 (Argentina)

quadriruga Forel, 1911 (Cambodia, Indonesia, Laos, Thailand)

quadrispinosa Roger, 1863 (Mexico)

queenslandica Forel, 1902 (Australia)

queenslandica froggatti Forel, 1902 (Australia)

queenslandica gilberti Emery, 1922 (Australia)

queenslandica rogans Forel, 1902 (Australia)

queenslandica scabrula Emery, 1914 (Australia)

R
ralumensis Forel, 1901 (New Guinea)

ramamy Blaimer, 2013 (Madagascar)

ranavalonae Forel, 1887 (Madagascar, Mayotte)

• Crematogaster descarpentriesi Santschi, 1928

• Crematogaster emmae Forel, 1891

• Crematogaster emmae laticeps Forel, 1892

• Crematogaster inops Forel, 1892

• Crematogaster ranavalonae paulinae Forel, 1892

• Crematogaster ranavalonae pepo Forel, 1922

ransonneti Mayr, 1868 (India, Sri Lanka)

raptor Longino, 2003 (Costa Rica)

rasoherinae Forel, 1891 (Comoros, Madagascar, Mayotte, Seychelles)

• Crematogaster rasoherinae brunnea Forel, 1907

• Crematogaster rasoherinae brunneola Emery, 1922

• Crematogaster voeltzkowi Forel, 1907

rauana Forel, 1907 (United Republic of Tanzania)

razana Blaimer, 2012 (Madagascar)

rectinota Forel, 1913 (Kenya, Zimbabwe)

recurva Emery, 1897 (New Guinea)

resulcata Bolton, 1995 (South Africa)

• Crematogaster subsulcata Santschi, 1932

reticulata Hosoishi, 2009 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Thailand)

retifera Santschi, 1926 (Zimbabwe)

rifelna Buren, 1968 (Mexico, United States)

rivai Emery, 1897 (Ethiopia)

rivai luctuosa Menozzi, 1930 (Somalia)

rochai Forel, 1903 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Venezuela)

• Crematogaster brevispinosa malevolens Santschi, 1919

rogenhoferi Mayr, 1879 (Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Sri Lanka, Thailand, Vietnam)

rogenhoferi costulata Emery, 1895 (Myanmar)

rogenhoferi fictrix Forel, 1911 (Borneo, Indonesia, Malaysia)

rogenhoferi lutea Emery, 1893 (Indonesia, Sri Lanka)

rogeri Emery, 1922 (Sri Lanka)

ronganensis Zhou, 2001 (China)

roslihashimi Feldhaar, Maschwitz & Fiala, 2016 (Borneo, Malaysia)

rothneyi Mayr, 1879 (Bangladesh, Cambodia, China, India, Indonesia, Myanmar, Pakistan, Philippines, Sri Lanka, Thailand, Vietnam)

• Crematogaster rothneyi civa Forel, 1902

rothneyi haputalensis Forel, 1913 (Sri Lanka)

rudis Emery, 1894 (Argentina, Bolivia, Brazil)

rufa (Jerdon, 1851) (India)

rufigena Arnold, 1958 (Zimbabwe)

rufotestacea Mayr, 1876 (Australia)

rufotestacea dentinasis Santschi, 1929 (Australia)

rugosa André, 1895 (Congo, Ivory Coast, Kenya)

rugosa nigriventris Santschi, 1919 (Ivory Coast)

rugosa rugaticeps Santschi, 1919 (Cameroun)

rugosior Santschi, 1910 (Congo)

ruspolii Forel, 1892 (Ethiopia)

ruspolii atriscapis Forel, 1915 (Democratic Republic of Congo)

russata Wheeler, W.M., 1925 (Bolivia)

russoi Menozzi, 1930 (Dominican Republic, Greater Antilles)

rustica Santschi, 1935 (Democratic Republic of Congo)

