Solenopsis geminata

The original distribution of S. geminata was Central America to the southern United States. It has been extended to many tropical and sub-tropical areas of the world by human activities. This species is well known as one of the several pestiferous Solenopsis "fire ants". In general S. geminata prefers to nest in open fields or sunny glades, avoiding the shade of deep woods. The nests are usually irregular, sandy craters of loose construction but sometimes rotten stumps are utilized as nesting sites. The ferocity of this ant is proverbial, for the activity of the workers when disturbed never fails to attract attention, however callous the observer (Creighton 1930).

Identification
A member of the Solenopsis geminata species-group.

Wetterer (2012) - Although S. geminata minors are very difficult to distinguish from minors of related fire ants, large majors have several distinctive cephalic characteristics that make them simple to identify. These include: (1) a disproportionally large, almost square head with parallel sides, (2) a deep longitudinal groove on the front of the head extending from a distinct medial indentation in the vertex, (3) black mandibles, often with all teeth worn off from use, and (4) short antennal scapes extending only about halfway to the occiput in the largest majors.

Distribution
In areas near Japan this species has been recorded from the Philippines and Taiwan. In Japan it was recorded from the Nansei Islands (around the U.S. Army base on Okinawa Island and at the radar site on Ie-jima Island). It was almost certainly introduced to these areas by human commerce. This species is recently found on Iwo Islands (Kubota, 1983; Terayama, 1999; Japanese Ant Image Database).

Distribution based on Regional Taxon Lists
Afrotropical Region: Cameroun, Gabon, Guinea, Liberia, South Africa, United Arab Emirates. Australasian Region: Australia, New Caledonia. Indo-Australian Region: American Samoa, Borneo, Cook Islands, Fiji, Guam, Hawaii, Indonesia, Kiribati, Malaysia, Marshall Islands, Micronesia (Federated States of), New Guinea, Northern Mariana Islands, Philippines, Samoa, Solomon Islands, Tonga, Vanuatu. Malagasy Region: Madagascar, Mauritius, Réunion. Nearctic Region: Canada, United States. Neotropical Region: Barbados, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, French Guiana, Galapagos Islands, Greater Antilles, Grenada, Guadeloupe, Guatemala, Guyana, Haiti, Honduras, Lesser Antilles, Mexico, Netherlands Antilles, Nicaragua, Panama, Puerto Rico, Saint Lucia, Suriname, Trinidad and Tobago, Venezuela. Oriental Region: Bangladesh, Cambodia, India, Laos, Nicobar Island, Pakistan, Sri Lanka, Thailand, Vietnam. Palaearctic Region: Canary Islands, China, Greece, Italy, Japan, Republic of Korea.

Biology
Wheeler (1905), Bahamas - very common on New Providence Island wherever there was soil or sand (West Bay, Nassau, Hog Island, Stanley, etc.). It constructs straggling moundlets with many entrances, garners seeds, but still retains its carnivorous instincts, stings fiercely - in short, exhibits all the traits which have gained for it the name of 'hormiga brava' in Cuba and of 'fire ant' in many other localities.

Deyrup, Davis & Cover (2000): There is no reason why a species could not be imported to a place where it is already native. Normally, one would assume that the native population, having evolved adaptive adjustments to the area, would exclude the representatives of the exotic population, or absorb and dilute the small number of immigrants beyond recognition. If, however, the exotic population has undergone natural selection that makes it better able to coexist with humans and take advantage of the resources and habitat modification that humans offer, the exotic population might be favored. Populations of S. geminata have been transported around the World; we have seen specimens from, among other places, a mid-Pacific islet, Johnson Atoll. Some of the confusing variation found among Florida populations could be due to the occurrence of one or more exotic populations overlaid on native populations.

Solenopsis geminata has been observed forming knotted balls of workers clinging to floating wood when their nests are flooded during the rainy season in the Brazilian Pantanal and the Colombian and Venezuelan Llanos (Jaffe, 1993).

Nipitwattanaphon et al. (2020) examined this species in Thailand. They found all examined colonies were polygynous with only a few queens. Queens from the same colonies were highly genetically related. Population structure was partitioned into two clusters. Pairwise FST values revealed very high genetic differentiation between colonies suggesting low gene flow among populations. This result suggests that queens were locally mated and founded colonies by a budding strategy. Isolation-by-distance among local populations was not significant.

Associations with other Organisms
S geminata is parasitized by numerous species of phorid flies.

Porter et al. (2018) studied the fire ant decapitating fly Brown and Morrison (Diptera: Phoridae) and its host specificity. This fly occurs in Texas and neighboring regions of Mexico (Plowes et al. 2009). This fly is 1 of more than 20 species of Pseudacteon decapitating flies known to parasitize tropical fire ants in their native range (Plowes et al. 2009). We found that P. bifidus was not attracted to non-Solenopsis ants presented in the field. Furthermore, no Pseudacteon species known to parasitize Solenopsis fire ants has been observed to parasitize ants in another genera despite extensive field observations.

