Spain



Gary D. Alpert: Museum of Comparative Zoology, Harvard University

The following valid species and subspecies of ants are known to occur in Spain. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Spanish ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Spanish Ants web page. Please advise if any corrections are warranted.

A

 * Amblyopone emeryi Saunders, 1890


 * Amblyopone gaetulica Baroni Urbani, 1978


 * Amblyopone impressifrons (Emery, 1869)


 * Anergates atratulus 	(Schenck, 1852)


 * Anochetus ghilanii (Spinola, 1853)


 * Aphaenogaster cardenai 	Espadaler, 1981


 * Aphaenogaster dulciniae 	Emery, 1924


 * Aphaenogaster gemella 	Roger, 1862


 * Aphaenogaster gibbosa (Latreille, 1798)


 * Aphaenogaster iberica 	Emery, 1908


 * Aphaenogaster senilis Mayr, 1853


 * Aphaenogaster senilis grata 	Santschi, 1933


 * Aphaenogaster splendida (Roger, 1859)


 * Aphaenogaster striativentris Forel, 1895


 * Aphaenogaster subterranea (Latreille, 1798)


 * Aphaenogaster striativentris 	Forel, 1895

B

 * Bothriomyrmex gibbus Soudek, 1925


 * Bothriomyrmex hispanicus 	Santschi, 1922


 * Bothriomyrmex meridionalis (Roger, 1863)


 * Bothriomyrmex saundersi 	Santschi, 1922

C

 * Camponotus aethiops (Latreille, 1798)


 * Camponotus amaurus 	Espadaler, 1997


 * Camponotus barbaricus Emery, 1905


 * Camponotus catalanus 	Emery, 1924


 * Camponotus cruentatus (Latreille, 1802)


 * Camponotus fallax (Nylander, 1856)


 * Camponotus figaro Collingwood & Yarrow, 1969


 * Camponotus foreli Emery, 1881


 * Camponotus gestroi Emery, 1878


 * Camponotus haroi 	Espadaler, 1997


 * Camponotus herculeanus (Linnaeus, 1758)


 * Camponotus lateralis (Olivier, 1792)


 * Camponotus ligniperdus (Latreille, 1802)


 * Camponotus micans (Nylander, 1856)


 * Camponotus piceus (Leach, 1825)


 * Camponotus pilicornis (Roger, 1859)


 * Camponotus ruber Emery, 1925


 * Camponotus sylvaticus (Olivier, 1792)


 * Camponotus truncatus (Spinola, 1808)


 * Camponotus universitatis Forel, 1890


 * Camponotus vagus (Scopoli, 1763)


 * Cardiocondyla batesii Forel, 1894


 * Cardiocondyla elegans 	Emery, 1869


 * Cardiocondyla emeryi Forel, 1881


 * Cardiocondyla mauritanica 	Forel, 1890


 * Cardiocondyla nigra Forel, 1905


 * Cataglyphis douwesi 	de Haro & Collingwood, 2000


 * Cataglyphis floricola 	Tinaut, 1993


 * Cataglyphis gadeai 	De Haro & Collingwood, 2003


 * Cataglyphis hispanicus 	Emery, 1906


 * Cataglyphis humeya 	Tinaut, 1991


 * Cataglyphis ibericus 	Emery, 1906


 * Cataglyphis piliscapus (Forel, 1901)


 * Cataglyphis rosenhaueri 	Santschi, 1925


 * Cataglyphis velox 	Santschi, 1929


 * Cataglyphis viaticus 	Fabricius, 1787


 * Chalepoxenus kutteri Cagniant, 1973


 * Chalepoxenus muellerianus (Finzi, 1922)


 * Crematogaster auberti 	Emery, 1869


 * Crematogaster fuentei 	Menozzi, 1922


 * Crematogaster inermis Mayr, 1862


 * Crematogaster laestrygon Emery, 1869


 * Crematogaster scutellaris (Olivier, 1792)


 * Crematogaster sordidula (Nylander, 1849)


 * Cryptopone ochracea (Mayr, 1855)

D

 * Dolichoderus quadripunctatus (Linnaeus, 1771)

