Formica fusca

In Korea this species is common in mountainous highlands of more than 1000 meters (Dong & Kim, 2019).

Identification
Black, legs brownish. Gula, occiput, mid femora and promesonotum without standing hairs - occasionally one or two weak pronotal hairs. Pubescent hairs on gaster longer than their interspace width. Frons with fine microsculpture. Length: 4.5-7.0 mm.

Mackay and Mackay (2002) - Metasternal lobe is absent or very poorly developed and usually has erect hairs around the metasternal cavity. It is concolorous black with abundant appressed pilosity. The underside of the head has no erect hairs, and the first tergite of the gaster (excluding the posterior edge) has only about 4 course, erect hairs.

Distribution
Holarctic.

Distribution based on Regional Taxon Lists
Nearctic Region: Canada. Oriental Region: India, Nepal, Pakistan. Palaearctic Region: Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Isle of Man, Latvia, Lithuania, Luxembourg, Malta, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland.

Foraging/Diet
Formica fusca collect honeydew. Novgorodova (2015b) investigated ant-aphid interactions of a dozen honeydew collecting ant species in Western Siberia pine and aspen-birch-pine forests (54°7´N, 83°06´E, 200 m, Novosibirsk) and mixed-grass-cereal steppes with aspen-birch groves (53°44´N, 78°02´E, 110 m, near Karasuk) in the Novosibirsk Region and coniferous forests in the northeastern Altai (north end of Lake Teletskoe, 51°48´N, 87°17´E, 434 m). All of the ants studied had workers that showed high fidelity to attending particular aphid colonies, i.e, individual foragers that collect honeydew tend to return to the same location, and group of aphids, every time they leave the nest. F. fusca showed no specialization beyond this foraging site fidelity. F. fusca tended Symydobius oblongus (Heyden), Chaitophorus populeti (Panzer) and Aphis craccivora Koch.

Europe
This is the common black ant of Europe. It nests variously in banks, under stones and in tree stumps along hedgerows and woodland borders. Workers are timid, fast moving and forage singly, predating small insects but also feeding on extra floral nectaries and on aphid honeydew. Colonies are usually small with up to 500 workers and one or a few queens present. Alatae are developed in June and July and fly off the nests in July and early August (Collingwood 1979).

This species is known to suffer from labial gland disease, a condition caused by an, in Spain (Espadaler & Riasol, 1981; Elton, 1991).

Other Insects

 * Host for the slave-making ants:
 * (Ruano et al., 2019; Seifert, 2018; de la Mora et al., 2021)
 * (Trager, 2013; Sulger et al., 2014; de la Mora et al., 2021)


 * Host for the temporary parasites:



Fungi
This ant is a host for the fungi , ,  (Espadaler & Santamaria, 2012) and (Małagocka et al., 2017).

Castes
Queens of different fecundity levels differed in their Cuticular Hydrocarbons profiles. This was associated with worker behaviour; the higher a queen’s fecundity, the more attention she received from workers (Hannonen et al. 2002).

Nomenclature

 *  fusca. Formica fusca Linnaeus, 1758: 580 (w.) EUROPE.
 * Latreille, 1802c: 159 (q.m.)
 * Wheeler, G.C. & Wheeler, J. 1953c: 164 (l.).
 * Combination in F. (Serviformica): Forel, 1913i: 361.
 * Senior synonym of barbata: Emery & Forel, 1879: 451; of flavipes, libera: Latreille, 1802c: 159; Smith, F. 1851: 3; Roger, 1863b: 13; of pallipes, rufipes: Dlussky, 1967a: 58; of tristis: Emery, 1892b: 162; of glebaria: Mayr, 1855: 346; Yarrow, 1954a: 230; Dlussky, 1967a: 58; Radchenko, 2007: 35.
 * Current subspecies: nominal plus alpicola, fuscolemani, hyrcana, maura, tombeuri.
 * See also: Emery, 1893i: 657; Wheeler, W.M. 1913f: 494; Donisthorpe, 1915d: 304; Stitz, 1939: 348; Dlussky, 1967a: 58; Dlussky & Pisarski, 1971: 148; Francoeur, 1973: 189; Tarbinsky, 1976: 180; Collingwood, 1979: 120; Kupyanskaya, 1990: 183; Atanassov & Dlussky, 1992: 260; Seppä, et al. 2011: 31.
 * Status as species: Schar et al., 2018: 6.
 * libera. Formica libera Scopoli, 1763: 313 (w.) AUSTRIA. Junior synonym of fusca: Latreille, 1802a: 159.
 * flavipes. Formica flavipes Geoffroy, in Fourcroy, 1785: 452 (m.) FRANCE. Junior synonym of fusca: Latreille, 1802c: 161.
 * barbata. Formica barbata Razoumowski, 1789: 225, fig. 12 (w.) SWITZERLAND. Junior synonym of fusca: Emery & Forel, 1879: 451.
 * tristis. Formica tristis Christ, 1791: 513, pl. 60, fig. 13 (m.) no locality given. Junior synonym of fusca: Emery, 1892b: 162.
 * glebaria. Formica glebaria Nylander, 1846a: 917, pl. 18, fig. 23 (w.q.) FINLAND. Foerster, 1850a: 31 (m.). Combination in F. (Serviformica): Forel, 1915d: 63. Subspecies of fusca: Emery & Forel, 1879: 451; Wheeler, W.M. 1908g: 408; Emery, 1909b: 196; Ruzsky, 1914b: 104; Forel, 1915d: 63; Emery, 1916b: 254; Santschi, 1925g: 353; Karavaiev, 1927c: 286; Stitz, 1939: 353; Novak & Sadil, 1941: 108; of rufibarbis: Karavaiev, 1936: 237. Status as species: Bondroit, 1918: 49; Müller, 1923: 139; Boven, 1947: 188. Junior synonym of cunicularia: Bernard, 1967: 296; Boven, 1977: 164; Agosti & Collingwood, 1987a: 59; of fusca: Mayr, 1855: 346; Yarrow, 1954a: 230; Dlussky, 1967a: 58; Radchenko, 2007: 35.
 * pallipes. Formica fusca var. pallipes Kuznetsov-Ugamsky, 1926b: 97 (w.) KAZAKHSTAN. [Unresolved junior primary homonym of pallipes Latreille, above.] Junior synonym of fusca: Dlussky, 1967a: 58; Tarbinsky, 1976: 180.
 * rufipes. Formica fusca var. rufipes Stitz, 1930: 238 (w.q.) RUSSIA. [Unresolved junior primary homonym of rufipes Fabricius, 1775: 391, above.] Junior synonym of fusca: Dlussky, 1967a: 58.

