Kenya



The following 462 valid species and subspecies of ants are known to occur in Kenya. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Kenys ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Kenya Ants web page. Please advise if any corrections are warranted.

A

 * Acropyga silvestrii Emery, 1915


 * Aenictus alluaudi Santschi, 1910


 * Aenictus crucifer Santschi, 1914


 * Aenictus decolor (Mayr, 1879)


 * Aenictus eugenii Emery, 1895


 * Aenictus fuscovarius Gerstacker, 1859


 * Aenictus fuscovarius magrettii Emery, 1892


 * Aenictus hamifer Emery, 1896


 * Aenictus rotundatus Mayr, 1901


 * Aenictus rotundatus guineensis Santschi, 1924


 * Aenictus steindachneri Mayr, 1901


 * Aenictogiton species


 * Anochetus africanus (Mayr, 1865)


 * Anochetus bequaerti Forel, 1913


 * Anochetus katonae Forel, 1907


 * Anochetus obscuratus Santschi, 1911


 * Anochetus maynei Forel, 1913


 * Anochetus pellucidus Emery, 1902


 * Anochetus rothschildi Forel, 1907


 * Anochetus sedilloti Emery, 1884


 * Anochetus traegaordhi Mayr, 1904


 * Anoplolepis custodiens (Smith, F., 1858)


 * Atopomyrmex mocquerysi André, 1889


 * Axinidris acholli Weber, 1941


 * Axinidris bidens Shattuck, 1991


 * Axinidris hypoclinoides (Santschi, 1919)


 * Axinidris icipe Snelling, 2007


 * Axinidris kakamegensis Shattuck, 1991


 * Axinidris luhya Snelling, 2007


 * Axinidris murielae Shattuck, 1991


 * Axinidris okekai Snelling, 2007

B

 * Baracidris pilosa Fernández, 2003

C

 * Calyptomyrmex brunneus Arnold, 1948


 * Calyptomyrmex clavatus Weber, 1952


 * Calyptomyrmex duhun Bolton, 1981


 * Calyptomyrmex piripilis Santschi 1923


 * Calyptomyrmex tensus Bolton, 1981


 * Camponotus acvapimensis Mayr, 1862


 * Camponotus aegyptiacus Emery, 1915


 * Camponotus bayeri Forel, 1913


 * Camponotus bertolonii Emery, 1895


 * Camponotus bottegoi Emery, 1895


 * Camponotus braunsi Mayr, 1895


 * Camponotus braunsi erythromelus Emery, 1896


 * Camponotus brutus Forel, 1886


 * Camponotus chrysurus Gerstäcker, 1871


 * Camponotus cinctellus (Gerstäcker, 1859)


 * Camponotus erinaceus Gerstäcker, 1871


 * Camponotus etiolipes Bolton, 1995


 * Camponotus ferreri cavisquamis Santschi, 1926


 * Camponotus flavomarginatus Mayr, 1862


 * Camponotus foraminosus Forel, 1879


 * Camponotus foraminosus chrysogaster Emery, 1895


 * Camponotus galla Forel, 1894


 * Camponotus grandidieri Forel, 1886


 * Camponotus jeanneli Santschi, 1914


 * Camponotus kersteni Gerstäcker, 1871


 * Camponotus lilianae Forel,, 1913


 * Camponotus maculatus (Fabricius, 1782)


 * Camponotus mayri Forel, 1879


 * Camponotus niveosetosus Mayr, 1862


 * Camponotus orinobates Santschi, 1919


 * Camponotus perrisii Forel, 1886


 * Camponotus pompeius Forel, 1886


 * Camponotus posticus Santschi, 1926


 * Camponotus robecchii Emery, 1892


 * Camponotus robecchii troglodytes Forel, 1894


 * Camponotus rufoglaucus Jerdon, 1851


 * Camponotus schoutedeni Forel, 1911


 * Camponotus sericeus (Fabricius,, 1798)


 * Camponotus rubripes (Latreille, 1802)


 * Camponotus sericeus (Fabricius, 1798)