F

 * Formica bruni Kutter, 1967


 * Formica cinerea Mayr, 1853


 * Formica cunicularia Latreille, 1798


 * Formica decipiens Bondroit, 1918


 * Formica dusmeti 	Emery, 1909


 * Formica exsecta Nylander, 1846


 * Formica foreli Bondroit, 1918


 * Formica frontalis 	Santschi, 1919


 * Formica fusca Linnaeus, 1783


 * Formica gagates Latreille, 1798


 * Formica gerardi Bondroit, 1917


 * Formica lemani Bondroit, 1917


 * Formica lugubris Zetterstedt, 1838


 * Formica paralugubris Seifert, 1996


 * Formica picea Nylander, 1846


 * Formica polyctena 	Foerster, 1850


 * Formica pratensis 	Retzius, 1783


 * Formica pressilabris Nylander, 1846


 * Formica rufa Linnaeus, 1758


 * Formica rufa obscurata 	Santschi, 1925


 * Formica rufibarbis Fabricius, 1793


 * Formica sanguinea Latreille, 1798


 * Formica selysi Bondroit, 1918


 * Formica subrufa Roger, 1859


 * Formicoxenus nitidulus (Nylander, 1846)

G

 * Goniomma baeticum 	Reyes, Espadaler & Rodriquez, 1987


 * Goniomma blanci (Andre, 1881)


 * Goniomma collingwoodi 	Espadaler, 1997


 * Goniomma compressisquama 	Tinaut, 1995


 * Goniomma decipiens 	Espadaler, 1997


 * Goniomma hispanicum 	André, 1883


 * Goniomma kugleri 	Espadaler, 1986

H

 * Harpagoxenus sublaevis (Nylander, 1849)


 * Hypoponera abaillei (Andre, 1881)


 * Hypoponera eduardi (Forel, 1894)


 * Hypoponera punctatissima (Roger, 1859)


 * Hypoponera ragusai (Emery, 1894)


 * Hypoponera ragusai santschii (Emery, 1909)

L

 * Lasius alienus (Foerster, 1850)


 * Lasius bicornis (Foerster, 1850)


 * Lasius brunneus (Latreille, 1798)


 * Lasius carniolicus Mayr, 1861


 * Lasius cinereus Seifert, 1992


 * Lasius citrinus Emery, 1922


 * Lasius distinguendus (Emery, 1916)


 * Lasius emarginatus (Olivier, 1792)


 * Lasius flavus (Fabricius, 1782)


 * Lasius fuliginosus (Latreille, 1798)


 * Lasius grandis Forel, 1909


 * Lasius jensi Seifert, 1982


 * Lasius lasioides (Emery, 1869)


 * Lasius meridionalis (Bondroit, 1920)


 * Lasius mixtus (Nylander, 1846)


 * Lasius myops Forel, 1894


 * Lasius neglectus Van Loon, Boomsma & Andrasfalvy, 1990


 * Lasius niger (Linnaeus, 1758)


 * Lasius paralienus Seifert, 1992


 * Lasius piliferus 	Seifert, 1992


 * Lasius platythorax 	Seifert, 1991


 * Lasius psammophilus Seifert, 1992


 * Lasius rabaudi (Bondroit, 1917)


 * Lasius sabularum (Bondroit, 1918)


 * Lasius umbratus (Nylander, 1846)


 * Lepisiota grauenfeldi (Mayr, 1855)


 * Lepisiota nigra (Dalla Torre, 1893)


 * Leptanilla charonea 	López, Martínez & Barandica, 1994


 * Leptanilla ortunoi 	López, Martínez & Barandica, 1994


 * Leptanilla plutonia 	López, Martínez & Barandica, 1994


 * Leptanilla reveilerii Emery, 1870


 * Leptanilla zaballosi 	López, Martínez & Barandica, 1994


 * Leptothorax acervorum (Fabricius, 1793)


 * Leptothorax gredleri Mayr, 1855


 * Leptothorax muscorum (Nylander, 1846)


 * Linepithema humile (Mayr, 1868)

M

 * Messor barbarus 	Linnaeus, 1767


 * Messor bouvieri Bondroit, 1918


 * Messor capitatus (Latreille, 1798)


