Temnothorax recedens

A widespread, mainly western Mediterranean species occurring in southern Europe, Turkey and NW Africa (Rigato & Toni, 2011). It shows very wide habitat preferences, being found in deciduous forests, bushes growing along water sources, phrygana and in mountains. Nests are most often in soil, sometimes under stones or in rocks crevices. Workers have been collected in litter or on rocks. Colonies are polygynous (Salata, Borowiec & Trichas, 2018).

Identification
Prebus (2017) - A member of the Palearctic clade.

Salata et al. (2018) - The only Temnothorax species known from Crete with a deep metanotal groove, big eyes, and reduced body sculpture (mesosoma and head almost entirely smooth and shiny).

Cretan populations of Temnothorax recedens are distinctly bicoloured, with the head, lateral parts of the mesosoma, and a large part of gaster always darker than the rest of the body. This differs from populations from eastern Greece, which have less contrasting colouration. Cretan specimens also have shorter propodeal spines than most populations in southern and central Greece. These characteristics of Cretan T. recedens are most similar to T. recedens populations from the western part of its range. Further study of the genetics and morphometrics of populations across the entire species' range is needed to test if Temnothorax recedens is a complex of cryptic species or a valid single, albeit variable, species.

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Armenia, Balearic Islands, Bulgaria, Croatia, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Israel, Italy, Malta, Montenegro, Portugal, Republic of Macedonia, Russian Federation, Slovenia, Spain, Switzerland, Tunisia, Turkmenistan.

Nomenclature

 *  recedens. Myrmica recedens Nylander, 1856b: 94, pl. 3, fig. 41 (w.) FRANCE. Roger, 1859: 258 (q.); Forel, 1874: 83 (m.). Combination in Temnothorax: Mayr, 1861: 68; in Leptothorax (Temnothorax): Forel, 1890a: lxxii; in Temnothorax: Bolton, 2003: 271. Senior synonym of pictus: Collingwood, 1978: 69; of ergatogyna, mordax: Dlussky & Soyunov, 1988: 31; of barbarus: Cagniant & Espadaler, 1997: 268. See also: Forel, 1915d: 27; Emery, 1916b: 186; Bernard, 1967: 186; Kutter, 1977c: 135; Atanassov & Dlussky, 1992: 126.
 * mordax. Leptothorax (Temnothorax) mordax Santschi, 1919e: 242 (q.) SPAIN. Junior synonym of recedens: Dlussky & Soyunov, 1988: 31.
 * pictus. Leptothorax (Temnothorax) pictus Emery, 1924c: 166 (w.) SPAIN. Junior synonym of recedens: Collingwood, 1978: 69.
 * barbarus. Leptothorax (Temnothorax) recedens var. barbarus Santschi, 1939d: 76 (w.q.) TUNISIA. Junior synonym of recedens: Cagniant & Espadaler, 1997: 268.
 * ergatogyna. Leptothorax (Temnothorax) ergatogyna Bernard, 1950a: 17, fig. 3 (w.q.) FRANCE. Combination in Temnothorax: Bernard, 1967: 186. Junior synonym of recedens: Dlussky & Soyunov, 1988: 31.

Worker
Salata et al. (2018) - based on Cretan specimens (n=15): HL: 0.637 ± 0.07 (0.503-0.745); HW: 0.518 ± 0.06 (0.408-0.603); SL: 0.623 ± 0.07 (0.491- 0.735); EL: 0.138 ± 0.01 (0.106-0.156); EW: 0.095 ± 0.01 (0.067-0.112); ML: 0.787 ± 0.116 (0.609-0.978); PSL: 0.129 ± 0.03 (0.089-0.179); SDL: 0.141 ± 0.03 (0.084- 0.19); PL: 0.293 ± 0.04 (0.212-0.358); PPL: 0.216 ± 0.03 (0.156-0.257); PH: 0.199 ± 0.03 (0.156-0.246); PPH: 0.193 ± 0.03 (0.154-0.243); HTL: 0.518 ± 0.08 (0.358- 0.603); PNW: 0.364 ± 0.05 (0.279-0.441); PW: 0.142 ± 0.02 (0.109-0.168); PPW: 0.213 ± 0.03 (0.156-0.257); HI: 81.2 ± 2.3 (77.7-85.9); SI1: 97.7 ± 3.9 (92.3-108.1); SI2: 120.3 ± 2.7 (115.6-125.8); MI: 65.0 ± 8.1 (49.3-75.0); EI1: 69.9 ± 8.7 (53.8-78.9); EI2: 15.0 ± 1.7 (12.8-17.9); TI: 102.4 ± 13.0 (87.1-132.0); PI: 147.6 ± 10.4 (128.1-167.1); PPI: 112.0 ± 10.7 (94.2-131.3).