 * Camponotus solon Forel, 1886


 * Camponotus somalinus André, 1887


 * Camponotus somalinus pattensis Forel, 1907


 * Camponotus thraso negus Forel, 1907


 * Camponotus viri Santschi, 1915


 * Camponotus vividus Smith, F., 1858


 * Camponotus zimmermanni pansus Santschi, 1926


 * Cardiocondyla emeryi Forel, 1891


 * Cardiocondyla shuckardi Forel, 1891


 * Cardiocondyla wroughtonii (Forel, 1890)


 * Carebara alluaudi Santschi, 1913


 * Carebara distincta (Bolton & Belshaw, 1993)


 * Carebara elmenteitae (Patrizi, 1948)


 * Carebara jeanneli Santschi, 1913


 * Carebara octata Bolton & Belshaw, 1993


 * Carebara perpusilla concedens Santschi, 1914


 * Carebara pisinna Bolton & Belshaw, 1993


 * Carebara polita (Santschi, 1914)


 * Carebara punctata Karavaiev, 1931


 * Carebara rara (Bolton & Belshaw, 1993)


 * Carebara thoracica (Weber, 1950)


 * Carebara vidua Smith, F., 1858


 * Cataulacus brevisetosus Forel, 1901


 * Cataulacus egenus Santschi, 1911


 * Cataulacus intrudens (Smith, F., 1876)


 * Cataulacus jeanneli Santschi, 1914


 * Cataulacus kenyensis Santschi, 1935


 * Cataulacus lujae Forel, 1911


 * Cataulacus moloch Bolton, 1982


 * Cataulacus pullus Santschi, 1910


 * Cataulacus pygmaeus André, 1890


 * Cataulacus striativentris Santschi, 1924


 * Cataulacus traegaordhi Santchi, 1914


 * Cataulacus vorticus Bolton, 1974


 * Cataulacus wissmannii Forel, 1894


 * Centromyrmex sellaris Mayr, 1896


 * Cerapachys braytoni Weber, 1949


 * Cerapachys foreli (Santschi, 1914)


 * Cerapachys kenyensis Consani, 1951


 * Cerapachys lamborni Crawley, 1923


 * Cerapachys niger Santschi, 1914


 * Cerapachys nitidulus Brown, 1975


 * Cerapachys nkomoensis (Forel, 1916)


 * Cerapachys vespula (Weber, 1949)


 * Crematogaster aegyptiaca turkanensis Santschi, 1935


 * Crematogaster africana Mayr, 1895


 * Crematogaster amabilis Santschi, 1911


 * Crematogaster amabilis retiaria Santschi, 1933


 * Crematogaster bequaerti modica Santschi, 1926


 * Crematogaster bequaerti mutabilis Santschi, 1914


 * Crematogaster buchneri uasina Santschi, 1935


 * Crematogaster castanea Smith, 1858


 * Crematogaster castanea aquila Forel, 1907


 * Crematogaster castanea bruta Santschi, 1926


 * Crematogaster castanea busschodtsi Emery,, 1899


 * Crematogaster castanea ferruginea Forel, 1892


 * Crematogaster castanea inversa Forel, 1907


 * Crematogaster castanea mediorufa Forel, 1907


 * Crematogaster castanea tricolor Gerstackker, 1859


 * Crematogaster castanea ulugurensis Forel, 1911


 * Crematogaster chiarinii Emery, 1881


 * Crematogaster chiarinii bayeri Santschi, 1926


 * Crematogaster clariventris Mayr, 1895


 * Crematogaster concava Emery, 1899


 * Crematogaster excisa lacustris Santschi, 1914


 * Crematogaster foraminiceps Santschi, 1913


 * Crematogaster foraminiceps mirmillo Santschi, 1935


 * Crematogaster gabonensis André, 1889


 * Crematogaster gambiensis André, 1889


 * Crematogaster gerstaeckeri Dalla Torre, 1892


 * Crematogaster gerstaeckeri pulla Wheeler, W.M., 1922


 * Crematogaster gerstaeckeri sjostedti Mayr, 1907


 * Crematogaster jeanneli Santschi, 1914


 * Crematogaster kachelibae Arnold, 1954


 * Crematogaster litoralis Arnold, 1955


 * Crematogaster luctans Forel, 1907


 * Crematogaster mimosae Santschi, 1914


 * Crematogaster mimosae tenuipilis Santschi, 1937


 * Crematogaster phoenix Santschi, 1921


 * Crematogaster pseudinermis Viehmeyer, 1923


 * Crematogaster rectinota Forel, 1913


 * Crematogaster rugosa André, 1895


 * Crematogaster stadelmanni Mayr, 1895


 * Crematogaster striatula Emery, 1892


 * Crematogaster striatula horatii Santschi, 1937


 * Crematogaster vulcania Santschi, 1913


 * Crematogaster wellmani Forel, 1909


 * Cyphoidris spinosa Weber, 1952

D

 *  Decamorium decem (Forel, 1913)


