Platythyrea parallela

Distribution based on Regional Taxon Lists
Australasian Region: Australia, New Caledonia. Indo-Australian Region: Borneo, Fiji, Guam, Indonesia, Malaysia, Micronesia (Federated States of), New Guinea, Northern Mariana Islands, Philippines, Samoa, Singapore, Solomon Islands. Malagasy Region: Seychelles. Oriental Region: Bangladesh, India, Laos, Sri Lanka, Thailand, Vietnam.

Nomenclature

 * . Ponera parallela Smith, F. 1859a: 143 (w.) INDONESIA (Aru Is).
 * Type-material: 2 syntype workers.
 * [Note: one worker is labelled as holotype and bears a Donisthorpe type-label. There is a second specimen in OXUM, from the same locality, and with the data label written in the same thickly-marked ink, but the method of mounting is not the same. Nevetheless, this specimen is also considered to belong to the type-series.]
 * Type-locality: Indonesia: Aru Is (A.R. Wallace).
 * Type-depository: OXUM.
 * [Misspelled as pararella by Phengsi, et al. 2018: 98 (in key).]
 * Donisthorpe, 1943d: 434 (m.); Wheeler, G.C. & Wheeler, J. 1976a: 42 (l.).
 * Combination in Platythyrea: Donisthorpe, 1932c: 454.
 * Unidenifiable taxon; incertae sedis in Ponerinae: Emery, 1911d: 116.
 * Status as species: Smith, F. 1861b: 44; Smith, F. 1863: 17; Mayr, 1863: 449; Smith, F. 1871a: 322; Dalla Torre, 1893: 41; Emery, 1901g: 566; Donisthorpe, 1932c: 454; Donisthorpe, 1943d: 434; Chapman & Capco, 1951: 48, 71 (combinations in Platythyrea, Ponera, respectively); Wilson, 1958a: 151; Wilson & Taylor, 1967: 20; Brown, 1975: 8, 47; Taylor & Brown, 1985: 37; Taylor, 1987a: 56; Dlussky, 1994: 53; Bolton, 1995b: 336; Dorow, 1996a: 77; Wetterer & Vargo, 2003: 416; Ghosh, et al. 2005: 9; Jaitrong & Nabhitabhata, 2005: 37; Clouse, 2007b: 260; Heterick, 2009: 136; Mohanraj, et al. 2010: 7; Pfeiffer, et al. 2011: 57; Sarnat & Economo, 2012: 161; Sarnat, et al. 2013: 73; Ramage, 2014: 154; Bharti, Guénard, et al. 2016: 53; Jaitrong, Guénard, et al. 2016: 43; Phengsi, et al. 2018: 98 (in key).
 * Senior synonym of annamita: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of aruana: Wilson, 1958a: 151; Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of australis: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 337.
 * Senior synonym of cephalotes: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 337.
 * Senior synonym of ceylonensis: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of coxalis: Wilson, 1958a: 151; Brown, 1975: 8; Bolton, 1995b: 337.
 * Senior synonym of cylindrica: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of egena: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of inconspicua: Brown, 1975: 8; Bolton, 1995b: 337.
 * Senior synonym of javana: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of pacifica: Wilson & Taylor, 1967: 20; Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of parva: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 337.
 * Senior synonym of philippinensis: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of pulchella: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of pusilla: Wilson, 1958a: 151; Brown, 1975: 8; Bolton, 1995b: 337; Ramage, 2014: 154.
 * Senior synonym of sechellensis: Brown, 1975: 9; Bolton, 1995b: 337; Dorow, 1996a: 77.
 * Senior synonym of subtilis: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of tritschleri: Brown, 1975: 9; Bolton, 1995b: 337.
 * Senior synonym of victoriae: Brown, 1975: 8; Bolton, 1995b: 337.
 * Senior synonym of wroughtonii: Brown, 1975: 8; Bolton, 1995b: 337.
 * Distribution:
 * Austral: Australia.
 * Malagasy: Seychelles.
 * Malesian: Fiji Is, French Polynesia, Indonesia (Ambon, Aru, Halmahera, Irian Jaya, Java, Kalimantan, Morotai, Sulawesi, Sumatra), Malaysia (Peninsula, Sabah, Sarawak), Micronesia, Papua New Guinea, Philippines (Guimaras, Luzon, Mindanao, Negros, Romblon), Samoa, Singapore, Solomon Is.
 * Oriental: India (+ Andaman Is), Laos, Sri Lanka, Thailand, Vietnam.
 * annamita. Platythyrea coxalis var. annamita Forel, 1911i: 215 (footnote) (w.) VIETNAM.
 * Type-material: holotype(?) worker.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: Vietnam (“Central Cochinchina”): Mois region (Dugas).
 * Type-depository: MHNG.
 * Subspecies of coxalis: Forel, 1911d: 378; Emery, 1911d: 29; Chapman & Capco, 1951: 47.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 336.
