Temnothorax schaumii

Mackay (2000) - This species nests in the bark of living trees, in branches, logs and oak galls of trees. It occurs in many habitats that range from from desert canyons in trees, to grasslands, to shaded deciduous forests. Smith (1924) found this to be the most common Temnothorax in Mississippi and Wesson and Wesson (1940) found this species to be common in oak trees in Ohio.

Identification
Mackay (2000) - A member of the Temnothorax schaumii species complex. These ants have 11-segmented antennae and are usually concolorous dark brown, but are occasionally concolorous yellow. The head is nearly completely covered with fine striae, which merge with the dense punctures. Occasionally there is a central strip, which is partly free of sculpture and somewhat shining. The top of the mesosoma is mostly punctate, with a few striae, the side of the mesosoma has numerous striae with punctures between them. The propodeal spines range from tiny angles to small spines, which are dull and rounded. The petiole and postpetiole are punctate and the node of the petiole is weakly truncate, with round edges.

The 11 segmented antenna and tiny propodeal spines separate this species from all other species with 11 segmented antennae in the subgenus, except Temnothorax whitfordi. It can be easily distinguished from Temnothorax whitfordi as the head and pronotum are predominantly punctate (predominantly smooth and shining in Temnothorax whitfordi, but the pronotum may be punctate as in Temnothorax schaumii). The punctures on the pronotum of Temnothorax schaumii are fine and completely cover the surfaces, whereas in Temnothorax whitfordi they are coarse and do not densely cover the surface. The small spines separate it from the others in the schaumii species complex.

Distribution
United States: Kansas, Indiana, Iowa, Missouri, Oklahoma, Texas, Arkansas, Louisiana, Illinois, Kentucky,Tennessee, Alabama, Mississippi, Ohio, Pennsylvania, Massachusetts, New Jersey, Washington D. C., West Virginia, Virginia, North Carolina, South Carolina, Georgia.

Distribution based on Regional Taxon Lists
Nearctic Region: United States.

Habitat
Ranges from from desert canyons in trees, to grasslands, to shaded deciduous forests

Abundance
common to abundant

Texas
Wheeler (1903) in notes about a Temnothorax schaumii synonym T. fortinodis:

"I have found only a single colony of this form at Austin. This had taken up its abode in an abandoned gall of Holcaspis cinerosus Basset on the live-oak (Quercus virginiana). It contained 143 workers, a single deiiJated queen and 35 larval in different stages. The latter were white and not greenish like the larval of L. obturator which inhabits the same kind of galls. The entrance to the fortinodis nest was a small round hole with much worn edges, evidently the modified exit of some parasite on the Holcaspis."

Nomenclature

 *  schaumii. Leptothorax schaumii Roger, 1863a: 180 (w.) U.S.A. Mayr, 1886d: 451 (m.); Wesson, L.G. & Wesson, R.G., 1940: 95 (q.); Wheeler, G.C. & Wheeler, J. 1960b: 23 (l.). Combination in L. (Myrafant): Smith, D.R. 1979: 1394; in Temnothorax: Bolton, 2003: 272. Senior synonym of fortinodis, gilvus, melanoticus: Creighton, 1950a: 271. See also: Wheeler, W.M. 1903c: 232; Mackay, 2000: 403.
 * fortinodis. Leptothorax fortinodis Mayr, 1886d: 452 (w.q.) U.S.A. Combination in L. (Myrafant): Smith, M.R. 1950: 30. Subspecies of schaumii: Wesson, L.G. & Wesson, R.G., 1940: 96. See also: Wheeler, W.M. 1903c: 233. Junior synonym of schaumii: Creighton, 1950a: 271.
 * gilvus. Leptothorax fortinodis var. gilvus Wheeler, W.M. 1903c: 235 (w.q.) U.S.A. Junior synonym of schaumii: Creighton, 1950a: 271.
 * melanoticus. Leptothorax fortinodis var. melanoticus Wheeler, W.M. 1903c: 235 (w.q.) U.S.A. Wesson, L.G. & Wesson, R.G., 1940: 94 (m.). Subspecies of fortinodis: Buren, 1944a: 287. Junior synonym of schaumii: Creighton, 1950a: 271.

Type Material
Mackay (2000) - Pennsylvania. The types could not be located in Roger's collection (Museum fur Naturkunde Zentralinstitutder Humboldt-Universitiitzu Berlin), and probably no longer exist.