 * Discothyrea mixta Brown, 1958


 * Discothyrea patrizii Weber, 1949


 * Dorylus affinis Shuckard, 1840


 * Dorylus affinis exilis Santschi, 1914


 * Dorylus atriceps (Smith, F., 1858)


 * Dorylus braunsi Emery, 1895


 * Dorylus brevipennis Emery, 1895


 * Dorylus buyssoni Santschi, 1910


 * Dorylus buyssoni conjugens Santschi, 1910


 * Dorylus conradti Emery, 1895


 * Dorylus depilis Emery,, 1895


 * Dorylus fimbriatus (Shuckard, 1840)


 * Dorylus fimbriatus laevipodex  Santschi, 1919


 * Dorylus fulvus (Westwood, 1839)


 * Dorylus fulvus badius Gerstacker, 1859


 * Dorylus helvolus Linnaeus, 1764


 * Dorylus kohli Wasmann, 1904


 * Dorylus moestus Emery, 1895


 * Dorylus nigricans burmeisteri (Shuckard, 1840)


 * Dorylus nigricans molestus (Gerstacker, 1859)


 * Dorylus stanleyi Forel, 1909


 * Dorylus wilverthi Emery, 1899

H

 * Hypoponera angustata (Santschi,, 1914)


 * Hypoponera cammerunensis (Santschi, 1914)


 * Hypoponera dulcis(Forel, 1907)


 * Hypoponera jeanneli Santschi, 1935


 * Hypoponera occidentalis (Bernard, 1953)


 * Hypoponera punctatissima (Roger, 1859)


 * Hypoponera ursa (Santschi, 1924)

L

 * Lepisiota affinis Santschi, 1937


 * Lepisiota canescens (Emery, 1897)


 * Lepisiota capensis (Mayr, 1862)


 * Lepisiota capensis anceps Forel, 1916


 * Lepisiota capensis guineensis (Mayr, 1902)


 * Lepisiota capensis simplex (Forel, 1892)


 * Lepisiota crinita (Mayr, 1895)


 * Lepisiota depressa Santschi, 1914


 * Lepisiota hirsuta Santschi, 1914


 * Lepisiota incisa (Forel, 1913)


 * Leptanilla species FHG 1


 * Leptogenys elegans Bolton, 1975


 * Leptogenys intermedia Emery, 19902


 * Leptogenys maxillosa (Smith, F., 1858)


 * Leptogenys pavesii Emery, 1892


 * Leptogenys regis Bolton, 1975


 * Leptogenys stuhlmanni Mayr, 1893

M

 * Melissotarsus emeryi Forel, 1907


 * Melissotarsus weissi Santschi, 1910


 * Meranoplus inermis Emery, 1895


 * Meranoplus magrettii André, 1884


 * Messor angularis Santschi, 1928


 * Messor cephalotes (Emery, 1895)


 * Messor ferreri Collingwood, 1993


 * Messor galla (Mayr, 1904)


 * Microdaceton exornatum Santschi, 1913


 * Microdaceton tibialis Weber, 1952


 * Monomorium afrum André, 1884


 * Monomorium arboreum Weber, 1943


 * Monomorium bicolor Emery, 1877


 * Monomorium cryptobium (Santschi, 1921)


 * Monomorium destructor (Jerdon,, 1851)


 * Monomorium draxocum Bolton, 1987


 * Monomorium elgonense (Santschi, 1935)


