Checklist of Pheidole species

The following species and subspecies belong to the genus Pheidole. Synonyms are listed under their senior names. For valid names only see Pheidole species and for distribution information see Pheidole species by Country.

A
aana Wilson & Taylor, 1967 (Samoa)

aberrans Mayr, 1868 (Argentina, Bolivia, Brazil, Paraguay, Peru)

• Pheidole aberrans diversiceps Santschi, 1916

• Pheidole aberrans fartilia Forel, 1913

• Pheidole aberrans mutica Emery, 1906

absurda Forel, 1886 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, United States)

• Pheidole ridicula Wheeler, W.M., 1916

acamata Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

acantha Eguchi, 2001 (Borneo, Indonesia, Malaysia)

accinota Wheeler, W.M., 1925 (Peru)

aciculata Wilson, 2003 (Brazil, Peru)

aculifera Wilson, 2003 (Panama)

acutidens (Santschi, 1922) (Argentina)

ademonia Wilson, 2003 (Ecuador, Guyana, Peru)

adrianoi Naves, 1985 (United States)

aeberlii Forel, 1894 (Senegal)

aeberlii erythraea Emery, 1901 (Eritrea, Ethiopia)

aenescens Wilson, 2003 (Brazil)

aequiseta Santschi, 1923 (Bolivia)

aglae Forel, 1913 (Borneo, Indonesia, Malaysia, New Guinea, Philippines, Thailand)

• Pheidole exasperata concordia Santschi, 1916

• Pheidole exasperata fusiformis Viehmeyer, 1914

• Pheidole exasperata polita Viehmeyer, 1914

agricola Wilson, 2003 (Mexico)

ajax Forel, 1899 (Colombia, Costa Rica, Nicaragua)

akermani Arnold, 1920 (South Africa)

alacris Santschi, 1923 (Argentina)

• Pheidole flavens mediorubra Santschi, 1933

alayoi Wilson, 2003 (Cuba, Greater Antilles)

albidula Santschi, 1928 (Democratic Republic of Congo)

albipes Wilson, 2003 (Guatemala, Honduras, Mexico)

alexeter Wilson, 2003 (Brazil)

alfaroi Emery, 1896 (Colombia, Costa Rica, Panama)

alienata Borgmeier, 1929 (Brazil)

allani Bingham, 1903 (China, India, Myanmar)

allarmata Wilson, 2003 (Brazil, Colombia, Ecuador, Peru, Suriname)

alpestris Wilson, 2003 (Ecuador)

alpinensis Forel, 1912 (Brazil)

alticola Wilson, 2003 (Colombia)

amabilis Wilson, 2003 (Costa Rica, Mexico)

amata Forel, 1901 (Colombia)

amazonica Wilson, 2003 (Ecuador, Peru)

amber Donisthorpe, 1941 (New Guinea)

ambigua Wilson, 2003 (Brazil)

ambonensis Karavaiev, 1930 (Indonesia)

ampla Forel, 1893 (Australia)

ampla mackayensis Forel, 1902 (Australia)

ampla parviceps Forel, 1915 (Australia)

ampla perthensis Crawley, 1922 (Australia)

amplificata Viehmeyer, 1914 (New Guinea)

anastasii Emery, 1896 (Belize, Costa Rica, Denmark, Guatemala, Honduras, Mexico, Nicaragua, Panama)

andrieui Santschi, 1930 (Mali, Sudan)

androsana Wheeler, W.M., 1905 (Bahamas, Dominican Republic)

angulicollis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

angulifera Wilson, 2003 (Colombia, Costa Rica)

angusta Forel, 1908 (Brazil)

• Pheidole angusta idulis Forel, 1912

angusticeps Wilson, 2003 (Costa Rica, El Salvador, Honduras)

• Pheidole gradifer Wilson, 2003

anima Wilson, 2003 (Honduras)

annemariae Forel, 1918 (Madagascar)

annexa Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

anthracina Forel, 1902 (Australia)

anthracina grandii Emery, 1914 (Australia)

anthracina orba Forel, 1902 (Australia)

antillana Forel, 1893 (Dominica, Lesser Antilles, St. Vincent)

antipodum (Smith, F., 1858) (Australia)

• Monomorium lippulum Wheeler, W.M., 1927

• Pheidole froggatti Forel, 1902

• Pheidole myops Forel, 1902

aper Forel, 1912 (Brazil)

aphrasta Zhou & Zheng, 1999 (China)

arachnion Wilson, 2003 (Costa Rica, Nicaragua)

• Pheidole iracunda Wilson, 2003

araneoides Wilson, 2003 (Brazil, Ecuador)

arboricola Wilson, 2003 (Costa Rica)

arcifera Santschi, 1925 (Brazil)

arctos Wilson, 2003 (Mexico)

areniphila Forel, 1910 (Botswana)

• Pheidole arenicola Forel, 1910

areniphila aurora Santschi, 1928 (United Republic of Tanzania)

argentina (Bruch, 1932) (Argentina)

arhuaca Forel, 1901 (Brazil, Colombia)

• Pheidole arhuaca bimons Forel, 1912

ariel Wilson, 2003 (Mexico)

aripoensis Wilson, 2003 (Trinidad and Tobago)

aristotelis Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

arnoldi Forel, 1913 (Zimbabwe)

arnoldi ballaensis Arnold, 1920 (Zimbabwe)

arnoldi rufescens Arnold, 1920 (Zimbabwe)

aspera Mayr, 1862 (South Africa)

asperata Emery, 1895 (India, Myanmar)

asperithorax Emery, 1894 (Bolivia, Brazil, Peru)

aspidata Eguchi & Bui, 2005 (Vietnam)

astur Wilson, 2003 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname)

athertonensis Forel, 1915 (Australia)

athertonensis cedarensis Forel, 1915 (Australia)

athertonensis tambourinensis Forel, 1915 (Australia)

atticola Forel, 1912 (Argentina)

atua Wilson & Taylor, 1967 (Samoa)

aurea Wilson, 2003 (United States)

aurivillii Mayr, 1896 (Cameroun, Guinea, Kenya, Uganda)

aurivillii attenuata Santschi, 1910 (Congo)

aurivillii kasaiensis Forel, 1911 (Democratic Republic of Congo)

aurivillii rubricalva Forel, 1915 (Democratic Republic of Congo)

auropilosa Mayr, 1887 (Argentina, Brazil, Paraguay)

avia Forel, 1908 (Brazil)

azteca Wilson, 2003 (Mexico)

B
bahai (Forel, 1922)

bajaensis Wilson, 2003 (Mexico)

bakeri Forel, 1912 (Cuba, Greater Antilles)

balcanica Seifert, 2016 (Bosnia and Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Turkey)

balzani Emery, 1894 (Bolivia, Venezuela)

bambusarum Forel, 1908 (Argentina, Brazil)

• Pheidole ogloblini Kusnezov, 1952

bandata Bharti, 2004 (India)

barbata Wheeler, W.M., 1908 (Mexico, United States)

barreleti Forel, 1903 (Sri Lanka)

barumtaun Donisthorpe, 1938 (New Guinea)

batrachorum Wheeler, W.M., 1922 (Central African Republic, Democratic Republic of Congo, Gabon)

beauforti Emery, 1911 (New Guinea)

bellatrix Wilson, 2003 (Colombia)

belli Mann, 1919 (Solomon Islands)

beloceps Wilson, 2003 (Costa Rica, Honduras, Mexico, Nicaragua)

bequaerti Forel, 1913 (Democratic Republic of Congo)

bergi Mayr, 1887 (Argentina, Brazil, Paraguay, Uruguay)

• Pheidole bergi pulliventris Santschi, 1929

• Pheidole bergi subparallela Emery, 1906

• Pheidole goetschi Santschi, 1939

bessonii Forel, 1891 (Madagascar)

bicarinata Mayr, 1870 (United States)

• Pheidole bicarinata vinelandica Forel, 1886

• Pheidole hayesi Smith, M.R., 1924

• Pheidole vinelandica buccalis Wheeler, W.M., 1908

• Pheidole vinelandica laeviuscula Emery, 1895

• Pheidole vinelandica longula Emery, 1895

• Pheidole vinelandica nebrascensis Forel, 1922

biconstricta Mayr, 1870 (Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Trinidad and Tobago, Venezuela)

• Pheidole biconstricta bicolor Emery, 1890

• Pheidole biconstricta burtoni Mann, 1916

• Pheidole biconstricta hybrida Emery, 1894

• Pheidole biconstricta rubicunda Emery, 1890

• Pheidole biconstricta surda Forel, 1912

• Pheidole holmgreni Wheeler, W.M., 1925

• Pheidole holmgreni festata Wheeler, W.M., 1925

• Pheidole radoszkowskii lallemandi Forel, 1901

bicornis Forel, 1899 (Costa Rica, Panama)

bidens Wilson, 2003 (Brazil)

bifurca Donisthorpe, 1941 (New Guinea)

bigote Longino, 2009 (Guatemala, Mexico)

bilimeki Mayr, 1870 (Bahamas, Belize, Colombia, Costa Rica, Cuba, Greater Antilles, Guatemala, Honduras, Mexico, Nicaragua, Panama, Venezuela)

• Pheidole anastasii cellarum Forel, 1908

• Pheidole anastasii johnsoni Wheeler, W.M., 1907

• Pheidole anastasii venezuelana Forel, 1905

• Pheidole floridana antoniensis Forel, 1901

• Pheidole floridana ares Forel, 1908

• Pheidole floridana deplanata Pergande, 1896

• Pheidole punctatissima annectens Wheeler, W.M., 1905

• Pheidole punctatissima insulana Wheeler, W.M., 1905

• Pheidole rectiluma Wilson, 2003

biloba (Karavaiev, 1935) (Cambodia)

binasifera Wilson, 2003 (Colombia)

binghamii Forel, 1902 (China, India, Vietnam)

biolleyi Forel, 1908 (Costa Rica, Mexico)

• Pheidole biolleyi tristani Forel, 1908

bison Wilson, 2003 (Bolivia)

blumenauensis Kempf, 1964 (Brazil)

• Pheidole diligens Santschi Santschi, 1923

bluntschlii Forel, 1911 (Indonesia, Thailand)

boliviana Wilson, 2003 (Bolivia, Colombia, Costa Rica, Panama)

• Pheidole mincana Wilson, 2003

• Pheidole scitula Wilson, 2003

boltoni Wilson, 2003 (Costa Rica)

• Pheidole humida Wilson, 2003

borgmeieri Kempf, 1972 (Brazil)

boruca Wilson, 2003 (Costa Rica, Honduras)

bos Forel, 1893 (Australia)

bos baucis Forel, 1910 (Australia)

bos eubos Forel, 1915 (Australia)

brachyops Wilson, 2003 (Costa Rica)

brandaoi Wilson, 2003 (Colombia)

branstetteri Longino, 2009 (Belize, Guatemala, Honduras, Mexico)

braueri Forel, 1897 (Seychelles)

brevicona Mayr, 1887 (Brazil)

• Pheidole kraepelini Forel, 1901

brevicornis Mayr, 1876 (Australia)

breviseta Santschi, 1919 (Argentina)

browni Wilson, 2003 (Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

bruchella Forel, 1915 (Argentina)

bruchi Forel, 1914 (Argentina, Paraguay)

bruesi Wheeler, W.M., 1911 (Grenada, Lesser Antilles)

brunnescens Santschi, 1929 (Brazil)

bucculenta Forel, 1908 (Brazil)

buchholzi Mayr, 1901 (Cameroun, Guinea)

buckleyi Smith, M.R., 1951

• Atta pennsylvanica Buckley, 1867

bufo Wilson, 2003 (Brazil, Colombia, Ecuador)

bula Sarnat, 2008 (Fiji)

bulliceps Wilson, 2003 (Ecuador)

bureni Wilson, 2003 (United States)

butteli Forel, 1913 (Borneo, Indonesia, Malaysia, Thailand)