Worker
2.5 Millim. lang, rothlich gelb, Hinflerleib an der Spitze etwas braunlich; die abstehenden Harchen sind keulenformig, kurz wie bei corticalis. Fuhler eilfgliedrig, ganz gelb. Mandibeln 4 – 5 zahnig, sehr undeutlicb gerunzelt. Clipeus ohne glanzenden Langseindruck. Kopf schwach langs gerunzelt, seitlich und hinten fein verworren oder fingerhutartig punktirt, schwach glanzend. Thorax dem von corticalis ganz ahnlich, oben ohne irgend einen Quereindruck, fingerhutartig punktirt, wenig glanzend, hinten in 2 kurze, an der Basis breite, 3-eckige Zahuchen endend. Stielchenglieder wie gewohnlich, das erste unten nit einem zahnformigen Vorsprung. Abdomen glatt, glanzend, Schienen onbehaart.

(Wheeler 1903) Length 2.5-2.75 mm, Mandibles 5-toothed. Clypeus convex witho:ut median impression; its anterior border rather straight. Antennre 11-jointed; scape reaching hardly to half way between the eye and the posterior angle of the head, funiculus with a distinctly 3-jointed club; first funicular joint almost as long as joints 2-5 together; joints 2-7 distinctly broader than long; terminal joint fully as long as the two preceding joints. Thorax rather short, flattened dorsally and laterally, broader in front than behind, with distinct and rather sharp humeral angles, and with a constriction at the mesoepinotal suture. Epinotal spines very short, dentiform, not longer than broad at their bases. Petiole seen from above oblong, 1 ½ times as long as broad, its sides parallel except at the peduncle which is narrower; in profile the anterior dorsal slope is concave and about the same length as the straight or somewhat convex posterior slope; ventral surface with a distinct tooth directed forward. Postpetiole scarcely half again as broad as the petiole, distinctly broader than long, oblong, with distinct though rounded anterior angles. Its dorsal surface is evenly semicircular in profile. Gaster of the usual shape, with small but distinct anterior angles.

Clypeus with sharp longitudinal rugae, two of which, near the middle, are more prominent than the others. Mandibles with distinct longitudinal rugae. Head, clypeus and mandibles with a silky luster, the first traversed by fine parallel rugae separated by rows of foveolate punctures, which are clearest in certain lights on the posterior lateral surfaces and cheeks. Thorax, petiole and. postpetiole opaque, covered uniformly with foveolate punctures. Gaster smooth and shining.

Hairs moderately numerous on the body, white, erect, clavate; short on the head and thorax, much longer on the gaster and of intermediate length on the pedicel. Hairs on the antennae and legs minute, non-clavate, appressed.

Yellowish-red, the edges of the mandibles black. Gaster in some specimens dark-brown thcroughout, in others yellow or with much of the base of the first segment yellow. Antennae and legs yellow, club and sometimes also the scape of the former, infuscated.

Male
(Wheeler 1903) Length 3.2 mm

Mandibles dentate, touching each other with their blades. Antennae 12-jointed, scape about as long as the first three joints of the funiculus together; funiculus from the second joint to the end of uniform thickness, filiform; second joint a little shorter than the third, shorter, in fact, than any of the succeeding joints. Instead of spines or teeth, the epinotum bears two indistinct elongate swellings. Radial cell of wings short and closed.

Mandibles rather smooth and shining, with scattered punctures near their inner edges. Clypeus moderately shining and very delicately longitudinally rugose. Cheeks and region between antennal insertions and eyes sharply striated longitudinally; front with delicate longitudinal rugae; vertex finely reticulate punctate. Thorax rather smooth and shining, median and posterior portions of mesonotum finely longitudinally rugose and in part obliquely. Petiole, postpetiole and gaster smooth and shining.

Pilosity sparse, tibim without suberect hairs.

Blackish-brown, pedicel and gaster darker. Mandibles, antennae except the brown scape, joints of legs, tarsi, and in part also the joints of the pedicel, yellow or reddish-yellow. Wings clear, hyaline.

Etymology
Commemorative. For the collector of the types, Prof. Schaum.