 * aruana. Platythyrea melancholica var. aruana Karavaiev, 1925a: 75 (in text) (w.) INDONESIA (Aru Is).
 * Type-material: 20 syntype workers.
 * Type-locality: Indonesia: Aru Is, Wammer I., Nr. 2646 (W. Karawaiew).
 * Type-depository: SIZK.
 * Junior synonym of parallela: Wilson, 1958a: 151; Brown, 1975: 9; Bolton, 1995b: 336.
 * australis. Platythyrea pusilla var. australis Forel, 1915b: 10 (w.) AUSTRALIA (Queensland).
 * Type-material: syntype workers (number not stated).
 * Type-localities: Australia: Queensland, Blackall Range (E. Mjöberg), and Mt Tambourine (E. Mjöberg).
 * Type-depositories: ANIC, MHNG, NHMB.
 * Wheeler, G.C. & Wheeler, J. 1971b: 1198 (l.).
 * Junior synonym of parallela: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 336.
 * cephalotes. Platythyrea cephalotes Viehmeyer, 1924a: 224 (w.) AUSTRALIA (New South Wales).
 * Type-material: holotype worker.
 * Type-locality: Australia: New South Wales, Trial Bay (H. Overbeck).
 * Type-depository: MNHU.
 * Junior synonym of parallela: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 336.
 * ceylonensis. Platythyrea ceylonensis Donisthorpe, 1941j: 202 (w.q.) SRI LANKA.
 * Type-material: 1 syntype worker, 1 syntype queen.
 * Type-locality: Sri Lanka (“Ceylon”): (no further data) (Thwaites).
 * Type-depository: BMNH.
 * Status as species: Chapman & Capco, 1951: 47.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 336.
 * coxalis. Platythyrea coxalis Emery, 1893e: 189 (footnote) (w.) WEST MALAYSIA.
 * Type-material: holotype worker.
 * Type-locality: Malaysia: Malacca peninsula (no collector’s name).
 * Type-depository: MSNG.
 * Viehmeyer, 1916a: 111 (q.).
 * Status as species: Emery, 1900d: 666; Emery, 1911d: 29; Viehmeyer, 1913: 145; Viehmeyer, 1916a: 111; Wheeler, W.M. & Chapman, 1925: 58; Chapman & Capco, 1951: 47; Baltazar, 1966: 240.
 * Junior synonym of parallela: Wilson, 1958a: 151; Brown, 1975: 8; Bolton, 1995b: 336.
 * cylindrica. Platythyrea coxalis var. cylindrica Forel, 1913k: 5, fig. A (q.) INDONESIA (Sumatra).
 * Type-material: holotype queen.
 * Type-locality: Indonesia: Sumatra, Tandjong Slamat, 1911-12 (H. v. Buttel-Reepen).
 * Type-depository: MNHU.
 * Subspecies of coxalis: Chapman & Capco, 1951: 48.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 336.
 * egena. Platythyrea pusilla var. egena Viehmeyer, 1916a: 111 (w.q.) SINGAPORE.
 * Type-material: syntype worker(s), syntype queen(s) (numbers not stated).
 * Type-locality: Singapore: (H. Overbeck).
 * Type-depository: MNHU.
 * Subspecies of pusilla: Chapman & Capco, 1951: 49.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 336.
 * inconspicua. Platythyrea inconspicua Mayr, 1870b: 961 (q.) SRI LANKA.
 * Type-material: holotype queen.
 * Type-locality: Sri Lanka (“Ceylon”) (Sichel)”.
 * [Note: type-locality recorded as South America by Emery, 1890a: 56, who said it was described “par erreur d’étiquette comme provenant de Ceylan”. Brown, 1975: 48, explains Emery’s error and establishes Sri Lanka as the correct locality.]
 * Type-depository: NHMW.
 * [Misspelled as conspicua by Mayr, 1884: 32.]
 * Emery, 1890a: 56 (w.).
 * Status as species: Emery, 1887b: 443; Dalla Torre, 1893: 27; Forel, 1899c: 4; Chapman & Capco, 1951: 48.
 * Junior synonym of punctata: Forel, 1901f: 335; Wheeler, W.M. 1908a: 123; Wheeler, W.M. 1908e: 401; Emery, 1911d: 30; Forel, 1912c: 35; Borgmeier, 1923: 53; Menozzi & Russo, 1930: 150; Borgmeier, 1937b: 219; Creighton, 1950a: 34; Kempf, 1964a: 141; Kempf, 1972a: 206.
 * Junior synonym of parallela: Brown, 1975: 8; Bolton, 1995b: 337.
 * javana. Platythyrea coxalis var. javana Forel, 1905c: 5 (w.q.) INDONESIA (Java).
 * Type-material: syntype worker(s) (number not stated), 1(?) syntype queen.
 * Type-locality: Indonesia: Java, Buitenzorg (= Bogor), 1904 (K. Kraepelin).
 * Type-depository: MHNG.
 * Subspecies of coxalis: Emery, 1911d: 29; Karavaiev, 1925a: 76; Chapman & Capco, 1951: 48.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337.
 * pacifica. Platythyrea pusilla var. pacifica Santschi, 1928a: 43 (w.) SAMOA (Upolu I.).
 * Type-material: 3 syntype workers.
 * Type-locality: Samoa: Upolu I., Apia, ix.1916 (Swale).
 * Type-depository: NHMB.
 * Subspecies of pusilla: Wheeler, W.M. 1935g: 9.