 * Monomorium exiguum Forel, 1894


 * Monomorium fasciatum Santschi, 1920


 * Monomorium hanneli Forel, 1907


 * Monomorium holothir Bolton, 1987


 * Monomorium iyenasu Bolton, 1987


 * Monomorium madecassum Forel, 1892


 * Monomorium malatu Bolton, 1987


 * Monomorium manir Bolton, 1987


 * Monomorium mictilis 	Forel, 1910


 * Monomorium mirandum Arnold, 1955


 * Monomorium modestum Santschi, 1914


 * Monomorium nitidiventre Emery, 1893


 * Monomorium osiridis Santschi, 1915


 * Monomorium pallidipes Forel, 1910


 * Monomorium paternum Bolton, 1987


 * Monomorium pharaonis (Linnaeus, 1758)


 * Monomorium rastractum Bolton, 1987


 * Monomorium robustior Forel, 1892


 * Monomorium rosae Santschi, 1920


 * Monomorium rotundatum Santschi, 1920


 * Monomorium spectrum Bolton, 1987


 * Monomorium speluncarum Santschi, 1914


 * Monomorium sutu Bolton, 1987


 * Monomorium vaguum Santschi, 1930


 * Monomorium valtinum Bolton, 1987


 * Monomorium westi Bolton, 1987


 * Myrmicaria distincta Santschi, 1925


 * Myrmicaria fusca consanguinea Santschi, 1925


 *  Myrmicaria natalensis eumenoides (Gerstacker, 1859)


 * Myrmicaria opaciventris Emery, 1893


 * Myrmicaria opaciventris congolensis Forel, 1909

N

 * Nesomyrmex angulatus Mayr, 1862


 * Nesomyrmex cataulacoides Snelling, 1992


 * Nesomyrmex evelynae (Forel, 1916)


 * Nesomyrmex innocens (Forel, 1913)


 * Nylanderia bourbonica (Forel, 1886)


 * Nylanderia weissi Santschi, 1910


 * Nylanderia zelotypa Santschi, 1915

O

 * Ocymyrmex nitidulus Emery, 1892


 * Ocymyrmex phraxus Bolton, 1981


 * Ocymyrmex robecchii Emery, 1892


 * Ocymyrmex weitzeckeri Emery, 1892


 * Odontomachus assiniensis Emery, 1892


 * Odontomachus troglodytes  Santschi, 1914


 * Oecophylla leakeyi Wilson & Taylor, 1964


 * Oecophylla longinoda (Latreille, 1802)


 * Oecophylla longinoda textor Wheeler, W.M., 1922

P

 * Pachycondyla ambigua André, 1890


 * Pachycondyla analis (Latreille, 1802)


 * Pachycondyla berthoudi (Forel, 1890)


 * Pachycondyla brunoi Forel, 1913


 * Pachycondyla caffraria (Smith,F., 1858)


 * Pachycondyla crassa (Emery, 1877)


 * Pachycondyla crassa  crassior Santschi, 1930


 * Pachycondyla crassa ilgii Forel, 1910


 * Pachycondyla darwinii africana (Forel, 1909)


 * Pachycondyla kenyensis Santschi, 1937


 * Pachycondyla kruegeri asina Santschi, 1912


 * Pachycondyla pachyderma Emery, 1901


 * Pachycondyla sennaarensis (Mayr, 1862)


 * Pachycondyla sharpi (Forel, 1901)


 * Pachycondyla sjostedti (Mayr, 1896)


 * Pachycondyla soror (Emery, 1899)


 * Pachycondyla subiridescens Wheeler, W.M., 1922


 * Pachycondyla suspecta (Santschi, 1914)


 * Pachycondyla talpa (André, 1890)


 * Pachycondyla tarsata (Fabricius, 1798)


 * Pachycondyla tarsata delagoensis (Emery, 1899)


 * Pachycondyla tarsata striatidens (Santschi, 1919)


 * Pachycondyla wroughtonii (Forel, 1901)


 * Paraparatrechina brunnella LaPolla & Cheng, 2010


 * Paraparatrechina umbranatis LaPolla & Cheng, 2010


 * Paraparatrechina weissi Santschi, 1910


 * Paratrechina longicornis (Latreille, 1802)


 * Phasmomyrmex wolfi (Emery, 1920)


 * Pheidole aurivillii Mayr, 1896


 * Pheidole crassinoda Emery, 1895


 * Pheidole crassinoda ruspolii Emery, 1897


 * Pheidole kitschneri Forel, 1910


 * Pheidole kohli Mayr, 1901


 * Pheidole megacephala (Fabricius, 1793)


