Temnothorax parvulus

Identification
Csösz et al. (2015) - A member of the parvulus species-complex. Temnothorax parvulus can be separated easily from members of other species-complexes by its homogenously areolate microsculpture on the head dorsum. The head sculpture of other species complexes may vary from smooth to coarsely rugulo-reticulate but is never homogenously areolate. In exceptional cases, long-spined species of the Temnothorax lichtensteini complex might exhibit a homogenous areolate sculpture on an extended area of the head dorsum, but T. parvulus is distinguished from T. lichtensteini and Temnothorax laconicus by its more erect propodeal spines (38–42° vs. ca. 20–25°). Temnothorax parvulus can also be safely separated from weakly sculptured, lightly colored Temnothorax tergestinus samples using slightly overlapping NOH/CS, SPTI/CS and SPWI/CS ratios (see S4 Table).

Temnothorax parvulus shares most its surface sculpture characters with other species belonging to this complex, Temnothorax ariadnae and Temnothorax helenae. Spine length ratio (SPST/CL) helps distinguish Temnothorax parvulus from T. ariadnae and T. helenae.

Non-overlapping values of SPST/CL for nest sample means of T. parvulus (mean: 0.282 [min: 0.259, max: 0.303]); and T. ariadnae (mean: 0.216 [min: 0199, max: 0.227]) provide a prefect means of separation. As 5% of nest samples of T. parvulus and T. helenae (mean: 0.234 [min: 0.189, max: 0.274] overlap in this ratio, the safest determination is provided by a discriminant (D5 = 0.0393_CL -0.0616_SPST +0.0892_PEW -0.0580_PPW -0.0338_SPWI -8.7356) formula, which separates 97.1% of single individuals and all nest samples of T. parvulus and T. helenae.

Distribution
Western Europe to the Black see coast and Turkey and from Italy and the Balkans to Central Europe.

Distribution based on Regional Taxon Lists
Palaearctic Region: Andorra, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Czech Republic, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Montenegro, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Turkey, Turkmenistan.

Nomenclature

 *  parvulus. Myrmica parvula Schenck, 1852: 103 (w.) GERMANY.
 * [Also described as new by Schenck, 1853: 187.]
 * Mayr, 1855: 448 (q.); Novák, in Kratochvíl, Novák & Šnoflák, 1944: 116 (m.).
 * Combination in Leptothorax: Mayr, 1855: 448.
 * Combination in Temnothorax: Bolton, 2003: 271.
 * Junior synonym of cingulata: Nylander, 1856b: 93.
 * Junior synonym of nylanderi: Smith, F. 1858b: 8; Roger, 1863b: 26.
 * Subspecies of tuberum: André, 1883a: 300.
 * Subspecies of nylanderi: Forel, 1874: 86; Dalla Torre, 1893: 126; Ruzsky, 1902d: 23; Stitz, 1914: 64; Forel, 1915d: 22 (in key); Emery, 1916b: 175; Menozzi, 1918: 85; Bondroit, 1918: 123; Menozzi, 1918: 85; Müller, 1923b: 95; Emery, 1924d: 255; Novák & Sadil, 1941: 91 (in key).
 * Status as species: Mayr, 1855: 448; Bernard, 1956a: 161; Bernard, 1959: 349; Bernard, 1967: 203 (redescription); Baroni Urbani, 1971c: 119; van Boven, 1977: 105; Arnol'di & Dlussky, 1978: 541 (in key); Agosti & Collingwood, 1987b: 274 (in key); Dlussky, Soyunov & Zabelin, 1990: 191; Casevitz-Weulersse, 1990c: 418 (in key); Atanassov & Dlussky, 1992: 140; Arakelian, 1994: 56; Radchenko, 1994d: 153 (in key); Bolton, 1995b: 242; Radchenko, 1995d: 4; Czechowski, et al. 2002: 52; Seifert, 2007: 241; Boer, 2010: 52; Czechowski, et al. 2012: 150; Csösz, Heinze & Mikó, 2015: 47 (redescription).

