Checklist of Temnothorax species

The following species and subspecies belong to the genus Temnothorax. Synonyms are listed under their senior names. For valid names only see Temnothorax species and for distribution information see Temnothorax species by Country.

A
abeli (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

adlerzi (Douwes, Jessen & Buschinger, 1988) (Greece)

adustus (Mackay, W.P., 2000) (Mexico, United States)

aeolius (Forel, 1911) (Greece, Israel, Turkey)

affinis (Mayr, 1855) (Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Cuba, Czech Republic, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Luxembourg, Montenegro, Netherlands, Poland, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland)

• Leptothorax tirolensis Gredler, 1858

africanus (Bernard, 1948) (Algeria)

alayoi (Baroni Urbani, 1978) (Cuba, Greater Antilles)

albipennis (Curtis, 1854) (Austria, Belgium, Czech Republic, France, Germany, Hungary, Iberian Peninsula, Italy, Netherlands, Poland, Portugal, Slovakia, Slovenia, Spain, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax tuberointerruptus Bondroit, 1918

albispinus (Wheeler, W.M., 1908) (Greater Antilles, Puerto Rico)

• Antillaemyrmex albispina pallipes Wheeler, W.M., 1931

alfacarensis Tinaut & Reyes-López, 2020 (Iberian Peninsula, Spain)

algerianus (Cagniant, 1968) (Algeria, Morocco)

algiricus (Forel, 1894) (Algeria, Balearic Islands, Croatia, Iberian Peninsula, Morocco, Spain, Tunisia)

• Leptothorax convexus timida Santschi, 1912

• Leptothorax gazella Santschi, 1923

• Leptothorax gazella monticola Santschi, 1923

alienus Schulz, Heinze & Pusch, 2007 (Italy)

alinae (Radchenko, 1994) (Russian Federation)

allardycei (Mann, 1920) (Bahamas, United States)

• Antillaemyrmex floridanus Wheeler, W.M., 1931

almeqeri Sharaf, 2019 (Saudi Arabia)

alpinus (Ruzsky, 1902) (Georgia, Russian Federation)

• Leptothorax werneri Radchenko, 1994

ambiguus (Emery, 1895) (Canada, United States)

• Leptothorax ambiguus pinetorum Wesson, L.G. & Wesson, R.G., 1940

• Leptothorax foveata Smith, M.R., 1934

americanus (Emery, 1895) (United States)

anacanthus (Santschi, 1912) (Morocco)

• Leptothorax hesperius inermis Santschi, 1909

anacanthus maximus (Santschi, 1931) (Morocco)

anaphalantus Snelling, Borowiec & Prebus, 2014 (Mexico, United States)

andersoni (Mackay, W.P., 2000) (Mexico, United States)

andrei (Emery, 1895) (Mexico, United States)

• Leptothorax angustulus bordagei Bondroit, 1918

• Leptothorax nitens heathii Wheeler, W.M., 1903

• Leptothorax nitens occidentalis Wheeler, W.M., 1903

• Leptothorax ocellatus Mackay, 2000

androsanus (Wheeler, W.M., 1905) (Bahamas, Cuba, Mexico)

angulinodis Csösz, Heinze & Mikó, 2015 (Greece)

angulohumerus Zhou, Huang, Yu & Liu, 2010 (China)

angustifrons Csösz, Heinze & Mikó, 2015 (Greece, Turkey)

angustulus (Nylander, 1856) (Balearic Islands, Croatia, France, Iberian Peninsula, Libya, Malta, Portugal, Republic of Macedonia, Spain)

angustulus angustinodis (Stitz, 1917) (Libya)

anira (Terayama & Onoyama, 1999) (Japan)

annexus (Baroni Urbani, 1978) (Mexico)

• Macromischa annectens Wheeler, W.M., 1931

annibalis (Santschi, 1909) (Tunisia)

annibalis numidicus (Santschi, 1929) (Tunisia)

anodonta (Arnoldi, 1977) (Armenia, Iran)

anodontoides (Dlussky & Zabelin, 1985) (Turkmenistan)

ansei Catarineu, Barberá & Reyes-López, 2017 (Iberian Peninsula, Spain)

antera (Terayama & Onoyama, 1999) (Japan)

antigoni (Forel, 1911) (Greece, Turkey)

arabicus Sharaf & Akbar, 2017 (Saudi Arabia)

arboreus Snelling, Borowiec & Prebus, 2014 (United States)

arcanus (Kutter, 1973) (Switzerland)

archangelskiji (Kuznetsov-Ugamsky, 1926) (Kazakhstan)

arenarius (Santschi, 1908) (Israel, Tunisia)

arenarius fusciventris (Santschi, 1911) (Tunisia)

argentipes (Wheeler, W.M., 1928) (China)

ariadnae Csösz, Heinze & Mikó, 2015 (Greece)

arimensis (Azuma, 1977) (Japan)

arnoldii Radchenko & Fedoseeva, 2015 (Russian Federation)

arpini (Tarbinsky, 1976) (Kyrgyzstan)

• Leptothorax narinicus Tarbinsky, 1976

artvinensis Seifert, 2006 (Turkey)

atlantis (Santschi, 1911) (Algeria, Iberian Peninsula, Portugal, Spain)

atlantis brunea (Santschi, 1918) (Algeria, Tunisia)

atlantis silvanus (Forel, 1907) (Algeria, Morocco)

atlantis suturalis (Santschi, 1931) (Morocco)

atlantis veneris (Santschi, 1931) (Tunisia)

atomus (Cagniant & Espadaler, 1997) (Morocco)

augusti (Baroni Urbani, 1978) (Mexico)

• Leptothorax petiolatus Forel, 1901

• Macromischa foreli Aguayo, 1931

auresianus (Santschi, 1929) (Algeria)

aveli (Bondroit, 1918) (Andorra, Balearic Islands, Democratic Peoples Republic of Korea, France, Hungary, Iberian Peninsula, Malta, Portugal, Republic of Korea, Spain)

