Checklist of Aphaenogaster species

The following species and subspecies belong to the genus Aphaenogaster. Synonyms are listed under their senior names. For valid names only see Aphaenogaster species.

A
aktaci Kiran & Tezcan, 2008 (Turkey)

albisetosa Mayr, 1886 (Mexico, United States)

• Novomessor cockerelli minor Enzmann, J., 1947

angulata Viehmeyer, 1922 (China)

annandalei Mukerjee, 1930 (India)

aquia (Buckley, 1867) (United States)

araneoides Emery, 1890 (Costa Rica, El Salvador, Guatemala, Honduras, Nicaragua, Panama)

• Aphaenogaster phalangium brevicollis Forel, 1899

ashmeadi (Emery, 1895) (United States)

• Aphaenogaster treatae harnedi Wheeler, W.M., 1919

atlantis Santschi, 1929 (Morocco)

B
balcanica (Emery, 1898) (Albania, Greece, Slovenia, Turkey)

baogong Terayama, 2009 (Taiwan)

barbara Shattuck, 2008 (Australia)

barbigula Wheeler, W.M., 1916 (Australia)

baronii Cagniant, 1988 (Morocco)

beccarii Emery, 1887 (China, India, Indonesia)

beesoni Donisthorpe, 1933 (India)

belti Forel, 1895 (Madagascar)

boulderensis Smith, M.R., 1941 (Mexico, United States)

boulderensis smithi Gregg, 1949 (United States)

burri (Donisthorpe, 1950) (Turkey)

C
caeciliae Viehmeyer, 1922 (China)

campana Emery, 1878 (Italy, Malta)

cardenai Espadaler, 1981 (Iberian Peninsula, Spain)

carolinensis Wheeler, W.M., 1915 (United States)

cavernicola Donisthorpe, 1938 (India)

cecconii Emery, 1894 (Greece)

cockerelli André, 1893 (Mexico, United States)

• Aphaenogaster sonorae Pergande, 1893

concolor Watanabe & Yamane, 1999 (China, Japan)

cristata (Forel, 1902) (India)

crocea André, 1881 (Algeria)

crocea croceoides Forel, 1890 (Algeria)

crocea lenis Santschi, 1911 (Tunisia)

crocea splendidoides Forel, 1890 (Algeria)

curiosa Santschi, 1933 (Morocco)

D
dejeani Cagniant, 1982 (Morocco)

depilis Santschi, 1911 (Algeria)

depilis afra Santschi, 1933 (Algeria)

depilis numida Santschi, 1933 (Tunisia)

depressa Bolton, 1995 (Turkey)

• Ceratopheidole perganderi Donisthorpe, 1950

dlusskyi Radchenko & Arakelian, 1991 (Armenia)

donann Watanabe & Yamane, 1999 (Japan)

dromedaria (Emery, 1900) (New Guinea)

• Aphaenogaster dromedarius nigra Donisthorpe, 1938

• Ischnomyrmex dromedarius fusca Emery, 1900

dulciniae Emery, 1924 (Iberian Peninsula, Spain)

E
edentula Watanabe & Yamane, 1999 (Japan)

ensifera Forel, 1899 (Mexico)

• Novomessor manni Wheeler, W.M. & Creighton, 1934

epirotes (Emery, 1895) (Albania, Bulgaria, Croatia, Greece, Israel, Italy, Montenegro, Serbia, Slovenia, Turkey)

erabu Nishizono & Yamane, 1990 (Japan)

espadaleri Cagniant, 1984 (Morocco)

exasperata Wheeler, W.M., 1921 (China)

F
fabulosa Arnoldi, 1968 (Turkmenistan)

fallax Cagniant, 1992 (Morocco)

famelica (Smith, F., 1874) (China, Democratic People's Republic of Korea, Japan)

• Aphaenogaster famelica frontosa Wheeler, W.M., 1928

• Aphaenogaster famelica ruida Wheeler, W.M., 1928

• Aphaenogaster vapida Wheeler, W.M., 1928

faureli Cagniant, 1969 (Algeria)

feae Emery, 1889 (China, India, Myanmar)

feae imbellis (Emery, 1900) (Indonesia)

feae nicobarensis (Forel, 1903) (Nicobar Island)

feae simulans Forel, 1915 (Indonesia)

fengbo Terayama, 2009 (Taiwan)

festae Emery, 1915 (Greece, Iraq, Turkey)

finzii Müller, 1921 (Bosnia and Herzegovina, Croatia, Greece, Italy, Montenegro, Serbia)

flemingi Smith, M.R., 1928 (United States)

• Aphaenogaster texana macrospina Smith, M.R., 1934

floridana Smith, M.R., 1941 (United States)

foreli Cagniant, 1995 (Morocco)

friederichsi Forel, 1918 (Madagascar)

fulva Roger, 1863 (United States)