Worker
Additional images can be found here

Queen
Additional images can be found here

Nomenclature

 * . Atta geminata Fabricius, 1804: 423 (q.) South America (no state data).
 * Type-material: syntype queens (number not stated).
 * [Note: according to Zimsen, 1964: 427, there are 5 syntypes.]
 * Type-locality: South America (“America meridionali”) (no further data).
 * Type-depositories: ZMUC, ZMUK.
 * [Misspelled as germinata by Wheeler, W.M. 1927e: 1.]
 * Roger, 1862c: 289 (w.m.); Mayr, 1867a: 109 (w.q.m.); Wheeler, W.M. 1900b: 21 (l.); Wheeler, G.C. & Wheeler, J. 1955c: 132 (l.); Crozier, 1970: 116 (k.).
 * Combination in Solenopsis: Mayr, 1863: 453; Roger, 1863b: 32.
 * [Combination in Monomorium: Mayr, 1884: 37 (error).]
 * Status as species: Roger, 1862c: 289; Roger, 1863b: 32, 49; Mayr, 1863: 453; Mayr, 1865: 108; Mayr, 1867a: 109 (redescription); Mayr, 1870b: 996 (in key); Smith, F. 1873: ix; Mayr, 1876: 111; Emery, 1878a: x (in list); Smith, F. 1879: 676; Forel, 1881: 10; Emery, 1883: 151; Forel, 1885b: 182; Mayr, 1886c: 365; Mayr, 1886d: 460; Cresson, 1887: 262; Emery, 1888c: 355; Emery, 1888e: 690; Rothney, 1889: 365; Emery, 1890a: 66; Emery, 1890b: 52; Forel, 1893g: 396; André, 1893b: 152; Dalla Torre, 1893: 76; Emery, 1893d: 89; Emery, 1893e: 191; Emery, 1893f: 243; Emery, 1893g: 266; Pergande, 1893: 35; von Jhering, 1894: 392; Emery, 1894k: 56; Forel, 1895b: 130; Emery, 1895c: 276; Emery, 1895k: 467; Emery, 1896g: 83 (in key); Emery, 1896h: 625; Mayr, 1897: 430; Forel, 1897b: 300; Forel, 1899c: 79; Emery, 1900d: 688; Emery, 1901f: 120; Emery, 1901g: 567; Forel, 1901c: 129; Rothney, 1903: 98; Bingham, 1903: 158; Wheeler, W.M. 1905b: 124; Emery, 1906c: 121; Forel, 1906d: 248; Forel, 1907e: 4; Forel, 1908b: 45; Forel, 1908c: 362; Forel, 1908e: 67; Wheeler, 1908a: 130; Wheeler, W.M. 1908e: 424; Wheeler, W.M. 1909b: 232; Forel, 1909a: 259, 268; Santschi, 1910c: 359; Wheeler, W.M. 1910g: 563; Santschi, 1911d: 3; Emery, 1911b: 531; Wheeler, W.M. 1911a: 23; Wheeler, W.M. 1911b: 169; Forel, 1912g: 4; Wheeler, W.M. 1913b: 493; Wheeler, W.M. 1913c: 115; Wheeler, W.M. 1913d: 240; Santschi, 1913c: 306; Stitz, 1913: 209; Forel, 1914e: 11; Bruch, 1914: 223; Emery, 1914f: 393; Wheeler, W.M. & Mann, 1914: 20; Mann, 1916: 447; Wheeler, W.M. 1916c: 3; Wheeler, W.M. 1916d: 324; Crawley, 1916b: 370; Wheeler, W.M. 1917g: 458; Luederwaldt, 1918: 42; Wheeler, W.M. 1918b: 24; Wheeler, W.M. 1919c: 272; Mann, 1920: 427; Emery, 1922e: 197; Wheeler, W.M. 1922a: 877; Wheeler, W.M. 1922c: 9; Mann, 1922: 30; Wheeler, W.M. 1923c: 4; Stitz, 1925: 119; Essig, 1926: 858; Stärcke, 1926: 84 (in key); Borgmeier, 1927c: 104; Wheeler, W.M. 1927b: 45; Creighton, 1930b: 59 (redescription); Menozzi & Russo, 1930: 158; Mukerjee, 1930: 154; Smith, M.R. 1930a: 3; Wheeler, W.M. 1932a: 10; Donisthorpe, 1932c: 463; Donisthorpe, 1933c: 534; Wheeler, W.M. 1933a: 62; Borgmeier, 1934: 102; Wheeler, W.M. 1935g: 26; Wheeler, W.M. 1936b: 200; Cole, 1937a: 99; Smith, M.R. 1937: 838; Wheeler, W.M. 1938: 252; Menozzi, 1942: 169; Eidmann, 1944: 450; Donisthorpe, 1946e: 31; Weber, 1948b: 82; Creighton, 1950a: 231; Smith, M.R. 1951a: 812; Chapman & Capco, 1951: 168; Cole, 1953g: 299; Smith, M.R. 1958c: 129; Kempf, 1961b: 507; Snelling, R.R. 1963: 7; Wilson, 1964b: 8; Baltazar, 1966: 260; Ettershank, 1966: 141; Smith, M.R. 1967: 357; Wilson & Taylor, 1967: 58; Taylor, 1967b: 1094; Kempf, 1972a: 235; Alayo, 1974: 14 (in key); Smith, D.R. 1979: 1385; Taylor, 1987a: 72; Zolessi, et al. 1988: 4; Deyrup, et al. 1989: 96; Brandão, 1991: 379; Ogata, 1991b: 110; Trager, 1991: 163 (redescription); Morisita, et al. 1992: 42; Collingwood, 1993b: 194; Perrault, 1993: 333; Dlussky, 1994: 54; Bolton, 1995b: 387; Wu, J. & Wang, 1995: 70; Tang, J., Li, et al. 1995: 71; Collingwood, Tigar & Agosti, 1997: 508; Tiwari, 1999: 58; Zhou, 2001b: 88; Wetterer, 2002: 129; Blard, et al. 2003: 131; Deyrup, 2003: 47; Imai, et al. 2003: 132; Lin & Wu, 2003: 66; Wetterer & Vargo, 2003: 417; Ghosh, et al. 2005: 33; Jaitrong & Nabhitabhata, 2005: 42; MacGown & Forster, 2005: 69; Clouse, 2007b: 249; Framenau & Thomas, 2008: 72; Terayama, 2009: 157; Mohanraj, et al. 2010: 7; Collingwood, et al. 2011: 438; Legakis, 2011: 15; Pfeiffer, et al. 2011: 51; Wetterer, 2011a: 21; Borowiec, L. & Salata, 2012: 534; Branstetter & Sáenz, 2012: 261; Guénard & Dunn, 2012: 53; Sarnat & Economo, 2012: 122; Sharaf & Aldawood, 2012: 10; Sarnat, et al. 2013: 72; Borowiec, L. 2014: 154; Ramage, 2014: 162; Bezděčková, et al. 2015: 122; Bharti, Guénard, et al. 2016: 44; Jaitrong, Guénard, et al. 2016: 38; Wetterer, et al. 2016: 17; Deyrup, 2017: 103; Salata & Borowiec, 2018c: 48; Fernández & Serna, 2019: 816; Lubertazzi, 2019: 172; Dias, R.K.S. et al. 2020: 90; Khachonpisitsak, et al. 2020: 122.
 * Senior synonym of bahiaensis: Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of cephalotes: Roger, 1863b: 32, 49; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of clypeata: Roger, 1862c: 289; Mayr, 1863: 453; Roger, 1863b: 32; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of coloradensis: Mayr, 1886c: 365; Mayr, 1886d: 460; Cresson, 1887: 259; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of diabola: Ettershank, 1966: 141; Kempf, 1972a: 236; Smith, D.R. 1979: 1386; Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of drewseni: Mayr, 1870b: 996 (footnote); Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of eduardi: Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of galapageia: Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of glaber: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1899c: 79; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of innota: Wheeler, W.M. 1922a: 877; Trager, 1991: 164; Bolton, 1995b: 387.
 * Senior synonym of laboriosus: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of laevissima: Donisthorpe, 1932c: 463; Trager, 1991: 163; Bolton, 1995b: 387.
 * Senior synonym of lincecumii: Emery, 1895c: 276; Wheeler, W.M. 1902f: 28; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387.
 * Senior synonym of mandibularis: Roger, 1862c: 289; Mayr, 1863: 453; Roger, 1863b: 32; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Wheeler, W.M. 1911f: 172; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of medusa: Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of mellea: Donisthorpe, 1932c: 455; Bolton, 1995b: 387.
 * Senior synonym of nigra: Creighton, 1930b: 59; Kempf, 1972a: 236; Trager, 1991: 163; Bolton, 1995b: 387.
 * Senior synonym of paleata: Roger, 1863b: 32, 49; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1895b: 130; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of perversa: Trager, 1991: 163; Bolton, 1995b: 388.
 * Senior synonym of polita: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1899c: 79; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * Senior synonym of rufa: Dalla Torre, 1893: 76; Ettershank, 1966: 141; Wilson & Taylor, 1967: 58; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Tiwari, 1999: 58; Zhou, 2001b: 88; Ramage, 2014: 162.
 * Senior synonym of saxicola: Emery, 1895c: 276; Wheeler, W.M. 1902f: 28; Emery, 1922e: 197; Creighton, 1930b: 59; Borgmeier, 1927c: 104; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Bolton, 1995b: 387.
 * Material of the nomen nudum minor referred here by Kempf, 1972a: 236.
 * Distribution [tramp species]
 * Afrotropical: Cameroon, Congo, Democratic Republic of Congo, Equatorial Guinea, Gabon, Guinea, Liberia, Nigeria, Senegal, Sierra Leone, South Africa.
 * Austral: Australia, New Caledonia, New Zealand.
 * Malagasy: Madagascar.
 * Malesian: Brunei, Christmas I., Cocos-Keeling Is, Cook Is, Fiji, French Polynesia, Hawaii, Mariana Is, Micronesia, Malaysia, Marshall Is, Indonesia, Papua New Guinea, Palau, Philippines, Samoa, Singapore, Solomon Is, Tonga, Tuvalu, Vanuatu.
 * Nearctic: Canada, U.S.A.
 * Neotropical: Anguilla, Antigua, Aruba, Bahamas, Barbados, Barbuda, Belize, Bolivia, Bonaire, Brazil, Cayman Is, Colombia, Costa Rica (+ Cocos Is), Cuba, Curaçao, Dominica, Dominican Republic, Ecuador (+ Galapagos), El Salvador, French Guiana, Grenada, Guadeloupe, Guatemala, Guyana, Haiti, Honduras, Jamaica, Martinique, Mexico (+ Revillagigedo Is), Montserrat, Nevis, Nicaragua, Panama, Peru, Puerto Rica, St Kitts, St Lucia, St Martin, St Vincent, Suriname, Trinidad, Turks & Caicos, Uruguay, Venezuela, Virgin Is.
 * Oriental: Bangladesh, China, India, Japan, Laos, Myanmar, Sri Lanka, Taiwan, Thailand, Vietnam.
 * Palaearctic: Britain, Cyprus, Greece, Italy, Netherlands, Saudi Arabia, Spain (Canary Is), United Arab Emirates.
 * Current subspecies: nominal plus micans.
 * bahiaensis. Solenopsis edouardi var. bahiaensis Santschi, 1925d: 237 (w.) BRAZIL (Bahia), VENEZUELA.
 * Type-material: 31syntype workers.
 * Type-localities: 30 workers Brazil: Bahia (Bondar), 1 worker Venezuela (no further data).
 * Type-depository: NHMB.
 * [Misspelled as bahiana by Borgmeier, 1927c: 104.]
 * Junior synonym of medusa: Creighton, 1930b: 68; Kempf, 1972a: 236.
 * Junior synonym of geminata: Trager, 1991: 163; Bolton, 1995b: 388.
 * cephalotes. Solenopsis cephalotes Smith, F. 1859a: 149 (s.w.) INDONESIA (Aru Is).
 * Type-material: 3 syntype workers.
 * Type-locality: Indonesia: Aru Is (A.R. Wallace).
 * Type-depository: OXUM.
 * Status as species: Smith, F. 1860b: 113; Smith, F. 1861b: 48; Smith, F. 1863: 21; Smith, F. 1871a: 333.
 * Junior synonym of saevissima: Mayr, 1863: 454.
 * Junior synonym of rufa: Forel, 1911e: 268; Emery, 1922e: 197; Donisthorpe, 1932c: 456; Creighton, 1950a: 231.
 * Junior synonym of geminata: Roger, 1863b: 32, 49; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 386; Zhou, 2001b: 88.
 * clypeata. Atta clypeata Smith, F. 1858b: 169 (q.m.) MEXICO (no state data).
 * Type-material: 3(?) syntype queens, 1 syntype male.
 * Type-locality: Mexico (no further data).
 * Type-depository: BMNH.
 * Status as species: Norton, 1868a: 66.
 * Junior synonym of geminata: Roger, 1862c: 289; Mayr, 1863: 453; Roger, 1863b: 32; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 386; Zhou, 2001b: 88.
 * coloradensis. Atta coloradensis Buckley, 1867: 346 (s.w.) U.S.A. (Texas).
 * Type-material: syntype workers (number not stated).
 * Type-locality: U.S.A.: Texas, nr Colorado river (S.B. Buckley).
 * Type-depository: unknown (no material known to exist).
 * [Note: Wheeler, W.M. 1902f: 29, comments that this taxon contains a mixture of Solenopsis geminata and a species of Pheidole.]
 * Junior synonym of geminata: Mayr, 1886c: 365; Mayr, 1886d: 460; Cresson, 1887: 259; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * diabola. Solenopsis geminata var. diabola Wheeler, W.M. 1908e: 424 (w.q.m.) U.S.A. (Texas, Arizona, California).
 * Type-material: syntype workers, syntype queens, syntype males (numbers not stated).
 * Type-localities: U.S.A.: Texas, Montopolis, Austin, Lampasas, and Langtry (W.M. Wheeler). Texas, Paris (A. Rucker), Texas, Dallas (W.D. Hunter), Texas, Granite Mt (H.W. Long), Texas, Plano (E.S. Tucker), Arizona, Tempe (W.M. Wheeler), California, Needles (W.M. Wheeler).