• Leptothorax aveli pici Bondroit, 1918

• Leptothorax berlandi Bondroit, 1918

• Leptothorax curvithorax Bondroit, 1918

• Leptothorax rabaudi Bondroit, 1918

aztecus (Wheeler, W.M., 1931) (Cuba, Guatemala, Mexico)

• Macromischa azteca maya Wheeler, W.M., 1937

B
baeticus (Emery, 1924) (Gibraltar, Iberian Peninsula, Spain)

banao (Fontenla Rizo, 1998) (Cuba)

barbouri (Aguayo, 1931) (Cuba, Greater Antilles)

barroi (Aguayo, 1931) (Cuba, Greater Antilles)

barryi (Cagniant, 1967) (Algeria)

basara (Terayama & Onoyama, 1999) (Japan)

bejaraniensis Reyes-López & Carpintero-Ortega, 2013 (Iberian Peninsula, Spain)

bermudezi (Wheeler, W.M., 1931) (Cuba, Greater Antilles)

• Croesomyrmex bermudezi mutabilis Wheeler, W.M., 1931

• Leptothorax anemicus Baroni Urbani, 1978

• Macromischa bermudezi gracilis Aguayo, 1932

bernardi (Espadaler, 1982) (Iberian Peninsula, Spain)

bestelmeyeri (Mackay, W.P., 2000) (United States)

bicolor (Mackay, W.P., 2000) (Mexico)

bikara (Terayama & Onoyama, 1999) (Japan)

bimbache Espadaler, 2007 (Canary Islands)

birgitae (Schulz, 1994) (Canary Islands)

blascoi (Espadaler, 1997) (Iberian Peninsula, Spain)

brackoi Salata & Borowiec, 2019 (Croatia, Greece)

bradleyi (Wheeler, W.M., 1913) (United States)

brauneri (Ruzsky, 1905) (Georgia, Russian Federation)

brevidentis Prebus, 2015 (Kenya)

brevispinosus (Mackay, W.P., 2000) (Mexico)

brevispinus (Chang & He, 2001) (China)

bristoli (Mackay, W.P., 2000) (Mexico, United States)

bruneri (Mann, 1924) (Cuba, Greater Antilles)

• Macromischa bruneri imitatrix Wheeler, W.M., 1937

• Macromischa bruneri ornatipes Wheeler, W.M., 1937

• Macromischa bruneri proxima Wheeler, W.M., 1937

brunneus (Cagniant, 1985) (Algeria, Morocco)

bucheti (Santschi, 1909) (Morocco)

bucheti festai (Emery, 1924) (Libya)

bucheti mixtus (Santschi, 1929) (Morocco)

bucheti tuneticus (Santschi, 1918) (Tunisia)

bugnioni (Forel, 1894) (Algeria)

bulgaricus (Forel, 1892) (Bulgaria, Greece, Israel, Republic of Macedonia)

bulgaricus cypridis (Santschi, 1930) (Cyprus)

C
cabrerae (Forel, 1893) (Canary Islands)

• Leptothorax cabrerae stitzi Santschi, 1918

• Leptothorax denticulatus Stitz, 1917

caesari (Espadaler, 1997) (Iberian Peninsula, Spain)

cagnianti (Tinaut, 1983) (Iberian Peninsula, Spain)

caguatan Snelling, Borowiec & Prebus, 2014 (Mexico, United States)

canescens (Santschi, 1908) (Canary Islands)

carinatus (Cole, 1957) (Mexico, United States)

cenatus (Bolton, 1982) (Kenya)

chandleri (Mackay, W.P., 2000) (United States)

ciferrii (Menozzi & Russo, 1930) (Dominican Republic, Greater Antilles, Lesser Antilles)

clypeatus (Mayr, 1853) (Austria, Bulgaria, Croatia, Czech Republic, France, Greece, Hungary, Iberian Peninsula, Poland, Romania, Russian Federation, Slovenia, Spain, Switzerland, Ukraine)

cokendolpheri (Mackay, W.P., 2000) (United States)

coleenae (Mackay, W.P., 2000) (United States)

conatensis Galkowski & Lebas, 2016 (France)

confucii (Forel, 1912) (Taiwan)

congruus (Smith, F., 1874) (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

continentalis Galkowski & Cagniant, 2017 (France, Spain)

convexus (Forel, 1894) (Algeria, Morocco)

• Leptothorax submuticus Emery, 1915

cordieri (Bondroit, 1918) (France)

cornibrevis (Collingwood, 1961) (Afghanistan)

corsicus (Emery, 1895) (Croatia, France)

corticalis (Schenck, 1852) (Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, Sweden, Switzerland, Ukraine)

crasecundus Seifert & Csősz, 2015 (Armenia, Bulgaria, Georgia, Greece, Romania, Russian Federation, Turkey, Ukraine)

crassispinus (Karavaiev, 1926) (Albania, Austria, Belarus, Bosnia and Herzegovina, Bulgaria, Croatia, Czech Republic, Estonia, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Montenegro, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Ukraine)

• Leptothorax nylanderi slavonicus Seifert, 1995

crassistriatus Salata, Borowiec & Trichas, 2018 (Greece)

creightoni (Mann, 1929) (Cuba, Greater Antilles)

creolus (Baroni Urbani, 1978) (Dominican Republic, Greater Antilles)

crepuscularis (Tinaut, 1994) (Iberian Peninsula, Spain)

cristinae (Espadaler, 1997) (Iberian Peninsula, Spain)

cuneinodis Radchenko, 2004 (Democratic Peoples Republic of Korea)

curtisetosus Salata & Borowiec, 2015 (Greece, Turkey)

curtulus (Santschi, 1929) (Morocco, Spain)

curtulus gentilis (Santschi, 1923) (Tunisia)

curvispinosus (Mayr, 1866) (United States)