 * Junior synonym of parallela: Wilson & Taylor, 1967: 20; Brown, 1975: 9; Bolton, 1995b: 337.
 * parva. Platythyrea parva Crawley, 1915a: 133 (w.) AUSTRALIA (Northern Territory).
 * Type-material: syntype workers (number not stated).
 * Type-locality: Australia: Northern Territory, Darwin, 1.iv.1913 (G.F. Hill).
 * Type-depositories: BMNH, MHNG, MCZC.
 * Junior synonym of parallela: Brown, 1975: 9; Taylor, 1987a: 56; Bolton, 1995b: 337.
 * philippinensis. Platythyrea coxalis var. philippinensis Viehmeyer, 1916b: 283 (w.) PHILIPPINES (Luzon I.).
 * Type-material: syntype workers (number not stated).
 * Type-locality: Philippines: Luzon, Los Baños (C.F. Baker).
 * Type-depositories: MCZC, MNHU.
 * Subspecies of coxalis: Wheeler, W.M. & Chapman, 1925: 58; Chapman & Capco, 1951: 48; Baltazar, 1966: 240.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337.
 * pulchella. Platythyrea pulchella Santschi, 1928h: 121 (w.) INDONESIA (Sumatra).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Sumatra, Medan, 17.vii.1920 (J.B. Corporaal).
 * Type-depository: NHMB.
 * Status as species: Chapman & Capco, 1951: 48.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337.
 * punctata. Platythyrea cribrinodis var. punctata Arnold, 1915: 24 (w.) ZIMBABWE.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Zimbabwe: Victoria Falls (G. Arnold).
 * Type-depository: SAMC.
 * [Junior secondary homonym of Pachycondyla punctata Smith, F. 1858b: 108.]
 * Replacement name: Platythyrea cribrinodis var. brevidentata Wheeler, W.M. 1922a: 759.
 * pusilla. Platythyrea pusilla Emery, 1893e: 188 (w.) INDONESIA (Ambon I.).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Ambon (Bedot & Pictet).
 * Type-depository: MSNG.
 * Wheeler, W.M. 1919e: 50 (q.).
 * Status as species: Emery, 1911d: 29; Viehmeyer, 1916a: 111; Wheeler, W.M. 1919e: 50; Cheesman & Crawley, 1928: 516; Wheeler, W.M. 1932d: 14; Wheeler, W.M. 1935g: 9; Chapman & Capco, 1951: 48.
 * Junior synonym of parallela: Wilson, 1958a: 151; Brown, 1975: 8; Bolton, 1995b: 337; Ramage, 2014: 154.
 * sechellensis. Platythyrea wroughtoni subsp. sechellensis Forel, 1912k: 159 (w.) SEYCHELLES IS (Praslin I.).
 * Type-material: holotype(?) worker.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: Seychelles: Praslin I. (H. Scott).
 * Type-depository: BMNH.
 * Subspecies of wroughtonii: Wheeler, W.M. 1922a: 1007.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337; Dorow, 1996a: 77.
 * subtilis. Platythyrea subtilis Emery, 1900d: 666 (footnote) (w.) BORNEO (Indonesia: Kalimantan, Laut I.).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Borneo (Kalimantan), Pulo Laut (Doherty).
 * Type-depository: MSNG.
 * Status as species: Emery, 1911d: 29; Wheeler, W.M. 1919e: 50; Chapman & Capco, 1951: 49.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337.
 * tritschleri. Platythyrea coxalis var. tritschleri Forel, 1901f: 338 (w.) INDONESIA (Sumatra).
 * Type-material: syntype workers (number not stated).
 * Type-locality: none given.
 * [Note: Emery, 1911d: 29, gives the type-locality as Sumatra; Brown, 1975: 48, expands this to Sumatra, Indrapura.]
 * Type-depository: MHNG.
 * Subspecies of coxalis: Emery, 1911d: 29; Chapman & Capco, 1951: 48.
 * Junior synonym of parallela: Brown, 1975: 9; Bolton, 1995b: 337.
 * victoriae. Platythyrea wroughtoni r. victoriae Forel, 1900d: 316 (w.) INDIA (Karnataka, West Bengal; also, no state data, “Inde septentrionale”).
 * Type-material: syntype workers (number not stated).
 * Type-localities: India: Bangalore (Rothney), Barrackpore (Rothney), and “Inde septentrionale” (Wroughton).
 * Type-depositories: MCZC, MHNG.
 * Status as species: Rothney, 1903: 96; Bingham, 1903: 75.
 * Subspecies of wroughtonii: Emery, 1911d: 30; Chapman & Capco, 1951: 49; Tiwari, 1999: 32 (error).
 * Junior synonym of parallela: Brown, 1975: 8; Bolton, 1995b: 337.
 * wroughtonii. Platythyrea wroughtonii Forel, 1900d: 315 (w.) INDIA (Kerala, Karnataka).
 * Type-material: syntype workers (number not stated).
 * Type-localities: India: Travancore (Ferguson), and Mysore (Lee).
 * Type-depository: MHNG.
 * Status as species: Bingham, 1903: 75; Emery, 1911d: 30; Wheeler, W.M. 1922a: 1007; Chapman & Capco, 1951: 49; Tiwari, 1999: 32 (error).
 * Junior synonym of parallela: Brown, 1975: 8; Bolton, 1995b: 337.