Head and gaster brown to dark brown. Mesosoma mostly in the same colouration as head, but with orange pronotum. Sometimes orange area covering some parts of mesonotum and propodeum, in extreme examples the whole mesosoma orange. Base of the first tergite of gaster with bright brown to orange spot. Antennae and legs with the same colouration as mesonotum. Femora with dark spots in central part.

Head oval, 1.2 times as long as wide, lateral surfaces below eyes straight, above eyes gently convex, posterior edges convex, occipital margin of head convex. Anterior margin of the clypeus slightly rounded, medial notch absent. Eyes big, oval, 1.5 times as long as wide. Antennal scape long, in lateral view slightly curved, 0.9 times as long as length of the head, in apex gradually narrowed, its base with small, triangular tooth, funiculus long, club 3 segmented. Surface of scape with very fine microreticulation, shiny, covered with dense, suberect to erect setae. Mandibles rounded with thick sparse, longitudinal striae, shiny. Clypeus shiny and smooth. Frontal carinae short, not extending beyond frontal lobes. Antennal fossa deep, smooth and shiny. Frontal lobes narrow, smooth and shiny. Head on the almost whole surface smooth and shiny. Sculpture, if occurs, limited to sparse, thin, longitudinal striation on the area between eyes and mandibles. Entire head bearing from suberect to erect, pale, long and thick setae.

Mesosoma elongate, 2.2 times as long as wide, slightly arched in profile. Metanotal groove deep. Pronotum convex on sides. Propodeal spines short, with wide base, directed upward, with acute tips. Dorsal surface of promesonotum smooth and shiny, sometimes with few longitudinal wrinkles. Dorsal surface of propodeum shiny, smooth or with slight reticulation. Lateral surfaces of pronotum smooth and shiny. Lateral surfaces of mesonotum and propodeum shiny, with slight, longitudinal striation or reticulation. Entire mesosoma bearing suberect to erect, pale, long and thick setae.

Petiole, in lateral view, with short peduncle, node high, with anterior and posteriol faces straight or convex, its dorsal surface arched. Postpetiole, in lateral view, regularly convex, apical half with gently convex sides. Peduncle and petiolar node and postpetiole dorsum shiny, with thin, microreticulation, area between rugae smooth, central parts of dorsal surfaces of petiolar node and postpetiole dorsum smooth. Petiole and postpetiole on dorsal surfaces bearing sparse, long, semierect to erect setae.

Gaster smooth and shiny, bearing erect, thin, pale setae. Legs short, shiny, with fine microreticulation. Tibiae 0.7 times as long as head length, bearing adpressed setae on the whole surface. Inner margins of tibia and femora with erect setae.