• Stenamma gallarum Patton, 1879

cuyaguateje (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

D
daidalosi Salata, Borowiec & Trichas, 2018 (Greece)

darlingtoni (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

• Macromischa opalina Wheeler, W.M., 1937

delaparti (Forel, 1890) (Algeria)

• Leptothorax emeryi Santschi, 1923

desertorum Dlussky & Soyunov, 1988 (Russian Federation, Turkmenistan)

desioi (Menozzi, 1939) (India, Pakistan)

desioi melanicus (Menozzi, 1939) (India)

dessyi (Menozzi, 1936) (Greece)

discoloratus (Arnoldi, 1977) (Russian Federation)

dissimilis (Aguayo, 1932) (Cuba, Greater Antilles)

• Macromischa chloana Wheeler, W.M., 1937

ditifet (Espadaler, 1997) (Morocco)

dlusskyi Radchenko, Yusupov & Fedoseeva, 2015 (Russian Federation)

duloticus (Wesson, L.G., 1937) (United States)

E
eburneipes (Wheeler, W.M., 1927) (China, Democratic Peoples Republic of Korea)

elmenshawyi Sharaf, Wachkoo & Hita Garcia, 2019 (Saudi Arabia)

emmae (Mackay, W.P., 2000) (Mexico, United States)

exilis (Emery, 1869) (Algeria, Balearic Islands, Croatia, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Italy, Malta, Montenegro, Portugal, Republic of Macedonia, Slovenia, Spain, Tunisia, Turkey)

• Leptothorax exilis boccaris Santschi, 1923

• Leptothorax exilis creticus Forel, 1910

• Leptothorax exilis darii Forel, 1911

• Leptothorax exilis nitidulus Santschi, 1923

• Leptothorax exilis specularis Emery, 1916

• Leptothorax tuberum ruficornis Emery, 1914

F
finzii (Menozzi, 1925) (Italy)

flavicornis (Emery, 1870) (Bulgaria, Croatia, France, Greece, Italy, Montenegro, Republic of Macedonia, Slovenia, Switzerland, Turkey)

flavidulus (Wheeler, W.M. & Mann, 1914) (Greater Antilles, Haiti)

flavispinus (André, 1883) (Gibraltar, Iberian Peninsula, Israel, Spain)

• Leptothorax nigrita curvispinosus André, 1881

flavispinus amilcaris (Santschi, 1939) (Tunisia)

flavispinus clivulus (Santschi, 1939) (Tunisia)

flavispinus crassispinus (Cagniant, 1970) (Tunisia)

flavispinus impurus (Santschi, 1939) (Tunisia)

flavispinus rufescens (Forel, 1907) (Tunisia)

foreli (Santschi, 1907) (Tunisia)

formosus (Santschi, 1909) (Iberian Peninsula, Morocco, Portugal, Spain)

• Leptothorax fuentei Santschi, 1919

fragosus (Mackay, W.P. & Buschinger, 2002) (Canada)

fultonii (Forel, 1902) (China, India)

fumosus (Ruzsky, 1923) (Russian Federation, Turkmenistan)

furunculus (Wheeler, W.M., 1909) (United States)

fuscatus (Mann, 1920) (Guatemala)

G
gaetulus (Santschi, 1923) (Morocco)

gallae (Smith, M.R., 1949) (United States)

georgicus Ward, Brady, Fisher & Schultz, 2014 (Armenia, Georgia)

• Leptothorax tamarae Radchenko, 1994

gibbifer (Baroni Urbani, 1978) (Cuba, Greater Antilles)

goniops (Baroni Urbani, 1978) (Mexico)

gordiagini (Ruzsky, 1902) (Bulgaria, Croatia, Greece, Italy, Montenegro, Russian Federation)

• Myrmetaerus microcellatus Soudek, 1925

• Myrmoxenus gordiagini menozzii Finzi, 1924

gracilicornis (Emery, 1882) (Canary Islands)

gracilicornis nivarianus (Santschi, 1925) (Canary Islands)

• Leptothorax nivarianus guancha Santschi, 1925

graecus (Forel, 1911) (Bulgaria, Croatia, Greece, Republic of Macedonia)

gredosi (Espadaler & Collingwood, 1982) (Andorra, France, Iberian Peninsula, Spain)

grouvellei (Bondroit, 1918) (France, Iberian Peninsula, Spain)

gundlachi (Wheeler, W.M., 1913) (Cuba, Greater Antilles)

• Croesomyrmex aguayoi Wheeler, W.M., 1931

• Croesomyrmex aguayoi archeri Wheeler, W.M., 1931

• Macromischa aguayoi natenzoni Aguayo, 1931

• Macromischa alayorum Fontenla Rizo, 1993

• Macromischa bierigi Santschi, 1931

H
hadrumetensis (Santschi, 1918) (Tunisia)

haira (Terayama & Onoyama, 1999) (Japan)

helenae Csösz, Heinze & Mikó, 2015 (Bulgaria, Greece, Turkey)

hengshanensis (Huang, Chen & Zhou, 2004) (China)

hesperius (Santschi, 1909) (Canary Islands)

himachalensis Bharti, Gul & Schulz, 2012 (India)

himalaicum Yusupov, Lakho & Dubovikoff, 2020 (Pakistan)

hispaniolae (Baroni Urbani, 1978) (Dominican Republic, Greater Antilles)

hispidus (Cole, 1957) (Mexico, United States)

huatuo Terayama, 2009 (Taiwan)

huehuetenangoi (Baroni Urbani, 1978) (Guatemala)

hyrcanus (Dubovikoff & Radchenko, 2010) (Azerbaijan)