Type Material


Ponera parallela

Holotype worker in. Labelled “Aru” (= Aru I., New Guinea) and with a Donisthorpe type-label. A second specimen in is from the same locality and has the data label written in the same thickly-marked ink, but the method of mounting isnot the same. Nevetheless, this may also be from the type-series.

Description
Brown (1975) provided the following:

Wilson (1958:151) began to assemble the synonymy of P. parallela during his study of the Melanesian ponerines, when he sank under it P. pusilla, P. coxalis, P. melancholica, and P. melancholica var. aruana, the last two names provisionally. P. melancholica originally was vaguely described by F. Smith in 1865 as a Pachycondyla, but Emery (1897: 556) surmised that the species was a Platythyrea, a placement in which he was followed by all later authors when the type of P. melancholica could not be located. However, the type is extant in the Oxford University Collection, where it was found by Barry Bolton, and I was able to examine it in London in 1973. It proves to be true Pachycondyla in the broad sense (Ectomomyrmex of recent authors), and so the species melancholica can be eliminated from consideration under Platythyrea. It will be dealt with in Part VI of this series.

Wilson was also able to compare digms of the parallela complex that I provided for him in 1955 with types of other related forms in European museums, and I have been able to study these and many others abroad and in MCZ in subsequent years, with the result that the synonymy of P. parallela is lengthened to include a surprising number of specific, subspecific, and varietal names.

As might be suspected from the length of the list of synonyms, P. parallela is a variable insect. The most conspicuous variation is seen in overall body size, as indicated by the length of the trunk (WL), which ranges from about 1.10 to about 2.50 mm in specimens representing more than 50 nest series assigned to this species that I have been able to examine carefully enough. In discussing individual synonyms below, I have given WL for at least one type specimen wherever I was able to make a suitable measurement. I am unable to split the size continuum into modal classes that seem to represent different species, and small, medium, and large forms are known to occur sympatrically at such relatively well sampled localities as the Cuernos Mts., near Dumaguete, Negros Oriental, Philippines, and on Ceylon. Within single nest series, however, size variation so far as known is relatively narrow in range.

Other variation, as noted by Wilson, occurs in the shape and size of the propodeal angles; in the length vs. width, dorsal convexity, and shape (of the posterodorsal margin) of the petiolar node; in the opacity of the background fine sculpture; in the number and distinctness of the superimposed coarser punctures; in the color of the legs, and other lesser traits. Some preliminary analysis of the length vs. width of the petiolar node by Wilson (1958:152) showed that the larger specimens tend to have disproportionately wider nodes. A rough check indicates that the additional data from the augmented series now available would probably widen Wilson's already fairly wide regression zone, but that the trend would remain the same. On the other hand, the sculptural variation does not seem to be closely correlated with size.

If more than one species is present in this complex, it will take much more refined analysis based on more and better material to reveal it. For the present, the burden of proof is shifted to the splitters, because I am considering the whole complex to represent a single species. Such a treatment only parallels the conclusions reached for other Platythyrea species in this study. My notes on the individual synonyms are repeated here, in the approximate chronological order of publication.

P. parallela type WL 2.15 mm.; tibiae dark. Sculpture weak, fairly smooth; larger punctures weak, obsolete on pronotum; tibiae dark. Type loc.: Aru Island.

P. inconspicua type: dealate queen, tibiae dark brown, WL 2.45 mm., is near the large end of the size range. On my request, Dr. Max Fischer of the Naturhistorisches Museum in Vienna kindly sent 2 specimens, each placed in the Mayr Collection as "inconspicua/G. Mayr Typus." Emery mentions these specimens as the two possessed by Mayr, who identified South American samples sent to him by Emery as P. inconspicua. One of Mayr's specimens now bears the tiny locality label, "Ceilan/Sichel," and on a larger label below this, apparently in Mayr's handwriting, "Cayenne/wolnicht/Ceilon." The writing is crude and was obviously done in haste; apparently Mayr decided, at some time after the original description, that the specimen was "wohl nicht" (probably not) from Ceylon, and its superficial similarity to the second "Typus," from Panama ("Pergand") further led him to believe that the locality of the first type should have been Cayenne. From this point, it was an easy step to identify Emery's specimens also as inconspicua.

