Checklist of Temnothorax species

The following species and subspecies belong to the genus Temnothorax. Synonyms are listed under their senior names. For valid names only see Temnothorax species.

A
abeli (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

adustus (Mackay, 2000) (Mexico, United States)

affinis (Mayr, 1855) (Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Georgia, Greece, Iberian Peninsula, Italy, Poland, Romania, Slovenia, Spain, Switzerland)

• Leptothorax tirolensis Gredler, 1858

• Leptothorax tuberoaffinis Bondroit, 1918

alayoi (Baroni Urbani, 1978) (Cuba, Greater Antilles)

alayorum (Fontenla Rizo, 1993) (Cuba)

albipennis (Curtis, 1854) (Belgium, Iberian Peninsula, Italy, Poland, Portugal, Spain, Switzerland, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax tuberointerruptus Bondroit, 1918

albispinus (Wheeler, W.M., 1908) (Greater Antilles, Puerto Rico)

• Antillaemyrmex albispina pallipes Wheeler, W.M., 1931

algiricus (Forel, 1894) (Algeria, Balearic Islands, Croatia, Iberian Peninsula, Morocco, Tunisia)

• Leptothorax angustulus silvanus Forel, 1907

• Leptothorax convexus timida Santschi, 1912

• Leptothorax gazella Santschi, 1923

• Leptothorax gazella monticola Santschi, 1923

algiricus trabutii (Forel, 1894) (Algeria)

• Leptothorax lindbergi Santschi, 1931

alienus Schulz, Heinze & Pusch, 2007 (Italy)

alinae (Radchenko, 1994) (Russian Federation)

allardycei (Mann, 1920) (Bahamas, United States)

• Antillaemyrmex floridanus Wheeler, W.M., 1931

alpinus (Ruzsky, 1902) (Georgia, Russian Federation)

ambiguus (Emery, 1895) (Canada, United States)

ambiguus foveatus (Smith, M.R., 1934)

ambiguus pinetorum (Wesson, L.G. & Wesson, R.G., 1940)

anacanthus (Santschi, 1912) (Morocco)

• Leptothorax hesperius inermis Santschi, 1909

anacanthus maximus (Santschi, 1931)

andersoni (Mackay, 2000) (Mexico, United States)

andrei (Emery, 1895) (Mexico, United States)

• Leptothorax nitens heathii Wheeler, W.M., 1903

• Leptothorax nitens occidentalis Wheeler, W.M., 1903

• Leptothorax ocellatus Mackay, 2000

androsanus (Wheeler, W.M., 1905) (Bahamas, Mexico)

angulohumerus Zhou, Huang, Yu & Liu, 2010 (China)

angustulus (Nylander, 1856) (Croatia, France, Iberian Peninsula, Libya, Malta, Portugal, Spain)

angustulus angustinodis (Stitz, 1917)

angustulus bordagei (Bondroit, 1918) (France)

anira (Terayama & Onoyama, 1999) (Japan)

annexus (Baroni Urbani, 1978) (Mexico)

• Macromischa annectens Wheeler, W.M., 1931

annibalis (Santschi, 1909) (Tunisia)

annibalis numidicus (Santschi, 1929) (Tunisia)

anodonta (Arnoldi, 1977) (Armenia, Iran)

anodontoides (Dlussky & Zabelin, 1985) (Turkmenistan)

antera (Terayama & Onoyama, 1999) (Japan)

antigoni (Forel, 1911) (Turkey)

arcanus (Kutter, 1973) (Switzerland)

archangelskiji (Kuznetsov-Ugamsky, 1926) (Kazakhstan)

arenarius (Santschi, 1908) (Israel, Tunisia)

arenarius fusciventris (Santschi, 1911) (Tunisia)

arenarius kruegeri (Kutter, 1928) (Libya)

argentipes (Wheeler, W.M., 1928) (China)

arimensis (Azuma, 1977) (Japan)

arpini (Tarbinsky, 1976) (Kyrgyzstan)

• Leptothorax narinicus Tarbinsky, 1976

artvinensis Seifert, 2006 (Turkey)

atlantis (Santschi, 1911) (Algeria, Iberian Peninsula, Portugal, Spain)

