Camponotus libanicus

Identification
Ionescu-Hirsch (2009) - C. libanicus belongs to the Camponotus kiesenwetteri species group (Radchenko, 1997c). The Israeli specimens vary in size and show allometric relationships between body measures. Major workers (n = 7) have a slightly wider than long head and a short scape (SL/HW = 0.86–1.01), while minor workers (n = 10) have a more elongate head (HW/HL = 0.91–0.96) and a relatively longer scape (SL/HW = 1.04–1.13). Petiolar scale thickness of C. libanicus is variable (0.29–0.35 mm), and its shape in lateral view varies from thin, anteriorly concave proximally and moderately convex distally, to thick and anteriorly strongly convex. C. libanicus specimens from Israel are similar to Camponotus nadimi from which they differ only by the anteriorly convex or truncated clypeus with slight indentation medially in major workers and gyne, as opposed to an anteromedially emarginated clypeus in C. nadimi, and by a generally thicker petiolar scale in media and minor workers (range = 0.29–0.35 mm, n = 12, as compared to 0.27–0.33 mm, n = 7). According to Emery (1915), C. libanicus is similar to Camponotus aegaeus from which it differs by the thickness of the petiolar scale. One minor worker of C. aegaeus from Turkey has a thin scale (0.26 mm), with the outline of the anterior surface straight proximally and slightly convex distally, identical in shape to the petiolar scale of a minor worker depicted by Emery (1915, fig. 2b, p. 5), and thinner than in the examined specimens of C. libanicus of similar size.

Israeli specimens show much greater variability of size than that described in Tohmé’s (1969b) redescription of the species, i.e., they are not monomorphic, as stated by Tohmé (1969b). Minor workers with thin petiolar scale cannot be distinguished from C. nadimi minors when collected in isolation.

Distribution
Ionescu-Hirsch (2009) - Eastern Mediterranean to Iran (Radchenko, 1997c; Paknia et al., 2008).

Distribution based on Regional Taxon Lists
Palaearctic Region: Cyprus, Greece, Iran, Israel, Lebanon, Turkey.

Nomenclature

 * . Camponotus libanicus André, 1881b: 54, pl. 3, figs. 14, 15 (w.) LEBANON.
 * Forel, 1889: 261 (w.); Forel, 1911d: 355 (q.); Tohmé, G. 1969: 11 (m.).
 * Combination in C. (Orthonotomyrmex): Forel, 1913d: 435; Santschi, 1921f: 310;
 * combination in C. (Myrmentoma): Emery, 1925b: 121.
 * Status as species: André, 1882a: 148 (in key); Forel, 1889: 261; Dalla Torre, 1893: 239; Emery, 1896d: 376 (in list); Forel, 1911d: 355; Emery, 1925b: 121; Finzi, 1930c: 24; Menozzi, 1936d: 304; Tohmé, G. 1969: 11; Agosti & Collingwood, 1987a: 58; Agosti & Collingwood, 1987b: 283 (in key); Kugler, J. 1988: 259; Bolton, 1995b: 108; Radchenko, 1996b: 1197 (in key); Radchenko, 1997b: 705; Petrov, 2006: 108 (in key); Paknia, et al. 2008: 153; Vonshak, et al. 2009: 39; Ionescu-Hirsch, 2010: 83; Paknia, et al. 2010: 31; Legakis, 2011: 31; Borowiec, L. & Salata, 2012: 479; Kiran & Karaman, 2012: 7; Karaman, C. & Aktaç, 2013: 50 (in key); Borowiec, L. 2014: 36; Tohmé, G. & Tohmé, 2014: 138; Lebas, et al. 2016: 136; Salata & Borowiec, 2018c: 43; Salata, Loss, et al. 2019: 93.
 * Senior synonym of nadimi: Salata, Loss, et al. 2019: 93.
 * Senior synonym of sahlbergi: Radchenko, 1996b: 1197 (in key); Radchenko, 1997b: 705; Kiran & Karaman, 2012: 7; Salata, Loss, et al. 2019: 93.
 * nadimi. Camponotus (Myrmentoma) nadimi Tohmé, G. 1969: 6, figs. 3, 4 (s.w.) LEBANON.
 * Status as species: Bolton, 1995b: 112; Vonshak, et al. 2009: 39; Ionescu-Hirsch, 2010: 84; Borowiec, L. 2014: 38; Tohmé, G. & Tohmé, 2014: 138.
 * Junior synonym of libanicus: Salata, Loss, et al. 2019: 93.
 * sahlbergi. Camponotus (Orthonotomyrmex) libanicus r. sahlbergi Forel, 1913d: 435 (s.w.) TURKEY.
 * Combination in C. (Myrmentoma): Emery, 1925b: 121.
 * Subspecies of libanicus: Emery, 1925b: 121; Bolton, 1995b: 121.
 * Junior synonym of libanicus: Radchenko, 1996b: 1197 (in key); Radchenko, 1997b: 705; Kiran & Karaman, 2012: 7.