S
sabatra Blaimer, 2010 (Madagascar)

sagei Forel, 1902 (China, India)

sagei laevinota Forel, 1902 (India)

sanguinea Roger, 1863 (Cuba)

sanguinea atavista Mann, 1920 (Cuba, Greater Antilles)

sanguinea sotolongoi Mann, 1920 (Cuba)

sanguinea torrei Wheeler, W.M., 1913 (Cuba)

santschii Forel, 1913 (Democratic Republic of Congo)

santschii clymene Forel, 1915 (South Africa)

saussurei Forel, 1899 (Mexico)

scapamaris Santschi, 1922 (Argentina)

scelerata Santschi, 1917 (Argentina, Brazil, Paraguay)

scelerata taperensis Borgmeier, 1937 (Brazil)

schencki Forel, 1891 (Madagascar)

schimmeri Forel, 1912 (Taiwan)

schmidti (Mayr, 1853) (Armenia, Austria, Bulgaria, Georgia, Greece, Hungary, Iran, Italy, Montenegro, Republic of Macedonia, Romania, Russian Federation, Slovenia, Turkey, Turkmenistan, Ukraine)

• Crematogaster auberti karawaewi Ruzsky, 1905

• Crematogaster scutellaris christowitchii Forel, 1892

• Crematogaster scutellaris medispina Forel, 1905

schultzei Forel, 1910 (Namibia, South Africa)

scita Forel, 1902 (Australia, Solomon Islands)

scita mixta Forel, 1902 (Australia)

scutellaris (Olivier, 1792) (Andorra, Austria, Balearic Islands, Bulgaria, Croatia, Denmark, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Israel, Italy, Malta, Montenegro, Portugal, Republic of Macedonia, Romania, Slovenia, Spain, Switzerland, Tunisia, Turkey, United Kingdom of Great Britain and Northern Ireland)

• Acrocoelia ruficeps Mayr, 1853

• Crematogaster scutellaris corsica Santschi, 1921

• Crematogaster scutellaris degener Santschi, 1937

• Crematogaster scutellaris grouvellei Bondroit, 1918

• Crematogaster scutellaris lichtensteini Bondroit, 1918

• Formica haematocephala Leach, 1825

• Myrmica rediana Gené, 1841

• Myrmica rubriceps Nylander, 1849

scutellaris alii Forel, 1907 (Algeria)

scutellaris nigra Krausse, 1912 (Italy)

scutellaris tenuispina Forel, 1902 (Algeria)

semperi Emery, 1893 (Philippines)

senegalensis Roger, 1863 (Gambia, Guinea, Madagascar, Mali, Niger, Oman, Saudi Arabia, Senegal, United Arab Emirates, Western Sahara, Yemen)

senegalensis goliathula Forel, 1922 (Angola)

sericea Forel, 1912 (Argentina, Brazil, Paraguay, Uruguay)

sewardi Forel, 1901 (Borneo, Indonesia, Krakatau Islands, Malaysia, Thailand, Vietnam)

sewellii Forel, 1891 (Comoros)

sewellii acis Forel, 1913 (Democratic Republic of Congo)

sewellii marnoi Mayr, 1895 (Sudan)

sikkimensis Forel, 1904 (India)

simboloni Hosoishi & Ogata, 2014 (Indonesia)

similis Stitz, 1911 (Democratic Republic of Congo)

simoni Emery, 1893 (Philippines)

sisa Blaimer, 2010 (Madagascar)

sjostedti Mayr, 1907 (Kenya, Mozambique, United Republic of Tanzania, Zimbabwe)

skounensis Soulié, 1961 (Cambodia)

snellingi Longino, 2003 (Costa Rica, Honduras)

solenopsides Emery, 1899 (Ivory Coast, United Republic of Tanzania)

solenopsides costeboriensis Santschi, 1919 (Ivory Coast)

solenopsides flavida Mayr, 1907 (United Republic of Tanzania)

solenopsides mandonbii Santschi, 1926 (Uganda)

solers Forel, 1910 (Namibia)

sordidula (Nylander, 1849) (Albania, Balearic Islands, Bulgaria, Croatia, France, Georgia, Gibraltar, Greece, Hungary, Iberian Peninsula, Israel, Italy, Montenegro, Portugal, Republic of Macedonia, Romania, Russian Federation, Slovenia, Spain, Switzerland, Turkey)

