Checklist of Dolichoderus species

The following species and subspecies belong to the genus Dolichoderus. Synonyms are listed under their senior names. For valid names only see Dolichoderus species and for distribution information see Dolichoderus species by Country.

A
abruptus (Smith, F., 1858) (Bolivia, Brazil, Colombia, Ecuador, Peru)

• Hypoclinea ursus Mayr, 1866

affinis Emery, 1889 (Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Vietnam)

• Dolichoderus affinis nigricans Emery, 1895

affinis glabripes Forel, 1895 (India)

affinis mus Santschi, 1920 (Vietnam)

albamaculus Shattuck & Marsden, 2013 (Australia)

andinus (Kempf, 1962) (Peru)

angusticornis Clark, 1930 (Australia)

attelaboides (Fabricius, 1775) (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)

• Dolichoderus attelaboides pulla Santschi, 1923

• Dolichoderus imbecillus Mann, 1916

• Dolichoderus imbecillus heterogaster Santschi, 1923

australis André, 1896 (Australia)

B
baenae Mackay, W.P., 1993 (Colombia, Ecuador)

beccarii Emery, 1887 (Borneo, Indonesia, Malaysia)

bidens (Linnaeus, 1758) (Brazil, Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)

• Dolichoderus auromaculatus Forel, 1885

• Dolichoderus bidens attenuatus Forel, 1903

• Dolichoderus bidens bahiana Santschi, 1921

• Formica perditor Fabricius, 1804

bispinosus (Olivier, 1792) (Belize, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Formica fungosa Fabricius, 1798

• Hypoclinea vestita Mayr, 1862

• Polyrhachis arboricola Norton, 1868

brevis Santschi, 1920 (Laos)

brevithorax Menozzi, 1928 (Indonesia)

burmanicus Bingham, 1903 (Myanmar)

butteli Forel, 1913 (Indonesia)

C
canopus Shattuck & Marsden, 2013 (Australia)

carbonarius Emery, 1895 (Malaysia)

carbonarius latisquamis Emery, 1900 (Indonesia)

clarki Wheeler, W.M., 1935 (Australia)

• Dolichoderus tristis Clark Clark, 1930

clusor Forel, 1907 (Australia)

cogitans Forel, 1912 (Bolivia, Brazil)

coniger (Mayr, 1870) (Borneo, Indonesia, Malaysia)

crawleyi Donisthorpe, 1917 (Singapore)

curvilobus (Lattke, 1987) (Colombia, Costa Rica)

cuspidatus (Smith, F., 1857) (Borneo, Indonesia, Malaysia, Malaysia, Thailand)

• Dolichoderus atra Menozzi, 1930

• Dolichoderus bifurcatus Karavaiev, 1926

• Dolichoderus cuspidatus surbecki Santschi, 1925

• Dolichoderus cuspidatus tenuispina Viehmeyer, 1922

D
dajiensis Wang, W. & Zheng, 2005 (China)

debilis Emery, 1890 (Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Panama, Suriname, Trinidad and Tobago, Venezuela)

• Dolichoderus debilis parabiotica Forel, 1912

• Dolichoderus debilis sieversi Forel, 1901

decollatus Smith, F., 1858 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)

• Dolichoderus capitatus Santschi, 1921

• Dolichoderus decollatus neglectus Menozzi, 1935

dentatus Forel, 1902 (Australia)

diversus Emery, 1894 (Brazil, Colombia, Costa Rica, Ecuador, Guyana, Mexico, Panama, Trinidad and Tobago, Venezuela)

• Dolichoderus championi Forel, 1899

• Dolichoderus championi ornatus Mann, 1916

• Dolichoderus championi taeniatus Wheeler, W.M., 1916

• Dolichoderus championi trinidadensis Forel, 1899

• Dolichoderus germaini garbei Forel, 1911

• Dolichoderus germaini leviusculus Emery, 1906

• Iridomyrmex mazaruni Donisthorpe, 1939

doloniger (Roger, 1862) (Bolivia, Colombia, Venezuela)

doriae Emery, 1887 (Australia)

E
epetreius (Lattke, 1987) (Venezuela)

erectilobus Santschi, 1920 (Thailand, Vietnam)

etus Shattuck & Marsden, 2013 (Australia)

extensispinus Forel, 1915 (Australia)