References based on Global Ant Biodiversity Informatics

 * Amstutz M. E. 1943. The ants of the Kildeer plain area of Ohio (Hymenoptera, Formicidae). The Ohio Journal of Science 43(4): 165-173.
 * Annotated Ant Species List Ordway-Swisher Biological Station. Downloaded at http://ordway-swisher.ufl.edu/species/os-hymenoptera.htm on 5th Oct 2010.
 * Belcher A. K., M. R. Berenbaum, and A. V. Suarez. 2016. Urbana House Ants 2.0.: revisiting M. R. Smith's 1926 survey of house-infesting ants in central Illinois after 87 years. American Entomologist 62(3): 182-193.
 * Carroll T. M. 2011. The ants of Indiana (Hymenoptera: Formicidae). Master's Thesis Purdue university, 385 pages.
 * Clark A. T., J. J. Rykken, and B. D. Farrell. 2011. The Effects of Biogeography on Ant Diversity and Activity on the Boston Harbor Islands, Massachusetts, U.S.A. PloS One 6(11): 1-13.
 * Coovert G. A. 2005. The Ants of Ohio (Hymenoptera: Formicidae). Ohio Biological Survey, Inc. 15(2): 1-207.
 * Coovert, G.A. 2005. The Ants of Ohio (Hymenoptera: Formicidae) Ohio Biological Survey Bulletin New Series Volume 15(2):1-196
 * Dash S. T. and L. M. Hooper-Bui. 2008. Species diversity of ants (Hymenoptera: Formicidae) in Louisiana. Conservation Biology and Biodiversity. 101: 1056-1066
 * Davis W. T., and J. Bequaert. 1922. An annoted list of the ants of Staten Island and Long Island, N. Y. Bulletin of the Brooklyn Entomological Society 17(1): 1-25.
 * Del Toro I., K. Towle, D. N. Morrison, and S. L. Pelini. 2013. Community Structure, Ecological and Behavioral Traits of Ants (Hymenoptera: Formicidae) in Massachusetts Open and Forested Habitats. Northeastern Naturalist 20: 1-12.
 * Del Toro, I. 2010. PERSONAL COMMUNICATION. MUSEUM RECORDS COLLATED BY ISRAEL DEL TORO
 * Deyrup M., C. Johnson, G. C. Wheeler, J. Wheeler. 1989. A preliminary list of the ants of Florida. Florida Entomologist 72: 91-101
 * Deyrup, M. 2003. An updated list of Florida ants (Hymenoptera: Formicidae). Florida Entomologist 86(1):43-48.
 * DuBois M. B. 1981. New records of ants in Kansas, III. State Biological Survey of Kansas. Technical Publications 10: 32-44
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-791
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-792
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-793
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-794
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-795
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-796
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-797
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-798
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-799
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-800
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-801
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-802
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-803
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-804
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-805
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-806
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-807
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-808
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-809
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-810
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-811
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-812
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-813
 * DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-814
 * Dubois, M.B. and W.E. Laberge. 1988. An Annotated list of the ants of Illionois. pages 133-156 in Advances in Myrmecology, J. Trager
 * Ellison A. M., and E. J. Farnsworth. 2014. Targeted sampling increases knowledge and improves estimates of ant species richness in Rhode Island. Northeastern Naturalist 21(1): NENHC-13NENHC-24.
 * Emery C. 1895. Beiträge zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8: 257-360.
 * Forster J.A. 2005. The Ants (hymenoptera: Formicidae) of Alabama. Master of Science, Auburn University. 242 pages.
