Tapinoma erraticum

This is a small black ant, agile and aggressive on disturbance. Colonies usually contain several hundred workers and many queens. Nests are constructed under stones or in bare ground in dry sunny places and loose earth solaria for brood incubation are often constructed during the early summer. This species is partly aphidicolous and partly carnivorous. Alatae are developed in June with flights occurring during July (Collingwood 1979). Pashaei Rad et al. (2018) found this species in Iran on the ground in a moderate to low rainfall area.

Identification
Dark brown to black: head wedge shaped widening posteriorly; clypeus with median notch as wide as or wider than deep. Dorsum of alitrunk and appendages without standing hairs. Body covered with adpressed white pubescence and very finely punctured. Antennae 12 segmented, palp formula 6,4. Length: 2.6-4.2 mm. (Collingwood 1979)

Distribution
Throughout Central Europe from Spain to the Caucasus and from the mountains of South Italy to North Germany (Collingwood 1979).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Andorra, Armenia, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, Canary Islands, Channel Islands, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Kazakhstan, Kyrgyzstan, Liechtenstein, Lithuania, Luxembourg, Malta, Monaco, Montenegro, Netherlands, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Turkmenistan, Ukraine, United Kingdom of Great Britain and Northern Ireland, Åland Islands.

Biology
This ant has been associated with the butterfly Glaucopsyche alexis (Obregon et al. 2015).

Guiliani et al. (2019) observed this species foraging on extrafloral nectaries of the invasive Reynoutria x bohemica (Polygonaceae) in Tuscany. The habitats examined were river banks and disturbed habitats.

Nomenclature

 *  erraticum. Formica erratica Latreille, 1798: 44 (w.q.m.) FRANCE.
 * Wheeler, G.C. & Wheeler, J. 1951: 197 (l.).
 * Combination in Tapinoma: Mayr, 1855: 373; Smith, F. 1855a: 111.
 * Status as species: Latreille, 1802c: 182; Schenck, 1852: 67; Smith, F. 1855a: 111; Mayr, 1855: 373 (redescription); Nylander, 1856b: 71; Smith, F. 1858b: 55; Mayr, 1861: 41 (in key); Smith, F. 1861a: 34; Roger, 1863b: 13; Mayr, 1863: 455; Mayr, 1865: 62; Smith, F. 1871b: 2; Forel, 1874: 60 (in key); André, 1874: 185 (in key); Mayr, 1877: 11; Emery, 1878a: ix (in list); Emery, 1878b: 49; Emery & Forel, 1879: 454; Mayr, 1880: 30; Saunders, E. 1880: 210; André, 1881b: 63; André, 1882b: 222 (in key); White, W.F. 1884: 256; Forel, 1886e: clxvii; Forel, 1889: 256; Nasonov, 1889: 26; Forel, 1890a: lxviii; Saunders, E. 1890: 203; Emery, 1891b: 15; Forel, 1892i: 308; Lameere, 1892: 65; Dalla Torre, 1893: 164; Medina, 1893: 105; Forel, 1894d: 13; Forel, 1895d: 227; Saunders, E. 1896: 27; Ruzsky, 1902d: 19; Ruzsky, 1903b: 308; Ruzsky, 1905b: 473; Forel, 1906c: 189; Wasmann, 1906: 115 (in key); Forel, 1907e: 15; Forel, 1909c: 104; Bondroit, 1910: 489; Karavaiev, 1910b: 46; Karavaiev, 1911: 9; Forel, 1911d: 350; Karavaiev, 1912a: 15; Krausse, 1912b: 165; Emery, 1913a: 40; Stitz, 1914: 78; Forel, 1915d: 43 (in key); Emery, 1916b: 215; Wheeler, W.M. & Mann, 1916: 172; Escherich, 1917: 329; Bondroit, 1918: 89; Santschi, 1919e: 246; Menozzi, 1921: 32; Santschi, 1921a: 114; Soudek, 1922: 59; Müller, 1923b: 111; Lomnicki, 1925b: 2; Emery, 1925d: 53 (redescription); Wheeler, W.M. 1926: 4; Kuznetsov-Ugamsky, 1927d: 34; Donisthorpe, 1927b: 200; Lomnicki, 1928: 6; Kuznetsov-Ugamsky, 1929b: 42; Karavaiev, 1930b: 147; Soudek, 1931: 11; Zimmermann, 1935: 43; Karavaiev, 1936: 169; Novák & Sadil, 1941: 96 (in key); Kratochvíl, in Kratochvíl, et al. 1944: 79; Donisthorpe, 1950e: 1062; Consani & Zangheri, 1952: 42; Ceballos, 1956: 309; Bernard, 1956b: 259; Baroni Urbani, 1964a: 6; Baroni Urbani, 1964b: 55; Baroni Urbani, 1964c: 159; Bernard, 1967: 257 (redescription); Baroni Urbani, 1968b: 475; Baroni Urbani, 1971c: 159; Pisarski, 1975: 28; Tarbinsky, 1976: 123 (redescription); Aktaç, 1977: 124; van Boven, 1977: 125; Kutter, 1977c: 181; Arnol’di & Dlussky, 1978: 547 (in key); Collingwood, 1978: 87 (in key); Collingwood, 1979: 35; Barquin Diez, 1981: 357; Schembri & Collingwood, 1981: 421; Seifert, 1984a: 151; Agosti & Collingwood, 1987b: 279 (in key); Dlussky, Soyunov & Zabelin, 1990: 169; Atanassov & Dlussky, 1992: 192; Hohmann, et al. 1993: 159; Shattuck, 1994: 143; Arakelian, 1994: 73; Bolton, 1995b: 400; Czechowski, et al. 2002: 14; Radchenko, 2007: 38; Seifert, 2007: 256; Casevitz-Weulersse & Galkowsky, 2009: 478; Lapeva-Gjonova, et al. 2010: 32; Boer, 2010: 16; Legakis, 2011: 23; Borowiec, L. & Salata, 2012: 537; Czechowski, et al. 2012: 66; Seifert, 2012a: 144; Berville, et al. 2013: 84; Borowiec, L. 2014: 167 (see note in bibliography); Radchenko, 2016: 88.
 * Senior synonym of atomus: Roger, 1863b: 13; Mayr, 1865: 62; Forel, 1874: 99 (in list); Emery & Forel, 1879: 454; Emery, 1925d: 53.
 * Senior synonym of bononiensis: Atanassov & Dlussky, 1992: 192; Radchenko, 2016: 88.