• Aphaenogaster fulva rubida Enzmann, J., 1947

fulva azteca Enzmann, J., 1947 (Mexico)

G
geei Wheeler, W.M., 1921 (China)

gemella (Roger, 1862) (Spain)

gemella marocana (Forel, 1903) (Morocco)

• Aphaenogaster gemella tingitana Santschi, 1933

georgica Arnoldi, 1968 (Georgia)

gibbosa (Latreille, 1798) (Andorra, Armenia, Bulgaria, France, Georgia, Greece, Iberian Peninsula, Iran, Israel, Montenegro, Portugal, Russian Federation, Saudi Arabia, Spain, The former Yugoslav Republic of Macedonia, Turkmenistan)

• Aphaenogaster gibbosa barcinensis Menozzi, 1922

• Aphaenogaster gibbosa chorassanica Arnol'di, 1968

• Aphaenogaster gibbosa muschtaidica Emery, 1908

• Aphaenogaster striola laevior Forel, 1892

• Atta striola Roger, 1859

gibbosa fiorii Emery, 1915 (Italy)

gibbosa homonyma Emery, 1921 (Tunisia)

gonacantha (Emery, 1899) (Madagascar)

gracillima Watanabe & Yamane, 1999 (Japan)

graeca Schulz, 1994 (Greece)

H
haarlovi Collingwood, 1961 (Afghanistan)

hesperia Santschi, 1911 (Canary Islands)

holtzi (Emery, 1898) (Turkey)

honduriana Mann, 1922 (Honduras)

huachucana Creighton, 1934 (Mexico, United States)

huachucana crinimera Cole, 1953 (Mexico, United States)

hunanensis Wu, J. & Wang, 1992 (China)

I
iberica Emery, 1908 (Balearic Islands, Canary Islands, Iberian Peninsula, Portugal, Spain)

• Aphaenogaster angusta Santschi, 1925

• Aphaenogaster conica Viehmeyer, 1914

• Aphaenogaster testaceopilosa vieirai Santschi, 1933

incurviclypea Wang, W. & Zheng, 1997 (China)

inermita Bolton, 1995 (Italy, Malta)

• Aphaenogaster testaceopilosa inermis Santschi, 1933

ionia Santschi, 1933 (Croatia, Greece, Italy, Malta, Slovenia)

• Aphaenogaster semipolita ionia Baroni Urbani, 1968

• Aphaenogaster testaceopilosa melitensis Santschi, 1933

iranica Kiran & Alipanah, 2013 (Iran)

irrigua Watanabe & Yamane, 1999 (Japan)

isekram Bernard, 1977 (Algeria)

italica Bondroit, 1918 (Italy)

J
januschevi Arnoldi, 1976 (Russian Federation)

japonica Forel, 1911 (China, Democratic People's Republic of Korea, Japan)

• Aphaenogaster smythiesi sinensis Wheeler, W.M., 1928

• Aphaenogaster verecunda Wheeler, W.M., 1928

K
kervillei Forel, 1910 (Israel, Lebanon)

kimberleyensis Shattuck, 2008 (Australia)

koniari Cagniant & Galkowski, 2013 (Morocco)

kumejimana Watanabe & Yamane, 1999 (Japan)

kurdica Ruzsky, 1905 (Armenia, Georgia, Iran, Russian Federation)

L
laevior Emery, 1887 (Indonesia, Sulawesi)

lamellidens Mayr, 1886 (United States)

• Aphaenogaster lamellidens nigripes Smith, M.R., 1923

ledouxi Tohmé, G., 1969 (Greece, Lebanon)

lepida Wheeler, W.M., 1930 (China, Taiwan)

• Aphaenogaster silvestrii Wheeler, W.M., 1929

• Aphaenogaster funkikoensis Creighton, 1950

• Aphaenogaster phillipi Menozzi, 1932

lesbica Forel, 1913 (Greece, Israel, Montenegro)

leveillei Emery, 1881 (Algeria)

longiceps (Smith, F., 1858) (Australia, China, India)

• Aphaenogaster longiceps flava Emery, 1921

• Stenamma longiceps ruginota Forel, 1902

loriai (Emery, 1897) (New Guinea)

• Aphaenogaster loriai atra Stitz, 1938

lustrans Smith, M.R., 1961 (New Guinea)

luteipes Watanabe & Yamane, 1999 (Japan)

M
maculifrons Kiran & Aktaç, 2008 (Turkey)

mariae Forel, 1886 (United States)

mauritanica Dalla Torre, 1893 (Tunisia)

mediterrae Shattuck, 2008 (Australia)

megommata Smith, M.R., 1963 (Mexico, United States)

messoroides Dlussky, 1990 (Turkmenistan)

mexicana (Pergande, 1896) (Mexico)

miamiana Wheeler, W.M., 1932 (United States)

miniata Cagniant, 1990 (Morocco)

minutula Watanabe & Yamane, 1999 (Japan)

muelleriana Wolf, 1915 (Albania, Croatia, Greece, Italy, Montenegro, Slovenia)

mutica Pergande, 1896 (Mexico)