F
feae Emery, 1889 (China, India, Myanmar, Thailand)

• Dolichoderus feae caligatus Wheeler, W.M., 1927

• Dolichoderus feae fuscus Emery, 1889

fernandezi Mackay, W.P., 1993 (Colombia)

ferrugineus Forel, 1903 (Bolivia, Brazil)

• Dolichoderus bidens inferior Mann, 1916

flatidorsus Zhou & Zheng, 1997 (China)

formosus Clark, 1930 (Australia)

• Dolichoderus occidentalis Clark, 1930

furcifer Emery, 1887 (Indonesia)

G
gagates Emery, 1890 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Suriname, Venezuela)

germaini Emery, 1894 (Brazil, Ecuador, Mexico, Paraguay, Peru)

ghilianii Emery, 1894 (Brazil, Peru)

gibbifer Emery, 1887 (Indonesia)

• Dolichoderus gibbifer gibbosior Forel, 1912

gibbus (Smith, F., 1861) (Indonesia, Sulawesi)

gordoni Shattuck & Marsden, 2013 (Australia)

goudiei Clark, 1930 (Australia)

H
haradae Mackay, W.P., 1993 (Brazil)

I
imitator Emery, 1894 (Bolivia, Brazil, Ecuador)

incisus Xu, Z., 1995 (China)

indrapurensis Forel, 1912 (Borneo, Indonesia, Malaysia)

indrapurensis nigrogaster Viehmeyer, 1922 (Borneo, Indonesia, Malaysia)

inermis Mackay, W.P., 1993 (Costa Rica)

inferus Shattuck & Marsden, 2013 (Australia)

inpai (Harada, 1987) (Brazil, Colombia, Ecuador)

J
jacobsoni Forel, 1909 (Indonesia)

K
kathae Shattuck & Marsden, 2013 (Australia)

kinabaluensis Dill, 2002 (Borneo, Indonesia, Malaysia)

L
lactarius (Smith, F., 1860) (Indonesia)

lamellosus (Mayr, 1870) (Argentina, Bolivia, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Paraguay, Suriname, Venezuela)

• Dolichoderus lamellosa missionensis Santschi, 1916

laminatus (Mayr, 1870) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Suriname, Venezuela)

• Dolichoderus laminatus luteiventris Emery, 1894

laotius Santschi, 1920 (China, Laos)

laurae Mackay, W.P., 1993 (Colombia)

lobicornis (Kempf, 1959) (Brazil, Colombia, Ecuador)

longicollis Mackay, W.P., 1993 (Colombia)

luederwaldti Santschi, 1921 (Brazil)

lugens Emery, 1894 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Mexico, Suriname)

lujae Santschi, 1923 (Brazil)

lutosus (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Suriname, Trinidad and Tobago, Venezuela)

• Dolichoderus lutosus nigriventris Forel, 1893

• Hypoclinea cingulata Mayr, 1862

M
magnipastor Dill, 2002 (Borneo, Indonesia, Malaysia)

mariae Forel, 1885 (United States)

• Dolichoderus mariae blatchleyi Wheeler, W.M., 1917

• Dolichoderus mariae davisi Wheeler, W.M., 1905

maschwitzi Dill, 2002 (Borneo, Indonesia, Malaysia)

mesonotalis Forel, 1907 (Brazil, Peru)

• Dolichoderus grandii Menozzi, 1924

• Dolichoderus simplex Forel, 1912

modiglianii Emery, 1900 (Indonesia)

• Dolichoderus modiglianii rubescens Emery, 1900

moggridgei Forel, 1886 (Afghanistan, China, India)

moggridgei bicolor Santschi, 1920 (India)

moggridgei lugubris Santschi, 1920 (India)

monoceros Emery, 1897 (New Guinea)

mucronifer (Roger, 1862) (French Guiana, Suriname)

• Dolichoderus spinicollis ensiger Forel, 1910

N
niger Crawley, 1922 (Australia)

nigricornis Clark, 1930 (Australia)

O
omacanthus (Kempf, 1972) (Brazil)

omicron Shattuck & Marsden, 2013 (Australia)

P
parvus Clark, 1930 (Australia)