Taxonomic Notes
Dong & Kim (2019): This species was previously known as Formica lemani in Korea but we treated this species as a misidentication of Formica fusca by the following characteristics: 1. No more than 6 erect hairs (Radchenko, 2005) on the pronotum of worker. 2. Workers with less than 3 hairs in mid femur. However, according to Dlussky and Pisarski (1971), there are some variable individuals without any setae on the pronotum, which requires close examination.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Aktaç, N.. "Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon." Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41 (1977): 115-135.
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * Alvarado M., and L. Galle. 2000. Ant assemblages associated with lowland forests in the southern part of the great Hungarian plain. Acta Zoologica Academiae Scientarum Hungaricae 46(2): 79-102.
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antonov I. A. 2012. Ant complexes of Baikalsk town. The Bulletin of Irkutsk State University 4: 143-146.
 * Antonov I. A. 2013. Ant Assemblages (Hymenoptera: Formicidae) of Cities of the Temperate Zone of Eurasia. Russian Journal of Ecology 44(6): 523526.
 * Antonov I. A., and A. S. Pleshanov. 2011. Ecological-geographical features of Myrmicafauna of Baikal Region. Bulletin of the Buryat State. Univ. 4: 104-108.
 * Arakelian G. R., S. O. Movsessian, and F. A. Chubarian. 1997. Ecological and faunistic review of ants (Hymenoptera: Formicidae) - Supplementary hosts of the trematode Dicrocoelium lanceatum in Armenia. Parasitologia 31(3): 239-244.
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * ArtDatabanken Bugs (via GBIG)
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2011. Iberomyrmex 3: 30-31.
 * Asociacion Iberica de Mirmecologia. 2012. List of species collected during the Taxomara Tres Cantos 2012. Iberomyrmex 4: 17-18.
 * Asociacion Iberica de Mirmecologia. 2014. List of species collected during the Taxomara 2014 Oviedo. Iberomyrmex 6: 23-24.
 * Assing V. 1989. Die Ameisenfauna (Hym.: Formicidae) nordwestdeutscher Calluna-Heiden. Drosera 89: 49-62.
 * Azcarate F. M., and B. Peco. 2012. Abandonment of grazing in a mediterranean grassland area: consequences for ant assemblages. Insect Conservation and Diversity 5: 279288.
 * Azuma M. 1955. A list of ants (Formicidae) from Hokkaido Is. Hyogo Biology 3:79-80.
 * Banert P, and B. Pisarski. 1972. Mrówki (Formicidae) Sudetów. Fragmenta Faunistica (Warsaw) 18: 345-359.
 * Bargum, K. and L. Sundstrom. 2007. Multiple Breeders, Breeder Shifts and Inclusive Fitness Returns in an Ant. Proceedings: Biological Sciences 274(1617):1547-1551
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
 * Baroni Urbani, C.. "Formiche dell'Italia appenninica (Studi sulla mirmecofauna d'Italia, III)." Memorie del Museo Civico di Storia Naturale di Verona 12 (1964): 149-172.
 * Baroni-Urbani 1969. Ant Communities of the High-Altitude Appennine Grasslands. Ecology 50(3): 488-492.
 * Barrett K. E. 1967. Ants in South Brittany. Entomologist's Record and Journal of Variation 79:112-116.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
 * Behr D., S. Lippke, and K. Colln. 1996. Zur kenntnis der ameisen von Koln (Hymenoptera, Formicidae). Decheniana-Beihefte (Bonn) 35: 215-232.
 * Behr D., and K. Colln. 1993. Zur ameisenfauna (Hymenoptera, Formicidae) von Gonnersdorf (Kr. Daun). Dendrocopos 20: 148-160.
 * Belskaya E., A. Gilev, and E. Belskii. 2017. Ant (Hymenoptera, Formicidae) diversity along a pollution gradient near the Middle Ural Copper Smelter, Russia. Environ Sci Pollut Res DOI 10.1007/s11356-017-8736-8
 * Bernadou A., G. Latil, V. Fourcassié, and X. Espadaler. 2006. Etude des communautés de fourmis d'une vallée andorrane. Union International pour l'Etude des Insectes Sociaux, Colloque annuel de la section francaise, 4pp.
 * Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
 * Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:5971
 * Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
 * Bernard F. 1960. Fourmis récoltées en Corse par J. Bonfils (1957). Compte Rendu Sommaire des Séances de la Société de Biogéographie 36: 108-114.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
 * Bernard F. 1976. Écologie des fourmis des grès d'Annot, comparées à celles de la Provence calcaire. Annales du Muséum d'Histoire Naturelle de Nice 3: 33-54.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bezdeckova K., and P. Bezdecka. 2009. Nejvetsi polykalicka kolonie Formica foreli (Hymenoptera: Formicidae) y Ceské republice. Acta rerum naturalium 7: 121126.
 * Bharti H., Y. P. Sharma, M. Bharti, and M. Pfeiffer. 2013. Ant species richness, endemicity and functional groups, along an elevational gradient in the Himalayas. Asian Myrmecology 5: 79-101.
 * Blacker N. C. 1989. The ants (Hymenoptera, Formicidae) of the Gower Peninsula, West Glamorgan, South Wales. Entomologist's Record and Journal of Variation 101: 261-266.
 * Blacker N. C. and C. A. Collingwood. 2002. Some significant new records of ants (Hymenoptera: Formicidae) from the Salisbury area, south Wiltshire, England, with a key to the British species of Lasius. British Journal of Entomology and Natural History 15: 25-46
 * Blacker N.C. 2007. Ants (Hym., Formicidae) in East Anglia-Additional Records from . Entomologist's Monthly Magazine 143: 69-90
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
 * Boer P., and J. Noordijk. 2004. De ruige gaststeekmier Myrmica hirsuta nieuw voor Nederland (Hymenoptera: Formicidae). Ned. Faun. Meded. 20: 25-32.
 * Bonaric J. C. 1971. Contribution a l'etude systematique et ecologique des formicides du Bas-Languedoc. PhD thesis Universite des sciences et techniques du Languedoc, 175 pages.
 * Bonaric J. C. 1971. Étude systématique et ecologique des fourmis de lHérault (suite). Ann. Soc. Hortic. Hist. Nat. Hérault 111: 119-126.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
 * Boven J. van 1949. Notes sur la faune des Hautes-Fagnes en Belgique. Bulletin et Annales de la Société Entomologique de Belgique 85: 135-143.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Bujan J., A. Brigic, Z. Sedlar, and R. Sostaric. 2015. Progressive vegetation succession of fen habitats promotes the lack of habitat specialist ants. Insectes Sociaux 62: 415-422.
 * Buschinger A. 1975. Die ameisenfauna des Bausenberges, der nordostlichen Eifel und Voreifel (Hym., Formicidae) mit einer quantitativen Auswertung von Fallenfangen. Beitrage Landespflege Rhid-Pfalz Beiheft 4: 251-273.
 * Cagniant, H. 1968. Liste preliminaire de fourmis forestieres d'Algerie. Resultats obtenus de 1963 a 1964. Bulletin de la Société d'Histoire Naturelle de Toulouse 104: 138-147
 * Cagniant, H.. "Deuxième liste de fourmis d'Algérie, récoltées principalement en forêt (Deuxième partie)." Bulletin de la Société d'Histoire Naturelle de Toulouse 106 (1970): 28-40.
