Guyana

The following 341 valid species and subspecies of ants are known to occur in Guyana. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Guyana ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Guyana Ants web page. Please advise if any corrections are warranted.

Acanthognathus

 * Acanthognathus brevicornis Smith, M.R., 1944
 * Acanthognathus lentus Mann, 1922

Acanthostichus

 * Acanthostichus brevicornis Emery, 1894

Acromyrmex

 * Acromyrmex coronatus globoculis 	Kempf, 1972
 * Acromyrmex hystrix Latreille, 1802
 * Acromyrmex octospinosus 	Reich, 1793
 * Acromyrmex landolti myersi 	Weber, 1937

Acropyga

 * Acropyga ayanganna 	LaPolla, 2004
 * Acropyga donisthorpei 	Weber, 1944
 * Acropyga exsanguis 	Wheeler, W.M., 1909
 * Acropyga fuhrmanni 	Forel, 1914
 * Acropyga goeldii 	Forel, 1893
 * Acropyga guianensis 	Weber, 1944
 * Acropyga panamensis 	Weber, 1944
 * Acropyga romeo 	LaPolla, 2004
 * Acropyga stenotes 	LaPolla, 2004

Allomerus

 * Allomerus octoarticulatus Mayr, 1878

Anochetus

 * Anochetus bispinosus 	Smith, F., 1858
 * Anochetus emarginatus 	Fabricius, 1804
 * Anochetus mayri 	Emery, 1884
 * Anochetus simoni 	Emery, 1890

Apterostigma

 * Apterostigma auriculatum 	Wheeler, W.M., 1925
 * Apterostigma dorotheae 	Weber, 1937
 * Apterostigma epinotale 	Weber, 1937
 * Apterostigma jubatum 	Wheeler, W.M., 1925
 * Apterostigma robustum 	Emery, 1896
 * Apterostigma urichii 	Forel, 1893
 * Apterostigma wasmannii 	Forel, 1892

Atta

 * Atta cephalotes 	Linnaeus, 1758
 * Atta laevigata 	Smith, F., 1858
 * Atta sexdens 	Linnaeus, 1758

Azteca

 * Azteca alfari 	Emery, 1893
 * Azteca chartifex
 * Azteca constructor 	Emery, 1896
 * Azteca delpini 	Emery, 1893
 * Azteca foveiceps 	Wheeler, W.M., 1921
 * Azteca gnava cayennensis 	Forel, 1912
 * Azteca instabilis 	Smith, F., 1862
 * Azteca jelskii 	Emery, 1893
 * Azteca schimperi 	Emery, 1893
 * Azteca sericea 	Mayr, 1866
 * Azteca trailii 	Emery, 1893
 * Azteca trailii tillandsiarum Wheeler, W.M., 1942
 * Azteca trigona mediops 	Forel, 1904
 * Azteca ulei cordiae 	Forel, 1904
 * Azteca velox Forel, 1899

Basiceros

 * Basiceros redux 	Donisthorpe, 1939
 * Basiceros singularis 	Smith, F., 1858

Brachymyrmex

 * Brachymyrmex aphidicola Forel, 1909
 * Brachymyrmex australis Forel, 1901
 * Brachymyrmex coactus Mayr, 1887
 * Brachymyrmex cordemoyi Forel, 1895
 * Brachymyrmex heeri Forel, 1874 (Santschi, 1923a)
 * Brachymyrmex pictus Mayr, 1887

