Lasius lasioides

Nests in the presence of trees, from forests to open areas with at least a few trees present, across a wide range of habitats.

Identification
Seifert (2020) - Lasius lasioides shows all diagnostic characters of the L. brunneus species complex. It can be separated from its semipatric relatives Lasius brunneus and Lasius silvaticus by smaller head width, longer scape and larger torulo-clypeal distance.

Confusion with Lasius himalayanus is excluded by zoogeography. The potentially sympatric Iranian desert species Lasius excavates differs by head shape and coloration. Coloration in L. lasioides: head, mesosoma, gaster, femora and tibiae dark to medium brown; tibio-femoral joint region, scape and (frequently) anterior margin of clypeus pale yellowish-brown.

The Holomediterranean population of L. lasioides is polymorphic; there is in particular an extreme variance in scape length. This polymorphism shows a rather clear geographic structuring. Exclusively the long-scaped morph, which corresponds to the types of L. lasioides, was found in Italy and Greece and the islands of Mallorca, Malta, and Sardinia. Only the short-scaped morph, which corresponds to the types of Lasius barbarous, was found in the Spanish mainland and on Cyprus. Sympatric occurrence of both morphs is observed in Morocco, Algeria, Tunisia, Syria, and Israel. The Iberian shortscaped population and the Italian long-scaped population are in contact in southern France.

Distribution
Seifert (2020) - Lasius lasioides is a Holomediterranean species and presumably the lack of records from Libya and Egypt are caused by a lack of sampling. The most southwestern site is in Morocco at 30.8°N, 8.8° W. The northern distributional border runs along 43.5° N in France, 44.6° N in Italy, 40.5°N in Greece and Turkey whereas the easternmost truly Mediterranean site is at 35.6°N, 36.2°E in Syria. The distribution farther east is poorly known. Samples from two sites in the East Anatolian highland (PR_N 502, 39.43°N, 39.88°E, 1800 m; PR_N 460, 38.63°N, 43.45°E, 2300 m) might possibly represent a separate population or even species with special adaptation to a cold winter climate. A sample from Ghaemshar / Iran (36.46°N, 52.86°E, 49 m) represents the easternmost known site and raises the question if there is a continuous distribution from Syria and Israel east to the northern Iran. The highest latitude site in the Moroccan Atlas Mountains is at 33.00°N, 5.07°W, at 2240 m. It may be expected to have colonized all sufficiently large islands in the Mediterranean Sea which have some tree stands.

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Balearic Islands, Croatia, France, Georgia, Greece, Iberian Peninsula, Iran, Israel, Italy, Malta, Montenegro, Morocco, Portugal, Republic of Macedonia, Spain, Turkey.

Biology
Rigato & Toni (2011) - Species known from Mediterranean Europe, Anatolia and along the Atlas mountains (Morocco). This is the first published record for Sardinia. Pashaei Rad et al. (2018) found this species in Iran on parkland ground in a moderate rainfall area.

Seifert (2020) - Lasius lasioides is apparently dependent on the presence of trees. It inhabits diverse types of broad-leafed or coniferous forest, both such with closed canopies or more open stands, as well as urban areas, gardens, pastures and road sides with at least a few scattered trees. The nests may be found under bark of the trees, on the ground in dead logs, or under stones. Their behavior is similar to L. brunneus: it is fugitive and not aggressive and workers prefer runways in crevices of bark or other surface structures. Development of alates differs throughout the whole geographic range in dependence from latitude and altitude. Eleven observations of alates occurred 30 April – 29 July, ten of these before 8 July.