 * Pheidole megacephala talpa Gerstäcker, 1871


 * Pheidole megacephala rotundata Forel, 1894


 * Pheidole nigeriensis Santschi, 1914


 * Pheidole prelli Forel, 1911


 * Pheidole pulchella Santschi, 1910


 * Pheidole punctulata Mayr, 1866


 * Pheidole scabriuscula Gerstäcker, 1871


 * Pheidole sculpturata Mayr, 1886


 * Pheidole sinaitica Mayr,, 1862


 * Pheidole speculifera Emery, 1877


 * Pheidole speculifera ascara Emery, 1901


 * Pheidole strator Bolton, 1975


 * Pheidole teneriffana Forel, 1893


 * Phrynoponera gabonensis (André, 1892)


 * Phrynoponera pulchella Bolton & Fisher, 2008


 * Plagiolepis alluaudi Emery, 1894


 * Plagiolepis brunni Mayr, 1895


 * Plagiolepis decora Santschi, 1914


 * Plagiolepis jouberti Forel, 1910


 * Plagiolepis pictipes Santschi, 1914


 * Platythyrea cribrinodis Gerstäcker, 1859


 * Platythyrea gracillima Wheeler, W.M., 1922


 * Platythyrea modesta Emery, 1899


 * Platythyrea schultzei Forel, 1910


 * Plectroctena cristata Emery, 1899


 * Plectroctena mandibularis Smith, F., 1858


 * Plectroctena strigosa Emery, 1899


 * Plectroctena subterranea Arnold, 1915


 * Polyrhachis gagates Smith, F., 1858


 * Polyrhachis latispina Emery, 1925


 * Polyrhachis lestoni Bolton, 1973


 * Polyrhachis medusa Forel, 1897


 * Polyrhachis militaris (Fabricius, 1782)


 * Polyrhachis schistacea Gerstäcker, 1859


 * Polyrhachis schlueteri Forel, 1886


 * Polyrhachis spinicola Forel, 1894


 * Polyrhachis viscosa Smith,F., 1858


 * Pristomyrmex africanus Karavaiev, 1931


 * Pristomyrmex cribrarius Arnold, 1926


 * Probolomyrmex guineensis Taylor, 1965


 * Proceratium toschii Consani, 1951


 * Psalidomyrmex procerus Emery, 1901


 * Pyramica bellatrix Bolton, 2000


 * Pyramica bequaerti (Santschi, 1923)


 * Pyramica cavinasis (Brown, 1950)


 * Pyramica concolor (Santschi, 1914)


 * Pyramica geoterra (Bolton, 1983)


 * Pyramica hensekta (Bolton, 1983)


 * Pyramica ludovici (Forel, 1904)


 * Pyramica lujae (Forel, 1902)


 * Pyramica mandibularis (Szabo, 1909)


 * Pyramica marginata (Santschi, 1914)


 * Pyramica maynei Forel, 1916


 * Pyramica minkara (Bolton, 1983)


 * Pyramica serrula (Santschi, 1910)


 * Pyramica simoni (Emery, 1895)


 * Pyramica sulumana (Bolton, 1983)


 * Pyramica tetragnatha Taylor, 1966


 * Pyramica thuvida (Bolton, 1983)


 * Pyramica truncatidens (Brown, 1950)

R

 * Rhoptromyrmex opacus Forel, 1909


 * Rhoptromyrmex transversinodis Mayr, 1901

S

 * Simopone conradti Emery, 1899


 * Simopone grandis Santschi, 1923


 * Simopone schoutedeni Santschi, 1923


 * Solenopsis africana Santschi, 1914


 * Solenopsis punctaticeps Mayr, 1865


 * Solenopsis punctaticeps cleptomana Santschi, 1914


 * Solenopsis saevissima itinerans Forel, 1911


 * Sphinctomyrmex species


 * Strumigenys arnoldi Forel, 1913


 * Strumigenys cacaoensis Bolton, 1983


 * Strumigenys dextra Brown, 1954


 * Strumigenys katapelta Bolton, 1983


 * Strumigenys korahyla Bolton, 1983


 * Strumigenys londianensis (Patrizi, 1946)