 * Messor celiae 	Reyes, 1985


 * Messor hispanicus 	Santschi, 1919


 * Messor ibericus 	Santschi, 1931


 * Messor lobicornis Forel, 1894


 * Messor lusitanicus Tinaut, 1985


 * Messor marocanus 	Santschi, 1927


 * Messor structor (Latreille, 1798)


 * Messor timidus 	Espadaler, 1997


 * Monomorium algiricum (Bernard, 1955)


 * Monomorium andrei Saunders, 1890


 * Monomorium andrei furunculum 	Emery, 1924


 * Monomorium carbonarium (Smith, F., 1858)


 * Monomorium destructor (Jerdon, 1851)


 * Monomorium mayri Forel, 1902


 * Monomorium monomorium Bolton, 1987


 * Monomorium pharaonis (Linnaeus, 1758)


 * Monomorium salomonis (Linnaeus, 1758)


 * Monomorium sichelii (Roger, 1862)


 * Monomorium subopacum 	Smith, F., 1858


 * Myrmecina graminicola (Latreille, 1802)


 * Myrmica aloba 	Forel, 1909


 * Myrmica bibikoffi Kutter, 1963


 * Myrmica curvithorax 	Bondroit, 1920


 * Myrmica karavajevi 	Arnol’di, 1930


 * Myrmica lemasnei 	Bernard, 1967


 * Myrmica lobulicornis Nylander, 1857


 * Myrmica rubra (Linnaeus, 17830


 * Myrmica ruginodis Nylander, 1846


 * Myrmica sabuleti Meinert, 1861


 * Myrmica scabrinodis Nylander, 1846


 * Myrmica schencki Viereck, 1903


 * Myrmica specioides Bondroit, 1918


 * Myrmica spinosior Santschi, 1931


 * Myrmica sulcinodis Nylander, 1846


 * Myrmica vandeli Bondroit, 1920


 * Myrmica wesmaeli Bondroit, 1918


 * Myrmica xavieri Radchenko, Elmes, & Savolainen, 2008


 * Myrmoxenus bernardi 	Espadaler, 1982


 * Myrmoxenus kraussei (Emery, 1915)


 * Myrmoxenus ravouxi (Andre, 1896)

N

 * Nylanderia jaegerskioeldi (Mayr, 1904)


 * Nylanderia vividula (Nylander, 1846)

O

 * Oxyopomyrmex santschii Forel, 1904


 * Oxyopomyrmex saulcyi 	Emery, 1889

P

 * Paratrechina longicornis (Latreille, 1802)


 * Pheidole pallidula (Nlyander, 1849)


 * Pheidole teneriffana Forel, 1893


 * Plagiolepis grassei LeMasne, 1956


 * [Plagiolepis pygmaea (Latreille, 1798)


 * Plagiolepis schmitzi Forel, 1895


 * Plagiolepis schmitzi pyrenaica Emery, 1921


 * Plagiolepis xene Starcke, 1936


 * Polyergus rufescens (Latreille, 1798)


 * Ponera coarctata 	Latreille, 1802


 * Ponera testacea 	Emery, 1895


 * Proceratium melinum 	(Roger, 1860)


 * Proformica ferreri 	Bondroit, 1918


 * Proformica longiseta 	Collingwood, 1978


 * Proformica nasuta (Nylander, 1856)


 * Pyramica argiola (Emery, 1869)


 * Pyramica baudueri 	Emery, 1875


 * Pyramica membranifera (Emery, 1869)


 * Pyramica tenuipilis (Emery, 1915)

R

 * Rossomyrmex minuchae 	Tinaut, 1981

S

 * Solenopsis fairchildi Wheeler, W.M., 1926


 * Solenopsis fugax (Latreille, 1798)


 * Solenopsis latro Forel, 1894


 * Solenopsis lusitanica Emery, 1915


 * Solenopsis monticola Bernard, 1950


 * Solenopsis nicaeensis Bernard, 1950


 * Solenopsis oraniensis Forel, 1894


 * Solenopsis orbula emery, 1875


 * Solenopsis robusta Bernard, 1950


 * Stenamma debile (Foerster, 1850)