 * Cagniant, H.. "Étude de fourmis récoltées par le Professeur H. Janetschek dans la Sierra Nevada." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 52 (1961): 104-117.
 * Cagniant, H.. Les peuplements de fourmis des forêts algériennes: écologie, biocénotique, essai biologique. Universite de Toulouse, 1973.
 * Cammell, M. E., M. J. Way, and M. R. Paiva. "Diversity and structure of ant communities associated with oak, pine, eucalyptus and arable habitats in Portugal." Insectes Sociaux 43 (1996): 37-46.
 * Carniel A. 1988. Contributo alla conoscenza della mirmecofauna del Cansiglio (Prealpi Carniche). Boll. Soc. Ent. Ital., Genova 119(3): 179-190.
 * Carniel A. 1998. Ricerche sulla mirmecofauna delle Prealpi Orobiche (Lombardia) (Insecta, Hymenoptera, Formicidae). Atti. Mus. Civ. Stor. Nat. Morbegno 9: 29-39.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Chen Y., C. W. Luo, H. W. Li, Y. J. Liu, H. F. Zheng, and F. C. Yang. 2013. Investigation of ant species and distribution on Wuliang Mountain. Journal of Henan Agricultural Sciences 42(5): 118-122.
 * Cherix D., and S. Higashi. 1979. Distribution verticale des fourmis dans le Jura vaudois et recensement prelimaire des bourdons (Hymenoptera, Formicidae et Apidae). Bull. Soc. Vaud. Sc. Nat. 356(74): 315-324.
 * Choi B.M., Kim, C.H., Bang, J.R. 1993. Studies on the distribution of ants (Formicidae) in Korea (13). A checklist of ants from each province (Do), with taxonomic notes. Cheongju Sabom Taehakkyo Nonmunjip (Journal of Cheongju National University of Education) 30: 331-380.
 * Colindre L. 2015. Les fourmis en Picardie: bilan 2014 (Hymenoptera/ Formicidae). Entomologiste Picard 26, 15 pages.
 * Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
 * Collingwood C. A. 1951. The distribution of ants in north-west Scotland. Scottish Naturalist 63: 45-49
 * Collingwood C. A. 1951. The distribution of ants in north-west Scotland. Scottish Naturalist 63: 45-49.
 * Collingwood C. A. 1955. Ants in S.W. Scotland. Entomologist's Record and Journal of Variation 67: 11-12.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1961. Ants in the Scottish Highlands. Scotish Naturalist 70: 12-21.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood C. A., and J. E. Satchell. 1956. The ants of the South Lake District. Journal of the Society for British Entomology 5: 159-164.
 * Collingwood C. A., and J. Hughes. 1987. Ant species in Yorkshire, England. Naturalist (Leeds) 112: 95-101.
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood C., and H. Heatwole. 2000. Ants from Northwestern China (Hymenoptera, Formicidae). Psyche 103 (1-2): 1-24.
 * Collingwood C.A. 1955. Ants in S.W. Scotland. Entomol.Rec. 67: 11-12
 * Collingwood C.A. 1959. Ants in the Scottish Highlands. The Scottish Naturalist. 70: 12-21
 * Collingwood C.A. 1961. Ants in Finland. Entomol. Rec. 73: 190-195
 * Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
 * Collingwood C.A. 1970. Formicidae (Hymenopter: Aculeata) of Nepal. Himalaya Khumbu Himal, 3: 371-388.
 * Collingwood C.A. and Satchell J.E. 1956. The Ants of the South Lake District. Journal of the Society for British Entomology. 5: 159-164
 * Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
 * Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
 * Collingwood, C. A. 1974. A revised list of Norwegian ants (Hymenoptera: Formicidae). Norsk Entomologisk Tidsskrift 21: 31-35.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Collingwood, C. A.. "A provisional list of Iberian Formicidae with a key to the worker caste." EOS (Revista española de entomología) Nº LVII (1978): 65-95.
 * Collingwood, C. A.. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Collingwood, C.A. 1958. A survey of Irish Formicidae. Proceedings of the Royal Irish Academy 59B:213-219
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S., B. Markó, and L. Gallé. 2001. Ants (Hymenoptera: Formicidae) of Stana Valley (Romania): Evaluation of the effectiveness of a myrmecological survey.  Entomologica Romanica 6 : 121-126.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Cuní, M.. "Excursión entomológica a varias localidades de la provincia de Gerona (Cataluña)." Anales de la Sociedad española de Historia Natural (1885): 51.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Czechowski W., B. Marko, A. Radchenko, and P. Slipinski. 2013. Long-term partitioning of space between two territorial species of ants (Hymenoptera: Formicidae) and their effect on subordinate species. Eur. J. Entomol. 110(2): 327337.
 * Czechowski, W. 2005. Integration raids in the Amazon ant Polyergus rufescens (Hymenoptera, Formicidae). Insectes Sociaux 52:103-104
 * Czechowski, W., and A. G. Radchenko. "Formica lusatica Seifert, 1997 (Hymenoptera: Formicidae), an ant species new to Finland, with notes on its biology and the description of males." Myrmecologische Nachrichten 8 (2006): 257-262.
 * Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
 * Dauber J., and D. Simmering. 2006. Ant assemblages in successional stages of Scotch Broom stands (Hymenoptera: Formicidae; Spermatophyta). Myrmecologische Nachrichten 9: 55-64.
 * Dauber, J., J. Bengtsson and L. Lenoir. 2006. Evaluating Effects of Habitat Loss and Land-Use Continuity on Ant Species Richness in Seminatural Grassland Remnants. Conservation Biology 20(4):1150-1160
 * De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
 * Dekoninck W., H. De Koninck, J. Y. Baugnee, and J. P. Maelfait. 2007. Ant biodiversity conservation in Belgian calcareous grasslands: active management is vital. Belg. J. Zool. 137 (2): 137-146.
 * Dekoninck W., K. Desender, and P. Grootaert. 2008. Establishment of ant communities in forests growing on former agricultural fields: Colonisation and 25 years of management are not enough (Hymenoptera: Formicidae). Eur. J. Entomol. 105: 681689.
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Dewes E. 2005. Ameisenerfassung im Waldschutzgebiet Steinbachtal/Netzbachtal. Abh. Delattinia 31: 89-118.
 * Diniz, M. A.. "Estado actual do conhecimento dos himenópteros de Portugal." Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 259 (1959): 1-42.
 * Donisthorpe H. 1914. Myrmecophilous notes for 1913. Entomologist's Record and Journal of Variation 26: 37-45.
 * Donisthorpe, H.. "A first instalment of the ants of Turkey." Annals and Magazine of Natural History (12)3 (1950): 1057-1067.
 * Downing H., and J. Clark. 2018. Ant biodiversity in the Northern Black Hills, South Dakota (Hymenoptera, Formicidae). Journal of the Kansas Entomological Society 91(2): 119-132.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Dusmet, J. M.. "Algunos insectos (la mayor parte himenópteros) cazados en Cataluña en 1925." Boletín de la Sociedad Entomológica de España (1927): 21.
 * Dusmet, J. M.. "Linneo y los himenópteros de España." Linneo en España. Homenaje a su segundo centenario Zaragoza (1907): 475.