 * Strumigenys omalyx Bolton, 1983


 * Strumigenys petiolata Bernard, 1953


 * Strumigenys rukha Bolton, 1983


 * Strumigenys stygia Santschi, 1913


 * Strumigenys tetraphanes Brown, 1954

T

 * Tapinolepis bothae Forel, 1907


 * Tapinolepis macrophthalma (Arnold, 1962)


 * Tapinoma acuminatum Forel, 1907


 * Tapinoma luridum connexum Santschi, 1914


 * Tapinoma luridum sokolovi Karavaiev, 1931


 * Tapinoma luteum (Emery, 1895)


 * Tapinoma schultzei (Forel, 1910)


 * Technomyrmex andrei Emery, 1899


 * Technomyrmex camerunensis Emery, 1899


 * Technomyrmex ilgi (Forel, 1910)


 * Technomyrmex moerens Santschi, 1913


 * Technomyrmex nigriventris Santschi, 1910


 * Technomyrmex pallipes (Smith, F., 1876)


 * Technomyrmex vapidus Bolton, 2007


 * Technomyrmex vitiensis Mann, 1921


 * Technomyrmex voeltzkowi (Forel, 1907)


 * Temnothorax cenatus Bolton, 1982


 * Terataner bottegoi Emery, 1896


 * Tetramorium aculeatum (Mayr, 1866)


 * Tetramorium altivagans Santschi, 1914


 * Tetramorium boehmei Hita Garcia & Fischer, 2010


 * Tetramorium boltoni Hita Garcia & Fischer, 2010


 * Tetramorium brevispinosum (Stitz, 1910)


 * Tetramorium caldarium (Roger, 1857)


 * Tetramorium camerunense Mayr, 1895


 * Tetramorium candidum Bolton, 1980


 * Tetramorium cristatum Stitz, 1910


 * Tetramorium delagoense Forel, 1894


 * Tetramorium dogieli Karavaiev, 1931


 * Tetramorium dumezi Menozzi, 1942


 * Tetramorium edouardi Forel, 1894


 * Tetramorium eminii (Forel, 1894)


 * Tetramorium gazense Arnold, 1958


 * Tetramorium humbloti Forel, 1891


 * Tetramorium intextum Santschi, 1914


 * Tetramorium kakamega Hita Garcia, Fischer & Peters, 2010


 * Tetramorium kestrum Bolton, 1980


 * Tetramorium laevithorax Emery, 1895


 * Tetramorium longicorne Forel, 1907


 * Tetramorium lucayanum Wheeler, W.M., 1905


 * Tetramorium metactum Bolton, 1980


 * Tetramorium miserabile Santschi, 1918


 * Tetramorium nigrum Forel, 1907


 * Tetramorium nodiferum (Emery, 1901)


 * Tetramorium occidentale (Santschi,, 1916)


 * Tetramorium pauper Forel, 1907


 * Tetramorium pinnipilum Bolton, 1980


 * Tetramorium pullulum Santschi, 1924


 * Tetramorium quadridentatum Stitz, 1910


 * Tetramorium rothschildi (Forel, 1907)


 * Tetramorium sericeiventre Emery, 1877


 * Tetramorium setigerum Mayr, 1901


 * Tetramorium shilohense Forel, 1913


 * Tetramorium simillimum (Smith, F., 1851)


 * Tetramorium snellingi Hita-Garcia, 2010


 * Tetramorium subcoecum Forel, 1907


 * Tetramorium tanaense Hita Garcia, Fischer & Peters, 2010


 * Tetramorium tenebrosum Arnold, 1926


 * Tetramorium tersum Santschi, 1911


 * Tetramorium viticola Weber, 1943


 * Tetramorium weitzeckeri Emery, 1895


 * Tetramorium zambezium Santschi, 1939


 * Tetramorium zapyrum Bolton, 1980


 * Tetramorium zonacaciae (Weber, 1943)


 * Tetraponera ambigua (Emery, 1895)


 * Tetraponera capensis (Smith, F., 1858)


 * Tetraponera emacerata Santschi, 1911


 * Tetraponera mocquerysi (André, 1890)


 * Tetraponera mocquerysi biozellata Karavaiev, 1931


 * Tetraponera natalensis (Smith, F., 1858)


 * Tetraponera ophthalmica (Emery, 1912)


 * Tetraponera parops Ward, 2006


 * Tetraponera penzigi (Mayr, 1907)