References based on Global Ant Biodiversity Informatics

 * Abril S., C. Gómez. 2012. Lista actualizada de las especies de hormigas de Menorca (Islas Baleares, España) y primera cita de Monomorium andrei Saunders, 1890 (Hymenoptera, Formicidae) de la isla Boletín de la Sociedad Entomológica Aragonesa 50: 403-407.
 * Abril S., and C. Gomez. 2013. Rapid assessment of ant assemblages in public pine forests of the central Iberian Peninsula. Forest Ecology and Management 293: 7984.
 * Acosta, F. J., M. D. Martínez Ibáñez, and M. A. Morales. "Contribución al conocimiento de la mirmecofauna del encinar peninsular. I." Boletín de la Asociación Española de Entomología 6 (2) (1983): 379-391.
 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * André, E. "Species des Hyménoptères d'Europe et d'Algérie. Les fourmis." in: André, Edm. 1881-1886. Species des Hyménoptères d'Europe et d'Algérie. Tome Deuxième. Beaune: Edmond André, 919 + 48 pp. Tomo II (1881): 1016.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2010. Iberomyrmex 3: 32-33.
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2011. Iberomyrmex 3: 30-31.
 * Asociacion Iberica de Mirmecologia. 2013. List of species collected during the Taxomara 2013. Iberomyrmex 4: 35-36.
 * Asociacion Iberica de Mirmecologia. 2015. List of species collected during the Taxomara Lisboa 2015 Iberomyrmex 7: 44-45.
 * Asociacion Iberica de Mirmecologia. 2016. List of species collected during the Taxomara Murcia 2016. Iberomyrmex 8: 48-49.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Beibl, J., A. Buschinger, S. Foitzik and J. Heinze. 2007. Phylogeny and phylogeography of the Mediterranean species of the parasitic ant genus Chalepoxenus and its Temnothorax hosts. Insectes Sociaux 54:189-199
 * Beibl, J., P. D'Ettorre and J. Heinze. 2007. Cuticular profiles and mating preference in a slave-making ant. Insectes Sociaux. 54:174-182
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard, F. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
 * Bernard, F. "Remarques sur le peuplement des Baléares en fourmis." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 47 (1956): 254-266.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., C. Galkowski, and S. Salata. 2015. What is Tetramorium semilaeve André, 1883? (Hymenoptera, Formicidae). ZooKeys 512:39-62.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2015. Pheidole symbioticaWasmann, 1909, an enigmatic supposed social parasite, is a nematodeinfested form of Pheidole pallidula(Nylander, 1849) (Hymenoptera: Formicidae: Myrmicinae). Sociobiology 62(2): 181-186.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec L., and S. Salata. 2017. New records of ants (Hymenoptera: Formicidae) from southern Portugal. Acta Entomologica Silesiana 25(3): 1-10.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Buschinger A. 1999. Bemerkenswerte ameisenfunde aus Sudtirol (Hymenoptera: Formicidae). Myrmecologische Nachrichten 3: 1-8.
 * Buschinger, A., K. Fischer, H. P. Guthy, K. Jessen, and U. Winter. "Biosystematic revision of Epimyrma kraussei, E. vandeli, and E. Foreli." Psyche 93 (1987): 253-276.
 * Buschinger, A., W. Ehrhardt, K. Fischer, and J. Ofer. "The slave-making ant genus Chalepoxenus (Hymenoptera, Formicidae). I. Review of literature, range, slave species." Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere (Jena) 115 (1988): 383-401.
 * Buschinger, A. "The Epimyrma species of Corsica." Spixiana 8 (3) (1985): 277-280.
 * Buschinger, A. "Workerless Epimyrma kraussei Emery 1915, the first parasitic ant of Crete." Psyche (Cambridge) 96 (1989): 69-74.
 * Cagniant, H. 1968. Liste preliminaire de fourmis forestieres d'Algerie. Resultats obtenus de 1963 a 1964. Bulletin de la Société d'Histoire Naturelle de Toulouse 104: 138-147
 * Cagniant, H. "Contribution à la connaissance des fourmis marocaines: Aphaenogaster baronii n. sp. (Hyménoptères, Formicoidea, Myrmicidae)." Bulletin de la Société d'Histoire Naturelle de Toulouse 124 (1988): 43-50.
