Checklist of Leptogenys species

The following species and subspecies belong to the genus Leptogenys. Synonyms are listed under their senior names. For valid names only see Leptogenys species and for distribution information see Leptogenys species by Country.

A
acutangula Emery, 1914 (New Caledonia)

• Leptogenys acutangula brevinoda Emery, 1914

acutirostris Santschi, 1912 (Madagascar)

adlerzi Forel, 1900 (Australia)

alamando Rakotonirina & Fisher, 2014 (Madagascar)

alatapia Rakotonirina & Fisher, 2014 (Madagascar)

alluaudi Emery, 1895 (Madagascar)

amazonica Borgmeier, 1930 (Brazil, Ecuador)

ambo Rakotonirina & Fisher, 2014 (Madagascar)

amon Bolton, 1975 (Cameroun)

amu Lattke, 2011 (Colombia)

andritantely Rakotonirina & Fisher, 2014 (Madagascar)

angusta (Forel, 1892) (Madagascar)

angustinoda Clark, 1934 (Australia)

anitae Forel, 1915 (Australia)

anjara Rakotonirina & Fisher, 2014 (Madagascar)

ankhesa Bolton, 1975 (Democratic Republic of Congo)

antillana Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

antongilensis Emery, 1899 (Madagascar)

arcirostris Santschi, 1926 (Madagascar)

arcuata Roger, 1861 (Barbados, Bolivia, Brazil, French Guiana, Grenada, Guadeloupe, Guyana, Lesser Antilles, Saint Vincent and the Grenadines, Suriname)

• Leptogenys arcuata deletangi Santschi, 1921

arnoldi Forel, 1913 (Zambia, Zimbabwe)

aspera (André, 1889) (Borneo, Indonesia, Malaysia, Vietnam)

assamensis Forel, 1900 (Bangladesh, India)

attenuata (Smith, F., 1858) (South Africa, Zimbabwe)

• Leptogenys jaegerskjoeldi Santschi, 1914

australis (Emery, 1888) (Argentina, Brazil, Uruguay)

• Leptogenys hanseni Borgmeier, 1930

avaratra Rakotonirina & Fisher, 2014 (Madagascar)

avo Rakotonirina & Fisher, 2014 (Madagascar)

B
barimaso Rakotonirina & Fisher, 2014 (Madagascar)

bellii Emery, 1901 (Eritrea, Ethiopia)

bezanozano Rakotonirina & Fisher, 2014 (Madagascar)

bidentata Forel, 1900 (Australia)

bifida Lattke, 2011 (Honduras)

binghamii Forel, 1900 (China, India, Myanmar)

birmana Forel, 1900 (Bangladesh, Cambodia, China, India, Myanmar, Thailand)

bituberculata Emery, 1901 (New Guinea)

bohlsi Emery, 1896 (Argentina, Brazil, Paraguay)

• Leptogenys bohlsi weiseri Santschi, 1925

borivava Rakotonirina & Fisher, 2014 (Madagascar)

borneensis Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia, Thailand)

breviceps Viehmeyer, 1914 (New Guinea)

• Euponera niger Donisthorpe, 1949

• Pseudoponera lubbocki Donisthorpe, 1938

bubastis Bolton, 1975 (Cameroun)

buyssoni Forel, 1907 (Ethiopia)

C
caeciliae Viehmeyer, 1912 (New Guinea)

camerunensis Stitz, 1910 (Cameroun, Democratic Republic of Congo)

carbonaria Lattke, 2011 (Colombia)

carinata Donisthorpe, 1943 (India)

castanea (Mayr, 1862) (South Africa, Zimbabwe)

• Leptogenys hewitti Santschi, 1932

• Leptogenys parva Forel, 1901

• Leptogenys parva bellua Forel, 1914

• Leptogenys parva dispar Santschi, 1914

centralis Wheeler, W.M., 1915 (Australia)

chalybaea (Emery, 1887) (Borneo, Cambodia, Indonesia, Malaysia, Thailand)

chamela Lattke, 2011 (Mexico)

chelifera (Santschi, 1928) (Australia)

chinensis (Mayr, 1870) (China, India, Philippines)

chrislaini Rakotonirina & Fisher, 2014 (Madagascar)

ciliata Lattke, 2011 (Ecuador)

clarki Wheeler, W.M., 1933 (Australia)

coerulescens Emery, 1895 (Madagascar)

comajojo Rakotonirina & Fisher, 2014 (Comoros)

confucii Forel, 1912 (Japan, Taiwan)

conigera (Mayr, 1876) (Australia)