 * Type-depository: MCZC.
 * Subspecies of geminata: Forel, 1908c: 362; Forel, 1909a: 250; Wheeler, W.M. 1909b: 232; Wheeler, W.M. 1910g: 563; Forel, 1912g: 4; Wheeler, W.M. 1915b: 397; Luederwaldt, 1918: 43; Emery, 1922e: 197; Borgmeier, 1927c: 105.
 * Junior synonym of rufa: Creighton, 1930b: 66; Creighton, 1950a: 231; Smith, M.R. 1951a: 812.
 * Junior synonym of geminata: Ettershank, 1966: 141; Kempf, 1972a: 236; Smith, D.R. 1979: 1386; Trager, 1991: 163; Bolton, 1995b: 387.
 * drewseni. Diplorhoptrum drewseni Mayr, 1861: 73 (diagnosis in key) (w.) ITALY.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Italy: (no further data) (Drewsen).
 * Type-depository: NHMW.
 * Combination in Solenopsis: Mayr, 1863: 453.
 * Status as species: Mayr, 1863: 453; Dours, 1873: 170; André, 1874: 200 (in key).
 * Junior synonym of geminata: Mayr, 1870b: 996 (footnote); Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 387; Zhou, 2001b: 88.
 * eduardi. Solenopsis eduardi Forel, 1912g: 12 (w.) COLOMBIA.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Colombia: Rio Frio (A.Forel).
 * Type-depository: MHNG.
 * [Misspelled as edouardi by Santschi, 1923c: 263, and others.]
 * Status as species: Emery, 1922e: 197; Santschi, 1923c: 263; Santschi, 1925b: 13; Borgmeier, 1927c: 104; Ettershank, 1966: 140 (error).
 * Subspecies of geminata: Creighton, 1930b: 67; Santschi, 1931c: 271; Kempf, 1972a: 236.
 * Junior synonym of geminata: Trager, 1991: 163; Bolton, 1995b: 387.
 * galapageia. Solenopsis geminata var. galapageia Wheeler, W.M. 1919c: 272 (w.q.) ECUADOR (Galapagos Is).
 * Type-material: syntype workers, syntype queens (numbers not stated, “several”).
 * Type-locality: Ecuador: Galapagos Is, Charles I., 1905-6 (F.X. Williams).
 * Type-depositories: CASC, MCZC.
 * Subspecies of geminata: Creighton, 1930b: 65; Wheeler, W.M. 1935g: 26; Ettershank, 1966: 141; Linsley & Usinger, 1966: 175; Kempf, 1972a: 236.
 * Junior synonym of geminata: Trager, 1991: 163; Bolton, 1995b: 387.
 * glaber. Myrmica glaber Smith, F. 1862b: 34 (w.) PANAMA.
 * Type-material: holotype (?) worker.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: Panama: (no further data) (R.W. Stretch).
 * Type-depository: BMNH.
 * Combination in Solenopsis: Roger, 1863b: 32.
 * Status as species: Roger, 1863b: 32; Mayr, 1863: 432.
 * Junior synonym of geminata: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1899c: 79; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 388; Zhou, 2001b: 88.
 * innota. Solenopsis geminata var. innota Santschi, 1915c: 257, fig. 6 (s.w.q.m.) GABON, LIBERIA, CONGO.
 * Type-material: syntype workers, syntype queens, syntype males (numbers not stated).
 * Type-localities: Gabon: Samkita, 1914 (F. Faure), Liberia: Monrovia (Delafosse), Congo: Ngoma (Galli-Valerio).
 * Type-depositories: MNHN, NHMB.
 * Subspecies of geminata: Emery, 1922e: 197.
 * Junior synonym of geminata: Wheeler, W.M. 1922a: 877; Trager, 1991: 164; Bolton, 1995b: 388.
 * laboriosus. Crematogaster laboriosus Smith, F. 1860b: 109 (q.) INDONESIA (Bacan I.).
 * Type-material: holotype queen.
 * Type-locality: Indonesia: Bachian (= Bacan I.), “Bac.23” (A.R. Wallace).
 * Type-depository: OXUM.
 * Combination in Solenopsis: Mayr, 1863: 453.
 * Status as species: Roger, 1863b: 37.
 * Junior synonym of rufa: Emery, 1922e: 197; Creighton, 1930b: 66; Creighton, 1950a: 231; Donisthorpe, 1932c: 463.
 * Junior synonym of geminata: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Emery, 1922e: 197; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 388; Zhou, 2001b: 88.
 * laevissima. Myrmica laevissima Smith, F. 1860b: 108 (w.) INDONESIA (Bacan I.).
 * Type-material: 2 syntype workers.
 * Type-locality: Indonesia: Bachian (= Bacan I.), “Bac.7”, “found in houses” (A.R. Wallace).
 * Type-depository: OXUM.
 * [Misspelled as levissima by Dalla Torre, 1893: 111.]
 * Status as species: Mayr, 1863: 433; Smith, F. 1871a: 325; Dalla Torre, 1893: 111; Wheeler, W.M. 1918b: 24; Chapman & Capco, 1951: 126.
 * Junior synonym of geminata: Donisthorpe, 1932c: 463; Trager, 1991: 163; Bolton, 1995b: 388.
 * lincecumii. Atta lincecumii Buckley, 1867: 344 (s.w.) U.S.A. (Texas).
 * Type-material: syntype workers (number not stated).
 * Type-locality: U.S.A.: Texas, Central Texas, near streams (S.B. Buckley).
 * Type-depository: unknown (no type-material is known to exist).
 * Combination in Aphaenogaster: Cresson, 1887: 260.
 * Status as species: Cresson, 1887: 260.
 * Junior synonym of geminata: Emery, 1895c: 276; Wheeler, W.M. 1902f: 28; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 388.
 * mandibularis. Solenopsis mandibularis Westwood, 1840b: 87, pl. 2, fig. 5 (w.) “America” (no state data).
 * Type-material: syntype workers (number not stated).
 * Type-locality: "America Aequinoctiali": (no further data) (D.L. Guilding).
 * Type-depository: OXUM.
 * Status as species: Smith, F. 1858b: 178.
 * Junior synonym of geminata: Roger, 1862c: 289; Mayr, 1863: 453; Roger, 1863b: 32; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Wheeler, W.M. 1911f: 172; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 389; Zhou, 2001b: 88.
 * medusa. Solenopsis geminata subsp. medusa Mann, 1916: 447, pl. 4, fig. 31 (w.) BRAZIL (Rio Grande do Norte, Ceará).
 * Type-material: syntype workers (numbers not stated, “a number of colonies”).
 * Type-localities: Brazil: Rio Grande do Norte, Ceará-Mirim, 1911 (W.M. Mann), Brazil: Ceará, Maranguapé Mts, 1911 (W.M. Mann).
 * Type-depositories: LACM, MCZC.
 * [Solenopsis geminata subsp. medusa Wheeler, W.M. 1915b: 397. Nomen nudum (attributed to Mann).]
 * Creighton, 1930b: 69 (q.).
 * Subspecies of geminata: Emery, 1922e: 197; Borgmeier, 1927c: 105; Creighton, 1930b: 68; Ettershank, 1966: 142; Kempf, 1972a: 236.
 * Junior synonym of geminata: Trager, 1991: 163; Bolton, 1995b: 389.
 * mellea. Myrmica mellea Smith, F. 1859a: 148 (w.) INDONESIA (Aru Is).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Aru Is, “Aru” (A.R. Wallace).
 * Type-depository: OXUM.
 * [Misspelled as mellae by Chapman & Capco, 1951: 127.]
 * Status as species: Mayr, 1863: 433; Smith, F. 1871a: 325; Dalla Torre, 1893: 112.
 * Junior synonym of geminata: Donisthorpe, 1932c: 455; Bolton, 1995b: 389.
 * nigra. Solenopsis geminata var. nigra Forel, 1908b: 45 (w.) COSTA RICA.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Costa Rica: Zent (P. Biolley).
 * Type-depository: MHNG.
 * Subspecies of geminata: Forel, 1909a: 268; Forel, 1913l: 225; Wheeler, W.M. 1915b: 397; Emery, 1922e: 197; Menozzi, 1927c: 268; Santschi, 1930e: 78; Menozzi, 1931b: 267; Santschi, 1939f: 162; Alayo, 1974: 34.
 * Junior synonym of geminata: Creighton, 1930b: 59; Kempf, 1972a: 236; Trager, 1991: 163; Bolton, 1995b: 389.
 * paleata. Myrmica paleata Lund, 1831a: 116 (footnote) (w.) BRAZIL (Minas Gerais).
 * Type-material: syntype workers (number not stated, “numerous”).
 * Type-locality: Brazil: Minas Gerais (Lund).
 * Type-depository: ZMUC.
 * [Also described as new by Lund, 1831b: 100.]
 * Junior synonym of geminata: Roger, 1863b: 32, 49; Mayr, 1865: 108; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1895b: 130; Trager, 1991: 163; Bolton, 1995b: 390; Zhou, 2001b: 88.
 * perversa. Solenopsis edouardi var. perversa Santschi, 1925b: 13 (s.w.q.) BRAZIL (Pernambuco).
 * Type-material: syntype workers, syntype queens (numbers not stated).
 * Type-locality: Brazil: Pernambuco, Tapera (no collector’s name, “sent by R.P. Wasermann”).
 * Type-depository: NHMB.
 * Subspecies of eduardi: Borgmeier, 1927c: 104.
 * Junior synonym of eduardi: Creighton, 1930b: 67; Kempf, 1972a: 236; Bolton, 1995b: 390.
 * polita. Myrmica polita Smith, F. 1862b: 34 (w.) PANAMA.
 * Type-material: holotype (?) worker.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: Panama: (no further data) (R.W. Stretch).
 * Type-depository: BMNH.
 * [Unresolved junior primary homonym of Myrmica polita Smith, F. 1860b: 108 (Bolton, 1995b: 390).]
 * Combination in Solenopsis: Roger, 1863b: 32.
 * Status as species: Roger, 1863b: 32; Mayr, 1863: 434.
 * Junior synonym of geminata: Mayr, 1863: 453; Mayr, 1886c: 362; Mayr, 1886d: 460; Dalla Torre, 1893: 76; Pergande, 1893: 35; Forel, 1899c: 79; Emery, 1922e: 197; Borgmeier, 1927c: 104; Creighton, 1930b: 59; Creighton, 1950a: 231; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 390; Zhou, 2001b: 88.
 * rufa. Atta rufa Jerdon, 1851: 106 (w.q.) INDIA (Kerala, Karnataka).
 * Type-material: syntype workers, syntype queens (numbers not stated).
 * Type-locality: India: “very common in Malabar, but is also found in the Carnatic”…..”often appears in houses” (T.C. Jerdon).
 * Type-depository: unknown (no type-material known to exist).
 * [Duplicated in Jerdon, 1854a: 48.]
 * Bingham, 1903: 158 (m.).
 * Combination in Solenopsis: Emery, 1892b: 166.
 * Status as species: Smith, F. 1858b: 163; Mayr, 1863: 397.
 * Subspecies of geminata: Forel, 1899a: 119; Forel, 1901b: 14; Bingham, 1903: 159; Forel, 1903a: 689; Forel, 1903d: 405; Wheeler, W.M. 1907a: 272; Forel, 1908a: 3; Wheeler, W.M. 1908c: 165; Forel, 1909a: 268; Forel, 1909e: 394; Wheeler, W.M. 1909d: 340; Forel, 1910d: 123; Wheeler, W.M. 1910g: 563; Forel, 1911e: 268; Wheeler, W.M. 1912a: 45; Forel, 1913k: 55; Emery, 1914f: 412; Crawley, 1915a: 135; Crawley, 1915b: 239; Wheeler, W.M. 1915b: 397; Viehmeyer, 1916b: 285; Santschi, 1919a: 326; Wheeler, W.M. 1919e: 84; Santschi, 1920h: 161; Emery, 1922e: 197; Santschi, 1924c: 99; Crawley, 1924: 399; Stitz, 1925: 119; Wheeler, W.M. 1927b: 45; Wheeler, W.M. 1927e: 1; Wheeler, W.M. 1927h: 90; Cheesman & Crawley, 1928: 518; Santschi, 1928h: 125; Wheeler, W.M. 1928c: 20; Creighton, 1930b: 66; Menozzi, 1930d: 327; Smith, M.R. 1930a: 3; Wheeler, W.M. 1930h: 68; Wheeler, W.M. 1932a: 10; Wheeler, W.M. 1932d: 16; Santschi, 1932b: 14; Wheeler, W.M. 1934h: 12; Donisthorpe, 1935: 633; Karavaiev, 1935a: 94; Wheeler, W.M. 1935g: 26; Wheeler, W.M. 1936f: 8; Cole, 1937a: 99; Creighton, 1950a: 231; Smith, M.R. 1951a: 812; Chapman & Capco, 1951: 168; Baltazar, 1966: 260.
 * Junior synonym of geminata: Dalla Torre, 1893: 76; Ettershank, 1966: 141; Wilson & Taylor, 1967: 58; Kempf, 1972a: 235; Smith, D.R. 1979: 1385; Trager, 1991: 163; Bolton, 1995b: 390; Tiwari, 1999: 57; Zhou, 2001b: 88; Ramage, 2014: 162.
 * saxicola. Myrmica (Monomorium) saxicola Buckley, 1867: 341 (w.) U.S.A. (Texas).
 * Type-material: syntype workers (number not stated).
 * Type-locality: U.S.A.: Texas, Buchanan County (S.B. Buckley).
 * Type-depository: unknown (no type-material is known to exist).
 * Combination in Monomorium: Cresson, 1887: 262.
 * Status as species: Cresson, 1887: 262; Dalla Torre, 1893: 116.
 * Junior synonym of geminata: Emery, 1895c: 276; Wheeler, W.M. 1902f: 28; Emery, 1922e: 197; Creighton, 1930b: 59; Borgmeier, 1927c: 104; Smith, M.R. 1951a: 812; Smith, D.R. 1979: 1385; Bolton, 1995b: 391.

