Camponotus kiesenwetteri

The species inhabits dry grassy areas and thin forests, building nests in soil, often under rocks (Radchenko 1997).

Identification
Salata et al. (2019) - Head, mesosoma, and gaster uniformly black; metanotal groove present, shallow; propodeum with distinct dentate protrusions; body densely punctate, appears dull; base of scape with extension; whole body bears long, thick, pale, dense and erect setae, and short appressed microsetae; petiolar scale thick.

The species can be easily separated by the following combination of characters: strongly sculptured body, mesosoma with metanotal groove and posterior margin of propodeum with distinct dentate protrusions, and antennal scape with distinct basal extension. Camponotus nitidescens and Camponotus schulzi both differ in having a partly shiny body, and Camponotus boghossiani differs in having a propodeum without apical protrusions and an antennal scape without basal extension.

Distribution
Salata et al. (2019) - Greece: Attica, North Aegean Islands, South Aegean Islands (Cyclades, Dodecanese), Central Greece, Crete (Chania, Heraklion, Lasithi, Rethymno), Ionian Islands, Central Macedonia, Eastern Macedonia and Thrace, Peloponnese; Cyprus; Turkey: Balıkesir, İzmir and Muğla.

Distribution based on Regional Taxon Lists
Palaearctic Region: Cyprus, Greece , Iran, Turkey.

Nomenclature

 * . Formica (Hypoclinea) kiesenwetteri Roger, 1859: 241 (w.m.) (m. excluded, see Mayr, 1861: 39) GREECE (Zakynthos (= Zante) I.).
 * [Misspelled as kiesenvetteri by André, 1874: 178 (in key).]
 * Forel, 1886e: clxi (q.m.).
 * Combination in Camponotus: Mayr, 1861: 36 (in key);
 * combination in C. (Orthonotomyrmex): Forel, 1914a: 273; Santschi, 1921f: 310;
 * combination in C. (Myrmentoma): Forel, 1912i: 92; Emery, 1925b: 121.
 * Status as species: Mayr, 1861: 36 (in key); Roger, 1861b: 164; Mayr, 1863: 399; Roger, 1863b: 1; André, 1874: 178 (in key); Emery & Forel, 1879: 449; Forel, 1879a: 94; André, 1882a: 150 (in key); Forel, 1886e: clx; Forel, 1889: 260; Dalla Torre, 1893: 237; Emery, 1894j: 10; Emery, 1896d: 376 (in list); Forel, 1910a: 23; Forel, 1911d: 355; Emery, 1915h: 2; Emery, 1925b: 121; Menozzi, 1936d: 304; Hamann & Klemm, 1976: 677; Aktaç, 1977: 126; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 283 (in key); Collingwood, 1993b: 195; Bolton, 1995b: 106; Radchenko, 1996b: 1197 (in key); Radchenko, 1997b: 705; Petrov, 2006: 108 (in key); Legakis, 2011: 31; Karaman, C. et al. 2011: 189; Borowiec, L. & Salata, 2012: 477; Kiran & Karaman, 2012: 7; Karaman, C. & Aktaç, 2013: 50 (in key); Borowiec, L. 2014: 35 (see note in bibliography); Lebas, et al. 2016: 136; Borowiec, L. & Salata, 2018: 4; Salata & Borowiec, 2018c: 43; Salata, Loss, et al. 2019: 92.
 * Senior synonym of cypria: Salata, Loss, et al. 2019: 92.
 * cypria. Camponotus (Orthonotomyrmex) kiesenwetteri var. cypria Emery, 1920c: 26 (w.) CYPRUS.
 * Combination in C. (Myrmentoma): Emery, 1925b: 121.
 * Subspecies of kiesenwetteri: Emery, 1925b: 121; Bolton, 1995b: 95.
 * Junior synonym of kiesenwetteri: Salata, Loss, et al. 2019: 92.

