Checklist of Camponotus species

The following species and subspecies belong to the genus Camponotus. Synonyms are listed under their senior names. For valid names only see Camponotus species and for distribution information see Camponotus species by Country.

A
abditus Forel, 1899 (Guatemala, Mexico)

abjectus Santschi, 1937 (Angola, Senegal)

abrahami Forel, 1913 (Lebanon, Turkey)

abscisus Roger, 1863 (Brazil, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

absquatulator Snelling, R.R., 2006 (Mexico, United States)

abunanus Mann, 1916 (Brazil)

acutirostris Wheeler, W.M., 1910 (Mexico, United States)

• Camponotus acutirostris clarigaster Wheeler, W.M., 1915

acvapimensis Mayr, 1862 (Cameroun, Eritrea, Ghana, Kenya, Liberia, Nigeria, Oman, Senegal, Togo, United Arab Emirates, Zimbabwe)

• Camponotus acwapimensis poultoni Forel, 1913

• Camponotus flavosetosus Donisthorpe, 1945

• Camponotus mombassae Forel, 1886

adami Forel, 1910 (Australia)

adenensis Emery, 1893 (United Arab Emirates, Yemen)

aegaeus Emery, 1915 (Bulgaria, Greece, Switzerland, Turkey)

aegyptiacus Emery, 1915 (Benin, Egypt, Eritrea, Guinea, Kenya, Kuwait, Oman, Senegal, Yemen)

• Camponotus maculatus infrasquameus Santschi, 1926

aeneopilosus Mayr, 1862 (Australia)

aeneopilosus flavidopubescens Forel, 1902 (Australia)

aequatorialis Roger, 1863 (Gabon, Guinea)

aequatorialis kohli Forel, 1915 (Democratic Republic of Congo)

aequitas Santschi, 1920 (South Africa, Zimbabwe)

aethiops (Latreille, 1798) (Albania, Andorra, Armenia, Austria, Balearic Islands, Bulgaria, China, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Malta, Montenegro, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Camponotus aethiops concava Dalla Torre, 1893

• Camponotus aethiops sylvaticoides Dalla Torre, 1893

• Camponotus maculatus glaber Emery, 1925

• Camponotus marginatus hyalinipennis Costa, 1884

• Camponotus sylvaticus sylvaticoaethiops Forel, 1874

• Formica angustata Latreille, 1798

• Formica marginata Latreille, 1798

• Formica nigrata Nylander, 1849

aethiops cachmiriensis Emery, 1925 (India)

aethiops escherichi Emery, 1925 (Turkey)

afflatus Viehmeyer, 1925 (Australia)

ager (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Suriname, Trinidad and Tobago, Venezuela)

agonius Santschi, 1915 (Democratic Republic of Congo, Guinea)

aguilerai Kusnezov, 1952 (Argentina)

aktaci Karaman, C., 2013 (Turkey)

alacer Forel, 1912 (Brazil)

albicoxis Forel, 1899 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

albipes Emery, 1893 (Sri Lanka)

albistramineus Wheeler, W.M., 1936 (Greater Antilles, Haiti)

albivillosus Zhou, 2001 (China)

alboannulatus Mayr, 1887 (Bolivia, Brazil, Venezuela)

alboannulatus montanus Emery, 1894 (Bolivia)

alboannulatus nessus Forel, 1912 (Brazil)

albocinctus (Ashmead, 1905) (New Guinea, Philippines)

albosparsus Bingham, 1903 (China, India, Japan)

alii Forel, 1890 (Algeria, United Arab Emirates)

alii auresi Emery, 1920 (Algeria)

alii dallatorrei Özdikmen, 2010 (Algeria)

• Camponotus alii concolor Dalla Torre, 1893

alii hoggarensis Santschi, 1929 (Algeria)

altivagans Wheeler, W.M., 1936 (Greater Antilles, Haiti)

amamianus Terayama, 1991 (Japan, Republic of Korea)

amaurus Espadaler, 1997 (Iberian Peninsula, Spain)

americana (Buckley, 1866)

americanus Mayr, 1862 (United States)

• Camponotus castaneus rufinasis Santschi, 1936

amoris Forel, 1904 (Brazil)

amphidus Santschi, 1926 (Zimbabwe)

anatolicus Karaman, C. & Aktac, 2013 (Turkey)

anderseni McArthur & Shattuck, 2001 (Australia)

andrei Forel, 1885 (Mexico)

andrei cholericus Wheeler, W.M., 1914 (Mexico)

andrewsi Donisthorpe, 1936 (Christmas Island)

andrius Dalla Torre, 1893 (Canary Islands, Greece)

andyyoungi McArthur, 2008 (Australia)

anezkae Klimes & McArthur, 2014 (New Guinea)

anguliceps Stitz, 1938 (Borneo, Indonesia)

angusticeps Emery, 1886 (South Africa)

angusticollis (Jerdon, 1851) (Bangladesh, Borneo, India, Nepal)

• Camponotus prismaticus Mayr, 1862

• Formica ardens Smith, F., 1858

• Formica impetuosa Smith, F., 1858

angusticollis sanguinolentus Forel, 1895 (Borneo, India)

annetteae McArthur & Shattuck, 2001 (Australia)

anningensis Wu & Wang, 1989 (China)

annulatus Karavaiev, 1929 (Indonesia)

anthrax Wheeler, W.M., 1911 (Mexico, United States)

apicalis (Mann, 1916) (Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, Trinidad and Tobago, Venezuela)

• Dendromyrmex apicalis filiae Weber, 1943

• Dendromyrmex apicalis guyanensis Santschi, 1921

• Dendromyrmex apicalis opaciceps Wheeler, W.M., 1923

• Dendromyrmex wheeleri Donisthorpe, 1937

apostolus Forel, 1901 (Costa Rica)

arabicus Collingwood, 1985 (Saudi Arabia)

arboreus (Smith, F., 1858) (Brazil, Ecuador, Panama, Paraguay)

• Camponotus adpressisetosus Forel, 1879

arcuatus Mayr, 1876 (Australia)

arcuatus aesopus Forel, 1907 (Australia)

arenatus Shattuck & McArthur, 2002 (Australia)

argus Santschi, 1935 (Democratic Republic of Congo)

arhuacus Forel, 1902 (Colombia)

armeniacus Arnoldi, 1967 (Armenia, Iran)

arminius Forel, 1910 (Mozambique, Yemen)

arminius bicontractus Wheeler, W.M., 1922 (South Africa)

armstrongi McAreavey, 1949 (Australia)

arnoldinus Forel, 1914 (South Africa)

arrogans (Smith, F., 1858) (Bangladesh, Borneo, India, Indonesia, Singapore, Thailand)

• Camponotus maculatus junctus Forel, 1892

aruensis Karavaiev, 1933 (Indonesia, New Guinea)

ashokai Karmaly & Narendran, 2006 (India)

asli Dumpert, 1989 (Malaysia)

atlantis Forel, 1890 (Canary Islands, Iran, Tunisia, United Arab Emirates)

atlantis atrioris Santschi, 1922 (Algeria)

atlantis hesperius Emery, 1893 (Canary Islands)

atlantis marocanus Emery, 1925 (Morocco)

atlantis nigrovarius Forel, 1894 (Algeria)

atra (Buckley, 1866)

atriceps (Smith, F., 1858) (Argentina, Barbados, Belize, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States, Uruguay, Venezuela)

• Camponotus abdominalis costaricensis Forel, 1899

• Camponotus abdominalis cupiens Forel, 1908

• Camponotus abdominalis depressidens Santschi, 1922

• Camponotus abdominalis epistomatus Santschi, 1936

• Camponotus abdominalis fuchsae Forel, 1908

• Camponotus abdominalis fumata Stitz, 1933

• Camponotus abdominalis laevilata Santschi, 1922

• Camponotus abdominalis rabidus Santschi, 1922

• Camponotus abdominalis romani Wheeler, W.M., 1923

• Camponotus abdominalis transvectus Wheeler, W.M., 1910

• Camponotus atriceps atricipitoesuriens Forel, 1879

• Camponotus atriceps stercorarius Forel, 1885

• Camponotus atriceps ustulatus Forel, 1885

• Camponotus taeniatus Roger, 1863

• Camponotus ustulatus mediopallidus Forel, 1901

• Camponotus vulpinus Mayr, 1862

• Formica abdominalis Fabricius Fabricius, 1804

• Formica esuriens Smith, F., 1858

• Formica fulvacea Norton, 1868

atriceps nocens Wheeler, W.M., 1911 (Grenada)

atricolor (Nylander, 1849) (Austria, Bulgaria, Greece, Hungary, Romania, Russian Federation)

atriscapus Santschi, 1926 (Democratic Republic of Congo)

atrox Emery, 1925 (Democratic Peoples Republic of Korea, Republic of Korea)

• Camponotus herculeanus cruentata Emery, 1925

• Camponotus herculeanus jeholensis Teranishi, 1936

• Camponotus herculeanus koreanus Teranishi, 1940

augustei Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

auratiacus Zhou, 2001 (China)

auratus Karavaiev, 1935 (Vietnam)

aurelianus Forel, 1912 (Indonesia)

aureopilus Viehmeyer, 1914 (New Guinea)

• Camponotus aureopilus velutinus Stitz, 1938

aureus Dumpert, 2006 (Thailand)

auricomus Roger, 1862 (Colombia, French Guiana, Greater Antilles, Guyana, Lesser Antilles, Mexico, Saint Vincent and the Grenadines, Suriname, Trinidad and Tobago)

auricomus lucianus Forel, 1899 (Lesser Antilles)

auricomus vincentensis Forel, 1899 (Lesser Antilles)

auriculatus Mayr, 1897 (Sri Lanka)

auriventris Emery, 1889 (Borneo, Myanmar, Thailand, Vietnam)

aurocinctus (Smith, F., 1858) (Australia)

• Camponotus midas Froggatt, 1896

aurofasciatus Santschi, 1915 (São Tomé & Principe)

auropubens Forel, 1894 (Mayotte, Mozambique, Zimbabwe)

auropubens absalon Santschi, 1915 (South Africa)

auropubens aldabrensis Forel, 1897 (Mozambique, Seychelles)

auropubens argentopubens Santschi, 1915 (Angola)

auropubens jacob Santschi, 1915 (Mozambique)

aurosus Roger, 1863 (Mauritius, Réunion)

autrani Forel, 1886 (Borneo, Indonesia)

avius Santschi, 1926 (Zimbabwe)

avius hertigi Santschi, 1937 (Angola)

B
badius (Smith, F., 1857) (Borneo, India, Singapore)

badius saginatus Stitz, 1925 (Borneo)

bakeri Wheeler, W.M., 1904 (United States)

baldaccii Emery, 1908 (Greece, Israel, Turkey)

balzani Emery, 1894 (Bolivia, Brazil, Ecuador, Paraguay)

banghaasi Emery, 1903 (Peru)

barbaricus Emery, 1905 (Gibraltar, Iberian Peninsula, Malta, Portugal, Spain)

barbaricus eubarbaricus Cagniant, 1970 (Algeria)

barbaricus xanthomelas Emery, 1905 (Algeria)

barbarossa Emery, 1920 (Cameroun)

barbarossa micipsa Wheeler, W.M., 1922 (Democratic Republic of Congo)

barbarossa sulcatinasis Santschi, 1926 (Democratic Republic of Congo)

barbatus Roger, 1863 (Borneo, Philippines, Sri Lanka)

barbatus infuscoides Bingham, 1903 (Sri Lanka)

barbatus samarus Santschi, 1932 (Borneo)

barbatus taylori Forel, 1892 (Bangladesh, China, India)

barbosus Baroni Urbani, 1971 (Philippines)

• Camponotus barbatus Menozzi Menozzi, 1926

baronii Alayo & Zayas Montero, 1977 (Cuba)

basuto Arnold, 1958 (Lesotho)

batesii Forel, 1895 (Madagascar)

bayeri Forel, 1913 (Democratic Republic of Congo, Kenya)

baynei Arnold, 1922 (South Africa)

beccarii Emery, 1887 (Indonesia)

bedoti Emery, 1893 (Borneo, China, Indonesia, Krakatau Islands, New Guinea, Solomon Islands)

beebei Wheeler, W.M., 1918 (Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)

belligerus Santschi, 1920 (Namibia)

bellus Forel, 1908 (Indonesia)

• Camponotus ornatus Forel, 1907

bellus adustus Viehmeyer, 1916 (Singapore)

bellus leucodiscus Wheeler, W.M., 1919 (Borneo, Philippines)

belumensis Dumpert, 1995 (Malaysia)

benguelensis Santschi, 1911 (Angola)

bermudezi Aguayo, 1932 (Cuba, Greater Antilles)

berthoudi Forel, 1879 (South Africa)

bertolonii Emery, 1895 (Kenya, South Africa)

bianconii Emery, 1895 (South Africa)

bidens Mayr, 1870 (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Suriname, Trinidad and Tobago)

bidens repressus Forel, 1912 (Brazil)

bifossus Santschi, 1917 (South Africa)

bigenus Santschi, 1919 (Australia)

binghamii Forel, 1894 (Myanmar)

biolleyi Forel, 1902 (Costa Rica)

bishamon Terayama, 1999 (Japan)

bispinosus Mayr, 1870 (Colombia, Ecuador)

bituberculatus André, 1889 (Guinea, Senegal, Sierra Leone)

blandus (Smith, F., 1858) (Argentina, Bolivia, Brazil, Costa Rica, French Guiana, Guyana, Honduras, Mexico, Paraguay, Suriname, Uruguay, Venezuela)

blandus aequinotus Forel, 1902 (Venezuela)

blandus denudatus Emery, 1903 (Bolivia)

blandus pronotalis Santschi, 1936 (Panama)

bocki Forel, 1907 (Bolivia)

boghossiani Forel, 1911 (Greece, Turkey)

boghossiani stenoticus Emery, 1915

• Camponotus kiesenwetteri angustatus Forel, 1889

bonariensis Mayr, 1868 (Argentina, Brazil, Paraguay, Uruguay)

bonariensis curtulus Santschi, 1936 (Brazil)

bonariensis garbei Santschi, 1922 (Brazil)

bonariensis luteolus Emery, 1906 (Argentina)

• Camponotus maculatus carolinus Forel, 1916

bonariensis parvulus Emery, 1894 (Brazil)

bonariensis tucumanus Santschi, 1922 (Argentina)

bonariensis weiseri Santschi, 1922 (Argentina)

borellii Emery, 1894 (Argentina)

borellii saltensis Emery, 1925 (Argentina)

bottegoi Emery, 1895 (Kenya, Somalia)

brachycephalus Santschi, 1920 (Vietnam)

branneri (Mann, 1916) (Brazil, Ecuador)

brasiliensis Mayr, 1862 (Brazil, Paraguay)

brasiliensis chacoensis Santschi, 1919 (Argentina)

brasiliensis clivius Emery, 1925 (Brazil)

brasiliensis costaricanus Özdikmen, 2010 (Costa Rica)

• Camponotus brasiliensis antennatus Santschi, 1925

braunsi Mayr, 1895 (Kenya, Mozambique, Zimbabwe)

braunsi candidus Santschi, 1926 (Tanzania)

braunsi epinotalis Santschi, 1916 (Zimbabwe)

braunsi erythromelus Emery, 1896 (Kenya, Somalia)

brettesi Forel, 1899 (Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Trinidad and Tobago)

brettesi canalis Forel, 1912 (Panama)

brevicollis Stitz, 1916 (Congo)

brevis Forel, 1899 (Colombia, Costa Rica, Ecuador, Panama)

brevis obscurifrons Santschi, 1936 (Panama)

brevis pauli Forel, 1908 (Costa Rica)

breviscapus Zhou, 2001 (China)

brevisetosus Forel, 1910 (South Africa)

britteni Donisthorpe, 1931 (United Kingdom of Great Britain and Northern Ireland)

brookei Forel, 1914 (South Africa)

bruchi Forel, 1912 (Argentina, Bolivia)