C
caffra Emery, 1895 (South Africa, United Republic of Tanzania)

caffra abyssinica Forel, 1910 (Eritrea, Ethiopia)

caffra amoena Forel, 1911 (Eritrea, Ethiopia)

caffra bayeri Forel, 1916 (Democratic Republic of Congo)

caffra montivaga Santschi, 1939 (Zimbabwe)

caffra senilifrons Wheeler, W.M., 1922 (Democratic Republic of Congo)

cairnsiana Forel, 1902 (Australia)

caldwelli Mann, 1921 (Fiji)

calens Forel, 1901 (Brazil, Colombia, Costa Rica, Honduras, Mexico, Panama, Peru, Venezuela)

californica Mayr, 1870 (Mexico, United States)

• Pheidole californica hagermani Cole, 1936

• Pheidole californica incenata Wheeler, W.M., 1915

• Pheidole californica nevadensis Wheeler, W.M., 1915

• Pheidole californica pyramidensis Emery, 1922

• Pheidole californica satura Wheeler, W.M., 1915

• Pheidole californica shoshoni Cole, 1933

• Pheidole oregonica Emery, 1895

calimana Wilson, 2003 (Colombia)

caltrop Wilson, 2003 (Costa Rica, Honduras, Panama)

cameroni Mayr, 1887 (Ecuador)

• Pheidole monticola Cameron, 1891

camilla Wilson, 2003 (Ecuador)

camptostela Kempf, 1972 (Brazil)

capellinii Emery, 1887 (China, India, Indonesia, Thailand, Vietnam)

• Pheidole attila Forel, 1913

• Pheidole lighti Wheeler, W.M., 1927

capensis Mayr, 1862 (Mozambique, South Africa)

capensis dregei Emery, 1895 (South Africa)

capensis modestior Santschi, 1919 (South Africa)

capensis reddersbergensis Forel, 1913 (South Africa)

capillata Emery, 1906 (Brazil, Peru)

caracalla Wilson, 2003 (Ecuador, Peru)

carapuna Mann, 1916 (Brazil, Colombia, Costa Rica, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Venezuela)

• Pheidole carapuna chaquimayensis Wheeler, W.M., 1925

carapunco Kusnezov, 1952 (Argentina)

cardiella Wilson, 2003 (Ecuador, Peru)

cardinalis Wilson, 2003 (Brazil)

caribbaea Wheeler, W.M., 1911 (Dominican Republic, Jamaica)

• Pheidole caribbaea sloanei Wheeler, W.M., 1917

carinata Wilson, 2003 (Costa Rica)

cariniceps Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

carinote Longino, 2009 (Costa Rica)

carrolli Naves, 1985 (United States)

casta Wheeler, W.M., 1908 (United States)

castanea (Smith, F., 1858) (Brazil)

cataphracta Wilson, 2003 (Colombia)

cataractae Wheeler, W.M., 1916 (Brazil, Guyana, Peru)

caulicola Wilson, 2003 (Costa Rica)

cavifrons Emery, 1906 (Argentina, Brazil, Uruguay)

• Pheidole arciruga Forel, 1908

• Pheidole cavifrons fuscipunctis Santschi, 1916

cavigenis Wheeler, W.M., 1915 (Mexico, United States)

ceibana Wilson, 2003 (Honduras)

celaena Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

centeotl Wheeler, W.M., 1914 (Mexico)

cerebrosior Wheeler, W.M., 1915 (Mexico, United States)

ceres Wheeler, W.M., 1904 (Mexico, United States)

• Pheidole ceres tepaneca Wheeler, W.M., 1914

cerina Wilson, 2003 (Costa Rica, Panama)

cervicornis Emery, 1900 (New Guinea)

ceylonica (Motschoulsky, 1863) (Sri Lanka)

chalca Wheeler, W.M., 1914 (Mexico)

chalcoides Wilson, 2003 (Ecuador, Mexico)

charazana Wilson, 2003 (Colombia)

cheesmanae Donisthorpe, 1941 (Indonesia, New Guinea)

chilensis Mayr, 1862 (Chile, Peru)

chloe Forel, 1908 (Costa Rica, Mexico)

chocoensis Wilson, 2003 (Colombia, Costa Rica, Honduras, Nicaragua, Panama)

christinae Fischer, Hita Garcia & Peters, 2012 (Uganda)

christopherseni Forel, 1912 (Colombia, Costa Rica, Honduras, Mexico, Panama)

chrysops Wilson, 2003 (Brazil, Paraguay)

cicatricosa Stitz, 1917

• Pheidole pallidula recticeps Menozzi, 1932

cielana Wilson, 2003 (Mexico)

cingulata (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

citrina Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

clavata (Emery, 1877) (Eritrea, Ethiopia)

claviscapa Santschi, 1925 (Bolivia, Brazil)

clementensis Gregg, 1969 (Mexico, United States)

clydei Gregg, 1950 (Mexico, United States)

clypeocornis Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

cocciphaga Borgmeier, 1934 (Colombia, Costa Rica, Dominican Republic, Greater Antilles, Guyana, Nicaragua, Puerto Rico, Suriname, Venezuela)

cockerelli Wheeler, W.M., 1908 (Mexico, United States)

coffeicola Borgmeier, 1934 (Brazil, Suriname)

colaensis Mann, 1921 (Fiji)

colobopsis Mann, 1916 (Brazil, Colombia, Guatemala, Honduras, Mexico, Peru, Venezuela)

colpigaleata Eguchi, 2006 (Vietnam)

comata Smith, F., 1858 (Borneo, Indonesia, Malaysia, Thailand)

• Pheidole multicoma Eguchi, 1999

concentrica Forel, 1902 (Australia)

concentrica recurva Forel, 1910 (Australia)

concinna Santschi, 1910 (Congo)

conficta Forel, 1902 (Australia)

confoedusta Wheeler, W.M., 1909 (Mexico)

constanciae Forel, 1902 (China, India)

constanciae nigra Forel, 1902 (India)

constipata Wheeler, W.M., 1908 (United States)

coonoorensis Forel, 1902 (India)

coracina Wilson, 2003 (Mexico)

cordiceps Mayr, 1868 (Argentina)

• Pheidole cordiceps lilianae Forel, 1912

cornicula Wilson, 2003 (Bolivia, Paraguay)

corticicola Santschi, 1910 (Congo, Guinea, Ivory Coast)

coveri Wilson, 2003 (Trinidad and Tobago, Venezuela)

cramptoni Wheeler, W.M., 1916 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

• Pheidole cramptoni petiolicola Wheeler, W.M., 1921

crassicornis Emery, 1895 (Mexico, United States)

crassinoda Emery, 1895 (Botswana, Kenya, Namibia, South Africa, Uganda)

crassinoda pluto Arnold, 1920 (Mozambique)

crassinoda ruspolii Emery, 1897 (Ethiopia, Kenya, Zimbabwe)

crassinoda sordidula Santschi, 1937 (United Republic of Tanzania)

creightoni Gregg, 1955 (United States)

crinita Wilson, 2003 (Ecuador)

crozieri Wilson, 2003 (Peru)

cryptocera Emery, 1900 (New Guinea)

cubaensis Mayr, 1862 (Cuba, Greater Antilles)

• Pheidole cubaensis grayi Mann, 1920

cuevasi Wilson, 2003 (Brazil, Peru)

cuitensis Forel, 1910 (Angola)

cuprina Wilson, 2003 (Peru)

cursor Wilson, 2003 (Brazil, Ecuador, Guyana, Peru)

cyrtostela Wilson, 2003 (Brazil, Paraguay)

D
dammermani Wheeler, W.M., 1924 (Indonesia)

daphne Wilson, 2003 (Costa Rica)

darlingtoni Wheeler, W.M., 1936 (Greater Antilles, Haiti)

darwini Fischer, Hita Garcia & Peters, 2012 (Gabon)

dasypyx Wilson, 2003 (Costa Rica, Panama)

davidsonae Wilson, 2003 (Peru)

davisi Wheeler, W.M., 1905 (Mexico, United States)

dea Santschi, 1921 (Democratic Republic of Congo)

debilis Longino, 2009 (Costa Rica)

decarinata Santschi, 1929 (Mali, Sudan)

decepticon Fischer & Fisher, 2013 (Comoros, Mayotte, Seychelles)

deceptrix Forel, 1899 (Belize, Guatemala, Honduras, Mexico)

• Pheidole chiapasana Wilson, 2003

• Pheidole variceps Wilson, 2003

defecta Santschi, 1923 (Guatemala)

deima Wilson, 2003 (Brazil, Ecuador, Peru)

delecta Forel, 1899 (Mexico)

delicata Wilson, 2003 (Bolivia)

deltea Eguchi, 2001 (Borneo, Indonesia, Malaysia)

demeter Wilson, 2003 (Peru)

dentata Mayr, 1886 (Mexico, United States)

• Leptothorax tennesseensis Cole, 1938

• Pheidole commutata Mayr, 1886

• Pheidole dentata faisonsica Forel, 1901

dentigula Smith, M.R., 1927 (Mexico, United States)

descolei Kusnezov, 1952 (Argentina, Brazil)

deserticola Forel, 1910 (Australia)

deserticola foveifrons Viehmeyer, 1924 (Australia)

desertorum Wheeler, W.M., 1906 (Mexico, United States)

• Pheidole desertorum comanche Wheeler, W.M., 1906

• Pheidole desertorum maricopa Wheeler, W.M., 1906

diabolus Wilson, 2003 (Colombia, Costa Rica)

diana Forel, 1908 (Costa Rica)

diffidens (Walker, 1859) (Sri Lanka)

diffusa (Jerdon, 1851) (India)

diligens (Smith, F., 1858) (Brazil, Peru)

dinophila Wilson, 2003 (Brazil)

dione Forel, 1913 (Argentina)

dispar (Forel, 1895) (Australia)

distincta Donisthorpe, 1943 (New Guinea)

distorta Forel, 1899 (Colombia)

diversipilosa Wheeler, W.M., 1908 (Mexico, United States)

dodo Fischer & Fisher, 2013 (Mauritius)

dolon Wilson, 2003 (Bolivia, Peru)

dorsata Wilson, 2003 (Colombia, Guyana)

dossena Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

drepanon Wilson, 2003 (Dominican Republic, Greater Antilles)

dryas Wilson, 2003 (Costa Rica)

dugasi Forel, 1911 (China, Thailand, Vietnam)

dumicola Wilson, 2003 (Mexico)

duneraensis Bharti, 2001 (India)

durionei Santschi, 1923 (Argentina)

dwyeri Gregg, 1969 (Mexico)

dyctiota Kempf, 1972 (Brazil)