 * Dvorak, L., P. BOGUSCH, I. MALENOVSKÝ, P. BEZDÌÈKA, K. BEZDÌÈKOVÁ, K. HOLÝ, P. LIKA, J. MACEK, L. ROLLER, M. RÍHA et al. "Hymenoptera of Hády Hill, near the city of Brno (Czech Republic), collected during the Third Czech-Slovak Hymenoptera meeting." Acta Musei Moraviae, Scientiae biologicae (Brno) 93 (2008): 53-92.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Emery C. 1869. Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli con descrizioni di specie nuove o meno conosciute. Ann. Accad. Aspir. Nat. Secunda Era 2: 1-26.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Emery, C.. "Beiträge zur Kenntniss der palaearktischen Ameisen." Öfversigt af Finska Vetenskaps-Societetens Förhandlingar (Helsinki) 20 (1898): 124-151.
 * Emery, C.. "Spicilegio mirmecologico." Bollettino della Societa Entomologica Italiana 33 (1901): 57-63.
 * Espadaler X., X. Roig, K. Gómez, and F. García. 2011. Formigues de les Planes de Son i mata de València (Hymenoptera, Formicidae) Treballs de la Institució Catalana d'Història Natural 16: 609-627.
 * Espadaler, X., and C. Ascaso. "Adición a las hormigas del Montseny (Barcelona)." Orsis 5 (1990): 141-147.
 * Espadaler, X., and J. M. Riasol Boixart. "Secretergates de Formica sp.: una morfologia de origen patologico en hormigas." Revista Ibérica de Parasitologia 41 (1981): 539-549.
 * Espadaler, X., and X. Roig. "Hormigas del Vall de Aneu." (2007).
 * Espadaler, X.. "Contribución al conocimiento de los formícidos (Hymenoptera, Formicidae) del Pirineo catalán." Tesis Universida (1979): 285 pp.
 * Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants  assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Finzi, B.. "Risultati scientifici della spedizione Ravasini-Lona in Albania. III. Formiche." Bollettino della Società Entomologica Italiana 55 (1923): 1-4.
 * Finzi, B.. "Secondo contributo alla conoscenza della fauna mirmecologica della Venezia Giulia." Bollettino della Società Entomologica Italiana 56 (1924): 120-123.
 * Forel A. 1890. Fourmis de Tunisie et de l'Algérie orientale. Annales de la Société Entomologique de Belgique 34: lxi-lxxvi.
 * Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
 * Forel A. 1906. Fourmis d'Asie mineure et de la Dobrudscha récoltées par M. le Dr. Oscar Vogt et Mme Cécile Vogt, Dr. méd. Ann. Soc. Entomol. Belg. 50: 187-190.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
 * Forel, A.. "Fourmis d'Espagne récoltées par M. O. Vogt et Mme Cécile Vogt, Docteurs en médecine." Annales de la Société Entomologique de Belgique 53 (1909): 103-106.
 * Forel, A.. "Südpalaearctische Ameisen." Mitteilungen der Schweizerischen Entomologischen Gessellschaft 9 (1895): 227-234.
 * Formidabel Database
 * Fowles, A.P. 1996. A provisional checklist of the invertebrates recorded from Wales. 2. Aculeate wasps, bees and ants (Hymenoptera: Aculeata). Countryside Council for Wales
 * Franch, J., and X. Espadaler. "Ants as colonizing agents of pine stumps in San Juan de la Peña (Huesca, Spain)." Vie et Milieu 38 (1988): 149-154.
 * Francoeur. A. 1973. Revision taxonomique des especes nearctiques du group fusca, genre Formica. Memoires de la Societe Entomologique du Quebec 3: 1-316.
 * Francois J. 1958. Contribution a l'etude ecologique des Formicides (Insectes, Hymenopteres) de la region Dijonnaise. Travaux du laboratoire de Zoologie et de la Station Aquicole Grimaldi de la Faculte des Sciences de Dijon 25, 39 pages.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Gadeau de Kerville H. 1922. Materiaux pour la Faune des Hymenopteres de la Normandie. Bull. Soc. Amis Sc. Nat. Rouen 1916-1921, 1922: 217-225.
 * Galkowski C. 2011. Une liste des fourmis (Hymenoptera, Formicidae) récoltées dans la région de Grasse, avec la mention dune nouvelle espèce de la faune de France. Bulletin de la Société linnéenne de Provence, 62 : 41-44.
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Lebas. 2016. Temnothorax conatensis nov. sp., décrite des Pyrenees-Orientales (France) (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 80-87.
 * Galkowski C., and P Wegnez. 2010. Myrmica constricta Karavaiev 1934, nouvelle espece pour la France (Hymenoptera, Formicidae). Bull. Soc. Ent. Mulhouse 66(3): 41-45.
 * Galle L. 1972. Study of ant-populations in various grassland ecosystems. Acta Biologica Szeged 18(1-4): 159-164.
 * Galle L. 1973. Formicidae populations of the ecosystems in the environs of Tiszafured. Tiscia, Szeged 7: 59-68.
 * Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
 * Galle L. 1993. Data to the ant fauna of the Bukk (Hymenoptera: Formicidae). Natural history of the national parks of Hungary 7: 445-448.
 * Galle L. 1997. Contribution to the ant fauna of Slovenia with special reference to the submediterranean and eudinaric regions. Annals for Istrian and Mediterranean studies 11: 209-214.
 * Galle L., and G. Szonyi. 1988. A check list of ants (Hymenoptera: Formicoidea) of a sandy grassland in Kiskunsag National Park (Hungary). Acta Biol. Szeged 34: 167-168.
 * Gallé L. 1990. Assembly of sand-dune forest ant communities. Memorabilia Zoologica 44: 1-6.
 * Gallé L. 1991. Structure and succession of ant assemblages in a north European sand dune area. Holarctic Ecology 14: 31-37.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Garcia F, Arnal J, Espadaler X. 2008. Primeros registros de Myrmica bibikoffi Kutter, 1963 (Hymenoptera: Formicidae) en la Península Ibérica. Heteropterus Revista de Entomologia 8: 211-215.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gaspar C. 1966. Etude myrmecologique des tourbieres dans les Hautes-Fagnes en Belgique (Hymenoptera, Formicidae). Rev. Ecol. Biol. Sol. 3(2): 301-312.
 * Gaspar C. 1966. Etude myrmecologique des tourbieres dans les Huates-Fagnes en Belgique (Hymenoptera, Formicidae). Rev. Ecol. Biol. Sol. 3(2): 301-312.
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Gaspar C., and C. Thirion. 1978. Modification des populations d'Hymenopteres sociaux dans les milieux anthropogenes. Memorabilia Zoologica 29: 61-77.
 * Gaspare Charles. 1965. Étude myrmécologique d'une région naturelle de Belgique: la Famenne Survey des Fourmis de la Région (Hymenoptera, Formicidae). Institut agronomique de l'Etat a' Gembloux. 32(4): 427-434.
 * Giacalone I., and M. Moretti. 2001. Contributo alla conoscenza della mirmecofauna (Hymenoptera: Formicidae) dei castagneti al Sud delle Alpi (ticino, Svizzera). Bollettino della Societa ticinese di Scienze naturali 89(1-2): 51-60.
 * Gilev A. V., I. V. Kuzmin, V. A. Stolbov, and S. D. Sheikin. 2012. Materials on the fauna and ecology of ants (formicidae) Southern part of the Tyumen region. Tyumen State University Herald 6: 86-91.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Glaser F., A. Freitag, and H. Martz. 2012. Ants (Hymenoptera: Formicidae) in the Münstertal (Val Müstair)  a hot spot of regional species richness between Italy and Switzerland. Gredleriana 12: 273 - 284.