Type Material
Csösz et al. (2015) - Lectotype: “L. parvulus # Sch” and “Lectotype Leptothorax parvulus Schenck, 1852 det. A. Schulz &M. Verhaagh 1999” (1# / SMF), [GER:parvulus-TYPE].

Worker
Csösz et al. (2015) - Body color: brown; yellow. Body color pattern: mesosoma, antenna and legs, waist and anterior region of 1st gastral tergite lighter than head dorsum and posterior region of gaster. Antenna color pattern: clava concolorous funicle. Absolute cephalic size: 488–586 μm (mean = 550, n = 29). Cephalic length vs. Maximum width of head capsule (CL/CWb): 1.147–1.214 (mean = 1.184). Postocular distance vs. cephalic length (PoOc/CL): 0.392–0.413 (mean = 0.405). Postocular sides of cranium contour frontal view orientation: converging posteriorly. Postocular sides of cranium contour frontal view shape: strongly convex. Vertex contour line in frontal view shape: straight. Vertex sculpture: main sculpture absent, ground sculpture areolate; main sculpture dispersed forked costate sculpture, ground sculpture areolate. Genae contour from anterior view orientation: converging. Gena contour line in frontal view shape: convex. Gena sculpture: rugoso-reticulate with areolate ground sculpture. Median region of antennal rim vs. frontal carina in frontal view structure: not fully overlapped by frontal carina. Concentric carinae laterally surrounding antennal foramen count: present. Eye length vs. absolute cephalic size (EL/CS): 0.237–0.262 (mean = 0.250). Frontal carina distance vs. absolute cephalic size (FRS/CS): 0.353–0.376 (mean = 0.361). Longitudinal carinae on median region of frons count: absent. Smooth median region on frons count: present. Antennomere count: 12. Scape length vs. absolute cephalic size (SL/CS): 0.763–0.796 (mean = 0.778). Facial area of the scape absolute setal angle: 0–15°. External area of the scape absolute setal angle: 30°. Ground sculpture of submedian area of clypeus: smooth. Median carina of clypeus count: present. Lateral carinae of clypeus count: present. Median anatomical line of propodeal spine angle value to Weber length in lateral view: 38–42°. Spine length vs. absolute cephalic size (SPST/CS): 0.278–0.331 (mean = 0.306). Minimum spine distance vs. absolute cephalic size (SPBA/CS): 0.273–0.312 (mean = 0.292). Maximum spine distance vs. absolute cephalic size (SPWI/CS): 0.353–0.415 (mean = 0.384). Apical spine distance vs. absolute cephalic size (SPTI/CS): 0.332–0.395 (mean = 0.364). Maximum mesosoma width vs. absolute cephalic size (MW/CS): 0.599–0.636 (mean = 0.618). Metanotal depression count: present. Metanotal depression shape: shallow. Dorsal region of mesosoma sculpture: rugulose with areolate ground sculpture. Lateral region of pronotum sculpture: areolate ground sculpture, main sculpture forked costate. Mesopleuron sculpture: areolate ground sculpture superimposed by dispersed rugulae. Metapleuron sculpture: areolate ground sculpture superimposed by dispersed rugulae. Frontal profile of petiolar node contour line in lateral view shape: concave. Dorsal profile of petiolar node contour line angle value to frontal profile of petiole contour line in lateral view: 105–110°. Anterodorsal rim of petiole count: absent medially.

Dorsal profile of petiolar node contour line in lateral view shape: slightly convex. Dorsal region of petiole sculpture: ground sculpture areolate, main sculpture dispersed rugose; ground sculpture areolate, main sculpture absent. Dorso-caudal petiolar profile contour line in lateral view shape: straight; concave. Dorsal region of postpetiole sculpture: ground sculpture areolate, main sculpture dispersed rugose; ground sculpture areolate, main sculpture absent.