 * Senior synonym of breve: Dlussky, Soyunov & Zabelin, 1990: 169; Radchenko, 2016: 88.
 * Senior synonym of caerulescens: Mayr, 1863: 455; Mayr, 1865: 62; Forel, 1874: 99 (in list); Emery & Forel, 1879: 454; Dalla Torre, 1893: 164; Emery, 1913a: 40.
 * Senior synonym of collina: Schenck, 1852: 67; Smith, F. 1855a: 111; Mayr, 1855: 373; Nylander, 1856b: 71; Smith, F. 1858b: 55; Mayr, 1863: 455; Roger, 1863b: 13; Mayr, 1865: 62; André, 1874: 202 (in list); Forel, 1874: 99 (in list); Emery & Forel, 1879: 454; Dalla Torre, 1893: 164; Ruzsky, 1905b: 473; Emery, 1913a: 40; Emery, 1925d: 53; Radchenko, 2016: 88.
 * Senior synonym of glabrella: Smith, F. 1855a: 111; Mayr, 1855: 373; Nylander, 1856b: 71; Smith, F. 1858b: 55; Mayr, 1863: 455; Roger, 1863b: 13; Mayr, 1865: 62; André, 1874: 202 (in list); Forel, 1874: 99 (in list); Emery & Forel, 1879: 454; Dalla Torre, 1893: 164; Ruzsky, 1905b: 473; Emery, 1913a: 40; Radchenko, 2007: 38; Radchenko, 2016: 88.
 * Senior synonym of tauridis: Dlussky, Soyunov & Zabelin, 1990: 169; Radchenko, 2016: 88.
 * Senior synonym of transcaucasica: Dlussky, Soyunov & Zabelin, 1990: 169; Radchenko, 2016: 88.
 * Current subspecies: nominal plus platyops.
 * atomum. Formica atomus Latreille, 1798: 44 (w.) FRANCE.
 * Combination in Tapinoma: Roger, 1863b: 13.
 * Subspecies of erraticum: Latreille, 1802c: 183 (in text); Emery, in Dalla Torre, 1893: 165; Emery, 1913a: 40; Bolton, 1995b: 399 (error).
 * Junior synonym of erraticum: Roger, 1863b: 13; Mayr, 1865: 62; Emery & Forel, 1879: 454; Emery, 1925d: 53.
 * caerulescens. Formica caerulescens Losana, 1834: 314, fig. 3 (w.) ITALY. Junior synonym of erraticum: Mayr, 1865: 62.
 * glabrella. Formica glabrella Nylander, 1849: 38 (footnote) (w.) RUSSIA. Junior synonym of erraticum: Smith, F. 1855a: 111; Mayr, 1855: 373; Radchenko, 2007: 38.
 * collina. Tapinoma collina Foerster, 1850a: 43 (w.q.) GERMANY. Junior synonym of erraticum: Schenck, 1852: 67.
 * bononiensis. Tapinoma erraticum var. bononiensis Emery, 1925d: 55 (w.q.) ITALY. Junior synonym of erraticum: Atanassov & Dlussky, 1992: 192.
 * breve. Tapinoma breve Emery, 1925d: 60, fig. 12 (w.) KAZAKHSTAN. Subspecies of erraticum: Kuznetsov-Ugamsky, 1927d: 34. Junior synonym of erraticum: Dlussky, Soyunov & Zabelin, 1990: 169.
 * tauridis. Tapinoma tauridis Emery, 1925d: 59, fig. 11 (w.q.m.) UKRAINE. Junior synonym of erraticum: Dlussky, Soyunov & Zabelin, 1990: 169. See also: Karavaiev, 1927d: 336.
 * transcaucasica. Tapinoma tauridis subsp. transcaucasica Karavaiev, 1927d: 337 (w.m.) KAZAKHSTAN. [First available use of Tapinoma simrothi subsp. karavaievi var. transcaucasica Karavaiev, 1926e: 187; unavailable name.] Junior synonym of erraticum: Dlussky, Soyunov, & Zabelin, 1990: 169.

Type Material
Seifert (2012) - A recent search conducted by J. Casevitz-Weulersse showed that there were no Tapinoma specimens in the Latreille collection of Muséum National d'Histoire Naturelle Paris (J. Casevitz-Weulersse, pers. comm. 13 June 2008). There is also no information that any of the taxonomists of the 20th century has ever seen an original specimen of Tapinoma erraticum. Hence, a complete loss of original material is highly probable or, at least, a reliable identification is impossible due to the useless original description and missing labelling of specimens. The presence of a minimum of seven Tapinoma species in Europe (SEIFERT 1984) and the fact that a safe identification of some species cannot be guaranteed for all future by descriptive statements and published information induces the need to fix a neotype. As the description of Latreille contains no diagnostic information, the neotype designation can only follow the rationale that (a) it should belong to the species widely considered as T. erraticum in Europe and (b) that the neotype comes as nearly as practicable from the original type locality. The locus typicus of T. erraticum is Brive-la-Gaillarde (France). I selected a neotype worker from a nest sample kindly collected by Christophe Galkowski about 11 km south from the city centre of Brive. The neotype is labelled "FRA: 45.0517°N, 1.5372°E, Nespouls-Faugère, 330 m, along road, under stone, leg. Galkowski 2008.07.02" and "Neotype Tapinoma erraticum (Latreille, 1798), design. B.Seifert 2010". It is stored in the collection of the Senckenberg Museum für Naturkunde Görlitz and has the following data: CS 836 μm, CL / CW 1.100, SL / CS 1.024, EL / CS 0.259, dAN / CS 0.279, ExCly / CS 8.87%, ExOcc / CS 1.42%, MGr / CS 3.58%, MW / CS 0.646 and ML / CS 1.347. Digital files with detailed z-stack photographs of the neotype in different positions are available on request from SMN Görlitz. There are 19 nestmate workers from the neotype nest in the collection of SMN Görlitz either mounted or stored in undenatured 99.8% ethanol.

Taxonomic Notes
Khalili-Moghadam et al. (2019) - A recent paper on Albanian ants by Wagner et al. (2018) suggested that southern populations from the Balkans recorded as Tapinoma erraticum represent a cryptic undescribed species. Parts of populations from areas east of the Balkans probably belong to the same cryptic taxon but prior to the formal description we applied the appellation T. erraticum to the Iranian populations.