 * Glaser F., T. Kopf, and K. H. Steiberger. 2003. Ameisen (Hymenoptera: Formicidae) im Frastanzer Ried und den Illauen (Vorarlberg, Österreich) Artenspektrum, Gefährdung und Schutzempfehlungen. Vorarlberger Naturschau 13: 287-310.
 * Goetsch, W.. "Beiträge zur Biologie spanischer Ameisen." EOS (Revista española de entomología) 18 (1942): 175-241.
 * Gomez K., D. Martinez, and X. Espadaler. 2018. Phylogeny of the ant genus Aphaenogaster (Hymenoptera: Formicidae) in the Iberian Peninsula, with the description of a new species. Sociobiology 65: 215-224.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Groc S., J. H. C. Delabie, R. Cereghino, J. Orivel, F. Jaladeau, J. Grangier, C. S. F. Mariano, and A. Dejean. 2007. Ant species diversity in the Grands Causses (Aveyron, France): In search of sampling methods adapted to temperate climates. C. R. Biologies 330: 913922.
 * Grzes I. M. 2009. Ant species richness and evenness increase along a metal pollution gradient in the Boles?aw zinc smelter area. Pedobiologia 53: 65-73.
 * Gulzar A. 2014. Classification and Distribution of ants in Kashgar, Xinjiang. Master's Thesis Shaanxi Normal University, 75 pages.
 * Guo X., Q. Lin, J. Cui, D. Gao, S. Xu, and Z. Sheng. 2014. Biodiversity and spatial distribution patterns of ant species in tea gardens of Chongqing. Chinese Journal of Eco-Agriculture DOI: 10.3724/SP.J.1011.2014.31052
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Haguet G., M. Chevrier, and E. Brunel. 2002. Les invertebres de la dune de Bon Abri, premier inventaire. Groupe d'Etude des Invertebres Armoricains, 37 pages.
 * Hayashida K. 1959. Ecological Distribution of Ants in Mt. Atusanupuri, An Active Volcano in Akan National Park, Hokkaido. Jour. Pac. Sci. Hokkaiao Univ. Ser. 4(14): 252-260.
 * Hayashida K. 1959. Ecological distribution of ants in Mt. Atusanupuri, an active volcano in Akan National Park, Hokkaido. Journal of the Faculty of Science, Hokkaido University. Series VI. Zoology 14:252-260.
 * Hayashida K. 1964. Studies on the ecological distribution of ants in Kutchan and its adjacent area. Journal of the Sapporo Otani Junior College 2: 107-129.
 * Hayashida K., and S. Maeda. 1960. Studies on the ecological distribution of ants in Akkeshi. Journ. Sc. Hokkaido Univ., IV. Zool., 14 (3) : 305-319.
 * Heinken, T. 2004. Migration of an Annual Myrmecochore: A Four Year Experiment with Melampyrum pratense L.. Plant Ecology 170(1):55-72
 * Helantera, H. and F.L.W. Ratnieks. 2009. Two independent mechanisms of egg recognition in worker Formica fusca ants. Behavioural Ecology and Sociobiology 63:573-580
 * Helantera, H. and L. Sundstrom. 2007. Worker policing and nest mate recognition in the ant Formica fusca. Behavioral Ecology and Sociobiology 61(8):1143-114
 * Helantera, H., S.J. Martin and F.L.W. Ratnieks. 2007. Prior Experience with Eggs Laid by Non-Nestmate Queens Induces Egg Acceptance Errors in Ant Workers. Behavioral Ecology and Sociobiology 62(2):223-228.
 * Hernandez Cuba O., C. Perez Banon, and M. A. Marcos Garcia. 2008. Los formicidos (Hymenoptera, Formicidae) del Parque natural de la Font Roja. Iberis 6: 9-20.
 * Hernandez Cuba O., J. L. Perez-Bote, and J. M. Garcia Jimenez. 2006. Los formicidos (Hymenoptera: Formicidae) del Parque Natural de Cornaldo (suroeste de la Peninsula Iberica). Boletin Sociedad Entomologica Aragonesa 38: 356-358.
 * Holgersen H. 1942. Ants of northern Norway (Hym., Form.). Tromso Mus. Årsh. 63(2): 1-34.
 * Holgersen H. 1943. Ant studies in Rogaland (south-western Norway). Avhandlingar utgitt av det Norske Videnskaps-Akademi i Oslo. I. Matematisk-Naturvidenskapelig Klasse 1943(7): 1-75.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-203.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-493.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-506.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-518.
 * Hosoishi, S., S. Yamane and K. Ogata. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13:345-350.
 * Hågvar S. 2005. Altitudinal zonation of ants (Formicidae) in a steep fjord landscape in Sogndal, Western Norway. Norw. J. Entomol. 52: 3-12.
 * Ihnatiuk O. A., and S. V. Stukalyuk. 2015. Degradation changes in the structure of multispecies associations of ants in urbanized areas. Russian Journal of Ecology 46(1): 109–115.
 * Jakubzik A., H. Kinkler, and K. Colln. 2010. Aculeate Hymenoptera from a Humid Biotope in Leverkusen-Steinbüchel. Decheniana (Bonn) 163: 145158.
 * Jeffery H. G. 1931. The Formicidae (or ants) of the Isle of Wight. Proceedings of the Isle of Wight Natural History and Archaeological Society 2: 125-128.
 * Kanizsai O., R. Gallé, and L. Gallé. 2009. Perception of spatial patchiness by ant assemblages (Hymenoptera: Formicidae). Tiscia 37: 3-7.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karavaiev V. 1912. Ameisen aus dem paläarktischen Faunengebiete. Rus. Entomol. Obozr. 12: 581-596.
 * Karavaiev V. 1926. Beiträge zur Ameisenfauna des Kaukasus, nebst einigen Bemerkungen über andere palaearktische Formen. (Schluss). Konowia 5: 187-199.
 * Karavaiev V. 1927. Übersicht der Ameisenfauna der Krim nebst einigen Neubeschreibungen. Konowia 5: 281-303.
 * Karavaiev V. 1931. Beitrag zur Ameisenfauna Jakutiens. (Auf Grund der Sammelergebnisse der Expeditionen der Wissenschaften der UdSSR., ausgeführt in den Jahren 1925 und 1926.). Zool. Anz. 94: 104-117.
 * Kim B.J. 1996. Synonymic list and distribution of Formicidae (Hymenoptera) in Korea. Entomological Research Bulletin Supplement 169-196.
 * Kim B.J., K.G. Kim, D.P. Ryu, J.H. Kim. 1995. Ants of Chindo island in Korea (Hymenoptera; Formicidae). The Korean Journal of Systematic Zoology 11(1): 101-113.
 * Kim et al. 1993. Systematic study of ants from Chejudo Province. Koran Journal of Entomology 23(3): 117-141.
 * Kim, Byung-Jin, Ky-Gyong Kim, Dong-Pyo Ryu and Joong-Hyon Kim. 1995. The Korean Journal of Systematic Zoology. 11(1):101-113.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Kozisek T. 1987. Ants (Formicoidea) of the Abrod State Nature Reserve. Ochrana Prírody 8: 205-208.
 * Kratochvíl J., V. Novák, and J. Snoflák. 1944. Mohelno. Soubor práci venoványch studiu vyznamne památky prírodní. 5. Hymenoptera - Aculeata. Formicidae - Apidae - Vespidae. Arch. Svazu Ochr. Prír. Domov. Moravé 6: 1-155.
 * Krausse, A. H.. "Ueber sardische Ameisen." Archiv für Naturgeschichte, Berlin A 78(7) (1912): 162-166.
 * Kvamme T. 1982. Atlas of the Formicidae of Norway (Hymenoptera: Aculeata). Insecta Norvegiae 2: 1-56.
 * Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur  Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
 * Laeger T., and R. Schultz. 2005. Ameisen (Hymenoptera: Formicidae) als Beifänge in Bodenfallen  wie genau spiegeln sie reale Abundanzverhältnisse wider? Myrmecologische Nachrichten 7: 17-24.
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Le Moli F., and A. Zaccone. 1995. Ricerche sulla mirmecofauna del Cansiglio (Prealpi Carniche). Soc. Ven. Sc. Nat. 20: 33-52.