• Leptogenys conigera mutans Forel, 1900

conradti Forel, 1913 (Cameroun, Ghana, Ivory Coast)

consanguinea Wheeler, W.M., 1909 (Guatemala, Mexico)

cordoba Lattke, 2011 (Mexico)

corniculans Lattke, 2011 (Brazil)

cracens Lattke, 2011 (Mexico)

crassicornis Emery, 1895 (Bangladesh, China, Myanmar)

crassinoda Arnold, 1926 (South Africa, Zimbabwe)

crudelis (Smith, F., 1858) (Brazil)

• Leptogenys rubicunda Borgmeier, 1930

crustosa Santschi, 1914 (Cameroun, Democratic Republic of Congo, Gabon, Guinea, Sudan)

• Leptogenys africanus Weber, 1942

• Leptogenys conradti rufipes Santschi, 1937

cryptica Bolton, 1975 (Ghana)

cuneata Lattke, 2011 (Ecuador, Panama, Peru)

D
dalyi Forel, 1900 (India)

darlingtoni Wheeler, W.M., 1933 (Australia)

davydovi Karavaiev, 1935 (China, Vietnam)

deborae Lattke, 2011 (Venezuela)

dentilobis Forel, 1900 (India)

diana Rakotonirina & Fisher, 2014 (Madagascar)

diatra Bolton, 1975 (Cameroun)

diminuta (Smith, F., 1857) (Australia, Bangladesh, Borneo, China, India, Indonesia, Malaysia, Myanmar, New Guinea, Philippines, Solomon Islands, Sri Lanka, Thailand, Vietnam)

• Leptogenys diminuta bismarckensis Forel, 1901

• Leptogenys diminuta hodgsoni Forel, 1900

• Leptogenys diminuta papuana Stitz, 1912

• Leptogenys diminuta santschii Mann, 1919

• Leptogenys diminuta stitzi Viehmeyer, 1924

• Leptogenys diminuta yarrabahna Forel, 1915

• Ponera ferox Smith, F., 1865

• Ponera simillima Smith, F. 1860:104 Smith, F., 1860

diminuta deceptrix Forel, 1901 (India)

diminuta diminutolaeviceps Forel, 1900 (India)

diminuta fruhstorferi Emery, 1896 (Indonesia)

diminuta laeviceps (Smith, F., 1857) (Borneo, India, Indonesia, Malaysia)

diminuta nongnongi Karavaiev, 1925 (Indonesia)

diminuta opacinodis (Emery, 1887) (Indonesia)

diminuta palliseri Forel, 1900 (India)

diminuta sarasinorum Forel, 1900 (Nepal)

diminuta striatula Emery, 1895 (Myanmar)

diminuta tjibodana Karavaiev, 1926 (Indonesia)

diminuta woodmasoni (Forel, 1886) (India)

donisthorpei Mann, 1922 (Honduras, Mexico)

drepanon Wilson, 1958 (New Guinea)

E
ebenina Forel, 1915 (Australia)

edsoni Rakotonirina & Fisher, 2014 (Madagascar)

elegans Bolton, 1975 (Cameroun, Ivory Coast, Kenya, Nigeria, Uganda)

elongata (Buckley, 1866) (Mexico, United States)

• Lobopelta mexicana Mayr, 1870

• Lobopelta septentrionalis Mayr, 1886

• Ponera texana Buckley, 1866

emeryi Forel, 1901 (New Guinea)

emiliae Forel, 1902 (India)

ergatogyna Wheeler, W.M., 1922 (Cameroun, Democratic Republic of Congo, Ghana, Uganda)

• Leptogenys cursor Arnold, 1954

erugata Lattke, 2011 (Colombia, Venezuela)

erythraea Emery, 1902 (Eritrea, Ethiopia)

excellens Bolton, 1975 (Democratic Republic of Congo)

excisa (Mayr, 1876) (Australia)