Borowiec & Salata (2020): Wagner et al. (2018) suggest that southeastern populations of Tapinoma erraticum belong to an independent species named provisionally as Tapinoma sp. Balc. Both taxa differ mostly in subtle characters in male genitalia, the southern taxon is also slightly larger than true T. erraticum. We re-examined our materials from Greece and Bulgaria and confirmed that in this region occurs only Tapinoma sp. Balc. The only confirmed localities of true T. erraticum from Balkan Peninsula are from Croatia, Montenegro, North Macedonia and Slovenia. With great probability populations from Jordan belong to the undescribed species. However, we cannot exclude occurrence of another cryptic species from this group in this region.

Worker
Seifert (2012) - Head, mesosoma and gaster covered by a rather dense pubescence. With maximum head length and width in visual plane, posterior margin of head in medium-sized and large specimens slightly excavated. Anteromedian clypeal excision usually at least as deep a wide. Eye contours deviating from ideally elliptic form, showing linear sections on median margin. Anterior margin of clypeus with a few standing setae, the two longest based near to the anterolateral margin of clypeal excision. Remaining surface of head capsule and dorsal mesosoma without standing setae, such are present on mandibles, coxae and ventral surface of gaster. Head, mesosoma and gaster dark blackish brown. Antennae, femora and tibiae dark brown. Tarsi and distal end of tibiae light yellowish brown.

Male
Seifert (2012) - (see fig. 3 in Seifert 1984 for genital in ventral view): excision of the subgenital plate much shallower than in Tapinoma subboreale. The caudal tips of subgenital plate rounded, much shorter and much less distant than in T. subboreale. Stipal tips, in contrast, widely separate and caudally strongly surpassing the tips of subgenital plate which is completely invisible in dorsal aspect.

References based on Global Ant Biodiversity Informatics

 * Abril S., C. Gómez. 2012. Lista actualizada de las especies de hormigas de Menorca (Islas Baleares, España) y primera cita de Monomorium andrei Saunders, 1890 (Hymenoptera, Formicidae) de la isla Boletín de la Sociedad Entomológica Aragonesa 50: 403-407.
 * Acosta, F. J., M. D. Martínez Ibáñez, and M. A. Morales. "Contribución al conocimiento de la mirmecofauna del encinar peninsular. I." Boletín de la Asociación Española de Entomología 6 (2) (1983): 379-391.
 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Aktaç, N. "Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon." Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41 (1977): 115-135.
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * Alonso de Medina, E., and X. Espadaler. "Nota sobre la entomofauna de la Sierra de Prades (Tarragona) (Formícidos)." Publicaciones del Departamento de Zoologia (Barcelona) 7 (1981): 67-71.
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnan X., M. Gracia, L. Comas, and J. Retana. 2009. Forest management conditioning ground ant community structure and composition in temperate conifer forests in the Pyrenees Mountains. Forest Ecology and Management 258(2): 5159.
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2011. Iberomyrmex 3: 30-31.
 * Asociacion Iberica de Mirmecologia. 2014. List of species collected during the Taxomara 2014 Oviedo. Iberomyrmex 6: 23-24.
 * Asociacion Iberica de Mirmecologia. 2015. List of species collected during the Taxomara Lisboa 2015 Iberomyrmex 7: 44-45.
 * Azcarate F. M., and B. Peco. 2012. Abandonment of grazing in a mediterranean grassland area: consequences for ant assemblages. Insect Conservation and Diversity 5: 279288.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. IV. La fauna mirmecologica delle isole Maltesi ed il suo significato ecologico e biogeografico. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 77: 408-559.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. V. Aspetti ecologici della Riviera del M. Cònero. Boll. Zool. 35: 39-76.
 * Baroni Urbani C. 1974. Studi sulla mirmecofauna d'Italia. XII. Le Isole Pontine. Fragm. Entomol. 9: 225-252.
 * Baroni Urbani C. 1977. Katalog der Typen von Formicidae (Hymenoptera) der Sammlung des Naturhistorischen Museums Basel (2. Teil). Mitt. Entomol. Ges. Basel (n.s.) 27: 61-102.
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
 * Baroni Urbani, C. "Catalogo delle specie di Formicidae d'Italia (Studi sulla mirmecofauna d'Italia X)." Memorie della Società Entomologica Italiana Volume 50 (1971): 5-287.
 * Baroni Urbani, C. "Formiche dell'Italia appenninica (Studi sulla mirmecofauna d'Italia, III)." Memorie del Museo Civico di Storia Naturale di Verona 12 (1964): 149-172.
 * Baroni Urbani, C. "Studi sulla mirmecofauna d'Italia. II. Formiche di Sicilia." Atti dell'Accademia Gioenia di Scienze Naturali in Catania (6) 16 (1964): 25-66.
 * Baroni Urbani, C. "Su alcune formiche raccolte in Turchia." Annuario dell'Istituto e Museo di Zoologia dell'Università di Napoli 16 (1964): 1-12.
 * Barquín, J. Las hormigas de Canarias. Taxonomia, ecologia y distribucion de los Formicidae In Secretariado de publicaciones de la Universidad de La Laguna, Colección Monografías Nº 3. Tenerife: Universidad de La Laguna, 1981.
 * Barrett K. E. 1967. Ants in South Brittany. Entomologist's Record and Journal of Variation 79:112-116.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
 * Baugnee J. Y., and J. F. Godeau. 2000. Signalement de Monomorium monomorium Bolton et de quelques autres fourmis à Belle-Ile-en-Mer, en Bretagne (Hymenoptera, Formicidae). Bulletin de la Société entomologique de France 105(2): 205-208.
 * Behr D., and K. Colln. 1993. Zur ameisenfauna (Hymenoptera, Formicidae) von Gonnersdorf (Kr. Daun). Dendrocopos 20: 148-160.
 * Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
 * Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:5971
 * Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1967. Recherches sur les fourmis des Monts-Dore. Annales de la Station biologique de Besse-en-Chandesse : 1-11.
 * Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
 * Bernard F. 1976. Écologie des fourmis des grès d'Annot, comparées à celles de la Provence calcaire. Annales du Muséum d'Histoire Naturelle de Nice 3: 33-54.
 * Bernard F. 1977. Écologie des fourmis du Parc national de Port-Cros. Bulletin du Muséum National d'Histoire Naturelle. Écologie Générale (3)36: 53-82.
 * Bernard F. 1978. Fourmis et milieu dans le massif des Maures. Vie et Milieu. Série C. Biologie Terrestre 27: 83-118.
 * Bernard, F. "Remarques sur le peuplement des Baléares en fourmis." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 47 (1956): 254-266.