 * Le Moli F., and M. R. Rose. 1991. Atti della Societa Italiana di Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milan. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 132:25-40.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis A. 1983. The Zoological Museum of the University of Athens 2. The collection of ants from Greece. Biologia Gallo-Hellenica 11(1): 85-87.
 * Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
 * Legakis Collection Database
 * Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
 * Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
 * Liu C., K. L. Dudley, Z. Xu, and E. P. Economo. 2017. Mountain metacommunities: climate and spatial connectivity shape ant diversity in a complex landscape. Ecography 10.1111/ecog.03067.
 * Liu X. 2012. Taxonomy, diversity and spatial distribution characters of the ant family Formicidae (Insecta: Hymenoptera) in southeastern Tibet. PhD Thesis 139 pages
 * Liu X., Z. H. Xu, X. Y. Zhou, N. N. Yu, and C. L. Zhang. 2011. Ant communities of West slope of Mount Demula and Bomi Valley in Southeastern Tibet. Forest Research 24(4): 458-463.
 * Liu X., Z. Xu, C. Zhang, N. Yu, and G. Xu. 2012. Distribution patterns of ants from West slope of Mount Demula and Bomi Valley in Southeastern Tibet. Journal of Northwest Forestry University 27(4): 77-82.
 * Liu X., Z. Xu, X. Zhou, N. Yu, and C. Zhang. 2011. Ant Communities of West Slope of Mount Demula and Bomi Valley in Southeastern Tibet. Forest Research 24(4): 458-463.
 * Livory A. 2003. Les fourmis de la Manche. L'Argiope 39: 25-49.
 * Livory A. Un Nouvel Inventaire: les Fourmis. Argiope 29: 27-34.
 * Lomnicki J. 1928. Spis mrówek Lwowa i okolicy. Ksiegi Pamiatkowej (Lecia Gimn. IV Jana Dlugosza Lwowie) 50: 1-10.
 * Lorinczi G. 2011. Density and spatial pattern of nests in sub-Mediterranean ground-dwelling ant communities (Hymenoptera: Formicidae). Community Ecology 12(1): 51-57.
 * Maavara V. 1953. Ants of Estonian SSR. ABIKS loodusevaatlejale 10: 1-44.
 * Majzlan O., and P. Devan. 2009. Selected insect groups (Hymenoptera, Neuroptera, Mecoptera, Raphidioptera) of the Rokoš Massif (Strážovské vrchy Mts.). Rosalia (Nitra), 20, p. 63–70.
 * Malozemova L. A. 1972. Ants of steppe forests, their distribution by habitats, and perspectives of their utilization for protection of forests (north Kazakhstan). [In Russian.]. Zoologicheskii Zhurnal 51: 57-68.
 * Marko B. 2008. Ants (Hymenoptera: Formicidae) of the Sand Dunes of Foieni Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Martínez Ibáñez, M. D.. "Algunos datos sobre malformaciones de hormigas." Actas de las VIII Jornadas de la Asociación Española de Entomología (1986): 1035-1041.
 * Medina, M.. "Catálogo provisional de las hormigas de Andalucía." Actas de la Sociedad Española de Historia Natural 20 (1891): 95-104.
 * Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
 * Menozzi C. 1939. Formiche dell'Himalaya e del Karakorum raccolte dalla Spedizione italiana comandata da S. A. R. il Duca di Spoleto (1929). Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano. 78: 285-345.
 * Menozzi, C.. "Beitrag zur Ameisenfauna des nördlichen und östlichen Spaniens. Aufzählung der von den Herren Dr. F. Haas (1914-1919) und Prof. A. Seitz (1923) gesammelten Arten." Senckenbergiana 9 (1927): 89-92.
 * Meyer M., C. Braunert, R. Gerend, and I. Schrankel. 2003. Résultats de l'excursion annuelle du groupe de travail entomologique de la Société des naturalistes luxembourgeois. Bull. Soc. Nat. luxemb. 103: 103-120.
 * Moretti M. P. Duelli, and M. K. Obrist. 2006. Biodiversity and resilience of arthropod communities after fire disturbance in temperate forests. Oecologia 149: 312327.
 * Muñoz-López M., T. Palomeque, J. A. Carrillo, J. Pons, A. Tinaut, and P. Lorite. 2012. A new taxonomic status for Iberoformica (Hymenoptera, Formicidae) based on the use of molecular markers. Journal of Zoological Systematics and Evolutionary Research 50: 30-37.
 * Nadig A. 1918. Alcune note sulla fauna dell'alta Valsesia. Formicidae. Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano 56: 331-341.
 * Navás, L.. "Una excursión científica a la Sierra Nevada." Butll. I Cat. D'Hist. Nat. (1902): 115.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Nielsen, M.G. 1986. Respiratory rates of ants from different climatic areas. Journal of Insect Physiology 32(2): 125-131
 * Novgorodova T. A., A. S. Ryabinin. 2015. Trophobiotic associations between ants (Hymenoptera, Formicidae) and aphids (Hemiptera, Aphidomorpha) in South Zauralye. News of Saratov University. Chemistry Series, Biology, Ecology 2(15): 98-107.
 * Nylander, W.. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * Núñez-Pérez, E.. "Bases para el desarrollo del control integrado de los pulgones (Hom. Aphididae) de los cultivos de la provincia de León." Universidad de León, Secretariado de Publicaciones Tesis doct (1992): 7 páginas y 4 microfichas. León.
 * O'Rourke F. J. 1948. The distribution and general ecology of the Irish Formicidae. Proceedings of the Royal Irish Academy. Section B: Biological, Geological, and Chemical Science 52: 383-410.
 * Ordóñez-Urbano C., J. Reyes-López, and S. Carpintero-Ortega. 2007. Estudio faunísidos de los formícidos (Hymenoptera: Formicidae) asociados a los bosques de ribera en la Provincia de Córdoba (España). Primeras Aportaciones. Boletin Sociedad Entomológica Aragonesa 40: 367-375.
 * Ovazza M. 1950. Contribution à la connaissance des fourmis des Pyrénées-Orientales. Récoltes de J. Hamon Vie et Milieu 1: 93-94.
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Park S.J., and B.J. Kim. 2002. Faunal comparison of ants among Cheongsando and other islands of South Sea in Korea. Korean Journal of Entomology 32(1): 7-12.
 * Park, Seong, Joon and Byung, and Kim, Jin. 2002. Faunal Comparison of Ants among Cheongsando and Other Islands of South Sea in Korea. Korean Jornal of Entomology. 32(1):7-12.
 * Passera L. 1967. Peuplement en fourmis terricoles du rebord Meridional des Causses jurassiques du Quercy: la lande calcaire a buis. Vie et milieu. Serie C, Biologie terrestre 18(1): 189-205.
 * Petal J. 1974. Analysis of a sheep pasture ecosystem in the Pieniny mountains (the Carpathians) XV. The effect of pasture management on ant population. Ekologia Polska 22(3/4): 679-692.
 * Petal J. 1994. Reaction of ant communities to degradations of forest habitats in the Karkonosze Mountains. Polish Academy of Sciences, Museum and Institute of Zoology, Memorabilia Zoologica 48: 171-179.
 * Petal J. M. 1963. Faune des fourmis de la reserve de tourbiere en projet a Rakowskie Bagno pres de Frampol (voivodie de Lublin). Annales Universitatis Mariae Curie-Sk?odowska 58(7): 143-174.
