Simopelta williamsi

Workers have been collected from leaf litter.

Distribution based on Regional Taxon Lists
Neotropical Region: Colombia, Costa Rica, Ecuador.

Habitat
Wet cloud forest at 570 (Wheeler, 1935) - 1450 meters elevation.

Castes
Known only from the worker caste.

Nomenclature

 *  williamsi. Simopelta williamsi Wheeler, W.M. 1935d: 14, fig. 3 (w.) ECUADOR. Combination in Belonopelta: Baroni Urbani, 1975b: 300; in Simopelta: Bolton, 1995b: 383. See also: Gotwald & Brown, 1967: 267; Mackay & Mackay, 2008: 324.

Worker
Length 3.3 mm.

Head nearly a fourth longer than broad, subrectangular, very slightly broader in front than behind, with nearly straight sides and posterior border. Eyes strongly convex, hemispherical, situated at the anterior fourth of the head. Mandibles rather broad, with evenly convex external and sharp masticatory borders, furnished with three teeth, the basal broad, truncated and separated by a toothless diastema from the two approximated and rather long apical teeth. Clypeus abrupt, perpendicular, bluntly carinate, its anterior border triangularly produced in the middle but without a spine. Frontal carinae narrow, subangular laterally; frontal groove shallow anteriorly, deep posteriorly, not reaching the middle of the head.

Antennae long; scapes som•ewhat arcuately bent, surpassing the posterior border of the head by a distance equal to their greatest diameter; first funicular joint nearly twice as long as broad, second decidedly broader than long, 3-10 longer than broad but the eighth to tenth only slightly so; terminal joint distinctly enlarged, glandiform, pointed, as long as the three preceding joints together. Thorax broad through the pronotum, which without the well-developed neck is as broad as long, with convex sides and less convex dorsum; promesonotal suture distinct and impressed; mesonotum laterally compressed, in profile straight and sloping to the distinct but shallow mesoepinotal impression; mesoepinotal suture obsolete; mesosternum with a small but distinct denticle near the insertion of the middle coxa; epinotum nearly twice as long as broad, scarcely broader behind than in front, strongly and transversely rounded above, in profile straight and horizontal, rounding posteriorly through a semicircular arc into the declivity, the lower portion of which is straight, perpendicular and submarginate on the sides. Petiole narrow, about one and one-half times as long as broad, broader behind than in front, with well-developed anterior teeth. In profile the node is higher than long and slightly higher behind than in front, with straight and perpendicular anterior and posterior surfaces, and feebly convex superior surface, the tooth depending from the ventral surface large and acutely angular. Postpetiole broader than long, not strongly truncated anteriorly and not marked off from the gaster by a constriction, so that the postpetiolar portion of the abdomen is short and oval and closely resembles that of Odontomachus. Sting well-developed. Legs slender.

Shining; the postpetiolar portion of the gaster more so than the remainder of the body; mandibles, head and thorax densely and finely reticulate or shagreened, with the following superimposed sculpture: head above and behind regularly reticulate-rugulose, anteriorly more longitudinally rugulose, on each side of the gula obliquely rugose, with several coarse punctures or foveolae; pronotum reticulately and concentrically, pleurae more strongly and sublongitudinally rugulose; mesonotum, base and declivity sharply, regularly and transversely rugulose or striate; sculpture of petiole more finely rugulose than that of the thorax, the rugules on the summit of the node being indistinctly concentric, on the sides longitudinal and on the anterior land posterior surfaces transverse. Postpetiolar portion of abdomen very smooth and shining, with sparse piligerous punctures. Antennal scapes more sharply, legs more delicately granular or shagreened.

Pilosity white, erect, of uneven length, longest and most abundant on the head, less so on the thorax, shortest on the abdomen; oblique and rather abundant on the antennae and legs. Pubescence present only on the funiculi.

Piceous or brown black; lower portions of epinotum more reddish; legs including coxae castaneous; tarsi, tip of gaster and sting yellowish.

Type Material
Described from six speciemns taken by Dr. F. X. Williams during December, 1922, at Naranjapata, Ecuador, at an altitude of 1850 ft.