 * Stenamma petiolatum Emery, 1897


 * Stenamma sardoum Emery, 1915


 * Stenamma striatulum Emery, 1895


 * Stenamma westwoodi Westwood, 1839


 * Strongylognathus caeciliae Forel, 1897


 * Strongylognathus huberi Forel, 1874


 * Strongylognathus testaceus (Schenck, 1852)

T

 * Tapinoma erraticum 	(Latreille, 1798)


 * Tapinoma madeirense Forel, 1895


 * Tapinoma melanocephalum (Fabricius, 1793)


 * Tapinoma nigerrimum (Nylander, 1856)


 * Tapinoma pygmaeum (Dufour, 1857)


 * Tapinoma simrothi Krausse-Heldrungen, 1911


 * Teleutomyrmex kutteri 	Tinaut, 1990


 * Teleutomyrmex schneideri Kutter, 1950


 * Temnothorax affinis (Mayr, 1855)


 * Temnothorax albipennis (Curtis, 1854)


 * Temnothorax angustulus (Nylander, 1856)


 * Temnothorax atlantis (Santschi, 1911)


 * Temnothorax baeticus 	(Emery, 1924)


 * Temnothorax berlandi (Bondroit, 1918)


 * Temnothorax blascoi 	Espadaler, 1997


 * Temnothorax caesari 	Espadaler, 1997


 * Temnothorax cagnianti 	Tinaut, 1983


 * Temnothorax clypeatus (Mayr, 1853)


 * Temnothorax corticalis (Schenck, 1852)


 * Temnothorax crepuscularis 	Tinaut, 1995


 * Temnothorax cristinae 	Espadaler, 1997


 * Temnothorax curtulus (Santschi, 1929)


 * Temnothorax flavispinus (Andre, 1881)


 * Temnothorax formosus (Santschi, 1909)


 * Temnothorax fuentei 	Santschi, 1919


 * Temnothorax gredosi 	Espadaler & Collingwood, 1982


 * Temnothorax grouvellei (Bondroit, 1918)


 * Temnothorax ibericus 	Menozzi, 1922


 * Temnothorax interruptus (Schenck, 1852)


 * Temnothorax kraussei (Emery, 1916)


 * Temnothorax leviceps (Emery, 1898)


 * Temnothorax lichtensteini (Bondroit, 1918)


 * Temnothorax longispinosus (Roger, 1863)


 * Temnothorax luteus (Forel, 1874)


 * Temnothorax massiliensis (Bondroit, 1918)


 * Temnothorax nadigi 	Kutter, 1925


 * Temnothorax naeviventris (Santschi, 1910)


 * Temnothorax niger 	Forel, 1894


 * Temnothorax nigriceps (Mayr,, 1855)


 * Temnothorax nylanderi (Foerster, 1850)


 * Temnothorax pardoi 	Tinaut, 1987


 * Temnothorax parvulus (Schenck, 1852)


 * Temnothorax platycephalus (Espadaler, 1997)


 * Temnothorax rabaudi (Bondroit, 1918)


 * Temnothorax racovitzai (Bondroit, 1918)


 * Temnothorax recedens 	(Nylander, 1856)


 * Temnothorax schaufussi (Forel, 1879)


 * Temnothorax specularis (Emery, 1916)


 * Temnothorax subcingulatus 	(Emery, 1924)


 * Temnothorax tristis (Bondroit, 1918)


 * Temnothorax tuberum (Fabricius, 175)


 * Temnothorax tyndalei 	(Forel, 1909)


 * Temnothorax unifasciatus (Latreille, 1798)


 * Temnothorax universitatis 	(Espadaler, 1997)


 * Tetramorium bicarinatum (Nylander,, 1846)


 * Tetramorium caespitum (Linnaeus, 1758)


 * Tetramorium caldarium (Roger, 1857)


 * Tetramorium forte 	Forel, 1904


 * Tetramorium impurum (Foerster, 1850)


 * Tetramorium lanuginosum Mayr, 1870


 * Tetramorium meridionale Emery, 1870


 * Tetramorium pyrenaeicum Roszler, 1937


 * Tetramorium semilaeve Andre, 1883