Camponotus

 * Camponotus ager 	Smith, F., 1858
 * Camponotus apicalis (Mann, 1916)
 * Camponotus atriceps 	Smith, F., 1858
 * Camponotus auricomus 	Roger, 1862
 * Camponotus beebei 	Wheeler, W.M., 1918
 * Camponotus bidens 	Mayr, 1870
 * Camponotus blandus 	Smith, F., 1858
 * Camponotus chartifex 	Smith, F., 1860
 * Camponotus corniculatus Wheeler, W.M., 1934
 * Camponotus fasciatellus 	Dalla Torre, 1892
 * Camponotus fastigatus 	Roger, 1863
 * Camponotus femoratus 	Fabricius, 1804
 * Camponotus ignestii Menozzi, 1935
 * Camponotus latangulus 	Roger, 1863
 * Camponotus leydigi 	Forel, 1886
 * Camponotus linnaei maccus Wheeler, W.M., 1934
 * Camponotus mocsaryi 	Forel, 1902
 * Camponotus nidulans (Smith, F., 1860)
 * Camponotus novogranadensis 	Mayr, 1870
 * Camponotus patimae 	Wheeler, W.M., 1942
 * Camponotus patimae dolentulus Wheeler, W.M., 1942
 * Camponotus picipes 	Olivier, 1792
 * Camponotus pittieri poenalis Wheeler, W.M., 1912
 * Camponotus platytarsus 	Roger, 1863
 * Camponotus rapax 	Fabricius, 1804
 * Camponotus rectangularis 	Emery, 1890
 * Camponotus renggeri 	Emery, 1894
 * Camponotus rufipes 	Fabricius, 1775
 * Camponotus sericeiventris 	Guérin-Méneville, 1838
 * Camponotus sexguttatus 	Fabricius, 1793
 * Camponotus simillimus 	Smith, F., 1862
 * Camponotus substitutus 	Emery, 1894
 * Camponotus zoc 	Forel, 1879

Carebara

 * Carebara anophthalma (Emery, 1906)
 * Carebara bicarinata Santschi, 1912
 * Carebara mayri 	Forel, 1901

Cephalotes

 * Cephalotes atratus 	Linnaeus, 1758
 * Cephalotes clypeatus 	Fabricius, 1804
 * Cephalotes depressus 	Klug, 1824
 * Cephalotes grandinosus 	Smith, F., 1860
 * Cephalotes maculatus 	Smith, F., 1876
 * Cephalotes manni 	Kempf, 1951
 * Cephalotes marginatus 	Fabricius, 1804
 * Cephalotes minutus 	Fabricius, 1804
 * Cephalotes opacus 	Santschi, 1920
 * Cephalotes pallens 	Klug, 1824
 * Cephalotes pallidoides 	De Andrade, 1999
 * Cephalotes pallidus 	De Andrade, 1999
 * Cephalotes palustris 	De Andrade, 1999
 * Cephalotes pavonii 	Latreille, 1809
 * Cephalotes placidus 	Smith, F., 1860
 * Cephalotes pusillus 	Klug, 1824
 * Cephalotes simillimus 	Kempf, 1951
 * Cephalotes spinosus 	Mayr, 1862
 * Cephalotes targionii 	Emery, 1894
 * Cephalotes umbraculatus 	Fabricius, 1804

Cheliomyrmex

 * Cheliomyrmex megalonyx 	Wheeler, W.M., 1921
 * Cheliomyrmex morosus 	Smith, F., 1859

Crematogaster

 * Crematogaster levior 	Longino, 2003
 * Crematogaster rochai 	Forel, 1903
 * Crematogaster tenuicula 	Forel, 1904
 * Crematogaster acuta 	Fabricius, 1804
 * Crematogaster abstinens 	Forel, 1899
 * Crematogaster brasiliensis arawak 	Weber, 1938
 * Crematogaster crinosa 	Mayr, 1862
 * Crematogaster curvispinosa 	Mayr, 1862
 * Crematogaster delitescens 	Wheeler, W.M., 1921
 * Crematogaster erecta 	Mayr, 1866
 * Crematogaster huberi 	Forel, 1907
 * Crematogaster limata 	Smith, F., 1858
 * Crematogaster longispina 	Emery, 1890
 * Crematogaster quadriformis 	Roger, 1863
 * Crematogaster stollii 	Forel, 1885

Cryptopone

 * Cryptopone guianensis 	Weber, 1939
 * Cryptopone holmgreni 	Wheeler, W.M., 1925

Cylindromyrmex

 * Cylindromyrmex striatus 	Mayr, 1870

Cyphomyrmex

 * Cyphomyrmex laevigatus 	Weber, 1938
 * Cyphomyrmex minutus 	Mayr, 1862
 * Cyphomyrmex rimosus 	Spinola, 1851

Daceton

 * Daceton armigerum 	Latreille, 1802

Dinoponera

 * Dinoponera gigantea (Perty, 1833)