I
ibericus (Menozzi, 1922) (Iberian Peninsula, Spain)

ikarosi Salata, Borowiec & Trichas, 2018 (Greece)

imias (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

incompletus Salata, Borowiec & Trichas, 2018 (Greece)

indra (Terayama & Onoyama, 1999) (Japan)

inermis (Forel, 1902) (India)

inquilinus Ward, Brady, Fisher & Schultz, 2014 (Ukraine)

• Chalepoxenus tauricus Radchenko, 1989

interruptus (Schenck, 1852) (Armenia, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Germany, Hungary, Iberian Peninsula, Montenegro, Netherlands, Norway, Poland, Republic of Macedonia, Romania, Russian Federation, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax junipereti Arnoldi, 1977

• Leptothorax nikitae Arnoldi, 1977

• Leptothorax tuberoaffinis Bondroit, 1918

• Leptothorax tuberum knipovitshi Karavaiev, 1916

• Myrmica simpliciuscula Nylander, 1856

iranicus (Radchenko, 1994) (Iran)

iris (Roger, 1863) (Cuba, Greater Antilles)

• Croesomyrmex iris rufithorax Wheeler, W.M., 1931

• Croesomyrmex iris tristis Wheeler, W.M., 1931

• Macromischa iris nigripes Santschi, 1930

isabellae (Wheeler, W.M., 1908) (Greater Antilles, Puerto Rico)

• Leptothorax hyperisabellae Baroni Urbani, 1978

• Macromischa isabellae mutica Smith, M.R., 1937

italicus (Consani, 1952) (Croatia, Italy, Slovenia)

ixili (Baroni Urbani, 1978) (Guatemala)

J
jailensis (Arnoldi, 1977) (Hungary, Russian Federation, Ukraine)

janushevi (Radchenko, 1994) (Tajikistan)

josephi (Mackay, W.P., 2000) (United States)

K
kashmirensis Bharti, Gul & Schulz, 2012 (India)

kaszabi (Pisarski, 1969) (Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

• Leptothorax tuberum sachalinensis Kupyanskaya, 1990

kemali (Santschi, 1934) (Greece, Turkey)

khatrii Yusupov, Lakho & Dubovikoff, 2020 (Pakistan)

kinomurai (Terayama & Onoyama, 1999) (Japan)

kipyatkovi Yusupov, Dubovikoff & Lopatina, 2020 (India)

kirghizicus (Tarbinsky, 1976) (Kyrgyzstan)

kiudiria (Espadaler, 1997) (Morocco)

korbi (Emery, 1924) (Bulgaria, Georgia)

koreanus (Teranishi, 1940) (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea)

kraussei (Emery, 1915) (Croatia, France, Germany, Gibraltar, Greece, Iberian Peninsula, Italy, Slovenia, Spain, Turkey)

• Epimyrma foreli Menozzi, 1921

• Epimyrma vandeli Santschi, 1927

kubira (Terayama & Onoyama, 1999) (Japan)

kuixing Terayama, 2009 (Taiwan)

kurilensis (Radchenko, 1994) (Russian Federation)

kutteri (Cagniant, 1973) (France, Iberian Peninsula, Spain)

L
laciniatus (Stitz, 1917) (Algeria)

laconicus Csősz, Seifert et al., 2013 (Greece)

laestrygon (Santschi, 1931) (Italy, Malta)

laetus (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

• Leptothorax similis Baroni Urbani, 1978

• Macromischa affinis Mann, 1920

• Macromischa affinis maerens Wheeler, W.M., 1937

• Macromischa affinis umbratipes Wheeler, W.M., 1937

lagrecai (Baroni Urbani, 1964) (Italy)

laurae (Emery, 1884) (Tunisia)

laurae antoniae (Cagniant, 1997) (Morocco)

laurae colettae (Santschi, 1915) (Tunisia)

laurae rosae (Santschi, 1915) (Tunisia)

leigong Terayama, 2009 (Taiwan)

leimu Terayama, 2009 (Taiwan)

leoni (Arnoldi, 1971) (Kazakhstan)

lereddei (Bernard, 1953) (Algeria)

leucacanthus (Baroni Urbani, 1978) (Mexico)

leviceps (Emery, 1898) (Italy, Spain)

• Leptothorax exilis dichroa Emery, 1916

leyeensis Zhou, Huang, Yu & Liu, 2010 (China)

lichtensteini (Bondroit, 1918) (Andorra, Austria, Balearic Islands, Bulgaria, Croatia, France, Greece, Iberian Peninsula, Italy, Liechtenstein, Montenegro, Republic of Macedonia, Slovenia, Spain, Turkey)

liebi (Mackay, W.P., 2000) (United States)

liviae (Agosti & Collingwood, 2011) (United Arab Emirates)

longipilosus (Santschi, 1912) (Tunisia)

longispinosus (Roger, 1863) (Canada, Iberian Peninsula, Spain, United States)

• Leptothorax longispinosus iowensis Buren, 1945

• Leptothorax longispinosus laeviceps Buren, 1944

lucidus Csösz, Heinze & Mikó, 2015 (Greece, Turkey)

luteus (Forel, 1874) (Albania, Austria, France, Greece, Iberian Peninsula, Israel, Romania, Spain, Switzerland, Syrian Arab Republic)

• Leptothorax tristis Bondroit, 1918

M
makora (Terayama & Onoyama, 1999) (Japan)

manni (Wheeler, W.M., 1914) (Cuba, Mexico)

maoerensis Zhou, Huang, Yu & Liu, 2010 (China)

marocanus (Santschi, 1909) (Morocco)

mauritanicus (Santschi, 1909) (Morocco)

mediterraneus Ward, Brady, Fisher & Schultz, 2014 (Andorra, Iberian Peninsula, Italy, Malta, Montenegro, Spain)

