Lasius lasioides

Nests in the presence of trees, from forests to open areas with at least a few trees present, across a wide range of habitats. In Greece this is a common species, known from all provinces. It prefers warm habitats, such as pastures, mediterranean bushes, luminous deciduous forests. In Achaia, a single nest was reported under a stone in a mountain pasture (Borowiec & Salata, 2021).

Identification
Seifert (2020) - Palaearctic Lasius s. str. species belonging to the Lasius brunneus species complex. It can be separated from its semipatric relatives Lasius brunneus and Lasius silvaticus by smaller head width, longer scape and larger torulo-clypeal distance.

Confusion with Lasius himalayanus is excluded by zoogeography. The potentially sympatric Iranian desert species Lasius excavatus differs by head shape and coloration. Coloration in L. lasioides: head, mesosoma, gaster, femora and tibiae dark to medium brown; tibio-femoral joint region, scape and (frequently) anterior margin of clypeus pale yellowish-brown.

The Holomediterranean population of L. lasioides is polymorphic; there is in particular an extreme variance in scape length. This polymorphism shows a rather clear geographic structuring. Exclusively the long-scaped morph, which corresponds to the types of L. lasioides, was found in Italy and Greece and the islands of Mallorca, Malta, and Sardinia. Only the short-scaped morph, which corresponds to the types of Lasius barbarus, was found in the Spanish mainland and on Cyprus. Sympatric occurrence of both morphs is observed in Morocco, Algeria, Tunisia, Syria, and Israel. The Iberian shortscaped population and the Italian long-scaped population are in contact in southern France.

Distribution
Seifert (2020) - Lasius lasioides is a Holomediterranean species and presumably the lack of records from Libya and Egypt are caused by a lack of sampling. The most southwestern site is in Morocco at 30.8°N, 8.8° W. The northern distributional border runs along 43.5° N in France, 44.6° N in Italy, 40.5°N in Greece and Turkey whereas the easternmost truly Mediterranean site is at 35.6°N, 36.2°E in Syria. The distribution farther east is poorly known. Samples from two sites in the East Anatolian highland (PR_N 502, 39.43°N, 39.88°E, 1800 m; PR_N 460, 38.63°N, 43.45°E, 2300 m) might possibly represent a separate population or even species with special adaptation to a cold winter climate. A sample from Ghaemshar / Iran (36.46°N, 52.86°E, 49 m) represents the easternmost known site and raises the question if there is a continuous distribution from Syria and Israel east to the northern Iran. The highest latitude site in the Moroccan Atlas Mountains is at 33.00°N, 5.07°W, at 2240 m. It may be expected to have colonized all sufficiently large islands in the Mediterranean Sea which have some tree stands.

This is a common species, known from all Greek provinces (Borowiec et al., 2022).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Azerbaijan, Balearic Islands, Croatia, France, Georgia, Greece, Iberian Peninsula, Iran, Israel, Italy, Malta, Montenegro, Morocco, Portugal, North Macedonia, Spain, Turkey.

Habitat
Seifert (1992) - Lasius lasioides is found in all habitats where at least few trees were present. Recorded were: a sand dune with trees (Malta), pastures with scattered trees which are often Quercus species (4 observations), open Quercus forests (2 observ.), dense Quercus forests (4 observ.), a dense Pinus nigra forest and a dense Abies pinsapo forest (Spain). Despite of similarities in habitat, there is only one observation of syntopic occurrence with L. brunneus in Spain.

Biology
Rigato & Toni (2011) - Species known from Mediterranean Europe, Anatolia and along the Atlas mountains (Morocco). This is the first published record for Sardinia. Pashaei Rad et al. (2018) found this species in Iran on parkland ground in a moderate rainfall area.

Seifert (2020) - Lasius lasioides is apparently dependent on the presence of trees. It inhabits diverse types of broad-leafed or coniferous forest, both such with closed canopies or more open stands, as well as urban areas, gardens, pastures and road sides with at least a few scattered trees. The nests may be found under bark of the trees, on the ground in dead logs, or under stones. Their behavior is similar to L. brunneus: it is fugitive and not aggressive and workers prefer runways in crevices of bark or other surface structures. Development of alates differs throughout the whole geographic range in dependence from latitude and altitude. Eleven observations of alates occurred 30 April – 29 July, ten of these before 8 July.