Discothyrea

 * Discothyrea denticulata 	Weber, 1939

Dolichoderus

 * Dolichoderus attelaboides 	Fabricius, 1775
 * Dolichoderus bidens 	Linnaeus, 1758
 * Dolichoderus bispinosus 	Olivier, 1792
 * Dolichoderus debilis 	Emery, 1890
 * Dolichoderus decollatus 	Smith, F., 1858
 * Dolichoderus diversus Emery, 1894
 * Dolichoderus gagates 	Emery, 1890
 * Dolichoderus lamellosus 	Mayr, 1870
 * Dolichoderus laminatus 	Mayr, 1870
 * Dolichoderus lugens 	Emery, 1894
 * Dolichoderus lutosus 	Smith, F., 1858
 * Dolichoderus rugosus 	Smith, F., 1858
 * Dolichoderus septemspinosus 	Emery, 1894
 * Dolichoderus spurius 	Forel, 1903

Dorymyrmex

 * Dorymyrmex brunneus 	Forel, 1908
 * Dorymyrmex pyramicus 	Roger, 1863

Eciton

 * Eciton burchellii cupiens 	Santschi, 1923
 * Eciton drepanophorum 	Smith, F., 1858
 * Eciton hamatum 	Fabricius, 1782
 * Eciton vagans 	Olivier, 1792

Ectatomma

 * Ectatomma brunneum 	Smith, F., 1858
 * Ectatomma edentatum 	Roger, 1863
 * Ectatomma lugens 	Emery, 1894
 * Ectatomma ruidum 	Roger, 1860
 * Ectatomma tuberculatum 	Olivier, 1792

Eurhopalothrix

 * Eurhopalothrix bolaui 	Mayr, 1870
 * Eurhopalothrix clypeata 	Brown & Kempf, 1960

Gigantiops

 * Gigantiops destructor 	Fabricius, 1804

Gnamptogenys

 * Gnamptogenys acuminata 	Emery, 1896
 * Gnamptogenys annulata 	Mayr, 1887
 * Gnamptogenys ericae 	Forel, 1912
 * Gnamptogenys gracilis 	Santschi, 1929
 * Gnamptogenys horni 	Santschi, 1929
 * Gnamptogenys minuta 	Emery, 1896
 * Gnamptogenys moelleri (Forel, 1912)
 * Gnamptogenys mordax 	Smith, F., 1858
 * Gnamptogenys regularis 	Mayr, 1870
 * Gnamptogenys striatula 	Mayr, 1884
 * Gnamptogenys strigata 	Norton, 1868
 * Gnamptogenys sulcata 	Smith, F., 1858
 * Gnamptogenys tortuolosa 	Smith, F., 1858

Hylomyrma

 * Hylomyrma blandiens 	Kempf, 1961
 * Hylomyrma immanis 	Kempf, 1973
 * Hylomyrma longiscapa 	Kempf, 1961
 * Hylomyrma reginae 	Kutter, 1977

Hypoponera

 * Hypoponera opaciceps 	Mayr, 1887

Labidus

 * Labidus coecus 	Latreille, 1802
 * Labidus praedator (Smith, F., 1858)
 * Labidus truncatidens 	Santschi, 1920

Lachnomyrmex

 * Lachnomyrmex platynodus Feitosa & Brandao, 2008

Leptogenys

 * Leptogenys arcuata 	Roger, 1861
 * Leptogenys gaigei 	Wheeler, W.M., 1923
 * Leptogenys guianensis Wheeler, W.M., 1923
 * Leptogenys langi 	Wheeler, W.M., 1923
 * Leptogenys linearis (Smith, F., 1858)
 * Leptogenys unistimulosa 	Roger, 1863

Linepithema

 * Linepithema iniquum 	(Mayr, 1870)

Mayaponera

 * Mayaponera constricta 	Mayr, 1884

Megalomyrmex

 * Megalomyrmex bituberculatus 	Fabricius, 1798
 * Megalomyrmex cuatiara Brandão, 1990
 * Megalomyrmex drifti 	Kempf, 1961
 * Megalomyrmex emeryi 	Forel, 1904
 * Megalomyrmex leoninus 	Forel, 1885
 * Megalomyrmex silvestrii 	Wheeler, W.M., 1909
 * Megalomyrmex wallacei Mann, 1916 66