References based on Global Ant Biodiversity Informatics

 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * Asociacion Iberica de Mirmecologia. 2012. List of species collected during the Taxomara Tres Cantos 2012. Iberomyrmex 4: 17-18.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
 * Baugnée J. Y. 2004. Observations récentes de la fourmi Leptothorax parvulus (Schenck, 1852) en Belgique (Hymenoptera Formicidae). Bulletin de la Société Royale Belge d'Entomologie 140: 101-103.
 * Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
 * Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:5971
 * Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
 * Bernard F. 1959. Les fourmis de l'île de Port-Cros. Contribution à l'écologie des anciennes forêts méditerranéennes. Vie Milieu 9: 340-360.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
 * Bernard F. 1976. Écologie des fourmis des grès d'Annot, comparées à celles de la Provence calcaire. Annales du Muséum d'Histoire Naturelle de Nice 3: 33-54.
 * Bernard F. 1978. Fourmis et milieu dans le massif des Maures. Vie et Milieu. Série C. Biologie Terrestre 27: 83-118.
 * Bernard, F. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
 * Berville L., M. Renucci, and E. Provost. 2012. Mise en place de protocoles de contrôle de la fourmi dArgentine (Linepithema humile) sur les îles de Port-Cros et de Porquerolles (Var, France). Sci. Rep. Port-Cros natl. Park, 26: 91-108.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bezdecka P., and K. Bezdeckova. 2012. Updated list of the ants of the Czech Republic. Pp 7-12. Bezd??ka P. & Bezd??ková K. (eds) 2012: Blanok?ídlí v ?eských zemích a na Slovensku 8, Chaloupky, 1.-3. ?ervna 2012, sborník abstrakt? z konference.  MVJ Jihlava, 37 pp.
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec M. L., and L. Borowiec. 2013. New data on the occurrence of ants (Hymenoptera: Formicidae) in Lower Silesia and other regions of Poland. Wiadomo?ci Entomologiczne 32: 49-57.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Buschinger A. 1975. Die ameisenfauna des Bausenberges, der nordostlichen Eifel und Voreifel (Hym., Formicidae) mit einer quantitativen Auswertung von Fallenfangen. Beitrage Landespflege Rhid-Pfalz Beiheft 4: 251-273.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse, J. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Colindre L. 2016. Les fourmis en Picardie: bilan des decouvertes et redecouvertes 2015 (Hymenoptera/ Formicidae). Entomologiste Picard 28: 8-10.
 * Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S, Heinze J, and I. Mikó. 2015. Taxonomic synopsis of the Ponto-Mediterranean ants of Temnothorax nylanderi species-group. PLoS ONE 10(11): e0140000. doi:10.1371/journal.pone.0140000
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Diniz, M. A. "Estado actual do conhecimento dos himenópteros de Portugal." Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 259 (1959): 1-42.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Espadaler X., F. Garcia, X. Roig, and R. Vila. 2013. Ants (Hymenoptera, Formicidae) from the Castell de Montesquiu park (Osona, north-east of the Iberian Peninsula). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 53: 223-227.
 * Espadaler X., M. Boieiro, R. Azedo, C. Bernardes, D. Figueiredo, and A. R. M. Serrano. 2008. Additions to the ant fauna of Portugal (Hymenoptera, Formicidae). Boletin Sociedad Entomologica Aragonesa 42: 349-351.
 * Espadaler, X., A. R. Azedo, C. Bernardes, D. Figueiredo, A. R. M. Serrano, and M. Boeiro. "Additions to the Ant fauna of Portugal (Hymenoptera: Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 349-351.
 * Foitzik S., and Heinze, J. 1999. Non-random size differences between sympatric species of the ant genus Leptothorax (Hymenoptera: Formicidae). Entomologia Generalis 24: 65-74.
 * Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
 * Forel, A. "Fourmis de Tunisie et de l'Algerie orientale recoltees et decrites par Auguste Forel." Annales de la Société Entomologique de Belgique 34 (1890): lxi-lxxvi.
