Camponotus herculeanus sachalinensis

This taxon is not in use as it is currently considered to be a junior synonym of Camponotus herculeanus.

Nomenclature

 * sachalinensis. Camponotus herculeanus var. sachalinensis Forel, 1904b: 38 (q.) MONGOLIA, RUSSIA.
 * Karavaiev, 1912b: 592 (m.).
 * Combination in C. (Camponotus): Emery, 1925b: 72.
 * As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 221.
 * Junior synonym of saxatilis: Kuznetsov-Ugamsky, 1928b: 4; Kuznetsov-Ugamsky, 1929a: 18.
 * Status as species: Collingwood, 1981: 29; Bolton, 1995b: 121; Collingwood & Heatwole, 2000: 12; Imai, et al. 2003: 39.
 * Subspecies of herculeanus: Ruzsky, 1905b: 222; Wheeler, W.M. 1906c: 325; Yano, 1910: 422; Karavaiev, 1912b: 592; Ruzsky, 1926: 108; Arnol'di, 1967: 1821; Pisarski, 1969a: 230; Pisarski, 1969b: 304; Dlussky & Pisarski, 1970: 86; Collingwood, 1976: 306; Pisarski & Krzysztofiak, 1981: 159; Kupyanskaya, 1986b: 96; Kupyanskaya, 1990: 166; Morisita, et al. 1991: 41; Radchenko, 1994b: 116 (in key); Radchenko, 1996b: 1203 (in key); Radchenko, 1997a: 555; Radchenko, 2005b: 158.
 * Senior synonym of altaica: Arnol'di, 1967: 1821.
 * Senior synonym of jacuticus: Arnol'di, 1967: 1821.
 * Junior synonym of herculeanus: Yasumatsu & Brown, 1957: 49; Schar et al., 2018: 6.

Taxonomic Notes
Aibek & Yamane (2009) - Various authors have treated this form as a subspecies of C. herculeanus or as a distinct species. Collingwood (1981) first raised this form to distinct species rank, but he did not mention any difference between the two. Radchenko (2005) argued that it is a subspecies of C. herculeanus, since sachalinensis and the typical form differ only in coloration, and intermediate specimens can be collected in Altai and Tuva. He mentioned that the two forms are indistinguishable also in the queen and male. The typical C. herculeanus is distinctly bicolored; especially in the queen caste the lateral part of mesonotum, whole propodeum, petiole and legs are reddish brown, while the queen of sachalinensis is almost wholly black except for legs. In the typical form, even the major worker has reddish brown alitrunk, petiole and legs, while the major worker of sachalinensis has a wholly black alitrunk; only propodeal declivity, petiole and legs are tinged with brown or red. Furthermore, in the worker of the typical form from Finland the pubescence on gastral tergites is longer and denser than in sachalinensis. Although the difference is minor, we tentatively follow Collingwood (1981) and Bolton (1995a) for the above-mentioned reason (see also Japanese Ant Database Group, 2003 for the Japanese population).