 * Berville L., A. Hefetz, X. Espadaler, A. Lenoir, M. Renucci, O. Blight, and E. Provost. 2013. Differentiation of the ant genus Tapinoma (Hymenoptera: Formicidae) from the Mediterranean Basin by specis-specific cuticular hydrocarbons profiles. Myrmecological News 18: 77-92.
 * Berville L., M. Renucci, and E. Provost. 2012. Mise en place de protocoles de contrôle de la fourmi dArgentine (Linepithema humile) sur les îles de Port-Cros et de Porquerolles (Var, France). Sci. Rep. Port-Cros natl. Park, 26: 91-108.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bigot L. 1959. Complement a l'inventaire de la faune entomologique de la Camargue 93eme note). La Terre et la Vie 106 : 149-157.
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
 * Boer P., W. Dekoninck, A. J. Van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten (Amsterdam) 63: 54-58.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
 * Bolu H., and I. Ozgen. 2018. Some observation about Formicidae (Hymenoptera) in almond agroecosystems in East-Southeast Anatolian region of Turkey. Mun. Ent. Zool. 13(2): 495-498.
 * Bonaric J. C. 1971. Contribution a l'etude systematique et ecologique des formicides du Bas-Languedoc. PhD thesis Universite des sciences et techniques du Languedoc, 175 pages.
 * Bonaric J. C. 1971. Étude systématique et ecologique des fourmis de lHérault (suite). Ann. Soc. Hortic. Hist. Nat. Hérault 111: 119-126.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec L., and S. Salata. 2017. New records of ants (Hymenoptera: Formicidae) from southern Portugal. Acta Entomologica Silesiana 25(3): 1-10.
 * Boulay, R., J.M. Fedriani, A.J. Manzaneda and X. Cerda. 2005. Indirect Effects of Alternative Food Resources in an Ant-Plant Interaction. Oecologia 144(1):72-79
 * Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
 * Boven J. van 1949. Notes sur la faune des Hautes-Fagnes en Belgique. Bulletin et Annales de la Société Entomologique de Belgique 85: 135-143.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Braschler, B. and B. Baur. 2005. Experimental Small-Scale Grassland Fragmentation Alters Competitive Interactions among Ant Species. Oecologia 143(2):291-300
 * Cagniant, H. 1968. Liste preliminaire de fourmis forestieres d'Algerie. Resultats obtenus de 1963 a 1964. Bulletin de la Société d'Histoire Naturelle de Toulouse 104: 138-147
 * Carpintero S., J. Reyes-Lopes, and G. M. Luque. 2011. Ant community structure under Retama sphaerocarpa shrubs in a semi-arid environment. Entomological Science 14: 147153.
 * Carpintero, S., A. Tinaut, J. Reyes, and L. Arias. "Estudio faunístico de los formícidos (Hymenoptera: Formicidae) del Parque Nacional de Doñana." Boletín de la Asociación Española de Entomología 25 (1-2) (2001): 133-152.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Cherix D., and S. Higashi. 1979. Distribution verticale des fourmis dans le Jura vaudois et recensement prelimaire des bourdons (Hymenoptera, Formicidae et Apidae). Bull. Soc. Vaud. Sc. Nat. 356(74): 315-324.
 * Chevrier M., and C. Mouquet C. 2005. Etude des peuplements des invertébrés des dunes de Bretagne. Rapport GRETIA, avenant au Contrat-Nature 2, Conseil Régional de Bretagne, Conseils Généraux des Côtes d'Armor, du Finistère et du Morbihan : 127 p.
 * Colindre L. 2015. Les fourmis en Picardie: bilan 2014 (Hymenoptera/ Formicidae). Entomologiste Picard 26, 15 pages.
 * Collingwood C. A. 1955. Ants in S.W. Scotland. Entomologist's Record and Journal of Variation 67: 11-12.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
 * Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
 * Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Collingwood, C. A. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Comín del Río, P., and Andrés de Haro. "Datos iniciales para un estudio ecológico de las hormigas de Menorca." Bolletí de la Societat d´Historia Natural de les Balears 24 (1980): 23-48.
 * Comín del Río, P., and X. Espadaler. "Ants of the Pityusic islands." In Biogeography of the Pityusic Islands, edited by H. Kuhbler, J. A. Alcover and C. Guerau, 278-231. The Hague: Junk, 1984.
 * Comín del Río, P. "Los Formícidos de Menorca. Contribución al estudio taxonómico, geográfico y biológico." Tesina de licenciatura Universida (1977): 135 pp.
 * Comín del Río, P. Estudio de los formícidos de Baleares: Contribución al estudio taxonómico, geográfìco y biológico. Palma de Mallorca: Universidad de las Islas Baleares, 1988.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Mem. Soc. Entomol. Ital. 31: 38-48.
 * Cournault, L. and J.-C. de Biseau. 2009. Hierarchical perception of fertility signals and nestmate recognition cues in two dolichoderine ants. Behavioral Ecology and Sociobiology 63:1635-1641
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S., B. Markó, and L. Gallé. 2001. Ants (Hymenoptera: Formicidae) of Stana Valley (Romania): Evaluation of the effectiveness of a myrmecological survey.  Entomologica Romanica 6 : 121-126.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Cuní, M. "Insectos observados en los alrededores de Barcelona." Anales de la Sociedad española de Historia Natural XVII (1888): 133.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
 * Dekoninck W., F. Hendrickx, M. Dethier, and J. P. Maelfait. 2010. Forest Succession Endangers the Special Ant Fauna of Abandoned Quarries along the River Meuse (Wallonia, Belgium). Restoration Ecology 18(5): 681690.
 * Dekoninck W., H. De Koninck, J. Y. Baugnee, and J. P. Maelfait. 2007. Ant biodiversity conservation in Belgian calcareous grasslands: active management is vital. Belg. J. Zool. 137 (2): 137-146.
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Delye G. 1983. Contribution a l'etude des peuplements des invertebres de milieux extremes: les fourmis des dunes littorales del aCamargue (B. d. Rh) et de l'Espiguette (Gard). Bull. Soc. Linneenne Provence 35: 121-124.
 * Dlussky G. M., O. S. Soyunov, and S. I. Zabelin. 1990. Ants of Turkmenistan. Ashkabad: Ylym Press, 273 pp.
 * Donisthorpe H. 1914. Myrmecophilous notes for 1913. Entomologist's Record and Journal of Variation 26: 37-45.