E
ectatommoides Wilson, 2003 (Colombia, Costa Rica, Ecuador, Panama)

ecuadorana Wilson, 2003 (Ecuador)

eidmanni Menozzi, 1926 (Brazil)

elecebra (Wheeler, W.M., 1904) (United States)

elegans Donisthorpe, 1938 (New Guinea)

elisae Emery, 1900 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

• Pheidole elisae nenia Forel, 1913

elongicephala Eguchi, 2008 (China, Vietnam)

embolopyx Brown, 1968 (Brazil, Ecuador, Peru)

emmae Forel, 1905 (Indonesia)

ensifera Forel, 1897 (Madagascar)

eowilsoni Longino, 2009 (Costa Rica, Honduras, Nicaragua)

eparmata Wilson, 2003 (Venezuela)

epetrion Wilson, 2003 (Ecuador)

epiphyta Longino, 2009 (Costa Rica)

erato Mann, 1919 (Solomon Islands)

erethizon Wilson, 2003 (Mexico)

eriophora Wilson, 2003 (Colombia)

ernsti Forel, 1912 (Taiwan)

erratilis Wilson, 2003 (Costa Rica, Nicaragua)

• Pheidole petersoni Wilson, 2003

escherichii Forel, 1910 (Eritrea, Ethiopia)

euryscopa Wilson, 2003 (Colombia)

exarata Emery, 1896 (Colombia, Costa Rica, Honduras, Nicaragua, Venezuela)

• Pheidole crassipes grantae Forel, 1908

exasperata (Mayr, 1866) (China)

excellens Mayr, 1862 (Ghana, Guinea)

excellens fulvobasalis Santschi, 1921 (Benin)

excellens weissi Santschi, 1910 (Congo)

excubitor Wilson, 2003 (Costa Rica, Mexico)

exigua Mayr, 1884 (Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Greater Antilles, Guyana, Mexico, Panama, Peru, Puerto Rico, Trinidad and Tobago, Venezuela)

exquisita Wilson, 2003 (Ecuador)

F
fabricator (Smith, F., 1858) (Brazil, Colombia, Venezuela)

• Atta nigriventris Smith, F., 1858

fadli Sharaf, 2007 (Egypt)

fallax Mayr, 1870 (Colombia, Costa Rica, Cuba, Guatemala, Guyana, Honduras, Jamaica, Nicaragua, Puerto Rico, Trinidad and Tobago, Venezuela)

• Pheidole fallax britoi Forel, 1912

• Pheidole fallax columbica Forel, 1886

• Pheidole fallax ovalis Forel, 1912

• Pheidole fallax rubens Forel, 1899

• Pheidole jelskii fallacior Forel, 1901

fantasia Chapman, 1963 (Borneo, Indonesia, Malaysia, Philippines)

fatigata Bolton, 1995 (New Guinea)

• Pheidole granulatus Donisthorpe, 1949

feae Emery, 1895 (China, India, Myanmar)

fera Santschi, 1925 (Brazil)

fergusoni Forel, 1902 (India)

fervens Smith, F., 1858 (Borneo, China, Cook Islands, Fiji, French Polynesia, Guam, India, Indonesia, Japan, Kiribati, Krakatau Islands, Malaysia, Marshall Islands, Mauritius, Micronesia (Federated States of), New Caledonia, Northern Mariana Islands, Palau, Philippines, Pitcairn, Samoa, Singapore, Taiwan, Thailand, Tokelau, Tonga, United States, Vanuatu, Vietnam)

• Pheidole amia Forel, 1912

• Pheidole cavannae Emery, 1887

• Pheidole javana Mayr, 1867

• Pheidole javana desucta Wheeler, W.M., 1929

• Pheidole javana dharmsalana Forel, 1902

• Pheidole javana dolenda Forel, 1912

• Pheidole javana soror Santschi, 1937

• Pheidole nodus azumai Santschi, 1941

• Pheidole oceanica nigriscapa Santschi, 1928

• Solenopsis pungens Smith, F., 1861

fervens pectinata Stitz, 1912 (Indonesia)

fervida Smith, F., 1874 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation, Vietnam)

fervida kwazana Teranishi, 1940 (Republic of Korea)

fimbriata Roger, 1863 (Argentina, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole diversa Smith, F., 1860

• Pheidole fimbriata tucumana Forel, 1913

• Pheidole smithii Dalla Torre, 1892

fiorii Emery, 1890 (Costa Rica, Nicaragua, Panama)

fissiceps Wilson, 2003 (Peru)

flavens Roger, 1863 (Argentina, Bahamas, Barbados, Belize, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, Galapagos Islands, Greater Antilles, Guatemala, Haiti, Honduras, Lesser Antilles, Madagascar, Mexico, Nicaragua, Panama, Paraguay, Peru, Trinidad and Tobago, United States)

• Pheidole exigua tuberculata Mayr, 1887

• Pheidole flavens gracilior Forel, 1901

• Pheidole flavens haytiana Forel, 1907

• Pheidole flavens spei Santschi, 1930

• Pheidole flavens vincentensis Forel, 1893

• Pheidole floridana aechmeae Wheeler, W.M., 1934

• Pheidole greggi Naves, 1985

flavens thomensis Emery, 1894

flaveria Zhou & Zheng, 1999 (China)

flavida Mayr, 1887 (Brazil)

flavifrons Wilson, 2003 (Brazil, Peru)

flavothoracica Viehmeyer, 1914 (New Guinea)

floricola Wilson, 2003 (Costa Rica, Ecuador, Honduras, Nicaragua)

floridana Emery, 1895 (Mexico, United States)

• Pheidole lauta Wheeler, W.M., 1908

foederalis Borgmeier, 1928

foreli Mayr, 1901 (South Africa)

foreli pubens Forel, 1910 (South Africa)

fortis Eguchi, 2006 (Thailand, Vietnam)

fossimandibula Longino, 2009 (Costa Rica)

foveolata Eguchi, 2006 (Vietnam)

fowleri Wilson, 2003 (Brazil)

fracticeps Wilson, 2003 (Brazil, Ecuador, Paraguay, Peru, Trinidad and Tobago)

fullerae Wilson, 2003 (Ecuador)

funki LaPolla, 2005 (Guyana)

funkikoensis Wheeler, W.M., 1929 (China, Taiwan)

furcata Sarnat, 2008 (Fiji)

furtiva Wilson, 2003 (United States)

fuscula Emery, 1900 (New Guinea)

G
gagates Wilson, 2003 (Ecuador, Peru)

gaigei Forel, 1914 (Colombia)

galba Wilson, 2003 (Peru)

gambogia Donisthorpe, 1948 (New Guinea)

gangamon Wilson, 2003 (Mexico)

gatesi (Wheeler, W.M., 1927) (China, Myanmar, Thailand, Vietnam)

gauthieri Forel, 1901 (Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama)

• Pheidole gauthieri oxymora Forel, 1912

gellibrandi Clark, 1934 (Australia)

geminata Wilson, 2003 (Ecuador)

gemmula Wilson, 2003 (Peru)

geraesensis Santschi, 1929 (Brazil)

germaini Emery, 1896 (Brazil)

gertrudae Forel, 1886 (Argentina, Bolivia, Brazil, Paraguay)

• Megalomyrmex humilis Borgmeier, 1930

• Pheidole gertrudae leonhardi Forel, 1901

• Pheidole gertrudae loretensis Santschi, 1933

ghatica Forel, 1902 (India)

ghigii Emery, 1900 (Borneo, Indonesia, Malaysia)

gibba Mayr, 1887 (Brazil)

• Pheidole gibba lopesi Borgmeier, 1937

gibbata Borgmeier, 1934 (Colombia, Suriname)

gigaflavens Wilson, 2003 (Argentina, Bolivia, Paraguay)

gigas Wilson, 2003 (Brazil)

gigliolii Menozzi, 1935

gilva Wilson, 2003 (Ecuador, Peru)

gilvescens Creighton & Gregg, 1955 (Mexico, United States)

glabrella Fischer, Hita Garcia & Peters, 2012 (Cameroun)

globularia Wilson, 2003 (Ecuador)

glomericeps Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

gnomus Wilson, 2003 (Ecuador)

godmani Forel, 1893 (Lesser Antilles, St. Vincent)

goeldii Forel, 1895 (Brazil)

gombakensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

gouldi Forel, 1886 (Costa Rica, Guatemala, Honduras, Mexico)

gracilescens (Smith, F., 1860) (Indonesia, Sulawesi)

gracilipes (Motschoulsky, 1863) (Sri Lanka)

grallatrix Emery, 1899 (Madagascar)

grandinodus Wilson, 2003 (Brazil)

granulata Pergande, 1896 (Mexico)

gravida Wilson, 2003 (Venezuela)

grayi Forel, 1902 (India)

grex Wilson, 2003 (Colombia)

grundmanni Smith, M.R., 1953 (United States)

guajirana Wilson, 2003 (Colombia)

guayasana Wilson, 2003 (Ecuador)

guerrerana Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

guilelmimuelleri Forel, 1886 (Argentina, Brazil, Colombia)

• Pheidole guilelmimuelleri gustavi Forel, 1911

guineensis (Fabricius, 1793) (Borneo, Guinea, Indonesia, Malaysia)

gulo Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

gymnoceras Longino, 2009 (Costa Rica, Nicaragua)

H
hainanensis Chen, Ye, Lu & Zhou, 2011 (China)

hamtoni Wilson, 2003 (Mexico)

harlequina Wilson, 2003 (Dominican Republic, Greater Antilles)

harrisonfordi Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Pheidole prolixa Wilson, 2003

• Pheidole ruida Wilson, 2003

• Pheidole tenebra Wilson, 2003

hartmeyeri Forel, 1907 (Australia)

haskinsorum Wilson, 2003 (Bolivia, Ecuador)

hasticeps Wilson, 2003 (Costa Rica, Mexico, Panama)

havilandi Forel, 1911 (Borneo, Indonesia, Malaysia)

• Pheidole havilandi sapuana Forel, 1911

• Pheidole havilandi selangorensis Forel, 1913

haywardi Kusnezov, 1952 (Argentina, Paraguay)

hazenae Wilson, 2003 (Colombia, Ecuador)

hecate Wheeler, W.M., 1911 (Jamaica)

• Pheidole hecate bruesi Wheeler, W.M., 1917

• Pheidole hecate malevola Wheeler, W.M., 1911

hector Wilson, 2003 (Costa Rica)

hedlundorum Wilson, 2003 (Venezuela)

heliosa Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

hercules Donisthorpe, 1941 (New Guinea)

heterothrix Santschi, 1923 (Argentina)

• Pheidole gavrilovi Kusnezov, 1952

hetschkoi Emery, 1896 (Brazil)

• Pheidole hetschkoi frivola Santschi, 1923

hewitti Santschi, 1932 (South Africa)

• Pheidole capensis hewitti Arnold, 1944

heyeri Forel, 1899 (Brazil)

• Pheidole guilelmimuelleri ultrix Forel, 1912

hierax Wilson, 2003 (Panama)

hirsuta Emery, 1896 (Costa Rica, Guatemala, Panama)

hirtula Forel, 1899 (Mexico)

• Pheidole vasliti acolhua Wheeler, W.M., 1914

hispaniolae Wilson, 2003 (Dominican Republic, Greater Antilles)

hizemops Wilson, 2003 (Costa Rica)

hoelldobleri Wilson, 2003 (Costa Rica)

hongkongensis Wheeler, W.M., 1928 (China, Thailand, Vietnam)

hoplitica Wilson, 2003 (Mexico, United States)

horni Emery, 1901 (Sri Lanka)

horribilis Wilson, 2003 (Ecuador, Peru)

hortensis Forel, 1913 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

hortonae Wilson, 2003 (Venezuela)

hospes Smith, F., 1865 (New Guinea)

hospita Bingham, 1903 (India)

huacana Wilson, 2003 (Ecuador, Peru)

huberi Forel, 1911 (Borneo, Indonesia, Malaysia)