• Leptothorax angustulus brunea Santschi, 1918

atomus (Cagniant & Espadaler, 1997) (Morocco)

augusti (Baroni Urbani, 1978) (Mexico)

• Leptothorax petiolatus Forel, 1901

• Macromischa foreli Aguayo, 1931

auresianus (Santschi, 1929) (Algeria)

aveli (Bondroit, 1918) (Andorra, Balearic Islands, Democratic People's Republic of Korea, France, Iberian Peninsula, Malta, Portugal, Spain)

• Leptothorax berlandi Bondroit, 1918

• Leptothorax curvithorax Bondroit, 1918

• Leptothorax rabaudi Bondroit, 1918

aveli pici (Bondroit, 1918) (France)

aztecus (Wheeler, W.M., 1931) (Guatemala, Mexico)

• Macromischa azteca maya Wheeler, W.M., 1937

B
baeticus (Emery, 1924) (Iberian Peninsula, Spain)

banao (Fontenla Rizo, 1998) (Cuba)

barbouri (Aguayo, 1931) (Cuba, Greater Antilles)

barroi (Aguayo, 1931) (Cuba, Greater Antilles)

barryi (Cagniant, 1967) (Algeria)

basara (Terayama & Onoyama, 1999) (Japan)

bermudezi (Wheeler, W.M., 1931) (Cuba, Greater Antilles)

• Croesomyrmex bermudezi mutabilis Wheeler, W.M., 1931

• Leptothorax anemicus Baroni Urbani, 1978

• Macromischa bermudezi gracilis Aguayo, 1932

bestelmeyeri (Mackay, 2000) (United States)

bicolor (Mackay, 2000) (Mexico)

bikara (Terayama & Onoyama, 1999) (Japan)

bimbache Espadaler, 2007 (Canary Islands)

blascoi (Espadaler, 1997) (Iberian Peninsula, Spain)

bradleyi (Wheeler, W.M., 1913) (United States)

brauneri (Ruzsky, 1905) (Georgia)

brevispinosus (Mackay, 2000) (Mexico)

brevispinus (Chang & He, 2001) (China)

bristoli (Mackay, 2000) (Mexico, United States)

bruneri (Mann, 1924) (Cuba, Greater Antilles)

• Macromischa bruneri imitatrix Wheeler, W.M., 1937

• Macromischa bruneri ornatipes Wheeler, W.M., 1937

• Macromischa bruneri proxima Wheeler, W.M., 1937

bucheti (Santschi, 1909) (Morocco)

bucheti festai (Emery, 1924) (Libya)

bucheti mixtus (Santschi, 1929)

bucheti tuneticus (Santschi, 1918) (Tunisia)

bugnioni (Forel, 1894) (Algeria)

bugnioni pupseli (Santschi, 1909) (Tunisia)

bulgaricus (Forel, 1892) (Bulgaria, Greece, Israel)

bulgaricus aeolius (Forel, 1911) (Israel, Turkey)

bulgaricus cypridis (Santschi, 1930) (Cyprus)

bulgaricus smyrnensis (Forel, 1911)

C
cabrerae (Forel, 1893) (Canary Islands)

• Leptothorax cabrerae stitzi Santschi, 1918

caesari (Espadaler, 1997) (Iberian Peninsula, Spain)

cagnianti (Tinaut, 1983) (Iberian Peninsula, Spain)

canescens (Santschi, 1908) (Canary Islands)

carinatus (Cole, 1957) (Mexico, United States)

cenatus (Bolton, 1982) (Kenya)

chandleri (Mackay, 2000) (United States)

ciferrii (Menozzi & Russo, 1930) (Dominican Republic, Greater Antilles, Lesser Antilles)

clypeatus (Mayr, 1853) (Austria, Bulgaria, Croatia, Czech Republic, Greece, Iberian Peninsula, Poland, Slovenia, Spain)

cokendolpheri (Mackay, 2000) (United States)

coleenae (Mackay, 2000) (United States)

confucii (Forel, 1912) (Taiwan)

congruus (Smith, F., 1874) (China, Democratic People's Republic of Korea, Japan, Republic of Korea, Russian Federation)

convexus (Forel, 1894) (Algeria, Morocco)