Nomenclature

 * . Prenolepis lasioides Emery, 1869b: 6, pl. 1, fig. 3 (w.q.m.) ITALY.
 * Combination in Lasius: Emery, 1869b: 25 (in text); Emery, 1870: 194;
 * combination in Lasius (Lasius): Forel, 1915d: 53;
 * combination in Donisthorpea: Donisthorpe, 1915d: 347;
 * combination in Formicina (Donisthorpea): Emery, 1916b: 241;
 * combination in Formicina: Bondroit, 1918: 27;
 * combination in Acanthomyops: Kuznetsov-Ugamsky, 1927e: 189;
 * combination in Acanthomyops (Donisthorpea): Donisthorpe, 1927b: 400;
 * combination in Lasius: Müller, 1923b: 126; Kuznetsov-Ugamsky, 1929b: 38.
 * As unavailable (infrasubspecific) name: Forel, 1904b: 386; Ruzsky, 1905b: 310.
 * Junior synonym of fumatus: Emery & Forel, 1879: 452.
 * [Note: the reference Emery & Forel, 1879: 452, reverses the seniority of lasioides established by Emery, 1870: 194, but does not affect the synonymy and is discounted here.]
 * Junior synonym of alienus: Wilson, 1955a: 77; Smith, D.R. 1979: 1435; Agosti & Collingwood, 1987a: 58; Arakelian, 1994: 117.
 * Subspecies of alienus: Dalla Torre, 1893: 182; Cagniant, 1964: 92; Forel, 1915d: 53 (in key); Emery, 1916b: 241; Menozzi, 1925d: 34; Zimmermann, 1935: 49; Finzi, 1936: 191.
 * Subspecies of niger: Forel, 1911d: 352; Forel, 1913d: 434; Forel, 1915d: 53 (in key); Donisthorpe, 1915d: 347; Emery, 1916b: 241; Santschi, 1921e: 170; Kulmatycki, 1922: 81; Finzi, 1923: 4; Emery, 1925b: 230; Schkaff, 1925: 275; Soudek, 1925b: 15; Donisthorpe, 1927b: 400; Kuznetsov-Ugamsky, 1927e: 189; Kuznetsov-Ugamsky, 1929b: 37; Finzi, 1936: 191; Bernard, 1945: 137.
 * Status as species: Emery, 1870: 194; Bondroit, 1918: 27; Stärcke, 1926: 124 (in key); Stärcke, 1944a: 155; Consani & Zangheri, 1952: 44; Seifert, 1992b: 8 (redescription); Bolton, 1995b: 223; Mei, 1995: 768; Poldi, et al. 1995: 7; Schembri & Collingwood, 1995: 156; Bračko, 2006: 149; Cagniant, 2006a: 194; Gratiashvili & Barjadze, 2008: 136; Paknia, et al. 2008: 154; Casevitz-Weulersse & Galkowsky, 2009: 484; Vonshak, et al. 2009: 40; Karaman, M.G. 2011: 85; Legakis, 2011: 27; Borowiec, L. & Salata, 2012: 501; Kiran & Karaman, 2012: 12; Borowiec, L. & Salata, 2013: 357; Borowiec, L. 2014: 87; Bračko, et al. 2014: 20; Lebas, et al. 2016: 218; Borowiec, L. & Salata, 2018: 6; Salata & Borowiec, 2018b: 157 (in key); Salata & Borowiec, 2018c: 46.
 * Senior synonym of barbarus: Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of fumatus: Emery, 1870: 194; Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of fuscula: Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1916b: 241; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Bolton, 1995b: 223; Seifert, 2020: 49.
 * Senior synonym of nigrobrunneus: Seifert, 2020: 49.
 * barbarus. Lasius alienus var. barbarus Santschi, 1931a: 11.
 * [First available use of Lasius niger st. lasioides var. barbara Santschi, 1921e: 170 (w.) MOROCCO, TUNISIA; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Emery, 1925b: 230; Bernard, 1945: 137.
 * Subspecies of brunneus: Santschi, 1936c: 208.
 * Junior synonym of alienus: Wilson, 1955a: 78; Smith, D.R. 1979: 1435.
 * Junior synonym of lasioides: Seifert, 1992b: 8; Bolton, 1995b: 221; Seifert, 2020: 49.
 * fumatus. Lasius fumatus Emery, 1869b: 26, fig. 3 (w.q.) ITALY.
 * Synonym of lasioides: Emery & Forel, 1879: 452.
 * Subspecies of niger: Emery & Forel, 1879: 452.
 * [Note: the species which is referred to as Prenolepis lasioides in the text of Emery, 1869b: 6, is referred to in the list of figures for that paper (p. 26) as Lasius fumatus. Both names refer to the same specimens and hence are automatically synonymous (Bolton, 1995b: 223).]
 * Junior synonym of lasioides: Emery, 1870: 194; Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1925b: 230; Smith, D.R. 1979: 1435; Seifert, 1992b: 8; Bolton, 1995b: 223; Seifert, 2020: 49.
 * fuscula. Prenolepis fuscula Emery, 1869b: 8, pl. 1, fig. 4 (w.) ITALY.
 * Combination in Lasius: Emery, 1869b: 25 (in text), 26 (in figure captions).
 * As unavailable (infrasubspecific) name: Emery & Forel, 1879: 452.
 * Junior synonym of lasioides: Dalla Torre, 1893: 182; Forel, 1915d: 53; Emery, 1916b: 241; Emery, 1925b: 230; Bolton, 1995b: 223; Smith, D.R. 1979: 1435; Seifert, 2020: 49.
 * nigrobrunneus. Acanthomyops (Donisthorpea) brunneus var. nigrobrunneus Donisthorpe, 1926b: 18 (w.) ITALY.
 * Subspecies of brunneus: Donisthorpe, 1927a: 8.
 * Junior synonym of brunneus: Wilson, 1955a: 47; Bernard, 1967: 358; Seifert, 1992b: 6; Bolton, 1995b: 224; Radchenko, 2016: 366.
 * Junior synonym of lasioides: Seifert, 2020: 49.