 * Donisthorpe, H. "A first instalment of the ants of Turkey." Annals and Magazine of Natural History (12)3 (1950): 1057-1067.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Dusmet, J. M. "Algunos formícidos y mutílidos de España." Actas de la Sociedad Española de Historia Natural Tomo XXVII (1899): 109.
 * Dvorak, L., P. BOGUSCH, I. MALENOVSKÝ, P. BEZDÌÈKA, K. BEZDÌÈKOVÁ, K. HOLÝ, P. LIKA, J. MACEK, L. ROLLER, M. RÍHA et al. "Hymenoptera of Hády Hill, near the city of Brno (Czech Republic), collected during the Third Czech-Slovak Hymenoptera meeting." Acta Musei Moraviae, Scientiae biologicae (Brno) 93 (2008): 53-92.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Emery C. 1878. Liste des fourmis de la collection de feu Camille van Volxem, avec la description d'une espèce nouvelle. Annales de la Société Entomologique de Belgique 21: viii-x.
 * Emery C. 1913. Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae. Genera Insectorum 137: 1-50.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Emery C. 1925. Revision des espèces paléarctiques du genre Tapinoma. Revue Suisse de Zoologie 32: 45-64.
 * Emery, C. "Alcune formiche dell'isola di Creta." Bull. Soc. Entomol. Ital. Resoc. Adun. 26 (1894): 7-10.
 * Emery, C. "Beiträge zur Kenntniss der palaearktischen Ameisen." Öfversigt af Finska Vetenskaps-Societetens Förhandlingar (Helsinki) 20 (1898): 124-151.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte seconda. Formiche dell'Europa e delle regioni limitrofe in Africa e in Asia." Annali del Museo Civico di Storia Naturale 12 (1878): 43-59.
 * Espadaler X., X. Roig, K. Gómez, and F. García. 2011. Formigues de les Planes de Son i mata de València (Hymenoptera, Formicidae) Treballs de la Institució Catalana d'Història Natural 16: 609-627.
 * Espadaler, X., J. Pujade-Villar, and A. Bernadou. "Contribució al coneixement de la taxonomia i la fenologia de les formigues (Hymenoptera: Formicidae) d'Andorra." Butlletí de l' Institució Catalana d'Història Natural 74 (2006): 81-90.
 * Espadaler, X., and C. Ascaso. "Adición a las hormigas del Montseny (Barcelona)." Orsis 5 (1990): 141-147.
 * Espadaler, X., and J. Reyes-López. "Dos nuevas localidades para una (¿rara?) hormiga endémica ibérica,Temnothorax blascoi (Espadaler, 1997) (Hymenoptera, Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 449-450.
 * Espadaler, X., and X. Roig. "Hormigas del Vall de Aneu." (2007).
 * Espadaler, X. "Contribución al conocimiento de los formícidos (Hymenoptera, Formicidae) del Pirineo catalán." Tesis Universida (1979): 285 pp.
 * Espadaler, X. "Formigues del Montseny." in: Terradas, J., Miralles, J. (eds.) El patrimoni biològic del Montseny Catàlegs d (1987): 101-103.
 * Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants  assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
 * Finzi, B. "Formiche dell'isola d'Elba e Monte Argentario." Bollettino della Società Entomologica Italiana 56 (1924): 12-15.
 * Finzi, B. "Risultati scientifici della spedizione Ravasini-Lona in Albania. III. Formiche." Bollettino della Società Entomologica Italiana 55 (1923): 1-4.
 * Forel A. 1886. Études myrmécologiques en 1886. Annales de la Société Entomologique de Belgique. 30: 131-215.
 * Forel A. 1890. Fourmis de Tunisie et de l'Algérie orientale. Annales de la Société Entomologique de Belgique 34: lxi-lxxvi.
 * Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
 * Forel A. 1904. Miscellanea myrmécologiques. Rev. Suisse Zool. 12: 1-52.
 * Forel A. 1904. Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Ezheg. Zool. Muz. 8: 368-388.
 * Forel A. 1906. Moeurs des fourmis parasites des genres Wheeleria et Bothriomyrmex. Revue Suisse de Zoologie 14: 51-69.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
 * Forel A. 1910. Glanures myrmécologiques. Ann. Soc. Entomol. Belg. 54: 6-32.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel, A. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
 * Forel, A. "Fourmis de Tunisie et de l'Algerie orientale recoltees et decrites par Auguste Forel." Annales de la Société Entomologique de Belgique 34 (1890): lxi-lxxvi.
 * Forel, A. "Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen." Annales de la Société Entomologique de Belgique 30 (1886): clix-clxviii.
 * Forel, A. "Südpalaearctische Ameisen." Mitteilungen der Schweizerischen Entomologischen Gessellschaft 9 (1895): 227-234.
 * Formidabel Database
 * Franch, J., and X. Espadaler. "Ants as colonizing agents of pine stumps in San Juan de la Peña (Huesca, Spain)." Vie et Milieu 38 (1988): 149-154.
 * Francois J. 1958. Contribution a l'etude ecologique des Formicides (Insectes, Hymenopteres) de la region Dijonnaise. Travaux du laboratoire de Zoologie et de la Station Aquicole Grimaldi de la Faculte des Sciences de Dijon 25, 39 pages.
 * Freitag A. 2013. Biodiversite 2010 en ville de Neuchatel: les fourmis (Hymenopteres, Formicidae). Bulletin de la societe Neuchateloise des sciences naturelles 133: 183-200.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Gadeau de Kerville H. 1922. Materiaux pour la Faune des Hymenopteres de la Normandie. Bull. Soc. Amis Sc. Nat. Rouen 1916-1921, 1922: 217-225.
 * Galkowski C. 2011. Une liste des fourmis (Hymenoptera, Formicidae) récoltées dans la région de Grasse, avec la mention dune nouvelle espèce de la faune de France. Bulletin de la Société linnéenne de Provence, 62 : 41-44.
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galle L. 1972. Study of ant-populations in various grassland ecosystems. Acta Biologica Szeged 18(1-4): 159-164.
 * Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
 * Galle L. 1993. Data to the ant fauna of the Bukk (Hymenoptera: Formicidae). Natural history of the national parks of Hungary 7: 445-448.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Gaspar C., and C. Thirion. 1978. Modification des populations d'Hymenopteres sociaux dans les milieux anthropogenes. Memorabilia Zoologica 29: 61-77.
 * Gaspare Charles. 1965. Étude myrmécologique d'une région naturelle de Belgique: la Famenne Survey des Fourmis de la Région (Hymenoptera, Formicidae). Institut agronomique de l'Etat a' Gembloux. 32(4): 427-434.