 * Cagniant, H. "Contribution á la connaissance des fourmis marocaines. Aphaenogaster gemella au Maroc: Nouvelle description d´Aphaenogaster gemella ssp. Marocana Forel (n. status) de la région de Tanger. Problemes biogéographiques soulevés par l'espèce A. gemella (Roger)." Bulletin de la Société d'Histoire Naturelle de Toulouse 125 (1989): 47-54.
 * Cagniant, H. "Étude de quelques fourmis marocaines. Statistique provisoire des Formicidae du Maroc." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 53 (1964): 83-118.
 * Cagniant, H. Les peuplements de fourmis des forêts algériennes: écologie, biocénotique, essai biologique. Universite de Toulouse, 1973.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Casevitz-Weulersse, J. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Ceballos, G. "Catálogo de los Himenópteros de España.", 295-321. Vol. Catálogo de los Himenópt. Madrid: Instituto Español de Entomologia, 1956.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood, C. A. 1993. A Comparitive Study of the Ant Fauna of Five Greek Islands. Biologia Gallo-hellenica. 20,1:191-197
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Collingwood, C. A. "A comparative study of the ant fauna of five Greek islands." Biologia Gallo-Hellenica 20 (1993): 191-197.
 * Comín del Río, P., and Andrés de Haro. "Datos iniciales para un estudio ecológico de las hormigas de Menorca." Bolletí de la Societat d´Historia Natural de les Balears 24 (1980): 23-48.
 * Comín del Río, P., and X. Espadaler. "Ants of the Pityusic islands." In Biogeography of the Pityusic Islands, edited by H. Kuhbler, J. A. Alcover and C. Guerau, 278-231. The Hague: Junk, 1984.
 * Comín del Río, P. "Los Formícidos de Menorca. Contribución al estudio taxonómico, geográfico y biológico." Tesina de licenciatura Universida (1977): 135 pp.
 * Comín del Río, P. Estudio de los formícidos de Baleares: Contribución al estudio taxonómico, geográfìco y biológico. Palma de Mallorca: Universidad de las Islas Baleares, 1988.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Dlussky G. M., O. S. Soyunov, and S. I. Zabelin. 1990. Ants of Turkmenistan. Ashkabad: Ylym Press, 273 pp.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Emery, C., and A. Forel. "Catalogue des Formicides d'Europe." Mitteilungen der Schweirzerischen Entomologischen Gesellschaft 5 (1879): 441-481.
 * Espadaler X. F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2009. Ants (Hymenoptera, Formicidae) from Mont-Rebei gorge (Pallars Jussà). Boletín Sociedad Entomológica Aragonesa 44: 393399.
 * Espadaler X., F. Garcia, X. Roig, and R. Vila. 2013. Ants (Hymenoptera, Formicidae) from the Castell de Montesquiu park (Osona, north-east of the Iberian Peninsula). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 53: 223-227.
 * Espadaler, X. and Cagniant, H. Plagiolepis Xene Starcke, The First Inquiline Ant From The Balearic Islands (Spain). Psyche. 98:351-354.
 * Espadaler, X. and H. Cagniant. 1991. Psyche. 98: 351-354.
 * Espadaler, X., and C. Restrepo. "Els gèneres Epimyrma Emery i Chalepoxenus Menozzi, formigues paràsites socials (Hymenoptera: Formicidae), a la Península Ibèrica. Estat actual del coneixement." Butlletí de la Institució Catalana d'Història Natural 49 (Sec. Z (1983): 123-126.
 * Espadaler, X., and H. Cagniant. "Plagiolepis xene Starckë, the first inquiline ant from the Balearic Islands (Spain)." Psyche 98 (4) (1992): 351-354.
 * Espadaler, X., and L. Lopez Soria. "Rareness of certain Mediterranean ant species: fact or artifact?" Insectes Sociaux 38 (1991): 365-377.
 * Espadaler, X., and X. Roig. "Ants from the Montnegre-Corredor Natural Park with description of the male lasius cinereus Seifert." Miscellanea Zoologica 23 (2) (2001): 45-53. Abstract
 * Foitzik S., and Heinze, J. 1999. Non-random size differences between sympatric species of the ant genus Leptothorax (Hymenoptera: Formicidae). Entomologia Generalis 24: 65-74.
