Cladomyrma maryatiae

This species was collected from Neonauclea species (sp. C of J. Moog; sp. 2 of D. Davidson) in Sabah and Sarawak. Specimens are recorded as displaying aggressive behavior (Davidson, label information), a trait which it shares with most other Cladomyrma species. [Two dubious records exist for collections from the plant Spatholobus oblongifolius, but those determinations are based solely on workers and should be regarded with some caution.]. (Agosti et al. 1999)

Identification
Agosti et al. (1999) placed Cladomyrma species into two groups based on the shape of the queen's petiole, raised (an erect node or scale) or depressed (dorsally truncate and low). Cladomyrma maryatiae belongs to the raised petiole group.

The coloration of the queen is variable, from dark brown to a lighter brown. However, this species is distinct from most others by the combination of the following characters: (1) absence of erect hairs on second and third gastral tergite, (2) raised petiole, (3) large metapleural gland orifice, and from the very similar aurochaetae by the absence of the long curved hairs on the gastral tergite 1, and the matte clypeus. All workers seen were small.

Distribution based on Regional Taxon Lists
Indo-Australian Region: Borneo, Indonesia, Malaysia.

Nomenclature

 *  maryatiae. Cladomyrma maryatiae Agosti, Moog & Maschwitz, 1999: 21, figs. 9, 11, 23, 35 (s.w.q.) BORNEO.

Worker
Major worker. AL 1.02–1.22, HL 0.98–1.08, HW 0.84–0.96, EL 0.20–0.22, SL 0.45–0.52, CI 85–89, EI 21–24, SI 51–56 (n = 8).

Small body size; coloration reddish brown; clypeus in lateral view smoothly curved; metapleural gland orifice large; gaster matte; gastral pubescence widely set (hairs of about same length as distance between their insertions).

Minor: AL 0.72–0.90, HL 0.70–0.80, HW 0.59–0.67, EL 0.14–0.16, SL 0.34–0.40, CI 83–87, EI 22–29, SI 58–61 (n = 8).

Small body size; body color dark brown, with head and gaster darker and genae slightly lighter colored; metapleural orifice covered with some long, erect setae; gastral pubescence very short and dense, hair longer than the space between them.

Queen
AL 2.10–2.28, HL 1.30–1.52, HW 1.06–1.14, EL 0.46–0.50, SL 0.64–0.74, CI 79–82, EI 42–45, SI 58–64 (n = 8). Holotype: AL 2.20, HL 1.43, HW 1.14, EL 0.48, SL 0.68, CI 80, EI 42, SI 60.

Clypeus slightly angulate in lateral view, matte; head elongated in frontal view; clypeus and frontal part of head reddish, head otherwise dark brown; head surface slightly shining; dorsal part of katepisternum matte and with a distinct pubescence; metapleural gland orifice large and open, in front a bunch of long erect hairs; petiole in lateral view squamiform, in hind view dorsally truncated; whole body covered with long, erect, golden hairs; on gastral tergites 2 and 3 erect hairs only at posterior margin and lateral parts; gastral tergites 2 and 3 with short, widely set appressed to semidecumbent pubescence; gastral tergite 1 with erect, golden hairs; gaster somewhat shining; head and alitrunk evenly dark brown to brown; medium to small body size.

Type Material
HOLOTYPE: E-MALAYSIA, Sabah, Ranau, Poring Hot Springs, 3/14/1995, Joachim Moog, 95-123, ex: Neonauclea sp., altitude ca. 700 m, mature colony,. PARATYPES: E-MALAYSIA, Sabah, Ranau, Poring Hot Springs, 3/12/1995, Joachim Moog, 95-119, ex: Neonauclea sp. C, altitude ca. 600 m, young colony: 195 workers in total; Sabah, Ranau, Poring Hot Springs, 3/12/1995, Joachim Moog, 95-120, ex: Neonauclea sp. ?C, colony size: ca. 760 workers in total; Sabah, vic. Danum Valley Res. Center, 85 km W Lahad Datu, 8/7/1991, Diane Davidson, 91-031, ex: Neonauclea sp. 2; AMNH,, , , , , , and collections of Agosti and Maschwitz.

References based on Global Ant Biodiversity Informatics

 * Agosti D., J. Moog, and U. Maschwitz. 1999. Revision of the Oriental Plant-ant genus Cladomyrma. American Museum Novitates 3283: 1-24.
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58