Camponotus novogranadensis

This species has recently been found, as a newly discovered introduction, in Estero, Lee County, Florida (Deyrup and Belmont 2013).

Identification
In the field C. novogranadensis is easily mistaken for Camponotus planatus. Both species are dark-colored, non-glossy, and small (majors are about 5mm long). Camponotus novogranadensis is black, with brown or yellowish brown antennae, clypeus, and the sides of the face above the mandibles. Camponotus planatus is usually bicolored, dark red with a black gaster, but occasionally completely black. Under the microscope these two species are conspicuously different. The clypeus of C. novogranadensis has a strong, sharp, median ridge, absent in C. planatus. The mesosoma of C. novogranadensis is covered with small, semiappressed hairs and large, sparse, curved, proclinate hairs, that of C. planatus moderately densely covered with short, sub-erect hairs. (Deyrup and Belmont 2013)

Distribution based on Regional Taxon Lists
Nearctic Region: United States. Neotropical Region: Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago.

Biology
Nests are in dead wood or hollow stems, usually in disturbed areas (Vasconcelos 1999, Longino 2002; Sanabria-Blandon and Chacon de Ulloa 2009; Ribas et al. 2012). It also occurs in the canopy of primary forest (Longino 2002). This species is listed as an indicator ant species of disturbed habitats (Ribas et al. 2012). In a study of effects of disturbance on ants in central Amazonia, Vasconcelos (1999) found ground-foraging C. novogranadensis in no mature forest plots, in 25% of plots in both old regrowth and new regrowth, and in 100% of plots in abandoned pasture. The diet of C. novogranadensis includes honeydew from Membracidae and Aetalionidae (Letourneau & Choe 1987), and nectar from extrafloral nectaries of orchids, which this species aggressively defends (Damon & Perez-Soriano 2005), and animal protein (in baits) (Vasconcelos 1999). Schmid et al. (2014) found this ant nesting in infructescences (the stem and remains of buds and fruits above the level of the water reservoir in the rosette) of the bromeliad Vriesea friburgensis on Santa Catarina Island, Brazil. De Oliveira et al. (2015) found a colony of C. novogranadensis opportunistically nesting in a Cecropia saxatilis tree (southwest Bahia, Brazil).

Nomenclature

 * . Camponotus novogranadensis Mayr, 1870a: 380 (s.w.) COLOMBIA (“New Granada”).
 * [Misspelled as novogrenadensis by Wheeler, W.M. 1916c: 14, Wheeler, W.M. 1942: 257.]
 * Emery, 1906c: 191 (q.); Wheeler, G.C. & Wheeler, J. 1953e: 192 (l.).
 * Combination in C. (Myrmamblys): Forel, 1912i: 90;
 * combination in C. (Neomyrmamblys): Santschi, 1921f: 311;
 * combination in C. (Myrmaphaenus): Emery, 1925b: 156.
 * Status as species: Forel, 1879a: 87; Mayr, 1884: 30; Emery, 1888c: 364; Emery, 1890b: 56; Dalla Torre, 1893: 245; Emery, 1894k: 62; Forel, 1895b: 103; Emery, 1896d: 375 (in list); Forel, 1899c: 149; Forel, 1902b: 171; Emery, 1903: 72; Emery, 1906c: 191; Forel, 1906d: 249; Forel, 1908b: 71; Forel, 1911c: 310; Forel, 1912i: 81; Donisthorpe, 1915d: 349; Wheeler, W.M. 1916c: 14; Mann, 1916: 479; Luederwaldt, 1918: 49; Mann, 1922: 53; Wheeler, W.M. 1922c: 16; Wheeler, W.M. 1923a: 5; Emery, 1925b: 156; Borgmeier, 1927c: 153; Donisthorpe, 1927b: 404; Borgmeier, 1934: 111; Wheeler, W.M. 1942: 257; Kempf, 1961b: 523; Kempf, 1970b: 340; Kempf, 1972a: 46; Bolton, 1995b: 114; Wild, 2007b: 28; Branstetter & Sáenz, 2012: 256; Bezděčková, et al. 2015: 113; Deyrup, 2017: 195; Mackay & Mackay, 2019: 764.
