Messor wasmanni

A grain harvesting species (Collingwood and Agosti 1996). This taxon belongs to a group of species widespread in the Middle East and Central Asia, where several to many still unreviewed forms occur. Its westernmost known range reaches Sardinia and Corsica. It is common in central and southern Italy, and in the Balkans (Rigato & Toni, 2011).

Identification
Collingwood and Agosti (1996) - HW 2.5-2.9; EL/HW 0.16-0.19. This species differs from Messor semirufus and related species by the shape of the pronotum which is mildly flattened dorsally and expanded laterally into a protuberant boss at each side. This is a constant and characteristic feature and most easily seen in slanted profile.

Distribution based on Regional Taxon Lists
Palaearctic Region: Bulgaria, Croatia, Greece, Iran, Israel, Italy, Lebanon, Malta, Montenegro, Oman, Republic of Macedonia, Romania, Slovenia, Syrian Arab Republic, Turkey.

Nomenclature

 * . Messor barbarus subsp. wasmanni Krausse, 1910: 524 (in text) (w.) ITALY (Sardinia).
 * [Also described as new by Krausse, 1911: 15.]
 * Finzi, 1929: 83 (q.); Finzi, 1933: 163 (m.).
 * As unavailable (infrasubspecific) name: Krausse, 1912b: 164; Emery, 1915a: 249; Emery, 1916b: 142; Santschi, 1917e: 90; Emery, 1921f: 71; Menozzi, 1921: 26; Emery, 1922c: 94.
 * Subspecies of barbarus: Emery, 1912f: 97.
 * Subspecies of semirufus: Santschi, 1927c: 230; Finzi, 1928c: 788; Finzi, 1929: 82; Finzi, 1930d: 312; Finzi, 1933: 163; Zimmermann, 1935: 17.
 * Subspecies of meridionalis: Bondroit, 1920b: 305; Baroni Urbani, 1964b: 30 (footnote); Baroni Urbani, 1969a: 331; Baroni Urbani, 1968b: 434; Baroni Urbani, 1971c: 59.
 * Junior synonym of semirufus: Bernard, 1967: 147; Baroni Urbani, 1974: 227.
 * Junior synonym of meridionalis: Collingwood, 1978: 68; Poldi, et al. 1995: 3.
 * Status as species: Casevitz-Weulersse, 1990b: 156 (in key); Collingwood, 1993b: 194; Bolton, 1995b: 258; Collingwood & Agosti, 1996: 322; Bračko, 2006: 137; Petrov, 2006: 92 (in key); Bračko, 2007: 17; Casevitz-Weulersse & Galkowski, 2009: 489; Karaman, M.G. 2011b: 40; Legakis, 2011: 11; Borowiec, L. & Salata, 2012: 516; Kiran & Karaman, 2012: 21; Borowiec, L. & Salata, 2013: 385 (in text); Borowiec, L. 2014: 114; Bračko, et al. 2014: 14; Lebas, et al. 2016: 300; Borowiec, L. & Salata, 2018: 6; Salata & Borowiec, 2018c: 46; Salata & Borowiec, 2019a: 66.

A very common grain-harvesting ant found on all of the Greek islands, on the southern Balkan mainland, in South Italy and the Mediterranean islands of Corsica, Sardinia and Sicily but not in Malta or the Balearics. It has also been taken locally in Tunisia but its occurrence so far south as Oman is a surprise. However, Tohme (1968) and Tohme & Tohme (1981) recorded it as the synonym Messor concolor from Lebanon and Syria respectively. (Collingwood and Agosti 1996)

Type Material
Collingwood and Agosti (1996) - Specimens labelled M. concolor cotype in both the Oxford Hope Museum and the NHMB are confirmed as M. wasmanni, M concolor being a junior synonym.