• Pheidole huberi perakensis Forel, 1911

huilana Wilson, 2003 (Colombia)

humeralis Wheeler, W.M., 1908 (United States)

humeridens Wilson, 2003 (Argentina)

hyatti Emery, 1895 (Mexico, United States)

• Pheidole hyatti ecitonodora Wheeler, W.M., 1908

• Pheidole hyatti solitanea Wheeler, W.M., 1915

• Pheidole vaslitii Pergande, 1896

I
iceni Fernández, 2011 (Colombia)

impressa Mayr, 1870 (Brazil)

impressa detrita Emery, 1890 (Venezuela)

impressiceps Mayr, 1876 (Australia, New Guinea)

impressiceps commista Forel, 1901 (New Guinea)

inca Wilson, 2003 (Peru)

incerta (Smith, F., 1863) (Indonesia)

incisa Mayr, 1870 (Colombia)

incurvata Viehmeyer, 1924 (Australia)

indagatrix Wilson, 2003 (Costa Rica, Honduras, Nicaragua, Panama)

indica Mayr, 1879 (Antigua and Barbuda, Balearic Islands, Bangladesh, Barbados, Canary Islands, China, Comoros, Cuba, Dominican Republic, Egypt, Eritrea, Greater Antilles, Greece, Iberian Peninsula, India, Iran, Israel, Japan, Kenya, Kuwait, Madagascar, Malta, Mauritius, Mayotte, Nepal, Oman, Peru, Republic of Korea, Saint Helena, Saudi Arabia, Seychelles, Spain, United Arab Emirates, United States)

• Pheidole indica himalayana Forel, 1902

• Pheidole indica rotschana Forel, 1902

• Pheidole striativentris Mayr, 1879

• Pheidole teneriffana Forel, 1893

• Pheidole teneriffana taina Aguayo, 1932

indosinensis Wheeler, W.M., 1928 (China, Vietnam)

industa Santschi, 1939 (Uruguay)

inermis Mayr, 1870 (Guatemala, Honduras, Mexico, Nicaragua)

infernalis Wilson, 2003 (Bolivia)

inflexa Santschi, 1923

innotata Mayr, 1866

innupta Menozzi, 1931 (Costa Rica, Peru)

inornata Eguchi, 2001 (Borneo, Indonesia, Malaysia, Thailand)

inquilina (Wheeler, W.M., 1903) (United States)

inscrobiculata Viehmeyer, 1916 (Malaysia, Philippines, Thailand)

insipida Forel, 1899 (Mexico)

inversa Forel, 1901 (Colombia)

irritans (Smith, F., 1858) (South Africa)

isis Mann, 1919 (Solomon Islands)

isis taki Mann, 1919 (Solomon Islands)

J
jacobsoni Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

• Pheidole orophila Eguchi, 2001

jaculifera Wilson, 2003 (Brazil)

jamaicensis Wheeler, W.M., 1908 (Cuba, Dominican Republic, Haiti, Jamaica)

janzeni Longino, 2009 (Costa Rica, El Salvador, Guatemala, Honduras, Mexico)

jeannei Wilson, 2003 (Brazil)

jelskii Mayr, 1884 (Argentina, Barbados, Brazil, Dominican Republic, French Guiana, Greater Antilles, Grenada, Haiti, Paraguay, Saint Lucia, Trinidad and Tobago)

• Pheidole fallax emiliae Forel, 1901

• Pheidole jelskii antillensis Forel, 1901

• Pheidole jelskii arenicola Emery, 1894

jivaro Wilson, 2003 (Ecuador, Panama)

jonas Forel, 1907 (Comoros, Madagascar, Mayotte)

jordanica Saulcy, 1874 (Egypt, Israel)

• Pheidole megacephala obtusa Stitz, 1917

• Pheidole schmitzi Forel, 1911

• Pheidole sinaitica laticeps Mayr, 1904

jubilans Forel, 1911 (Indonesia)

jucunda Forel, 1885 (China, India)

jucunda fossulata Forel, 1902 (China, India)

jujuyensis Forel, 1913 (Argentina)

juniperae Wilson, 2003 (United States)

K
karolmorae Longino, 2009 (Costa Rica, Honduras, Nicaragua)

karolsetosa Longino, 2009 (Costa Rica)

katonae Forel, 1907 (United Republic of Tanzania)

kava Fischer, Sarnat & Economo, 2016 (Fiji)

kikutai Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

kitschneri Forel, 1910 (Kenya, South Africa)

knowlesi Mann, 1921 (Fiji)

• Pheidole knowlesi extensa Mann, 1921

kochi (Emery, 1911) (New Guinea)

kohli Mayr, 1901 (Kenya, South Africa)

komori Fischer & Fisher, 2013 (Comoros, Mozambique)

koshewnikovi Ruzsky, 1905 (Kyrgyzstan)

• Pheidole pallidula arenarum Ruzsky, 1905

• Pheidole pallidula orientalis Müller, 1923

kugleri Wilson, 2003 (Peru)

kukrana Wilson, 2003 (Costa Rica, Guatemala, Nicaragua, Venezuela)

kuna Wilson, 2003 (Panama)

kusnezovi Wilson, 2003 (Argentina)

• Eriopheidole symbiotica Kusnezov, 1952

L
laelaps Wilson, 2003 (Colombia, Ecuador)

laevicolor Eguchi, 2006 (Vietnam)

laevifrons Mayr, 1887 (Brazil)

laevinota Forel, 1908 (Argentina, Brazil)

laevithorax Eguchi, 2008 (Vietnam)

laeviventris Mayr, 1870 (Colombia)

laevivertex Forel, 1901 (Mexico)

lagunculinoda Longino, 2009 (Guatemala, Mexico)

laidlowi Mann, 1916 (Brazil, Ecuador, Peru)

lamellinoda Forel, 1902 (India)

lamia Wheeler, W.M., 1901 (United States)

laminata Emery, 1900 (New Guinea)

lancifera Wilson, 2003 (Brazil, Peru)

lanigera Wilson, 2003 (Peru, Venezuela)

lanuginosa Wilson, 1984 (India)

laselva Wilson, 2003 (Colombia, Costa Rica, Ecuador, Honduras, Nicaragua, Panama, Venezuela)

• Pheidole ebenina Wilson, 2003

laticornis Wilson, 2003 (Costa Rica, Venezuela)

laticrista Santschi, 1916 (Argentina)

latinoda Roger, 1863 (Sri Lanka)

latinoda angustior Forel, 1902 (India, Sri Lanka)

latinoda major Forel, 1885 (India)

latinoda peradeniyae Forel, 1911 (Sri Lanka)

lattkei Wilson, 2003 (Venezuela)

laudatana Wilson, 2003 (Dominica)

leloi Eguchi & Bui, 2016 (Vietnam)

lemnisca Wilson, 2003 (Colombia, Ecuador)

lemur Forel, 1912 (Brazil)

leoncortesi Longino, 2009 (Mexico)

leonina Wilson, 2003 (Brazil, Peru)

leptina Wilson, 2003 (Brazil, Peru)

liengmei Forel, 1894 (Mozambique)

liengmei malindana Forel, 1913 (Zimbabwe)

liengmei micrartifex Forel, 1913 (Zimbabwe)

liengmei shinsendensis Forel, 1913 (Democratic Republic of Congo)

lignicola Mayr, 1887 (Argentina, Brazil, Paraguay)

• Pheidole lignicola levocciput Forel, 1908

lilloi (Kusnezov, 1952) (Argentina)

liteae Forel, 1910 (Australia)

littoralis Cole, 1952 (United States)

lobulata Emery, 1900 (New Guinea)

loki Fischer & Fisher, 2013 (Mayotte)

lokitae Forel, 1913 (Borneo, Indonesia, Malaysia, Philippines)

longiceps Mayr, 1876 (Australia)

longiceps doddi Forel, 1910 (Australia)

longiceps frontalis Forel, 1902 (Australia)

longicornis Emery, 1888 (Indonesia)

longinoi Wilson, 2003 (Costa Rica)

longior Santschi, 1933 (Argentina)

longipes (Latreille, 1802) (Borneo, French Guiana, India, Indonesia, Malaysia, Nicobar Island, Thailand)

longiscapa Forel, 1901 (Brazil, Venezuela)

• Pheidole longiscapa martensis Forel, 1914

longiseta Wilson, 2003 (Brazil, Peru)

longispinosa Forel, 1891 (Madagascar)

longispinosa scabrata Forel, 1895 (Madagascar)

lourothi Wilson, 2003 (Costa Rica)

lovejoyi Wilson, 2003 (Brazil)

lucaris Wilson, 2003 (Brazil, Costa Rica)

lucida Forel, 1895 (Madagascar)

lucioccipitalis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

lucretii Santschi, 1923 (Brazil)

lupus Wilson, 2003 (Ecuador, Peru)

lustrata Wilson, 2003 (Costa Rica)

luteipes Emery, 1914 (New Caledonia)

luteipes obvia Emery, 1914 (New Caledonia)

lutzi Forel, 1905 (Brazil)

• Pheidole lutzi heinzi Forel, 1913

M
macclendoni Wheeler, W.M., 1908 (United States)

machetula Wilson, 2003 (Peru)

mackayi Wilson, 2003 (Mexico)

macracantha Wilson, 2003 (Panama)

macromischoides Wilson, 2003 (Cuba)

macrops Wilson, 2003 (Mexico)

maculifrons Wheeler, W.M., 1929 (Philippines)

madecassa Forel, 1892 (Madagascar)

madrensis Wilson, 2003 (Mexico)

magna Eguchi, 2006 (Vietnam)

magrettii Emery, 1887 (Indonesia, Malaysia, Thailand)

• Pheidole nodgii maxwellensis Forel, 1913

maja Forel, 1886 (Guatemala)

makilingi Viehmeyer, 1916 (Philippines)

malabarica (Jerdon, 1851) (India)

malinsii Forel, 1902 (China, India, Sri Lanka)

mallota Wilson, 2003 (Panama)

mamore Mann, 1916 (Brazil, Guyana, Peru, Suriname)

manteroi Emery, 1897 (New Guinea)

mantilla Wilson, 2003 (Costa Rica, Panama)

manuana Wilson, 2003 (Peru)

manukana Eguchi, 2001 (Borneo, Indonesia, Malaysia)

marcidula Wheeler, W.M., 1908 (Mexico, United States)

maufei Arnold, 1920 (Zimbabwe)

mayri Forel, 1894 (Senegal)

medioflava Donisthorpe, 1941 (New Guinea)

megacephala (Fabricius, 1793) (Aruba, Australia, Bahamas, Barbados, Bermuda, Borneo, Brazil, Canary Islands, Cape Verde, China, Colombia, Cook Islands, Costa Rica, Cuba, Dominican Republic, Ecuador, Egypt, Fiji, French Guiana, Greater Antilles, Greece, Guam, Guatemala, Guinea, Guyana, Haiti, Hawaii, Honduras, India, Indonesia, Iran, Italy, Ivory Coast, Japan, Kenya, Kiribati, Lesser Antilles, Madagascar, Malaysia, Mali, Marshall Islands, Mauritius, Mayotte, Mexico, Micronesia (Federated States of), Montenegro, Mozambique, Netherlands Antilles, New Caledonia, New Guinea, New Zealand, Nicaragua, Nicobar Island, Niue, Northern Mariana Islands, Oman, Palau, Panama, Peru, Philippines, Puerto Rico, Republic of Macedonia, Romania, Rwanda, Réunion, Saint Lucia, Saint Martin (French part), Samoa, Senegal, Seychelles, Sierra Leone, Solomon Islands, Suriname, Thailand, Tokelau, Tonga, Trinidad and Tobago, Uganda, United Arab Emirates, United Kingdom of Great Britain and Northern Ireland, United States, Vanuatu, Venezuela, Vietnam, Wallis and Futuna Islands, Zimbabwe)