• Leptothorax angustulus kraussei Emery, 1916

megalops (Hamann & Klemm, 1967) (Sudan, United Arab Emirates)

mekira Terayama & Kubota, 2011 (Japan)

melas (Espadaler, Plateaux & Casevitz-Weulersse, 1984) (France)

melleus (Forel, 1904) (Kazakhstan, Kyrgyzstan, Mongolia, Turkmenistan, Uzbekistan)

• Leptothorax balchashensis Arnoldi, 1971

• Leptothorax melleus csikii Pisarski, 1969

melnikovi (Ruzsky, 1905) (Georgia, Russian Federation)

messiniaensis Salata & Borowiec, 2019 (Greece)

mexicanus (Mackay, W.P., 2000) (Mexico)

michali Radchenko, 2004 (Democratic Peoples Republic of Korea)

microreticulatus Bharti, Gul & Schulz, 2012 (India, Pakistan)

mimeuri (Cagniant, 1997) (Morocco)

minotaurosi Salata, Borowiec & Trichas, 2018 (Greece)

minozzii (Santschi, 1922) (Italy)

• Leptothorax silvestrii Menozzi, 1921

minutissimus (Smith, M.R., 1942) (United States)

mirabilis (Espadaler & Cagniant, 1996) (Morocco)

miserabilis (Santschi, 1918) (Tunisia)

mitsukoae Terayama & Yamane, 2013 (Japan)

mongolicus (Pisarski, 1969) (China, Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

monjauzei (Cagniant, 1968) (Algeria)

morea Csősz, Salata & Borowiec, 2018 (Albania, Bosnia and Herzegovina, Croatia, Greece, Montenegro, Serbia, Turkey)

morongo Snelling, Borowiec & Prebus, 2014 (Mexico, United States)

mortoni (Aguayo, 1937) (Cuba, Greater Antilles)

mpala Prebus, 2015 (Kenya)

muellerianus (Finzi, 1922) (Bulgaria, Croatia, Greece, Iberian Peninsula, Italy, Portugal, Slovenia, Spain, Switzerland, Turkey)

• Chalepoxenus gribodoi Menozzi, 1923

• Chalepoxenus insubricus Kutter, 1950

• Chalepoxenus siciliensis Kutter, 1973

myersi (Wheeler, W.M., 1931) (Cuba, Greater Antilles)

myrmiciformis Snelling, Borowiec & Prebus, 2014 (Mexico, United States)

N
nadigi (Kutter, 1925) (Andorra, Armenia, Bulgaria, Georgia, Iberian Peninsula, Iran, Iran, Poland, Republic of Macedonia, Spain, Switzerland)

• Leptothorax caucasicus Arnoldi, 1977

• Leptothorax hasardaghi Dlussky & Zabelin, 1985

naeviventris (Santschi, 1910) (Iberian Peninsula, Spain, Tunisia)

naeviventris kefensis (Santschi, 1929) (Tunisia)

nassonovi (Ruzsky, 1895) (Democratic Peoples Republic of Korea, Kyrgyzstan, Mongolia, Republic of Korea, Russian Federation, Ukraine)

• Leptothorax galeatus Wheeler, W.M., 1927

• Leptothorax nassonovi firssovi Kuznetsov-Ugamsky, 1928

• Leptothorax nassonovi subnudus Ruzsky, 1905

neminan (Espadaler, 1997) (Morocco)

neomexicanus (Wheeler, W.M., 1903) (Mexico, United States)

nevadensis (Wheeler, W.M., 1903) (Mexico, United States)

• Leptothorax eldoradensis Wheeler, W.M., 1915

• Leptothorax lindae Mackay, W.P., 2000

• Leptothorax maryanae Mackay, W.P., 2000

• Leptothorax melanderi Wheeler, W.M., 1909

niger (Forel, 1894) (Andorra, France, Greece, Iberian Peninsula, Israel, Spain)

• Leptothorax glaber Santschi, 1925

nigricans (Baroni Urbani, 1978) (Cuba, Greater Antilles)

• Macromischa melanocephala Wheeler, W.M., 1931

nigriceps (Mayr, 1855) (Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Montenegro, Netherlands, Poland, Republic of Macedonia, Romania, Slovakia, Slovenia, Spain, Switzerland)

• Leptothorax tuberum tuberonigriceps Forel, 1874

nigritus (Emery, 1878) (Algeria, Georgia)

nigritus rupestris (Forel, 1909) (Algeria)

nigritus saharensis (Santschi, 1929) (Algeria)

nigritus salambo (Santschi, 1912) (Tunisia)

nipensis (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

nitens (Emery, 1895) (Mexico, United States)

• Leptothorax melinus Mackay, W.P., 2000

• Leptothorax nitens mariposa Wheeler, W.M., 1917

nitidiceps (Dalla Torre, 1893) (Algeria, Morocco, Tunisia)

nordmeyeri (Schulz, 1997) (India)

normandi (Santschi, 1912) (Tunisia)

nuwuvi Snelling, Borowiec & Prebus, 2014 (United States)

nylanderi (Foerster, 1850) (Andorra, Armenia, Austria, Belgium, Bulgaria, Denmark, France, Georgia, Germany, Iberian Peninsula, Jersey, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Russian Federation, Spain, Sweden, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax nylanderi nylanderotuberum Ruzsky, 1902

• Leptothorax tuberum nylanderocorticalis Forel, 1874

• Myrmica cingulata Schenck, 1852

nylanderi nylanderonigriceps (Stitz, 1939) (Germany)

O
obliquicanthus (Cole, 1953) (Mexico, United States)

obscurior (Dalla Torre, 1893) (Algeria, Morocco)

• Leptothorax exilis retifer Santschi, 1929

obturator (Wheeler, W.M., 1903) (United States)

ocarinae (Baroni Urbani, 1978) (Mexico)

opaciabdomin (Chang & He, 2001) (China)

oraniensis (Forel, 1894) (Algeria, Tunisia)