A close examination of the original (and only real) type of P. inconspicua shows that it is really an Old World tropical member of the genus, and in fact is a large specimen of P. parallela. In the first place, the masticatory borders of the mandibles are clearly denticulate or crenulate (in the American species, the border is toothless and cultrate). In the second place, the palpi are very short, and are segmented 3,2, as in other P. parallela examined, large and small. Mayr's Panama specimen, on the other hand, has much longer palpi (extending past the posterior margin of the buccal cavity), and the maxillary palpi are 6 merous, the labials 4 merous, as in other members of the punctata group. The Panama specimen is in fact a specimen of P. pilosula, and cannot be considered as in any way a type of P. inconspicua; it was never mentioned in the original description because it had not even been collected by then.

P. inconspicua is therefore removed from the synonymy of P. punctata, where Emery had assigned it, and is placed as a new synonym of parallela.

P. pusilla and P. coxalis were described in the same paper at a time when the correct ,generic placement of Ponera parallela F. Smith was still unknown, and by this time, Emery had already considered P. inconspicua to be a neotropical species (see above). Wilson (1968:151) examined the holotypes of Emery's two species in 1955 and left type compared digms in MCZ, one of which also compared with P. parallela, and the other with P. wroughtonii, its subspecies victoriae, and P. parva. His synonymy of pusilla and coxalis under parallela (1958) seems to me to be correct. The type locality of P. pusilla is Amboina, and for P. coxalis is Perak, Malaya.

The types of P. wroughtonii and P. wroughtonii victoriae were examined in the Forel Collection in Geneva. WL of a wroughtonii specimen was 1.36 mm, and victoriae is a little larger (MCZ syntype WL 1.40 mm). The differences cited by Forel to separate these forms are very vague, and I was unable to appreciate them in the type specimens; these both seem to me to be small sized parallela. The type localities for wroughtonii are Travancore and Mysore (S peninsular India); for victoriae, they are Bangalore (in Mysore), Barrackpore (N of Calcutta), and "Inde septentrionale." Both forms have the tibiae light yellow.

The types of P. subtilis and P. pusilla were compared directly in the Emery Collection (MCSN, Genoa), and I could find no important differences between them; the subtilis type is perhaps a little more heavily punctate on the sides of the propodeum and petiolar node. A syntype specimen of P. wroughtonii from "Bangalore" in the Emery Collection compares well with these types also. P. subtilis type loc.: Pulau Laut, SE Borneo (Kalimantan).

In the Kodeco Tract west of Batulitjin (SE Kalimantan), inland from Pulau Laut, in 1972, I collected several samples of the P. parallela complex, including workers of fairly large size (WL 1.81 1.90 mm) and one very small stray worker, the smallest I have seen (WL 1.11 mm), with a fairly short, heavily punctured node with distinct blunt teeth behind. While on the subject of size, I should also mention a single, very large, worker individual (WL 2.50 mm) taken by C. F. Clagg at 5000 6000 feet elevation on Mt. Apo, Mindanao. It is hard to believe that this relative giant is simply a large specimen of P. parallela, but it is almost exactly like that species; its eyes are a bit small (greatest diameter 0.13 mm) for such a large specimen, and its second funicular segment is about 1 1/2 times as long as broad (usually about as broad as long in parallela), but in other characters, including 3,2 palpal segmentation, it fits P. parallela.

P. coxalis var. tritschleri is a variety intermediate in size between the types of P. parallela and P. wroughtonii with the lobules of the posterior border of the petiolar node almost completely obsolete. Type locality Indrapura, Sumatra (MHN  Geneva).

P. coxalis var. javana is similar to the foregoing in size, but the tendency to form 3 low lobes on the posterior border of the petiole is better expressed. In javana, the larger punctures are weak and difficult to see. Type locality is Buitenzorg (now Bogor), Java. Both of the above varieties have reddish brown tibiae (MHNG).

P. coxalis var. annamita is a little larger than the foregoing (WL 2.12 mm), but seems to belong to the same species. Sculpture fairly weak; tibiae brown. Type loc.: Region of the Mois, central Cochin China. (MHNG).

P. wroughtonii subsp. sechellensis has WL 1.55 mm. It has weak, feebly shining sculpture, with small, widely spaced punctures; tibiae yellow. Type loc.: Praslin, Seychelles Is. (BMNH London.)

P. coxalis var. cylindrica is a medium sized queen (WL 1.67 mm) with larger punctures distinct, but reduced in size on pronotum; tibiae brown. Type loc.: Tandjong Slamat, E Sumatra. (MNHU Berlin.)

P. pusilla var. australis is a small form (not measured) similar to wroughtonii types; tibiae with yellowish extensor surfaces. Type loc.: Blackall Range and Tambourine Mt., S Queensland. (MHN Geneva.)

Platythyrea parva is like the wroughtonii types, but head a bit broader; tibiae with extensor surfaces lightened and yellowish; WL 1.34 and 1.40 on holotype and paratype measured. Type loc.: Darwin, N Australia. (BMNH London, MHNG and MCZC.)