• Camponotus distinguendus boliviana Stitz, 1933

bruchi lysistrata Santschi, 1919 (Argentina)

bruchi titicacensis Emery, 1920 (Peru)

brullei (Smith, F., 1858) (Canary Islands)

• Formica carinata Brullé, 1840

bruneiensis Viehmeyer, 1922 (Borneo, Brunei Darussalam)

brunni Forel, 1901 (Japan)

brutus Forel, 1886 (Angola, Cameroun, Ivory Coast, Kenya, Sierra Leone)

bryani Santschi, 1928 (Fiji)

buchholzi Mayr, 1902 (Cameroun)

buddhae Forel, 1892 (India, Iran, Kyrgyzstan, Russian Federation, Turkey, Turkmenistan)

• Camponotus fedtschenkoi obliquipilosa Emery, 1920

bugnioni Forel, 1899 (Colombia)

burgeoni Santschi, 1926 (Democratic Republic of Congo)

burtoni Mann, 1916 (Brazil)

butteli Forel, 1905 (Madagascar)

buttikeri Arnold, 1958 (South Africa)

C
cacicus Emery, 1903 (Brazil, Ecuador, Peru)

caesar Forel, 1886 (Angola, Guinea)

caesar imperator Emery, 1899 (Cameroun)

caffer Emery, 1895 (South Africa)

callistus Emery, 1911 (Bolivia, Ecuador)

callistus bradleyi Wheeler, W.M., 1934 (Ecuador, Peru)

callmorphus Stitz, 1923 (Namibia)

calvus Emery, 1920 (Malaysia)

cambouei Forel, 1891 (Madagascar)

camelinus (Smith, F., 1857) (Borneo, India, Indonesia, Singapore, Thailand)

• Camponotus senilis Mayr, 1862

camelus Emery, 1883 (New Caledonia)

cameranoi Emery, 1894 (Argentina, Brazil, Paraguay)

cameranoi eugaster Santschi, 1922 (Brazil)

cameranoi inlex Forel, 1901 (Brazil)

cameratus Viehmeyer, 1925 (Australia)

cameroni Forel, 1892 (China, Ecuador)

• Camponotus mayri Cameron Cameron, 1891

candiotes Emery, 1894 (Greece, Turkey)

canescens Mayr, 1870 (Bolivia, Brazil, Colombia, Costa Rica, Panama, Trinidad and Tobago)

canescens antennatus Santschi, 1920 (Bolivia)

canescens stomatus Santschi, 1925 (Panama)

capito Mayr, 1876 (Australia)

capito ebeninithorax Forel, 1915 (Australia)

capperi Forel, 1899 (Greater Antilles, Jamaica)

capperi corticalis Forel, 1899 (Jamaica)

capperi formosulus Wheeler, W.M., 1917 (Jamaica)

capperi subdepilis Wheeler, W.M., 1917 (Jamaica)

capperi unctulus Wheeler, W.M., 1917 (Jamaica)

caracalla Forel, 1912 (Brazil)

carbo Emery, 1877 (Eritrea, Ethiopia, Oman)

carbo occidentalis Mayr, 1902 (Ghana)

carbonarius (Latreille, 1802) (Greater Antilles)

carin Emery, 1889 (Borneo, China, Myanmar, Philippines)

carin babiensis Emery, 1925 (Indonesia)

carin tenuisquamis Forel, 1901 (Indonesia)

carin tipunus Forel, 1913 (Taiwan)

caryae (Fitch, 1855) (China, Mexico, United States)

casicus Santschi, 1920 (French Guiana)

castaneus (Latreille, 1802) (United States)

• Camponotus clarus Mayr, 1862

• Formica mellea Say Say, 1836

castanicola Donisthorpe, 1943 (Philippines)

• Camponotus castanea Stitz, 1925

catalanus Emery, 1924 (Spain)

cecconii Emery, 1908 (Cyprus, Greece, Iran)

cerberulus Emery, 1920 (Mexico)

ceriseipes Clark, 1938 (Australia)

cervicalis Roger, 1863 (Madagascar)

cervicalis gaullei Santschi, 1911 (Madagascar)

ceylonicus Emery, 1925 (Sri Lanka)

• Colobopsis testacea Bingham, 1903

chalceoides Clark, 1938 (Australia)

chalceus Crawley, 1915 (Australia)

championi Forel, 1899 (Costa Rica, Mexico, Panama)

chapini Wheeler, W.M., 1922 (Democratic Republic of Congo)

chapini ganzii Weber, 1943 (Sudan)

chartifex (Smith, F., 1860) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Nicaragua, Panama, Suriname)

• Dendromyrmex madeirensis melinoni Santschi, 1921

chazaliei Forel, 1899 (Lesser Antilles)

cheesmanae Donisthorpe, 1932 (Colombia)

chilensis (Spinola, 1851) (Argentina, Chile, Peru)

chloroticus Emery, 1897 (Australia, Cook Islands, Fiji, Guam, Marshall Islands, Micronesia (Federated States of), New Caledonia, New Guinea, Niue, Northern Mariana Islands, Samoa, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Camponotus irritans chlorogaster Emery, 1920

• Camponotus maculatus sanctaecrucis Mann, 1919

chongqingensis Wu & Wang, 1989 (China)

christi Forel, 1886 (Madagascar)

christi ambustus Forel, 1892 (Madagascar)

christi ferrugineus Emery, 1899 (Madagascar)

christi foersteri Forel, 1886 (Madagascar)

christi maculiventris Emery, 1895 (Madagascar)

christmasensis McArthur, 2008 (Australia)

christophei Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

christopherseni Forel, 1912 (Guatemala, Panama)

chromaiodes Bolton, 1995 (United States)

• Formica ferruginea Fabricius Fabricius, 1798

chrysurus Gerstäcker, 1871 (Kenya)

chrysurus acutisquamis Mayr, 1902 (Cameroun)

chrysurus apellis Wheeler, W.M., 1922 (Angola)

chrysurus securifer Emery, 1920 (Cameroun)

chrysurus yvonnae Forel, 1920 (Democratic Republic of Congo)

churchetti McArthur, 2008 (Australia)

cilicicus Emery, 1908 (Iraq, Israel, Turkey)

cillae Forel, 1912 (Brazil)

cinctellus (Gerstäcker, 1859) (Eritrea, Kenya, Mozambique, Rwanda, Tanzania, Zimbabwe)

• Camponotus venustus Mayr, 1867

cinctellus belliceps Santschi, 1939 (Democratic Republic of Congo)

cinerascens (Fabricius, 1787) (India, Philippines)

cinereus Mayr, 1876 (Australia)

cinereus amperei Forel, 1913 (Australia)

cinereus notterae Forel, 1907 (Australia)

cingulatus Mayr, 1862 (Bolivia, Brazil, Paraguay, Uruguay)

• Camponotus cingulatus bambusarum Forel, 1902

• Camponotus cingulatus brunneiventris Santschi, 1920

• Camponotus cingulatus damocles Forel, 1909

• Camponotus cingulatus fissa Stitz, 1933

• Camponotus cingulatus myster Santschi, 1922

• Camponotus rufipes lessonai Emery, 1894

circularis Mayr, 1870 (Colombia)

circularis rufitibia Emery, 1920 (Peru)

circumspectus (Smith, F., 1861) (Indonesia, Sulawesi)

clarior Forel, 1902 (Australia)

claripes Mayr, 1876 (Australia)

• Camponotus claripes piperatus Wheeler, W.M., 1933

claripes inverallensis Forel, 1910 (Australia)

claripes nudimalis Forel, 1913 (Australia)

claripes orbiculatopunctatus Viehmeyer, 1925 (Australia)

clarithorax Creighton, 1950 (Mexico, United States)

claviscapus Forel, 1899 (Belize, Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Trinidad and Tobago, Venezuela)

claviscapus lehmanni Forel, 1911 (Nicaragua)

claviscapus occultus Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

claviscapus subcarinatus Forel, 1899 (Guatemala)

cleobulus Santschi, 1919 (South Africa)

clerodendri (Emery, 1887) (Borneo, Indonesia)

clypeatus Mayr, 1866 (Brazil)

cocosensis Wheeler, W.M., 1919 (Cocos (Keeling) Islands)

cognatocompressus Forel, 1904

coloratus Forel, 1904 (Colombia, Costa Rica, Mexico, Paraguay)

compositor Santschi, 1922 (Bolivia, Brazil)

compressiscapus André, 1889 (Senegal, Sierra Leone)

compressus (Fabricius, 1787) (Bangladesh, Borneo, Canary Islands, China, India, Liechtenstein, Nepal, Philippines, United Arab Emirates)

• Camponotus quadrilaterus Roger Roger, 1863

• Formica callida Smith, F., 1858

• Formica indefessa Sykes, 1835

compressus bernardi Özdikmen, 2010 (Algeria)

• Camponotus compressus nitens Bernard, 1953

compressus martensi Forel, 1907 (Algeria)

compressus occipitalis Stitz, 1917 (Algeria)

compressus probativus Santschi, 1921 (Mozambique)

compressus rectinotus Santschi, 1921 (Libya)

concavus Kim & Kim, 1994 (Greece, Republic of Korea)

concolor Forel, 1891 (Madagascar)

• Camponotus gallienii Forel, 1916

confluens Forel, 1913 (Democratic Republic of Congo)

confluens bequaerti Forel, 1913 (Democratic Republic of Congo)

confluens trematogaster Santschi, 1915 (Democratic Republic of Congo)

confucii Forel, 1894 (China, India)

congolensis Emery, 1899 (Democratic Republic of Congo, Guinea, Ivory Coast, Liberia, Sierra Leone, Zimbabwe)

congolensis weissi Santschi, 1911 (Congo)

coniceps Santschi, 1926 (Congo)

conicus (Mayr, 1876) (Tonga)

conithorax Emery, 1914 (Australia, New Guinea, Vanuatu)

conradti Forel, 1914 (Cameroun)

conradti fimbriatipes Santschi, 1920 (Cameroun)

consanguineus (Smith, F., 1861) (Indonesia, Sulawesi)

consectator (Smith, F., 1858) (Australia)

consobrinus (Erichson, 1842) (Australia)

• Camponotus dimidiatus Roger, 1863

• Camponotus nigriceps obniger Forel, 1902

conspicuus (Smith, F., 1858) (Argentina, Bahamas, Brazil, Costa Rica, Cuba, Greater Antilles, Guatemala, Jamaica, Lesser Antilles, Mexico, Panama, Uruguay)

conspicuus inaequalis Roger, 1863 (Cuba)

conspicuus sharpi Forel, 1893 (Mexico)

conspicuus williamsi Forel, 1916 (Lesser Antilles)

conspicuus zonatus Emery, 1894 (Costa Rica, Ecuador, Galapagos Islands, Honduras, Mexico, Panama)

constructor Forel, 1899 (Costa Rica, Ecuador, Panama)

contractus Mayr, 1872 (Borneo)

contractus buttesi Forel, 1902 (Malaysia)

contractus scortechinii Emery, 1888 (Malaysia)

conulus Mayr, 1870 (Colombia)

convexiclypeus Mackay, W.P., 1997 (Peru)

coptobregma Kempf, 1968 (Brazil)

corallinus (Roger, 1863) (Philippines)

cordiceps Santschi, 1939 (Argentina)

cordincola Wheeler, W.M., 1934 (Panama)

coriolanus Forel, 1912 (Brazil)

• Camponotus scrobifer Borgmeier, 1928

corniculatus Wheeler, W.M., 1934 (Guyana)

cornis Wang, C. & Wu, 1994 (China)

coruscus (Smith, F., 1862) (Colombia, Costa Rica, Guatemala, Honduras, Mexico, Panama, Venezuela)

coruscus astilbus Stitz, 1938 (Suriname)

coruscus fulgens Forel, 1885 (Guatemala)

cosmicus (Smith, F., 1858) (Mozambique, South Africa, Zimbabwe)

cotesii Forel, 1893 (China, India)

cowlei Froggatt, 1896 (Australia)

coxalis (Smith, F., 1859) (Indonesia, New Guinea)

crassicornis Emery, 1920 (Bolivia, Brazil)

crassisquamis Forel, 1902 (Bangladesh, India)

crassus Mayr, 1862 (Argentina, Brazil, Paraguay, Trinidad and Tobago)

• Camponotus crassus picticornis Santschi, 1936

• Camponotus flexus Mayr, 1862

• Camponotus vezenyii Forel, 1907

crassus amazonicus Santschi, 1922 (Brazil)

crassus delabiatus Santschi, 1925 (Brazil)

crawleyi Emery, 1920 (Cameroun)

crenatus Mayr, 1876 (Australia)

crepusculi Arnold, 1922 (Zimbabwe)

cressoni André, 1887 (Bolivia, Brazil, Mexico)

cressoni purensis Forel, 1912 (Brazil)

criniticeps Menozzi, 1939 (Ethiopia)

crispulus Santschi, 1922 (Argentina)

cristatus Mayr, 1866 (Fiji)

• Camponotus cristatus nagasau Mann, 1921

crozieri McArthur & Leys, 2006 (Australia)

crucheti Santschi, 1911 (Angola)

cruentatus (Latreille, 1802) (Andorra, France, Gibraltar, Iberian Peninsula, Monaco, Morocco, Portugal, Spain)

• Camponotus cruentatus cruentior Santschi, 1931

• Camponotus cruentatus lindbergi Santschi, 1931

• Formica gigas Leach, 1825

• Formica opaca Nylander, 1856

cruentatus asper Menozzi, 1925 (Australia)

cuauhtemoc Snelling, R.R., 1988 (Mexico, Mexico, United States)

cubangensis Forel, 1901 (Angola)

cubangensis dofleini Forel, 1911 (South Africa)

• Camponotus mayri ledieui Forel, 1916

culmicola Wheeler, W.M., 1905 (Bahamas)

culmicola haweisi Mann, 1920 (Bahamas)

cuneidorsus Emery, 1920 (Costa Rica, Guatemala, Honduras, Mexico, Peru)

cuneiscapus Forel, 1910 (Namibia, South Africa)

curviscapus Emery, 1896 (Costa Rica, Mexico)

custodulus Emery, 1911 (New Guinea)

cylindricus (Fabricius, 1798) (Borneo, Mauritius, Thailand)

• Colobopsis doriae Mayr, 1872

cyrtomyrmodes Donisthorpe, 1941 (New Guinea)

D
daitoensis Terayama, 1999 (Japan)

dalmasi Forel, 1899 (Colombia)

dalmaticus (Nylander, 1849) (Bulgaria, Croatia, Greece, Israel, Montenegro, Republic of Macedonia, Slovenia, Switzerland, Turkey)

darlingtoni Wheeler, W.M., 1934 (Australia)

• Camponotus rottnesti Donisthorpe, 1941

darwinii Forel, 1886 (Madagascar, Seychelles)

darwinii rubropilosus Forel, 1891 (Madagascar)

darwinii themistocles Forel, 1910 (Madagascar)

debellator Santschi, 1926 (Zimbabwe)

decipiens Emery, 1893 (Mexico, United States)