• Leptothorax rusticus Santschi, 1918

• Leptothorax rusticus chobauti Santschi, 1923

• Leptothorax rusticus sergensis Santschi, 1929

orchidus Zhou, Huang, Yu & Liu, 2010 (China)

oxianus (Ruzsky, 1905) (Kyrgyzstan, Uzbekistan)

• Leptothorax oxianus juglandeti Arnoldi, 1976

• Leptothorax talassicus Tarbinsky, 1976

oxynodis (Mackay, W.P., 2000) (United States)

P
paiute Snelling, Borowiec & Prebus, 2014 (United States)

pakistanensis Rasheed, Bodlah, Magomedovich, Fareen, Bodlah, Prebus & Wachkoo, 2020

pallidipes (Santschi, 1910) (Tunisia)

pallidus (Collingwood, 1961) (Afghanistan)

palustris (Deyrup & Cover, 2004) (United States)

pamiricus (Ruzsky, 1902) (Pakistan, Russian Federation)

pan (Santschi, 1936) (Morocco)

pardoi (Tinaut, 1987) (France, Gibraltar, Iberian Peninsula, Morocco, Portugal, Spain)

parvulus (Schenck, 1852) (Andorra, Armenia, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Montenegro, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Turkey, Turkmenistan)

pastinifer (Emery, 1894) (Bahamas, Cuba, Greater Antilles)

• Leptothorax pastoris Baroni Urbani, 1978

• Macromischa lucayensis Forel, 1901

• Macromischa pastinifera opacipes Wheeler, W.M., 1905

pelagosanus (Müller, 1923) (Croatia, Italy, Republic of Macedonia)

peninsularis (Wheeler, W.M., 1934) (Mexico)

pergandei (Emery, 1895) (Mexico, United States)

• Leptothorax floridanus Emery, 1895

• Leptothorax manni Wesson, L.G., 1935

• Leptothorax pergandei flavus Smith, M.R., 1929

personatus (Cagniant, 1987) (Morocco)

peyerimhoffi (Santschi, 1929) (Algeria)

pilagens Seifert, Kleeberg, Feldmeyer, Pamminger, Jongepier & Foitzik, 2014 (United States)

pisarskii Radchenko, 2004 (China, Democratic Peoples Republic of Korea)

platycephalus (Espadaler, 1997) (Iberian Peninsula, Spain)

platycnemis (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

poeyi (Wheeler, W.M., 1913) (Cuba, Greater Antilles)

• Macromischa poeyi rugiceps Aguayo, 1932

politus (Smith, M.R., 1939) (Mexico, United States)

porphyritis (Roger, 1863) (Cuba, Greater Antilles)

• Macromischa jaumei Santschi, 1930

• Macromischa porphyritis latispina Wheeler, W.M., 1931

productus (Santschi, 1918) (Canary Islands, Morocco)

proteii Salata, Borowiec & Trichas, 2018 (Greece)

pseudandrei Snelling, Borowiec & Prebus, 2014 (United States)

pulchellus (Emery, 1894) (Lesser Antilles)

punctaticeps (Mackay, W.P., 2000) (Mexico)

punctatissimus (Mackay, W.P., 2000) (Mexico)

punctithorax (Mackay, W.P., 2000) (Mexico)

punicans (Roger, 1863) (Cuba, Greater Antilles)

pupseli (Santschi, 1909) (Tunisia)

purpuratus (Roger, 1863) (Cuba, Greater Antilles)

Q
quasimodo Snelling, Borowiec & Prebus, 2014 (United States)

R
racovitzai (Bondroit, 1918) (Balearic Islands, France, Gibraltar, Iberian Peninsula, Montenegro, Portugal, Spain, Switzerland)

• Leptothorax massiliensis Bondroit, 1918

ravouxi (André, 1896) (Austria, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Iberian Peninsula, Italy, Poland, Russian Federation, Slovenia, Spain, Switzerland)

• Epimyrma goesswaldi Menozzi, 1930

recedens (Nylander, 1856) (Albania, Armenia, Balearic Islands, Bulgaria, Croatia, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Israel, Italy, Malta, Montenegro, Portugal, Republic of Macedonia, Russian Federation, Slovenia, Spain, Switzerland, Tunisia, Turkmenistan)

• Leptothorax ergatogyna Bernard, 1950

• Leptothorax mordax Santschi, 1919

• Leptothorax pictus Emery, 1924

• Leptothorax recedens barbarus Santschi, 1939

reduncus (Wang, M. & Wu, J., 1988) (China)

reticulatus (Chang & He, 2001) (China)

risii (Forel, 1892) (Canary Islands)

• Leptothorax elongatus Santschi, 1919

• Leptothorax risii armouri Wheeler, W.M., 1927

• Leptothorax risii globulinodis Santschi, 1909

rogeri Emery, 1869 (Greece)

rothneyi (Forel, 1902) (India)

rottenbergii (Emery, 1870) (Canary Islands, Croatia, Greece, Italy)

rottenbergii ernesti (Santschi, 1909) (Algeria)

rottenbergii scabriosus (Santschi, 1909) (Algeria, Canary Islands)

rougeti (Bondroit, 1918) (France)

rudis (Wheeler, W.M., 1917) (United States)

rufus Prebus, 2015 (Kenya)

rugatulus (Emery, 1895) (Canada, Mexico, United States)

• Leptothorax curvispinosus annectens Wheeler, W.M., 1903

• Leptothorax rugatulus brunnescens Wheeler, W.M., 1917

• Leptothorax rugatulus cockerelli Wheeler, W.M., 1917

• Leptothorax rugatulus dakotensis Wheeler, G.C. & Wheeler, E.W., 1944

• Leptothorax rugatulus mediorufus Wheeler, W.M., 1917

ruginosus Zhou, Huang, Yu & Liu, 2010 (China)

rugithorax (Mackay, W.P., 2000) (Mexico)

rugosus (Mackay, W.P., 2000) (Mexico)

rugulosus (Mackay, W.P., 2000) (Mexico)