Type Material


Crematogaster laboriosus

Holotype queen in. Labelled “Bac. 23.”

Myrmica laevissima

Two worker syntypes in. Labelled “Bac. 7.”

Myrmica mellea

Holotype worker in. Labelled “Aru.”

Solenopsis cephalotes

Three worker syntypes (on a single card) in. Labelled “Aru.” (= Aru I. New Guinea).

Two specimens in, labelled by Donisthorpe as syntypes of cephalotes, have “Men.” (= Menado, Sulawesi) as their locality. Smith gives the type-locality as “Aru.” All the specimens, from both localities, appear to be S. geminata (Fabricius).

Worker
Trager (1991) - MEASUREMENTS AND INDICES: HL 1.06-2.20, HWO.98-2.33, SLO.78-1.l4, EL 0.15-0.29, PW 0.57-1.06, AL 1.18-2.08, CI 92-108, SI 47-84, 01 11-16. N = 34.

WORKER DIAGNOSIS. Head (ffv) subquadrate to subtrapezoidal (sides often divergent or flaring anteriorly, especially in specimens from southern Central America and eastern South America), with sides straight to weakly convex and parallel to weakly divergent anteriad (sides weakly convergent anteriad in specimens from southern Texas), often slightly indented just anterior to eyes; posterior border with deep angular median emargination between two nearly hemispherical lobes ("temples"); emargination 1.0-1.5 x as wide as distance between apices of frontal lobes; median clypeal tooth lacking or (rarely) rudimentary; carinal teeth thick at base, strongly protruding, clypeal border between them concave; mandibles thick and strongly curved mesad, especially in largest individuals; mandibular teeth present in all individuals upon eclosion, but often worn off through seed-milling by larger individuals, such that apices dulled or flattened; mandibular costulae complete in smaller majors, to irregular and largely obsolete in larger majors; eye (lv) appearing small relative to hypertrophied head, greatest diameter with 9-11 facets, least diameter with 7-9; largest majors rarely with median ocellus more or less well developed; scapes (ffv) short, curved, scape failing to reach apices of occipital lobes by 0.3-0.5 x SL; pronotum with rounded, at most faintly angular anterolateral corners; promesonotal suture conspicuous, approximately right-angular to weakly obtuse-angular, raised as a small boss at most anterior point; promesonotal profile (lv) formed of 2 convexities meeting at anterior mesonotal boss, pronotal profile more strongly convex and at most feebly angular; anteroventral border of mesopleuron thickened, often bearing one or more spine-like, triangular, lobate or rectangular projecting flanges; metanotal impression marked, set off by a ridge at its juncture with propodeum; propodeal profile more or less diamond-shaped, with dorsum flat to weakly concave; descending though obtuse, carinate angles to weakly convex declivous face; petiolar peduncle as long as or a little longer than base of node; profile of petiolar node cuneate with a relatively sharp crest; postpetiole 1.02-1.08 x as wide as petiole.

Piligerous foveolae of head and thoracic dorsum conspicuous and abundant, 0.025 or more in diameter, those near mouthparts and on sides ofhead sometimes elongate; on larger specimens, mesopleuron largely unsculptured to feebly rugose, but coarsely rugose along pleural suture and near edges of sclerite, especially anteroventrally (in smaller specimens, mesopleuron striate-punctate to coarsely rugose); dorsum of propodeum with a pair of irregular dorsolateral carinae, these best developed at juncture of dorsal and declivous faces; posteriorly, transverse striae or rugae may occur on dorsal, concave surface between carinae; area surrounding propodeal spiracle encircled by coarse, irregular rugosity; declivous face of propodeum with transverse rugae contiguous with those of metapleuron on lower portion, but on upper part more neatly aligned than, and not always contiguous with those on side of propodeum; lateral carinae usually obsolescent on all but uppermost portion of propodeal declivous face; petiolar peduncle transversely striate; base of node areolate-punctate; petiolar ventral process consisting of 1 or 2 small teeth, or rarely, a very narrow, transparent flange; dorsum and anterior face of petiolar node sparsely punctate foveolate, sometimes dorsum with transverse striation like that on posterior face, dorsal margin weakly scalloped; posterior face of petiolar node transversely striate to weakly rugose-areolate below, sparsely sculptured, or less often sculptured as below near top, though less coarsely; sides of postpetiole rugose-punctate; venter of postpetiole dull, coarsely punctate; dorsum ofpostpetiole weakly scalloped, usually shiny and unsculptured or with a weaker version of sculpture below; posterior face of postpetiole transversely rugose-punctate.

Pilosity of head and promesonotum abundant, 0.13--0.37 mm in length; some pilosity often present on meso- and metapleuron.

Color highly variable, though generally fairly consistent within a colony; ranging from concolorous orange-red with only posterior portion of gaster dark brown (var. rufa), to nearly concolorous brownish black with only head near base of mandibles and appendages (especially distally) reddish-brown (var. nigra). Smaller workers tend to be darker and more uniformly colored than bigger ones. Darker S. geminata are possibly limited to or at least prefer more humid microhabitats, and ecological conditions during rearing may be at least partly responsible for adult coloration, but this needs study. Redder variants often are, or at least appear less sculptured than darker forms, but are more likely to have mesopleural flanges. However, I have studied samples from single colonies with virtually the entire range of color and sculpture described above, and S. geminata individuals may have any possible combination of color and sculpture.

Queen
Creighton (1930) - Length 7.5-8 mm.

Head, exclusive of the mandibles, one-sixth broader than long, quadrate, a little wider behind the eyes than in front of them, the sides very feebly convex from the eyes to the occipital angles, straight or nearly straight in front of the eyes and meeting the anterior border of the head at a sharp angle. Occipital angles well-marked, the occiput flat with a narrow and shallow median impression, occipital furrow clearly defined, frontal furrow short, clearly marked only for about half the distance from the median ocellus to the base of the frontal lobes, thereafter becoming shallow and indistinct. Ocelli large and prominent. Clypeus feebly projecting, carinal teeth very stout and rather blunt, the edge of the clypeus between them with a shallow concave impression ; lateral denticles small, often poorly defined and in some cases represented only by a sinuousity in the edge of the clypeus. Mandibles strongly bent but less so than in the major worker, the masticatory border with three large teeth and usually the rudiment of a fourth. Eyes large, strongly convex, irregularly oval in outline, their posterior border reaching a point half way between the occiput and the anterior border of the head. The antennal scape in repose just reaches the lateral ocellus. Funicular joints and club as in the major worker.

Thorax robust, elliptical, its maximum width three-fifths of its length, only slightly narrower than the head (the eyes excluded). Seen in profile the mesonotum shows a straight posterior half and a convex anterior portion which overhangs the pronotum. Scutellum as high as the mesonotum, slightly convex with a short, perpendicular posterior face. Angle of the epinotum well-defined but very obtuse, the basal and declivious faces of about equal length. Mesosternum large and subglobose beneath.

Petiolar nodes very similar to those of the major worker except that the peduncle is thicker, the node of the petiole slightly lower and the postpetiole bears on either side an obtuse, somewhat conical, ventral projection with a small opening at its summit (this condition is sometimes found in the major worker but in that caste the conical projection is usually absent and the opening occurs as a small tubercle on the side of the node). Seen from above the nodes are very strongly transverse and of approximately equal width. Abdomen as in the major worker. Wings hyaline with yellow veins.

Punctures smaller and less numerous than in the major worker. Somewhat larger on the head than on the thorax and abdomen. Body hairs long, golden and erect, somewhat longer on the head than elsewhere, longest on the anterior edge of the clypeus. Mandibles with a few coarse, indistinct striae, epinotuin almost completely covered with fine wavy striae, petiolar nodes, except their summits which are shining, striato-rugulose. For the rest smooth and shining. The color varies from a clear yellowish brown with the front of the head, the mesosternum and the appendages paler and the mandibles and the posterior half of the abdomen castaneous, to a deep castaneous brown with only the extreme anterior portion of the head yellowish brown.

Male
Creighton (1930) - Length 5.8 mm.

Head trapezoidal, its maximum width (including the eyes) approximately one-fourth greater than its length. Eyes very large, strongly convex and oval in outline, occupying more than one-half the. side of the head, their anterior border reaching the insertion of the mandible. Ocelli very large and prominent, the lateral ocelli which mark the boundary of the occiput with a shallow concave impression between them. Anterior edge of the clypeus approximately straight; seen from the side the clypeus shows a blunt, beak-like central lobe. Mandibles small, linear, bidentate. Antennal scape about one and one-half times as long as broad, roughly cylindrical ; first funicular joint sub globose, broader than the scape or the following joint; second funicular joint more than twice as long as broad, third joint one and one-half times as long as broad, the remaining joints all more than twice as long as broad and progressively decreasing in width.

Thorax bulky, elliptical, its greatest width two-thirds of its length, only slightly less than twice as wide as the head (eyes included). Seen from the side the anterior part of the mesonotum is greatly swollen and overhangs the pronotum which is so much displaced that the head of the insect appears to be attached to the ventral surface of the thorax. Epinotum rather rounded, the basal face strongly convex transversely and slightly convex longitudinally, declivious face flat and virtually perpendicular. Node of the petiole in profile low but*with an acute summit, the anterior face not sharply separated from the thick peduncle, the posterior face perpendicular. Seen from behind the summit of the node shows a broad, shallow median impression. Postpetiole in profile as high as the node of the petiole, about one and one-half times as high as long with a long, backward sloping anterior face, a rounded summit and a very declivious posterior face. The conical lateral projections are even stronger than in the female. Seen from above both nodes are very transverse, the postpetiole is approximately three times as broad as long· and one-sixth wider than. he node of the petiole. First gastric segment truncate at the base but not impressed. Wings hyaline, the veins clear yellow.

Punctures fine and fairly numerous, the hairs which they bear long, thin, golden, erect or sub erect and of uniform length over the body, those on the legs shorter and stiffer; antennae without long hairs but clothed with a dense short pubescence. Base of the epinotum, area between the eye and the insertion of the antenna and the area between the ocelli stria to-granulate. Base of the petiolar nodes granulate. For the rest smooth and shining. Color yellowish brown to piceous brown, the antennae and legs pale yellow.