• Camponotus fallax rasilis Wheeler, W.M., 1910

declivus Santschi, 1922 (Mexico)

dedalus Forel, 1911 (Indonesia)

dedalus moeschiellus Forel, 1912 (Indonesia)

deletangi Santschi, 1920 (Bolivia)

densopilus Shattuck, 2005 (Indonesia, New Guinea)

dentatus (Mayr, 1866) (Fiji)

depressiceps Forel, 1879 (Brazil)

depressus Mayr, 1866 (Brazil, Paraguay)

desantii Santschi, 1915 (Benin)

descarpentriesi Santschi, 1926 (Madagascar)

desectus (Smith, F., 1860) (New Guinea)

detritus Emery, 1886 (Namibia, South Africa)

devestivus Wheeler, W.M., 1928 (China, Japan)

dewitzii Forel, 1886 (Democratic Republic of Congo)

dicksoni Arnold, 1948 (South Africa)

difformis Stitz, 1938 (New Guinea)

dimorphus Emery, 1894 (Bolivia, Brazil, Paraguay)

diplopunctatus Emery, 1915 (Eritrea, Ethiopia)

diplopunctatus subconvexus Viehmeyer, 1923 (Sudan)

discolor (Buckley, 1866) (United States)

discors Forel, 1902 (Australia)

discors angustinodus Emery, 1925 (Australia)

discors laetus Emery, 1925 (Australia)

discors yarrabahensis Forel, 1915 (Australia)

distinguendus (Spinola, 1851) (Argentina, Chile, Peru)

• Camponotus distinguendus denudata Emery, 1906

• Camponotus distinguendus santacruziana Emery, 1906

• Camponotus distinguendus tenuipubens Santschi, 1916

divergens Mayr, 1887 (Brazil)

diversipalpus Santschi, 1922 (Brazil)

dolabratus Menozzi, 1927 (Costa Rica)

dolendus Forel, 1892 (China, India, Nepal)

dolichoderoides Forel, 1911 (Borneo, Indonesia, Thailand)

donisthorpei Emery, 1920 (Cameroun)

donnellani Shattuck & McArthur, 2002 (Australia)

dorycus (Smith, F., 1860) (China, Indonesia, New Guinea)

dorycus confusus Emery, 1887 (Australia, New Guinea)

dorycus recticeps Stitz, 1938 (New Guinea)

dracocephalus Stitz, 1938 (Borneo, Indonesia)

dromas Santschi, 1919 (Australia)

dromedarius Forel, 1891 (Madagascar)

dromedarius pulcher Forel, 1892 (Madagascar)

druryi Forel, 1886 (Tanzania)

dryandrae McArthur, 1996 (Australia)

dufouri Forel, 1891 (Central African Republic, Madagascar)

dufouri imerinensis Forel, 1891 (Madagascar)

dumetorum Wheeler, W.M., 1910 (Mexico, United States)

E
eastwoodi McArthur, 1996 (Australia)

echinoploides Forel, 1891 (Madagascar)

edmondi André, 1887 (Comoros, Madagascar)

edmondi ernesti Forel, 1891 (Madagascar)

egregius (Smith, F., 1858) (Brazil)

elegans Forel, 1902 (Australia)

elevatus Forel, 1899 (Costa Rica, Mexico)

ellioti Forel, 1891 (Madagascar)

ellioti relucens Santschi, 1911 (Madagascar)

elysii Mann, 1919 (Solomon Islands)

emarginatus Emery, 1886 (South Africa)

emeryodicatus Forel, 1901 (Brazil, Ecuador, Paraguay, Zimbabwe)

• Camponotus emeryi Forel, 1901 Forel, 1901

emeryodicatus decessor Forel, 1908 (Brazil)

empedocles Emery, 1920 (United Arab Emirates, Zimbabwe)

enigmaticus Mackay, W.P., Mackay, M. & Mackay, E.E., 2004 (Nicaragua)

eperiamorum Clouse, 2007 (Micronesia (Federated States of))

ephippiatus Viehmeyer, 1916 (Singapore)

ephippium (Smith, F., 1858) (Australia)

ephippium narses Forel, 1915 (Australia)

equus Stitz, 1932 (Indonesia)

eremicus Wheeler, W.M., 1915 (Australia)

erigens Forel, 1894 (Algeria)

erigens subconcolor Emery, 1920 (Algeria)

erinaceus Gerstäcker, 1871 (Kenya, Tanzania)

errabundus Arnold, 1949 (Zimbabwe)

erythrocephalus Clouse, 2007 (Micronesia (Federated States of))

erythrostoma Emery, 1920 (Brazil)

esau Forel, 1915 (Australia)

essigi Smith, M.R., 1923 (Mexico, Trinidad and Tobago, United States)

• Camponotus nevadensis Gregg, 1973

• Camponotus pertusus Mackay, W.P., 1997

ethicus Forel, 1897 (Madagascar)

etiolatus Wheeler, W.M., 1904 (Mexico, United States)

etiolipes Bolton, 1995 (Kenya, Mozambique, Zimbabwe)

• Formica longipes Gerstäcker Gerstäcker, 1859

eugeniae Forel, 1879 (Lesotho, Mozambique, South Africa, Zimbabwe)

eugeniae amplior Forel, 1913 (Democratic Republic of Congo)

eurynotus Forel, 1907 (Bolivia, Brazil, Ecuador)

evae Forel, 1910 (Australia)

evae zeuxis Forel, 1915 (Australia)

evansi Crawley, 1920 (Iraq)

excavatus Donisthorpe, 1948 (New Guinea)

excisus Mayr, 1870 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Panama, Trinidad and Tobago)

exiguoguttatus Forel, 1886 (China, India)

exsectus Emery, 1900 (Borneo, Indonesia)

extensus Mayr, 1876 (Australia)

ezotus Bolton, 1995 (Australia)

• Camponotus erythropus Viehmeyer Viehmeyer, 1925

F
fabricator (Smith, F., 1858) (Saint Helena)

falco Forel, 1902 (Colombia)

fallatus Bolton, 1995 (Brazil)

• Camponotus escherichi Menozzi, 1926

fallax (Nylander, 1856) (Albania, Andorra, Armenia, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Liechtenstein, Lithuania, Luxembourg, Malta, Montenegro, Netherlands, Poland, Portugal, Republic of Korea, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)

• Camponotus fallax pageti Hamann & Klemm, 1976

• Camponotus marginatus kamensis Ruzsky, 1903

• Camponotus marginatus ruzskyi Emery, 1898

fasciatellus Dalla Torre, 1892 (Bolivia, Colombia, French Guiana, Guyana, Mexico, Suriname)

• Camponotus fasciatus Mayr Mayr, 1870

fasciatus (Mayr, 1867) (Borneo, Indonesia)

fastigatus Roger, 1863 (Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Suriname, Trinidad and Tobago)

fastigatus barbiculus Santschi, 1922 (Bolivia)

fastigatus schmalzi Emery, 1894 (Brazil)

fastigatus verae Forel, 1908 (Brazil)

favorabilis Santschi, 1919 (Zimbabwe)

fayfaensis Collingwood, 1985 (Saudi Arabia, United Arab Emirates)

fedtschenkoi Mayr, 1877 (Afghanistan, Armenia, China, Iran, Kazakhstan, Kyrgyzstan, Russian Federation, Turkmenistan)

• Camponotus fedtschenkoi adi Stitz, 1930

• Camponotus tichomirovi Ruzsky, 1905

fellah Dalla Torre, 1893 (Egypt, Eritrea, Iran, Iraq, Israel, Kuwait, Oman, Turkey, United Arab Emirates)

femoratus (Fabricius, 1804) (Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago, Venezuela)

fergusoni McArthur, 2003 (Australia)

ferreri Forel, 1913 (Cameroun)

ferreri akka Forel, 1916 (Democratic Republic of Congo)

ferreri cavisquamis Santschi, 1926 (Kenya)

fervidus Donisthorpe, 1943 (Borneo)

• Formica fervens Smith, F. Smith, F., 1857

festai Emery, 1894 (Greece, Israel, Lebanon, Turkey)

• Camponotus maculatus caeciliae Forel, 1906

festinatus (Buckley, 1866) (Mexico, United States)

• Camponotus fumidus pubicornis Emery, 1893

• Camponotus fumidus spurcus Wheeler, W.M., 1910

festinus (Smith, F., 1857) (Borneo)

festinus cetegus Forel, 1911 (Borneo, Indonesia)

festinus diligens (Smith, F., 1857) (Malaysia)

festinus eximius Emery, 1900 (Borneo, Indonesia)

festinus inezae Forel, 1913 (Malaysia)

festinus simaluranus Forel, 1915 (Indonesia)

fiebrigi Forel, 1906 (Paraguay)

fieldeae Forel, 1902 (Australia)

fieldellus Forel, 1910 (Australia)

figaro Collingwood & Yarrow, 1969 (Iberian Peninsula, Portugal, Spain)

fijianus Özdikmen, 2010 (Fiji)

• Camponotus dentatus humeralis Wheeler, W.M., 1934

flavescens (Fabricius, 1793) (French Guiana)

flavicomans Clouse, 2007 (Palau)

flavocassis Donisthorpe, 1941 (New Guinea)

flavocrines Donisthorpe, 1941 (New Guinea)

flavolimbatus Viehmeyer, 1922 (Samoa)

• Camponotus buxtoni Santschi, 1928

• Camponotus conithorax nautarum Santschi, 1919

flavomarginatus Mayr, 1862 (Eritrea, Ghana, Guinea, Kenya, Oman, Rwanda, Somalia, South Africa, United Arab Emirates)

• Camponotus micans albisectus Emery, 1892

fletcheri Donisthorpe, 1942 (Sri Lanka)

floridanus (Buckley, 1866) (United States)

• Camponotus atriceps yankee Forel, 1885

florius Santschi, 1926 (Congo)

foleyi Santschi, 1939 (Algeria)

foleyi ajjer Bernard, 1953 (Algeria)

foleyi fezzanensis Bernard, 1953 (Algeria)

foleyi grasi Bernard, 1953 (Algeria)

foleyi lelubrei Bernard, 1953 (Algeria)

foleyi pseudocompressus Özdikmen, 2010 (Algeria)

• Camponotus foleyi hoggarensis Bernard, 1953

foleyi rufescens Bernard, 1953 (Algeria)

foraminosus Forel, 1879 (Cape Verde, Guinea, Kenya, Mozambique, Oman, Senegal)

foraminosus chrysogaster Emery, 1895 (Ethiopia, Kenya)

foraminosus cuitensis Forel, 1901 (Angola)

foraminosus deductus Santschi, 1915 (Nigeria)

foraminosus dorsalis Santschi, 1926 (Tanzania)

foraminosus flavus Stitz, 1916 (Congo)

foraminosus honorus Forel, 1910 (Ethiopia)

foreli Emery, 1881 (Algeria, Greece, Iberian Peninsula, Portugal, Spain)

foreli tingitanus Santschi, 1921 (Morocco)

formiciformis Forel, 1885 (Belize, Ecuador, Guatemala, Mexico)

formosensis Wheeler, W.M., 1927 (Taiwan)

fornasinii Emery, 1895 (Mozambique)

fragilis Pergande, 1893 (Mexico, United States)

fraseri McArthur, 2008 (Australia)

friedae Forel, 1912 (China, Japan, Taiwan)

• Camponotus friedae amia Forel, 1912

froggatti Forel, 1902 (Australia)

frontalis Pergande, 1896 (Mexico)

fryi Mann, 1916 (Brazil)

fugax Forel, 1902 (Jamaica)

fulvopilosus (De Geer, 1778) (Botswana, Lesotho, Namibia, South Africa)

• Camponotus fulvopilosus detritoides Forel, 1910

• Camponotus fulvopilosus flavopilosus Emery, 1895

• Formica pilosa Olivier, 1792

• Formica rufiventris Fabricius Fabricius, 1804

fumidus Roger, 1863 (Argentina, Bahamas, Brazil, Lesser Antilles, Mexico, Venezuela)

fumidus dominicensis Wheeler, W.M., 1913 (Dominican Republic)

fumidus fraterculus Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

fumidus haytianus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus illitus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus imbecillus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus soulouquei Forel, 1901 (Haiti)

fumidus toltecus Forel, 1899 (Mexico)

furvus Santschi, 1911 (Congo)

fuscivillosus Xiao & Wang, 1989 (China)

fuscocinctus Emery, 1888 (Brazil)

fuscus Kim & Kim, 1994 (Republic of Korea)

G
gabonensis Santschi, 1926 (Gabon)

galla Forel, 1894 (Eritrea, Ethiopia, Kenya)

• Camponotus foraminosus latinotus Forel, 1907

gallagheri Collingwood & Agosti, 1996 (Oman)

gambeyi Emery, 1883 (New Caledonia)

gambeyi marthae Forel, 1894 (New Caledonia)

gasseri (Forel, 1894) (Australia, New Guinea)

• Camponotus gasseri caloratus Wheeler, W.M., 1934

• Camponotus gasseri lysias Forel, 1913

• Camponotus gasseri obtusitruncatus Forel, 1902

geayi Santschi, 1922 (French Guiana)

genatus Santschi, 1922 (Brazil)

gentingensis Dumpert, 1995 (Malaysia)

geralensis Emery, 1920 (Argentina, Bolivia, Brazil, Paraguay)

• Camponotus luederwaldti Santschi, 1922

• Camponotus macrocephalus Emery Emery, 1894

gerberti Donisthorpe, 1949 (Madagascar)

germaini Emery, 1903 (Brazil, Paraguay)

• Camponotus germaini tacuruensis Emery, 1911

gestroi Emery, 1878 (Algeria, Armenia, Bulgaria, Gibraltar, Greece, Iberian Peninsula, Iran, Iraq, Israel, Italy, Spain, Turkey)

• Camponotus gestroi creticus Forel, 1886

gestroi ponariensis Santschi, 1921 (Algeria)

gibber Forel, 1891 (Madagascar)

gibbinotus Forel, 1902 (Australia)

gibbosus Karavaiev, 1929 (Indonesia)

gigas (Latreille, 1802) (Borneo, Philippines, Thailand)

gigas borneensis Emery, 1887 (Borneo)

gilviceps Roger, 1863 (Borneo)

• Camponotus smithi Donisthorpe, 1932

• Formica ruficeps Smith, F. Smith, F., 1857

gilviventris Roger, 1863 (Cuba, Greater Antilles)

gilviventris refectus Wheeler, W.M., 1937 (Cuba)

gilviventris renormatus Wheeler, W.M., 1937 (Cuba)

glabrisquamis Emery, 1911 (New Guinea)

godmani Forel, 1899 (Brazil, Mexico, Panama, Trinidad and Tobago)

godmani palliolatus Wheeler, W.M., 1922 (Trinidad and Tobago)

goeldii Forel, 1894 (Brazil)

gombaki Dumpert, 1986 (Malaysia)

gouldi Forel, 1886 (Madagascar)

gouldianus Forel, 1922 (Australia)

grandidieri Forel, 1886 (Comoros, Kenya, Madagascar, Mauritius, Mayotte, Mozambique, Namibia, Seychelles, Zimbabwe)

grandidieri atrabilis Santschi, 1915 (Madagascar)

grandidieri comorensis Santschi, 1915 (Comoros)

grandidieri eumendax Özdikmen, 2010 (South Africa)

• Camponotus foraminosus mendax Emery, 1895

grandidieri ruspolii Forel, 1892 (Somalia)

greeni Forel, 1911 (Sri Lanka)

gretae Forel, 1902 (Myanmar)

guanchus Santschi, 1908 (Canary Islands)

guayapa Kusnezov, 1952 (Argentina)

guidae McArthur, 2007 (Australia)

guizhouensis Wang, M., 1992 (China)

gundlachi Mann, 1920 (Cuba, Greater Antilles)

guppyi Mann, 1919 (Solomon Islands)

guttatus Emery, 1899 (Cameroun, Guinea, Niger)

guttatus minusculus Viehmeyer, 1914 (Tanzania)