 * Current subspecies: nominal plus modestior.

References based on Global Ant Biodiversity Informatics

 * Adams B. J., S. A. Schnitzer, and S. P. Yanoviak. 2019. Connectivity explains local ant community structure in a Neotropical forest canopy: a large-scale experimental approach. Ecology 100(6): e02673.
 * Ahuatzin D. A., E. J. Corro, A. Aguirre Jaimes, J. E. Valenzuela Gonzalez, R. Machado Feitosa, M. Cezar Ribeiro, J. Carlos Lopez Acosta, R. Coates, W. Dattilo. 2019. Forest cover drives leaf litter ant diversity in primary rainforest remnants within human-modified tropical landscapes. Biodiversity and Conservation 28(5): 1091-1107.
 * Basset Y., L. Cizek, P. Cuenoud, R. K. Didham, F. Guilhaumon, O. Missa, V. Novotny, F. Odegaards, T. Roslin, J. Schmidl et al. 2012. Arthropod diversity in a tropical forest. Science 338(6113): 1481-1484.
 * Benitez-Malvido, J., W. Dattilo, A. P. Martinez-Falcon, C. Duran-Barron, J. Valenzuela, S. Lopez, and R. Lombera. 2016. The multiple impacts of tropical forest fragmentation on arthropod biodiversity and on their patterns of interactions with host plants. Plos One 11: e0146461.
 * Buren W. F. 1983. Artificial faunal replacement for imported fire ant control. Florida Entomologist 66: 93-100.
 * Carvalho K. S., M. A. F. Carneiro, I. S. Nascimento, A. K. Saha, and E. M. Bruna. Lower ant diversity on earth mounds in a semi-arid ecosystem: natural variation or a sign of degradation? Sociobiology 63(4): 1022-1030.
 * Castano-Meneses, G., M. Vasquez-Bolanos, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha, and I. Alcala-Martinez. 2015. Avances de Formicidae de Mexico. Universidad Nacional Autonoma de Mexico.
 * Coelho M. S., G. W. Fernandes, J. C. Santos, and J. H. C. Delabie. 2009. Ants (Hymenoptera: Formicidae) as bioindicators of land restoration in a Brazilian Atlantic forest fragment. Sociobiology 54(1): 51-63.
 * Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
 * Davidson D. W., S. C. Cook, R. R. Snelling and T. H. Chua. 2003. Explaining the Abundance of Ants in Lowland Tropical Rainforest Canopies. Science 300: 969-972.
 * Dejean, A., S. Durou, I. Olmsted, R.R. Snelling and J. Orivel. 2003. Nest Site Selection by Ants in a Flooded Mexican Mangrove, with Special Reference to the Epiphytic Orchid Myrmecophila christinae. Journal of Tropical Ecology 19(3) :325-331
 * Dejean, A., S. Durou, I. Olmsted, R.R. Snelling and J. Orivel. 2003. Nest Site Selection by Ants in a Flooded Mexican Mangrove, with Special Reference to the Epiphytic Orchid Myrmecophila christinae. Journal of Tropical Ecology 19(3):325-331
 * Del Toro, I., M. Vázquez, W.P. Mackay, P. Rojas and R. Zapata-Mata. Hormigas (Hymenoptera: Formicidae) de Tabasco: explorando la diversidad de la mirmecofauna en las selvas tropicales de baja altitud. Dugesiana 16(1):1-14.
 * Deyrup M., and R. A. Belmont. 2013. First record of a Florida population of the Neotropical carpenter ant Camponotus novogranadensis (Hymenoptera: Formicidae). Florida Entomologist 96(1): 283-285.
 * Emery C. 1894. Estudios sobre las hormigas de Costa Rica. Anales del Museo Nacional de Costa Rica 1888-1889: 45-64.
 * Emery C. 1896. Formiche raccolte dal dott. E. Festa nei pressi del golfo di Darien. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 11(229): 1-4.