References based on Global Ant Biodiversity Informatics

 * Achury R., and A.V. Suarez. 2017. Richness and composition of ground-dwelling ants in tropical rainforest and surrounding landscapes in the Colombian Inter-Andean valley. Neotropical Entomology https://doi.org/10.1007/s13744-017-0565-4
 * Adams, R.M.M. and J.T. Longino. 2007. Nesting biology of the arboreal fungus-growing ant Cyphomyrmex cornutus and behavioral interactions with the social-parasitic ant Megalomyrmex mondabora. Insectes Sociaux 54:136-143
 * Addison D. S., I. Bartoszek, V. Booher, M. A. Deyrup, M. Schuman, J. Schmid, and K. Worley. 2016. Baseline surveys for ants (Hymenoptera: Formicidae) of the western Everglades, Collier County, Florida. Florida Entomologist 99(3): 389-394.
 * Ahuatzin D. A., E. J. Corro, A. Aguirre Jaimes, J. E. Valenzuela Gonzalez, R. Machado Feitosa, M. Cezar Ribeiro, J. Carlos Lopez Acosta, R. Coates, W. Dattilo. 2019. Forest cover drives leaf litter ant diversity in primary rainforest remnants within human-modified tropical landscapes. Biodiversity and Conservation 28(5): 1091-1107.
 * Alatorre-Bracamontes, C.E. and M Vasquez-Bolanos. 2010. Lista comentada de las hormigas (Hymenoptera: Formicidae) del norte de México. Dugesiana 17(1):9-36
 * Alayo D. P. 1974. Introduccion al estudio de los Himenopteros de Cuba. Superfamilia Formicoidea. Academia de Ciencias de Cuba. Instituto de Zoologia. Serie Biologica no.53: 58 pp. La Habana.
 * Amat-G G., M. G. Andrade-C. and F. Fernández. (eds.) 1999. Insectos de Colombia. Volumen II. Bogotá: Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 433 pp. 131975
 * André E. 1893. Description de quatre espèces nouvelles de fourmis d'Amérique. Rev. Entomol. (Caen) 12: 148-152.
 * Annotated Ant Species List Ordway-Swisher Biological Station. Downloaded at http://ordway-swisher.ufl.edu/species/os-hymenoptera.htm on 5th Oct 2010.
 * Armbrecht, I., and I. Perfecto. 2003. Litter-twig dwelling ant species richness and predation potential within a forest fragment and neighboring coffee plantations of contrasting habitat quality in Mexico. Agriculture, Ecosystems & Environment 97: 107-115.
 * Ascunce, M. S., S. M. Valles, D. H. Oi, D. Shoemaker, R. Plowes, L. Gilbert, and S. Sanchez-Pena. 2010. Molecular diversity of the microsporidium Kneallhazia solenopsae reveals an expanded host range among fire ants in North America. Journal of invertebrate Pathology 105: 279-288.
 * Barberena-Arias M. F., and T. M. Aide. 2003. Species Diversity and Trophic Composition of Litter Insects During Plant Secondary Succession. Caribbean Journal of Science 39(2): 161-169.
 * Basset Y., L. Cizek, P. Cuenoud, R. K. Didham, F. Guilhaumon, O. Missa, V. Novotny, F. Odegaards, T. Roslin, J. Schmidl et al. 2012. Arthropod diversity in a tropical forest. Science 338(6113): 1481-1484.
 * Boer P. 2019. Ants of Curacao, species list. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/SPECIES%20LIST%20CURACAO.html
 * Boer P. 2019. Ants of Saba, species list. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/SPECIES%20LIST%20SABA.html
 * Borgmeier T. 1934. Contribuição para o conhecimento da fauna mirmecológica dos cafezais de Paramaribo, Guiana Holandesa (Hym. Formicidae). Archivos do Instituto de Biologia Vegetal (Rio de Janeiro) 1: 93-111.
 * Braman C. A., and B. T. Forschler. 2018. Survey of Formicidae attracted to protein baits on Georgia’s Barrier Island dunes. Southeastern Naturalist 17(4): 645-653.
 * Brandao, C.R.F. 1991. Adendos ao catalogo abreviado das formigas da regiao neotropical (Hymenoptera: Formicidae). Rev. Bras. Entomol. 35: 319-412.
 * Buren W. F. 1982. Red imported fire ant now in Puerto Rico. Florida Entomologist 65: 188-189.
 * Cancino, E.R., D.R. Kasparan, J.M.A. Coronado Blanco, S.N. Myartseva, V.A. Trjapitzin, S.G. Hernandez Aguilar and J. Garcia Jimenez. 2010. Himenópteros de la Reserva El Cielo, Tamaulipas, México. Dugesiana 17(1):53-71
 * Castano-Meneses, G., M. Vasquez-Bolanos, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha, and I. Alcala-Martinez. 2015. Avances de Formicidae de Mexico. Universidad Nacional Autonoma de Mexico.
 * Castaño-Meneses G., P. Gonzalez Alvarado, Y. Granados Calixto, L. J. Lopez-Damian, and E. Beltran Sachez. 2010. Hormigas (Hymenoptera: Formicidae) presentes en el area de distribucion de lagartijas cornudas del genero Phrynosoma (Lacertilia: Phrynosomatidae) en el cerro del Tepetlayo, Eduardo Neri, Guerrero, Mexico. Entomologia Mexicana 9: 79-83.
 * Castelazo, C.D. and V. R.Gray, 1998. Numero y variacion estacional de asociaciones hormiga-planta en un bosque montano bajo de Veracruz, Mexico. Acta Zoologica Mexicana 73:44-55
 * Chacon de Ulloa P., G. I. Jaramillo, and M. M. Lozano. 2006. Hormigas urbanas en el departamento del Valle del Cauca, Colombia. Rev. Acad. Colomb. Cienc. 30 (116): 435-441.
 * Chanatasig-Vaca C. I., E. Huerta Lwanga, P. Rojas Fernandez, A. Ponce-Mendoza, J. Mendoza Vega, A. Moron Rios, H. Van Del Wal, and B. B. Dzib-Castillo. 2011. Effecto uso de suelo en las hormigas (Formicidae: Hymenoptera) de Tikinmul, Campeche, Mexico. Acta Zoológica Mexicana 27(2): 441-461.
 * Clouse R. 1999. Leaf-Litter Inhabitants of a Brazilian Pepper Stand in Everglades National Park. The Florida Entomologist. 82: 388-403
 * Coelho M. S., G. W. Fernandes, J. C. Santos, and J. H. C. Delabie. 2009. Ants (Hymenoptera: Formicidae) as bioindicators of land restoration in a Brazilian Atlantic forest fragment. Sociobiology 54(1): 51-63.
 * Cokendolpher J.C., Reddell J.R., Taylor S.J, Krejca J.K., Suarez A.V. and Pekins C.E. 2009. Further ants (Hymenoptera: Formicidae) from caves of Texas [Hormigas (Hymenoptera: Formicdae) adicionales de cuevas de Texas]. Texas Memorial Museum Speleological Monographs, 7. Studies on the cave and endogean fauna of North America, V. Pp. 151-168
 * Crawley W. C. 1916. Ants from British Guiana. Ann. Mag. Nat. Hist. 8(17): 366-378.
 * Creighton W. S. 1930. The New World species of the genus Solenopsis (Hymenop. Formicidae). Proceedings of the American Academy of Arts and Sciences 66: 39-151.
 * Culver D. C. 1974. Species packing in Caribbean and North Temperate ant communities. Ecology 55(5): 974-988.
 * Cupul-Magana F. G., and M. Vasquez-Bolanos. 2019. New records of ants (Hymenoptera: Formicidae) from María Cleofas Island, Gulf of California, Mexico. Dugesiana 26(1): 27-29.
 * Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
 * Del Toro, I., M. Vázquez, W.P. Mackay, P. Rojas and R. Zapata-Mata. Hormigas (Hymenoptera: Formicidae) de Tabasco: explorando la diversidad de la mirmecofauna en las selvas tropicales de baja altitud. Dugesiana 16(1):1-14.
 * Deyrup M., C. Johnson, G. C. Wheeler, J. Wheeler. 1989. A preliminary list of the ants of Florida. Florida Entomologist 72: 91-101
 * Deyrup M., L. Davis, and S. Buckner. 1998. Composition of the ant fauna of three Bahamian islands. Proceedings of the seventh symposium on the natural history of the Bahamas. 23-32. Bahamian Field Station, San Salvador, Bahamas
 * Deyrup, M. and J. Trager. 1986. Ants of the Archbold Biological Station, Highlands County, Florida (Hymenoptera: Formicidae). Florida Entomologist 69(1):206-228
 * Diaz-Castelazo, C., and V. Rico-Gray, 1998. Numero y variacion estacional de asociaciones hormiga-planta en un bosque montano bajo de Veracruz, Mexico. Acta Zoologica Mexicana 73: 45-55.
 * Dominguez D. F., M. Bustamante, R. Albuja, A. Castro, J. E. Lattke, and D. A. Donoso. 2016. Codigos de barras (COI barcodes) para hormigas (Hymenoptera: Formicidae) de los bosques secos del sur del Ecuador. Ecosistemas 25(2): 76-78.
 * Donoso D. A., G. Onore, G. Ramon, and J. E. Lattke. 2014. Invasive ants of continental Ecuador, a first account. REMCB 35: 133-141.
 * Emery C. 1890. Voyage de M. E. Simon au Venezuela (Décembre 1887 - Avril 1888). Formicides. Annales de la Société Entomologique de France (6)10: 55-76.
 * Emery C. 1896. Formiciden, gesammelt in Paraguay von Dr. J. Bohls. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 9: 625-638.
 * Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
 * Escalante J. A. 1975. Hormigas de la Provincia de la Convencionm Cusco. Revista Peruana de Entomologia 18:125-126.
 * Escalante J. A. 1976. Hormigas del valle de K'Osnipata (Paucartambo, Cusco). Revista Peruana de Entomologia 107-108.
 * Escalante J. A. 1979. Notas sobre las hormigas del Cusco. Rev. Per. Ent. 22(1): 111-112.
 * Feener D.H.,Jr. 1987. Response of Pheidole morrisi to two species of enemy ants, and a general model of defense behavior in Pheidole. Journal of the Kansas Entomological Society 60: 569-575.
 * Feitosa, R.M., R.R. Hora, J.H.C. Delabie, J. Valenzuela and D. Fresneau. 2008. A new social parasite in the ant genus Ectatomma F. Smith (Hymenoptera, Formicidae, Ectatomminae). Zootaxa 1713:47-52.
 * Fernandes I., and J. de Souza. 2018. Dataset of long-term monitoring of ground-dwelling ants (Hymenoptera: Formicidae) in the influence areas of a hydroelectric power plant on the Madeira River in the Amazon Basin. Biodiversity Data Journal 6: e24375.
 * Fernandes, P.R. XXXX. Los hormigas del suelo en Mexico: Diversidad, distribucion e importancia (Hymenoptera: Formicidae).
 * Fernandez Garcia I., and J. L. Fontenla. 2005. Nuevas adiciones a la entomofauna del Area Protegida Mil Cumbres, Pinar del Río, Cuba. Cocuyo 15: 20-22.
 * Fernandez Triana J. L., H. Sariol, M. A. Vega Castillo, S. Ricardo, M. Gonzalez, and E. P. Ferrer. 2002. Datos preliminares dobre la biodiversidad del orden Hymenoptera en la provincia Granma, Cuba. Bol. S.E.A. 31: 43-48.
 * Fernández Triana J. L., J. L. Fontenla, E. Portuondo Ferrer, and J. A. Genaro. 2005. Especies de himenópteros registrados en el Parque Nacional La Bayamesa, Cuba, 17-22 de junio del 2003 y 2-10 de febrero del 2004. In Maceira F., D., A.Fong G., W. S. Alverson, y/and T. Wachter, eds. 2005. Cuba: Parque Nacional La Bayamesa. Rapid Biological Inventories Report 13. The Field Museum, Chicago.
 * Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Flores, C. P., J. A. R. Garza, and A. E. C. Ramirez. 2004. Conocimiento Tlapaneco de hormigas (Hymenoptera: Formicidae) de Malinaltepec, Guerrero, Mexico. Etnobiologia 4: 19-28.
 * Flores-Maldonado K. Y., S. A. Phillips, and G. Sanchez-Ramos. 1999. The myrmecofauna (Hymenoptera: Formicidae) along an altitudinal gradient in the Sierra Madre Oriental of Northeastern Mexico. The Southwestern Naturalist 44(4): 457-461.