 * Forel, A. "Fourmis de la faune méditerranéenne récoltées par MM. U. et J. Sahlberg." Revue Suisse de Zoologie 21 (1913): 427-438.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Galkowski C. 2011. Une liste des fourmis (Hymenoptera, Formicidae) récoltées dans la région de Grasse, avec la mention dune nouvelle espèce de la faune de France. Bulletin de la Société linnéenne de Provence, 62 : 41-44.
 * Garcia F., X. Espadaler, and K. Gomez. 2009. First record of Amblyopone impressifrons (Emery, 1869) from the Iberian Peninsula and of Proceratium melinum (Roger, 1860) from Catalonia (Hymenoptera, Formicidae). Boletín Sociedad Entomológica Aragonesa 45: 357-360.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Gasperini, R. "Notizia sulla fauna imenotterologadalmata, II, Formicidae-Mutillidae-Scoliadae-Sapygidae-Pompilidae-Sphegidae-Chrysididae." Anuario Dalmatico 4 (1887): 143-160.
 * Gomez C., D. Casellas, J. Oliveras, and J. M. Bas. 2003. Structure of ground-foraging ant assemblages in relation to land-use change in the northwestern Mediterranean region. Biodiversity and Conservation 12: 21352146.
 * Gomez K. 2017. Two species of exotic ants (Hymenoptera: Formicidae) new to Malta. Boletin de la Sociedad Entomologica Aragonesa (S.E.A.) 61: 233-235.
 * Gomez, C., P. Pons and J.M. Bas. 2003. Effects of the Argentine ant Linepithema humile on Seed Dispersal and Seedling Emergence of Rhamnus alaternus. Ecography 26(4):532-538
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Gómez C., and S. Abril. 2011. Selective logging in public pine forests of the central Iberian Peninsula: Effects of the recovery process on ant assemblages Forest Ecology and Management 262: 1061-1066.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis Collection Database
 * Lombarte, A., S. Romero, and Andrés de Haro. "Contribución al conocimiento faunístico de los formicidos de la Sierra de Collserola (Barcelona)." Orsis 4 (1989): 125-140.
 * Mei M. 1992. A survey of the socially parasitic ant genera Epimyrma Emery, 1915 and Chalepoxenus Menozzi, 1922 in Italy (Hymenoptera, Formicidae, Myrmicinae). Insectes Sociaux. 39: 145-156.
 * Müller, G. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Obregon R., J. Lopez, J. L. Reyes-Lopez. 2014. A catalogue of the ant fauna (Hymenoptera, Formicidae) of Sierra Mágina (Jaén, southern Spain). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 370374.
 * Ordóñez-Urbano C., J. Reyes-López, and S. Carpintero-Ortega. 2007. Estudio faunísidos de los formícidos (Hymenoptera: Formicidae) asociados a los bosques de ribera en la Provincia de Córdoba (España). Primeras Aportaciones. Boletin Sociedad Entomológica Aragonesa 40: 367-375.
 * Passera L. 1967. Peuplement en fourmis terricoles du rebord Meridional des Causses jurassiques du Quercy: la lande calcaire a buis. Vie et milieu. Serie C, Biologie terrestre 18(1): 189-205.
 * Peru L. 1999. Quelques ajouts et corrections a un catalogue des fourmis de la region Centre (Hymenoptera, Formicidae). Bulletin de l'Entomologie Tourangelles 20(1): 9-12.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Pisarski B. 1982. Ants (Hymenoptera: Formicoidea) of Warsaw and Mazovia. Memorabilia Zool. 36: 73-90.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Prado e Castro C., M. D. Garcia, C. Palma, and M. D. Martinez-Ibanez. 2014. First report on sarcosaprophagous Formicidae from Portugal (Insecta: Hymenoptera). Annales de la Société entomologique de France 50(1): 51-58.
 * Restrepo, C., X. Espadaler, and Andrés de Haro. "Contribución al conocimiento faunístico de los formícidos del Macizo de Garraf (Barcelona)." Orsis (Organismes i Sistemes) 1 (1986): 113-129.
 * Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
 * Reyes López, J. L., and G. Luque García. "Nuevas citas de Monomorium andrei Saunders, 1890 (Hymenoptera: Formicidae) para la Península Ibérica y Baleares." Boletín de la Asociación Española de Entomología 27 (1-4) (2003): 221-222.
 * Reyes-López, J., J. López-Tirado, and R. Obregón-Romero. "Nuevas citas de hormigas parásitas y esclavistas (Hym., Formicidae) para Sierra Mágina (Andalucía)." Boletín de la Asociación Española de Entomología 35, no. 3-4 (2011): 497-501.