• Leptogenys excisa major Forel, 1910

exigua Crawley, 1921 (Australia)

exudans (Walker, 1859) (Sri Lanka)

F
falcigera Roger, 1861 (Comoros, Guam, Hawaii, Madagascar, Marshall Islands, Mauritius, Mayotte, Micronesia (Federated States of), Northern Mariana Islands, Palau, Philippines, Sri Lanka)

• Leptogenys insularis Smith, F., 1879

fallax (Mayr, 1876) (Australia)

famelica Emery, 1896 (Bolivia, Costa Rica)

fasika Rakotonirina & Fisher, 2014 (Madagascar)

ferrarii Forel, 1913 (Zimbabwe)

• Leptogenys ferrari dentatula Santschi, 1915

fiandry Rakotonirina & Fisher, 2014 (Madagascar)

foraminosa Lattke, 2011 (Colombia, Costa Rica)

foreli Mann, 1919 (New Guinea, Solomon Islands, Vanuatu)

• Leptogenys walkeri Donisthorpe, 1942

fortior Forel, 1900 (Australia)

fotsivava Rakotonirina & Fisher, 2014 (Madagascar)

foveonates Lattke, 2011 (Venezuela)

foveopunctata Mann, 1921 (Fiji)

fugax Mann, 1921 (Fiji)

furtiva Arnold, 1926 (Zimbabwe)

G
gagates Mann, 1922 (Honduras)

gaigei Wheeler, W.M., 1923 (Brazil, Colombia, Ecuador, French Guiana, Guyana, Suriname)

• Leptogenys gaigei defuga Wheeler, W.M., 1923

• Leptogenys melzeri Borgmeier, 1930

gatu Lattke, 2011 (Colombia, Costa Rica, Panama)

glabra Lattke, 2011 (Colombia)

gorgona Lattke, 2011 (Colombia, Ecuador)

gracilis Emery, 1899 (Comoros, Madagascar)

grandidieri Forel, 1910 (Madagascar)

guianensis Wheeler, W.M., 1923 (Brazil, French Guiana, Guyana)

guineensis Santschi, 1914 (Ghana, Guinea)

H
hackeri Clark, 1934 (Australia)

harmsi Donisthorpe, 1935 (Christmas Island)

havilandi Forel, 1901 (South Africa)

hebrideana Wilson, 1958 (Vanuatu)

hemioptica Forel, 1901 (Indonesia)

hezhouensis Zhou, 2001 (China)

honduriana Mann, 1922 (Honduras)

honoria Bolton, 1975 (Cameroun)

huapingensis Zhou, 2001 (China)

humiliata Mann, 1921 (Fiji)

hysterica Forel, 1900 (Borneo, India, Indonesia, Malaysia, Sri Lanka, Thailand)

I
iheringi Forel, 1911 (Brazil)

imerinensis Forel, 1892 (Madagascar)

imperatrix Mann, 1922 (Costa Rica, Ecuador, Guatemala, Honduras, Panama)

incisa Forel, 1891 (Madagascar, Mozambique)

indigatrix Wilson, 1958 (New Guinea)

ingens Mayr, 1866 (Colombia, Venezuela)

intermedia Emery, 1902 (Botswana, Kenya, Mozambique, South Africa, Zambia, Zimbabwe)

• Leptogenys brevinodis deflocata Santschi, 1926

• Leptogenys nitida adpressa Forel, 1914

• Leptogenys nitida aena Forel, 1914

• Leptogenys nitida brevinodis Forel, 1915

• Leptogenys nitida capensis Baroni Urbani, 1971

• Leptogenys nitida gracilis Santschi, 1914

• Leptogenys nitida grandior Forel, 1915

• Leptogenys nitida insinuata Santschi, 1914

• Leptogenys nitida speculans Santschi, 1926

• Leptogenys tenuis Stitz, 1911

• Ponera nitida Smith, F. 1858 Smith, F., 1858

intricata Viehmeyer, 1924 (Australia)

iridescens (Smith, F., 1857) (Borneo, Indonesia, Malaysia, Philippines, Thailand)

iridescens currens Forel, 1901 (Borneo, Indonesia, Malaysia)

iridipennis (Smith, F., 1858) (India)

ixta Lattke, 2011 (Mexico)