Nomenclature

 * . Prenolepis lasioides Emery, 1869b: 6, pl. 1, fig. 3 (w.q.m.) ITALY.
 * Combination in Lasius: Emery, 1869b: 25 (in text); Emery, 1870: 194;
 * combination in Lasius (Lasius): Forel, 1915d: 53;
 * combination in Donisthorpea: Donisthorpe, 1915d: 347;
 * combination in Formicina (Donisthorpea): Emery, 1916b: 241;
 * combination in Formicina: Bondroit, 1918: 27;
 * combination in Acanthomyops: Kuznetsov-Ugamsky, 1927e: 189;
 * combination in Acanthomyops (Donisthorpea): Donisthorpe, 1927b: 400;
 * combination in Lasius: Müller, 1923b: 126; Kuznetsov-Ugamsky, 1929b: 38.
 * As unavailable (infrasubspecific) name: Forel, 1904b: 386; Ruzsky, 1905b: 310.
 * Junior synonym of fumatus: Emery & Forel, 1879: 452.
 * [Note: the reference Emery & Forel, 1879: 452, reverses the seniority of lasioides established by Emery, 1870: 194, but does not affect the synonymy and is discounted here.]
 * Junior synonym of alienus: Wilson, 1955a: 77; Smith, D.R. 1979: 1435; Agosti & Collingwood, 1987a: 58; Arakelian, 1994: 117.
 * Subspecies of alienus: Dalla Torre, 1893: 182; Cagniant, 1964: 92; Forel, 1915d: 53 (in key); Emery, 1916b: 241; Menozzi, 1925d: 34; Zimmermann, 1935: 49; Finzi, 1936: 191.
 * Subspecies of niger: Forel, 1911d: 352; Forel, 1913d: 434; Forel, 1915d: 53 (in key); Donisthorpe, 1915d: 347; Emery, 1916b: 241; Santschi, 1921e: 170; Kulmatycki, 1922: 81; Finzi, 1923: 4; Emery, 1925b: 230; Schkaff, 1925: 275; Soudek, 1925b: 15; Donisthorpe, 1927b: 400; Kuznetsov-Ugamsky, 1927e: 189; Kuznetsov-Ugamsky, 1929b: 37; Finzi, 1936: 191; Bernard, 1945: 137.
 * Status as species: Emery, 1870: 194; Bondroit, 1918: 27; Stärcke, 1926: 124 (in key); Stärcke, 1944a: 155; Consani & Zangheri, 1952: 44; Seifert, 1992b: 8 (redescription); Bolton, 1995b: 223; Mei, 1995: 768; Poldi, et al. 1995: 7; Schembri & Collingwood, 1995: 156; Bračko, 2006: 149; Cagniant, 2006a: 194; Gratiashvili & Barjadze, 2008: 136; Paknia, et al. 2008: 154; Casevitz-Weulersse & Galkowsky, 2009: 484; Vonshak, et al. 2009: 40; Karaman, M.G. 2011: 85; Legakis, 2011: 27; Borowiec, L. & Salata, 2012: 501; Kiran & Karaman, 2012: 12; Borowiec, L. & Salata, 2013: 357; Borowiec, L. 2014: 87; Bračko, et al. 2014: 20; Lebas, et al. 2016: 218; Borowiec, L. & Salata, 2018: 6; Salata & Borowiec, 2018b: 157 (in key); Salata & Borowiec, 2018c: 46.
 * Senior synonym of barbarus: Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of fumatus: Emery, 1870: 194; Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of fuscula: Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1916b: 241; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of nigrobrunneus: Seifert, 2020: 49.
 * barbarus. Lasius alienus var. barbarus Santschi, 1931a: 11.
 * [First available use of Lasius niger st. lasioides var. barbara Santschi, 1921e: 170 (w.) MOROCCO, TUNISIA; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Emery, 1925b: 230; Bernard, 1945: 137.
 * Subspecies of brunneus: Santschi, 1936c: 208.
 * Junior synonym of alienus: Wilson, 1955a: 78; Smith, D.R. 1979: 1435.
 * Junior synonym of lasioides: Seifert, 1992b: 8; Bolton, 1995b: 221; Seifert, 2020: 49.
 * fumatus. Lasius fumatus Emery, 1869b: 26, fig. 3 (w.q.) ITALY.
 * Synonym of lasioides: Emery & Forel, 1879: 452.
 * Subspecies of niger: Emery & Forel, 1879: 452.
 * [Note: the species which is referred to as Prenolepis lasioides in the text of Emery, 1869b: 6, is referred to in the list of figures for that paper (p. 26) as Lasius fumatus. Both names refer to the same specimens and hence are automatically synonymous (Bolton, 1995b: 223).]
 * Junior synonym of lasioides: Emery, 1870: 194; Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * fuscula. Prenolepis fuscula Emery, 1869b: 8, pl. 1, fig. 4 (w.) ITALY.
 * Combination in Lasius: Emery, 1869b: 25 (in text), 26 (in figure captions).
 * As unavailable (infrasubspecific) name: Emery & Forel, 1879: 452.
 * Junior synonym of lasioides: Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1916b: 241; Emery, 1925b: 230; Bolton, 1995b: 223; Smith, D.R. 1979: 1435; Seifert, 2020: 49.
 * nigrobrunneus. Acanthomyops (Donisthorpea) brunneus var. nigrobrunneus Donisthorpe, 1926b: 18 (w.) ITALY.
 * Subspecies of brunneus: Donisthorpe, 1927a: 8.
 * Junior synonym of brunneus: Wilson, 1955a: 47; Bernard, 1967: 358; Seifert, 1992b: 6; Bolton, 1995b: 224; Radchenko, 2016: 366.
 * Junior synonym of lasioides: Seifert, 2020: 49.

Type Material
Lectotype, an alate gyne, labelled by Mayr ”Neapel Emery“, ”Collect. G.Mayr“, ”L. fumatus m., lasioides m., det. Emery“ and ”lectotype, des. E.O.Wilson“; 1 paralectotype worker labelled by Mayr ”Neapel Col. G.Mayr“, ”L. fumatus m., lasioides m., det. Emery“ and ”Typus“; both specimens in.

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