H
habereri Forel, 1911 (Japan, Taiwan)

haematocephalus Emery, 1903 (Peru)

haereticus Santschi, 1914 (Nigeria)

hagensii Forel, 1886 (Madagascar)

hannani Forel, 1899 (Greater Antilles, Jamaica)

• Camponotus hannani willardi Forel, 1899

hapi Weber, 1943 (Sudan)

haroi Espadaler, 1997 (Iberian Peninsula, Spain)

hartogi Forel, 1902 (Australia)

• Camponotus ferruginipes Crawley, 1922

hastifer Emery, 1911 (New Guinea)

havilandi Arnold, 1922 (South Africa)

heathi Mann, 1916 (Brazil)

heathi gilvigaster Wheeler, W.M., 1923 (Brazil)

hedwigae Forel, 1912 (Brazil)

helleri Emery, 1903 (Ecuador, Paraguay, Peru)

hellmichi Menozzi, 1935 (Chile)

helvus Xiao & Wang, 1989 (China)

hemichlaena Yasumatsu & Brown, 1951 (Japan)

herculeanus (Linnaeus, 1758) (Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Canada, China, Croatia, Czech Republic, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kazakhstan, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Netherlands, Norway, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United States)

• Camponotus herculeanus caucasicus Arnoldi, 1967

• Camponotus herculeanus eudokiae Ruzsky, 1926

• Camponotus herculeanus montanus Ruzsky, 1904

• Camponotus herculeanus nadigi Menozzi, 1922

• Camponotus herculeanus shitkowi Ruzsky, 1904

• Camponotus herculeanus whymperi Forel, 1902

• Formica atra Zetterstedt, 1838

• Formica castanea Lepeletier de Saint-Fargeau, 1835

• Formica intermedia Zetterstedt, 1838

hermanni Emery, 1911 (Brazil)

heros Santschi, 1926 (Tanzania)

heteroclitus Forel, 1895 (Madagascar)

hildebrandti Forel, 1886 (Madagascar)

hildebrandti dichromothrix Emery, 1920 (Madagascar)

himalayanus Forel, 1893 (India, Nepal)

hippocrepis Emery, 1920 (Bolivia, Ecuador)

hirtus Karaman, C. & Aktac, 2013 (Turkey)

hispidus Emery, 1906 (Paraguay)

hoelldobleri Cagniant, 1991 (Morocco)

holosericeus Emery, 1889 (China, India, Myanmar)

holzi Forel, 1921 (Ecuador)

honaziensis Karaman, C. & Aktac, 2013 (Croatia, Cyprus, Greece, Republic of Macedonia, Spain, Turkey, Turkey)

hoplites Emery, 1914 (New Caledonia)

horni Clark, 1930 (Australia)

horrens Forel, 1910 (Philippines)

horripilus Emery, 1900 (Indonesia)

horseshoetus Datta & Raychaudhuri, 1985 (India)

hosei Forel, 1911 (Borneo, New Guinea)

hosei mimus Forel, 1911 (Borneo)

hospes (Emery, 1884) (Borneo, Indonesia)

hospes adultus Viehmeyer, 1916 (Malaysia)

hova Forel, 1891 (Comoros, Madagascar, Mayotte, Mozambique, Oman, Seychelles)

hova becki Santschi, 1923 (Madagascar)

hova boivini Forel, 1891 (Seychelles)

hova cemeryi Özdikmen, 2010 (Madagascar)

• Camponotus hova luteolus Emery, 1925

hova fairmairei Santschi, 1911 (Madagascar)

hova fulvus Emery, 1894 (Comoros, Seychelles)

hova hovahovoides Forel, 1892 (Madagascar)

hova hovoides Dalla Torre, 1893 (Madagascar)

hova mixtellus Dalla Torre, 1893 (Seychelles)

hova obscuratus Emery, 1925 (Madagascar)

hova pictiventris Mayr, 1901 (South Africa)

hova radamae Forel, 1891 (Mayotte, Mayotte)

howensis Wheeler, W.M., 1927 (Australia, Lord Howe Island)

humeralis Emery, 1920 (Peru)

humerus Wang, C. & Wu, 1994 (China)

humilior Forel, 1902 (Australia)

hunteri Wheeler, W.M., 1910 (United States)

husseini Dietrich, 2004 (Jordan)

hyatti Emery, 1893 (Mexico, United States)

• Camponotus sayi bicolor Pergande, 1894

• Camponotus sayi californica Emery, 1925

hypoclineoides Wheeler, W.M., 1919 (Borneo, Indonesia)

I
icarus Forel, 1912 (Indonesia)

ignestii Menozzi, 1935 (Guyana)

iheringi Forel, 1908 (Brazil, Paraguay)

• Camponotus iheringi bajulus Emery, 1925

• Camponotus iheringi latinota Forel, 1909

ilgii Forel, 1894 (Ethiopia, Senegal)

imitator Forel, 1891 (Madagascar)

imitator resinicola Santschi, 1911 (Madagascar)

immaculatus Forel, 1892 (Madagascar)

immigrans Santschi, 1914 (Cameroun)

importunus Forel, 1911 (Mozambique, South Africa)

impressilabris Stitz, 1938 (Malaysia)

impressus (Roger, 1863) (United States)

improprius (Forel, 1879) (Colombia)

inca Emery, 1903 (Brazil, Peru)

inca rector Santschi, 1923 (Brazil)

incensus Wheeler, W.M., 1932 (United States)

inconspicuus Mayr, 1872 (Borneo, Indonesia)

indefinitus Karavaiev, 1929 (Indonesia)

indeflexus (Walker, 1859) (Sri Lanka)

indicatus Santschi, 1922 (French Guiana)

inflatus Lubbock, 1880 (Australia)

• Camponotus aurofasciatus Wheeler, W.M. Wheeler, W.M., 1915

innexus Forel, 1902 (Australia)

innocens Forel, 1909 (Paraguay)

insipidus Forel, 1893 (Australia)

integellus Forel, 1899 (Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Peru, Venezuela)

• Camponotus rudigenis Emery, 1903

interjectus Mayr, 1877 (Afghanistan, China, Iran, Iraq, Israel, Kazakhstan, Kyrgyzstan, Turkmenistan)

intrepidus (Kirby, W., 1819) (Australia)

• Camponotus magnus Mayr, 1862

• Formica agilis Smith, F., 1858

intrepidus bellicosus Forel, 1902 (Australia)

invidus Forel, 1892 (India)

ionius Emery, 1920 (Bulgaria, Greece, Turkey)

iridis Santschi, 1922 (Brazil)

irritabilis (Smith, F., 1857) (Borneo)

irritabilis winkleri Forel, 1911 (Borneo)

irritans (Smith, F., 1857) (Borneo, China, India, Malaysia, Nicobar Island, Sri Lanka, Vietnam)

• Camponotus agnatus Roger, 1863

irritans carensis Emery, 1920 (Nicobar Island)

irritans carinifer Viehmeyer, 1916 (Singapore)

irritans cliens Forel, 1911 (Indonesia)

irritans croceomaculatus Emery, 1914 (New Caledonia)

irritans curtus Emery, 1900 (Indonesia)

irritans fatuus Forel, 1886 (Indonesia, New Guinea)

irritans hongkongensis Forel, 1912 (China)

irritans inferior Emery, 1925 (Borneo)

irritans kubaryi Mayr, 1876 (Palau)

irritans melanogaster Stitz, 1938 (Micronesia (Federated States of))

irritans pallidus (Smith, F., 1857) (Bangladesh, Borneo, Philippines)

irritans procax Santschi, 1919 (Vietnam)

irritans sumatranus Özdikmen, 2010 (Indonesia)

• Camponotus irritans puberula Emery, 1920

irritans tinctus (Smith, F., 1858) (Myanmar)

isabellae Forel, 1909 (Sri Lanka)

itoi Forel, 1912 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea)

itoi kwansienensis Viehmeyer, 1922 (China)

iwoensis Terayama & Kubota, 2011 (Japan)

J
jaliensis Dalla Torre, 1893 (Greece, Israel)

janeti Forel, 1895 (Australia)

janforrestae McArthur & Shattuck, 2001 (Australia)

japonicus Mayr, 1866 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Philippines, Republic of Korea, Russian Federation)

• Camponotus herculeanus manczshuricus Emery, 1925

• Camponotus japonicus miltotus Wheeler, W.M., 1929

• Camponotus japonicus sanguinea Karavaiev, 1929

• Camponotus japonicus wui Wheeler, W.M., 1929

• Camponotus pennsylvanicus aterrimus Emery, 1895

jeanneli Santschi, 1914 (Kenya)

jejuensis Kim & Kim, 1986 (Republic of Korea)

jianghuaensis Xiao & Wang, 1989 (China)

jizani Collingwood, 1985 (Oman, Saudi Arabia)

johnclarki Taylor, 1992 (Australia)

• Notostigma sanguinea Clark, 1930

judithmorrisae McArthur, 2008 (Australia)

juliae Emery, 1903 (Costa Rica)

K
kadi Mann, 1921 (Fiji)

• Camponotus schmeltzi trotteri Mann, 1921

kaguya Terayama, 1999 (Japan)

karawaiewi Menozzi, 1926 (Philippines)

kattensis Bingham, 1903 (India)

kaura Snelling, R.R. & Torres, 1998 (Greater Antilles, Puerto Rico)

kefir Ionescu-Hirsch, 2010 (Israel)

keiferi Wheeler, W.M., 1934 (Mexico)

keihitoi Forel, 1913 (China, Japan)

• Camponotus caryae teranishii Wheeler, W.M., 1928

• Camponotus tokyoensis Teranishi, 1915

kelleri Forel, 1886 (Madagascar)

kelleri invalidus Forel, 1897 (Madagascar)

keralensis Karmaly & Narendran, 2006 (India)

kersteni Gerstäcker, 1871 (Kenya)

khaosokensis Dumpert, 2006 (Thailand)

• Camponotus hoelldobleri Dumpert Dumpert, 2006

kiesenwetteri (Roger, 1859) (Greece, Iran, Turkey)

kiesenwetteri cyprius Emery, 1920 (Cyprus)

kiusiuensis Santschi, 1937 (Japan, Republic of Korea)

klaesii (Forel, 1886) (Indonesia)

klugii Emery, 1895 (South Africa)

knysnae Arnold, 1922 (South Africa)

kollbrunneri Forel, 1910 (Ethiopia)

kolthoffi Stitz, 1934 (China)

kopetdaghensis Dlussky & Zabelin, 1985 (Iran, Russian Federation, Turkmenistan)

korthalsiae Emery, 1887 (Indonesia)

korthalsiae concilians Forel, 1915 (Indonesia)

• Camponotus jacobsoni Forel, 1915

koseritzi Emery, 1888 (Bolivia, Brazil)

kraepelini Forel, 1901 (Bolivia)

kugleri Ionescu-Hirsch, 2010 (Israel)

kunigamiensis Terayama, 2013 (Japan)

kurdistanicus Emery, 1898 (China, Iran, Iraq, Israel, Russian Federation, Turkey)

kutteri Forel, 1915 (Indonesia)

kutterianus Baroni Urbani, 1972 (Cuba, Greater Antilles)

L
laconicus Emery, 1920 (Greece)

laevigatus (Smith, F., 1858) (Canada, Mexico, United States)

lamarckii Forel, 1892 (India, Nepal)

lamborni Donisthorpe, 1933 (Malawi)

lameerei Emery, 1898 (Kyrgyzstan, Russian Federation, Turkmenistan)

• Camponotus fallax barbatus Vashkevich, 1926

• Camponotus fallax barbiger Donisthorpe, 1941

laminatus Mayr, 1866 (Fiji)

lancifer Emery, 1894 (Brazil)

landolti Forel, 1879 (Brazil, Colombia, Costa Rica, Mexico, Panama)

langi Wheeler, W.M., 1922 (Democratic Republic of Congo)

langi jejunus Santschi, 1926 (Democratic Republic of Congo)

laotsei Wheeler, W.M., 1921 (China)

largiceps Wu & Wang, 1989 (China)

lasiselene Wang, C. & Wu, 1994 (China, Thailand)

latangulus Roger, 1863 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)

latebrosus (Walker, 1859) (Sri Lanka)

lateralis (Olivier, 1792) (Albania, Algeria, Andorra, Armenia, Balearic Islands, Bulgaria, Croatia, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Malta, Monaco, Montenegro, Portugal, Republic of Macedonia, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Camponotus kosswigi Donisthorpe, 1950

• Camponotus lateralis armouri Wheeler, W.M., 1926

• Camponotus lateralis balearis Santschi, 1929

• Formica axillaris Spinola, 1808

• Formica bicolor Latreille Latreille, 1798

• Formica hemipsila Foerster, 1850

• Formica melanogastes Latreille, 1802

• Formica pallidinervis Brullé, 1833

lateralis cypridis Santschi, 1939 (Cyprus)

lateralis purius Santschi, 1929 (Algeria)

lateralis rhodius Santschi, 1934

lauensis Mann, 1921 (Fiji)

laurenti Santschi, 1939 (Algeria)

lautus (Say, 1836)

leae Wheeler, W.M., 1915 (Australia)

legionarium Santschi, 1911 (Madagascar)

lenkoi Kempf, 1960 (Brazil)

leonardi Emery, 1889 (Borneo, China, India, Myanmar, Philippines, Thailand)

• Colobopsis pubescens Mayr, 1862

• Formica pilosa Smith, F. Smith, F., 1857

leonardi gracilentus Viehmeyer, 1916 (Philippines, Singapore)

leonardi griseus Karavaiev, 1929 (Indonesia)

leptocephalus Emery, 1923 (Brazil)

lespesii Forel, 1886 (Brazil, Colombia, Mexico, Panama, Paraguay)

• Camponotus lespesi melancholicus Emery, 1894

leucophaeus (Smith, F., 1861) (Indonesia, Sulawesi)

leveillei Emery, 1895 (Madagascar)

levuanus Mann, 1921 (Fiji)

leydigi Forel, 1886 (Argentina, Brazil, French Guiana, Guyana, Paraguay, Suriname)

• Camponotus nobilis Santschi, 1916

libanicus André, 1881 (Greece, Iran, Israel, Lebanon, Turkey)

• Camponotus libanicus sahlbergi Forel, 1913

ligeus Donisthorpe, 1931 (Ethiopia)

lighti Wheeler, W.M., 1927 (China)

ligniperda (Latreille, 1802) (Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)

• Camponotus herculeanus herculeanoligniperdus Forel, 1874

• Formica obsoleta Christ Christ, 1791

ligniperda afer Stärcke, 1942 (Czech Republic)

ligniperda nigrescens Gösswald, 1932 (Germany)

lilianae Forel, 1913 (Democratic Republic of Congo, Kenya)

lilianae cornutus Forel, 1913 (Democratic Republic of Congo)

limbiventris Santschi, 1911 (Tanzania)

lindigi Mayr, 1870 (Belize, Colombia, Costa Rica, Greater Antilles, Honduras, Nicaragua, Panama, Trinidad and Tobago, Venezuela)

linnaei Forel, 1886 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Peru)