• Atta testacea Smith, F., 1858

• Formica edax Forskål, 1775

• Myrmica agilis Smith, F., 1857

• Myrmica laevigata Smith, F., 1855

• Myrmica suspiciosa Smith, F., 1859

• Myrmica trinodis Losana, 1834

• Oecophthora perniciosa Gerstäcker, 1859

• Oecophthora pusilla Heer, 1852

• Pheidole janus Smith, F., 1858

• Pheidole laevigata Mayr, 1862

• Pheidole megacephala scabrior Forel, 1891

• Pheidole picata Forel, 1891

• Pheidole picata bernhardae Emery, 1915

• Pheidole picata gietleni Forel, 1905

megacephala costauriensis Santschi, 1914 (Ghana, Guinea)

megacephala duplex Santschi, 1937 (Angola)

megacephala ilgi Forel, 1907 (Ethiopia)

megacephala impressifrons Wasmann, 1905 (Eritrea, South Africa)

• Pheidole megacephala impressiceps Wasmann, 1904

megacephala melancholica Santschi, 1912 (Guinea, Ivory Coast)

megacephala nkomoana Forel, 1916 (Democratic Republic of Congo)

megacephala rotundata Forel, 1894 (Kenya, Mozambique, Zimbabwe)

megacephala speculifrons Stitz, 1911 (United Republic of Tanzania)

megacephala talpa Gerstäcker, 1871 (Kenya)

megatron Fischer & Fisher, 2013 (Comoros)

meihuashanensis Li & Chen, 1992 (China)

meinerti Forel, 1905 (Brazil, Ecuador, Guyana, Peru, Venezuela)

• Pheidole mimula Wheeler, W.M., 1916

meinertopsis Wilson, 2003 (Guyana, Peru)

melanogaster Donisthorpe, 1943 (New Guinea)

melastomae Wilson, 2003 (Colombia, Ecuador, Panama)

mendanai Mann, 1919 (Solomon Islands)

mendicula Wheeler, W.M., 1925 (Brazil, Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Guadeloupe, Panama, Peru, Trinidad and Tobago)

mentita Santschi, 1914 (Guinea)

mentita pullata Santschi, 1914 (Guinea)

mera Wilson, 2003 (Guatemala, Mexico, United States)

merimbun Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines)

mesomontana Longino, 2009 (Costa Rica)

metallescens Emery, 1895 (United States)

• Pheidole metallescens splendidula Wheeler, W.M., 1908

metana Wilson, 2003 (Colombia, Ecuador)

micon Wilson, 2003 (Colombia, Venezuela)

micridris Wilson, 2003 (Ecuador, Peru)

microgyna Wheeler, W.M., 1928 (Guyana)

microps Wilson, 2003 (Ecuador, Peru)

micula Wheeler, W.M., 1915 (Mexico, United States)

midas Wilson, 2003 (Ecuador, Peru, Trinidad and Tobago, Venezuela)

militicida Wheeler, W.M., 1915 (Mexico, United States)

minax Wilson, 2003 (Peru)

minensis Santschi, 1923 (Brazil)

minima Mayr, 1901 (Cameroun, Guinea)

minima catella Santschi, 1914 (Nigeria)

minima faurei Santschi, 1920 (Gabon)

minima malelana Wheeler, W.M., 1922 (Democratic Republic of Congo)

minor (Jerdon, 1851) (India)

minuscula Bernard, 1953 (Guinea, Iraq, Israel, Saudi Arabia)

minutula Mayr, 1878 (Brazil, Ecuador, French Guiana, Guyana, Panama, Peru)

• Pheidole minutula folicola Forel, 1904

• Pheidole minutula personata Santschi, 1929

mirabilis Wilson, 2003 (Colombia)

miseranda Wheeler, W.M., 1924 (Indonesia, Krakatau Islands)

mittermeieri Wilson, 2003 (Bolivia)

mixteca Wilson, 2003 (Mexico)

mjobergi Forel, 1915 (Australia)

modiglianii Emery, 1900 (Borneo, Indonesia, Malaysia)

moerens Wheeler, W.M., 1908 (Dominican Republic, Greater Antilles, Haiti, Puerto Rico, United States)

• Pheidole moerens creola Wheeler, W.M. & Mann, 1914

• Pheidole moerens dominicensis Wheeler, W.M., 1913

moffetti Wilson, 2003 (French Guiana)

monstrosa Wilson, 2003 (Brazil)

montana Eguchi, 1999 (Borneo, Indonesia, Malaysia)

monteverdensis Wilson, 2003 (Costa Rica)

mooreorum Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole fariasana Wilson, 2003

morelosana Wilson, 2003 (Mexico)

morrisii Forel, 1886 (United States)

• Pheidole morrisi impexa Wheeler, W.M., 1908

• Pheidole morrisii vanceae Forel, 1901

moseni Wheeler, W.M., 1925 (Brazil)

mosenopsis Wilson, 2003 (Bolivia, Paraguay)

multidens Forel, 1902 (China, India)

multispina Wilson, 2003 (Costa Rica, Panama)

mus Forel, 1902 (India, Niue)

mutisi Fernández & Wilson, 2008 (Colombia)

mylognatha Wheeler, W.M., 1922 (Democratic Republic of Congo)

N
nana Emery, 1894 (Brazil)

naoroji Forel, 1902 (India)

napoensis Wilson, 2003 (Ecuador)

nasifera Wilson, 2003 (Peru)

nasutoides Hölldobler & Wilson, 1992 (Costa Rica)

navigans Forel, 1901 (Mexico)

naylae Wilson, 2003 (Cuba, Greater Antilles)

nebulosa Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole scabriventris Wilson, 2003

nemoralis Forel, 1892 (Madagascar)

nemoralis petax Forel, 1895 (Madagascar)

neokohli Wilson, 1984 (Democratic Republic of Congo)

• Anergatides kohli Wasmann, 1915

neolongiceps Brown, 1950 (Cuba, Greater Antilles)

• Pheidole longiceps Aguayo Aguayo, 1932

neolongiscapa Özdikmen, 2010 (China)

• Pheidole longiscapa Zhou & Zheng Zhou & Zheng, 1999

neoschultzi LaPolla, 2006 (Guyana)

• Pheidole schultzi LaPolla, 2005

nesiota Wilson, 2003 (Brazil)

nietneri Emery, 1901 (China, Sri Lanka)

nigella Emery, 1894 (Bolivia, Peru)

• Pheidole dimidiata Emery, 1894

nigeriensis Santschi, 1914 (Kenya, Nigeria, Uganda)

nigricula Wilson, 2003 (Costa Rica, Honduras, Nicaragua)

nigritella Bernard, 1953 (Guinea)

nimba Bernard, 1953 (Cameroun, Guinea)

nindi Mann, 1919 (Guam, Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands, Solomon Islands)

nitella Wilson, 2003 (Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Peru)

nitidicollis Emery, 1896 (Belize, Brazil, Costa Rica, Guatemala, Honduras, Jamaica, Mexico, Nicaragua)

• Pheidole sagana Wheeler, W.M., 1934

nitidula Emery, 1888 (Argentina, Brazil)

• Pheidole nitidula daguerrei Santschi, 1931

• Pheidole perversa Forel, 1908

• Pheidole perversa richteri Forel, 1909

• Pheidole strobeli Emery, 1906

• Pheidole strobeli misera Santschi, 1916

• Pheidole strobeli silvicola Borgmeier, 1927

njassae Viehmeyer, 1914 (United Republic of Tanzania)

njassae legitima Santschi, 1916 (United Republic of Tanzania)

• Pheidole njassae sculptior Viehmeyer, 1914

noar Wilson, 2003 (Dominican Republic, Greater Antilles)

nodgii Forel, 1905 (Indonesia)

nodgii verlatenensis Wheeler, W.M., 1937 (Indonesia, Krakatau Islands)

nodifera (Smith, F., 1858) (China, Thailand, Vietnam)

nodus Smith, F., 1874 (Bangladesh, India, Indonesia, Japan, Republic of Korea, Sri Lanka, Taiwan, Thailand, Vietnam)

• Pheidole nodus flebilis Santschi, 1937

• Pheidole nodus praevexata Wheeler, W.M., 1929

• Pheidole rhombinoda Mayr, 1879

• Pheidole rhombinoda formosensis Forel, 1913

• Pheidole rhombinoda micantiventris Mayr, 1897

• Pheidole rhombinoda stella Forel, 1911

• Pheidole rhombinoda taprobanae Forel, 1902

• Pheidole treubi Forel, 1905

nubicola Wilson, 2003 (Mexico)

nubila Emery, 1906 (Argentina)

nuculiceps Wheeler, W.M., 1908 (Mexico, United States)

O
oaxacana Wilson, 2003 (Guatemala, Mexico)

obnixa Forel, 1912 (Argentina)

obrima Wilson, 2003 (Venezuela)

obscurifrons Santschi, 1925 (Argentina)

obscurithorax Naves, 1985 (Argentina, Brazil, Paraguay, United States)

obtusopilosa Mayr, 1887 (Argentina, Brazil, Uruguay)

obtusospinosa Pergande, 1896 (Mexico, United States)

• Pheidole arizonica Santschi, 1911

• Pheidole subdentata Pergande, 1896

occipitalis André, 1890 (Guinea, Senegal, Sierra Leone)

occipitalis adami Santschi, 1939 (Guinea)

occipitalis neutralis Santschi, 1914 (Guinea)

oceanica Mayr, 1866 (Australia, Cook Islands, Fiji, French Polynesia, Guam, Kiribati, Marshall Islands, Micronesia (Federated States of), New Caledonia, New Guinea, Niue, Northern Mariana Islands, Samoa, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Pheidole oceanica boraborensis Wheeler, W.M., 1932

• Pheidole oceanica pattesoni Mann, 1919

• Pheidole oceanica upoluana Santschi, 1928

ocellata Zhou, 2001 (China)

ochracea Eguchi, 2008 (China, Vietnam)

oculata (Emery, 1899) (Madagascar)

oliveirai Wilson, 2003 (Colombia)

olsoni Wilson, 2003 (Costa Rica, Guatemala, Honduras, Panama)

onifera Mann, 1921 (Fiji)

onyx Wilson, 2003 (Costa Rica)

opaciventris Mayr, 1876 (Australia)

optiva Forel, 1901 (Mexico)

orbica Forel, 1893 (Puerto Rico, St. Vincent)

oswaldi Forel, 1891 (Madagascar)

oswaldi decollata Forel, 1892 (Madagascar)

otisi Wilson, 2003 (Costa Rica, Honduras)

oxyops Forel, 1908 (Argentina, Brazil, Paraguay)

• Pheidole genalis Borgmeier, 1929

• Pheidole oxyops regia Forel, 1908

P
paiute Gregg, 1959 (United States)

palenquensis Wilson, 2003 (Ecuador)

pallidula (Nylander, 1849) (Afghanistan, Albania, Algeria, Andorra, Armenia, Austria, Balearic Islands, Bulgaria, Canary Islands, Croatia, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Iran, Israel, Italy, Kazakhstan, Kyrgyzstan, Malta, Monaco, Montenegro, Portugal, Republic of Macedonia, Romania, Russian Federation, Slovenia, Spain, Turkey, Turkmenistan)