 * Petal J. M. 1967. Contribution a la connaissance des fourmis (Formicidae, Hymenoptera) de la region de Lublin. VII. Associations des fourmis des milieux de tourbieres, de forets et de dunes aux environs de Libiszow (dist. De Parczew). Annales Universitatis Mariae Curie-Sklodowska Lublin-Polonia 22(9): 117-130.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z., B. Petrov, D. Milicic, T. Karan-Znidarsic. 2007. Contribution to the Myrmecofauna (Hymenoptera: Formicidae) of East and South Serbia. Acta Zoologica Bulgarica 59(3): 295-299.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Pisarski B. 1964. Fauna Mrowek Afganistanu. Bibliogr. k. 160-166, Nieoprawiony maszynopis pracy, Praca doktorska. Instytut Zoologiczny PAN, 1964, Bibliogr. p. 160-166
 * Pisarski B. 1967. Fourmis (Hymenoptera: Formicidae) d'Afghanistan récoltées par M. Dr. K. Lindberg. Annales Zoologici (Warsaw) 24: 375-425.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Punttila P., Y. Haila, J. Niemela, and T. Pajunen. 1994. Ant communities in fragments of old-growth taiga and managed surroundings. Ann. Zool. Fennici 31: 131-144.
 * Punttila P., Y. Haila, and H. Tukia. 1996. Ant communities in taiga clearcuts: habitat effects and species interactions. Ecography 19: 16-28.
 * Punttila P., and Y. Haila. 1996. Colonisation of a burned forest by ants in the southern Finnish Boreal forest. Silva Fennica 30(4): 421-435.
 * Pusvaskyte O. 1979. Myrmecofauna of the Lituanian SSR. Acta Entomologica Lituanica 4: 99-105.
 * Radchenko A. G. 2007. The ants (Hymenoptera, Formicidae) in the collection of William Nylander. Fragmenta Faunistica (Warsaw) 50: 27-41.
 * Ran H., and S. Y. Zhou. 2012. Checklist of chinese ants: formicomorph subfamilies (Hymenoptera: Formicidae) II. Journal of Guangxi Normal University: Natural Science Edition 30(4): 81-91.
 * Rasheed M. T., I. Bodlah, A. G. Fareen, A. A. Wachkoo, X. Huang, and S. A. Akbar. 2019. A checklist of ants (Hymenoptera: Formicidae) in Pakistan. Sociobiology 66(3): 426-439.
 * Reznikova Z. I. 2003. Distribution patterns of ants in different natural zones and landscapes in Kazakhstan and West Siberia along a meridian trend. Euroasian Entomological Journal 2(4): 235-342.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
 * Rosenhauer, W. G.. "Die Thiere Andalusiens." Erlangen (1856).
 * Ruzsky M. 1916. On zoological research in Yeniseisk province, work of summer of 1915. Izv. Imp. Tomsk. Univ. 65 (3rd p part: 1-21.
 * Ruzsky M. 1920. Ants of Kamchatka. Izv. Inst. Issled. Sib. 2: 76-80
 * Rzeszowski K., H. Babik, W. Czechowski, and B. Marko. 2013. Ants (Hymenoptera: Formicidae) of Chelmowa Gora in the Swietokrzyski National Park. Fragmenta Faunistica 56(1): 1-15.
 * Saaristo M. I. 1990. New provincial records for twelve ant species (Hymenoptera, Formicidae) from Finland. Entomologica Fennica 1: 191-192.
 * Salata S. 2014. Ants (Hymenoptera: Formicidae) of the National Park of the Stołowe Mts. Przyroda Sudetow 17: 161-172.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Salgueiro, J.. "Primeiros registros de Formicídeos (Hymenoptera) para dúas áreas protegidas portuguesas. Adiçao de duas novas especies á mirmecofauna de Portugal." Boletín de la Sociedad Entomológica Aragonesa 32 (2003): 109-110.
 * Salgueiro, J.. "Variaçao anual em três comunidades de formicídeos da Serra da Estrela. Adiçao de um género novo e de duas espécies novas para portugal." Boletín de la Asociación Española de Entomología 26 (3-4) (2002): 121-131.
 * Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
 * Santschi F. 1925. Fourmis d'Espagne et autres espéces paléartiques EOS (Revista española de entomología) 1: 339-360.
 * Santschi F. 1926. Travaux scientifiques de l'Armée d'Orient (1916-1918). Fourmis. Bulletin du Muséum National d'Histoire Naturelle 32: 286-293.
 * Santschi, F.. "Fourmis d'Espagne et autres espéces paléartiques." EOS (Revista española de entomología) 1 (1925): 339-360.
 * Santschi, F.. "Fourmis d'Espagne et des Canaries." Boletín de la Real Sociedad española de Historia natural (Madrid) 19 (1919): 241-248.
 * Santschi, F.. "Fourmis du Portugal." Memórias e Estudos do Museu Zoológico da Universidade de Coimbra 59 (1932): 1-3.
 * Santschi, F.. "Inventa entomologica itineris Hispanici et Maroccani, quod a. 1926 fecerunt Harald et Håkan Lindberg. Fourmis du Bassin Méditerranéen occidental et du Maroc récoltées par MM. Lindberg." Societas Scientiarum Fennica (Helsingfors) 3 (14) (1931): 1-13.
 * Santschi, F.. "Liste de fourmis d'Espagne recueilliés par Mr. J. M. Dusmet." Boletín de la Sociedad Entomológica de España 15 (1932): 69-74.
 * Saunders, E.. "Hymenoptera aculeata from Majorca and Spain. II Spain (1901-1902)." Transactions of the Entomological Society of London (1904): 624-660.
 * Schar S., G Talavera, X. Espadaler, J. D. Rana, A. A. Andersen, S. P. Cover, and R. Vila. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 00: 1-12.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Scupola A. 2016. First records of the ant genus Formica Linnaeus 1758 (Hymenoptera Formicidae) from the coastal pine woods of Salento Peninsula (Apulia, southern Italy). Thalassia Salentina 38: 137-141.
 * Seifert B. 2007. Die Ameisen Mittel- und Nordeuropas. Tauer: lutra Verlags- und Vertriebsgesellschaft, 368 pp.
 * Sellier Y., C. Galkowski, C. Lebas, and P. Wegnez. 2016. Découverte de Temnothorax pardoi (Tinaut, 1987) dans la réserve naturelle nationale du Pinail (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 106-113.
 * Slipinski P., B. Marko, K. Rzeszowski, H. Babik, and W. Czechowski. 2014. Lasius fuliginosus(Hymenoptera: Formicidae) shapes local ant assemblages. North-Western Journal of Zoology 10(2): 404-412.
 * Slipinski P., M. Zmihorski, and W. Czechowski. 2012. Species diversity and nestedness of ant assemblages in an urban environment. Eur. J. Entomol. 109: 197206.
 * Somogyi A. A., G. Lorinczi, J. Kovacs, and I. E. Maak. 2017. Structure of ant assemblages in planted poplar (Populus alba) forests and the effect of the common milkweek (Asclepias syriaca). Acta Zoologica Academiae Scientiarum Hungaricae 63(4): 443–457.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Stiprais M. 1973. Materi?li par R?gas kukai?u faunu. - Latvijas Entomologs, 15: 30-32.
 * Stitz, H.. "Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae." Jena G. Fischer (1939): 428 pp.
 * Stukalyuk S. V. 2015. Structure of the ant assemblages (Hymenoptera, Formicidae) in the broad-leaved forests of Kiev. Entomological Review 95(3): 370–387.
 * Stukalyuk S. V. 2017. Stratification of the ant species (Hymenoptera, Formicidae) in the urban broadleaf woodlands of the city of Kiev. Entomological Review 97(3): 320-343.
 * Stumper R. 1953. Etudes myrmecologiques. XI. Fourmis luxembourgeoises. Bulletin Soc. Nat. luxemb. 57: 122-135.
 * Stärcke A. 1942. Ants, collected by Dr. C. F. Engelhard at Stockmarknes (Island Hadsekoy, Vesterale Archipelago circa 68°50' Lat. N. Norway) and between 64° and 66° Lat. N. on the western coast of Norway, July 1932. Entomologische Berichten (Amsterdam) 11: 21-23.