N
nadigi Santschi, 1923 (Morocco)

nana Wheeler, W.M., 1932 (United States)

O
obsidiana (Mayr, 1861) (Georgia, Greece, Iran, Israel, Montenegro, Russian Federation)

occidentalis (Emery, 1895) (Canada, United States)

• Aphaenogaster subterranea borealis Wheeler, W.M., 1915

opposita (Say, 1836) (United States)

osimensis Teranishi, 1940 (Japan)

ovaticeps (Emery, 1898) (Greece, Italy)

P
pallescens Walker, 1871 (Egypt)

pallida (Nylander, 1849) (Bulgaria, Italy)

pallida laurenti Santschi, 1939 (Algeria)

patruelis Forel, 1886 (Mexico, United States)

• Aphaenogaster patruelis willowsi Wheeler, W.M., 1933

• Stenamma patruele bakeri Wheeler, W.M., 1904

patruelis carbonaria Pergande, 1894 (Mexico)

perplexa Smith, M.R., 1961 (New Guinea)

• Aphaenogaster quadrispina wheeleri Donisthorpe, 1938

phalangium Emery, 1890 (Colombia, Costa Rica, Honduras, Panama)

• Aphaenogaster araneoides canalis Enzmann, J., 1947

• Aphaenogaster araneoides inermis Forel, 1899

• Aphaenogaster araneoides nitidiventris Forel, 1912

phillipsi Wheeler, W.M. & Mann, 1916 (Israel, Jordan, Occupied Palestinian Territory)

picea (Wheeler, W.M., 1908) (United States)

picena Baroni Urbani, 1971 (Italy)

polyodonta Zhou, 2001 (China)

poultoni Crawley, 1922 (Australia)

praedo Emery, 1908 (Morocco)

praedo ellipsoida Santschi, 1933 (Morocco)

praenoda Santschi, 1933 (Morocco)

praenoda confinis Santschi, 1933 (Morocco)

projectens Donisthorpe, 1947 (New Guinea)

pumilopuncta Zhou, 2001 (China)

punctaticeps Mackay, 1989 (United States)

punctithorax Cole, 1938 (United States)

pusilla Enzmann, J., 1947 (United States)

pythia Forel, 1915 (Australia, New Guinea)

Q
quadrispina Emery, 1911 (New Guinea)

R
radchenkoi Kiran, Aktaç & Tezcan, 2008 (Turkey)

reichelae Shattuck, 2008 (Australia)

relicta Wheeler, W.M. & Mann, 1914 (Haiti)

relicta epinotalis Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

rhaphidiiceps (Mayr, 1877) (Kazakhstan, Turkmenistan)

rifensis Cagniant, 1994 (Morocco)

rothneyi (Forel, 1902) (China, India)

rudis Enzmann, J., 1947 (United States)

rugulosa Watanabe & Yamane, 1999 (Japan)

rupestris Forel, 1909 (Algeria)

S
sagei (Forel, 1902) (China, India)

sagei pachei (Forel, 1906) (India, Nepal)

saharensis Bernard, 1953 (Algeria)

sangiorgii (Emery, 1901) (Greece, The former Yugoslav Republic of Macedonia)

sardoa Mayr, 1853 (Algeria, Italy)

• Aphaenogaster sardoa longispina Wolf, 1915

sardoa anoemica Santschi, 1910 (Morocco)

schmidti Karavaiev, 1912 (Republic of Korea)

schurri (Forel, 1902) (China, India)

semipolita (Nylander, 1856) (Georgia, Greece, Italy)

senilis Mayr, 1853 (Balearic Islands, Canary Islands, France, Iberian Peninsula, Italy, Portugal, Spain)

• Aphaenogaster senilis acoreensis Santschi, 1933

• Aphaenogaster senilis occidua Santschi, 1933

• Aphaenogaster testaceopilosa fuentei Medina, 1893

senilis disjuncta Santschi, 1933 (Morocco)

senilis grata Santschi, 1933 (Spain)

sicardi Cagniant, 1990 (Morocco)

sicula Emery, 1908 (Italy, Malta, Montenegro)

simonellii Emery, 1894 (Greece)

simonellii sporadis Santschi, 1933 (Greece)

smythiesii (Forel, 1902) (China, India, Nepal)

smythiesii prudens (Forel, 1902) (India, Nepal)

spinosa Emery, 1878 (Italy)