P. pusilla var. egena: WL 1.51 mm; tibiae more or less pale. Type loc.: Singapore. (MNHU Berlin.)

P. coxalis var. philippinensis: WL 1.75, 1.80 mm; tibiae brown. Type loc.: Los Bafios, Laguna Prov., S Luzon (MNHU Berlin, MCZ). Dr. J. W. Chapman assembled large series of P. parallela in the Philippines, although many of the specimens were molded or otherwise damaged during World War II. Most of his samples come from the Cuernos Mts. near Dumaguete, Negros Oriental, 1500 to 4000 feet elevation. These vary somewhat in body size and have the legs brown, reddish brown, or with tibiae somewhat lightened in color. Other Philippine samples are from Luzon: Baguio (Chapman); Los Bafios (F. X. Williams); Mt. Makiling (A. L. Cambre); Mindanao: Davao City (A. Reyes), Momungan (no collector), Miatan, Katipunan (D. Empeso); Romblon, Cabrador I. (L. Morato). Borneo: Kuching, Sarawak, (Hewitt); Sandakan (C. F. Baker); Mt. Dulit, N. Borneo, 3000 ft (E. Mjoberg). Gilolo I.: Kalam (C. S. Banks). S. Celebes: Balampesoang Forest, near Tanete (W. L. Brown).

P. cephalotes: WL 2.00 mm; tibiae pale. Weakly shining in most lights, but some small punctures extend onto pronotum. Petiolar node L 0.63, W 0.60 mm, posterodorsal border of node sinuate as seen from above. Eyes a little smaller than in New Guinea parallela. Type loc.: "Walde be.i Trial Bay," New South Wales, Australia. (MNHU, Berlin.) Additional material from N.S.W.: Jenolan Caves (J. C. Wiburd), Bulga (W. W. Froggatt), Dorrigo (W. Heron), Royal National Park, 25 m elevation (E. S. Ross and D. Q. Cavagnaro), as well as Brisbane, Queensland (H. Hacker) is much like the cephalotes type. Smaller specimens come from the McIlwraith Range, Cape York Peninsula, Queensland (P. J. Darlington).

P. melancholica var. aruana was synonymized with parallela by Wilson provisionally because the type was unavailable. The type locality is Wammar in the Aru Archipelago, and Aru is the type locality of ''parallela'. Upon reading Karavaiev's description again, I can only conclude that Wilson's synonymy is reasonable even in the absence of the aruana type.

P. pulchella: the type in NM Basel is an alate queen from Medan, Sumatra (Corporaal). Node rather short, only slightly longer than broad, but with a sinuate posterior margin as seen from above. Scapes and tibiae whitish, prominent against black of remainder of body.

P. pusilla var. pacifica was put into the synonymy of P. parallela by Wilson and Taylor (1967:20), with good reason in my opinion. Type loc.: Apia, Upolu, Samoa.

P. ceylonensis: WL 1.86 mm; tibiae brown. Type loc.: Ceylon. (BMNH London.) This last synonym in the long list is just a fairly large parallela queen and workers. Other specimens from Ceylon are from Yakkala (KLA Perera), and Soans and I took a series of slightly larger body size (WL 1.70 1.74 mm) in Kerala State, S India: Peria Reserve, Cannanore Dist., 4 April 1969, tropical evergreen forest, ca. 900 m. Other samples from the SE Asian mainland are: Thailand: E slope, Doi Sutep, 260 m, 15 July 1962 (WL 1.47 mm) and Khao Yai National Park, 750 m, 26 July 1962 (WL 2.06 mm), E. S. Ross and D. Q Cavagnaro. Malaysia: Pahang, below The Gap, about 850 m (R. H. Crozier), in hill forest. The MCZ also has a sample from Wai Lima, Lampong, S. Sumatra (Karny).