 * Emery C. 1903. Intorno ad alcune specie di Camponotus dell'America Meridionale. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna (n.s.)7: 62-81.
 * Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
 * Escalante Gutiérrez J. A. 1993. Especies de hormigas conocidas del Perú (Hymenoptera: Formicidae). Revista Peruana de Entomología 34:1-13.
 * Fernandes I., and J. de Souza. 2018. Dataset of long-term monitoring of ground-dwelling ants (Hymenoptera: Formicidae) in the influence areas of a hydroelectric power plant on the Madeira River in the Amazon Basin. Biodiversity Data Journal 6: e24375.
 * Fernandes T. T., R. R. Silva, D. Rodrigues de Souza-Campana, O. Guilherme Morais da Silva, and M. Santina de Castro Morini. 2019. Winged ants (Hymenoptera: Formicidae) presence in twigs on the leaf litter of Atlantic Forest. Biota Neotropica 19(3): http://dx.doi.org/10.1590/1676-0611-bn-2018-0694
 * Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
 * Forel A. 1902. Quatre notices myrmécologiques. Annales de la Société Entomologique de Belgique 46: 170-182.
 * Forel A. 1908. Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles 44: 35-72.
 * Forel A. 1911. Ameisen des Herrn Prof. v. Ihering aus Brasilien (Sao Paulo usw.) nebst einigen anderen aus Südamerika und Afrika (Hym.). Deutsche Entomologische Zeitschrift 1911: 285-312.
 * Forel A. 1912. Formicides néotropiques. Part VI. 5me sous-famille Camponotinae Forel. Mémoires de la Société Entomologique de Belgique. 20: 59-92.
 * Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
 * Gomes E. C. F., G. T. Ribeiro, T. M. S. Souza, and L. Sousa-Souto. 2014. Ant assemblages (Hymenoptera: Formicidae) in three different stages of forest regeneration in a fragment of Atlantic Forest in Sergipe, Brazil. Sociobiology 61(3): 250-257.
 * Gove, A. D., J. D. Majer, and V. Rico-Gray. 2009. Ant assemblages in isolated trees are more sensitive to species loss and replacement than their woodland counterparts. Basic and Applied Ecology 10: 187-195.
 * Gutierrez-Martínez P. R., and D. Arauz Naranjo. 2016. Hormigas (Hymenoptera: Formicidae) en el bosque seco del Parque Nacional Diria, Costa Rica. Cuadernos de Investigación UNED 8(2): 185-188.
 * INBio Collection (via Gbif)
 * Kempf W. W. 1961. A survey of the ants of the soil fauna in Surinam (Hymenoptera: Formicidae). Studia Entomologica 4: 481-524.
 * Kempf W. W. 1978. A preliminary zoogeographical analysis of a regional ant fauna in Latin America. 114. Studia Entomologica 20: 43-62.
 * Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
 * Kost, C., and M. Heil. 2005. Increased availability of extrafloral nectar reduces herbivory in Lima bean plants (Phaseolus lunatus, Fabaceae). Basic and Applied Ecology 6: 237-248.
 * Landero-Torres I., M. A. Garcia-Martinez, M. E. Galindo-Tovar, O. R. Leyva-Ovalle, H. E. Lee-Espinosa, J. Murguia-Gonzalez, and J. Negrin-Ruiz. 2014. An ornamental Heliconias crop as a reservoir of the native myrmecofauna: a case of tropical horticulture in Central Veracruz, Mexico. Southwestern Entomologist 39(1): 135-146.
 * Leponce M., J. H. C. Delabie, J. Orivel, J. Jacquemin, M. Calvo Martin, and A. Dejean. 2019. Tree-dwelling ant survey (Hymenoptera, Formicidae) in Mitaraka, French Guiana, in Touroult J. (ed.), “Our Planet Reviewed” 2015 large-scale biotic survey in Mitaraka, French Guiana. Zoosystema 41 (10): 163-179.