• Camponotus angulatus Mayr, 1870

• Camponotus angulicollis Emery, 1887

linnaei comoedus Wheeler, W.M., 1934 (Mexico)

linnaei maccus Wheeler, W.M., 1934 (French Guiana, Guyana)

linnaei muticus Forel, 1899 (Costa Rica)

liogaster Santschi, 1932 (South Africa)

lividicoxis Viehmeyer, 1925 (Australia)

loa Mann, 1919 (Solomon Islands)

loa belli Mann, 1919 (Solomon Islands)

longi Forel, 1902 (India)

longiceps (Smith, F., 1863) (China, Indonesia)

longideclivis McArthur, 1996 (Australia)

longifacies McArthur, 2003 (Australia)

longipalpis Santschi, 1926 (Democratic Republic of Congo)

longipilis Emery, 1911 (Bolivia, Peru)

loweryi McArthur, 1996 (Australia)

lownei Forel, 1895 (Australia)

• Formica nitida Lowne Lowne, 1865

lubbocki Forel, 1886 (Madagascar)

lubbocki christoides Forel, 1891 (Madagascar)

lubbocki rectus Forel, 1891 (Madagascar)

lucayanus Wheeler, W.M., 1905 (Bahamas)

luctuosus (Smith, F., 1858) (Indonesia)

luteiventris Emery, 1897 (New Guinea)

luteus (Smith, F., 1858) (India)

lutzi Forel, 1905 (Brazil)

M
maafui Mann, 1921 (Fiji)

macarangae Dumpert, 1996 (Borneo, Indonesia)

macareaveyi Taylor, 1992 (Australia)

• Camponotus sanguinea McAreavey, 1949

maccooki Forel, 1879 (Mexico, United States)

macilentus Smith, F., 1877 (Ecuador, Galapagos Islands)

macilentus albemarlensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

macilentus altinotus Stitz, 1932 (Galapagos Islands, Mexico)

macilentus barringtonensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus bindloensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus castellanus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus duncanensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus hoodensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus jacobensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus narboroensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus pervicus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus sapphirinus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus vulcanalis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus wollebaeki Stitz, 1932 (Galapagos Islands)

mackayensis Forel, 1902 (Australia)

macrocephalus (Erichson, 1842) (Australia)

• Camponotus fictor Forel, 1902

• Camponotus fictor augustulus Viehmeyer, 1925

• Colobopsis rufifrons semicarinata Forel, 1895

macrochaeta Emery, 1903 (Brazil, Ecuador)

macromischoides Fontenla Rizo, 1997 (Cuba)

maculatus (Fabricius, 1782) (Algeria, Angola, Benin, Cameroun, Central African Republic, Chad, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Iran, Ivory Coast, Kenya, Lesotho, Liberia, Madagascar, Mauritius, Mayotte, Micronesia (Federated States of), Mozambique, Namibia, Oman, Palau, Réunion, Senegal, Sierra Leone, South Africa, Sudan, São Tomé & Principe, Tanzania, Trinidad and Tobago, Uganda, Zimbabwe)

• Camponotus maculatus atramentarius Forel, 1904

• Camponotus maculatus ballioni Forel, 1904

• Camponotus maculatus boera Santschi, 1925

• Camponotus maculatus cavallus Santschi, 1911

• Camponotus maculatus cluisoides Forel, 1913

• Camponotus maculatus conakryensis Emery, 1920

• Camponotus maculatus erythraea Emery, 1920

• Camponotus maculatus flavifemur Santschi, 1937

• Camponotus maculatus flavominor Emery, 1925

• Camponotus maculatus hannae Santschi, 1919

• Camponotus maculatus hieroglyphicus Santschi, 1917

• Camponotus maculatus intonsus Emery, 1905

• Camponotus maculatus liocnemis Emery, 1905

• Camponotus maculatus lividior Santschi, 1911

• Camponotus maculatus lohieri Emery, 1915

• Camponotus maculatus manzer Forel, 1910

• Camponotus maculatus mathildae Forel, 1910

• Camponotus maculatus melanocnemis Santschi, 1911

• Camponotus maculatus miserabilis Santschi, 1914

• Camponotus maculatus nubis Weber, 1943

• Camponotus maculatus proletaria Baroni Urbani, 1971

• Camponotus maculatus radamoide Forel, 1891

• Camponotus maculatus sarmentus Emery, 1920

• Camponotus maculatus schereri Forel, 1911

• Camponotus maculatus schultzei Forel, 1912

• Camponotus maculatus semispicatus Emery, 1920

• Camponotus maculatus sudanicus Weber, 1943

• Camponotus maculatus thomensis Santschi, 1920

• Camponotus maculatus tuckeri Santschi, 1932

• Camponotus maculatus zumpti Santschi, 1937

• Camponotus sexpunctatus liengmei Forel, 1894

• Formica cognata Smith, F., 1858

• Formica lacteipennis Smith, F., 1858

maculatus foveolatus Stitz, 1925 (Philippines)

maculatus obfuscatus Viehmeyer, 1916 (Singapore)

maculatus subnudus Emery, 1889 (Myanmar, Philippines)

maculatus sylvaticomaculatus Dalla Torre, 1893 (Turkey)

maculatus ugandensis Santschi, 1923 (Uganda)

magister Santschi, 1925 (Morocco)

magister tibestiensis Santschi, 1932 (Libya)

maguassa Wheeler, W.M., 1922 (Democratic Republic of Congo)

malleensis McArthur, 2007 (Australia)

manni Wheeler, W.M., 1934 (Fiji)

marcens Forel, 1907 (Australia)

marianensis Clouse, 2007 (Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands)

maritimus Ward, 2005 (United States)

markli Dumpert, 2004 (Thailand)

maschwitzi Dumpert, 2006 (Borneo, Indonesia)

massiliensis Schmitz, 1950 (France)

massinissa Wheeler, W.M., 1922 (Democratic Republic of Congo)

mathildeae Smith, M.R., 1949 (Mexico)

maxwellensis Forel, 1913 (Malaysia)

maynei Forel, 1916 (Democratic Republic of Congo)

mayri Forel, 1879 (Kenya, Mozambique, South Africa, Zimbabwe)

mayri chimporensis Santschi, 1930 (Angola)

medeus Emery, 1920 (Peru)

medeus fulvulus Emery, 1920 (Peru)

megalonyx Wheeler, W.M., 1919 (Borneo, India)

• Camponotus chalaconotus Menozzi, 1929

melanocephalus Roger, 1863 (Venezuela)

melanoticus Emery, 1894 (Argentina, Bolivia, Brazil, Ecuador, Mexico, Paraguay)

melanoticus catharinae Santschi, 1939 (Brazil)

melanoticus flavopubens Emery, 1925 (Mexico)

melanoticus kempfi Özdikmen, 2010 (Brazil)

• Camponotus melanoticus nigrescens Santschi, 1939

melanoticus paranaensis Santschi, 1939 (Brazil)

melanoticus publicola Forel, 1912 (Brazil)

melanoticus valerius Santschi, 1922 (Brazil)

melanus Dumpert, 1995 (Borneo, Malaysia)

melichloros Kirby, W.F., 1888 (Christmas Island)

melinus Mackay, W.P., 1997 (Mexico)

mendax Forel, 1895 (India)

• Camponotus pseudolus Forel, 1907

mendax integer Forel, 1895 (Sri Lanka)

micans (Nylander, 1856) (Algeria, Iberian Peninsula, Monaco, Portugal, Spain)

micans asniensis Santschi, 1939 (Morocco)

michaelseni Forel, 1907 (Australia)

micragyne Dumpert, 1995 (Malaysia)

microps Snelling, R.R., 2006 (Mexico, United States)

micrositus Wheeler, W.M., 1937 (Cuba, Greater Antilles)

mina Forel, 1879 (Mexico, United States)

• Camponotus erythropus Pergande Pergande, 1893

• Camponotus mina zuni Wheeler, W.M., 1910

minimus Crawley, 1922 (Australia)

minozzii Emery, 1920 (Peru)

minus Wang, C. & Wu, 1994 (China)

mirabilis Emery, 1903 (Bolivia, Brazil, Peru)

mississippiensis Smith, M.R., 1923 (United States)

misturus (Smith, F., 1857) (Borneo)

• Formica exasperata Smith, F., 1857

misturus fornaronis Forel, 1892 (Bangladesh, Borneo, India)

mitis (Smith, F., 1858) (China, India, Sri Lanka)

mocquerysi Emery, 1899 (Madagascar)

mocsaryi Forel, 1902 (Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname)

moderatus Santschi, 1930 (Ethiopia)

modoc Wheeler, W.M., 1910 (Canada, Mexico, United States)

moelleri Forel, 1912 (Brazil)

moeschi Forel, 1910 (Borneo, Indonesia)

moeschi lygaeus Viehmeyer, 1916 (Singapore)

molossus Forel, 1907 (Australia)

monardi Santschi, 1930 (Angola)

monju Terayama, 1999 (Japan)

montivagus Forel, 1885 (Guatemala, Mexico)

• Camponotus nitidus nuperus Wheeler, W.M., 1914

morosus (Smith, F., 1858) (Argentina, Chile)

mozabensis Emery, 1899 (Algeria)

• Camponotus foreli impolitus Forel, 1902

mucronatus Emery, 1890 (Costa Rica, Guatemala, Honduras, Mexico)

mucronatus formaster Wheeler, W.M., 1934 (Guatemala, Mexico)

mucronatus hirsutinasus Wheeler, W.M., 1934 (Guatemala)

mucronatus santschii Forel, 1899 (Colombia)

mus Roger, 1863 (Argentina, Bolivia, Brazil, Ecuador, Mexico, Paraguay, Trinidad and Tobago, Uruguay)

• Camponotus ogloblini Santschi, 1934

mus mendozanus Santschi, 1922 (Argentina)

mus mustelus Emery, 1920 (Bolivia)

mussolinii Donisthorpe, 1936 (New Guinea)

mutilatus (Smith, F., 1859) (Indonesia)

mutilatus stitzi Özdikmen, 2010 (New Guinea)

• Camponotus mutilatus luteiventris Stitz, 1938

mystaceus Emery, 1886 (Botswana, Namibia, South Africa, Zimbabwe)

mystaceus exsanguis Forel, 1910 (Namibia)

mystaceus kamae Forel, 1910 (Namibia)

N
nacerdus Norton, 1868 (Mexico)

nadimi Tohmé, G., 1969 (Israel, Lebanon)

naegelii Forel, 1879 (Brazil)

namacola Prins, 1973 (Namibia)

nasicus Forel, 1891 (Madagascar)

nasutus Emery, 1895 (Namibia, South Africa, Zimbabwe)

nasutus fenestralis Santschi, 1937 (Namibia)

nasutus pretiosus Arnold, 1922 (Zimbabwe)

nasutus quadridentatus Santschi, 1937 (Namibia)

nasutus quinquedentatus Forel, 1910 (South Africa)

nasutus subnasutus Arnold, 1922 (South Africa)

natalensis (Smith, F., 1858) (Lesotho, South Africa, Zimbabwe)

natalensis corvus Forel, 1879 (South Africa)

natalensis diabolus Forel, 1879 (South Africa)

natalensis fulvipes Forel, 1914 (South Africa)

natalensis politiceps Santschi, 1914 (South Africa)

navigator Wilson & Taylor, 1967 (Samoa)

nawai Ito, 1914 (Japan, Republic of Korea)

nearcticus Emery, 1893 (Canada, United States)

• Camponotus fallax pardus Wheeler, W.M., 1917

• Camponotus fallax tanquaryi Wheeler, W.M., 1917

• Camponotus marginatus minutus Emery, 1893

nepos Forel, 1912 (Brazil)

newzealandicus Donisthorpe, 1940 (New Zealand)

nicobarensis Mayr, 1865 (Bangladesh, China, India, Nicobar Island, Thailand, Vietnam)

nicobarensis monticola Bingham, 1903 (Myanmar, Philippines)

nicobarensis rabbani Emery, 1925 (Myanmar)

nidulans (Smith, F., 1860) (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Suriname, Venezuela)

• Camponotus fabricii Roger, 1862

• Camponotus fabricii acoma Forel, 1908

• Dendromyrmex chartifex felis Wheeler, W.M., 1916

• Dendromyrmex chartifex mamoreensis Mann, 1916

• Dendromyrmex fabricii io Weber, 1943

• Dendromyrmex fabricii niger Weber, 1943

• Dendromyrmex fabricii pictus Wheeler, W.M., 1923

• Dendromyrmex fabricii rufescens Wheeler, W.M., 1916

• Dendromyrmex mamoreensis vestita Santschi, 1921

• Dendromyrmex nidulans nigripes Wheeler, W.M., 1916

nigricans Roger, 1863 (Philippines)

nigricans enganensis Forel, 1916 (Indonesia, Singapore)

• Camponotus nigricans nitidiceps Viehmeyer, 1916

nigricans pantiensis Santschi, 1932 (Indonesia)

nigriceps (Smith, F., 1858) (Australia)

• Camponotus consobrinus perthianus Wheeler, W.M., 1933

nigriceps lividipes Emery, 1887 (Australia)

nigrifrons (Mayr, 1870) (Tonga)

nigripes Dumpert, 1995 (Georgia)

nigroaeneus (Smith, F., 1858) (Australia)

• Camponotus nigroaeneus divus Forel, 1907

nigronitidus Azuma, 1951 (Japan)

nipponensis Santschi, 1937 (Democratic Peoples Republic of Korea, Japan, Republic of Korea)

nipponicus Wheeler, W.M., 1928 (China, Japan)

nirvanae Forel, 1893 (India)

nitens Mayr, 1870 (Brazil, Colombia)

nitens fuhrmanni Forel, 1921 (Colombia)

nitidescens Forel, 1889 (Greece)

nitidior (Santschi, 1921) (Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Dendromyrmex fabricii isthmicus Weber, 1943

niveosetosus Mayr, 1862 (Botswana, Kenya, Namibia, South Africa)

niveosetosus irredux Forel, 1910 (South Africa)

niveosetosus madagascarensis Forel, 1886 (Madagascar)

normatus Forel, 1899 (Costa Rica)

nosibeensis André, 1887 (Madagascar)

novaeboracensis (Fitch, 1855) (Canada, United States)

• Camponotus herculeanus rubens Emery, 1925

• Camponotus ligniperdus pictus Forel, 1886

novaehollandiae Mayr, 1870 (Australia, New Caledonia, Solomon Islands)

• Camponotus villosus Crawley, 1915

novogranadensis Mayr, 1870 (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States)

novogranadensis modestior Forel, 1904 (Costa Rica)

nutans Mayr, 1867 (Indonesia)

nutans cleliae Santschi, 1926 (Indonesia)

nylanderi Emery, 1921 (United Arab Emirates)

• Formica pallens Nylander, 1849

nywet Bolton, 1995 (Brazil)

• Formica nana Smith, F. Smith, F., 1858

O
oasium Forel, 1890 (Afghanistan, Algeria, Egypt, Iran, Iraq, Kazakhstan, Mali, Oman, Saudi Arabia, Turkey, Turkmenistan, United Arab Emirates)

• Camponotus bactrianus Pisarski, 1967

• Camponotus maculatus ruzskyellus Forel, 1922

• Camponotus thoracicus ninivae Pisarski, 1971

obliquus Smith, M.R., 1930 (United States)

oblongus (Smith, F., 1858) (Bangladesh, Myanmar)

oblongus binominatus Forel, 1916 (India)

• Camponotus oblongus opaca Forel, 1907

obreptivus Forel, 1899 (Costa Rica)

obscuripes Mayr, 1879 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

obscuriventris Cagniant, 1991 (Morocco)

obtritus Emery, 1911 (Peru)

occasus Emery, 1920 (Cape Verde)

oceanicus (Mayr, 1870) (Fiji)

ocreatus Emery, 1893 (Mexico, United States)

• Camponotus acutirostris primipilaris Wheeler, W.M., 1910

oculatior Santschi, 1935 (Democratic Republic of Congo)

odiosus Forel, 1886 (Indonesia)

oertzeni Forel, 1889 (Greece, Iran, Montenegro, Russian Federation, Turkey)