Monomorium

 * Monomorium floricola 	Jerdon, 1851
 * Monomorium pharaonis 	Linnaeus, 1758

Mycetomoellerius

 * Mycetomoellerius farinosus (Emery, 1894)
 * Mycetomoellerius guianensis (Weber, 1937)
 * Mycetomoellerius opulentus (Mann, 1922)
 * Mycetomoellerius relictus (Borgmeier, 1934)
 * Mycetomoellerius urichii (Forel, 1893)
 * Mycetomoellerius verrucosus (Borgmeier, 1948)

Mycetophylax

 * Mycetophylax bigibbosus (Emery, 1894)
 * Mycetophylax conformis Mayr, 1884
 * Mycetophylax faunulus (Wheeler, W.M., 1925)

Mycocepurus

 * Mycocepurus goeldii 	Forel, 1893
 * Mycocepurus smithii 	Forel, 1893

Myrmelachista

 * Myrmelachista donisthorpei 	Wheeler, W.M., 1934
 * Myrmelachista flavida 	Wheeler, W.M., 1934
 * Myrmelachista guyanensis 	Wheeler, W.M., 1934

Myrmicocrypta

 * Myrmicocrypta buenzlii 	Borgmeier, 1934
 * Myrmicocrypta elisabethae 	Weber, 1937
 * Myrmicocrypta guianensis 	Weber, 1937
 * Myrmicocrypta infuscata 	Weber, 1946
 * Myrmicocrypta spinosa 	Weber, 1937
 * Myrmicocrypta squamosa uncinata 	Mayr, 1887
 * Myrmicocrypta unidentata 	Weber, 1937

Neivamyrmex

 * Neivamyrmex angustinodis 	Emery, 1888
 * Neivamyrmex bohlsi 	Emery, 1896
 * Neivamyrmex emersoni 	Wheeler, W.M., 1921
 * Neivamyrmex falciferus 	Emery, 1900
 * Neivamyrmex gibbatus 	Borgmeier, 1953
 * Neivamyrmex guerinii 	Shuckard, 1840
 * Neivamyrmex guyanensis 	Santschi, 1916
 * Neivamyrmex halidaii 	Shuckard, 1840
 * Neivamyrmex iridescens 	Borgmeier, 1950
 * Neivamyrmex maxillosus 	Emery, 1900
 * Neivamyrmex megathrix 	Kempf, 1961
 * Neivamyrmex pilosus 	(Smith, F. 1858)
 * Neivamyrmex postangustatus 	Borgmeier, 1934
 * Neivamyrmex punctaticeps 	Emery, 1894
 * Neivamyrmex walkerii 	Westwood, 1842

Neoponera

 * Neoponera apicalis 	Latreille, 1802
 * Neoponera carinulata (Roger, 1861)
 * Neoponera cavinodis (Mann, 1916)
 * Neoponera commutata 	Roger, 1860
 * Neoponera crenata 	Roger, 1861
 * Neoponera foetida 	Linnaeus, 1758
 * Neoponera inversa 	Smith, F., 1858
 * Neoponera laevigata 	Smith, F., 1858
 * Neoponera obscuricornis 	Emery, 1890
 * Neoponera procidua 	Emery, 1890
 * Neoponera unidentata 	Mayr, 1862
 * Neoponera villosa 	Fabricius, 1804

Nesomyrmex

 * Nesomyrmex anduzei 	Weber, 1943
 * Nesomyrmex echinatinodis 	Forel, 1886
 * Nesomyrmex pleuriticus 	Kempf, 1959
 * Nesomyrmex rutilans 	Kempf, 1958
 * Nesomyrmex spininodis 	Mayr, 1887

Nomamyrmex

 * Nomamyrmex esenbeckii 	Westwood, 1842
 * Nomamyrmex hartigii 	Westwood, 1842

Nylanderia

 * Nylanderia caeciliae 	Forel, 1899
 * Nylanderia fulva 	Mayr, 1862
 * Nylanderia lietzi 	Forel, 1908

Ochetomyrmex

 * Ochetomyrmex neopolitus 	Fernández, 2003
 * Ochetomyrmex semipolitus 	Mayr, 1878

Octostruma

 * Octostruma amrishi (Makhan, 2007)
 * Octostruma balzani (Emery, 1894)
 * Octostruma batesi (Emery, 1894)