 * Francois J. 1958. Contribution a l'etude ecologique des Formicides (Insectes, Hymenopteres) de la region Dijonnaise. Travaux du laboratoire de Zoologie et de la Station Aquicole Grimaldi de la Faculte des Sciences de Dijon 25, 39 pages.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Garcia F., X. Espadaler, and K. Gomez. 2009. First record of Amblyopone impressifrons (Emery, 1869) from the Iberian Peninsula and of Proceratium melinum (Roger, 1860) from Catalonia (Hymenoptera, Formicidae). Boletín Sociedad Entomológica Aragonesa 45: 357-360.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Ghahari H., and C. A. Collingwood. 2013. A study on the ants (Hymenoptera: Vespoidea: Formicidae) from Western Iran. Acta Phytopathologica et Entomologica Hungarica 48 (1): 155164.
 * Giacalone I., and M. Moretti. 2001. Contributo alla conoscenza della mirmecofauna (Hymenoptera: Formicidae) dei castagneti al Sud delle Alpi (ticino, Svizzera). Bollettino della Societa ticinese di Scienze naturali 89(1-2): 51-60.
 * Glaser F. 2004. Distribution and conservation of ants (Hymenoptera, Formicidae) in the floodplain of the river Etsch / Adige (Southern Tyrol, Italy). Gredleriana 4: 203-246.
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Kiran K. C. Karaman, A. Lapeva-Gjonova, and V. Aksoy. 2017. Two new species of the "ultimate" parasitic ant genus Teleutomyrmex KUTTER, 1950 (Hymenoptera: Formicidae) from the Western Palaearctic. Myrmecological News 25: 145-155.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
 * Legakis Collection Database
 * Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
 * Marko B. 2008. Ants (Hymenoptera: Formicidae) of the Sand Dunes of Foieni Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Mei, M. "Nuovi reperti de formicidi per l'Italia centrale (Hymenoptera, Formicidae)." Bollettino dell'Associazione Romana di Entomologia 37 (1984): 49-58.
 * Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
 * Meyer M., C. Braunert, R. Gerend, and I. Schrankel. 2003. Résultats de l'excursion annuelle du groupe de travail entomologique de la Société des naturalistes luxembourgeois. Bull. Soc. Nat. luxemb. 103: 103-120.
 * Moretti M. P. Duelli, and M. K. Obrist. 2006. Biodiversity and resilience of arthropod communities after fire disturbance in temperate forests. Oecologia 149: 312327.
 * Ottonetti L., L. Tucci, and G. Santini. 2006. Recolonization Patterns of Ants in a Rehabilitated Lignite Mine in Central Italy: Potential for the Use of Mediterranean Ants as Indicators of Restoration Processes. Restoration Ecology 14(1): 6066.
 * Paknia O., A. Radchenko, H. Alipanah, and M. Pfeiffer. 2008. A preliminary checklist of the ants (Hymenoptera: Formicidae) of Iran. Myrmecological News 11: 151-159.
 * Petrov I. Z. 2004. A list of currently known ant species (Formicidae, Hymenoptera) of Serbia. Arch. Biol. Sci., Belgrade 56(3-4): 121-125.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Plateaux L. 2000. Les fourmis de Lorraine. Bull. Soc. Ent. Lorr. 11-16.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
 * Salata S., P. Zurawlew, and J. K. Kowalczyk. 2018. New distribution records of several ant species (Hymenoptera: Formicidae) in Poland. Wiadomosci Entomologiczne 37(1): 46-53.
 * Salata S., and L. Borowiec. 2014. New localities of some rare ant species (Hymenoptera: Formicidae). WIAD. ENTOMOL., 33 (1): 77-79.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Schmitz H. 1950. Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit. Brotéria Ser. Cienc. Nat. 19: 12-16.
 * Schmitz, H. "Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit." Brotéria, Ciências Naturais 19 (1950): 12-16.
 * Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
 * Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
 * Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Tausan I. 2014. Temnothorax parvulus (Schenck, 1852)(Hymenoptera: Formicidae) in Romania. Brukenthal, Acta Musei 9: 511-514.
 * Vogrin, V. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Wegnez P. 2014. Premières captures de Lasius distinguendus Emery, 1916 et de Temnothorax albipennis (Curtis, 1854) au Grand-Duché de Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société royale belge d’Entomologie 150 (2014) : 168-171.
 * Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
 * Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
 * Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.