• Camponotus karawajewi Radchenko, 1996

oertzeni kappariensis Dalla Torre, 1893

oetkeri Forel, 1910 (Australia)

oetkeri voltai Forel, 1913 (Australia)

ogasawarensis Terayama & Satoh, 1990 (Japan)

olivieri Forel, 1886 (Angola)

olivieri concordius Santschi, 1915 (Angola)

olivieri delagoensis Forel, 1894 (Mozambique)

olivieri freyeri Santschi, 1915 (Madagascar, Seychelles)

olivieri infelix Santschi, 1915 (South Africa)

olivieri lemma Forel, 1886 (Angola)

olivieri moshianus Santschi, 1915 (Tanzania)

olivieri osiris Forel, 1911 (Democratic Republic of Congo)

olivieri patersoni Arnold, 1959 (Botswana)

olivieri pax Menozzi, 1942 (Cameroun)

olivieri sorptus Santschi, 1915 (Democratic Republic of Congo)

olivieri tenuipilis Santschi, 1915 (South Africa)

olivieri zaireicus Özdikmen, 2010 (Democratic Republic of Congo)

• Camponotus olivieri nitidior Santschi, 1926

ominosus Forel, 1911 (Sri Lanka)

opaciceps Roger, 1863 (Brazil, Grenada)

opaciventris Mayr, 1879 (India)

orinobates Santschi, 1919 (Kenya)

orinus Dumpert, 1995 (Malaysia)

• Camponotus montanus Dumpert, 1989

orites Santschi, 1919 (South Africa)

• Camponotus orinodromus Santschi, 1919

orthocephalus Emery, 1894 (Argentina, Bolivia, Brazil, Costa Rica, Ecuador, Peru)

orthodoxus Santschi, 1914 (Guinea, Ivory Coast)

ostiarius Forel, 1914 (South Africa)

ovaticeps (Spinola, 1851) (Chile)

overbecki Viehmeyer, 1916 (Singapore)

owensae Shattuck & McArthur, 2002 (Australia)

oxleyi Forel, 1902 (Australia)

P
pachylepis Emery, 1920 (Bolivia, Peru)

palkura McArthur, 2007 (Australia)

pallens (Le Guillou, 1842) (French Polynesia)

pallescens Mayr, 1887 (Brazil)

pallidiceps Emery, 1887 (Australia)

palmyrensis Tohmé, G. & Tohmé, H., 2000 (Syrian Arab Republic)

palpatus Emery, 1897 (New Guinea)

panamensis Fernández, 2002 (Panama)

papago Creighton, 1953 (Mexico, United States)

parabarbatus Bharti & Wachkoo, 2014 (India)

paradoxus (Mayr, 1866) (Brazil)

• Camponotus paradoxus janitor Forel, 1908

parius Emery, 1889 (Bangladesh, China, India, Myanmar)

patimae Wheeler, W.M., 1942 (Guyana)

patimae dolentulus Wheeler, W.M., 1942 (Guyana)

pavidus (Smith, F., 1860) (Indonesia)

pawseyi McArthur, 2003 (Australia)

peleliuensis Clouse, 2007 (Palau)

pellarius Wheeler, W.M., 1914 (Mexico)

pellax Santschi, 1919 (Australia)

pellitus Mayr, 1862 (Brazil, Paraguay)

• Camponotus pellitus scintillans Forel, 1901

pennsylvanicus (De Geer, 1773) (Bermuda, Canada, United States)

• Camponotus herculeanus herculeanopennsylvanicus Forel, 1879

• Camponotus herculeanus mahican Emery, 1925

• Formica semipunctata Kirby, W., 1837

peperi Forel, 1913 (Guatemala)

perjurus Shattuck & McArthur, 2002 (Australia)

perneser Bolton, 1995 (Indonesia, Sulawesi)

• Tapinoma nitida Smith, F., 1860

perrisii Forel, 1886 (Angola, Kenya)

perrisii densipunctatus Stitz, 1916 (Democratic Republic of Congo)

perrisii insularis Stitz, 1916 (Equatorial Guinea)

• Camponotus foraminosus annobonensis Santschi, 1920

perrisii jucundus Santschi, 1911 (Congo)

perrisii nigeriensis Santschi, 1914 (Nigeria)

perroti Forel, 1897 (Madagascar)

perroti aeschylus Forel, 1913 (Madagascar)

personatus Emery, 1894 (Argentina, Brazil, Paraguay)

• Camponotus personatus aloysii Emery, 1925

peseshus Bolton, 1995 (Australia)

• Camponotus nitidiceps Viehmeyer, 1925

petersii Emery, 1895 (Botswana, Lesotho, Mozambique, South Africa, Zimbabwe)

petersii janus Forel, 1911 (South Africa)

pexus Santschi, 1929 (Morocco)

philwardi McArthur, 2008 (Australia)

phragmaticola Donisthorpe, 1943 (India)

phytophilus Wheeler, W.M., 1934 (Mexico)

piceatus Norton, 1868 (Mexico)

piceus (Leach, 1825) (Albania, Algeria, Andorra, Armenia, Austria, Balearic Islands, Belgium, Bulgaria, China, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Lebanon, Malta, Montenegro, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine)

• Camponotus ebeninus Emery, 1869

• Camponotus lateralis ebneri Finzi, 1930

• Camponotus lateralis rectus Forel, 1892

• Formica foveolata Mayr, 1853

• Formica merula Losana, 1834

picipes (Olivier, 1792) (Brazil, Ecuador, French Guiana, Greater Antilles, Guyana, Haiti, Mexico, Suriname)

picipes guatemalensis Forel, 1885 (Guatemala)

picipes jamaicensis Wheeler, W.M., 1917 (Jamaica)

picipes jason Forel, 1912 (Bolivia)

picipes pilosulus Emery, 1925 (Mexico)

picipes plombyi Wheeler, W.M. & Mann, 1914 (Haiti)

picipes spengleri Forel, 1908 (Brazil)

pictipes Forel, 1891 (Madagascar)

pictostriatus Karavaiev, 1933 (Indonesia)

pilicornis (Roger, 1859) (France, Gibraltar, Iberian Peninsula, Portugal, Spain)

• Camponotus lichtensteini Bondroit, 1918

pilicornis siculus Grandi, 1935 (Italy, Republic of Macedonia)

piliventris (Smith, F., 1858) (Australia)

pitjantjatarae McArthur, 2003 (Australia)

pittieri Forel, 1899 (Brazil, Costa Rica, Lesser Antilles)

pittieri fuscogaster Emery, 1920 (Peru)

pittieri poenalis Wheeler, W.M., 1921 (Guyana)

placidus (Smith, F., 1858) (Indonesia)

planatus Roger, 1863 (Colombia, Costa Rica, Cuba, Ecuador, El Salvador, Guadeloupe, Guatemala, Honduras, Mexico, Nicaragua, Panama, United States, Venezuela)

planatus acaciae Emery, 1920 (Costa Rica)

planatus colombicus Forel, 1899 (Colombia)

planatus continentis Forel, 1901 (Mexico)

planatus esdras Forel, 1916 (Mexico)

planitae Santschi, 1929 (Algeria, Tunisia)

planus Smith, F., 1877 (Ecuador, Galapagos Islands)

planus fernandinensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus fidelis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus hephaestus Wheeler, W.M., 1933 (Ecuador, Galapagos Islands)

planus indefessus Wheeler, W.M., 1919 (Galapagos Islands)

planus isabelensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus peregrinus Emery, 1893 (Ecuador, Galapagos Islands)

planus pinzonensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus sansalvadorensis Wheeler, W.M., 1924 (Ecuador, Galapagos Islands)

planus santacruzensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

platypus Roger, 1863 (Philippines)

platytarsus Roger, 1863 (French Guiana, Guyana, Mexico, Suriname)

plutus Santschi, 1922 (French Guiana)

poecilus Emery, 1893 (Indonesia)

politae (Wu & Wang, 1994) (China)

polymorphicus Mackay, W.P., López-Castro & Fernández, 2002 (Colombia)

polynesicus Emery, 1896 (Fiji)

• Camponotus janus Mann, 1921

• Camponotus janussus Bolton, 1995

• Camponotus maudella Mann, 1921

• Camponotus maudella seemanni Mann, 1921

• Camponotus mayriella Mann, 1921

• Colobopsis carinata Mayr, 1870

pompeius Forel, 1886 (Gabon, Guinea, Kenya)

pompeius cassius Wheeler, W.M., 1922 (Democratic Republic of Congo)

pompeius iota Santschi, 1926 (Congo)

pompeius marius Emery, 1899 (Cameroun)

postangulatus Emery, 1911 (Peru)

postcornutus Clark, 1930 (Australia)

posteropilus Shattuck, 2005 (New Guinea)

posticus Santschi, 1926 (Democratic Republic of Congo, Kenya)

postoculatus Forel, 1914 (South Africa)

pressipes Emery, 1893 (Borneo, Philippines)

pressipes errans Forel, 1913 (Indonesia)

propinquellus Emery, 1920 (Peru)

propinquus Mayr, 1887 (Brazil)

• Camponotus paris Forel, 1912

propinquus baretoi Santschi, 1922 (Argentina)

prosseri Shattuck & McArthur, 2002 (Australia)

prostans Forel, 1910 (Australia)

prosulcatus Santschi, 1935 (Democratic Republic of Congo)

pseudoirritans Wu & Wang, 1989 (China)

pseudolendus Wu & Wang, 1989 (China)

puberulus Emery, 1897 (Eritrea, Somalia)

pudorosus Emery, 1925 (Mexico, United States)

pulchellus Forel, 1894 (New Caledonia)

pullatus Mayr, 1866 (Mexico)

pulvinatus Mayr, 1907 (Tanzania)

punctaticeps (Mayr, 1867) (Indonesia)

punctatissimus Forel, 1907 (Myanmar)

punctatus Forel, 1912 (Argentina, Brazil)

punctiventris Emery, 1920 (Australia)

punctulatus Mayr, 1868 (Argentina, Bolivia, Brazil, Mexico, Paraguay)

punctulatus andigenus Emery, 1903 (Peru)

punctulatus brevibarbis Santschi, 1939 (Bolivia)

punctulatus chubutensis Forel, 1913 (Argentina, Brazil)

punctulatus cruentus Santschi, 1922 (Argentina)

punctulatus hybridus Forel, 1901 (Argentina, Brazil)

punctulatus imberbis Emery, 1906 (Argentina)

punctulatus lillii Forel, 1908 (Brazil)

punctulatus lizeri Santschi, 1922 (Bolivia)

punctulatus minutior Forel, 1886 (Argentina)

punctulatus pergandei Emery, 1925 (Mexico)

• Camponotus punctulatus ruficornis Pergande, 1896

punctulatus tenuibarbis Santschi, 1939 (Argentina)

puniceps Donisthorpe, 1942 (India)

pupillus Santschi, 1939 (Algeria, Morocco)

putatus Forel, 1892 (Madagascar)

pylartes Wheeler, W.M., 1904 (United States)

pylartes fraxinicola Smith, M.R., 1923 (United States)

pylorus Santschi, 1920 (Vietnam)

Q
quadriceps (Smith, F., 1859) (Indonesia)

• Formica angulata Smith, F., 1859

• Formica nitida Smith, F. Smith, F., 1859

quadriceps convexior Viehmeyer, 1914 (Indonesia)

quadriceps curvatus Stitz, 1911 (New Guinea)

quadriceps nanulus Emery, 1925 (New Guinea)

• Camponotus quadriceps nana Emery, 1911

quadrimaculatus Forel, 1886 (Comoros, Mayotte)

quadrimaculatus opacatus Emery, 1925 (Madagascar)

• Camponotus quadrimaculatus opaca Emery, 1899

quadrimaculatus sellaris Emery, 1895 (Mayotte)

quadrinotatus Forel, 1886 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Camponotus marginatus nigricolor Ruzsky, 1926

• Camponotus marginatus rubicolor Ruzsky, 1925

quadrisectus (Smith, F., 1858) (Borneo, Philippines)

quadrisectus hians Forel, 1907 (Singapore)

quadrisectus margaritae Forel, 1907 (Singapore)

quercicola Smith, M.R., 1954 (Mexico, United States)

R
radiatus Forel, 1892 (India)

radovae Forel, 1886 (Madagascar)

radovae radovaedarwinii Forel, 1891 (Madagascar)

ramulorum Wheeler, W.M., 1905 (Bahamas, Cuba)

ramulorum marcidus Wheeler, W.M., 1905 (Bahamas)

ramulorum mestrei Wheeler, W.M., 1913 (Cuba, Greater Antilles)

ramulorum vernulus Wheeler, W.M., 1936 (Haiti)

rapax (Fabricius, 1804) (Brazil, Ecuador, French Guiana, Guyana, Suriname)

raphaelis Forel, 1899 (Costa Rica, Guatemala, Honduras, Mexico)

reaumuri Forel, 1892 (Madagascar)

rebeccae Forel, 1913 (Israel, Syrian Arab Republic)

reburrus Mackay, W.P., 2013 (Ecuador)

rectangularis Emery, 1890 (Belize, Brazil, Costa Rica, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Suriname)

rectangularis aulicus Wheeler, W.M., 1933 (Guatemala, Mexico)

rectangularis rubroniger Forel, 1899 (El Salvador, Mexico)

rectangularis setipes Forel, 1904 (Trinidad and Tobago)

rectangularis sordidatus Wheeler, W.M., 1934 (Peru)

rectangularis willowsi Wheeler, W.M., 1933 (Mexico)

rectithorax Forel, 1895 (Guatemala, Mexico, Nicaragua)

• Camponotus nitida Norton, 1868

reepeni Forel, 1913 (Indonesia)

reevei Arnold, 1922 (South Africa)

reichardti Arnoldi, 1967 (China, Kyrgyzstan, Uzbekistan)

reichenspergeri Santschi, 1926 (Indonesia)

reinaldi Kempf, 1960 (Brazil)

renggeri Emery, 1894 (Argentina, Bolivia, Brazil, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, Uruguay)

repens Forel, 1897 (Madagascar)

• Mayria madagascarensis Forel, 1886

reticulatus Roger, 1863 (Borneo, Micronesia (Federated States of), Northern Mariana Islands, Palau, Seychelles, Sri Lanka, Wallis and Futuna Islands)

reticulatus fullawayi Wheeler, W.M., 1912 (Guam)

reticulatus gestiens Forel, 1915 (Indonesia)

reticulatus imparilis Forel, 1915 (Indonesia)

reticulatus jagori Stitz, 1925 (Philippines)

reticulatus latitans Forel, 1893 (India)

reticulatus sericellus Viehmeyer, 1916 (Singapore)

reticulatus yerburyi Forel, 1893 (Sri Lanka)