 * Sushko G. G. 2013. The ants (hymenoptera, formicidae) bogs Belarusian Lakeland. Proceedings of the international. scientific-practical. conference, Vitebsk, 21-22 November 2013 - Vitebsk, 2013. - P. 215-216.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Sveum P. 1979. Notes on the distribution of some Norwegian ant species (Hymenoptera, Formicidae). Fauna Norv. ser. B. 26: 10-11.
 * Szujecki A., J. Szyszko, S. Mazur, and S. Perlinski. 1978. A succession of the ants (Formicidae) on afforested arable land and forest soils. Memorabilia Zoologica 29: 183-189.
 * Taheri A., J. Reyes-Lopez, and N. Bennas. 2014. Contribution a l'etude de la fauna myrmecologique du parc national de Talassemtane (nord du Maroc): biodiversite, biogeographie et especes indicatrices. Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 225236.
 * Tak N., and N. S. Rathore. 2008. Insecta: Hymenoptera: Formicidae. Fauna of Pin Valley National Park, Conservation Area Series,Zool. Surv. India 34: 53-60.
 * Tang J., Li S., Huang E., Zhang B. and Chen Y.. 1995. Hymenoptera: Formicidae (1). Economic Insect Fauna of China 47: 1-133.
 * Tausan I., J. Dauber, M. R. Tricia, and B. Marko. 2017. Succession in ant communities (Hymenoptera: Formicidae) in deciduous forest clear-cuts – an Eastern European case study. European Journal of Entomology 114: 92-100.
 * Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
 * Terayama. M. 2004. Geological and ecological distribution of Japanese ants communities. (translated from Japanese) Reports of the Saitama Prefecture Animal Research Association. 48:24
 * Thapa V. K. 2000. An Inventory of Nepal's Insects, Vol. III. IUCN Nepal, Kathmandu, xi + 475 pp.
 * Ticha, K. 2005. Inventarizacní pruzkum mravencu (Hymenoptera: Formicidae) NPP Svarec. 2005. Acta Rerum Naturalium 1:127-130.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Vagalinski B., and A. Lapeva-Gjonova. 2012. The ants (Hymenoptera, Formicidae) of Plana Mountain (Bulgaria). Historia naturalis bulgarica 20: 87-101.
 * Varela-Hernandez, F., M. Rocha-Ortega, R. W. Jones, and W. P. Mackay. 2016. Insectos: Hormigas (Formicidae) del estado de Queretaro, Mexico. Pages 397-404 in W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
 * Vele A., J. Holisa, J. Tragnerova. 2015. Ant succession on burned areas in forested landscape: a case study from the Bohemian Switzerland National Forest. Zpravy Lesnickeho Vyzkumu 60(1): 47-52.
 * Vele A., J. Holusa, J. Frouz, and O. Konvicka. 2011. Local and landscape drivers of ant and carabid beetle communities during spruce forest succession. European Journal of Soil Biology 47: 349-356.
 * Vele A., J. Holusa, J. Horak. 2016. Ant abundance increases with clearing size. J For Res 21: 110–114.
 * Vele A., J. Holusa, and J. Frouz. 2009. Sampling for ants in different-aged spruce forests: A comparison of methods. European Journal of Soil Biology 45(4): 1-6.
 * Vlasakova, B., J. Raabova, T. Kyncl, P. Dostal, M. Kavarova, P. Kovar and T. Herben. 2009. Ants accelerate succession from mountain grassland towards spruce forest. Journal of Vegetation Science 20: 577587
 * Vogrin, V.. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Wegnez P. 2014. Premières captures de Lasius distinguendus Emery, 1916 et de Temnothorax albipennis (Curtis, 1854) au Grand-Duché de Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société royale belge d’Entomologie 150 (2014) : 168-171.
 * Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
 * Wegnez P. 2018. Premières decouvertes de Myrmica bibikoffi Kutter, 1963 et de Ponera testacea Emery, 1895, au Luxembourg (Hymenoptera: Formicidae)Bulletin de la Société royale belge d’Entomologie 154: 263–272.
 * Wegnez P., D. Ignace, E. Lommelen, M. Hardy, J. Bogaert, and C. Nilsson. 2015. Redécouverte de Teleutomyrmex schneideriKutter, 1950 dans les Alpes françaises (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 52-57.
 * Wegnez P., and A. Ronk. 2017. Découverte de Camponotus herculeanus (Linnaeus, 1758) et signalement de quelques autres espèces rares de fourmis au Luxembourg (Hymenoptera : Formicidae). Bulletin de la Société des naturalistes luxembourgeois 119 : 153–159.
 * Wegnez P., and F. Mourey. 2016. Formica uralensis Ruzsky, 1895 une espèce encore présente en France mais pour combien de temps ? (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 152: 72-80.
 * Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
 * Wheeler W. M. 1907. The origin of slavery among ants. Popular Science Monthly 71: 550-559.
 * Wheeler W. M. 1909. Observations on some European ants. Journal of the New York Entomological Society 17: 172-187.
 * Wiezik M. 2006. Ants (Hymenoptera:  Formiciadae)  of  the  Sitno  National  Nature  Reserve (tiavnické vrchy Protected Landscape Area). Naturae Tutela 10: 73-77.
 * Wiezik M. 2007. Ants (Hymenoptera: Formicidae) of mountain and alpine ecosystems at Southern part of Krá?ovoho?ské Tatry Mts. Naturae Tutela 11: 85-90.
 * Wiezik M. 2008. Vertical distribution of ants from alpine habitats at Nízke Tatry mts. Naturae Tutela 12: 61-67.
 * Wiezik M., A. Wiezikova, and J. Murin. 2008. Ant assemblages at horná orava peatbogs. Naturae Tutela 12: 55-60.
 * Wiezik M., M. Kozon, and A. Wiezikova. 2013. Ants of  selected  peat  bog  habitats  at  the  Horná Orava protected area. Naturae Tutela 17(1): 57-63.
 * Wiezik M., M. Svitok, A. Wiezikova, and M. Dovciak. 2013. Shrub encroachment alters composition and diversity of ant communities in abandoned grasslands of western Carpathians. Biodivers Conserv 22: 23052320.
 * Wlodarczyk T. 2010. Ant species composition in relation to forest associations in Szczecin Landscape Park. Polish Journal of Entomology 79: 91-99.
 * Yarrow I. H. H. 1954. The British ants allied to Formica fusca L. (Hym., Formicidae). Trans. Soc. Br. Entomol. 11: 229-244.
 * Yu N., Z. Xu, C. Zhang, X. Liu, and Q. Li. 2012. Ant Communities from East Slope of Mount Demola to Zayu Valley in Southeastern Tibet. Scientia Silvae Sinicae 48(10): 163-169.
 * Yusopov Z. M. 2010. To the level of scrutiny of ants fauna (Hymenoptera, Formicidae from Kabardino-Balkarian state high-mountainous reserve (Central Caucasus). Proceedings of the Samara Scientific Center of RAS 12(5): 1464-1466.
 * Zhang C., Z. Xu, N. Yu, Q. He, and X. Liu. 2012. Distribution Patterns of Ant Species on East Slope of Mount Demola and Zayu Valley in Southeastern Tibet. Journal of Northeast Forestry University 40(3): 87-92
 * Zhuytszyuan D. 2016. The ants (Hymenoptera, Formicidae) Nizhne-Svirsky reserve and their environmental features. Master's thesis Saint Petersburg State University.
 * Zryanin V. A., and T. A. Zryanina. 2007. New data on the ant fauna Hymenoptera, Formicidae in the middle Volga River Basin. Uspekhi Sovremennoi Biologii 127(2): 226-240.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.
 * del Campo Gracia P., M. D. Martinez Ibanez, A. Tinaut, and S. Montagud Alario. 2014. Faunistic study of the Formicidae (Hymenoptera, Formicidae) from the Comunitat Valenciana (Spain). Boln. Asoc. esp. Ent. 38 (1-2): 33-65.