Worker
Seifert (1992) - HL 848.1 ± 42.6 (64), HL/HW 1.0688 ± 0.0147 (64), SL/HL 0.9482 ± 0.0269 (64), CLCA 0.38 ± 0.14 (63), PDCL 31.41 ± 7.79 (63), nHS 0.0, nHHT 0.0, nBH 3.23 ± 1.27 (64), nUH 0.20 ± 0.34 (64), UHL/HL 0.0227 ± 0.0361 (64), PNHL/HL 0.1130 ±0.Ql08 (63).

Head: HL/HW (900) 1.059, SL/HL (900) 0.942. Mandibles normally 7-toothed (7.02 ± 0.53, 6-8, n = 27). Frontal line clearly visible. Carina well developed in center of clypeus, in anterior and posterior part frequently absent, lateral clypeal profile very convex. Genal setae completely absent. Mesosoma: reminiscent of L. brunneus; propodeum in lateral view with very straight posterior slope and conic dome, which is frequently slightly higher than promesonotum, mesopropodeal depression on average deeper than in L. brunneus. ScaIe: in anterior view wider than in L. brunneus and with moderately convex sides, always at least slightly emarginated; scale in lateral view thin and with convex anterior and straight posterior profile. Scape and hindtibia: seta and pubescence condition as in L. brunneus. Surface characters: very similar to L. brunneus; dorsal head with rather dilute, very appressed (0-10°) pubescence, PLF 20-26 µm; cuticular surface mildly shining, with weak micropunctures and weak microreticulum. Frontal pronotum with fine microreticulum and less dilute pubescence compared to head. Clypeal pubescence very sparse. Colour: head, meso soma, gaster, femora and tibiae dark to medium brown; tibio-femoral joint region, scape and (frequently) anterior margin of clypeus pale yellowish brown.

Queen
Seifert (1992) - N=7: HL 1234.9 ± 26.8, HW 1396.3 ± 50.4, ML 2470.9 ± 64.7, HL/HW 0.885 ± 0.0243, SL/HL 0.887 ± 0.0137, SL/HW 0.785 ± 0.0191, MH/ML 0.462 ± 0.0172, PDCL 20.86 ± 4.71, nHS 0.0, nHHT 0.0, nBH 1.14 ± 0.80, nUH 0.71 ± 1.07, PNHL 139.1 ± 12.4. Lectotype (n=1) NHMW: HL 1225, HW 1401, ML 2535, HLlHW 0.874, SLlHL 0.883, SLIHW 0.772, MH/ML0.471, PDCL 18.6, nHS 0, nHHT 0, nB H 0.5, nUH 0, PNHL 118, MW/ML 0.632.

Head: short, occipital margin straight; frontal groove fully developed from frontal triangle to midocellus, set in the middle of a shallow trough; mandibles in the 4 specimens with fully visible masticatory border 7-toothed. Genal setae completely absent. Mesosoma: strongly flattened, much wider than high. Scale: wide, clearly emarginate and with more rounded dorsal corners compared to L. brunneus. Scape and hindtibia: seta and pubescence condition as in worker. Surface characters: frontal head in overall appearance very smooth and moderately shining, micropunctures slightly weaker than in L. brunneus, weak microreticulum; pubescence 10-20°, PLF 30-45 !lm. Colour: head, mesosoma and gaster dark reddish brown; scape and tibiae yellowish.

Type Material
Seifert (2020) - Lectotype, an alate gyne, labelled by Mayr ”Neapel Emery“, ”Collect. G.Mayr“, ”L. fumatus m., lasioides m., det. Emery“ and ”lectotype, des. E.O.Wilson“; 1 paralectotype worker labelled by Mayr ”Neapel Col. G.Mayr“, ”L. fumatus m., lasioides m., det. Emery“ and ”Typus“; both specimens in.

Seifert (1992) - The lectotype queen in the NHMW is in good condition and is labelled: 'L. fumatus m., lasioides m., det. Emery, Neapel, Emery, Collect. G. Mayr' and 'Lectotype, des. E. O. Wilson.' The lectotype is probably the only original material of Emery that is still available (a search in the Emery collection in was negative). The specimen compares well with the worker attributed queens from Malta, Mallorca, and Algeria.

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