References based on Global Ant Biodiversity Informatics

 * Basu P. 1994. Ecology of ground foraging ants in a tropical evergreen forest in Western Ghats, India. PhD Thesis, School of ecology and environmental sciences, Pondichery University, India. 155 pages.
 * Brown W. L., Jr. 1975. Contributions toward a reclassification of the Formicidae. V. Ponerinae, tribes Platythyreini, Cerapachyini, Cylindromyrmecini, Acanthostichini, and Aenictogitini. Search Agric. (Ithaca N. Y.) 5(1): 1-115.
 * CSIRO Collection
 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Cheesman L. E., and W. C. Crawley. 1928. A contribution towards the insect fauna of French Oceania. - Part III. Formicidae. Ann. Mag. Nat. Hist. 10(2): 514-525.
 * Clouse R. M. 2007. The ants of Micronesia (Hymenoptera: Formicidae). Micronesica. 39: 171-295.
 * Clouse, R.M. 2007. The ants of Micronesia (Hymenoptera: Formicidae), Micronesica 39(2): 171-295.
 * Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
 * Dahl F. 1901. Das Leben der Ameisen im Bismarck-Archipel, nach eigenen Beobachtungen vergleichend dargestellt. Mitt. Zool. Mus. Berl. 2: 1-70.
 * Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
 * Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
 * Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
 * Dias R. K. S., and K. R. K. Anuradha Kosgamage. 2012. Occurrence and species diversity of ground-dwelling worker ants (Family: Formicidae) in selected lands in the dry zone of Sri Lanka. J. Sci. Univ. Kelaniya 7: 55-72.
 * Dias, R.K.S. 2006. Overview of ant research in Sri Lanka: 2000-2004. ANeT Newsletter 8:7-10
 * Donisthorpe H. 1941. Descriptions of new ants (Hym., Formicidae) from various localities. Annals and Magazine of Natural History (11)8: 199-210.
 * Donisthorpe H. 1941. Descriptions of new species of ants from New Guinea. Annals and Magazine of Natural History (11)7: 129-144.
 * Donisthorpe H. 1947. Ants from New Guinea, including new species and a new genus. Annals and Magazine of Natural History (11)13: 577-595.
 * Donisthorpe H. 1948. A fourth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)1: 131-143.
 * Donisthorpe H. 1948. A third instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (11)14: 589-604.
 * Donisthorpe H. 1950. An eighth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)3: 338-341.
 * Donisthorpe, Horace. 1943. The Ants of Waigeu Island, North Dutch New Guinea. The Annals and Magazine of Natural History 11 (10): 433-475.
 * Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). [concl.]. Ann. Mus. Civ. Stor. Nat. 25(5): 427-473.
 * Emery C. 1893. Formicides de l'Archipel Malais. Revue Suisse de Zoologie 1: 187-229.
 * Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 14:565-580.
 * Emery C. 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1-125.
 * Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine)." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 5, no. 25 (1887): 427-473.
 * Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
 * Emery, C. "Voyage de MM. Bedot et Pictet dans l'Archipel Malais. Formicides de l'Archipel Malais." Revue Suisse de Zoologie 1 (1893): 187-229.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Fontanilla A. M., A. Nakamura, Z. Xu, M. Cao, R. L. Kitching, Y. Tang, and C. J. Burwell. 2019. Taxonomic and functional ant diversity along tropical, subtropical, and subalpine elevational transects in southwest China. Insects 10, 128; doi:10.3390/insects10050128
 * Forel A. 1900. Les Formicides de l'Empire des Indes et de Ceylan. Part VII. J. Bombay Nat. Hist. Soc. 13: 303-332.
 * Forel A. 1901. Formiciden aus dem Bismarck-Archipel, auf Grundlage des von Prof. Dr. F. Dahl gesammelten Materials. Mitt. Zool. Mus. Berl. 2: 4-37.
 * Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
 * Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
 * Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
 * Forel A. 1915. Results of Dr. E. Mjöbergs Swedish Scientific Expeditions to Australia 1910-13. 2. Ameisen. Ark. Zool. 9(16): 1-119
 * Ghosh S. N., S. Sheela, and B. G. Kundu. 2005. Ants (Hymenoptera: Formicidae) of Rabindra Sarovar, Kolkata. Records of the Zoological Survey of India. Occasional Paper 234: 1-40.
 * Hashimoto Y., Y. Morimoto, and M. Mohamed. 2003. Species List of Ground and Leaf Litter Ants Collected in Lower Kinabatangan. Pp 13-18. In Lower Kinabatangan Scientific Expedition 2002, 176 pp. ISBN-13: 983-2369-11-8
 * Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
 * Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
 * Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
 * Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
 * Janda M., G. D. Alpert, M. L. Borowiec, E. P. Economo, P. Klimes, E. Sarnat, and S. O. Shattuck. 2011. Cheklist of ants described and recorded from New Guinea and associated islands. Available on http://www.newguineants.org/. Accessed on 24th Feb. 2011.
 * Jennings J. T., L. Krogmann, and C. Burwell. 2013. Review of the hymenopteran fauna of New Caledonia with a checklist of species. Zootaxa 3736(1): 1-53.
 * Karavaiev V. 1926. Ameisen aus dem Indo-Australischen Gebiet. Treubia 8: 413-445.
 * Karavaiev, V. 1925. Ponerinen (Fam. Formicidae) aus dem Indo-Australischen Gebiet. Konowia 4:69-81.
 * Latumahina F., M. Borovanska, N. S. Putra, and M. Janda. 2015. Ants of Ambon Island  diversity survey and checklist. ZooKeys 472: 4357.