 * Frye J. A., T. Frye, and T. W. Suman. 2014. The ant fauna of inland sand dune communities in Worcester County, Maryland. Northeastern Naturalist, 21(3): 446-471.
 * General D., and L. Thompson. 2008. Ants of Arkansas Post National Memorial: How and Where Collected. Journal of the Arkansas Academy of Science 62: 52-60.
 * General D.M. & Thompson L.C. 2007. Ants (Hymenoptera: Formicidae) of Arkansas Post National Memorial. Journal of the Arkansas Acaedemy of Science. 61: 59-64
 * Gregg R. E. 1945 (1944). The ants of the Chicago region. Annals of the Entomological Society of America 37: 447-480
 * Guénard B., K. A. Mccaffrey, A. Lucky, and R. R. Dunn. 2012. Ants of North Carolina: an updated list (Hymenoptera: Formicidae). Zootaxa 3552: 1-36.
 * Headley A. E. 1943. The ants of Ashtabula County, Ohio (Hymenoptera, Formicidae). The Ohio Journal of Science 43(1): 22-31.
 * Hill, J.G. 2006. Ants collected at Okatibbee Lake, Lauderdale County, Mississippi
 * Ipser R. M. 2004. Native and exotic ants (Hymenoptera: Formicidae) of Georgia: Ecological Relationships with implications for development of biologically-based management strategies. Doctor of Philosophy thesis, University of Georgia. 165 pages.
 * Ipser, R.M., M.A. Brinkman, W.A. Gardner and H.B. Peeler. 2004. A Survey of Ground-Dwelling Ants (Hymenoptera: Formicidae) in Georgia. The Florida Entomologist 87(3) 253-260.
 * Ivanov, K. 2019. The ants of Ohio (Hymenoptera, Formicidae): an updated checklist. Journal of Hymenoptera Research 70: 65–87.
 * Ivanov K., L. Hightower, S. T. Dash, and J. B. Keiper. 2019. 150 years in the making: first comprehensive list of the ants (Hymenoptera: Formicidae) of Virginia, USA. Zootaxa 4554 (2): 532–560.
 * Lynch J. F. 1988. An annotated checklist and key to the species of ants (Hymenoptera: Formicidae) of the Chesapeake Bay region. The Maryland Naturalist 31: 61-106
 * Lynch J. F., and A. K. Johnson. 1988. Spatial and temporal variation in the abundance and diversity of ants (Hymenoptera: Formicidae) in the soild and litter layers of a Maryland forest. American Midland Naturalist 119(1): 31-44.
 * MacGown J. A., J. G. Hill, R. L. Brown, T. L. Schiefer, J. G. Lewis. 2012. Ant diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi. Mississippi Agricultural and Forestry Experiment Station Bulletin 1197: 1-30
 * MacGown J. A., J. G. Hill, and M. Deyrup. 2009. Ants (Hymenoptera: Formicidae) of the Little Ohoopee River Dunes, Emanuel County, Georgia. J. Entomol. Sci. 44(3): 193-197.
 * MacGown J. A., J. G. Hill, and R. L. Brown. 2010.  Native and exotic ant in Mississippi state parks.  Proceedings:  Imported Fire Ant Conference, Charleston, South Carolina, March 24-26, 2008: 74-80.
 * MacGown J. A., and R. L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
 * MacGown, J.A. and JV.G. Hill. Ants of the Great Smoky Mountains National Park (Tennessee and North Carolina).
 * MacGown, J.A. and R.L. Brown. 2006. Observations on the High Diversity of Native Ant Species Coexisting with Imported Fire Ants at a Microspatial Scale in Mississippi. Southeastern Naturalist 5(4):573-586
 * MacGown, J.A. and R.L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A., J.G. Hill, R.L. Brown and T.L. 2009. Ant Diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi Report #2009-01. Schiefer. 2009.
 * MacGown. J. 2011. Ants collected during the 25th Annual Cross Expedition at Tims Ford State Park, Franklin County, Tennessee
 * Mackay W. P. 2000. A review of the New World ants of the subgenus Myrafant, (genus Leptothorax) (Hymenoptera: Formicidae). Sociobiology 36: 265-444.
 * Mackay W. P., and E. E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
 * Mackay, W.P. and E. *Mackay, W. P. and E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Edwin Mellen Press, Lewiston, NY.
 * Mahon M. B., K. U. Campbell, and T. O. Crist. 2017. Effectiveness of Winkler litter extraction and pitfall traps in sampling ant communities and functional groups in a temperate forest. Environmental Entomology 46(3): 470–479.
 * Mann H. R., E. Rowe, J. Selfridge, and D. L. Price. 2018. Leaf litter and arboreal ants (Hymenoptera: Formicidae) in a Mid-Atlantic Forest. Northeastern Naturalist 25(2): 341-354.