 * Gasperini, R. "Notizia sulla fauna imenotterologadalmata, II, Formicidae-Mutillidae-Scoliadae-Sapygidae-Pompilidae-Sphegidae-Chrysididae." Anuario Dalmatico 4 (1887): 143-160.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Goetsch, W. "Beiträge zur Biologie spanischer Ameisen." EOS (Revista española de entomología) 18 (1942): 175-241.
 * Gomez K. 2017. Two species of exotic ants (Hymenoptera: Formicidae) new to Malta. Boletin de la Sociedad Entomologica Aragonesa (S.E.A.) 61: 233-235.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Groc S., J. H. C. Delabie, R. Cereghino, J. Orivel, F. Jaladeau, J. Grangier, C. S. F. Mariano, and A. Dejean. 2007. Ant species diversity in the Grands Causses (Aveyron, France): In search of sampling methods adapted to temperate climates. C. R. Biologies 330: 913922.
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Haguet G., M. Chevrier, and E. Brunel. 2002. Les invertebres de la dune de Bon Abri, premier inventaire. Groupe d'Etude des Invertebres Armoricains, 37 pages.
 * Hauschteck-Jungen E., and H. Jungen. 1983. Ant chromosomes. II. Karyotypes of western palearctic species. Insectes Soc. 30: 149-164.
 * Hohmann H., F. La Roche, G. Ortega, and J. Barquín. 1993. Bienen, Wespen und Ameisen der Kanarischen Inseln. Veröff. Überseemus. Bremen Naturwiss. 12: 14.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-203.
 * Hosoishi, S., S. Yamane and K. Ogata. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13:345-350.
 * Jeffery H. G. 1931. The Formicidae (or ants) of the Isle of Wight. Proceedings of the Isle of Wight Natural History and Archaeological Society 2: 125-128.
 * Karaman C., K. Kiran, and V. Aksoy. 2014. New records of the genus Strumigenys Smith, 1860 (Hymenoptera, Formicidae) from Black Sea region of Turkey. Trakya University Journal of Natural Sciences, 15(2): 59-63.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karaman M. G., and G. S. Karaman. 2003. Contribution to the knowledge of the ants (Hymenoptera, Formicidae) from Serbia. The Montenegrin Academy of Sciences and Arts. Glasnik of the Section of Natural Sciences 15: 39-58.
 * Karavaiev V. 1927. Übersicht der Ameisenfauna der Krim nebst einigen Neubeschreibungen. Konowia 5: 281-303.
 * Khalili-Moghadam A., L. Borowiec, and A. Nemati. 2019. New records of ants (Hymenoptera: Formicidae) from the Chaharmahal va Bakhtiari Province of Iran with taxonomic comments. Polish Journal of Entomology 88 (2): 163–182.
 * Kiran K. C. Karaman, A. Lapeva-Gjonova, and V. Aksoy. 2017. Two new species of the "ultimate" parasitic ant genus Teleutomyrmex KUTTER, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic. Myrmecological News 25: 145-155.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Korlevic, A. "Prilozi fauni hrvatskih opnokrilaca." Glasn. Hrv. Narav. Dr. 5 (1890): 189-250.
 * Kozisek T. 1987. Ants (Formicoidea) of the Abrod State Nature Reserve. Ochrana Prírody 8: 205-208.
 * Krausse, A. H. "Ueber sardische Ameisen." Archiv für Naturgeschichte, Berlin A 78(7) (1912): 162-166.
 * Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
 * Kuznetsov G. T. 1990. Comparative analysis of Hymenoptera (Formicidae) population on altitudinal zones of central Kopetdag. Izvestiya Akademii Nauk Turkmenskoi SSR. Seriya Biologicheskikh Nauk 1990(3): 64-67.
 * Lapeva-Gjonova A., and K. Kiran. 2012. Ant fauna (Hymenoptera, Formicidae) of Strandzha (Istranca) Mountain and adjacent Black Sea coast. North-western journal of Zoology 8(1): 72-84.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis A. 1983. First contribution to the study of the ants (Hymenoptera, Formicidae) of the Zagori Region (Epirus, Greece): an annotated list of species Entomologia Hellenica 1: 3-6.
 * Legakis A. 1983. The Zoological Museum of the University of Athens 2. The collection of ants from Greece. Biologia Gallo-Hellenica 11(1): 85-87.
 * Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
 * Legakis Collection Database
 * Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
 * Livory A. 2003. Les fourmis de la Manche. L'Argiope 39: 25-49.
 * Lomnicki J. 1928. Spis mrówek Lwowa i okolicy. Ksiegi Pamiatkowej (Lecia Gimn. IV Jana Dlugosza Lwowie) 50: 1-10.
 * Lomnicki, J. "Une contribution à la connaissance de la faune des fourmis des îles Baléares." Polskie Pismo Entomologiczne - Bulletin Entomologique de Pologne 4 (1925): 1-3.
 * Lorinczi G. 2011. Lasius (Chthonolasius) nitidigaster Seifert, 1996 -a new ant species (Hymenoptera: Formicidae) for the Hungarian fauna. Natura Somogyiensis 19: 223-228.
 * Marikovsky P. I. 1979. Ants of the Semireche Desert. [In Russian.]. Alma Ata: Nauka, 263 pp.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Markó B., and S. Csősz. 2002. Die europäischen Ameisenarten (Hymenoptera: Formicidae) des Hermannstädter (Sibiu, Rumänien) Naturkundemuseums I.: Unterfamilien Ponerinae, Myrmicinae und Dolichoderinae. Annales Historico-Naturales Musei Nationalis Hungarici 94: 109-121.
 * Martinez J. J. 2008. Firebreaks in planted pine forests in Israel: patches for Mediterranean Bata ants. Vie et Milieu 58(3/4): 233-236.
 * Martorell, M. "Catálogos sinonímicos de insectos encontrados en Cataluña." Barcelona (1879).
 * Martínez Ibáñez, M. D., and A. Tinaut. "Nuevas especies de formícidos (Hymenoptera, Formicidae) para la Sierra de Albarracín (Teruel)." Real Sociedad Española de Historia Natural Tomo extra (1996): 174-177.
 * Medina, M. "Catálogo provisional de las hormigas de Andalucía." Actas de la Sociedad Española de Historia Natural 20 (1891): 95-104.
 * Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
 * Menozzi C. 1921. Formiche dei dintorni di Sambiase di Calabria. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici. 15: 24-32.