 * Mann W. M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology 63:273-391.
 * Mann William. 1916. The Ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology at Harvard College 63(7): 273-391
 * Mann, W.M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology of Harvard College 63: 273-391
 * Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
 * Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
 * Musthak Ali T. M. 1991. Ant Fauna of Karnataka-1. Newsletter of IUSSI Indian Chapter 5(1-2): 1-8.
 * Narendra A., H. Gibb, and T. M. Ali. 2011. Structure of ant assemblages in Western Ghats, India: role of habitat, disturbance and introduced species. Insect Conservation and diversity 4(2): 132-141.
 * Neville, P. J., O'Dowd, D. J., and Yen, A. L. 2008. Issues and implications for research on disturbed oceanic islands illustrated through an ant survey of the Cocos (Keeling) Islands. J Insect Conserv. 12:313-323.
 * Neville, P.J., D.J. O'Dowd and A.L. Yen. 2008. Issues and implications for research on disturbed oceanic islands illustrated through an ant survey of the Cocos (Keeling) Islands. Insect Conserv. 12. 313-323.
 * Overbeck H. 1924. A List of Ants from Singapore and Neighbouring Places. Journal of the Malayan Branch of the Royal Asiatic Society 2 (1): 25-40.
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
 * Pfeiffer, M., H. Cheng Tuck, and T. Chong Lay. 2008. Exploring arboreal ant community composition and co-ccurrence patterns in plantations of oil palm Elaeis guineensis in Borneo and Peninsular Malaysia. Ecography 31(1): 21-32.
 * Rizali A., D. J. Lohman, D. Buchori, L. Budi Prasetyo, H. Triwidodo, M. M. Bos, S. Yamane, and C. H. Schulze. 2009. Ant communities on small tropical islands: effects of island size and isolation are obscured by habitat disturbance and tramp ant species. Journal of Biogeography 37(2): 229-236.
 * Room, P.M. 1975. Relative Distributions of Ant Species in Cocoa Plantations in Papua New Guinea Relative Distributions of Ant Species in Cocoa Plantations in Papua New Guinea. Journal of Applied Ecology 12(1):47-61
 * Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
 * Smith F. 1861. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Ceram, Celebes, Ternate, and Gilolo. [part]. Journal and Proceedings of the Linnean Society of London. Zoology 6: 36-48.
 * Smith F. 1863. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Mysol, Ceram, Waigiou, Bouru and Timor. Journal and Proceedings of the Linnean Society of London. Zoology 7: 6-48.
 * Snelling R. R. 1998. Insect Part 1: The social Hymenoptera. In Mack A. L. (Ed.) A Biological Assessment of the Lakekamu Basin, Papua New Guinea, RAP 9. 189 ppages
 * Snelling R. R. 2000. Ants of the Wapoga river area, Irian Jaya, Indonesia. In Mack, Andrew L. and Leeanne E. Alonso (eds.). 2000. A Biological Assessment of the Wapoga River Area of Northwestern Irian Jaya, Indonesia. RAP Bulletin of Biological Assessment 14, Conservation International, Washington, DC.
 * Taylor R. W. 1987. A checklist of the ants of Australia, New Caledonia and New Zealand (Hymenoptera: Formicidae). CSIRO (Commonwealth Scientific and Industrial Research Organization) Division of Entomology Report 41: 1-92.
 * Tiwari R. N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4): 1-96.
 * Tiwari, R.N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4):1-96
 * Varghese T. 2004. Taxonomic studies on ant genera of the Indian Institute of Science campus with notes on their nesting habits. Pp. 485-502 in : Rajmohana, K.; Sudheer, K.; Girish Kumar, P.; Santhosh, S. (eds.) 2004. Perspectives on biosystematics and biodiversity. Prof. T.C. Narendran commemoration volume. Kerala: Systematic Entomology Research Scholars Association, xxii + 666 pp.
 * Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
 * Viehmeyer H. 1913. Ameisen aus dem Kopal von Celebes. Stettiner Entomologische Zeitung 74: 141-155.
 * Viehmeyer H. 1914. Papuanische Ameisen. Deutsche Entomologische Zeitschrift 1914: 515-535.
 * Viehmeyer H. 1916. Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv für Naturgeschichte (A)81(8):108-168.
 * Viehmeyer H. 1916. Ameisen von den Philippinen und anderer Herkunft (Hym.). Entomologische Mitteilungen. Berlin-Dahlem 5: 283-291.
 * Watanasit S., and N. Noon-anant. 2005. Ants at Ton Nga Chang Wildlife Sanctuary, Songkhla. Sonklanakarin J. Sci. Technolo. 27(2): 267-280.
 * Way M. J., and B. Bolton. 1997. Competition between ants for coconut palm nesting sites. Journal of Natural History 31: 439-455.
 * Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
 * Wheeler W. M., and J. W. Chapman. 1925. The ants of the Philippine Islands. Part I, Dorylinae and Ponerinae. Philipp. J. Sci. 28: 47-73.
 * Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
 * Wheeler, William Morton.1935.Checklist of the Ants of Oceania.Occasional Papers 11(11): 3-56
 * Wilson E. O. 1958. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bulletin of the Museum of Comparative Zoology 118: 101-153.
 * Wilson E. O. 1959. Some ecological characteristics of ants in New Guinea rain forests. Ecology 40: 437-447.
 * Wilson E.O. 1959. Adaptive shift and dispersal in a tropical ant fauna. Evolution 13(1): 122-144.
 * Wilson Edward O. 1959. Adaptive Shift and Dispersal in a Tropical Ant Fauna. Evolution 13(1): 122-144
 * Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
 * Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.