Strumigenys lalassa

Occurs in wet forest habitats, from lowlands to cloud forest. It is most abundant in mid-elevation and cloud forest sites, decreasing in abundance at lower elevations. It inhabits leaf litter on the forest floor. (Longino, Ants of Costa Rica)

Identification
Bolton (2000) - A member of the gundlachi-complex in the Strumigenys gundlachi group. Strumigenys lalassa is closest related to Strumigenys nubila; characters linking the two are noted there. Differentiation is easy as lalassa has the postpetiole disc reticulate-punctate, lacks an apicoscrobal hair, and has no standing hairs on the mesonotum. In nubila the postpetiole disc is smooth with sparse fine longitudinal costulae, an apicoscrobal hair is present, and standing hairs occur on the mesonotum.

Longino (Ants of Costa Rica) - Mandibles in full-face view linear, elongate and narrow; ventral surface of petiole without spongiform tissue; leading edge of scape with freely projecting hairs; inner margin of mandible without a tooth or distinctly enlarged denticle at or near the midlength; labral lobes short, trigger hairs at apices of lobes long; outer margins of mandibles bowed; mandibles long (MI 67).

Distribution based on Regional Taxon Lists
Neotropical Region: Colombia, Costa Rica, Ecuador, Panama.

Nomenclature

 *  lalassa. Pyramica lalassa Bolton, 2000: 189, fig. 134 (w.) PANAMA. Combination in Strumigenys: Baroni Urbani & De Andrade, 2007: 122

Worker
HOLOTYPE. TL 2.6, HL 0.66, HW 0.55, CI 83, ML 0.46, MI 70, SL 0.28, SI 51, PW 0.34, AL 0.66. Characters of gundlachi complex. Inner margin of mandible shallowly convex in proximal half, concave in distal half. Apex with 2 intercalary denticles between apicodorsal and apicoventral teeth. Preapical dentition of 5 - 8 teeth and denticles in total. Proximal of the apicodorsal tooth are two denticles, followed by two small spiniform teeth of which the distal is shorter than the proximal; sometimes a minute denticle occurs between these two teeth. Proximal of the longest spiniform preapical tooth are 1 - 4 denticles, located on the remainder of the concave section of the margin; some may be very poorly defined, low and obtuse. Eye with 5 ommatidia in the longest row. Apicoscrobal hair absent. Cephalic dorsum densely clothed with short spatulate ground-pilosity that is closely applied and directed anteriorly, but without standing hairs of any form. Pronotal humeral hair short and stout, flattened or expanded apically. Promesonotal dorsum with spatulate ground-pilosity but without any standing hairs. Disc of postpetiole reticulate-punctate. First gastral tergite and sternite smooth, the former with short basigastral costulae.

PARATYPE. TL 2.6-2.9, HL 0.66-0.74, HW 0.58-0.60, CI 81-88, ML 0.46-0.48, MI 66-72, SL 0.28-0.31, SI 48 - 52, PW 0.36-0.40, AL 0.66-0.76 (3 measured). As holotype but eye with 4 - 6 ommatidia in longest row.

The holotype and most other material entirely lacks standing hairs on the cephalic dorsum, but occasionally a widely separated pair of very short standing hairs is present close to the occipital margin.

Type Material
Holotype worker, Panama: Bocas del Toro, 12-14.vii.1987, 1050 m., 8°47'N, 82°12'W, premontane rain forest (D.M. Olson).

Paratypes. 1 worker with same data as holotype; 2 workers, Costa Rica: Heredia Prov., La Selva, 14.iii.1987, 50 m., 10026'N, 83°59'W, #241 (D. M. Olson); 6 workers, Prov. Heredia, 17 km. S Pto Viejo, 10°l8'N, 84°02'W, 550 m., 12.ix.1985, # 1088-s, wet forest litter sample (J. Longino); 4 workers, Prov. Puntarenas, Monteverde, 1500 m., 10°l8'N, 84°48'W, 14.xii.1987, # 1980-s, cloud forest litter sample, ground (J. Longino) (, BMNH,, ).

References based on Global Ant Biodiversity Informatics

 * Bolton, B. 2000. The Ant Tribe Dacetini. Memoirs of the American Entomological Institute 65
 * Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
 * Longino J. T. 2013. Ants of Nicargua. Consulted on 18 Jan 2013. https://sites.google.com/site/longinollama/reports/ants-of-nicaragua
 * Longino J. T. L., and M. G. Branstetter. 2018. The truncated bell: an enigmatic but pervasive elevational diversity pattern in Middle American ants. Ecography 41: 1-12.
 * Longino J. T., and R. K. Colwell. 2011. Density compensation, species composition, and richness of ants on a neotropical elevational gradient. Ecosphere 2(3): 16pp.
 * Longino J. et al. ADMAC project. Accessed on March 24th 2017 at https://sites.google.com/site/admacsite/
 * Patrick M., D. Fowler, R. R. Dunn, and N. J. Sanders. 2012. Effects of Treefall Gap Disturbances on Ant Assemblages in a Tropical Montane Cloud Forest. Biotropica 44(4): 472478.
 * Sosa-Calvo J., S. O. Shattuck, and T. R. Schultz. 2006. Dacetine ants of Panama: new records and description of a new species. Proceedings of the Entomological Society of Washington 108: 814-821.