• Oecophthora subdentata Mayr, 1853

• Pheidole pallidula obscura Santschi, 1936

• Pheidole symbiotica Wasmann, 1909

• Xenoaphaenogaster inquilina Baroni Urbani, 1964

pallidula selenia Özdikmen, 2010 (Algeria)

• Pheidole pallidula inermis Stitz, 1917

pallidula tasdelenia Özdikmen, 2010 (Tunisia)

• Pheidole pallidula tristis Forel, 1907

pampana Santschi, 1929 (Argentina)

paraensis Wilson, 2003 (Brazil)

pararugiceps Longino, 2009 (Costa Rica)

parasitica Wilson, 1984 (India)

pariana Wilson, 2003 (Venezuela)

parva Mayr, 1865 (Borneo, China, India, Indonesia, Japan, Malaysia, Mauritius, Myanmar, Philippines, Saudi Arabia, Seychelles, Sri Lanka, Taiwan, Thailand, United Arab Emirates, Vietnam)

• Pheidole bugi Wheeler, W.M., 1919

• Pheidole flavens farquharensis Forel, 1907

• Pheidole parva decanica Forel, 1902

• Pheidole rinae mala Forel, 1911

• Pheidole rinae tipuna Forel, 1912

• Pheidole sauteri Wheeler, W.M., 1909

• Pheidole tarda Donisthorpe, 1947

• Pheidole voeltzkowii Forel, 1894

parvicorpus Eguchi, 2001 (Borneo, Indonesia, Malaysia)

peckorum Wilson, 2003 (Ecuador)

pedana Wilson, 2003 (Brazil, Peru)

pegasus Sarnat, 2008 (Fiji)

peguensis Emery, 1895 (Myanmar)

pelor Wilson, 2003 (United States)

peltastes Wilson, 2003 (Brazil, Ecuador)

penetralis Smith, F., 1863 (Indonesia)

pepo Wilson, 2003 (Colombia)

peregrina Wheeler, W.M., 1916 (Brazil)

• Pheidole impariceps Santschi, 1923

• Pheidole impariceps fusciclava Santschi, 1923

perkinsi Wilson, 2003 (Dominica, Lesser Antilles)

perpilosa Wilson, 2003 (United States)

perpusilla Emery, 1894 (Brazil, Colombia, Costa Rica, Guatemala, Guyana, Mexico, Nicaragua, Panama, Trinidad and Tobago)

• Pheidole decem Forel, 1901

• Pheidole emersoni Wheeler, W.M., 1922

• Pheidole flavens breviscapa Forel, 1899

perryorum Wilson, 2003 (Brazil)

peruviana Wilson, 2003 (Ecuador, Peru)

petrensis Wilson, 2003 (Mexico)

phanigaster Longino, 2009 (Mexico)

philemon Forel, 1910 (Solomon Islands)

philippi Emery, 1915 (Eritrea, Ethiopia)

phipsoni Forel, 1902 (India)

pholeops Wilson, 2003 (Brazil, Ecuador, Peru)

picea (Buckley, 1867)

piceonigra Emery, 1922 (Costa Rica, Mexico)

• Pheidole picea Mayr Mayr, 1870

picobarva Longino, 2009 (Costa Rica)

pidax Wilson, 2003 (Colombia)

pieli Santschi, 1925 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Thailand, Vietnam)

• Pheidole rinae incensa Wheeler, W.M., 1928

pilifera (Roger, 1863) (Mexico, United States)

• Pheidole pennsylvanica Roger, 1863

• Pheidole pilifera artemisia Cole, 1933

• Pheidole pilifera coloradensis Emery, 1895

• Pheidole pilifera septentrionalis Wheeler, W.M., 1908

• Pheidole xerophila pacifica Wheeler, W.M., 1915

pilispina Wilson, 2003 (Panama)

piliventris (Smith, F., 1858) (Brazil)

pilosior Wilson, 2003 (United States)

pinealis Wheeler, W.M., 1908 (Mexico, United States)

pinicola Wilson, 2003 (Mexico)

plagiaria Smith, F., 1860 (Borneo, China, Indonesia, Krakatau Islands, Malaysia, Myanmar, Philippines, Thailand, Vietnam)

• Pheidole divergens Mayr, 1867

• Pheidole peguensis yomensis Forel, 1903

plagiaria moica Forel, 1911 (Vietnam)

plagiaria palawanica Stitz, 1925 (Philippines)

plagiaria rectilineata Viehmeyer, 1916 (Indonesia, Sulawesi)

planidorsum Eguchi, 2001 (Borneo, Indonesia, Malaysia, Vietnam)

planifrons Santschi, 1920 (Thailand, Vietnam)

plato Wilson, 2003 (Ecuador)

platypus Crawley, 1915 (Australia)

plebecula Forel, 1899

plinii Forel, 1911 (Borneo, Indonesia, Malaysia, Singapore)

polita Emery, 1894 (Bolivia, Brazil, Peru)

• Pheidole wheeleri Mann, 1916

polymorpha Wilson, 2003 (Mexico)

porcula Wheeler, W.M., 1908 (Mexico, United States)

poringensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

portalensis Wilson, 2003 (Mexico, United States)

potosiana Wilson, 2003 (Costa Rica, Mexico)

praeses Wilson, 2003 (Brazil)

praeusta Roger, 1863 (Colombia)

prattorum Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

prelli Forel, 1911 (Kenya)

prelli ingenita Santschi, 1928 (Ivory Coast)

prelli redbankensis Forel, 1913 (Mozambique, Zimbabwe)

pronotalis Forel, 1902 (India, Sri Lanka)

prostrata Wilson, 2003 (Costa Rica, Panama)

protaxi Oliveira & Lacau, 2015

protea Forel, 1912 (Indonesia, Thailand)

protensa Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

providens (Sykes, 1835) (India)

proxima Mayr, 1876 (Australia, New Zealand)

proxima bombalensis Forel, 1910 (Australia)

proxima transversa Forel, 1902 (Australia)

psammophila Creighton & Gregg, 1955 (Mexico, United States)

psilogaster Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

pubiventris Mayr, 1887 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Mexico)

• Pheidole indistincta Forel, 1899

• Pheidole pubiventris cearensis Forel, 1901

• Pheidole pubiventris nevadensis Forel, 1901

• Pheidole pubiventris timmii Forel, 1901

• Pheidole pubiventris variegata Emery, 1896

pugnax Dalla Torre, 1892 (Costa Rica, El Salvador, Honduras, Panama)

• Pheidole radowszkowskii militaris Emery, 1890

pulchella Santschi, 1910 (Cameroun, Congo, Kenya)

• Pheidole niapuana Wheeler, W.M., 1922

• Pheidole pulchella achantella Santschi, 1939

pullula Santschi, 1911 (Argentina)

punctatissima Mayr, 1870 (Belize, Colombia, Costa Rica, Denmark, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Pheidole punctatissima napaea Wheeler, W.M., 1934

punctithorax Borgmeier, 1929 (Brazil)

punctulata Mayr, 1866 (Comoros, Eritrea, Guinea, Ivory Coast, Kenya, Madagascar, Mayotte, Mozambique, Namibia, Seychelles, South Africa, Zimbabwe)

punctulata atrox Forel, 1913 (Democratic Republic of Congo)

• Pheidole inquilina Forel Forel, 1914

punctulata spinosa Forel, 1891 (Madagascar)

punctulata subatrox Santschi, 1937 (Democratic Republic of Congo)

purpurascens Emery, 1897 (New Guinea)

purpurea Longino, 2009 (Guatemala, Honduras, Mexico)

puttemansi Forel, 1911 (Brazil)

• Pheidole fallax janeirensis Forel, 1912

pygmaea Wilson, 2003 (Colombia)

pyriformis Clark, 1938 (Australia)

Q
quadrensis Forel, 1900 (Borneo, Indonesia, Malaysia)

quadriceps Wilson, 2003 (Colombia)

quadricuspis Emery, 1900 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

quadriprojecta Smith, M.R., 1947 (New Guinea)

quadrispinosa (Smith, F., 1865) (Indonesia, New Guinea)

• Pheidolacanthinus armatus Smith, F., 1865

• Pheidole smithi Emery, 1887

quercicola Wilson, 2003 (Mexico)

quiaccana Wheeler, W.M., 1925 (Bolivia)

quinata Eguchi, 2000 (Borneo, Indonesia, Malaysia)

R
rabo Forel, 1913 (Borneo, China, Indonesia, Malaysia, Thailand, Vietnam)

• Pheidole concinna Wheeler, W.M. Wheeler, W.M., 1928

• Pheidole tsailuni Wheeler, W.M., 1929

radoszkowskii Mayr, 1884 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Greater Antilles, Guatemala, Guyana, Honduras, Jamaica, Lesser Antilles, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Venezuela)

• Pheidole medialis Wilson, 2003

• Pheidole radoszkowskii luteola Forel, 1893

• Pheidole radoszkowskii opacissima Forel, 1901

• Pheidole radoszkowskii parvinoda Forel, 1912

• Pheidole radowszkowskii acuta Emery, 1894

• Pheidole radowszkowskii australis Emery, 1890

• Pheidole triconstricta paranana Santschi, 1925

ragnax Fischer & Fisher, 2013 (Mayotte)

rebeccae Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

reclusi Forel, 1899 (Colombia)

recondita Clouse, 2007 (Guam, Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands)

rectisentis Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

rectispina Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico)

rectitrudis Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

reflexans Santschi, 1933 (Argentina, Paraguay)

reichenspergeri Santschi, 1923 (Brazil)

renae Wilson, 2003 (Costa Rica)

retivertex Eguchi, 2001 (Borneo, Indonesia, Malaysia)

retronitens Santschi, 1930 (Democratic Republic of Congo)

rhea Wheeler, W.M., 1908 (Mexico, United States)

rhinoceros Forel, 1899 (Colombia, Costa Rica, Honduras, Mexico, Nicaragua, Panama)

rhinomontana Longino, 2009 (Costa Rica, Honduras, Nicaragua)

rhytifera Wilson, 2003 (Venezuela)

rinae Emery, 1900 (Borneo, Indonesia, Malaysia, Thailand, Vietnam)

risii Forel, 1892 (Argentina, Brazil)

riveti Santschi, 1911 (Ecuador)

roberti Forel, 1902 (China, India)

rochai Forel, 1912 (Brazil)

rogeri Emery, 1896 (Costa Rica, Honduras)

rogersi Forel, 1902 (Bangladesh, India)

rogersi taylori Forel, 1902 (India)

rohani Santschi, 1925 (Angola)

rohani monardi Santschi, 1930 (Angola)

rohani pellax Santschi, 1930 (Angola)

roosevelti Mann, 1921 (Fiji)

rosae Forel, 1901 (Argentina, Brazil)

• Pheidole silvestrii Emery, 1906

rosula Wilson, 2003 (Argentina)

rotundiceps Wilson, 2003 (Ecuador)

roushae Wilson, 2003 (Belize, Guatemala, Honduras, Mexico)

rubiceps Wilson, 2003 (Venezuela)

rudigenis Emery, 1906 (Argentina, Brazil, Paraguay)

• Pheidole bucolica Santschi, 1929

ruficeps (Smith, F., 1861) (Indonesia, Sulawesi)

• Pheidole mordax Smith, F., 1863

rufipilis Forel, 1908 (Brazil)

