Ward, Philip. S.

BIOGRAPHICAL NOTE
Office: 381A Briggs Hall Lab: 381B Briggs Hall Phone: (530) 752-0486 Email: psward@ucdavis.edu Fax: (530) 752-1537 Education: B.Sc. Biology, Queens University, Canada (1973) Ph.D. Zoology, University of Sydney, Australia (1979) Appointment: 25% Instruction and Research 75% Organized Research Teaches: Systematics, biogeography and evolution of ants; ant-plant mutualisms; phylogeny and speciation. Research Interests: Systematics and evolutionary biology of ants; genetic structure of social insect populations.

PUBLICATIONS

 * [[Media:Baroni Urbani Bolton Ward 1992.pdf|Baroni Urbani, C.; Bolton, B.; Ward, P. S. 1992. The internal phylogeny of ants (Hymenoptera: Formicidae). Syst. Entomol. 17: 301-329 PDF]]


 * [[Media:Blaimer et al. 2015 BMC EvoBio.pdf|Blaimer, B. B.; Brady, S. G.; Schultz, T. R.; Lloyd, M. W.; Fisher, B. L.; Ward, P. S. 2015. Phylogenomic methods outperform traditional multi-locus approaches in resolving deep evolutionary history: a case study of formicine ants. BMC Evolutionary Biology 15:257. PDF]]


 * [[Media:Brady et al 2014.pdf|Brady, S. G.; Fisher, B. L.; Schultz, T. R.; Ward, P. S. 2014. The rise of army ants and their relatives: diversification of specialized predatory doryline ants. BMC Evolutionary Biology 2014 14:93, 14 pp. (doi:10.1186/1471-2148-14-93). '''PDF]]


 * [[Media:Brady & Ward 1999.pdf|Brady, S. G., J. Gadau and P. S. Ward. 1999. Is the ant genus Camponotus paraphyletic? Pages 27-28 in Austin, A. D. and P. J. DeBarro. Fourth International Hymenopterists Conference: program and abstracts. International Society of Hymenopterists, Glen Osmond, SA. 96 pp PDF']]


 * [[Media:Brady & Ward 2005.pdf|Brady, S.G. & Ward, P.S. 2005. Morphological phylogeny of army ants and other dorylomorphs. Systematic Entomology 30: 592-618. PDF]]


 * [[Media:Brady et al 2006.pdf|Brady, S.G., Schultz, T.R., Fisher, B.L. & Ward, P.S. 2006. Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences U.S.A. 103: 18172-18177. PDF]]


 * [[Media:Gadau et al 1999.pdf|Gadau, J.; Brady, S. G.; Ward, P. S. 1999.  Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Ann. Entomol. Soc. Am. 92(4): 514-522 PDF]]


 * Johnson, B. R.; Borowiec, M. L.; Chiu, J. C.; Lee, E. K.; Atallah, J.; Ward, P. S. 2013. Phylogenomics resolves evolutionary relationships among ants, bees, and wasps. Current Biology 23:2058-2062. [2013-10-21]
 * [[Media:Johnson & Ward- 2002.pdf| Johnson, R. A.; Ward, P. S. 2002. Biogeography and endemism of ants (Hymenoptera: Formicidae) in Baja California, Mexico: a first overview. Journal of Biogeography 29:1009-1026. [2002-08] PDF]]


 * [[Media:Lucky & Ward 2010.pdf|Lucky, A.; Ward, P. S. 2010. Taxonomic revision of the ant genus Leptomyrmex Mayr (Hymenoptera: Formicidae). Zootaxa 2688:1-67. [2010-11-25] PDF]]


 * [[Media:Olson D Ward P 1996.pdf|Olson, D. M.; Ward, P. S. 1996. The ant fauna (Hymenoptera: Formicidae) of Kirindy Forest (tropical dry forest) in western Madagascar. Pp. 161-164 in Ganzhorn, J. U., Sorg, J.-P. (eds.) Ecology and economy of a tropical dry forest in Madagascar. Primate Report 46-1. Göttingen: German Primate Center (DPZ), 382 pp. PDF]]


 * [[Media:Ward 1980a.pdf|Ward, P. S. 1980a. A systematic revision of the Rhytidoponera impressa group (Hymenoptera: Formicidae) in Australia and New Guinea. Aust. J. Zool. 28: 475-498 PDF]]