J
jeanettei Tiwari, 2000 (India)

jeanneli Santschi, 1914 (United Republic of Tanzania)

johary Rakotonirina & Fisher, 2014 (Madagascar)

josephi Mackay, W.P. & Mackay, E.E., 2004 (Costa Rica, Panama)

K
karawaiewi Santschi, 1928 (Indonesia)

keysseri Viehmeyer, 1914 (New Guinea)

khammouanensis Roncin & Deharveng, 2003 (Laos)

khaura Bolton, 1975 (Central African Republic, Nigeria)

kiche Lattke, 2011 (Guatemala)

kitteli (Mayr, 1870) (Bangladesh, Borneo, China, India, Indonesia, Malaysia, Thailand, Vietnam)

kitteli altisquamis Forel, 1900 (China, Myanmar)

kitteli laevis (Mayr, 1879) (Indonesia)

kitteli minor Forel, 1900 (India)

kitteli siemsseni Viehmeyer, 1922 (China)

kitteli transiens Forel, 1911 (Borneo, Indonesia, Malaysia)

kraepelini Forel, 1905 (Cambodia, China, Indonesia, Thailand, Vietnam)

kraepelini baccha Santschi, 1919 (Vietnam)

kraepelini esae Forel, 1912 (Indonesia)

kraepelini nero Forel, 1913 (Indonesia)

L
laeviterga Zhou et al., 2012 (China)

langi Wheeler, W.M., 1923 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela)

laozii Xu, 2000 (China)

lattkei Bharti & Wachkoo, 2013 (India)

lavavava Rakotonirina & Fisher, 2014 (Madagascar)

leiothorax Prins, 1965 (Mozambique)

letilae Mann, 1921 (Fiji)

linda Lattke, 2011 (Colombia)

linearis (Smith, F., 1858) (Brazil, French Guiana, Guyana, Suriname)

• Leptogenys dasygyna Wheeler, W.M., 1923

lohahela Rakotonirina & Fisher, 2014 (Madagascar)

longensis Forel, 1915 (Australia)

longiceps Santschi, 1914 (Ghana, Senegal)

longiscapa Donisthorpe, 1943 (India)

lucida Rakotonirina & Fisher, 2014 (Madagascar)

lucidula Emery, 1895 (Cambodia, China, India, Myanmar)

• Leptogenys huangdii Xu, 2000

luederwaldti Forel, 1913 (Brazil)

• Leptogenys anacleti Borgmeier, 1930

M
mactans Bolton, 1975 (South Africa)

magna Forel, 1900 (Australia)

malama Rakotonirina & Fisher, 2014 (Madagascar)

mangabe Rakotonirina & Fisher, 2014 (Madagascar)

manja Rakotonirina & Fisher, 2014 (Madagascar)

manni Wheeler, W.M., 1923 (United States)

manongarivo Rakotonirina & Fisher, 2014 (Madagascar)

mastax Bolton, 1975 (Ghana, Ivory Coast)

mavaca Lattke, 2011 (Brazil, Venezuela)

maxillosa (Smith, F., 1858) (Barbados, Brazil, Comoros, Cuba, Democratic Republic of Congo, Kenya, Lesser Antilles, Madagascar, Mauritius, Namibia, Oman, Philippines, Réunion, Saudi Arabia, Seychelles, South Africa, Zimbabwe)

• Formica vinsonnella Dufour, 1864

• Leptogenys cribrata Emery, 1895

• Leptogenys falcata Roger, 1861

maya Lattke, 2011 (Mexico)

mayotte Rakotonirina & Fisher, 2014 (Mayotte)

melena Lattke, 2011 (Venezuela)

mengzii Xu, 2000 (China)

meritans (Walker, 1859) (Sri Lanka)

microps Bolton, 1975 (Guinea, Ivory Coast)

minima Lattke, 2011 (Brazil)

mjobergi Forel, 1915 (Australia)

modiglianii Emery, 1900 (Indonesia)

moelleri (Bingham, 1903) (India)

montuosa Lattke, 2011 (Costa Rica, Panama)

mucronata Forel, 1893 (Lesser Antilles)

mucronata columbica Forel, 1901 (Colombia)

mutabilis (Smith, F., 1861) (Borneo, Indonesia, Malaysia, Philippines, Sulawesi, Thailand)

myops (Emery, 1887) (Borneo, Indonesia, Malaysia, Thailand)

