Tetraponera pilosa

Habitats from which it has been recorded include second-growth rainforest, rainforest edge, “lower montane mixed dipterocarp forest”, and “rainforest along creek beds”. Colonies have been collected in dead twigs of Hevea brasiliensis, Vitex pubescens, Piper sp., and unidentifed vines and trees. (Ward 2001)

Identification
Ward (2001) - Tetraponera pilosa can be easily recognised by the features listed in the species-group diagnosis (next paragraph) and in the keys. In comparison with other Asian Tetraponera species, it shows relatively little variation in morphology.

The pilosa group contains a single, taxonomically isolated species whose workers are characterised by the presence of ocelli, large eyes (REL >0.40), subangulate humeri, shallow mesopropodeal impression, and dense punctate sculpture. The anteromedial lobe of the clypeus is poorly developed and standing pilosity is relatively sparse. Queens show the same traits (although the presence of ocelli is not discriminating since these are present in all Tetraponera queens). Males have very distinctive terminalia: the hypopygium is subrectangular in shape; the paramere bears a pair of digitiform lobes at its posterior end; and the aedeagus is subtriangular, with a row of fine teeth on the posteroventral margin.

Distribution based on Regional Taxon Lists
Indo-Australian Region: Borneo, Indonesia, Malaysia, Philippines, Singapore. Oriental Region: India, Nicobar Island, Thailand, Vietnam.

Nomenclature

 *  pilosa. Pseudomyrma pilosa Smith, F. 1858b: 160 (w.) BORNEO. Forel, 1915a: 23 (q.); Viehmeyer, 1916a: 117 (m.). Combination in Sima: Emery, 1887b: 449; in Tetraponera: Wheeler, W.M. 1919e: 65. Senior synonym of nicobarensis: Ward, 2001: 648.
 * nicobarensis. Sima nicobarensis Forel, 1903d: 402 (w.) INDIA (Nicobar Is). Combination in Tetraponera: Chapman & Capco, 1951: 80. Junior synonym of pilosa: Ward, 2001: 648.

Worker
Ward (2001) - HW 1.14-1.51, HL 1.28-1.70, LHT 0.93-1.31, CI 0.86-0.93, FCI 0.09-0.13, REL 0.43-0.49, REL2 0.49-0.56, SI 0.56-0.59, SI3 1.02-1.18, FI 0.43-0.49, PLI 0.48-0.53, PWI 0.45-0.52, PDI 1.01-1.08, LHT/HW 0.81-0.87, CSC 5-9, MSC 3-6.

Relatively large species, with broad head (CI >0.85); clypeus short, the anteromedial lobe broad, flat, and weakly developed; distance between frontal carinae less than or equal to maximum scape width; eyes large and prominent; ocelli present; profemur relatively robust (see FI values); pronotum with lateral margins well developed, and with humeri subangulate, when seen in dorsal view; mesopropodeal impression consisting of a short, transverse furrow, only moderately incised; propodeum slightly higher than wide, the dorsal face broad and flattened, and rounding gradually into the declivitous face; petiole as illustrated, long and low, with gently sloping anterodorsal and posterodorsal faces, and with prominent recurved anteroventral tooth; petiole about twice as long as broad; postpetiole as long as or slightly longer than broad; metabasitarsal sulcus present, lying in a thin strip of slightly darkened cuticle, occupying about 0.5-0.6x the length of the basitarsus. Head, mesosoma and petiole densely punctate to reticulopunctate, and subopaque; punctures becoming finer on postpetiole and gaster. Standing pilosity relatively sparse (see CSC and MSC values), absent from propodeum; short, appressed pubescence common on body, becoming denser posteriorly, and forming a thick mat on abdominal tergite IV Body dark brown, mandibles, antennae and apices of legs tending to be lighter medium-brown.

Type Material
Ward (2001) - Syntype (unique?), worker, “Borneo”.

References based on Global Ant Biodiversity Informatics

 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Crawley W.C. 1924. Ants from Sumatra, with biological notes by Edward Jacobson. Annals and Magazine of Natural History (9)13: 380-409
 * Dlussky G. M.; Radchenko A. G. 1990. The ants (Hymenoptera, Formicidae) of Vietnam. Subfamily Pseudomyrmicinae. Subfamily Myrmicinae (tribes Calyptomyrmecini, Meranoplini, Cataulacini). Pp. 119-125 in: Akimov, I. A.; Emelianov, I. G.; Zerova, M. D. et al. (eds.) 1990. News of faunistics and systematics. [In Russian.]. Kiev: Naukova Dumka, 184 pp.
 * Eguchi K.; Bui T. V.; Yamane S. 2011. Generic synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), part I  Myrmicinae and Pseudomyrmecinae. Zootaxa 2878: 1-61.
 * Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine)." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 5, no. 25 (1887): 427-473.
 * Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
 * Floren A., W. Wetzel, and M. Staab. 2013. The contribution of canopy species to overall ant diversity (Hymenoptera: Formicidae) in temperate and tropical ecosystems.  Myrmecological News 19: 65-74.
 * Forel A. 1903. Les fourmis des îles Andamans et Nicobares. Rapports de cette faune avec ses voisines. Rev. Suisse Zool. 11: 399-411.
 * Forel A. 1911. Die Ameisen des K. Zoologischen Museums in München. Sitzungsber. Math.-Phys. Kl. K. Bayer. Akad. Wiss. Münch. 11: 249-303.
 * Forel A. 1912. Einige neue und interessante Ameisenformen aus Sumatra etc. Zool. Jahrb. Suppl. 15: 51-78.
 * Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
 * Hashimoto Y., M. Mohamed, and H. Sakata. 1999. The ants (Hymenoptera: Formicidae) of the Tabin Wildlife Reserve, Sabah. Tabin Scientific Expedition 69-74.
 * Hashimoto Y., and M. Mohamed. 2004. A preliminary survey of ant fauna at Crocker Range Park. In: Maryati Mohamed, Zulhazman Hamzah, T. Tachi & J. Nais (eds.). Crocker Range scientific expedition 2002. Kota Kinabalu: Universiti Malaysia Sabah, pp. 51-71.
 * Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
 * Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
 * Karavaiev V. 1933. Ameisen aus dem Indo-Australischen Gebiet, VII. (Schluss). Konowia 12: 260-271.
 * Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
 * Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
 * Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
 * Santschi F. 1932. Résultats scientifiques du voyage aux Indes orientales néerlandaises de LL. AA. RR. le Prince et la Princesse Léopold de Belgique. Hymenoptera. Formicidae. Mémoires du Musée Royal d'Histoire Naturelle de Belgique. (2)4: 11-29.
 * Tanaka H. O., S. Yamane, and T. Itioka. 2012. Effects of a fern-dwelling ant species, Crematogaster difformis, on the ant assemblages of emergent trees in a Bornean tropical rainforest. Ann. Entomol. Soc. Am. 105(4): 592-598.
 * Viehmeyer H. 1916. Ameisen von den Philippinen und anderer Herkunft (Hym.). Entomologische Mitteilungen. Berlin-Dahlem 5: 283-291.
 * Ward P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy 15: 589-665.
 * Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebrate Taxonomy 15:589-665.
 * Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.