 * Longino J. T. 2013. Ants of Honduras. Consulted on 18 Jan 2013. https://sites.google.com/site/longinollama/reports/ants-of-honduras
 * Longino J. et al. ADMAC project. Accessed on March 24th 2017 at https://sites.google.com/site/admacsite/
 * Longino, J.T. 2010. Personal Communication. Longino Collection Database
 * Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
 * Lutinski J. A., B. C. Lopes, and A. B. B.de Morais. 2013. Diversidade de formigas urbanas (Hymenoptera: Formicidae) de dez cidades do sul do Brasil. Biota Neotrop. 13(3): 332-342.
 * Maes, J.-M. and W.P. MacKay. 1993. Catalogo de las hormigas (Hymenoptera: Formicidae) de Nicaragua. Revista Nicaraguense de Entomologia 23.
 * Mann W. M. 1922. Ants from Honduras and Guatemala. Proceedings of the United States National Museum 61: 1-54.
 * Maravalhas J., and H. L. Vasconcelos. 2014. Revisiting the pyrodiversitybiodiversity hypothesis: long-term fire regimes and the structure of ant communities in a Neotropical savanna hotspot. Journal of Applied Ecology 51: 1661-1668.
 * Marinho C. G. S., R. Zanetti, J. H. C. Delabie, M. N. Schlindwein, and L. de S. Ramos. 2002. Ant (Hymenoptera: Formicidae) Diversity in Eucalyptus (Myrtaceae) Plantations and Cerrado Litter in Minas Gerais, Brazil. Neotropical Entomology 31(2): 187-195.
 * Mentone T. O., E. A. Diniz, C. B. Munhae, O. C. Bueno, and M. S. C. Morini. 2011. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop. 11(2): http://www.biotaneotropica.org.br/v11n2/en/abstract?inventory+bn00511022011.
 * Mentone T.O., M. S. C. Morini, L. Souza, and S. M. P. Braga. 2009. Hymenoptera communities in an agroecosyste using direct seedling in Southeastern Brazil. Sociobiology 53(2B): 473-486.
 * Miranda P. N., F. B. Baccaro, E. F. Morato, M. A. Oliveira. J. H. C. Delabie. 2017. Limited effects of low-intensity forest management on ant assemblages in southwestern Amazonian forests. Biodivers. Conserv. DOI 10.1007/s10531-017-1368-y
 * Mirmecofauna de la reserva ecologica de San Felipe Bacalar
 * Munhae C. B., Z. A. F. N. Bueno, M. S. C. Morini, and R. R. Silva. 2009. Composition of the Ant Fauna (Hymenoptera: Formicidae) in Public Squares in Southern Brazil. Sociobiology 53(2B): 455-472.
 * Nascimento Santos M., J. H. C. Delabie, and J. M. Queiroz. 2019. Biodiversity conservation in urban parks: a study of ground-dwelling ants (Hymenoptera: Formicidae) in Rio de Janeiro City. Urban Ecosystems https://doi.org/10.1007/s11252-019-00872-8
 * Neves F. S., R. F. Braga, M. M. do Espirito-Santo, J. H. C. Delabie, G. Wilson Fernandes, and G. A. Sanchez-Azofeifa. 2010. Diversity of Arboreal Ants In a Brazilian Tropical Dry Forest: Effects Of Seasonality and Successional Stage. Sociobiology 56(1): 1-18.
 * Novais, S. M., W. D. DaRocha, N. Calderon-Cortes, and M. Quesada. 2017. Wood-boring beetles promote ant nest cavities: extended effects of a twig-girdler ecosystem engineer. Basic and Applied Ecology 24: 53-59.
 * Oliveira Mentone T. de, E. A. Diniz, C. de Bortoli Munhae, O. Correa Bueno and M. S. de Castro Morini. 2012. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop 11(2): 237-246.
 * Oliveira R. F., L. C. de Almeida, D. R. de Suza, C. Bortoli Munhae, O. C. Bueno, and M. S. de Castro Morini. 2012. Ant diversity (Hymenoptera: Formicidae) and predation by ants on the different stages of the sugarcane borer life cycle Diatraea saccharalis (Lepidoptera: Crambidae). Eur. J. Entomol. 109: 381387.