• Camponotus bedoti motschulskii Emery, 1893

rhamses Santschi, 1915 (Egypt)

rhamses completus Santschi, 1930 (Ethiopia)

riedeli Pisarski, 1971 (Iraq)

riehlii (Roger, 1863) (Cuba, Greater Antilles, United States)

robecchii Emery, 1892 (Eritrea, Kenya, Namibia, Somalia, Yemen, Zimbabwe)

robecchii abyssinicus Santschi, 1913 (Ethiopia)

robecchii assumptionis Crawley, 1921 (Seychelles)

robecchii dispar Stitz, 1923 (Namibia)

robecchii rhodesianus Forel, 1913 (Zimbabwe)

robecchii troglodytes Forel, 1894 (Botswana, Kenya, Mozambique, Namibia)

robertae Santschi, 1926 (Ivory Coast)

robustus Roger, 1863 (Madagascar)

roeseli Forel, 1910 (Madagascar)

rosariensis Forel, 1912 (Argentina, Paraguay)

rothneyi Forel, 1893 (Bangladesh, India)

rothneyi krafti Forel, 1901 (Singapore)

rothneyi makilingi Viehmeyer, 1916 (Philippines)

rothneyi taivanae Forel, 1913 (Taiwan)

rotumanus Wilson & Taylor, 1967 (Fiji)

rotundinodis Santschi, 1935 (Democratic Republic of Congo)

rotundus Klimes & McArthur, 2014 (New Guinea)

roubaudi Santschi, 1911 (Congo)

royi Shattuck & Janda, 2009 (New Guinea)

ruber Emery, 1925 (Algeria, Balearic Islands, Portugal, Spain)

rubidus Xiao & Wang, 1989 (China)

rubiginosus Mayr, 1876 (Australia)

rubripes (Latreille, 1802) (Kenya, Sierra Leone)

rubrithorax Forel, 1899 (Mexico)

rubrithorax nigrithorax Stitz, 1937 (Mexico)

rudis McArthur, 2003 (Australia)

ruficornis Emery, 1895 (Chile)

rufifemur Emery, 1900 (Borneo, India, Indonesia, Thailand)

rufifemur obscurus Stitz, 1925 (Philippines)

rufifrons (Smith, F., 1860) (Fiji, Indonesia, New Guinea)

rufifrons leucopus Emery, 1914 (New Caledonia)

rufigaster Menozzi, 1928 (Indonesia)

rufipes (Fabricius, 1775) (Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Suriname, Uruguay, Venezuela)

• Camponotus cajurensis Luederwaldt, 1918

• Camponotus rufipes alpina Emery, 1906

• Camponotus rufipes magnifica Forel, 1913

• Camponotus rufipes subrufescens Santschi, 1929

• Formica herrichi Mayr, 1853

• Formica merdicola Lund, 1831

rufoglaucus (Jerdon, 1851) (Bangladesh, Borneo, Cambodia, Canary Islands, China, India, Kenya, Namibia, Nepal, Rwanda, Sri Lanka, Thailand, Vietnam, Zimbabwe)

• Camponotus redtenbacheri Mayr, 1862

rufoglaucus controversus Santschi, 1916 (Tanzania)

• Camponotus rufoglaucus flavopilosus Viehmeyer, 1913

rufoglaucus feae Emery, 1882 (Canary Islands)

• Camponotus feae erythropus Stitz, 1917

• Camponotus feae excelsus Stitz, 1917

• Formica pubescens Brullé Brullé, 1840

rufoglaucus latericius Stitz, 1923 (Namibia)

rufoglaucus syphax Wheeler, W.M., 1922 (Democratic Republic of Congo)

rufoglaucus tenuis Forel, 1907 (India)

rufoglaucus zanzibaricus Forel, 1911 (Tanzania)

rufoglaucus zulu Emery, 1895 (South Africa)

rufonigrus Shattuck & McArthur, 2002 (Australia)

rufus Crawley, 1925 (Australia)

rusticus Santschi, 1916 (Argentina)

S
sacchii Emery, 1899 (Somalia)

sachalinensis Forel, 1904 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation)

• Camponotus herculeanus altaica Ruzsky, 1915

• Camponotus herculeanus jacuticus Karavaiev, 1929

sadinus Mann, 1921 (Fiji)

salvini Forel, 1899 (Costa Rica, Mexico, Panama)

samius Forel, 1889 (Bulgaria, Greece, Turkey)

• Camponotus maculatus spagnolinii Emery, 1920

samueli McArthur, 2008 (Australia)

sanctaefidei Dalla Torre, 1892 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Panama, Paraguay, Peru)

• Camponotus quadrilaterus Mayr Mayr, 1887

sanctaefidei convexinodis Wheeler, W.M., 1934 (Brazil)

sanctaefidei coronatus Santschi, 1922 (Brazil)

sanctaefidei hondurianus Mann, 1922 (Honduras)

sanctaefidei leonhardi Forel, 1901 (Bolivia)

sanctaefidei weberi Wheeler, W.M., 1935 (Trinidad and Tobago)

• Camponotus sanctaefidei darlingtoni Wheeler, W.M., 1934

sanctus Forel, 1904 (Afghanistan, Greece, Iran, Israel, Turkey)

• Camponotus compressus cosensis Finzi, 1939

• Camponotus maculatus symiensis Forel, 1910

sanguinifrons Viehmeyer, 1925 (Australia, New Guinea)

sankisianus Forel, 1913 (Democratic Republic of Congo)

sannini Tohmé, G. & Tohmé, H., 1999 (Greece, Israel, Lebanon)

sansabeanus (Buckley, 1866) (Mexico, United States)

• Camponotus maculatus bulimosus Wheeler, W.M., 1910

• Camponotus maculatus torrefactus Emery, 1925

santosi Forel, 1908 (Bahamas, Cuba, Greater Antilles)

santosi pazosi Santschi, 1922 (Cuba)

satan Wheeler, W.M., 1919 (Borneo, Indonesia)

saundersi Emery, 1889 (Borneo, Myanmar, Thailand)

saundersi krama Forel, 1912 (Indonesia)

saussurei Forel, 1879 (Lesser Antilles)

saxatilis Ruzsky, 1895 (China, Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

sayi Emery, 1893 (Mexico, United States)

scabrinodis Arnold, 1924 (Zimbabwe)

scalaris Forel, 1901 (South Africa)

schaefferi Wheeler, W.M., 1909 (United States)

schmeltzi Mayr, 1866 (Fiji)

• Camponotus schmeltzi loloma Mann, 1921

schmitzi Stärcke, 1933 (Borneo, Brunei Darussalam)

schneei Mayr, 1903 (Marshall Islands)

schoedli Dumpert, 2006 (Thailand)

schoutedeni Forel, 1911 (Benin, Democratic Republic of Congo, Kenya, Zimbabwe)

scipio Forel, 1908 (Paraguay)

scipio insignis Santschi, 1922 (Paraguay)

scissus Mayr, 1887 (Brazil)

scotti McArthur, 2003 (Australia)

scratius Forel, 1907 (Australia)

scratius nuntius Forel, 1907 (Australia)

sculptor Santschi, 1920 (Argentina)

sedulus (Smith, F., 1857) (Borneo, Indonesia)

selene (Emery, 1889) (China, India, Myanmar)

selene obtusatus (Emery, 1895) (Myanmar)

sellidorsatus Prins, 1973 (South Africa)

semirufus Emery, 1925 (China, Kazakhstan, Kyrgyzstan, Turkmenistan)

semitestaceus Snelling, R.R., 1970 (Mexico, United States)

semoni Forel, 1905 (Venezuela)

senex (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

• Tapinoma tomentosa Norton, 1868

senkakuensis Terayama, 2013 (Japan)

septentrionalis (Buckley, 1866)

sericatus Mayr, 1887 (Brazil)

sericeiventris (Guérin-Méneville, 1838) (Argentina, Bolivia, Brazil, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Uruguay)

• Formica cuneata Perty, 1833

sericeiventris holmgreni Wheeler, W.M., 1931 (Bolivia)

sericeiventris otoquensis Wheeler, W.M., 1931 (Panama)

sericeiventris pontifex Santschi, 1936 (Mexico)

• Camponotus sericeiventris imperator Wheeler, W.M., 1934

sericeiventris rex Forel, 1907 (Costa Rica, Guatemala, Mexico)

sericeiventris satrapus Wheeler, W.M., 1931 (Brazil, Colombia, Costa Rica, Ecuador, Honduras, Panama, Peru)

sericeus (Fabricius, 1798) (Algeria, Benin, Burkina Faso, Egypt, Eritrea, Gambia, India, Israel, Ivory Coast, Kenya, Kenya, Kuwait, Mali, Mauritius, Oman, Pakistan, Saudi Arabia, Senegal, Sri Lanka, Thailand, United Arab Emirates, Zimbabwe)

• Formica aurulenta Latreille, 1802

• Formica obtusa Smith, F., 1858

• Formica pyrrhocephala Motschoulsky, 1863

sericeus euchrous Santschi, 1926 (Niger, Sudan)

sericeus peguensis Emery, 1895 (Myanmar)

sericeus sanguiniceps Donisthorpe, 1942 (Myanmar)

sericeus sulgeri Santschi, 1913 (Congo)

serotinus Cagniant, 1996 (Morocco)

sesquipedalis Roger, 1863 (Sri Lanka)

setitibia Forel, 1901 (Borneo, Indonesia)

setosus Shattuck & McArthur, 2002 (Australia)

seurati Emery, 1920 (Algeria)

severini Forel, 1909 (Borneo, Indonesia)

sexguttatus (Fabricius, 1793) (Argentina, Barbados, Bolivia, Brazil, Colombia, Costa Rica, Dominican Republic, Ecuador, French Guiana, Greater Antilles, Guyana, Haiti, Honduras, Lesser Antilles, Nicaragua, Panama, Paraguay, Peru, Puerto Rico, Saint Lucia, Suriname, United States, Uruguay)

• Camponotus sexguttatus fusciceps Emery, 1906

• Formica albofasciata Smith, F., 1862

• Formica bimaculata Smith, F., 1858

• Formica ruficeps Fabricius Fabricius, 1804

sexguttatus albotaeniolatus Forel, 1921 (Ecuador)

sexguttatus antiguanus Wheeler, W.M., 1923 (Antigua and Barbuda)

sexguttatus basirectus Wheeler, W.M., 1923 (Brazil)

sexguttatus biguttatus Emery, 1894 (Bolivia)

• Camponotus sexguttatus grenadensis Forel, 1897

sexguttatus decorus (Smith, F., 1858) (Brazil)

sexguttatus montserratensis Wheeler, W.M., 1923 (Lesser Antilles)

sexguttatus ornatus Emery, 1894 (Bolivia)

sexguttatus perturbans Kutter, 1931 (Colombia)

sexguttatus unitaeniatus Wheeler, W.M., 1923 (Peru)

sexpunctatus Forel, 1894 (Mozambique)

shaqualavensis Pisarski, 1971 (Iran, Iraq, Turkey)

• Camponotus arnoldi Pisarski, 1971

• Camponotus babau Bolton, 1995

shohki Terayama, 1999 (Japan)

sibreei Forel, 1891 (Madagascar)

sicheli Mayr, 1866 (Algeria, Balearic Islands, Canary Islands, Gibraltar, Iberian Peninsula, Malta)

sicheli niger Emery, 1925 (Algeria)

siemsseni Forel, 1901 (China, India, Indonesia)

sikorai Emery, 1920 (Madagascar)

silvestrii Emery, 1906 (Brazil)

silvicola Forel, 1902 (Argentina, Colombia, Ecuador, Paraguay)

simillimus (Smith, F., 1862) (Brazil, Colombia, Ecuador, French Guiana, Guyana, Panama, Suriname)

simillimus atratior Santschi, 1922 (Brazil)

simillimus impatibilis Forel, 1899 (Costa Rica)

simillimus indianus Forel, 1879 (Colombia, Panama)

simillimus riograndensis Emery, 1888 (Brazil)

simoni Emery, 1893 (Sri Lanka)

simpsoni McArthur, 2003 (Australia)

simulans Forel, 1910 (Namibia, Zimbabwe)

simulator Forel, 1915 (Australia)

simus Emery, 1908 (Democratic Republic of Congo)

simus manidis Forel, 1909 (Congo, Democratic Republic of Congo)

sinaiticus Ionescu-Hirsch, 2010 (Egypt, Israel)

singularis (Smith, F., 1858) (Borneo, China, Indonesia, Myanmar, Nepal, Thailand, Vietnam)

• Cataglyphis birmana Collingwood, 1962

singularis rufomaculatus Donisthorpe, 1941 (Borneo)

sklarus Bolton, 1995 (Myanmar)

• Camponotus similis Donisthorpe, 1933

smithianus Wheeler, W.M., 1919 (Borneo, Indonesia)

snellingi Bolton, 1995 (United States)

• Camponotus fallax pavidus Snelling, R.R., 1988

socius Roger, 1863 (Brazil, United States)

• Camponotus socius osceola Wheeler, W.M., 1932

socorroensis Wheeler, W.M., 1934 (Mexico)

socrates Forel, 1904 (India)

solenobius Menozzi, 1926 (Philippines)

solon Forel, 1886 (Angola, Benin, Guinea, Ivory Coast, Kenya, Senegal)

solon chiton Emery, 1925 (Senegal)

solon jugurtha Emery, 1925 (Democratic Republic of Congo)

somalinus André, 1887 (Eritrea, Kenya, Oman, Somalia)

somalinus curtior Forel, 1894 (Ethiopia)

somalinus pattensis Forel, 1907 (Kenya)

sommeri (Forel, 1894) (New Caledonia)

spanis Xiao & Wang, 1989 (China)

spenceri Clark, 1930 (Australia, Borneo, China)

• Camponotus reticulatus Kirby, W.F. Kirby, W.F., 1896

sphaericus Roger, 1863 (Cuba)

sphaericus cardini Mann, 1920 (Cuba)

sphaericus rufipilis Aguayo, 1932 (Cuba, Greater Antilles)

sphaericus sphaeralis Roger, 1863 (Cuba)

sphenocephalus Emery, 1911 (Peru)

sphenoidalis Mayr, 1870 (Colombia)

spinitarsus Emery, 1920 (Australia)

spinolae Roger, 1863 (Chile)

spissinodis Forel, 1909 (Algeria)

• Camponotus lateralis crassinodis Forel, 1894

sponsorum Forel, 1910 (Australia)

staryi Pisarski, 1971 (Iran, Iraq)

stefani McArthur, 2007 (Australia)

storeatus Forel, 1910 (Namibia, South Africa)

strangulatus Santschi, 1911 (Madagascar)

• Camponotus hova maculatoides Emery, 1920

striatipes Dumpert, 1995 (Borneo, Malaysia)

striatus (Smith, F., 1862) (Costa Rica, Honduras, Mexico, Panama)

• Camponotus alfaroi Emery, 1890

• Dolichoderus granulatus Pergande, 1896

strictus (Jerdon, 1851) (Borneo, India)

subbarbatus Emery, 1893 (United States)

subcircularis Emery, 1920 (Bolivia)

subnitidus Mayr, 1876 (Australia)

subnitidus famelicus Emery, 1887 (Australia)

subnitidus longinodis Forel, 1915 (Australia)

subpilus Shattuck, 2005 (Indonesia, New Guinea)

substitutus Emery, 1894 (Argentina, Brazil, Chile, Colombia, French Guiana, Guatemala, Guyana, Mexico, Panama, Paraguay, Suriname, Uruguay, Venezuela)

• Camponotus melanoticus multipilis Forel, 1907

substitutus clarus Stitz, 1938 (Argentina)

substitutus pullulus Emery, 1925 (Argentina)

subtilis (Smith, F., 1860) (Indonesia)

subtruncatus Borgmeier, 1929 (Brazil)

sucki Forel, 1901 (Borneo, Indonesia)

suffusus (Smith, F., 1858) (Australia)

• Camponotus schencki Mayr, 1862

suffusus bendigensis Forel, 1902 (Australia)

sylvaticus (Olivier, 1792) (Bulgaria, Canary Islands, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Luxembourg, Portugal, Spain, Turkey)

• Formica castanipes Leach, 1825

sylvaticus basalis Smith, F., 1878 (India)

• Camponotus maculatus lobinieri Forel, 1902

sylvaticus paradichrous Emery, 1925 (India)