N
namana Rakotonirina & Fisher, 2014 (Madagascar)

namoroka Rakotonirina & Fisher, 2014 (Madagascar)

navua Mann, 1921 (Fiji)

nebra Bolton, 1975 (Cameroun)

neutralis Forel, 1907 (Australia)

nigricans Lattke, 2011 (Brazil, Colombia, Ecuador)

nitens Donisthorpe, 1943 (New Guinea)

nuserra Bolton, 1975 (Ghana, Ivory Coast)

O
oaxaca Lattke, 2011 (Mexico)

occidentalis Bernard, 1953 (Guinea)

• Leptogenys ferrarii bernardi Baroni Urbani, 1971

• Leptogenys ferrarii sulcinodis Bernard, 1953

optica Viehmeyer, 1914 (New Guinea)

orchidioides Lattke, 2011 (Guatemala)

oresbia Wilson, 1958 (Solomon Islands)

oswaldi (Forel, 1891) (Madagascar)

P
pangui Xu, 2000 (China)

panops Lattke, 2011 (French Guiana)

papuana Emery, 1897 (New Guinea)

paraense Lattke, 2011

parvula Emery, 1900 (Borneo, Indonesia, Malaysia, Vietnam)

pavesii Emery, 1892 (Cameroun, Comoros, Kenya, Madagascar, Mayotte, Somalia, Sudan, United Republic of Tanzania)

• Leptogenys maxillosa sericeus Weber, 1942

peninsularis Mann, 1926 (Mexico)

peringueyi Forel, 1913 (South Africa)

peruana Lattke, 2011 (Peru)

peuqueti (André, 1887) (Bangladesh, China, India, Myanmar, Philippines, Sri Lanka, Vietnam)

• Leptogenys minchinii Forel, 1900

• Leptogenys watsoni Forel, 1900

phylloba Lattke, 2011 (Colombia, Ecuador)

pilaka Rakotonirina & Fisher, 2014 (Madagascar)

pinna Lattke, 2011 (Costa Rica)

piroskae Forel, 1910 (Eritrea, Ethiopia)

pittieri Lattke, 2011 (Venezuela)

podenzanai (Emery, 1895) (Australia)

• Prionogenys podenzanai malandensis Forel, 1915

pompiloides (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

princeps Bolton, 1975 (Ghana)

processionalis (Jerdon, 1851) (India, Sri Lanka, Vietnam)

• Lobopelta distinguenda andrei Emery, 1887

• Ponera ocellifera Roger, 1861

processionalis distinguenda (Emery, 1887) (Borneo, Indonesia, Malaysia)

pruinosa Forel, 1900 (Philippines, Sri Lanka)

pubiceps Emery, 1890 (Barbados, Colombia, Cuba, Puerto Rico, Venezuela)

pubiceps cubaensis Santschi, 1930 (Cuba, Greater Antilles)

pubiceps vincentensis Forel, 1901 (Lesser Antilles)

pucuna Lattke, 2011 (Ecuador)

punctata Emery, 1914 (New Caledonia)

punctaticeps Emery, 1890 (Costa Rica, Cuba, Dominican Republic, Greater Antilles, Grenada, Haiti, Lesser Antilles, Panama)

• Leptogenys ambigua Santschi, 1931

punctiventris (Mayr, 1879) (Cambodia, India, Philippines, Thailand)

purpurea (Emery, 1887) (New Guinea)

pusilla (Emery, 1890) (Colombia, Costa Rica, Mexico, Panama)

Q
quadrata Lattke, 2011 (Colombia, Costa Rica, Ecuador)

quiriguana Wheeler, W.M., 1923 (Belize, Guatemala, Mexico)

quirozi Lattke, 2011 (Mexico)

R
rabebe Rakotonirina & Fisher, 2014 (Madagascar)

rabesoni Rakotonirina & Fisher, 2014 (Madagascar)

ralipra Rakotonirina & Fisher, 2014 (Madagascar)

rasila Lattke, 2011 (Ecuador)

ravida Bolton, 1975 (Democratic Republic of Congo)

reggae Lattke, 2011 (Jamaica)

regis Bolton, 1975 (Kenya, United Republic of Tanzania)

ridens Forel, 1910 (Madagascar)

ritae Forel, 1899 (Colombia, Ecuador, Panama, Trinidad and Tobago, Venezuela)