 * [[Media:Ward 1984.pdf|Ward, P. S. 1984. A revision of the ant genus Rhytidoponera (Hymenoptera: Formicidae) in New Caledonia. Aust. J. Zool. 32: 131-175]]


 * [[Media:Ward 1985b.pdf|Ward, P. S. 1985b. The Nearctic species of the genus Pseudomyrmex (Hymenoptera: Formicidae). Quaest. Entomol. 21: 209-246 PDF]]


 * [[Media:Ward 1986.pdf|Ward, P. S. 1986. Functional queens in the Australian greenhead ant, Rhytidoponera metallica (Hymenoptera: Formicidae). Psyche (Cambridge) 93:1-12. [1986-10-10] PDF]]


 * [[Media:Ward 1988.pdf|Ward, P. S. 1988. Mesic elements in the western Nearctic ant fauna: taxonomic and biological notes on Amblyopone, Proceratium, and Smithistruma (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society. 61:102-124. PDF]]


 * [[Media:Ward 1989a.pdf|Ward, P. S. 1989a. Systematic studies on pseudomyrmecine ants: revision of the Pseudomyrmex oculatus and P. subtilissimus species groups, with taxonomic comments on other species. Quaest. Entomol. 25: 393-468 PDF]]


 * [[Media:Ward 1990.pdf|Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Syst. Entomol. 15: 449-489 PDF]]


 * [[Media:Ward 1991.pdf|Ward, P. S. 1991. Phylogenetic analysis of pseudomyrmecine ants associated with domatia-bearing plants. Pp. 335-352 in: Huxley, C. R., Cutler, D. F. (eds.)  Ant-plant interactions.  Oxford: Oxford University Press, xviii + 601 pp. PDF]]


 * [[Media:Ward 1992.pdf|Ward, P. S. 1992. Ants of the genus Pseudomyrmex (Hymenoptera: Formicidae) from Dominican amber, with a synopsis of the extant Antillean species. Psyche (Camb.) 99: 55-85 PDF]]


 * [[Media:Ward 1993.pdf|Ward, P. S. 1993. Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae). J. Hym. Res. 2: 117-168 PDF]]


 * [[Media:Ward 1994.pdf|Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Syst. Entomol. 19: 159-175 PDF]]


 * [[Media:Ward 1996.pdf|Ward, P. S. 1996. A new workerless social parasite in the ant genus Pseudomyrmex (Hymenoptera: Formicidae), with a discussion of the origin of social parasitism in ants. Syst. Entomol. 21: 253-263 PDF]]


 * Ward, P. S. 1999a. Deceptive similarity in army ants of the genus Neivamyrmex (Hymenoptera: Formicidae): taxonomy, distribution and biology of N. californicus (Mayr) and N. nigrescens (Cresson).  J. Hym. Res. 8: 74-97


 * [[Media:Ward 1999b.pdf|Ward, P. S. 1999b. Systematics, biogeography and host plant associations of the Pseudomyrmex viduus group (Hymenoptera: Formicidae), Triplaris- and Tachigali-inhabiting ants.  Zool. J. Linn. Soc. 126: 451-540 PDF]]


 * [[Media:Ward 2000a.pdf|Ward, P. S. 2000a. On the identity of Pheidole vaslitii Pergande (Hymenoptera: Formicidae), a neglected ant from Baja California. J. Hym. Res. 9: 85-98 PDF]]


 * [[Media:Ward 2001.pdf| Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebr. Taxon.  15: 589-665 PDF]]


 * [[Media:Ward 2005.pdf|Ward, P.S. 2005. A synoptic review of the ants of California. Zootaxa 936: 1-68. PDF]]


 * [[Media:Ward 2006.pdf|Ward, P.S. 2006. The ant genus Tetraponera in the Afrotropical region: synopsis of species groups and revision of the T. ambigua-group. Myrmecologische Nachrichten. 8:119-130. PDF]]


 * [[Media:Ward 2007a.pdf|Ward, P.S. 2007a. Phylogeny, classification, and species-level taxonomy of ants. Zootaxa 1668: 549-563. PDF]]