• Acrocoelia mayri Mayr, 1853

• Crematogaster sordidula caspica Emery, 1912

• Crematogaster sordidula flachii Forel, 1895

• Orthocrema esterelana Bernard, 1978

sordidula aeolia Forel, 1911 (Turkey)

sordidula marocana Santschi, 1921 (Morocco)

sordidula molongori Weber, 1943 (Sudan)

sordidula osmanica Santschi, 1921 (Turkey)

sorokini Ruzsky, 1905 (Iran, Kazakhstan, Turkmenistan)

sotobosque Longino, 2003 (Brazil, Colombia, Costa Rica, Ecuador, Panama, Peru, Venezuela)

stadelmanni Mayr, 1895 (Central African Republic, Guinea, Ivory Coast, Kenya, Liberia)

stadelmanni anguliceps Stitz, 1916 (Cameroun)

stadelmanni angustata Mayr, 1895 (Cameroun)

stadelmanni dolichocephala Santschi, 1911 (Congo)

stadelmanni gracilenta Viehmeyer, 1922 (Cameroun)

stadelmanni intermedia Mayr, 1896 (Cameroun)

stadelmanni ovinodis Stitz, 1916 (Democratic Republic of Congo)

stadelmanni schereri Forel, 1911 (Liberia)

stadelmanni spissata Santschi, 1937 (Democratic Republic of Congo)

steinheili Forel, 1881 (Bahamas, Barbados, Cuba, Dominican Republic, Greater Antilles, Haiti, Lesser Antilles, Puerto Rico, United States Virgin Islands)

steinheili argentina Forel, 1915 (Argentina)

stenocephala Emery, 1922 (Mali, Senegal)

stigmata Santschi, 1914 (Nigeria)

stigmatica Forel, 1911 (Brazil)

stollii Forel, 1885 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Nicaragua, Panama, Peru, Venezuela)

• Crematogaster hyperphyes Kusnezov, 1953

• Crematogaster stolli autruni Mann, 1916

• Crematogaster stolli guianensis Crawley, 1916

• Crematogaster stollii amazonensis Forel, 1904

• Tranopeltoides parvispina Wheeler, W.M., 1922

storki Hosoishi & Ogata, 2016 (Indonesia)

striaticeps Forel, 1909 (Algeria, Saudi Arabia)

striatula Emery, 1892 (Cameroun, Ivory Coast, Kenya, Sierra Leone)

striatula benitensis Santschi, 1910 (Congo)

striatula horatii Santschi, 1937 (Kenya)

striatula langi Santschi, 1926 (Democratic Republic of Congo)

striatula obstinata Santschi, 1911 (Congo)

striatula omega Santschi, 1935 (Democratic Republic of Congo)

subcircularis Mayr, 1879 (Borneo, Indonesia, Malaysia, Philippines)

subdentata Mayr, 1877 (Afghanistan, Armenia, China, Georgia, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Russian Federation, Turkmenistan, Ukraine)

• Crematogaster subdentata flavicapilla Ruzsky, 1905

subdentata kaschgariensis Forel, 1901 (Kazakhstan, Pakistan)

subdentata kuznetzovi Stärcke, 1935 (Kazakhstan)

subnuda Mayr, 1879 (China, India, Pakistan, Sri Lanka)

subnuda discinodis Emery, 1893 (Singapore)

subnuda nigrosubnuda Özdikmen, 2010 (Taiwan)

• Crematogaster subnuda formosae Wheeler, W.M., 1909

subtonsa Santschi, 1925 (Brazil)

suehiro Terayama, 1999 (Japan)

• Crematogaster miroku Terayama, 2013

sumichrasti Mayr, 1870 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Crematogaster atitlanica Wheeler, W.M., 1936

• Crematogaster sumichrasti maya Wheeler, W.M., 1936

• Crematogaster sumichrasti surdior Forel, 1885

sundalandensis Hosoishi & Ogata, 2016 (Indonesia, Malaysia)

T
tanakai Hosoishi & Ogata, 2009 (Borneo, Indonesia, Malaysia)

tarsata Smith, F., 1865 (Indonesia, New Guinea)

tavaratra Blaimer, 2013 (Madagascar)

telolafy Blaimer, 2012 (Madagascar)

tenuicula Forel, 1904 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guyana, Panama, Peru, Venezuela)

tenuipilis Santschi, 1937 (Kenya, United Republic of Tanzania)

teranishii Santschi, 1930 (Japan, Republic of Korea)

terminalis (Shuckard, 1838) (United Kingdom of Great Britain and Northern Ireland)

tetracantha Emery, 1887 (New Guinea)

thalia Forel, 1911 (Paraguay)

theta Forel, 1911 (Democratic Republic of Congo)

togoensis Donisthorpe, 1945 (Togo)

torosa Mayr, 1870 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, United States)