 * Rigato S., and I. Toni. 2011. Short notes 21. Hymenoptera, Formicidae. Pp. 873-882 in: Nardi, G.; Whitmore, D.; Bardiani, M.; Birtele, D.; Mason, F.; Spada, L.; Cerretti, P. (eds.) 2011. Biodiversity of Marganai and Montimannu (Sardinia). Research in the framework of the ICP Forests network. Conservazione Habitat Invertebrati, 5. Sommacampagna, Verona: Cierre Edizioni, 896 pp.
 * Roig X., X. Espadaler, R. Cusco, F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2008. Ants in gypsum areas. First record from Catalonia (Iberian Peninsula) of Cardiocondyla batesii Forel (Hymenoptera, Formicidae) with an updated list of Catalan Formicidae. Boletin Sociedad Entomologica Aragonesa 42: 189-192.
 * Ruiz Heras P., M. D. Martinez Ibanez, F. J. Cabrero-Sanudo, and M. A. Vazquez Martinez. 2011. Primeros datos de Formícidos (Hymenoptera, Formicidae) en parques urbanos de Madrid. Boletín de la Asociación española de Entomología, 35(1-2): 87-106.
 * Rybka T. S. 2008. Peculiarities of entomocomplex forming in ecosystems of Yalta mountian-forest nature reserve. Visnyk of Dnipropetrovsk University. Biology. Ecology. 16(2): 141146.
 * Salata S., L. Borowiec, and A.Trichas. 2018. Taxonomic Revision of the Cretan Fauna of the Genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae), with Notes on the Endemism of Ant Fauna of Crete. Annales Zoologici (Warsaw) 68(4): 769-808.
 * Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
 * Salata S., and L. Borowiec. 2018. A new species of the ant genus Lasius Fabricius, 1804 from Crete (Hymenoptera, Formicidae). ZooKeys 789: 139–159.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
 * Santschi, F. 1918. Leptothorax nouveaux de l'Afrique mineure. Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord 9: 31-38.
 * Santschi, F. "Leptothorax nouveaux de l'Afrique Mineure." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 9 (1918): 31-38.
 * Schembri, Stephen P. and Cedric A. Collingwood. A Revision of the Myrmecofauna of the Maltese Islands. 417-442.
 * Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
 * Scupola I. 2009. Le formiche delle Isole Egadi (Sicilia) (Hymenoptera, Formicidae). Bollettino del Museo Civico di Storia Naturale di Verona Botanica Zoologia 33: 97-103
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Taheri A., J. Reyes-Lopez, and N. Bennas. 2014. Contribution a l'etude de la fauna myrmecologique du parc national de Talassemtane (nord du Maroc): biodiversite, biogeographie et especes indicatrices. Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 225236.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Tohme G. 1996. Formicidae. Etude de la diversité biologique n° 4 . Ministère de lAgriculture à Beyrouth (Eds.). P85-87.
 * Tohme G., and H. Tohme. 2014. Nouvelles liste des especes de fourmis du Liban (Hymenoptera, Formicoidea). Lebanese Science Journal 15(1): 133-141.
 * Vonshak M., and A. Ionescu-Hirsch. 2009. A checklist of the ants of Israel (Hymenoptera: Formicidae). Israel Journal of Entomology 39: 33-55.
 * Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
 * de Haro, Andrés, C. A. Collingwood, and P. Comín del Río. "Prospección mirmecológica por Ibiza y Formentera (Baleares)." Orsis 2 (1986): 115-120.
 * de Haro, Andrés, and C. A. Collingwood. "Formícidos de las Sierras de Prades-Montsant, Sierras de Cavalls-Alfara-Montes Blancos (Tarragona)." Boletín de la Estación Central de Ecología 10 (1981): 55-58.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por la península Tingitana al norte del Rif (Marruecos)." Orsis (Organismes i Sistemes) 12 (1997): 93-99.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por las sierras de Aitana-Alfaro y los cabos de la Nao-San Antonio (Alicante) y su comparación con la fauna balear y de Córcega-Cerdeña." Orsis 3 (1988): 165-172.
 * de Haro, Andrés. "Contribución al conocimiento de los formícidos del cabo de Creus (Gerona)." Pirineos 117 (1982): 49-57.