• Pheidole pubiventris impia Santschi, 1923

• Pheidole rufipilis divexa Forel, 1908

rugaticeps Emery, 1877 (Burkina Faso, Eritrea, Ethiopia, Mali, Oman, Senegal, South Africa)

rugaticeps arabs Emery, 1881 (Eritrea, Yemen)

rugatula Santschi, 1933 (Argentina, Paraguay)

rugiceps Wilson, 2003 (Belize, Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

rugifera Eguchi, 2001 (Borneo, Indonesia, Malaysia)

rugithorax Eguchi, 2008 (Vietnam)

rugosa Smith, F., 1858 (Sri Lanka)

rugulosa Gregg, 1959 (Mexico, United States)

rutilana Wilson, 2003 (Brazil)

ryukyuensis Ogata, 1982 (Japan)

S
sabahna Eguchi, 2000 (Borneo, Indonesia, Malaysia)

sabella Wilson, 2003 (Ecuador, Peru)

sabina Wilson, 2003 (Mexico)

sagax Wilson, 2003 (Colombia, Ecuador, Nicaragua, Venezuela)

sagei Forel, 1902 (China, India, Nepal)

sagittaria Wilson, 2003 (Costa Rica)

sarawakana Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

sarcina Forel, 1912 (Brazil, Ecuador)

sarpedon Wilson, 2003 (Ecuador)

sauberi Forel, 1905 (Borneo, Indonesia, Krakatau Islands, Malaysia, New Guinea, Philippines, Thailand)

saxicola Wheeler, W.M., 1922 (Democratic Republic of Congo)

sayapensis Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

scabriuscula Gerstäcker, 1871 (Kenya)

scalaris Wilson, 2003 (Costa Rica, Ecuador, Nicaragua, Trinidad and Tobago, Venezuela)

scapulata Santschi, 1923 (Argentina, Bolivia, Paraguay)

schmalzi Emery, 1894 (Bolivia, Brazil, Venezuela)

• Pheidole rata Borgmeier, 1929

• Pheidole rehi Forel, 1900

schoedli Eguchi, Hashimoto & Malsch, 2006 (Borneo, Indonesia, Malaysia)

schoutedeni Forel, 1913 (Democratic Republic of Congo)

schoutedeni platycephala Stitz, 1916 (Democratic Republic of Congo)

schultzei Forel, 1910 (Botswana)

schultzei ebangana Santschi, 1937 (Angola)

schultzei gwaaiensis Forel, 1913 (Zambia, Zimbabwe)

schultzei woodvalensis Arnold, 1960 (Zimbabwe)

schwarzmaieri Borgmeier, 1939 (Argentina, Brazil)

sciara Cole, 1955 (Mexico, United States)

scimitara Wilson, 2003 (Brazil, Peru)

sciophila Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole proserpina Wheeler, W.M., 1908

• Pheidole sciophila semilaevicephala Smith, M.R., 1934

scolioceps Wilson, 2003 (Brazil, Ecuador, Guyana, Peru)

scrobifera Emery, 1896 (Costa Rica, Honduras, Nicaragua, Panama)

sculptior Forel, 1893 (Barbados, Colombia, Puerto Rico, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole tayrona Wilson, 2003

sculpturata Mayr, 1866 (Guinea, Iran, Ivory Coast, Kenya, Namibia, Oman, South Africa, United Arab Emirates)

sculpturata areolata Forel, 1911 (South Africa)

sculpturata berthoudi Forel, 1894 (South Africa)

sculpturata dignata Santschi, 1915 (Congo)

sculpturata particeps Santschi, 1921 (United Republic of Tanzania)

sculpturata rhodesiana Forel, 1913 (Zimbabwe)

sculpturata welgelegenensis Forel, 1913 (Democratic Republic of Congo)

sculpturata zambesiana Forel, 1913 (Zambia, Zimbabwe)

sebofila Longino, 2009 (Costa Rica)

securigera Wilson, 2003 (Colombia)

seeldrayersi Forel, 1910 (Argentina, Bolivia, Colombia)

selathorax Zhou, 2001 (China)

seligmanni Wilson, 2003 (Colombia)

semidea Fischer, Hita Garcia & Peters, 2012 (Nigeria)

semilaevis Forel, 1901 (Brazil)

senex Gregg, 1952 (Mexico, United States)

• Pheidole pilifera anfracta Cole, 1952

senilis Santschi, 1929 (Brazil)

sensitiva Borgmeier, 1959 (Brazil, Colombia, Costa Rica, Honduras, Nicaragua, Panama, Peru)

sepulchralis Bingham, 1903 (India)

sericella Viehmeyer, 1914 (New Guinea)

servilia Wilson, 2003 (Colombia)

setosa Fischer, Hita Garcia & Peters, 2012 (Democratic Republic of Congo)

setsukoae Wilson, 2003 (Mexico)

severini Forel, 1904 (French Guiana, Suriname)

sexdentata Donisthorpe, 1948 (New Guinea)

sexspinosa Mayr, 1870 (Cook Islands, French Polynesia, Micronesia (Federated States of), New Guinea, Niue, Palau, Samoa, Solomon Islands, Tokelau, Tuvalu, Vanuatu, Wallis and Futuna Islands)

• Pheidole sexspinosa adamsoni Wheeler, W.M., 1932

sexspinosa biroi Emery, 1900 (New Guinea)

sexspinosa fuscescens Emery, 1900 (New Guinea, Solomon Islands)

sharpi Forel, 1902 (India)

sharpi hoogwerfi Forel, 1902 (India)

sicaria Wilson, 2003 (Costa Rica)

sigillata Wilson, 2003 (Brazil, Paraguay)

sikorae Forel, 1891 (Madagascar)

sikorae litigiosa Forel, 1892 (Madagascar)

similigena Wheeler, W.M., 1937 (Cuba)

simoni Emery, 1893 (Philippines)

simonsi Wilson, 2003 (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

simplex Wheeler, W.M., 1925 (Brazil, Peru)

simplispinosa Sarnat, 2008 (Fiji)

sinaitica Mayr, 1862 (Egypt, Eritrea, Iran, Iraq, Israel, Kenya, United Arab Emirates)

sinaitica santschii Forel, 1907 (Tunisia)

singaporensis Özdikmen, 2010 (India, Philippines, Singapore)

• Myrmica longipes Smith, F., 1857

singaporensis conicollis (Emery, 1900) (Indonesia)

singaporensis continentis (Forel, 1911) (Myanmar)

singaporensis pseudola Forel, 1915 (Indonesia)

singularis Smith, F., 1863 (Indonesia, New Guinea)

sinica (Wu & Wang, 1992) (China)

sitiens Wilson, 2003 (United States)

skwarrae Wheeler, W.M., 1934 (Mexico)

smythiesii Forel, 1902 (China, India, Nepal, Thailand, Vietnam)

• Pheidole bhavanae Bingham, 1903

• Pheidole smythiesii bengalensis Forel, 1902

socrates Forel, 1912 (Brazil, French Guiana, Guyana, Suriname)

soesilae Makhan, 2007 (Suriname)

soritis Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole sitarches Wheeler, W.M., 1908

• Pheidole sitarches campestris Creighton, 1950

• Pheidole sitarches rufescens Wheeler, W.M., 1908

• Pheidole sitarches transvarians Wheeler, W.M., 1908

sospes Forel, 1908 (Argentina, Brazil, Colombia, Ecuador, Peru)

• Pheidole minutissima Kusnezov, 1952

spadonia Wheeler, W.M., 1915 (Mexico, United States)

sparsisculpta Longino, 2009 (Costa Rica)

spathicornis Wilson, 2003 (Mexico)

spathifera Forel, 1902 (Bangladesh, China, India, Thailand)

spathifera aspatha Forel, 1902 (India)

spathifera yerburyi Forel, 1902 (India, Myanmar, Sri Lanka)

spathipilosa Wilson, 2003 (Costa Rica, Nicaragua)

specularis Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

speculifera Emery, 1877 (Cameroun, Eritrea, Ethiopia, Guinea, Kenya, Mali)

speculifera ascara Emery, 1901 (Eritrea, Ethiopia, Kenya)

speculifera bispecula Santschi, 1930 (Democratic Republic of Congo)

speculifera cubangensis Forel, 1901 (Angola)

sperata Forel, 1915 (Indonesia)

sphaerica Wilson, 2003 (Colombia)

spilota Wilson, 2003 (Colombia)

spinicornis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

spininodis Mayr, 1887 (Argentina, Brazil, Paraguay)

• Pheidole hohenlohei Emery, 1888

• Pheidole spielbergi Emery, 1888

• Pheidole spininodis bruta Santschi, 1934

• Pheidole spininodis lucifuga Santschi, 1923

• Pheidole spininodis pencosensis Forel, 1914

• Pheidole spininodis solaris Santschi, 1929

spinoda (Smith, F., 1858) (Australia)

spinulosa Forel, 1910 (Lesotho, Zimbabwe)

spinulosa conigera Forel, 1910 (South Africa)

spinulosa messalina Forel, 1910 (Lesotho)

squalida Santschi, 1910 (Congo)

steinheili Forel, 1901 (Colombia)

stigma Wilson, 2003 (Ecuador)

stomachosa Wheeler, W.M., 1917 (Jamaica)

strator Forel, 1910 (Eritrea, Ethiopia, Kenya)

strator fugax Arnold, 1920 (Mozambique)

strator tabida Menozzi, 1933 (Uganda)

striata Donisthorpe, 1947 (New Guinea)

striaticeps Mayr, 1870 (Costa Rica, Guatemala, Honduras, India, Mexico)

strigosa Wilson, 2003 (Brazil)

stulta Forel, 1886 (Brazil, Costa Rica, Panama)

• Pheidole championi Forel, 1899

• Pheidole championii sima Forel, 1912

styrax Wilson, 2003 (Venezuela)

subaberrans (Kusnezov, 1952) (Argentina)

subarmata Mayr, 1884 (Argentina, Bahamas, Barbados, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, French Guiana, Greater Antilles, Guatemala, Honduras, Mexico, Panama, Paraguay, Puerto Rico, St. Vincent, Trinidad and Tobago)

• Pheidole cornutula Emery, 1890

• Pheidole cornutula dentimentum Santschi, 1929

• Pheidole cornutula imbecillis Emery, 1906

• Pheidole hondurensis Mann, 1922

• Pheidole subarmata borinquenensis Wheeler, W.M., 1908

• Pheidole subarmata elongatula Forel, 1893

• Pheidole subarmata nassavensis Wheeler, W.M., 1905

• Pheidole subarmata nefasta Santschi, 1929

submonticola Eguchi, 2001 (Borneo, Indonesia, Malaysia)

subnuda Wilson, 2003 (Colombia, Venezuela)

subreticulata Emery, 1894 (Brazil)

subsphaerica Wilson, 2003 (Colombia)

sulcaticeps Roger, 1863 (Bangladesh, China, India, Sri Lanka)

sulcaticeps punensis Forel, 1902 (India)

sulcaticeps vellicans Forel, 1911 (Sri Lanka)

superba Wilson, 2003 (Colombia, Panama)

susannae Forel, 1886 (Argentina, Barbados, Brazil, Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Paraguay, Puerto Rico, Trinidad and Tobago, Venezuela)

• Pheidole incisa evoluta Borgmeier, 1929

• Pheidole partita Mayr, 1887

• Pheidole susannae atricolor Forel, 1901

• Pheidole susannae obscurior Forel, 1886

susanowo Onoyama & Terayama, 1999 (Japan)

sykesii Forel, 1902 (India)

synanthropica Longino, 2009 (Costa Rica, Honduras, Mexico, Nicaragua)

synarmata Wilson, 2003 (Brazil, Costa Rica, El Salvador, Guyana, Honduras, Nicaragua, Venezuela)