 * Meyer M., C. Braunert, R. Gerend, and I. Schrankel. 2003. Résultats de l'excursion annuelle du groupe de travail entomologique de la Société des naturalistes luxembourgeois. Bull. Soc. Nat. luxemb. 103: 103-120.
 * Mortazavi Z. S., H. Sadeghi, N. Aktac, L. Depa, L. Fekrat. 2015. Ants (Hymenoptera: Formicidae) and their aphid partners (Homoptera: Aphididae) in Mashhad region, Razavi Khorasan Province, with new records of aphids and ant species for fauna of Iran. Halteres 6:4-12.
 * Müller, G. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Nemet E., Z. Czekes, I. Tausan, and B. Marko. 2012. Contribution to the knowledge of the myrmecofauna of the Cefa Nature Park (North-Western Romania). Acta Scientiarum Transylvanica 20(1): 61-72.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Nieves-Aldrey J. L., F. Fontal-Cazalla, A. M. Garrido-Torres, and C. Rey del Castillo. 2003. Inventorio de Hymenoptera (Hexapoda) en el ventorrillo: un rico enclave de biodiversidad en la Sierra de Guadarrama (Espana Central). Graellsia 59(2-3): 25-43.
 * Nylander, W. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * Ordóñez-Urbano C., J. Reyes-López, and S. Carpintero-Ortega. 2007. Estudio faunísidos de los formícidos (Hymenoptera: Formicidae) asociados a los bosques de ribera en la Provincia de Córdoba (España). Primeras Aportaciones. Boletin Sociedad Entomológica Aragonesa 40: 367-375.
 * Ortiz, F. J., and A. Tinaut. "Introducción al conocimiento de las hormigas de la provincia de Almería." Publicaciones del Instituto de Estudios Almerienses. Boletín (Ciencias) 8 (1988): 223-231.
 * Ovazza M. 1950. Contribution à la connaissance des fourmis des Pyrénées-Orientales. Récoltes de J. Hamon Vie et Milieu 1: 93-94.
 * Ovazza M. 1953. Contributions a l'etude biologique de la Camargue, Formicides. Vie et Milieu 4(4) : 751-753.
 * Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
 * Paknia O., and M. Pfeiffer. 2011. Hierarchical partitioning of ant diversity: implications for conservation of biogeographical diversity in arid and semi-arid areas. Diversity and Distributions 17: 122-131.
 * Paknia O., and M. Pfeiffer. 2014. Niche-based processes and temporal variation of environment drive beta diversity of ants (Hymenoptera: Formicidae) in dryland ecosystems of Iran. Myrmecological News 20: 15-23.
 * Paraschivescu D. 1972. Fauna mirmecologica din zonele saline ale Romaniei. Studii si Cercetari de Biologie. Seria Zoologie 24: 489-495.
 * Pardo, X., J. Tormos, and A. Sendra. "Mirmecofauna de les Suredes valencianes." Miscellanea Zoologica 9 (1986): 251-256.
 * Pascual, M. R. "Estudio taxonómico y ecológico de los Formícidos de las Sierras de Alfacar, La Yedra, Huétor y Harana." Tesis Doctoral Universida (1986): 264 pp.
 * Pashaei Rad S., B. Taylor, R. Torabi, E. Aram, G. Abolfathi, R. Afshari, F. Borjali, M. Ghatei, F. Hediary, F. Jazini, V. Heidary Kiah, Z. Mahmoudi, F. Safariyan, and M. Seiri. 2018. Further records of ants (Hymenoptera: Formicidae) from Iran. Zoology in the Middle East 64(2): 145-159.
 * Passera L. 1967. Peuplement en fourmis terricoles du rebord Meridional des Causses jurassiques du Quercy: la lande calcaire a buis. Vie et milieu. Serie C, Biologie terrestre 18(1): 189-205.
 * Passera L. 1977. Peuplement myrmecologique du cordon littoral du Languedoc-Roussillon modifications anthropiques. Vie et Milieu 27: 249-265.
 * Pekar, S. 2009. Capture Efficiency of an Ant-Eating Spider, Zodariellum asiaticum (Araneae: Zodariidae), from Kazakhstan. Journal of Arachnology 37(3):388-391
 * Peru L. 1999. Quelques ajouts et corrections a un catalogue des fourmis de la region Centre (Hymenoptera, Formicidae). Bulletin de l'Entomologie Tourangelles 20(1): 9-12.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z. 2010. Contribution to the myrmecofauna (Hymenoptera: Formicidae) of Krusevac and its surroundings. Acta entomologica serbica, 2010, 15(1): 145-148.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Poldi B. 1980. Imenotteri formicidi della Brughiera di Rovasenda (Piemonte). In: Poldi, B. 1980. Quaderni sulla "Struttura delle zoocenosi terrestri". 1. La brughiera pedemontana. III. Serie AQ/1/110. Collana del programma finalizzato "Promozione della qualità dell'ambiente". Roma: Consiglio Nazionale delle Ricerche, pp. 7-19.
 * Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
 * Reyes-Lopez J., S. Carpintero. 2014. Comparison of the exotic and native ant communities (Hymenoptera: Formicidae) in urban green areas at inland, coastal and insular sites in Spain. Eur. J. Entomol. 111(3): 421428.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
 * Salata S., L. Borowiec, and A.Trichas. 2018. Taxonomic Revision of the Cretan Fauna of the Genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with Notes on the Endemism of Ant Fauna of Crete. Annales Zoologici (Warsaw) 68(4): 769-808.
 * Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
 * Santschi F. 1910. Contributions à la faune entomologique de la Roumanie. Formicides capturées par Mr. A. L. Montandon. Buletinul Societatii Române de Stiinte 19: 648-652.
 * Santschi, F. "Notes sur les fourmis paléarctiques. II. Fourmis d'Asie Mineure récoltées par M. H. Gadeau de Kerville." Boletín de la Real Sociedad española de Historia natural (Madrid) 21 (1921): 110-116.
 * Saunders, E. "Aculeate Hymenoptera collected by J. J. Walker, at Gibraltar and in North Africa. (Part I - Heterogyna)." Entomologists' Monthly Magazine XXVI (1890): 201-206 y 289-291.
 * Schembri, Stephen P. and Cedric A. Collingwood. A Revision of the Myrmecofauna of the Maltese Islands. 417-442.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
 * Seifert B. 2012. Clarifying naming and identification of the outdoor species of the ant genus Tapinoma Förster, 1850 (Hymenoptera: Formicidae) in Europe north of the Mediterranean region with description of a new species. Myrmecological News 16: 139-147.