 * Flores-Maldonado, K. Y., S. A. Phillips-Jr, and G. Sanchez-Ramos. 1999. The myrmecofauna (Hymenoptera: Formicidae) along an altitudinal gradient in the Sierra Madre Oriental of Northeastern Mexico. The Southwestern Naturalist 44: 457-461.
 * Fontanla Rizo J.L. 1997. Lista preliminar de las hormigas de Cuba. Cocuyo 6: 18-21.
 * Fontenla J. L. 2005. Species of ants (Formicidae) recorded during the rapid biological inventory of the Zapata Peninsula, 8-15 September 2002. In: Kirkconnell P., A., D. F. Stotz, y / and J. M. Shopland, eds. 2005. Cuba: Península de Zapata. Rapid Biological Inventories Report 07. The Field Museum, Chicago
 * Fontenla J. L. 2005. Species of ants (Formicidae) recorded in the Sierra de Cubitas and adjacent areas, Camagüey Province, 16-19 September 2002. In: Díaz, L., M., W. S. Alverson, A. Barreto V., y / and T. Wachter. 2006. Cuba: Camagüey, Sierra de Cubitas. Rapid Biological Inventories Report 08. The Field Museum, Chicago
 * Fontenla J. L., and J. Alfonso-Simonetti. 2018. Classification of Cuban ants (Hymenoptera: Formicidae) into functional groups. Poeyana Revista Cubana de Zoologia 506: 21-30.
 * Fontenla Rizo J. L. 1993. Composición y estructura de comunidades de hormigas en un sistema de formaciones vegetales costeras. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 441: 1-19.
 * Fontenla Rizo J. L. 1993. Mirmecofauna de Isla de la Juventud y de algunos cayos del archipielago cubano. Poeyana. Instituto de Ecologia y Sistematica, Academia de Ciencias de Cuba 444:1-7.
 * Fontenla Rizo J. L. 1997. Lista preliminar de las hormigas de Cuba (Hymenoptera: Formicidae). Cocuyo 6: 18-21.
 * Fontenla Rizo J. L., and L. M. Hernández. 1993. Relaciones de coexistencia en comunidades de hormigas en un agroecosistema de caña de azúcar. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 438: 1-16.
 * Forel A. 1897. Quelques Formicides de l'Antille de Grenada récoltés par M. H. H. Smith. Transactions of the Entomological Society of London. 1897: 297-300.
 * Forel A. 1906. Fourmis néotropiques nouvelles ou peu connues. Annales de la Société Entomologique de Belgique 50: 225-249.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1908. Ameisen aus Sao Paulo (Brasilien), Paraguay etc. gesammelt von Prof. Herm. v. Ihering, Dr. Lutz, Dr. Fiebrig, etc. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 58: 340-418.
 * Forel A. 1908. Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles 44: 35-72.
 * Forel A. 1909. Ameisen aus Guatemala usw., Paraguay und Argentinien (Hym.). Deutsche Entomologische Zeitschrift 1909: 239-269.
 * Forel A. 1912. Formicides néotropiques. Part IV. 3me sous-famille Myrmicinae Lep. (suite). Mémoires de la Société Entomologique de Belgique. 20: 1-32.
 * Forster J.A. 2005. The Ants (hymenoptera: Formicidae) of Alabama. Master of Science, Auburn University. 242 pages.
 * Frago E., E. Portuondo Ferrer, J. L. Fernandez Triana, O. Sariego, and J. G. Fonseca. 2010. Entomofauna del Parque Nacional 'Desembarco del Granma", Cuba Suroriental. Boletin de la Sociedad Entomologica Aragonesa 46: 355-362.
 * Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
 * Galkowski C. 2016. New data on the ants from the Guadeloupe (Hymenoptera, Formicidae). Bull. Soc. Linn. Bordeaux 151, 44(1): 25-36.
 * Garcia M. A. The vulnerability of leaflitter ants to forest disturbances in the islands of Puerto Rico, Greater Antilles. Novitates Caribaea 13: 74-91.
 * Garcia-Martinez M. A., V. Vanoye-Eligio, O. R. Leyva-Ovalle, P. Zetina-Cordoba, M. J. Aguilar-Mendez, and M. Rosas-Mejia. 2019. Diversity of ants (Hymenoptera: Formicidae) in a sub-montane and sub-tropical cityscape of Northeastern Mexico. Sociobiology 66(3): 440-447.
 * General D., and L. Thompson. 2008. New distributional records of ants in Arkansas. Journal of the Arkansas Academy of Science 62: 148-150.
 * Gotzek, D., H. J. Axen, A. V. Suarez, S. Helms Cahan, and D. Shoemaker. 2015. Global invasion history of the tropical fire ant: a stowaway on the first global trade routes. Molecular Ecology 24: 374-388.
 * Gove, A. D., J. D. Majer, and V. Rico-Gray. 2009. Ant assemblages in isolated trees are more sensitive to species loss and replacement than their woodland counterparts. Basic and Applied Ecology 10: 187-195.
 * Groc S., J. H. C. Delabie, F. Fernandez, F. Petitclerc, B. Corbara, M. Leponce, R. Cereghino, and A. Dejean. 2017. Litter-dwelling ants as bioindicators to gauge the sustainability of small arboreal monocultures embedded in the Amazonian rainforest. Ecological Indicators 82: 43-49.
 * Guénard B., K. A. Mccaffrey, A. Lucky, and R. R. Dunn. 2012. Ants of North Carolina: an updated list (Hymenoptera: Formicidae). Zootaxa 3552: 1-36.
 * Hernandez, F. Varela and G. Castano-Meneses. 2010. Checklist, Biological Notes and Distribution of Ants (Hymenoptera: Formicidae) from Barranca de Metztitlán Biosphere Reserve, Hidalgo, Mexico. Sociobiology 56(2):397-434
 * Horvitz, C.C. and A.J. Beattie. 1980. Ant Dispersal of Calathea (Marantaceae) Seeds by Carnivorous Ponerines (Formicidae) in a Tropical Rain Forest. American Journal of Botany 67(3):321-326.
 * Horvitz, C.C. and D.W. Schemske. 1986. Seed Dispersal of a Neotropical Myrmecochore: Variation in Removal Rates and Dispersal Distance. Biotropica 18(4):319-323.
 * INBio Collection (via Gbif)
 * Ipser R. M. 2004. Native and exotic ants (Hymenoptera: Formicidae) of Georgia: Ecological Relationships with implications for development of biologically-based management strategies. Doctor of Philosophy thesis, University of Georgia. 165 pages.
 * Jaffe, K., Mauleon, H. and Kermarrec A. 1990. Predatory Ants of Diaprepes Abbreviatus (Coleoptera: Curculionidae) In Citrus Groves In Martinique and Guadeloupe, F.W.I. Florida Entomologist. 73(4):684-687.
 * Jaffe, Klaus and Lattke, John. 1994. Ant Fauna of the French and Venezuelan Islands in the Caribbean in Exotic Ants, editor D.F. Williams. 182-190.
 * Jeanne R. J. 1979. A latitudinal gradient in rates of ant predation. Ecology 60(6): 1211-1224.
 * Johnson C. 1986. A north Florida ant fauna (Hymenoptera: Formicidae). Insecta Mundi 1: 243-246
 * Johnson R. Personnal Database. Accessed on February 5th 2014 at http://www.asu.edu/clas/sirgtools/resources.htm
 * Kempf W. W. 1961. A survey of the ants of the soil fauna in Surinam (Hymenoptera: Formicidae). Studia Entomologica 4: 481-524.
 * Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
 * King J. R. 2007. Patterns of co-occurrence and body size overlap among ants in Florida's upland ecosystems. Ann. Zool. Fennici. 44: 189-201
 * Klotz, J.H., J.R. Mangold, K.M. Vail, L.R. Davis Jr., R.S. Patterson. 1995. A survey of the urban pest ants (Hymenoptera: Formicidae) of Peninsular Florida. Florida Entomologist 78(1):109-118
 * Koptur, S., M. Palacios-Rios, C. Diaz-Castelazo, W. P. Mackay, and V. Rico-Gray. 2013. Nectar secretion on fern fronds associated with lower levels of herbivore damage: field experiments with a widespread epiphyte of Mexican cloud forest remnants. Annals of Botany 111: 1277-1283.
 * Kusnezov N. 1963. Zoogeografia de las hormigas en sudamerica. Acta Zoologica Lilloana 19: 25-186
 * Larsen, A., and S. M. Philpott. 2010. Twig-nesting ants: the hidden predators of the coffee berry borer in Chiapas, Mexico. Biotropica 42: 342-347.
 * Lattke J. E., and M. Velez, and N. Aguirre. 2016. Survey of ants in dry forests of Southwestern Ecuador (Hymenoptera: Formicidae). Sociobiology 63(3): 909-918.
 * Longino J. T. 2013. Ants of Nicargua. Consulted on 18 Jan 2013. https://sites.google.com/site/longinollama/reports/ants-of-nicaragua
 * Longino J. T. L., and M. G. Branstetter. 2018. The truncated bell: an enigmatic but pervasive elevational diversity pattern in Middle American ants. Ecography 41: 1-12.
 * Longino J. et al. ADMAC project. Accessed on March 24th 2017 at https://sites.google.com/site/admacsite/
 * Lopes M. C., G. P. A. Lamarre, C. Baraloto, P. V. A. Fine, A. Vincentini, and F. B. Baccaro. 2019. The Amazonas-trap: a new method for sampling plant-inhabiting arthropod communities in tropical forest understory. Entomologia Experimentalis et Applicata https://doi.org/10.1111/eea.12797
 * Lopez, A. S., M. Vasquez-Bolanos, and G. A. Q. Rocha. 2015. Hormigas (Hymenoptera: Formicidae) del Cerro de la Culebra, Arandas, Jalisco, Mexico. Dugesiana 19: 151-155.
 * Lubertazzi D. and Tschinkel WR. 2003. Ant community change across a ground vegetation gradient in north Floridas longleaf pine flatwoods. 17pp. Journal of Insect Science. 3:21
 * Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
 * MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
 * Mackay, W.P., S. Porter, D. Gonzalez, A. Rodriguez, H. Armendedo, A. Rebeles and S.B. Vinson. 1990. A comparison of monogyne and polygyne populations of the tropical fire ant, Solenopsis geminata, (Hymenoptera: Formicidae) in Mexico, Journal of the Kansas Entomological Society 63(4):611-615.
 * Maes, J.-M. and W.P. MacKay. 1993. Catalogo de las hormigas (Hymenoptera: Formicidae) de Nicaragua. Revista Nicaraguense de Entomologia 23.
 * Mann W. M. 1916. The Stanford Expedition to Brazil, 1911, John C. Branner, Director. The ants of Brazil. Bulletin of the Museum of Comparative Zoology 60: 399-490
 * Mann W. M. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403-439.
 * Mann W. M. 1922. Ants from Honduras and Guatemala. Proceedings of the United States National Museum 61: 1-54.
 * McDonald D. L., D. R. Hoffpauir, and J. L. Cook. 2016. Survey yields seven new Texas county records and documents further spread of Red Imported Fire Ant, Solenopsis invicta Buren. Southwestern Entomologist, 41(4): 913-920.
 * Medina U. C. A., F. Fernandez, and M. G. Andrade-C. 2010. Insectos: escarabajos coprofagos, hormigas y mariposas. Capitulo 6. Pp 197-215. En: Lasso, C. A., J. S. Usma, F. Trujillo y A. Rial (eds.). 2010. Biodiversidad de la cuenca del Orinoco: bases científicas para la identificación de áreas prioritarias para la conservación y uso sostenible de la biodiversidad. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, WWF Colombia, Fundación Omacha, Fundación La Salle e Instituto de Estudios de la Orinoquia (Universidad Nacional de Colombia). Bogotá, D. C., Colombia.