• Crematogaster arizonensis Wheeler, W.M., 1908

• Crematogaster brevispinosa tumulifera Forel, 1899

transiens Forel, 1913 (Democratic Republic of Congo)

transvaalensis Forel, 1894 (Botswana, South Africa)

transvaalensis hammi Arnold, 1920 (Zimbabwe)

trautweini Viehmeyer, 1914 (Cameroun)

travancorensis Forel, 1902 (China, India)

treubi Emery, 1896 (Brunei Darussalam, Cambodia, China, Indonesia, Laos, Malaysia, Myanmar, Thailand)

• Crematogaster buddhae sic Forel, 1907

• Crematogaster millardi Forel, 1902

• Crematogaster treubi vastatrix Forel, 1911

treubi apilis Forel, 1913 (Taiwan)

tricolor Gerstäcker, 1859 (Comoros, Kenya, Mayotte, Mozambique)

tsisitsilo Blaimer, 2013 (Madagascar)

tumidula Emery, 1900 (Borneo, Indonesia, Malaysia, Thailand)

U
udo Forel, 1905 (Indonesia)

unciata Santschi, 1925 (Brazil)

uruguayensis Santschi, 1912 (Argentina, Uruguay)

ustiventris Menozzi, 1935 (New Guinea)

V
vacca Forel, 1911 (Borneo, Indonesia, Malaysia)

• Crematogaster stethogompha Wheeler, W.M., 1919

• Crematogaster stethogompha detritinodis Wheeler, W.M., 1919

vagula Wheeler, W.M., 1928 (China, Japan, Republic of Korea)

vandeli Soulié, 1961 (Cambodia)

vandermeermohri Menozzi, 1930 (Indonesia)

vermiculata Emery, 1895 (Mexico, United States)

• Crematogaster colei Buren, 1968

• Crematogaster opuntiae Buren, 1968

• Crematogaster rossi Buren, 1968

vicina André, 1893 (Greater Antilles, Jamaica)

• Crematogaster vicina wighti Wheeler, W.M., 1908

victima Smith, F., 1858 (Argentina, Bolivia, Brazil, Cuba, Paraguay, Peru, Uruguay, Venezuela)

victima alegrensis Forel, 1901 (Brazil)

vidua Santschi, 1928 (Ivory Coast)

vieti Hosoishi & Ogata, 2016 (Vietnam)

vitalisi Menozzi, 1925 (Philippines)

volamena Blaimer, 2012 (Madagascar)

vulcania Santschi, 1913 (Kenya, Zimbabwe)

W
walshi Forel, 1902 (China, India, Sri Lanka, Vietnam)

warburgi Menozzi, 1933 (Iran, Israel)

wardi Longino, 2003 (Costa Rica)

wasmanni Santschi, 1910 (Democratic Republic of Congo)

weberi Emery, 1911 (New Guinea)

wellmani Forel, 1909 (Angola, Ghana, Guinea, Kenya, Sierra Leone, Zimbabwe)

• Crematogaster boxi Donisthorpe, 1945

• Crematogaster wellmani retusa Santschi, 1916

• Sphaerocrema dewasi Bruneau de Miré, 1966

wellmani luciae Forel, 1913 (Cameroun)

wellmani weissi Santschi, 1910 (Congo)

werneri Mayr, 1907 (Guinea, Uganda)

werneri cacozela Santschi, 1914 (Guinea)

werneri pasithea Santschi, 1915 (Benin)

wheeleri Mann, 1919 (Solomon Islands)

whitei Wheeler, W.M., 1915 (Australia)

wilwerthi Santschi, 1910 (Democratic Republic of Congo)

wilwerthi confusa Santschi, 1911 (Congo)

wilwerthi fauconneti Forel, 1910 (Democratic Republic of Congo)

wroughtonii Forel, 1902 (China, India)

X
xerophila (Wheeler, W.M., 1915) (Australia)

xerophila exigua Wheeler, W.M., 1915 (Australia)

Y
yaharai Hosoishi & Ogata, 2019 (Cambodia)

yamanei Hosoishi & Ogata, 2009 (Indonesia)

yappi Forel, 1901 (Malaysia)

Z
zavattarii Menozzi, 1926 (Nigeria)

zavattarii edmeeae Santschi, 1937 (Congo)

zoceensis Santschi, 1925 (China)

zonacaciae Weber, 1943 (Sudan)

=Fossil Species=

A
aurora LaPolla & Greenwlalt, 2015

P
praecursor Emery, 1891

primitiva Radchenko & Dlussky, 2019