Odontomachus

 * Odontomachus biumbonatus 	Brown, 1976
 * Odontomachus chelifer 	Latreille, 1802
 * Odontomachus haematodus 	Linnaeus, 1758
 * Odontomachus hastatus 	Fabricius, 1804
 * Odontomachus laticeps 	Roger, 1861
 * Odontomachus mayi 	Mann, 1912
 * Odontomachus scalptus 	Brown, 1978

Pachycondyla

 * Pachycondyla crassinoda 	Latreille, 1802
 * Pachycondyla harpax 	Fabricius, 1804
 * Pachycondyla impressa 	Roger, 1861

Paraponera

 * Paraponera clavata 	Fabricius, 1775

Paratrachymyrmex

 * Paratrachymyrmex bugnioni (Forel, 1912)
 * Paratrachymyrmex cornetzi (Forel, 1912)
 * Paratrachymyrmex diversus (Mann, 1916)
 * Paratrachymyrmex phaleratus (Wheeler, W.M., 1925)

Paratrechina

 * Paratrechina longicornis 	Latreille, 1802

Pheidole

 * Pheidole ademonia 	Wilson, 2003
 * Pheidole astur 	Wilson, 2003
 * Pheidole carapuna 	Mann, 1916
 * Pheidole cataractae Wheeler, W.M., 1916
 * Pheidole cramptoni Wheeler, W.M., 1916
 * Pheidole cursor 	Wilson, 2003
 * Pheidole dorsata Wilson, 2003
 * Pheidole exigua 	Mayr, 1884
 * Pheidole fallax Mayr, 1870
 * Pheidole fimbriata 	Roger, 1863
 * Pheidole funki 	LaPolla, 2005
 * Pheidole gauthieri Forel, 1901
 * Pheidole longiscapa Forel, 1901
 * Pheidole mamore 	Mann, 1916
 * Pheidole megacephala 	Fabricius, 1793
 * Pheidole meinerti 	Forel, 1905
 * Pheidole meinertopsis 	Wilson, 2003
 * Pheidole microgyna 	Wheeler, W.M., 1928
 * Pheidole minutula Mayr, 1878
 * Pheidole neoschultzi LaPolla, 2006
 * Pheidole perpusilla 	Emery, 1894
 * Pheidole radoszkowskii 	Mayr, 1884
 * Pheidole scolioceps 	Wilson, 2003
 * Pheidole socrates 	Forel, 1912
 * Pheidole synarmata 	Wilson, 2003
 * Pheidole tachigaliae Wheeler, W.M., 1921
 * Pheidole transversostriata Mayr, 1887

Platythyrea

 * Platythyrea angusta 	Forel, 1901
 * Platythyrea punctata 	Smith, F., 1858
 * Platythyrea sinuata 	Roger, 1860

Pogonomyrmex

 * Pogonomyrmex naegelii Forel, 1878

Prionopelta

 * Prionopelta antillana 	Forel, 1909

Probolomyrmex

 * Probolomyrmex kelleri Oliveira & Feitosa, 2019

Procryptocerus

 * Procryptocerus attenuatus 	Smith, F., 1876
 * Procryptocerus hirsutus 	Emery, 1896
 * Procryptocerus hylaeus 	Kempf, 1951
 * Procryptocerus pictipes 	Emery, 1896
 * Procryptocerus subpilosus 	Smith, F., 1860

Pseudomyrmex

 * Pseudomyrmex beccarii 	Menozzi, 1935
 * Pseudomyrmex concolor 	Smith, F., 1860
 * Pseudomyrmex cubaensis (Forel, 1901)
 * Pseudomyrmex curacaensis 	Forel, 1912
 * Pseudomyrmex elongatulus 	Dalla Torre, 1892
 * Pseudomyrmex elongatus 	Mayr, 1870
 * Pseudomyrmex faber 	Smith, F., 1858
 * Pseudomyrmex filiformis 	Fabricius, 1804
 * Pseudomyrmex gracilis 	Fabricius, 1804
 * Pseudomyrmex laevigatus 	Smith, F., 1877
 * Pseudomyrmex maculatus 	Smith, F., 1855
 * Pseudomyrmex malignus 	Wheeler, W.M., 1921
 * Pseudomyrmex niger 	Donisthorpe, 1940
 * Pseudomyrmex nigrocinctus 	Emery, 1890
 * Pseudomyrmex oculatus 	Smith, F., 1855
 * Pseudomyrmex penetrator 	Smith, F., 1877
 * Pseudomyrmex spiculus 	Ward, 1989
 * Pseudomyrmex tenuis 	Fabricius, 1804
 * Pseudomyrmex tenuissimus 	Emery, 1906
 * Pseudomyrmex termitarius 	Smith, F., 1855
 * Pseudomyrmex triplaridis (Forel, 1904)
 * Pseudomyrmex unicolor Smith, F., 1855
 * Pseudomyrmex urbanus 	Smith, F., 1877
 * Pseudomyrmex viduus (Smith, F., 1858)