 * Menozzi C, Russo G. 1930. Contributo alla conoscenza della mirmecofauna della Repubblica Dominicana (Antille). Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici. 24: 148-173.
 * Menozzi C. 1927. Formiche raccolte dal Sig. H. Schmidt nei dintorni di San José di Costa Rica. Entomologische Mitteilungen. Berlin-Dahlem. 16: 266-277.
 * Menozzi C. 1929. Formiche di Cuba e delle Isole Canarie raccolte dal Prof. Filippo Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura, Portici. 23: 1-5.
 * Mentone T. O., E. A. Diniz, C. B. Munhae, O. C. Bueno, and M. S. C. Morini. 2011. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop. 11(2): http://www.biotaneotropica.org.br/v11n2/en/abstract?inventory+bn00511022011.
 * Michigan State University, The Albert J. Cook Arthropod Research Collection. Accessed on January 7th 2014 at http://www.arc.ent.msu.edu:8080/collection/index.jsp
 * Mirmecofauna de la reserva ecologica de San Felipe Bacalar
 * Moody J. V., O. F. Francke, and F. W. Merickel. 1981. The distribution of fire ants, Solenopsis (Solenopsis) in Western Texas (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 54(3): 469-480.
 * Moody J. V., and O. F. Francke. 1982. The Ants (Hymenoptera, Formicidae) of Western Texas Part 1: Subfamily Myrmicinae. Graduate Studies Texas Tech University 27: 80 pp.
 * Moreau C. S., M. A. Deyrup, and L. R. David Jr. 2014. Ants of the Florida Keys: Species Accounts, Biogeography, and Conservation (Hymenoptera: Formicidae). J. Insect Sci. 14(295): DOI: 10.1093/jisesa/ieu157
 * Morrison L. W. 1998. A review of Bahamian ant (Hymenoptera: Formicidae) biogeography. Journal of Biogeography 25: 561-571.
 * Morrison, L.W. 2002. Long-Term Impacts of an Arthropod-Community Invasion by the Imported Fire Ant, Solenopsis invicta. Ecology 83(8):2337-2345
 * Nascimento Santos M., J. H. C. Delabie, and J. M. Queiroz. 2019. Biodiversity conservation in urban parks: a study of ground-dwelling ants (Hymenoptera: Formicidae) in Rio de Janeiro City. Urban Ecosystems https://doi.org/10.1007/s11252-019-00872-8
 * Navarro, E.V. Vergara, H. Echavarria Sanchez, F.J. Serna Cardona. 2007. Hormigas (Hymenoptera: Formicidae) asociadas al arboretum de la Universidad Nacional de Colombia, sede Medellin. Boletín Sociedad Entomológica Aragonesa 40:497-505.
 * Nestel, D., and F. Dickschen. 1990. The foraging kinetics of ground ant communities in different Mexican coffee agroecosystems. Oecologia 84: 58-63.
 * O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
 * Oliveira Mentone T. de, E. A. Diniz, C. de Bortoli Munhae, O. Correa Bueno and M. S. de Castro Morini. 2012. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop 11(2): 237-246.
 * Osorio-Perez K., M. F. Barberena-Arias, and T. M. Aide. 2007. Changes in Ant Species Richness and Composition During Plant Secondary Succession in Puerto Rico. Caribbean Journal of Science 43(2): 244-253.
 * Ottonetti L., L. Tucci, F. Frizzi, G. Chelazzi, and G. Santini. 2010. Changes in ground-foraging ant assemblages along a disturbance gradient in a tropical agricultural landscape. Ethology Ecology & Evolution 22: 7386.
 * Peck S. B. 1974. The Invertebrate Fauna of Tropical American Caves, Part II: Puerto Rico, An Ecological and Zoogeographic Analysis. Biotropica 6(1): 14-31.
 * Perez-Gelabert D. E. 2008. Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography. Zootaxa 1831:1-530.
 * Perez-Sanchez A. J., J. E. Lattke, and A. L. Viloria. 2012. Patterns of Ant (Hymenoptera: Formicidae) Richness and Relative Abundance along an Aridity Gradient in Western Venezuela. Neotrop Entomol DOI 10.1007/s13744-012-0096-y
 * Perfecto I., and J. Vandermeer. 1996. Microclimatic changes and the indirect loss of ant diversity in a tropical agroecosystem. Oecologia 108: 577-582.
 * Philpott S. M., I. Perfecto, and J. Vandermeer. 2006. Effects of management intensity and season on arboreal ant diversity and abundance in coffee agroecosystems. 15: 139-155.
 * Philpott, S.M., P. Bichier, R. Rice, and R. Greenberg. 2007. Field testing ecological and economic benefits of coffee certification programs. Conservation Biology 21: 975-985.
 * Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
 * Plowes R. M., J. G. Dunn, and L. E. Gilbert. 2007. The urban fire ant paradox: native fire ants persist in an urban refuge while invasive fire ants dominate natural habitats. Biological Invasions 9: 825-836.
 * Portuondo E. F., and J. L. Reyes. 2002. Mirmecofauna de los macizos montañosos de Sierra Maestra y Nipe-Sagua-Baracoa. Cocuyo 12: 10-13
 * Portuondo E. F., and J. L. Reyes. 2006. Species of Hymenoptera recorded in the Reserva Ecológica Pico Mogote, Santiago de Cuba Province, 20-25 September 2002. In: Maceira F., D., A. Fong G., y/and W. S. Alverson, eds. 2006. Cuba: Pico Mogote. Rapid Biological Inventories Report 09. The Field Museum, Chicago.
 * Portuondo E. F., and J. L. Reyes. 2006. Species of hymenopterans recorded in Siboney-Juticí Ecological Reserve, Santiago de Cuba Province, compiled from collections during the rapid inventory of 27-28 September 2002. Fong G., A., D. Maceira F., W. S. Alverson, y / and J. M. Shopland, eds. 2005. Cuba: Siboney-Juticí. Rapid Biological Inventories Report 10. The Field Museum, Chicago.
 * Portuondo Ferrer E., and J. L. Fernández Triana. 2005. Species of hymenopterans (bees, wasps, and ants) recorded in Alejandro de Humboldt National Park, from literature records, revision of the collection at BIOECO, and collections before and during the rapid inventory, 12-22 February 2004. In Fong G., A., D. Maceira F., W. S. Alverson, y/and T. Wachter, eds. 2005. Cuba: Parque Nacional Alejandro de Humboldt. Rapid Biological Inventories Report 14. The Field Museum, Chicago.
 * Portuondo Ferrer, E. and J. Fernandez Triana. Biodiversidad del orden Hymenoptera en Los Macizos Montanosos de Cuba Oriental. Boletin S.E.A. 35:121-136.
 * Quiroz Robledo L. N., and J. E. Valenzuela Gonzalez. 1993. Contribucion al conocimiento de la mirmecofauna del estado de Hidalgo, Mexico (Hymenoptera: Formicidae). En: Villavicencio-Nieto (ed) Flora y Fauna del Estado de Hidalgo. Universidad Autónoma de Hidalgo. p. 340-393. ISBN 968-63 40-36-X
 * Radoszkowsky O. 1884. Fourmis de Cayenne Française. Trudy Russkago Entomologicheskago Obshchestva 18: 30-39.
 * Reddell J. R., and J. C. Cokendolpher. 2001. Ants (Hymenoptera: Formicidae) from caves of Belize, Mexico, and California and Texas (U.S.A.) Texas. Texas Memorial Museum Speleological Monographs 5: 129-154.
 * Reyes, J. L. "Inventario de la colección de hormigas (Hymenoptera: Formicidae) del Centro Oriental de Ecosistemas y Biodiversidad, Santiago de Cuba, Cuba." Boletín de la Sociedad Aragonesa 36 (2005): 279-283.
 * Reynoso-Campos J. J., J. A. Rodriguez-Garza, and M. Vasquez-Bolanos. 2015. Hormigas (Hymenoptera: Formicidae) de la Isla Cozumel, Quintana Roo, Mexico (pp. 27-39). En: Castaño Meneses G., M. Vásquez-Bolaños, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha e I. Alcalá-Martínez (Coords.). Avances de Formicidae de  México.  UNAM,  Universiad  de  Guadalajara, Guadalajara, Jalisco.
 * Rico-Gray, V. 1993. Use of plant-derived food resources by ants in the dry tropical lowlands of coastal Veracruz, Mexico. Biotropica 25(3):301-315.
 * Rico-Gray,V., J.G. Garcia-Franco, M. Palacios-Rios, C. Diaz-Castelazo, V. Parra-Tabla and J.A. Navarro. 1998. Geographical and Seasonal Variation in the Richness of Ant-Plant Interactions in Mexico. Biotropica 30(2):190-200.
 * Rios-Casanova, L., A. Valiente-Banuet, and V. Rico-Gray. (2004). Las hormigas del Valle de Tehuacan (Hymenoptera: Formicidae): una comparacion con otras zonas aridas de Mexico. Acta Zoologica Mexicana 20: 37-54.
 * Rojas Fernandez P. 2010. Capítulo 24. Hormigas (Insecta: Hymenoptera: Formicidae). In: Diversidad Biológica de Veracruz. Volumen Invertebrados. CONABIO-Gobierno del Estado de Veracruz.
 * Rojas M. L. 2012. Fauna de insectos en cayos del golfo de Ana María, Cuba. Rev. Invest. Mar. 32(2): 66-72.
 * Rojas, J.C., J. Valle Mora, A. Virgen, Y. Brindis and L. Cruz Lopez. 2004. Variacion espacial y temporal del perfil de hidrocarburos cuticulares de colonias de Solenopsis geminata (F.) (Hymenoptera: Formicidae) de Soconusco, Chiapas, Mexico. Folia Entomol. Mex. 43(3): 271-278.
 * Saarinen, E.V. and J.C. Daniels.2006. Miami blue butterfly larvae (Lepidoptera: Lycaenidae) and ants (Hymenoptera: Formicidae): New information on the symbionts of an endangered taxon. Florida Entomologist 89(1): 69-74
 * Salcido-Lopez A., M. Vásquez-Bolaños, and G.A. Quiroz-Rocha. 2012. Hormigas (Hymenoptera: Formicidae) del Cerro de la Culebra, Arandas, Jalisco, Mexico. Dugesiana 19(2): 151-155.
 * Salinas P. J. 2010. Catalogue of the ants of the Táchira State, Venezuela, with notes on their biodiversity, biogeography and ecology (Hymenoptera: Formicidae: Amblyioponinae, Ponerinae, Proceratiinae, Myrmicinae, Ecitoninae, Formicinae, Pseudomyrmecinae, Dolichoderinae). Boletín de la SEA 47: 315-328.
 * Sanchez-Pena, S.R., M.C. Chacon-Cardosa and D. Resendez-Perez. 2009. Identification of fire ants (Hymenoptera: Formicidae) from Northeastern Mexico with morphology and molecular markers. Florida Entomologist 92(1):107-115.
 * Santos P. P., A. Vasconcelos, B. Jahyny, and J. H. C. Delabie. 2010. Ant fauna (Hymenoptera, Formicidae) associated to arboreal nests of Nasutitermes spp. (Isoptera, Termitidae) in a cacao plantation in southeastern Bahia, Brazil. Revista Brasileira de Entomologia 54(3): 450-454.
 * Santschi F. 1923. Solenopsis et autres fourmis néotropicales. Revue Suisse de Zoologie 30: 245-273.
 * Santschi F. 1925. Nouveaux Formicides brésiliens et autres. Bulletin et Annales de la Société Entomologique de Belgique 65: 221-247.
 * Santschi F. 1925. Nouvelles fourmis brésiliennes. Annales de la Société Entomologique de Belgique. 64: 5-20.
 * Santschi F. 1930. Quelques fourmis de Cuba et du Brésil. Bulletin. Société Entomologique d'Egypte. 14: 75-83.
 * Santschi F. 1931. Fourmis de Cuba et de Panama. Revista de Entomologia (Rio de Janeiro). 1: 265-282.
 * Santschi F. 1939. Résultats scientifiques des croisières du navire-école belge, "Mercator". XIV. Formicidae s. lt. Mémoires du Musée Royal d'Histoire Naturelle de Belgique. (2)15: 159-167.
 * Smith F. 1941. A list of the ants of Washington State. The Pan-Pacific Entomologist 17(1): 23-28.
 * Smith M. R. 1924. An annotated list of the ants of Mississippi (Hym.) (continued from page 54). Entomological News 35: 77-85.