 * Seifert B. D. D'Eustacchio, B. E. Kaufmann, M. Centorame, and M. Modica. 2017. Four species within the supercolonial ants of the Tapinoma nigerrimum complex revealed by integrative taxonomy (Hymenoptera: Formicidae). Myrmecological News 24: 123-144.
 * Seifert, B. "A method for differentiation of the female castes of Tapinoma ambiguum Emery and Tapinoma erraticum (Latr.) and remarks on their distribution in Europe north of the Mediterranean region ." Faun. Abh. Staatl. Mus. Tierk. Dresden 11 (1984): 151-155.
 * Sellier Y., C. Galkowski, C. Lebas, and P. Wegnez. 2016. Découverte de Temnothorax pardoi (Tinaut, 1987) dans la réserve naturelle nationale du Pinail (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 106-113.
 * Shattuck S. O. 1994. Taxonomic catalog of the ant subfamilies Aneuretinae and Dolichoderinae (Hymenoptera: Formicidae). University of California Publications in Entomology 112: i-xix, 1-241.
 * Sommer F. 1984. Etude de la myrmécofaune de la réserve de la Massane. Laboratoire Arago, Banyuls sur Mer (66), 29 pages.
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Soyer B. 1951. Les fourmis des collines des environs de Marseille. Bull Soc Linn Provence 18: 3-6.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Stitz, H. "Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae." Jena G. Fischer (1939): 428 pp.
 * Stukalyuk S. V. 2016. Crematogaster subdentata Mayr, 1877, a potentially invasive species of ant (Hymenoptera, Formicidae) new to the fauna of Crimea. Entomological Review 95(8)
 * Stumper R. 1953. Etudes myrmecologiques. XI. Fourmis luxembourgeoises. Bulletin Soc. Nat. luxemb. 57: 122-135.
 * Suay-Cano, V. A., A. Tinaut, and J. Selfa. "Las hormigas (Hymenoptera, Formicidae) asociadas a pulgones (Hemiptera, Aphididae) en la provincia de Valencia." Graellsia 58(1) (2002): 21-37.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Tinaut, A. "Contribución al conocimiento de los formícidos del Parque Nacional de Doñana." Boletín de la Asociación Española de Entomología 15 (1991): 57-63.
 * Trigos Peral G., and J. L. Reyes-Lopez. 2015. New record of Temnothorax algiricus (Hymenoptera, Formicidae) from the Iberian Peninsula and ten additional new records for the ant fauna from Málaga, with preliminary list of species from Natural Park Sierras de Tejeda, Almijara y Alhama (Andalusia, Spain). Boln. Asoc. esp. Ent. 39 (3-4): 411-416.
 * Trigos Peral G., and J. Reyes Lopez. 2016. Ants as bioindicators for monitoring of urban green in the south of the Iberian Peninsula. Proposal of functional groups. Iberomyrmex 8: 37-38.
 * Trigos-Peral G., and J. L. Reyes-Lopez. 2013. Primera relacion de los formicidos (Hymenoptera, Formicidae) de la Reserva Natural Lagunas de Campillos (Malaga). Boln. Asoc. esp. Ent. 37 (3-4): 217-224.
 * Urbieta Balado A., A. Palanca Soler, C. Castan Lanaspa, and X. Espadaler. 1984. Ants in the supraforestal pastures of the Tendenera massif. Documents dEcologie Pyrénéenne III-IV: 223-224.
 * Vanaclocha, P., C. - Monzó, K. Gómez, D. Tortosa, T. Pina, P. Castañera, and A. Urbaneja. "Hormigas (Hymenoptera: Formicidae) presentes en el suelo de los cítricos de la provincia de Valencia." Phytoma Nº 171 Ago (2005): 14-22.
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vogrin, V. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Wegnez P. 2014. Premières captures de Lasius distinguendus Emery, 1916 et de Temnothorax albipennis (Curtis, 1854) au Grand-Duché de Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société royale belge d’Entomologie 150 (2014) : 168-171.
 * Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
 * Wegnez P. 2018. Premières decouvertes de Myrmica bibikoffi Kutter, 1963 et de Ponera testacea Emery, 1895, au Luxembourg (Hymenoptera: Formicidae)Bulletin de la Société royale belge d’Entomologie 154: 263–272.
 * Wegnez P., and A. Ronk. 2017. Découverte de Camponotus herculeanus (Linnaeus, 1758) et signalement de quelques autres espèces rares de fourmis au Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société des naturalistes luxembourgeois 119 : 153–159.
 * Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
 * Wheeler W. M. and W. M. Mann. 1916. The ants of the Phillips Expedition to Palestine during 1914. Bulletin of the Museum of Comparative Zoology 60: 167-174.
 * Wheeler W. M., and W. M. Mann. 1916. The ants of the Phillips Expedition to Palestine during 1914. Bulletin of the Museum of Comparative Zoology 60: 167-174.
 * Wheeler, W. M. "Ants of the Balearic Islands." Folia Myrmecologica et Termitologica 1 (1926): 1-6.
 * Wiezik M. 2006. Ants (Hymenoptera:  Formiciadae)  of  the  Sitno  National  Nature  Reserve (tiavnické vrchy Protected Landscape Area). Naturae Tutela 10: 73-77.
 * Wiezik M., A. Wiezikova, and M. Svitok. 2010. Effects of secondary succession in abandoned grassland on the activity of ground-foraging ant assemblages (Hymenoptera: Formicidae). Acta Soc. Zool. Bohem. 74: 153-160
 * Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
 * de Biseau, J.-C., Y. Quinet, L. Deffernez and J.M. Pasteels. 1997. Explosive food recruitment as a competitive strategy in the ant Myrmica sabuleti (Hymenoptera, Formicidae). Insectes Sociaux 44:59-73
 * de Haro, Andrés, C. A. Collingwood, and P. Comín del Río. "Prospección mirmecológica por Ibiza y Formentera (Baleares)." Orsis 2 (1986): 115-120.
 * de Haro, Andrés, and C. A. Collingwood. "Formícidos de las Sierras de Prades-Montsant, Sierras de Cavalls-Alfara-Montes Blancos (Tarragona)." Boletín de la Estación Central de Ecología 10 (1981): 55-58.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Andalucia." Boletín de la Estación Central de Ecología 6 (12) (1977): 85-90.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Extremadura (España) y Sao Brás-Almodovar, Alcácer do Sal, Serra da Estrela (Portugal)." Boletim da Sociedade Portuguesa de Entomologia Suplemento 3(1) (1992): 95-104.