T
tachigaliae Wheeler, W.M., 1921 (French Guiana, Guyana)

tachirana Wilson, 2003 (Venezuela)

taipoana Wheeler, W.M., 1928 (China, Vietnam)

taivanensis Forel, 1912 (Taiwan, Thailand)

tambopatae Wilson, 2003 (Brazil, Colombia, Peru)

tandjongensis Forel, 1913 (China, Indonesia, Thailand, Vietnam)

tanyscapa Wilson, 2003 (Costa Rica, Nicaragua, Panama)

tarchon Wilson, 2003 (Argentina)

tasmaniensis Mayr, 1866 (Australia)

tasmaniensis continentis Forel, 1902 (Australia)

taurus Emery, 1906 (Argentina)

tawauensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

templaria Forel, 1902 (India)

templaria euscrobata Forel, 1913 (Sri Lanka)

tenebricosa Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

tenerescens Wheeler, W.M., 1922 (Trinidad and Tobago)

tennantae Wilson, 2003 (Colombia, Costa Rica, Panama, Peru, Venezuela)

tenuicephala Longino, 2009 (Costa Rica)

tenuiclavata Donisthorpe, 1943 (New Guinea)

tenuinodis Mayr, 1901 (Namibia, South Africa)

tenuinodis bothae Forel, 1901 (South Africa)

tenuinodis robusta Stitz, 1923 (Namibia)

tenuinodis sipapomae Arnold, 1920 (Zimbabwe)

tenuis Wilson, 2003 (Colombia)

tepicana Pergande, 1896 (Mexico, United States)

• Pheidole carbonaria Pergande, 1896

• Pheidole kingi André, 1898

• Pheidole kingi instabilis Emery, 1901

• Pheidole kingi torpescens Wheeler, W.M., 1915

• Pheidole rugifrons Pergande, 1896

• Pheidole townsendi André, 1898

tepuicola Wilson, 2003 (Venezuela)

termitobia Forel, 1901 (Brazil)

termitophila Forel, 1904 (Guinea, Liberia, Mali, Senegal, Sudan)

termitophila liberiensis Forel, 1911 (Liberia)

terraceensis Bharti, 2001 (India)

terresi Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

• Pheidole terresi illota Wheeler, W.M. & Mann, 1914

terribilis Wilson, 2003 (Bolivia, Brazil, Suriname)

tetra Creighton, 1950 (Mexico, United States)

tetracantha Emery, 1897 (New Guinea)

tetrica Forel, 1913 (Brazil)

tetroides Wilson, 2003 (Mexico)

texana Wheeler, W.M., 1903 (United States)

texticeps Wilson, 2003 (Costa Rica, Honduras, Mexico, Nicaragua, Panama)

• Pheidole perdiligens Wilson, 2003

thrasys Wilson, 2003 (Honduras, Panama)

tigris Wilson, 2003 (Colombia, Panama)

tijucana Borgmeier, 1927 (Brazil)

tillandsiarum Wheeler, W.M., 1934 (Mexico)

tisiphone Wheeler, W.M., 1911 (Guatemala, Mexico)

titanis Wheeler, W.M., 1903 (Mexico, United States)

tjibodana Forel, 1905 (Borneo, Cambodia, China, Indonesia, Malaysia, Philippines, Thailand, Vietnam)

tobini Wilson, 2003 (Ecuador, Peru)

tolteca Forel, 1901 (Mexico)

torosa Wilson, 2003 (Mexico)

trachyderma Emery, 1906 (Argentina)

trageri Wilson, 2003 (Brazil)

tragica Wheeler, W.M., 1934 (Mexico)

traini Wilson, 2003 (Costa Rica, Panama)

transfigens Forel, 1911 (New Guinea)

transversostriata Mayr, 1887 (Barbados, Brazil, Colombia, Guyana, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole lacerta Wheeler, W.M., 1922

• Pheidole transversostriata nigridens Forel, 1901

trapezoidea Viehmeyer, 1914 (Australia)

tricarinata Santschi, 1914 (Guinea)

tricolor Donisthorpe, 1949 (New Guinea)

triconstricta Forel, 1886 (Argentina, Bolivia, Brazil, Grenada, Paraguay, Uruguay)

• Pheidole radoszkowskii discursans Forel, 1912

• Pheidole radoszkowskii saviozae Forel, 1911

• Pheidole triconstricta ambulans Emery, 1906

• Pheidole triconstricta hebe Santschi, 1923

• Pheidole triconstricta rosariensis Forel, 1913

trinitatis Wilson, 2003 (Trinidad and Tobago)

triplex Wilson, 2003 (Ecuador, Trinidad and Tobago)

tristicula Wilson, 2003 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Peru, Trinidad and Tobago)

tristis (Smith, F., 1858) (Argentina, Brazil, Paraguay)

• Atta fumipennis Smith, F., 1858

• Atta rubra Smith, F., 1858

• Pheidole crassipes Mayr, 1887

• Pheidole emeryi Mayr, 1887

• Pheidole emeryi alsia Santschi, 1923

tristops Wilson, 2003 (Colombia)

truncula Wilson, 2003 (Costa Rica)

tschinkeli Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua, Venezuela)

tumida Eguchi, 2008 (China, Vietnam)

turneri Forel, 1902 (Australia)

tuxtlasana Wilson, 2003 (Guatemala, Honduras, Mexico)

tysoni Forel, 1901 (Mexico, United States)

U
ulothrix Wilson, 2003 (Costa Rica, Panama)

ululevu Fischer, Sarnat & Economo, 2016 (Cook Islands, Fiji)

umbonata Mayr, 1870 (Fiji, French Polynesia, Guam, Kiribati, Micronesia (Federated States of), New Caledonia, Niue, Northern Mariana Islands, Palau, Samoa, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Pheidole umbonata zimmermani Wheeler, W.M., 1936

umbonata fusciventris Emery, 1900 (New Guinea)

umphreyi Wilson, 2003 (Costa Rica, Panama)

uncagena Sarnat, 2008 (Fiji)

unicornis Wilson, 2003 (Colombia, Ecuador)

upeneci Forel, 1913 (Borneo, Indonesia, Malaysia)

ursus Mayr, 1870 (Belize, Costa Rica, Guatemala, Honduras, Mexico)

• Macromischa cressoni André, 1887

• Pheidole ursus gracilinoda Forel, 1904

V
vafella Wheeler, W.M., 1925 (Bolivia)

vafra Santschi, 1923 (Argentina, Brazil, Colombia, Costa Rica, French Guiana, Nicaragua, Panama, Paraguay, Trinidad and Tobago, Venezuela)

• Pheidole idiota Santschi, 1923

valens Wilson, 2003 (Brazil)

vallicola Wheeler, W.M., 1915 (Mexico, United States)

vallifica Forel, 1901 (Brazil, Colombia, Costa Rica, Honduras, Paraguay)

vanderveldi Forel, 1913 (Democratic Republic of Congo)

variabilis Mayr, 1876 (Australia)

variabilis aliena Emery, 1914 (New Caledonia)

variabilis croceithorax Emery, 1914 (New Caledonia)

variabilis latigena Forel, 1907 (Australia)

variabilis mediofusca Forel, 1902 (Australia)

variabilis ocior Forel, 1915 (Australia)

variabilis ocyma Forel, 1915 (Australia)

variabilis parvispina Forel, 1902 (Australia)

variabilis praedo Forel, 1902 (Australia)

variabilis redunca Crawley, 1915 (Australia)

variabilis rugocciput Forel, 1902 (Australia)

variabilis rugosula Forel, 1902 (Australia, New Zealand)

variolosa Emery, 1892 (Eritrea, Somalia)

vatu Mann, 1921 (Fiji)

veletis Wilson, 2003 (Bolivia, Brazil, Panama)

velox Emery, 1887 (Indonesia)

venatrix Wilson, 2003 (Colombia)

verricula Wilson, 2003 (Costa Rica, Honduras)

vestita Wilson, 2003 (Costa Rica)

veteratrix Forel, 1891 (Madagascar)

veteratrix angustinoda Forel, 1892 (Madagascar)

victima Santschi, 1929 (Brazil)

victoris Forel, 1913 (Zimbabwe)

vieti Eguchi, 2008 (China, Vietnam)

vigilans (Smith, F., 1858) (Australia, New Zealand, Norfolk Island)

• Pheidole ampla norfolkensis Wheeler, W.M., 1927

• Pheidole ampla parallela Forel, 1902

• Pheidole ampla yarrensis Forel, 1902

• Pheidole dolichocephala André, 1896

violacea Wilson, 2003 (Costa Rica)

virago Wheeler, W.M., 1915 (United States)

viriosa Wilson, 2003 (Ecuador)

vistana Forel, 1914 (Mexico, United States)

• Pheidole grallipes Wheeler, W.M., 1916

• Pheidole susannae longipes Pergande, 1896

vomer Wilson, 2003 (Colombia)

vorax (Fabricius, 1804) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Trinidad and Tobago)

• Pheidole cephalica Smith, F., 1858

• Pheidole opaca Mayr, 1862

• Pheidole opaca apterostigmoides Weber, 1943

• Pheidole opaca incrustata Forel, 1908

• Pheidole opaca sarrita Forel, 1908

vulcan Fischer & Fisher, 2013 (Comoros)

vulgaris Eguchi, 2006 (China, India, Thailand, Vietnam)

W
walkeri Mann, 1922 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole arietans Wilson, 2003

• Pheidole glyphoderma Wilson, 2003

• Pheidole triumbonata Wilson, 2003

wallacei Mann, 1916 (Brazil)

wardi Wilson, 2003 (Belize, Guatemala, Honduras, Mexico)

watsoni Forel, 1902 (Bangladesh, China, India, Myanmar, Nicobar Island)

weiseri Santschi, 1923 (Argentina)

wheelerorum Mackay, W.P., 1988 (United States)

wiesei Forel, 1910 (Australia)

williamsi Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

• Pheidole williamsi seymourensis Wheeler, W.M., 1924

wilsoni Mann, 1921 (Fiji)

wolfringi Forel, 1908 (Brazil)

woodmasoni Forel, 1885 (Bangladesh, India)

wroughtonii Forel, 1902 (India)

X
xanthocnemis Emery, 1914 (New Caledonia)

xanthogaster Wilson, 2003 (Colombia, Ecuador, Peru)

xerophila Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole xerophila tucsonica Wheeler, W.M., 1908

xocensis Forel, 1913 (Zimbabwe)

xocensis bulawayensis Forel, 1913 (Zimbabwe)

xyston Wilson, 2003 (Guatemala, Mexico)

Y
yaqui Creighton & Gregg, 1955 (Mexico, United States)

yeensis Forel, 1902 (China, Myanmar, Thailand, Vietnam)

yucatana Wilson, 2003 (Mexico)

Z
zelata Wilson, 2003 (Bolivia)

zeteki Smith, M.R., 1947 (Ecuador, Panama, Peru, Trinidad and Tobago)

zhoushanensis Li & Chen, 1992 (China)

zoceana Santschi, 1925 (China, Thailand, Vietnam)

zoster Wilson, 2003 (Brazil, Peru)

=Fossil Species=

C
cordata (Holl, 1829) (Baltic amber)

P
primigenia Baroni Urbani, 1995 (Dominican amber)

R
rasnitsyni Dubovikoff, 2011 (Baltic amber)

T
tertiaria Carpenter, 1930

tethepa Wilson, 1985 (Dominican amber)