 * Orsolon-Souza G., C. E. L. Esberard, A. J. Mayhe-Nunes, A. B. Vargas, S. Veiga-Ferreira, and E. Folly-Ramos. 2011. Comparison between Winklers extractor and pitfall traps to estimate leaf litter ants richness (Formicidae) at a rainforest site in southest Brazil. Braz. J. Biol. 71(4): 873-880.
 * Ottonetti L., L. Tucci, F. Frizzi, G. Chelazzi, and G. Santini. 2010. Changes in ground-foraging ant assemblages along a disturbance gradient in a tropical agricultural landscape. Ethology Ecology & Evolution 22: 7386.
 * Pereira M. C., J. H. C. Delabie, Y. R. Suarez, and W. F. Antonialli Junior. 2013. Spatial connectivity of aquatic macrophytes and flood cycle influence species richness of an ant community of a Brazilian floodplain. Sociobiology 60(1): 41-49.
 * Peres M. C. L. 2012. Efeito de clareiras naturais sobre as assembleias de formigas (Formicidae) e aranhas (Araneae) num trecho de Mata Atlantica. PhD thesis, Universidade Federal da Bahia, 111 pages.
 * Perfecto I., and J. Vandermeer. 1996. Microclimatic changes and the indirect loss of ant diversity in a tropical agroecosystem. Oecologia 108: 577-582.
 * Philpott, S. M. 2006. Ant patchiness: a spatially quantitative test in coffee agroecosystems. Naturwissenschaften 93: 386-392.
 * Philpott, S.M., P. Bichier, R. Rice, and R. Greenberg. 2007. Field testing ecological and economic benefits of coffee certification programs. Conservation Biology 21: 975-985.
 * Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
 * Radoszkowsky O. 1884. Fourmis de Cayenne Française. Trudy Russkago Entomologicheskago Obshchestva 18: 30-39.
 * Ramos L. S., R. Z. B. Filho, J. H. C. Delabie, S. Lacau, M. F. S. dos Santos, I. C. do Nascimento, and C. G. S. Marinho. 2003. Ant communities (Hymenoptera: Formicidae) of the leaf-litter in cerrado stricto sensu areas in Minas Gerais, Brazil. Lundiana 4(2): 95-102.
 * Ramos L. de S., C. G. S. Marinho, R. Zanetti, and J. H. C. Delabie. 2003. Impacto de iscas formicidas granuladas sobre a mirmecofauna não-alvo em eucaliptais segundo duas formas de aplicacação / Impact of formicid granulated baits on non-target ants in eucalyptus plantations according to two forms of application. Neotropical Entomology 32(2): 231-237.
 * Reynoso-Campos J. J., J. A. Rodriguez-Garza, and M. Vasquez-Bolanos. 2015. Hormigas (Hymenoptera: Formicidae) de la Isla Cozumel, Quintana Roo, Mexico (pp. 27-39). En: Castaño Meneses G., M. Vásquez-Bolaños, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha e I. Alcalá-Martínez (Coords.). Avances de Formicidae de  México.  UNAM,  Universiad  de  Guadalajara, Guadalajara, Jalisco.
 * Ribas C. R., J. H. Schoereder, M. Pic, and S. M. Soares. 2003. Tree heterogeneity, resource availability, and larger scale processes regulating arboreal ant species richness. Austral Ecology 28(3): 305-314.
 * Santos P. P., A. Vasconcelos, B. Jahyny, and J. H. C. Delabie. 2010. Ant fauna (Hymenoptera, Formicidae) associated to arboreal nests of Nasutitermes spp. (Isoptera, Termitidae) in a cacao plantation in southeastern Bahia, Brazil. Revista Brasileira de Entomologia 54(3): 450-454.