 * Smith M. R. 1930. A list of Florida ants. Florida Entomologist 14: 1-6.
 * Smith M. R. 1934. A list of the ants of South Carolina. Journal of the New York Entomological Society 42: 353-361.
 * Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
 * Smith M. R. 1937. The ants of Puerto Rico. Journal of Agriculture of the University of Puerto Rico 20: 819-875.
 * Smith M. R. 1942. The relationship of ants and other organisms to certain scale insects on coffee in Puerto Rico. Journal of Agriculture of the University of Puerto Rico 26: 21-27.
 * Smith M. R. 1965. House-infesting ants of the eastern United States. Their recognition, biology, and economic importance. United States Department of Agriculture. Technical Bulletin 1326: 1-105.
 * Smith M. R., and W. A. Morrison. 1916. South Carolina ants. Entomological News 27: 110-111.
 * Snelling R. 1993. Ants of Guana Island, British Virgin Islands. Notes From Underground 8: 11-12.
 * Snelling R. R. 2005. Wasps, ants, and bees: aculeate Hymeoptera. Pp. 283-296 in: Lazell, J. 2005. Island. Fact and theory in nature. Berkeley: University of California Press, xx + 382 pp.
 * Solomon, S.E. and A.S. Mikheyev. 2005. The ant (Hymenoptera: Formicidae) fauna of Cocos Island, Costa Rica. Florida Entomologist 88(4):415-423
 * Soubeyran Y. 2008. Especes exotiques envahissantes dans les collectivites francaises d'Outre-mer: Etat des lieux et recommandations. Collection Planète Nature. Comité français de l’UICN, Paris, France.
 * Torres J.A. 1984. Niches and Coexistence of Ant Communities in Puerto Rico: Repeated Patterns. Biotropica 16(4): 284-295.
 * Torres, I. L., M. A. G. Martinez, H. O. Rivera, M. E. G. Tovar, H. E. L. Espinosa, and J. M. Gonzalez. 2018. Comparacion de dos muestreos de hormigas del suelo en la barranca de Metlac, Fortin de las Flores, Veracruz, Mexico. Revista Cientifica UDO Agricola 10: 173-178.
 * Torres, Juan A. and Roy R. Snelling. 1997. Biogeography of Puerto Rican ants: a non-equilibrium case?. Biodiversity and Conservation 6:1103-1121.
 * Trager J. C. 1991. A revision of the fire ants, Solenopsis geminata group (Hymenoptera: Formicidae: Myrmicinae). Journal of the New York Entomological Society 99: 141-198
 * Trager, J. and C.Johnson. 1985. A slave-making ant in Florida: Polyergus lucidus with observations on the natural history of its host Formica archboldi (Hymenoptera: Formicidae). The Florida Entomologist 68(2):261-266.
 * Ulyssea M. A., and C. R. F. Brandao. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217224.
 * Ulysséa M. A., C. R. F. Brandão. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217-224.
 * Valdes-Rodriguez S., P. Chacon de Ulloa, and I. Armbrecht. 2014. Soil ant species in Gorgona Island, Colombian Pacific. Rev. Biol. Trop. 62 (1): 265-276.
 * Van Pelt A. F. 1948. A Preliminary Key to the Worker Ants of Alachua County, Florida. The Florida Entomologist 30(4): 57-67
 * Van Pelt A. F. 1956. The ecology of the ants of the Welaka Reserve, Florida (Hymenoptera: Formicidae). American Midland Naturalist 56: 358-387
 * Van Pelt A. F. 1966. Activity and density of old-field ants of the Savannah River Plant, South Carolina. Journal of the Elisha Mitchell Scientific Society 82: 35-43.
 * Varela-Hernandez, F., M. Rocha-Ortega, R. W. Jones, and W. P. Mackay. 2016. Insectos: Hormigas (Formicidae) del estado de Queretaro, Mexico. Pages 397-404 in W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
 * Varela-Hernandez, F., M. Rocha-Ortega, W. P. Mackay, and R. W. Jones. 2016. Lista preliminar de las hormigas (Hymenoptera: Formicidae) del estado de Queretaro, Mexico. Pages 429-435 in . W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
 * Vargas R. P. 1964. Clave para identificar los Formicidae de la provincia de Chiclayo. Revista Peruana de Entomologia 7(1): 98-102.
 * Vasconcelos H. L., J. M. S. Vilhena, W. E. Magnusson, and A. L. K. M. Albernaz. 2006. Long-term effects of forest fragmentation on Amazonian ant communities. Journal of Biogeography 33: 1348-1356.
 * Vasconcelos, H.L., J.M.S. Vilhena, W.E. Magnusson and A.L.K.M. Albernaz. 2006. Long-term effects of forest fragmentation on Amazonian ant communities. Journal of Biogeography 33:1348-1356
 * Vasquez-Bolanos, M. 2007. Una especie nueva del genero Tetramorium Mayr (Hymenoptera: Formicidae) de Mascota, Jalisco, Mexico. Dugesiana 14(2):93-97.
 * Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
 * Vázquez Moreno L. L., Y. M. Brito, J. A. Simonetti, D. Moreno Rodriguez, and A. A. Nunez. 2009. Diversidad de especies de hormigas (Hymenoptera: Formicidae) en cafetales afectados por Hypothenemus hampei Ferrari (Coleoptera: Curculionidae: Scolytinae). Fitosanidad 13(3): 163-168.
 * Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
 * Wauters N., W. Dekoninck, and D. Fournier. 2018. Introduction history and genetic diversity of the invasive ant Solenopsis geminata in the Galapagos Islands. Biological Invasions https://doi.org/10.1007/s10530-018-1769-1
 * Weber N. A. 1938. The food of the giant toad, Bufo marinus (L.), in Trinidad and British Guiana with special reference to the ants. Annals of the Entomological Society of America 31: 499-503.
 * Weber N. A. 1948. Studies on the fauna of Curaçao, Aruba, Bonaire and the Venezuelan islands: No. 14. Ants from the Leeward Group and some other Caribbean localities. Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen 5: 78-86.
 * Wetterer J. K. 2011. Worldwide Spread of the tropical fire ant, Solenopsis geminata (Hymenoptera: Formicidae). Myrmecological News 14: 21-35.
 * Wetterer J. K., and C. D. Lombard. 2010. Fire Ants (Hymenoptera: Formicidae) along an Important Sea Turtle Nesting Beach on St. Croix, USVI. Florida Entomologist 93(3): 449-450.
 * Wetterer J. K., M. J. Liles, J. M. Sermeno, L. Serrano Cervantes, E. E. Echeverria, R. M. Estrada Hernandez, A. Henriquez, D. Perez, D. A. Sanchez Garcia, C. E. Gomez Peralta, R. Lopez Sorto, and G. Melendez. 2016. Predaceous fie ants (Hymenoptera: Formicidae) at sea turtle (Testudines: Cheloniidae) nestig beaches and hatcheries in El Salvador. Florida Entomologist 99(1): 106-109.
 * Wetterer J.K. and J.L.W. Keularts. 2008. Population explosion of the hairy crazy ant, Paratrechina pubens (Hymenoptera: Formicidae), on St. Croix, US Virgin Islands. Florida Entomologist 91(3): 423-427.
 * Wetterer, J.K. and J.L.W. Keularts. 2008. Population Explosion of the Hairy Crazy Ant, Paratrechina pubens (Hymenoptera: Formicidae), on St. Croix, US Virgin Islands. The Florida Entomologist 91(3):423-427
 * Wetterer, James K. and O'Hara, Brandon C. 2002. Ants (Hymenoptera: Formicidae) of the Dry Tortugas, the Outermost Florida Keys. Florida Entomoloist 85(2):303-307.
 * Wheeler W. M. 1905. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21: 79-135.
 * Wheeler W. M. 1907. A collection of ants from British Honduras. Bulletin of the American Museum of Natural History 23: 271-277.
 * Wheeler W. M. 1908. The ants of Jamaica. Bulletin of the American Museum of Natural History 24: 159-163.
 * Wheeler W. M. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117-158.
 * Wheeler W. M. 1908. The ants of Texas, New Mexico and Arizona. (Part I.). Bulletin of the American Museum of Natural History 24: 399-485.
 * Wheeler W. M. 1913. Ants collected in Georgia by Dr. J. C. Bradley and Mr. W. T. Davis. Psyche (Cambridge) 20: 112-117.
 * Wheeler W. M. 1913. The ants of Cuba. Bulletin of the Museum of Comparative Zoology 54: 477-505.
 * Wheeler W. M. 1916. Ants collected in British Guiana by the expedition of the American Museum of Natural History during 1911. Bulletin of the American Museum of Natural History 35: 1-14.
 * Wheeler W. M. 1917. Jamaican ants collected by Prof. C. T. Brues. Bulletin of the Museum of Comparative Zoology 61: 457-471.
 * Wheeler W. M. 1918. Ants collected in British Guiana by Mr. C. William Beebe. Journal of the New York Entomological Society 26: 23-28.
 * Wheeler W. M. 1919. The ants of Tobago Island. Psyche (Cambridge) 26: 113.
 * Wheeler W. M. 1922. The ants of Trinidad. American Museum Novitates 45: 1-16.
 * Wheeler W. M. 1932. A list of the ants of Florida with descriptions of new forms. J. N. Y. Entomol. Soc. 40: 1-17.
 * Wheeler W. M. 1937. Ants mostly from the mountains of Cuba. Bulletin of the Museum of Comparative Zoology. 81: 439-465.
 * Wheeler W. M., and W. M. Mann. 1914. The ants of Haiti. Bulletin of the American Museum of Natural History 33: 1-61.
 * Wheeler, William Morton. 1911. Additions to the Ant-Fauna of Jamaica. Bulletin American Museum of Natural History. 30:21-29.
 * Wheeler, William Morton. 1916. Ants Collected in Trinidad by Professor Roland Thaxter, Mr. F. W. Urich, and Others. Bulletin of the Museum of Comparitive Zoology at Harvard University. 40(8):322-330
 * Wheeler, William Morton. 1923. Report on the Ants. The University of Iowa Studies in Natural History. 10(3):3-9.
 * Wheeler, William Morton. 1933. Formicidae of the Templeton Crocker Expedition. California Academy of Sciences. 21(6):57-64.
 * Wheeler, William Morton. 1934. Some Ants From The Bahama Islands. Psyche. 41(4):230-232.
 * Wheeler, William Morton. 1936. Ants From Hispaniola and Mona Island. Bulletin: Museum of Comparative Zoology at Harvard College. 80(2):192-211.
 * Whitcomb W. H., H. A. Denmark, A. P. Bhatkar, and G. L. Greene. 1972. Preliminary studies on the ants of Florida soybean fields. Florida Entomologist 55: 129-142.
 * Wilson E. O. 1962. Behavior of Daceton armigerum (Latreille), with a classification of self-grooming movements in ants. Bulletin of the Museum of Comparative Zoology 127: 403-421.
 * Zolessi L. C. de, Y. P. Abenante, and M. E. de Philippi. 1988. Lista sistematica de las especies de Formicidos del Uruguay. Comun. Zool. Mus. Hist. Nat. Montev. 11: 1-9.
 * Zolessi L. C. de; Y. P. de Abenante, and M. E. Philippi. 1989. Catálogo sistemático de las especies de Formícidos del Uruguay (Hymenoptera: Formicidae). Montevideo: ORCYT Unesco, 40 + ix pp.
 * de Zolessi, L.C., Y.P. de Abenante and M.E. Phillipi. 1989. Catalago Systematico de las Especies de Formicidos del Uruguay (Hymenoptera: Formicidae). Oficina Regional de Ciencia y Technologia de la Unesco para America Latina y el Caribe- ORCYT. Montevideo, Uruguay
 * do Nascimento, I.C. 2006. Fenologia dos Voos de Acasalamento em Formigas Tropicais