Taxonomic Notes
Seifert (2019): This taxon has been described from Greece. The type specimen depicted in antweb.org under FOCOL2486 shows a much stronger, more reticulate-foveolate sculpture and much denser pubescence on all body surfaces than observed in any member of the C. lateralis group. This results in a matte surface appearance at smaller magnifications. Yet, C. kiesenwetteri is certainly related to this group considering several body shape characters.

Type Material
Salata et al. (2019) - Syntype workers, Greece [Syntype workers of Formica (Hypoclinea) kiesenwetteri, AntWeb, (FOCOL2486 and FOCOL2487)

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) of Samos Island, Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-13.
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Collingwood, C. A. 1993. A Comparitive Study of the Ant Fauna of Five Greek Islands. Biologia Gallo-hellenica. 20,1:191-197
 * Collingwood, C. A.. "A comparative study of the ant fauna of five Greek islands." Biologia Gallo-Hellenica 20 (1993): 191-197.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Emery C. 1920. Studi sui Camponotus. Bullettino della Società Entomologica Italiana 52: 3-48.
 * Emery, C.. "Alcune formiche dell'isola di Creta." Bull. Soc. Entomol. Ital. Resoc. Adun. 26 (1894): 7-10.
 * Emery, C.. "Escursioni zoologiche del Dr. Enrico Festa nell'Isola di Rodi. XII. Formiche." Bollettino del Museo di Zoologia ed Anatomia Comparatadella Reale Università di Torino 30 (1915): 1-7.
 * Emery, C.. "Spicilegio mirmecologico." Bollettino della Societa Entomologica Italiana 33 (1901): 57-63.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel, A.. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
 * Forel, A.. "Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen." Annales de la Société Entomologique de Belgique 30 (1886): clix-clxviii.
 * Hamann H. H. F., and W. Klemm. 1976. Ergebnisse der von Dr. O. Paget und Dr. E. Kritscher auf Rhodos durchgeführten zoologischen Exkursionen. XVI. Formicidae. Annalen des Naturhistorischen Museums in Wien 80: 669-679.
 * Karaman C., N. Aktac, and K. Kiran. 2009. Ants of the genus CamponotusMayr, 1861 (Hymenoptera: Formicidae) in the Kaz Mountains, Turkey, with descriptions of sexuals of Camponotus candiotes Emery, 1894 and Camponotus ionius Emery, 1920. Tubitak 35(2): 183-197.
 * Legakis A. 1983. Aspects of the distribution of ants in an insular mediterranean ecosystem (Cyclades Is., Greece) Rapports et Procès-Verbaux des Réunions de la Commission Internationale pour l'Exploration Scientifique de la Mer Méditerranée. Monaco 28: 121-122.
 * Legakis A. 1983. The Zoological Museum of the University of Athens 2. The collection of ants from Greece. Biologia Gallo-Hellenica 11(1): 85-87.
 * Legakis A. 1989. Aspects of the distribution of ants in an insular mediterranean ecosystem (Cyclades Isl., Greece). Rev. Ecol. Biol. Sol. 26 (3): 363-369.
 * Legakis Collection Database
 * Radchenko A. G. 1997. A review of ants of the subgenus Myrmentoma, Genus Camponotus (Hymenoptera, Formicidae), from Asian Palaearctic. Entomological Review 77(8): 1050-1059.
 * Salata S., A. C. Loss, C. Karaman, K. Kiran, and L. Borowiec. 2019. Review of the Camponotus kiesenwetteri group (Hymenoptera, Formicidae) in the Aegean with the description of a new species. ZooKeys 899: 85-107.
 * Salata S., L. Borowiec, and A.Trichas. 2018. Taxonomic Revision of the Cretan Fauna of the Genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with Notes on the Endemism of Ant Fauna of Crete. Annales Zoologici (Warsaw) 68(4): 769-808.
 * Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
 * Salata S., and L. Borowiec. 2018. A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae). ZooKeys 789: 139–159.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Seifert B. 2019. A taxonomic revision of the members of the Camponotus lateralis species group (Hymenoptera: Formicidae) from Europe, Asia Minor and Caucasia. Soil Organisms 91:7–32.