Aphaenogaster enspelensis

Distribution
This taxon was described from.

Nomenclature

 * †. †Aphaenogaster enspelensis Jessen, 2020: 22, figs. 9a, b (q.) GERMANY.

Queen
Position: Head, alitrunk from dorsolateral, petiole and postpetiole from anterolateral. Gaster missing.

Colour: Black.

BL about 11.32. Head longer than wide, with distinct, rounded occipital corners. Posterior margin of head is indicating a median depression. Clypeus slightly convex in profile. Median part of clypeus with longitudinal striae, anterior clypeus margin here smooth. Eyes located slightly below mid length of head. Mandibles strongly developed, sub-triangular. Apex of left mandible missing. Due to compaction masticatory margin is bended downwards, only three triangle-shaped teeth can be identified. Based on estimation, there should be more than six triangular-shaped teeth. Shape and size of apical tooth is not clear either. Scape slightly surpasses head posteriorly. Head sculpture with wide coarse striae, visible at frons, at clypeus, between antennal sockets and eyes, and at gena. Alitrunk robust, high, arched. Metanotum well developed. Propodeal spines stout with broad base. Gradually descending posterior face of propodeum transversely striated. Scutum and scutellum sculptured rugosely. Tibial spurs on foreleg brushed or combed. At first and second tarsal, segment of protarsus bristles preserved. Wing venation with closed cells mcu, 1+2r and rm. Shape of cell mcu is trapezoid. The vein rs-m does not appear as a direct vertical extension of vein 2r-rs, but branches distally slightly offset from the vein RS. The results is a very short vein 4RS. In all other aspects, the description of the wing venation of the genus applies. Petiole stout with round bar-like peduncle and high distinct node. Petiole’s position is not clear; the node might have been twisted. Anterior face of node steep, tapered and rounded at top. Top of node with longitudinal respectively transverse grooves, depending whether the petiole has been twisted or not. Posterior face of node less steep. Peduncle broadly connected with alitrunk. Postpetiole disarticulated from petiole. Helcium almost round in cross section, its anterior margin is thickened, ventrally with a little invers notch. Node of postpetiole round, domeshaped, not tapered, slightly constricted towards gaster.

Holotype NHMM-PE1997/5555-LS: BLw/ oG: 7.50, HL: 1.98, HW*: 1.82, ED: 0.54, GeL: 0.51, ML: 1.07, MML: 0.75, SL: 1.84, AL: 3.36, AH: 2.42, FWL: 9.15, ScuL: 2.05, ScutL: 0.64, HiTL: 2.11, PH: 0.87, PPL: 0.46 (excl. helcium), PPH: 0.78, HeH: 0.44. Wing venation: 2M-Cu: 0.84, 1Cu: 0.84, 1M: 0.47, m-cu: 0.57, 1RS+M: 0.47, RS+M: 0.51. Indices: CI: 92.0, SI: 101.0, IED/HL: 27.52, IHL/AL: 58.87, Imcu: 56.14, I2RS+M/1RS+M: 109.38, I2RS+M/m-cu: 108.85, I2RS+M/2M+Cu: 61.40. (Because of perspective shift, no measurement is given for PL; it has only been estimated for calculating BL).

Type Material
Holotype: NHMM-PE1997/5555-LS, winged gyne. Type locality and horizon: Enspel Oilshale, Rhineland-Palatinate, Germany. Enspel Formation, Upper Oligocene, MP 28; (24.56–24.79 Ma, Mertz et al. 2007), layer S14.

Etymology
Named after the village Enspel in the Westerwald Mountains (Rhineland-Palatinate, Germany), territory of the Lake Enspel Fossil-Lagerstätte. <!--Un-comment this template when adding type specimen data