• Aphaenogaster testaceopilosum nitida Krausse, 1912

• Aphaenogaster testaceopilosum romana Santschi, 1933

spinosa etrusca Baroni Urbani, 1969 (Italy)

splendida (Roger, 1859) (Armenia, Croatia, Greece, Iberian Peninsula, Iran, Israel, Malta, Occupied Palestinian Territory, Spain)

splendida rugosoferruginea Forel, 1889 (Greece)

splendida transcaucasica Karavaiev, 1926 (Georgia)

striativentris Forel, 1895 (Iberian Peninsula, Spain)

strioloides Forel, 1890 (Algeria, Greece)

subcostata Viehmeyer, 1922 (Turkey)

subexaperata Zhou, 2001 (China)

subterranea (Latreille, 1798) (Albania, Andorra, Armenia, Balearic Islands, Belgium, Bulgaria, China, Croatia, Czech Republic, France, Georgia, Greece, Iberian Peninsula, Malta, Montenegro, Poland, Slovenia, Spain, The former Yugoslav Republic of Macedonia, Ukraine)

subterranea ichnusa Santschi, 1925 (Italy)

subterranea valida Wheeler, W.M., 1915 (United States)

subterraneoides Emery, 1881 (Croatia, Greece, Israel, Montenegro)

• Aphaenogaster pallida cypriotes Emery, 1908

subterraneoides armeniaca Arnoldi, 1968 (Armenia)

swammerdami Forel, 1886 (Madagascar)

swammerdami clara (Santschi, 1915) (Madagascar)

swammerdami curta Forel, 1891 (Madagascar)

swammerdami spinipes Santschi, 1911 (Madagascar)

syriaca Emery, 1908 (Iran, Israel, Lebanon)

• Aphaenogaster schmitzi syria Forel, 1910

syriaca schmitzi Forel, 1910 (Israel)

T
takahashii Wheeler, W.M., 1930 (China, Taiwan)

tennesseensis (Mayr, 1862) (United States)

• Atta laevis Mayr, 1862

• Myrmica subrubra Buckley, 1867

• Stenamma tennesseense ecalcaratum Emery, 1895

testaceopilosa (Lucas, 1849) (Algeria, Georgia, Greece)

testaceopilosa cabylica Stitz, 1917 (Algeria)

testaceopilosa canescens (Emery, 1895) (Algeria)

texana Wheeler, W.M., 1915 (Mexico, United States)

• Aphaenogaster silvestrii Menozzi, 1929

• Aphaenogaster texana furvescens Wheeler, W.M., 1915

theryi Santschi, 1923 (Morocco)

tibetana Donisthorpe, 1929 (China, Tibet)

tinauti Cagniant, 1992 (Morocco)

tipuna Forel, 1913 (Japan, Taiwan)

tokarainsulana Watanabe & Yamane, 1999 (China, Japan)

torossiani Cagniant, 1988 (Morocco)

treatae Forel, 1886 (United States)

• Aphaenogaster treatae wheeleri Mann, 1915

treatae pluteicornis Wheeler, G.C. & Wheeler, E.W., 1934 (United States)

turkestanica Arnoldi, 1976 (Kazakhstan)

U
uinta Wheeler, W.M., 1917 (United States)

ujhelyii Szabó, 1910 (Tunisia)

umphreyi Deyrup & Davis, 1998 (United States)

W
wangtian Terayama, 2009 (Taiwan)

wangye Terayama, 2009 (Taiwan)

weigoldi Viehmeyer, 1922 (China)

weulersseae Cagniant, 1989 (Morocco)

wilsoni Cagniant, 1988 (Morocco)

X
xuatian Terayama, 2009 (Taiwan)

=Fossil Species=

A
amphioceanica De Andrade, 1995 (Dominican Amber (Miocene))

antiqua Dlussky, 2002 (Rovno amber (Eocene))

archaica (Meunier, 1915) (Germany (Oligocene))

avita Fujiyama, 1970 (Japan (Miocene))

D
donisthorpei Carpenter, 1930 (United States)

dumetora (Lin, 1982) (China (Miocene))

L
lapidescens Zhang, 1989 (China (Miocene))

livida (Heer, 1867) (Croatia)

longaeva Scudder, 1877 (Canada (Miocene))

M
maculata Théobald, 1937 (France (Oligocene))

maculipes Théobald, 1937 (France (Oligocene))

mayri Carpenter, 1930 (United States)

mersa Wheeler, W.M., 1915 (Baltic Amber (Eocene))

O
oligocenica Wheeler, W.M., 1915 (Baltic Amber (Eocene))

P
paludosa Zhang, 1989 (China (Miocene))

pannonica Bachmayer, 1960 (Austria (Pliocene))

praerelicta De Andrade, 1995 (Mexican amber)

S
shanwangensis (Hong, 1984) (China (Miocene))

sommerfeldti Mayr, 1868 (Baltic Amber (Eocene))