• Leptogenys pusilla panamana Wheeler, W.M., 1923

• Leptogenys venatrix Forel, 1899

roberti Forel, 1900 (Bangladesh, India)

roberti coonoorensis Forel, 1900 (India)

rouxi (Emery, 1914) (New Caledonia)

rufa Mann, 1922 (Belize, Honduras, Mexico)

rufida Zhou et al., 2012 (China)

rugosopunctata Karavaiev, 1925 (Indonesia)

S
sagaris Wilson, 1958 (New Caledonia)

sahamalaza Rakotonirina & Fisher, 2014 (Madagascar)

santacruzi Lattke, 2011 (Ecuador)

saussurei (Forel, 1891) (Madagascar)

schwabi Forel, 1913 (South Africa, Zimbabwe)

serrata Lattke, 2011 (Colombia, Panama)

sianka Lattke, 2011 (Mexico)

sjostedti Forel, 1915 (Australia)

socorda Lattke, 2011 (Colombia)

sonora Lattke, 2011 (Mexico)

spandax Bolton, 1975 (Ghana)

stenocheilos (Jerdon, 1851) (India)

sterops Bolton, 1975 (Ivory Coast)

strator Bolton, 1975 (Democratic Republic of Congo)

strena Zhou, 2001 (China)

striatidens Bolton, 1975 (Cameroun)

stuhlmanni Mayr, 1893 (Comoros, Eritrea, Kenya, Mozambique, Sudan, United Republic of Tanzania)

• Leptogenys comorensis Forel, 1907

stygia Bolton, 1975 (Nigeria)

suarensis Emery, 1895 (Madagascar)

sulcinoda (André, 1892) (Gabon)

sunzii Xu & He, 2015

T
tama Lattke, 2011 (Colombia)

tatsimo Rakotonirina & Fisher, 2014 (Madagascar)

terroni Bolton, 1975 (Cameroun)

testacea (Donisthorpe, 1948) (Ghana)

tiobil Lattke, 2011 (Venezuela)

titan Bolton, 1975 (Cameroun)

toeraniva Rakotonirina & Fisher, 2014 (Madagascar)

toxeres Lattke, 2011 (Costa Rica)

transitionis Bharti & Wachkoo, 2013 (India)

tricosa Taylor, 1969 (Australia)

• Dorylozelus mjobergi Forel, 1915

triloba Emery, 1901 (New Guinea)

trilobata Santschi, 1924 (Cameroun, Democratic Republic of Congo)

truncata Mann, 1919 (Solomon Islands)

truncatirostris Forel, 1897 (Madagascar, Mayotte)

tsingy Rakotonirina & Fisher, 2014 (Madagascar)

turneri Forel, 1900 (Australia)

U
unistimulosa Roger, 1863 (Brazil, Colombia, Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago, Venezuela)

• Leptogenys unistimulosa bahiana Santschi, 1929

• Leptogenys unistimulosa trinidadensis Forel, 1901

V
variabilis Rakotonirina & Fisher, 2014 (Madagascar)

varicosa Stitz, 1925 (Philippines)

vatovavy Rakotonirina & Fisher, 2014 (Madagascar)

vindicis Bolton, 1975 (Cameroun)

violacea Donisthorpe, 1942 (New Guinea)

vitiensis Mann, 1921 (Fiji)

vitsy Rakotonirina & Fisher, 2014 (Madagascar)

voeltzkowi Forel, 1897 (Madagascar)

vogeli Borgmeier, 1933 (Brazil)

volcanica Lattke, 2011 (Colombia, Costa Rica, Panama)

W
wheeleri Forel, 1901 (Mexico, Panama)

Y
yandii Xu & He, 2015

yerburyi Forel, 1900 (Sri Lanka)

yocota Lattke, 2011 (Honduras)

Z
zapyxis Bolton, 1975 (Ghana)

zhuangzii Xu, 2000 (China)

zohy Rakotonirina & Fisher, 2014 (Madagascar)

=Fossil Species=

L
lacerata Zhang, 1989