References based on Global Ant Biodiversity Informatics

 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. IV. La fauna mirmecologica delle isole Maltesi ed il suo significato ecologico e biogeografico. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 77: 408-559.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. V. Aspetti ecologici della Riviera del M. Cònero. Boll. Zool. 35: 39-76.
 * Baroni Urbani C. 1969. Studi sulla mirmecofauna d'Italia. VIII. L'Isola di Giannutri ed alcuni scogli minori dell'arcipelago Toscano. Atti Soc. Toscana Sci. Nat. Mem. Ser. B 75: 325-338.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., C. Galkowski, and S. Salata. 2015. What is Tetramorium semilaeve André, 1883? (Hymenoptera, Formicidae). ZooKeys 512:39-62.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) of Samos Island, Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-13.
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J. 1990. Étude systématique de la myrmécofaune corse (Hymenoptera, Formicidae) (Première partie). Bull. Mus. Natl. Hist. Nat. Sect. A Zool. Biol. Écol. Anim. (4) 12: 135-163.
 * Casevitz-Weulersse J. 1992. La myrmecofaune de la reserve naturelle de Scandola, inventaire spécifique (1984/85-1991). Trav. Sci. Parc nat. Res. Nat. Corse, Fr, 36: 85-108.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse, J.. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Collingwood, C. A. 1993. A Comparitive Study of the Ant Fauna of Five Greek Islands. Biologia Gallo-hellenica. 20,1:191-197
 * Collingwood, C. A. and D. Agosti. 1996. Formicidae (Insects: Hymenoptera) of Saudi Arabia (Part 2) Fauna of Saudi Arabia 15: 300-385.
 * Collingwood, C. A., and Donat Agosti. "Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2)." Fauna of Saudi Arabia 15 (1996): 300-385.
 * Collingwood, C. A.. "A comparative study of the ant fauna of five Greek islands." Biologia Gallo-Hellenica 20 (1993): 191-197.
 * Collingwood, C.A. and D. Agosti. 1996. Formicidae (Insects: Hymenoptera) of Saudi Arabia (Part 2) Fauna of Saudi Arabia, 15: 300-385.
 * Consani M. 1947. Reperti corologici sulle formiche italiane. Redia 32: 179-182.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Finzi B. 1929. Le forme italiane del genere Messor. Bollettino della Società Entomologica Italiana 61: 75-94.
 * Finzi, B.. "Quarto contributo alla conoscenza della fauna mirmecologica della Venezia Giulia." Bollettino della Società Entomologica Italiana 60 (1928): 128-130.
 * Finzi, B.. "Raccolte entomologiche nell'Isola di Capraia fatte da C. Mancini e F. Capra (1927-1931). II. Formicidae." Memorie della Società Entomologica Italiana 11 (1933): 162-165.
 * Gomez K. 2017. Two species of exotic ants (Hymenoptera: Formicidae) new to Malta. Boletin de la Sociedad Entomologica Aragonesa (S.E.A.) 61: 233-235.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Krausse, A. H.. "Ueber sardische Ameisen." Archiv für Naturgeschichte, Berlin A 78(7) (1912): 162-166.
 * Legakis Collection Database
 * Müller, G.. "Faunisticna istrazivanja sjeverodalmatinskih otoka: Dugi otok i Kornati (1925-1927): Orthopteroidea, Coleoptera i Formicidae." Acta biol. 1 (1957): 187-218.
 * Müller, G.. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Rigato S., and I. Toni. 2011. Short notes 21. Hymenoptera, Formicidae. Pp. 873-882 in: Nardi, G.; Whitmore, D.; Bardiani, M.; Birtele, D.; Mason, F.; Spada, L.; Cerretti, P. (eds.) 2011. Biodiversity of Marganai and Montimannu (Sardinia). Research in the framework of the ICP Forests network. Conservazione Habitat Invertebrati, 5. Sommacampagna, Verona: Cierre Edizioni, 896 pp.
 * Salata S., L. Borowiec, and A.Trichas. 2018. Taxonomic Revision of the Cretan Fauna of the Genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with Notes on the Endemism of Ant Fauna of Crete. Annales Zoologici (Warsaw) 68(4): 769-808.
 * Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
 * Salata S., and L. Borowiec. 2018. A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae). ZooKeys 789: 139–159.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary contributions toward a revision of Greek Messor Forel, 1890 (Hymenoptera: Formicidae). Turkish Journal of Zoology 43: 52-67.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Santschi, F. 1917. Races et variétés nouvelles du Messor barbarus L. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 8: 89-94.
 * Santschi, F.. "Races et variétés nouvelles du Messor barbarus L." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 8 (1917): 89-94.
 * Santschi, F.. "Revision des Messor du groupe instabilis Sm. (Hymenopt.)." Boletín de la Real Sociedad española de Historia natural (Madrid) 27 (1927): 225-250.
 * Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
 * Sharaf M. R., B. L. Fisher, H. M. Al Dhafer, A. Polaszek, and A. S. Aldawood. 2018. Additions to the ant fauna (Hymenoptera: Formicidae) of Oman: an updated list, new records and a description of two new species. Asian Myrmecology 10: e010004
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vesnic A., and R. Skrijelj. 2014. Biodiversity and variation in species of the genus Messor Forel, 1890 (Hymenoptera, Formicidae) in the Mediterranean part of Bosnia and Herzegovina. Works of the Faculty of Forestry University of Sarajevo 1: 37-44.
 * Zimmermann, S.. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.