• Dolichoderus glauerti Wheeler, W.M., 1934

pastorulus Dill, 2002 (Borneo, Indonesia, Malaysia)

patens (Mayr, 1870) (Borneo, Indonesia, Malaysia)

patens pubiventris Emery, 1900 (Indonesia, Philippines)

piceus Mackay, W.P., 1993 (Colombia)

pilinomas Dill, 2002 (Borneo, Indonesia, Malaysia)

pilosus Zhou & Zheng, 1997 (China)

plagiatus (Mayr, 1870) (Canada, Mexico, United States)

• Dolichoderus borealis Provancher, 1888

• Dolichoderus plagiatus inornatus Wheeler, W.M., 1905

pustulatus Mayr, 1886 (United States)

• Dolichoderus plagiatus beutenmuelleri Wheeler, W.M., 1904

Q
quadridenticulatus (Roger, 1862) (Bolivia, Brazil, Ecuador, Peru, Venezuela)

• Dolichoderus analis Emery, 1894

• Dolichoderus gibbosus gibbosoanalis Forel, 1922

• Dolichoderus gibbosus integra Forel, 1911

• Dolichoderus gibbosus nitidior Emery, 1894

• Formica gibbosa Smith, F. Smith, F., 1858

quadripunctatus (Linnaeus, 1771) (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Croatia, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Liechtenstein, Lithuania, Luxembourg, Monaco, Montenegro, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine)

• Dolichoderus quadripunctatus unicolor Ruzsky, 1905

quadripunctatus kratochvili Novak, 1941 (Czech Republic)

R
reflexus Clark, 1930 (Australia)

• Dolichoderus armstrongi McAreavey, 1949

rosenbergi Forel, 1911 (Ecuador)

rufescens Mann, 1912 (Brazil)

rufotibialis Clark, 1930 (Australia)

rugocapitus Zhou, 2001 (China)

rugosus (Smith, F., 1858) (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname)

rutilus Shattuck & Marsden, 2013 (Australia)

S
sagmanotus Xu, Z., 2001 (China)

satanus Bolton, 1995 (Brazil)

• Formica obscura Smith, F., 1858

scabridus Roger, 1862 (Australia)

• Dolichoderus scabridus ruficornis Santschi, 1916

• Polyrhachis foveolatus Lowne, 1865

schulzi Emery, 1894 (Brazil, Colombia, Costa Rica)

• Dolichoderus biolleyi Forel, 1908

• Dolichoderus schulzi columbica Forel, 1914

scrobiculatus (Mayr, 1876) (Australia)

semiorbis Shattuck & Marsden, 2013 (Australia)

semirugosus (Mayr, 1870) (Borneo, Indonesia, Malaysia, Thailand)

septemspinosus Emery, 1894 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Venezuela)

setosus (Kempf, 1959) (Brazil)

shattucki Mackay, W.P., 1993 (Colombia, Ecuador, Panama)

sibiricus Emery, 1889 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation)

• Dolichoderus abietis Kono & Sugihara, 1939

• Dolichoderus quadripunctatus japonicus Yoshioka, 1939

• Dolichoderus quadripunctatus yoshiokae Wheeler, W.M., 1933

• Dolichoderus sinensis Wheeler, W.M., 1921

• Dolichoderus sinensis atriceps Wheeler, W.M., 1928

siggii Forel, 1895 (Thailand)

smithi Mackay, W.P., 1993 (Bolivia)

spinicollis (Latreille, 1817) (Brazil, Colombia, Venezuela)

spurius Forel, 1903 (French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago)

• Dolichoderus bidens albatus Viehmeyer, 1922

squamanodus Xu, Z., 2001 (China)

sulcaticeps (Mayr, 1870) (Borneo, Indonesia, Malaysia, Thailand, Vietnam)

• Dolichoderus sulcaticeps nigriventris Santschi, 1920

sundari Tiwari, 2000 (India)

superaculus (Lattke, 1987) (Colombia)

T
taprobanae (Smith, F., 1858) (Borneo, China, India, Indonesia, Laos, Malaysia, Sri Lanka, Vietnam)

• Dolichoderus semirufus André, 1887

• Dolichoderus taprobanae obscuripes Santschi, 1920

• Dolichoderus taprobanae tonkina Santschi, 1920

• Formica ingruens Walker, 1859

• Hypoclinea gracilis Motschoulsky, 1863

taprobanae borneensis Forel, 1911 (Borneo, Indonesia, Malaysia)

taprobanae ceramensis Stitz, 1912 (Indonesia)

taprobanae friedrichsi Forel, 1914 (Vietnam)

taprobanae gracilipes (Mayr, 1879) (India)

taprobanae siamensis Forel, 1911 (Thailand)

taschenbergi (Mayr, 1866) (Canada, United States)