Taxonomic Notes
Seifert (2019): This taxon has been described from Lebanon. The type specimens depicted in antweb.org under CASENT0913700 shows an absent metanotal depression, a very convex overall dorsal profile line of mesosoma as well as a much stronger sculpture and much denser pubescence on all body surfaces than observed in any member of the C. lateralis group.

Worker
Ionescu-Hirsch (2009) - TL = 5.1–7.3, HL = 1.31–1.80, HW = 1.21–1.82, EL = 0.32–0.41, SL = 1.35–1.64, ML = 2.19–2.54, PW = 1.17–1.50, mTbL = 1.02–1.39, hTbL = 1.45–1.85 (n = 20). Gyne: TL = 11.6, HL = 2.23, HW = 2.11, EL = 0.51, SL = 1.72, ML = 3.98, PW = 2.09, mTbL = 1.56, hTbL = 2.34 (n = 1).

References based on Global Ant Biodiversity Informatics

 * Bolu H., and I. Ozgen. 2018. Some observation about Formicidae (Hymenoptera) in almond agroecosystems in East-Southeast Anatolian region of Turkey. Mun. Ent. Zool. 13(2): 495-498.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Emery, C.. "Escursioni zoologiche del Dr. Enrico Festa nell'Isola di Rodi. XII. Formiche." Bollettino del Museo di Zoologia ed Anatomia Comparatadella Reale Università di Torino 30 (1915): 1-7.
 * Finzi B. 1930. Hymenopteren aus Palästina und Syrien. (Zoologische Studienreise von R. Ebner 1928 mit Unterstützung der Akademie der Wissenschaften in Wien.) Formicidae. Sitzungsberichte der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse. Abteilung I 139:22-24.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel, A.. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
 * Forel, A.. "Fourmis de la faune méditerranéenne récoltées par MM. U. et J. Sahlberg." Revue Suisse de Zoologie 21 (1913): 427-438.
 * Ionescu-Hirsch A. 2009. An annotated list of Camponotus of Israel (Hymenoptera: Formicidae), with a key and descriptions of new species. ISRAEL JOURNAL OF ENTOMOLOGY 39: 5798.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
 * Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
 * Paknia, O., A. G. Radchenko, and M. Pfeiffer. "New records of ants (Hymenoptera: Formicidae) from Iran." Asian Myrmecology 3, no. 29-38 (2010).
 * Radchenko A. G. 1997. A review of ants of the subgenus Myrmentoma, Genus Camponotus (Hymenoptera, Formicidae), from Asian Palaearctic. Entomological Review 77(8): 1050-1059.
 * Salata S., A. C. Loss, C. Karaman, K. Kiran, and L. Borowiec. 2019. Review of the Camponotus kiesenwetteri group (Hymenoptera, Formicidae) in the Aegean with the description of a new species. ZooKeys 899: 85-107.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Seifert B. 2019. A taxonomic revision of the members of the Camponotus lateralis species group (Hymenoptera: Formicidae) from Europe, Asia Minor and Caucasia. Soil Organisms 91:7–32.
 * Shiran E., M. S. Mossadegh, and M. Esfandiari. 2012. Mutualistic ants (Hymenoptera: Formicidae) associated with aphids in central and southwestern parts of Iran. J. Crop Prot. 2013, 2 (1): 1-12.
 * Tohme G. 1996. Formicidae. Etude de la diversité biologique n° 4 . Ministère de lAgriculture à Beyrouth (Eds.). P85-87.
 * Tohmé, G.. "Description d'espèces nouvelles de fourmis au Liban (Hymenoptera Formicoidea)." Publications de l'Universite Libanaise, Section des Sciences Naturelles 7 (1969): 1-15.
 * Vonshak M., and A. Ionescu-Hirsch. 2009. A checklist of the ants of Israel (Hymenoptera: Formicidae). Israel Journal of Entomology 39: 33-55.