 * Menke S. B., E. Gaulke, A. Hamel, and N. Vachter. 2015. The effects of restoration age and prescribed burns on grassland ant community structure. Environmental Entomology http://dx.doi.org/10.1093/ee/nvv110
 * Menke S. B., and N. Vachter. 2014. A comparison of the effectiveness of pitfall traps and winkler litter samples for characterization of terrestrial ant (Formicidae) communities in temperate savannas. The Great Lakes Entomologist 47(3-4): 149-165.
 * Moody J. V., and O. F. Francke. 1982. The Ants (Hymenoptera, Formicidae) of Western Texas Part 1: Subfamily Myrmicinae. Graduate Studies Texas Tech University 27: 80 pp.
 * Nuhn, T.P. and C.G. Wright. 1979. An Ecological Survey of Ants (Hymenoptera: Formicidae) in a Landscaped Suburban Habitat. American Midland Naturalist 102(2):353-362
 * O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
 * O'Neill J.C. and Dowling A.P.G. 2011. A Survey of the Ants (hymenoptera: Formicidae) of Arkansas and the Ozark Mountains. An Undergraduate Honors, University of Arkansas. 18pages.
 * Parys K. A., M. L. Gimmel, and S. J. Johnson. 2013. Checklist of Insects Associated with Salvinia minima Baker in Louisiana, USA. Check List 9(6): 14881495.
 * Resasco J., S. L. Pelini, K. L. Stuble, N. J. Sanders, R. R. Dunn, S. E. Diamond, A. M. Ellison, N. J. Gotelli, and D. J. Levey. 2014. Using Historical and Experimental Data to Reveal Warming Effects on Ant Assemblages. PLoS ONE 9(2): e88029. doi:10.1371/journal.pone.0088029
 * Roeder K. A., and D. V. Roeder. 2016. A checklist and assemblage comparison of ants (Hymenoptera: Formicidae) from the Wichita Mountains Wildlife Refuge in Oklahoma. Check List 12(4): 1935.
 * Smith M. R. 1924. An annotated list of the ants of Mississippi (Hym.). Entomological News 35: 47-54.
 * Smith M. R. 1934. Dates on which the immature or mature sexual phases of ants have been observed (Hymen.: Formicoidea) (continued from page 251). Entomological News 45: 264-267.
 * Smith M. R. 1935. A list of the ants of Oklahoma (Hymen.: Formicidae). Entomological News 46: 235-241.
 * Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
 * Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
 * Turner C. R., and J. L. Cook. 1998. The ants (Hymenoptera: Formicidae) of the Caddo Lake region of northeast Texas. Texas Journal of Science 50: 171-173.
 * Tynes J. S., and R. E. Hutchins. 1964. Studies of plant-nesting ants in east central Mississippi. American Midland Naturalist 72(1): 152-156.
 * Van Pelt A., and J. B. Gentry. 1985. The ants (Hymenoptera: Formicidae) of the Savannah River Plant, South Carolina. Dept. Energy, Savannah River Ecology Lab., Aiken, SC., Report SRO-NERP-14, 56 p.
 * Van Pelt, A. 1983. Ants of the Chisos Mountains, Texas (Hymenoptera: Formicidae) . Southwestern Naturalist 28:137-142.
 * Wang C., J. Strazanac and L. Butler. 2000. Abundance, diversity and activity of ants (Hymenoptera: Formicidae) in oak-dominated mixed Appalachian forests treated with microbial pesticides. Environmental Entomology. 29: 579-586
 * Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
 * Wheeler G. C., J. N. Wheeler, and P. B. Kannowski. 1994. Checklist of the ants of Michigan (Hymenoptera: Formicidae). The Great Lakes Entomologist 26(4): 297-310
 * Wheeler G. C., and J. Wheeler J. 1989. A checklist of the ants of Oklahoma. Prairie Naturalist 21: 203-210.
 * Wheeler W. M. 1903. A revision of the North American ants of the genus Leptothorax Mayr. Proceedings of the Academy of Natural Sciences of Philadelphia. 55: 215-260.
 * Wheeler W. M. 1905. An annotated list of the ants of New Jersey. Bulletin of the American Museum of Natural History. 21: 371-403.
 * Wheeler W. M. 1917. A list of Indiana ants. Proceedings of the Indiana Academy of Science 26: 460-466.
 * Wheeler, G.C. and J. Wheeler. 1985. A checklist of Texas ants. Prairie Naturalist 17:49-64.
 * Wheeler, G.C., J. Wheeler and P.B. Kannowski. 1994. CHECKLIST OF THE ANTS OF MICHIGAN (HYMENOPTERA: FORMICIDAE). Great Lakes Entomologist 26:1:297-310
 * Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
 * Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71