• Dolichoderus taschenbergi aterrimus Wheeler, W.M., 1915

• Dolichoderus taschenbergi gagates Wheeler, W.M., 1905

• Dolichoderus taschenbergi wheeleriella Forel, 1916

thoracicus (Smith, F., 1860) (Borneo, Cambodia, China, India, Indonesia, Laos, Malaysia, Philippines, Singapore, Sulawesi, Thailand, Vietnam)

• Dolichoderus bituberculatus emarginata Santschi, 1920

• Hypoclinea bituberculata Mayr, 1862

• Hypoclinea sellaris Roger, 1863

thoracicus bilikanus Santschi, 1925 (Indonesia)

thoracicus borneonensis (Roger, 1863) (Borneo, Indonesia, Malaysia)

thoracicus lacciperdus Santschi, 1925 (Cambodia)

thoracicus levior Karavaiev, 1926 (Indonesia)

thoracicus nasutus Karavaiev, 1935 (Cambodia)

thoracicus rufescens Stitz, 1925 (Borneo, Indonesia, Malaysia)

tricolor Emery, 1914 (New Caledonia)

tricornis Emery, 1897 (New Guinea)

tridentanodus Ortega-De Santiago & Vásquez-Bolaños, 2012 (Mexico)

tristis Mann, 1916 (Brazil, Peru)

tuberifer Emery, 1887 (Indonesia, Thailand)

turneri Forel, 1902 (Australia)

U
utriensis Ortiz & Fernández, 2011 (Colombia)

V
validus (Kempf, 1959) (Costa Rica, Ecuador, Panama)

varians Mann, 1916 (Brazil, Colombia, Ecuador)

voraginosus Mackay, W.P., 1993 (Brazil)

Y
ypsilon Forel, 1902 (Australia)

=Fossil Species=

A
affectus Théobald, 1937

antiquus Carpenter, 1930

B
balticus (Mayr, 1868)

brevicornis Dlussky, 2002

brevipalpis Dlussky, 2008

brevipennis Dlussky, 2008

bruneti Théobald, 1937

C
caribbaeus (Wilson, 1985)

coquandi Théobald, 1937

cornutus (Mayr, 1868)

D
dibolius Wilson, 1985

dlusskyi LaPolla & Greenwlalt, 2015

E
elegans Wheeler, W.M., 1915

evolans Zhang, J., 1989

explicans (Förster, 1891)

G
granulinotus Dlussky, 2008

H
heeri Dlussky & Putyatina, 2014

I
intermedius Mackay, W.P., 1993

J
jiaoyanshanensis (Hong, 1985)

K
kohlsi Dlussky & Rasnitsyn, 2002

kutscheri Dlussky, 2008

kutschlinicus (Deichmüller, 1881)

L
lacinius Zhang, J., 1989

longipennis (Mayr, 1868)

longipilosus Dlussky, 2002

lucidus Dlussky, 2008

luridivenosus Zhang, J., Sun & Zhang, X., 1994

M
mesosternalis Wheeler, W.M., 1915

N
nanus Dlussky, 2002

O
obliteratus (Scudder, 1877)

oviformis Théobald, 1937

P
passalomma Wheeler, W.M., 1915

perkovskyi Dlussky, 2008

pilipes Dlussky, 2008

pinguis Dlussky, Rasnitsyn & Perfilieva, 2015

polessus Dlussky, 2002

polonicus Dlussky, 2002

primitivus (Wilson, 1985)

prolaminatus (Wilson, 1985)

punctatus Dlussky, 2008

R
robustus Dlussky, 2002

rohweri Carpenter, 1930

S
sculpturatus (Mayr, 1868)

T
tavridus Perfilieva, Dubovikoff & Dlussky, 2017

tertiarius (Mayr, 1868)

transversipetiolaris Zhang, J., Sun & Zhang, X., 1994

V
vectensis Donisthorpe, 1920

vexillarius Wheeler, W.M., 1915

vlaskini Dlussky, 2008

Z
zherichini Dlussky, 2002