 * Schmid V. S., S. Langner, J. Steiner, and A. Zillikens. 2014. Inflorescences of the Bromeliad Vriesea friburgensis as Nest Sites and Food Resources for Ants and Other Arthropods in Brazil. Psyche http://dx.doi.org/10.1155/2014/396095
 * Schmidt F. A., and R. R. C. Solar. Is it important to collect hypogaeic ants? How to collect them? Biológico, São Paulo 69(2): 267-270.
 * Schmidt, F.A. and R.R.C Solar. 2010. Hypogaeic pitfall traps: methodological advances and remarks to improve the sampling of a hidden ant fauna. Insectes Sociaux 57:261-266.
 * Schoereder J. H., T. G. Sobrinho, M. S. Madureira, C. R. Ribas, and P. S. Oliveira. 2010. The arboreal ant community visiting extrafloral nectaries in the Neotropical cerrado savanna. Terrestrial Arthropod Reviews 3: 3-27.
 * Silva F. H. O., J. H. C. Delabie, G. B. dos Santos, E. Meurer, and M. I. Marques. 2013. Mini-Winkler Extractor and Pitfall Trap as Complementary Methods to Sample Formicidae. Neotrop Entomol 42: 351358.
 * Siqueira de Castro F., A. B. Gontijo, P. de Tarso Amorim Castro, and S. Pontes Ribeiro. 2012. Annual and Seasonal Changes in the Structure of Litter-Dwelling Ant Assemblages (Hymenoptera: Formicidae) in Atlantic Semideciduous Forests. Psyche doi:10.1155/2012/959715
 * Siqueira de Castro F., A. B. Gontijo, W. Duarte da Rocha, and S. Pontes Ribeiro. 2011. As comunidades de formigas de serapilheira nas florestas semidecíduas do Parque Estadual do Rio Doce, Minas Gerais. MG.BIOTA, Belo Horizonte 3(5): 5-24.
 * Smith M. A., W. Hallwachs, D. H. Janzen. 2014. Diversity and phylogenetic community structure of ants along a Costa Rican elevational gradient. Ecography 37(8): 720-731.
 * Sobrinho T., J. H. Schoereder, C. F. Sperber, and M. S. Madureira. 2003. Does fragmentation alter species composition in ant communities (Hymenoptera: Formicidae)? Sociobiology 42(2): 329-342.
 * Suguituru S. S., M. Santina de Castro Morini, R. M. Feitosa, and R. Rosa da Silva. 2015. Formigas do Alto Tiete. Canal 6 Editora 458 pages
 * Ulyssea M. A., and C. R. F. Brandao. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217224.
 * Ulysséa M. A., C. R. F. Brandão. 2013. Ant species (Hymenoptera, Formicidae) from the seasonally dry tropical forest of northeastern Brazil: a compilation from field surveys in Bahia and literature records. Revista Brasileira de Entomologia 57(2): 217-224.
 * Vasconcelos, H.L. and J.M.S. Vilhena. 2006. Species turnover and vertical partitioning of ant assemblages in the Brazilian Amazon: A comparison of forests and savannas. Biotropica 38(1):100-106.
 * Vasconcelos, H.L., J.M.S. Vilhena, W.E. Magnusson and A.L.K.M. Albernaz. 2006. Long-term effects of forest fragmentation on Amazonian ant communities. Journal of Biogeography 33:1348-1356
 * Vasquez-Bolanos M. 2011. Checklist of the ants (Hymenoptera: Formicidae) from Mexico. Dugesiana 18(1): 95-133.
 * Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
 * Wheeler W. M. 1916. Ants collected in British Guiana by the expedition of the American Museum of Natural History during 1911. Bulletin of the American Museum of Natural History 35: 1-14.
 * Wheeler W. M. 1922. The ants of Trinidad. American Museum Novitates 45: 1-16.
 * Wheeler W. M. 1942. Studies of Neotropical ant-plants and their ants. Bulletin of the Museum of Comparative Zoology 90: 1-262.
 * de Souza D. R., S. G. dos Santos, C. de B. Munhae, and M. S. de C. Morini. 2012. Diversity of Epigeal Ants (Hymenoptera: Formicidae) in Urban Areas of Alto Tietê. Sociobiology 59(3): 703-117.