 * [[Media:Ward 2007b.pdf|Ward, P.S. 2007b. The ant genus Leptanilloides: discovery of the male and evaluation of phylogenetic relationships based on DNA sequence data (pp. 637-649). In Snelling, R.R., Fisher, B.L. & Ward, P.S. (eds). Advances in ant systematics: homage to E.O. Wilson – 50 years of contributions. Memoirs of the American Entomological Institute 80: 690 pp. PDF]]


 * [[Media:Ward 2009.pdf|Ward, P.S. 2009. The ant genus Tetraponera in the Afrotropical region: the T. grandidieri group. Journal of Hymenoptera Research. 18:285-304. PDF]]


 * [[Media:Ward 2014phylogeny and evolution of ants.pdf|Ward, P. S. 2014. The phylogeny and evolution of ants. Annual Review of Ecology, Evolution, and Systematics 45:23-43. PDF]]


 * [[Media:Ward 2017.pdf|Ward, P. S. 2017. A review of the Pseudomyrmex ferrugineus and Pseudomyrmex goeldii species groups: acacia-ants and relatives (Hymenoptera: Formicidae). Zootaxa 4227(4):524-542. PDF]]


 * [[Media:Ward, P.S., Blaimer, B.B. et al. 2016. A revised phylogenetic classification of the ant subfamily Formicinae.pdf|Ward, P.S., Blaimer, B.B., Fisher, B.L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex. Zootaxa 4072 (3): 343–357 (doi 10.11646/zootaxa.4072.3.4).]]


 * Ward, P. S.; Bolton, B.; Shattuck, S. O.; Brown, W. L., Jr. 1996. A bibliography of ant systematics. Univ. Calif. Publ. Entomol. 116: 1-417.


 * [[Media:Ward & Brady 2003.pdf|Ward, P., and Brady, S. 2003. Phylogeny and biogeography of the ant subfamily Myrmeciinae (Hymenoptera: Formicidae). Invertebrate Systematics 17(3): 361-386 PDF]]


 * [[Media:Ward & Brady 2009.pdf|Ward, P.S. & Brady, S.G. 2009. Rediscovery of the ant genus Amyrmex Kusnezov and its transfer from Dolichoderinae to Leptanilloidinae. Zootaxa 2063: 46-54. PDF]]


 * [[Media:Ward Brady Fisher Schultz 2010.pdf|Ward, P.S., Brady, S.G., Fisher, B.L. & Schultz, T.R. 2010. Phylogeny and biogeography of Dolichoderinae ants: effects of data partitioning and relict taxa on historical inference. Systematic Biology 59: 342-362. PDF]]


 * [[Media:Ward et al 2015.pdf|Ward, P. S., S. G. Brady, B. L. Fisher, and T. R. Schultz. 2015. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology. 40:61-81. ]]


 * [[Media:Ward & Downie 2005.pdf|Ward, P.S. & Downie, D.A. 2005. The ant subfamily Pseudomyrmecinae: phylogeny and evolution of big-eyed arboreal ants. Systematic Entomology 30: 310-335. PDF]]
 * [[Media:Ward, P.S., Fisher, B.L. 2016. Tales of dracula ants the evolutionary history of the ant subfamily Amblyoponinae.pdf|Ward, P.S. & Fisher, B.L. 2016. Tales of dracula ants: the evolutionary history of the ant subfamily Amblyoponinae (Hymenoptera: Formicidae). Systematic Entomology, 41, 683–693 (DOI: 10.1111/syen.12186).]]


 * [[Media:Ward & Sumnicht 2012.pdf| Ward, P. S.; Sumnicht, T. P. 2012. Molecular and morphological evidence for three sympatric species of Leptanilla (Hymenoptera: Formicidae) on the Greek island of Rhodes. Myrmecological News 17:5-11. PDF]]


 * [[Media:Ward & Taylor 1981.pdf|Ward, P.S. & Taylor, R.W. 1981. Allozyme variation, colony structure and genetic relatedness in the primitive ant Nothomyrmecia macrops Clark. Journal of the Australian Entomological Society 20: 177-183. [24.viii.1981.] PDF]]
 * Wetterer, J. K.; Ward, P. S.; Wetterer, A. L.; Longino, J. T.; Trager, J. C.; Miller, S. E. 2000. Ants (Hymenoptera: Formicidae) of Santa Cruz Island, California. Bulletin of the Southern California Academy of Sciences 99:25-31.