S
sallei (Guérin-Méneville, 1852) (Dominican Republic, Greater Antilles, Haiti)

• Macromischa sallei haytiana Wheeler, W.M. & Mann, 1914

• Macromischa sallei opacinoda Wheeler, W.M., 1931

• Myrmica russula Nylander, 1856

salvini (Forel, 1899) (Costa Rica, Guatemala, Honduras, Mexico, Panama)

salvini obscurior (Forel, 1899) (Panama)

santra (Terayama & Onoyama, 1999) (Japan)

santschii (Forel, 1905) (Algeria, Morocco, Tunisia)

• Leptothorax flavispinus boiteli Santschi, 1929

• Leptothorax flavispinus facetus Santschi, 1939

sardous (Santschi, 1909) (Italy)

satunini (Ruzsky, 1902) (Armenia, Georgia, Kyrgyzstan, Russian Federation)

saxatilis Schulz, Heinze & Pusch, 2007 (Italy)

saxonicus (Seifert, 1995) (Bulgaria, Czech Republic, Germany, Greece, Hungary, Montenegro, Poland, Slovenia)

schaufussi (Forel, 1879) (Iberian Peninsula, Spain)

schaumii (Roger, 1863) (United States)

• Leptothorax fortinodis Mayr, 1886

• Leptothorax fortinodis gilvus Wheeler, W.M., 1903

• Leptothorax fortinodis melanoticus Wheeler, W.M., 1903

schmittii (Wheeler, W.M., 1903) (Mexico, United States)

schoedli Seifert, 2006 (Turkey)

schurri (Forel, 1902) (India)

schwarzi (Mann, 1920) (Cuba, Greater Antilles)

semenovi (Ruzsky, 1903) (Kyrgyzstan, Russian Federation, Turkmenistan)

semiruber (André, 1881) (Bulgaria, Greece, Israel, Republic of Macedonia)

• Leptothorax rottenbergi balcanica Santschi, 1909

• Leptothorax rottenbergi jesus Forel, 1913

senectutis (Baroni Urbani, 1978) (Cuba, Greater Antilles)

sentosus Ward, Brady, Fisher & Schultz, 2014 (Kazakhstan)

• Leonomyrma spinosa Arnoldi, 1968

serviculus (Ruzsky, 1902) (Democratic Peoples Republic of Korea, Mongolia, Republic of Korea, Russian Federation)

sevanensis (Arnoldi, 1977) (Armenia)

shannxiensis Zhou, Huang, Yu & Liu, 2010 (China)

shelkovnikovi (Karavaiev, 1926) (Russian Federation)

silvestrii (Santschi, 1911) (Mexico, United States)

simesno (Espadaler, 1997) (Morocco)

similis Csösz, Heinze & Mikó, 2015 (Turkey)

simlensis (Forel, 1904) (India)

singularis (Radchenko, 1994) (Tajikistan)

skwarrae (Wheeler, W.M., 1931) (Mexico)

smithi (Baroni Urbani, 1978) (United States)

• Leptothorax wheeleri Smith, M.R., 1929

smyrnensis (Forel, 1911) (Greece, Turkey)

solerii (Menozzi, 1936) (Greece)

solidinodus Prebus, 2015 (Kenya)

sordidulus (Müller, 1923) (Austria, Bosnia and Herzegovina, Bulgaria, Croatia, Germany, Greece, Hungary, Italy, Montenegro, Republic of Macedonia, Slovenia)

• Leptothorax carinthiacus Bernard, 1957

• Leptothorax mulleri Santschi, 1926

spinosior (Forel, 1901) (China, Japan, Republic of Korea)

spinosus (Forel, 1909) (Algeria, Morocco)

• Leptothorax maurus Santschi, 1921

splendens (Wheeler, W.M., 1905) (Bahamas, Cuba, Greater Antilles)

• Macromischa scabripes Mann, 1920

• Macromischa scabripes brunneipes Wheeler, W.M., 1937

splendidiceps (Baroni Urbani, 1968) (Malta)

squamifer (Roger, 1863) (Cuba, Greater Antilles)

• Macromischa squamifera atrinodis Mann, 1920

steinbergi (Arnoldi, 1971) (Kazakhstan)

stenotyle (Cole, 1956) (United States)

• Leptothorax angustinodus Cole, 1956

stollii (Forel, 1885) (Guatemala)

striatulus (Stitz, 1937) (Guatemala, Honduras, Mexico)

striatus Zhou, Huang, Yu & Liu, 2010 (China)

strymonensis Csősz, Salata & Borowiec, 2018 (Albania, Bosnia and Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Serbia, Turkey)

stumperi (Kutter, 1950) (France, Switzerland)

subcingulatus (Emery, 1924) (Iberian Peninsula, Spain)

subditivus (Wheeler, W.M., 1903) (Costa Rica, Guatemala, Mexico, United States, Venezuela)

• Macromischa flavitarsis Mann, 1920

• Macromischa laevissima Wheeler, W.M., 1911

• Macromischa luciliae Mann, 1935

suberis (Forel, 1894) (Algeria)

subtilis Csösz, Heinze & Mikó, 2015 (Greece, Turkey)

susamyri (Dlussky, 1965) (Kyrgyzstan)

T
taivanensis (Wheeler, W.M., 1929) (China, Taiwan)

tamarae (Arnoldi, 1968) (Georgia)

tamriensis Ajerrar, Gómez, Bouharroud, Zaafrani & Cagniant, 2018 (Morocco)

tarbinskii (Arnoldi, 1976) (Kyrgyzstan)

• Leptothorax longipilosus Tarbinsky, 1976

tauricus (Ruzsky, 1902) (Armenia, Bulgaria, Greece, Russian Federation, Ukraine)