Pseudoponera

 * Pseudoponera stigma 	Fabricius, 1804

Rasopone

 * Rasopone arhuaca Forel, 1901

Rogeria

 * Rogeria alzatei Kugler, C., 1994
 * Rogeria blanda (Smith, F., 1858)
 * Rogeria curvipubens Emery, 1894
 * Rogeria innotabilis Kugler, C., 1994
 * Rogeria lirata Kugler, 1994
 * Rogeria micromma Kempf, 1961
 * Rogeria prominula Kugler C., 1994
 * Rogeria scobinata Kugler, 1994
 * Rogeria tsumani LaPolla & Sosa-Calvo, 2006

Sericomyrmex

 * Sericomyrmex bondari Borgmeier, 1937
 * Sericomyrmex lutzi Wheeler, W.M., 1916
 * Sericomyrmex mayriForel, 1912
 * Sericomyrmex parvulus Forel, 1912
 * Sericomyrmex saussurei Emery, 1894

Solenopsis

 * Solenopsis altinodis 	Forel, 1912
 * Solenopsis brevicornis 	Emery, 1888
 * Solenopsis corticalis Forel, 1881
 * Solenopsis franki Forel, 1908
 * Solenopsis geminata 	Fabricius, 1804
 * Solenopsis globularia (Smith, F., 1858)
 * Solenopsis helena Emery, 1895
 * Solenopsis minutissima 	Emery, 1906
 * Solenopsis picea Emery, 1896
 * Solenopsis saevissima 	Smith, F., 1855
 * Solenopsis sulfurea 	Roger, 1862

Strumigenys

 * Strumigenys acarai (Sosa-Calvo, Schultz & LaPolla, 2010)
 * Strumigenys alberti 	Forel, 1893
 * Strumigenys carinithorax 	Borgmeier, 1934
 * Strumigenys crassicornis Mayr, 1887
 * Strumigenys dahlanae Sosa-Calvo, Schultz & LaPolla, 2010
 * Strumigenys denticulata 	Mayr, 1887
 * Strumigenys depressiceps Weber, 1934
 * Strumigenys dolichognatha Weber, 1934
 * Strumigenys elongata 	Roger, 1863
 * Strumigenys godmani 	Forel, 1899
 * Strumigenys mariae Sosa-Calvo, Schultz & LaPolla 2010
 * Strumigenys perparva 	Brown, 1958
 * Strumigenys precava 	Brown, 1954
 * Strumigenys royi (Sosa-Calvo, Schultz & LaPolla, 2010)
 * Strumigenys thaxteri (Wheeler, W.M., 1916)
 * Strumigenys waiwai (Sosa-Calvo, Schultz & LaPolla, 2010)

Talaridris

 * Talaridris mandibularis 	Weber, 1941

Tapinoma

 * Tapinoma melanocephalum 	Fabricius, 1793

Tetramorium

 * Tetramorium bicarinatum (Nylander, 1846)

Thaumatomyrmex

 * Thaumatomyrmex atrox 	Weber, 1939

Tranopelta

 * Tranopelta gilva 	Mayr, 1866

Typhlomyrmex

 * Typhlomyrmex clavicornis 	Emery, 1906
 * Typhlomyrmex rogenhoferi 	Mayr, 1862

Wasmannia

 * Wasmannia rochai 	Forel, 1912
 * Wasmannia scrobifera 	Kempf, 1961
 * Wasmannia sigmoidea 	Mayr, 1884

Uncertain species
The following species was described from Guyana but its true identity is currently unknown.
 * Pheidole gigliolii Menozzi, 1935