T
tahatensis Santschi, 1929 (Algeria)

taino Snelling, R.R. & Torres, 1998 (Cuba, Greater Antilles, Puerto Rico)

taipingensis Forel, 1913 (Malaysia)

tameri Bolton, 1995 (Cameroun, New Guinea, Sudan)

• Camponotus tricolor Weber Weber, 1943

taniae Mackay, W.P. & Delsinne, 2009 (Paraguay)

tashcumiri Tarbinsky, 1976 (Kyrgyzstan, Mongolia)

tasmani Forel, 1902 (Australia)

tauricollis Forel, 1894 (Mozambique)

tejonia (Buckley, 1866)

tenuipes (Smith, F., 1857) (Borneo, Indonesia)

tenuiscapus Roger, 1863 (Argentina, Brazil, Uruguay)

tepicanus Pergande, 1896 (Mexico)

terbimaculatus Emery, 1920 (Brazil)

terebrans (Lowne, 1865) (Australia)

• Camponotus latrunculus victoriensis Santschi, 1928

• Camponotus myoporus Clark, 1938

• Formica testaceipes Smith, F. Smith, F., 1858

tergestinus Müller, 1921 (Hungary, Italy, Romania, Slovenia, Turkey)

termitarius Emery, 1902 (Argentina, Brazil, Paraguay)

terricola Karavaiev, 1929 (Indonesia)

territus Santschi, 1939 (Argentina)

testaceus Emery, 1894 (Brazil)

texanus Wheeler, W.M., 1903 (United States)

texens Dumpert, 1986 (Malaysia)

textor Forel, 1899 (Costa Rica, Honduras, Panama)

thadeus Shattuck, 2005 (Australia)

thales Forel, 1910 (Lesotho)

thomasseti Forel, 1912 (Seychelles)

thoracicus (Fabricius, 1804) (Algeria, Iran, Kuwait, Oman, Saudi Arabia, Turkey, United Arab Emirates, Yemen)

• Camponotus sylvaticus dichrous André, 1882

thraso Forel, 1893 (Sri Lanka)

thraso agricola Forel, 1910 (South Africa)

thraso assabensis Emery, 1925 (Ethiopia)

thraso diogenes Forel, 1909 (Sri Lanka)

thraso montinanus Santschi, 1926 (Zimbabwe)

thraso nefasitensis Menozzi, 1931 (Ethiopia)

thraso negus Forel, 1907 (Ethiopia, Kenya, Zimbabwe)

thysanopus Wheeler, W.M., 1937 (Cuba, Greater Antilles)

tilhoi Santschi, 1926 (Chad)

timidus (Jerdon, 1851) (India)

tonduzi Forel, 1899 (Costa Rica, Panama)

tonkinus Santschi, 1925 (China)

torrei Aguayo, 1932 (Cuba, Greater Antilles)

tortuganus Emery, 1895 (Mexico, United States)

toussainti Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

traegaordhi Santschi, 1914 (South Africa)

traegaordhi fumeus Santschi, 1926 (Zimbabwe)

traili Mayr, 1878 (Bolivia, Brazil, Colombia, Panama, Venezuela)

• Dendromyrmex graciliceps Viehmeyer, 1916

• Dendromyrmex madeirensis Mann, 1916

• Dendromyrmex madeirensis romani Wheeler, W.M., 1923

• Dendromyrmex traili rufogaster Mann, 1916

trajanus Forel, 1912 (Indonesia)

transvaalensis Arnold, 1948 (South Africa)

transvaalensis griqua Arnold, 1952 (South Africa)

transvaalensis prinsi Özdikmen, 2010 (Zimbabwe)

• Camponotus transvaalensis arnoldi Prins, 1965

trapeziceps Forel, 1908 (Brazil, Paraguay)

trapeziceps prosaicus Santschi, 1922 (Brazil)

trapezoideus Mayr, 1870 (Brazil, Colombia)

trepidulus Creighton, 1965 (Mexico, United States)

triangulatus Klimes & McArthur, 2014 (New Guinea)

tricolor (Stitz, 1912) (New Guinea)

tricoloratus Clark, 1941 (Australia)

trietericus Menozzi, 1926 (Philippines)

trifasciatus Santschi, 1926 (South Africa)

triodiae McArthur, 2009 (Australia)

tripartitus Mayr, 1887 (Brazil)

• Camponotus tripartitus collegianus Santschi, 1929

tristis Clark, 1930 (Australia)

triton Wheeler, W.M., 1934 (Bahamas)

tritschleri Forel, 1912 (Indonesia)

truebi Forel, 1910 (Taiwan)

truebi arnoldi Forel, 1912 (Taiwan)

truebi genaiai Santschi, 1928 (Taiwan)

truncatus (Spinola, 1808) (Albania, Algeria, Andorra, Armenia, Austria, Balearic Islands, Bulgaria, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Israel, Italy, Malta, Montenegro, Netherlands, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Formica fuscipes Mayr, 1853

tumidus Crawley, 1922 (Australia)

turkestanicus Emery, 1887 (Afghanistan, China, Georgia, Iran, Kazakhstan, Kyrgyzstan, Turkmenistan)

• Camponotus attila Stärcke, 1935

• Camponotus maculatus kaschgariensis Forel, 1901

turkestanus André, 1882 (Afghanistan, China, Israel, Kazakhstan, Kyrgyzstan, Mongolia, Russian Federation, Turkmenistan)

• Camponotus maculatus ferganensis Pisarski, 1967

• Camponotus maculatus flavonitidus Emery, 1925

• Camponotus maculatus ruzskyi Vashkevich, 1924

U
ulcerosus Wheeler, W.M., 1910 (Mexico, United States)

• Camponotus bruesi Wheeler, W.M., 1910

ulei Forel, 1904 (Peru)

ulvarum Forel, 1899 (Costa Rica)

umbratilis Wheeler, W.M., 1934 (Fiji)

universitatis Forel, 1890 (France, Germany, Iberian Peninsula, Italy, Spain, Switzerland)

urichi Forel, 1899 (Brazil, Trinidad and Tobago)

urichi folicola Forel, 1904 (Brazil)

urichi sculnus Forel, 1904 (Brazil)

ursus Forel, 1886 (Madagascar)

ustus Forel, 1879 (Colombia, Dominican Republic, Greater Antilles, Grenada, Haiti, Puerto Rico)

• Camponotus larvigerus Wheeler, W.M. & Mann, 1914

• Camponotus larvigerus maculifrons Menozzi & Russo, 1930

• Camponotus ustus depolitus Wheeler, W.M., 1936

• Camponotus ustus furnissi Wheeler, W.M. & Mann, 1914

• Camponotus ustus sublautus Wheeler, W.M. & Mann, 1914

• Camponotus ustus ulysses Forel, 1907

V
vafer Wheeler, W.M., 1910 (Mexico, United States)

vagulus Forel, 1908 (Brazil, Paraguay)

vagus (Scopoli, 1763) (Albania, Algeria, Andorra, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Turkey, Ukraine)

• Camponotus vagus kodorica Forel, 1913

• Formica fuscoptera Geoffroy, 1785

• Formica pubescens Fabricius Fabricius, 1775

vagus ifranensis Cagniant, 1987 (Morocco)

valdeziae Forel, 1879 (South Africa, Zimbabwe)

vanispinus Xia & Zheng, 1997 (China)

varians Roger, 1863 (India, Sri Lanka)

variegatus (Smith, F., 1858) (Bangladesh, China, Hawaii, Micronesia (Federated States of), Turkey)

• Camponotus variegatus hawaiensis Emery, 1920

variegatus ambonensis Karavaiev, 1930 (Indonesia)

variegatus bacchus (Smith, F., 1858) (Sri Lanka)

• Formica ventralis Smith, F., 1858

variegatus batta Menozzi, 1930 (Indonesia)

variegatus cleon Forel, 1913 (Indonesia)

variegatus comottoi Emery, 1887 (Myanmar)

variegatus crassinodis Forel, 1892 (Myanmar, Philippines)

variegatus dulcis Dalla Torre, 1893 (China, Myanmar, Philippines)

variegatus flavotestaceus Donisthorpe, 1948 (New Guinea)

variegatus fuscithorax Dalla Torre, 1893 (India)

variegatus infuscus Forel, 1892 (India)

variegatus intrans Forel, 1911 (Sri Lanka)

variegatus proles Emery, 1925 (Laos)

variegatus somnificus Forel, 1902 (India)

variegatus stenonotus Stitz, 1938 (Borneo)

varius Donisthorpe, 1943 (India)

varus Forel, 1910 (Angola)

velox (Jerdon, 1851) (India)

versicolor Clark, 1930 (Australia)

vespertinus Arnold, 1960 (Zimbabwe)

vestitus (Smith, F., 1858) (Benin, Burkina Faso, Guinea-Bissau, Ivory Coast, Mali, Namibia, Senegal, South Africa, Zimbabwe)

vestitus anthracinus Santschi, 1930 (Angola)

vestitus bombycinus Santschi, 1930 (Angola)

vestitus comptus Santschi, 1926 (Ivory Coast)

vestitus intuens Santschi, 1926 (Angola, Eritrea)

vestitus lujai Santschi, 1930 (Democratic Republic of Congo)

vestitus pectitus Santschi, 1930 (Angola)

vestitus perpectitus Santschi, 1926 (Benin)

vestitus strophiatus Santschi, 1926 (South Africa)

vicinus Mayr, 1870 (Canada, Mexico, United States)

victoriae Arnold, 1959 (South Africa)

viehmeyeri Forel, 1911 (New Guinea)

vigilans (Smith, F., 1858) (Borneo, China, Indonesia)

vinosus (Smith, F., 1858) (Brazil)

viri Santschi, 1915 (Kenya)

virulens (Smith, F., 1861) (Indonesia, Sulawesi)

vitiensis Mann, 1921 (Borneo, Fiji)

vitiosus Smith, F., 1874 (China, Japan, Republic of Korea)

• Camponotus itoi tokioensis Ito, 1912

• Camponotus tokioensis atrigenatus Santschi, 1937

• Camponotus tokioensis inconstans Santschi, 1937

vitreus (Smith, F., 1860) (Australia, Borneo, China, India, Indonesia, New Guinea, Nicobar Island, Philippines, Thailand)

• Camponotus angustata siggii Forel, 1893

• Formica incursor Smith, F., 1860

• Prenolepis adlerzii Forel, 1886

vitreus angustulus Emery, 1925 (Bangladesh, Singapore)

• Colobopsis angustata Mayr, 1870

vitreus carinatus Stitz, 1938 (New Guinea)

vitreus latinotus Stitz, 1925 (Borneo)

vitreus oebalis Forel, 1911 (Indonesia)

vitreus praeluteus Karavaiev, 1929 (Indonesia)

vitreus praerufus Emery, 1900 (Borneo, Indonesia)

vitreus vittatulus Forel, 1913 (Indonesia)

vittatus Forel, 1904 (Brazil)

vividus (Smith, F., 1858) (Gambia, Guinea, Ivory Coast, Kenya, Liberia, Mozambique, Senegal, Sierra Leone, Uganda, Zimbabwe)

• Formica laboriosa Smith, F., 1858

vividus cato Forel, 1913 (Democratic Republic of Congo, Uganda)

• Camponotus vividus laevithorax Menozzi, 1924

vividus meinerti Forel, 1886 (Angola, Chad)

vividus reginae Forel, 1901 (Angola)

vividus semidepilis Wheeler, W.M., 1922 (Democratic Republic of Congo)

voeltzkowii Forel, 1894 (Madagascar)

vogti Forel, 1906 (Greece, Iran, Israel, Turkey)

vulpus Santschi, 1926 (Democratic Republic of Congo)

W
walkeri Forel, 1893 (Australia)

walkeri bardus Forel, 1910 (Australia)

wanangus Klimes & McArthur, 2014 (New Guinea)

wasmanni Emery, 1893 (Bangladesh, China, India)

wasmanni mutilarius Emery, 1893 (Myanmar)

wedda Forel, 1908 (Sri Lanka)

weismanni Forel, 1901 (New Guinea)

weissflogi Dumpert, 2006 (Malaysia)

wellmani Forel, 1909 (Angola, Guinea)

wellmani gamma Santschi, 1926 (Democratic Republic of Congo)

wellmani rufipartis Forel, 1916 (Democratic Republic of Congo)

werthi Forel, 1908 (Kerguelen Islands)

werthi skaifei Arnold, 1959 (South Africa)

westermanni Mayr, 1862 (Argentina, Brazil, Paraguay)

westermanni fulvicornis Emery, 1903 (Brazil)

wheeleri Mann, 1916 (Brazil, Ecuador)

whitei Wheeler, W.M., 1915 (Australia)

• Camponotus scutellus Clark, 1930

wiederkehri Forel, 1894 (Australia)

• Camponotus denticulatus Kirby, W.F., 1896

• Camponotus latrunculus Wheeler, W.M., 1915

• Camponotus wiederkehri lucidior Forel, 1910

wildae Donisthorpe, 1948 (New Guinea)

woodroffeensis McArthur, 2008 (Australia)

wroughtonii Forel, 1893 (India, Nepal)

wytsmani Emery, 1920 (Bolivia, Ecuador)

X
xanthogaster Santschi, 1925 (Brazil)

xanthopilus Shattuck, 2005 (New Guinea)

xerxes Forel, 1904 (Iran, Iraq, Israel, Oman, Qatar, Saudi Arabia, Turkey, Turkmenistan, United Arab Emirates, Uzbekistan)

• Camponotus compressus irakensis Menozzi, 1927

xingdoushanensis Wang, W. & Chen, 2003 (China)

xuthus Emery, 1925 (Australia)

Y
yala Kusnezov, 1952 (Argentina)

yamaokai Terayama & Satoh, 1990 (Japan)

yambaru Terayama, 1999 (Japan)

yessensis Yasumatsu & Brown, 1951 (Japan)

yiningensis Wang, C. & Wu, 1994 (China)

yogi Wheeler, W.M., 1915 (Mexico, United States)

Z
zenon Forel, 1912 (Brazil)

zenon criton Emery, 1925 (Brazil)

zimmermanni Forel, 1894 (Ethiopia)

zimmermanni pansus Santschi, 1926 (Kenya)

zoc Forel, 1879 (Colombia, French Guiana, Guatemala, Guyana, Suriname, Trinidad and Tobago, Venezuela)

=Fossil Species=

A
ambon Zhang, 1989

ampullosus Zhang, 1989

B
brodiei Donisthorpe, 1920

C
cockerelli (Donisthorpe, 1920) (Late Eocene to Early Oligocene)

compactus Förster, 1891

crozei Riou, 1999

curviansatus Zhang, 1989

F
fuscipennis Carpenter, 1930

G
gracilis Zhang, 1989

H
heracleus (Heer, 1849)

I
induratus (Heer, 1849)

J
juliae Pierce & Gibron, 1962 Pierce & Gibron, 1962

L
lignitus (Germar, 1837)

longiventris Théobald, 1937

longus Zhang, 1989

M
mengei Mayr, 1868 (Baltic amber)

microcephalus Carpenter, 1930

microthoracus Zhang, 1989

N
novotnyi Samsinak, 1967

O
obesus Piton, 1935

oeningensis (Heer, 1849)

P
palaeopterus (Zhang, 1989)

penninervis Théobald, 1937

petrifactus Carpenter, 1930

pictus Zhang, Sun & Zhang, 1994

pinguiculus (Heer, 1849)

plenus Zhang, 1989

S
shanwangensis Hong, 1984

T
theobaldi Özdikmen, 2010

tokunagai Naora, 1933

U
ullrichi Bachmayer, 1960

V
vehemens Förster, 1891

vetus Scudder, 1877