• Leptothorax pithyusae Arnoldi, 1977

• Leptothorax pithyusae babuganicus Arnoldi, 1977

tebessae (Forel, 1890) (Algeria)

tebessae gapsensis (Santschi, 1929) (Tunisia)

tembotovi Radchenko & Yusupov, 2015 (Russian Federation)

tenuisculptus (Baroni Urbani, 1978) (Mexico)

tenuispinus (Santschi, 1911) (Tunisia)

tergestinus (Finzi, 1928) (Bulgaria, Czech Republic, France, Germany, Greece, Hungary, Italy, Montenegro, Poland, Slovenia)

terricola (Mann, 1920) (Cuba, Greater Antilles)

terrigena (Wheeler, W.M., 1903) (Mexico, United States)

tesquorum (Arnoldi, 1977) (Georgia, Russian Federation, Turkmenistan)

• Leptothorax elderensis Dlussky & Zabelin, 1985

texanus (Wheeler, W.M., 1903) (Mexico, United States)

• Leptothorax texanus davisi Wheeler, W.M., 1905

theryi (Santschi, 1921) (Morocco)

tianpeng Terayama, 2009 (Taiwan)

tianschanicus (Tarbinsky, 1976) (Kyrgyzstan)

torrei (Aguayo, 1931) (Cuba, Greater Antilles, United States)

totonicapani (Baroni Urbani, 1978) (Guatemala)

trabutii (Forel, 1894) (Algeria)

• Leptothorax lindbergi Santschi, 1931

tramieri (Cagniant, 1983) (Morocco)

triangularis Salata & Borowiec, 2019 (Greece)

tricarinatus (Emery, 1895) (United States)

tricolor (Santschi, 1925) (Morocco)

tuberum (Fabricius, 1775) (Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, France, Georgia, Germany, Hungary, Iberian Peninsula, Iran, Jersey, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Korea, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland)

• Formica tuberosa Latreille, 1802

• Leptothorax melanocephalus Emery, 1870

• Leptothorax nigriceps pyrenaeus Bondroit, 1918

• Leptothorax pratostepposus Arnoldi, 1977

• Leptothorax tuberum acutinodis Arnoldi, 1977

• Leptothorax tuberum alaicus Tarbinsky, 1976

• Leptothorax tuberum ciscaucasicus Arnoldi, 1977

• Leptothorax tuberum nigricephala Karavaiev, 1930

• Leptothorax tuberum stipaceus Ruzsky, 1905

turcicus (Santschi, 1934) (Greece, Turkey)

turritellus (Espadaler, 1997) (Morocco)

tuscaloosae (Wilson, 1951) (United States)

tyndalei (Forel, 1909) (Algeria, Iberian Peninsula, Morocco, Spain)

• Leptothorax balli Santschi, 1939

• Leptothorax cervantesi Santschi, 1919

U
unifasciatus (Latreille, 1798) (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Channel Islands, Croatia, Czech Republic, Denmark, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Israel, Italy, Jersey, Kazakhstan, Liechtenstein, Lithuania, Luxembourg, Malta, Montenegro, Morocco, Netherlands, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Turkmenistan, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax anoplogynus Emery, 1869

• Leptothorax tuberum brauneri Karavaiev, 1937

• Leptothorax tuberum brauniella Radchenko, 1995

• Leptothorax tuberum kirillovi Ruzsky, 1905

• Leptothorax tuberum salina Karavaiev, 1937

• Leptothorax tuberum unifasciatointerruptus Forel, 1874

• Leptothorax unifasciatus ucrainicus Arnoldi, 1977

• Leptothorax volubilis Santschi, 1929

unifasciatus obenbergeri (Sadil, 1939) (Czech Republic)

unifasciatus staegeri (Bondroit, 1918) (Switzerland)

unifasciatus unifasciatonigricepoides Novak & Sadil, 1941 (Czech Republic)

unifasciatus unifasciatonigriceps (Novak & Sadil, 1941) (Germany)

universitatis (Espadaler, 1997) (Iberian Peninsula, Spain)

usunkul (Ruzsky, 1924) (Russian Federation)

V
variabilis Salata, Borowiec & Trichas, 2018 (Greece)

versicolor (Roger, 1863) (Cuba)

• Macromischa lugens Roger, 1863

• Macromischa lugens nigra Santschi, 1931

versicolor guanahacabibes (Fontenla Rizo, 1995) (Cuba, Greater Antilles)

villarensis (Aguayo, 1931) (Cuba, Greater Antilles)

• Leptothorax williami Baroni Urbani, 1978

• Macromischa manni Wheeler, W.M., 1931

violaceus (Mann, 1924) (Cuba, Greater Antilles)

volgensis (Ruzsky, 1905) (Russian Federation)

W
wardi Snelling, Borowiec & Prebus, 2014 (United States)

wheeleri (Mann, 1920) (Cuba, Greater Antilles)

• Macromischa wheeleri petri Aguayo, 1931

whitfordi (Mackay, W.P., 2000) (Mexico, United States)

wollastoni (Donisthorpe, 1940) (Portugal)

wroughtonii (Forel, 1904) (India)

wui (Wheeler, W.M., 1929) (China)

X
xanthos Radchenko, 2004 (China, Democratic Peoples Republic of Korea)

Y
yanwan Terayama, 2009 (Taiwan)

Z
zabelini (Radchenko, 1989) (Turkmenistan)

zaleskyi (Sadil, 1953) (Czech Republic)

zhejiangensis Zhou, Huang, Yu & Liu, 2010 (China)

=Fossil Species=

G
glaesarius (Wheeler, W.M., 1915)

gracilis (Mayr, 1868)

H
hystriculus (Wheeler, W.M., 1915)

L
longaevus (Wheeler, W.M., 1915)

P
petiolatus (Mayr, 1868)

placivus (Wheeler, W.M., 1915)

praecreolus (De Andrade, 1992)