• Leptothorax submuticus Emery, 1915

cornibrevis (Collingwood, 1961) (Afghanistan)

corticalis (Schenck, 1852) (Bulgaria, Czech Republic, Denmark, Georgia, Germany, Greece, Iberian Peninsula, Poland, Portugal, Romania, Slovenia, Spain, Sweden, Ukraine)

crassispinus (Karavaiev, 1926) (Bulgaria, Croatia, Czech Republic, Germany, Montenegro, Poland, Romania, Slovakia, Slovenia, Ukraine)

• Leptothorax nylanderi slavonicus Seifert, 1995

creightoni (Mann, 1929) (Cuba, Greater Antilles)

creolus (Baroni Urbani, 1978) (Dominican Republic, Greater Antilles)

crepuscularis (Tinaut, 1994) (Iberian Peninsula, Spain)

cristinae (Espadaler, 1997) (Iberian Peninsula, Spain)

cuneinodis Radchenko, 2004 (Democratic People's Republic of Korea)

curtulus (Santschi, 1929) (Morocco, Spain)

curtulus gentilis (Santschi, 1923) (Tunisia)

curvispinosus (Mayr, 1866) (United States)

• Stenamma gallarum Patton, 1879

cuyagueteje (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

D
darlingtoni (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

• Macromischa opalina Wheeler, W.M., 1937

delaparti (Forel, 1890) (Algeria)

• Leptothorax emeryi Santschi, 1923

desertorum Dlussky & Soyunov, 1988 (Russian Federation, Turkmenistan)

desioi (Menozzi, 1939) (India)

desioi melanicus (Menozzi, 1939) (India)

dessyi (Menozzi, 1936) (Greece)

discoloratus (Arnoldi, 1977) (Russian Federation)

dissimilis (Aguayo, 1932) (Cuba, Greater Antilles)

• Macromischa chloana Wheeler, W.M., 1937

ditifet (Espadaler, 1997) (Morocco)

duloticus (Wesson, L.G., 1937) (United States)

E
eburneipes (Wheeler, W.M., 1927) (China, Democratic People's Republic of Korea)

emmae (Mackay, 2000) (United States)

exilis (Emery, 1869) (Algeria, Croatia, France, Georgia, Greece, Italy, Malta, Montenegro, Slovenia, Tunisia, Turkey)

• Leptothorax exilis boccaris Santschi, 1923

• Leptothorax exilis creticus Forel, 1910

• Leptothorax exilis darii Forel, 1911

• Leptothorax exilis nitidulus Santschi, 1923

• Leptothorax tuberum ruficornis Emery, 1914

F
finzii (Menozzi, 1925) (Italy)

flavicornis (Emery, 1870) (Croatia, France, Italy, Montenegro, Slovenia, The former Yugoslav Republic of Macedonia)

flavidulus (Wheeler, W.M. & Mann, 1914) (Greater Antilles, Haiti)

flavispinus (André, 1883) (Iberian Peninsula, Israel, Spain)

• Leptothorax nigrita curvispinosus André, 1881

flavispinus amilcaris (Santschi, 1939) (Tunisia)

flavispinus clivulus (Santschi, 1939) (Tunisia)

flavispinus impurus (Santschi, 1939) (Tunisia)

flavispinus rufescens (Forel, 1907) (Tunisia)

foreli (Santschi, 1907) (Tunisia)

formosus (Santschi, 1909) (Iberian Peninsula, Morocco, Portugal, Spain)

fragosus (Mackay & Buschinger, 2002) (Canada)

fuentei (Santschi, 1919) (Spain)

fultonii (Forel, 1902) (China, India)

fumosus (Ruzsky, 1923) (Russian Federation, Turkmenistan)

furunculus (Wheeler, W.M., 1909) (United States)

fuscatus (Mann, 1920) (Guatemala)

G
gaetulus (Santschi, 1923) (Morocco)

gallae (Smith, M.R., 1949) (United States)

gibbifer (Baroni Urbani, 1978) (Cuba, Greater Antilles)

goniops (Baroni Urbani, 1978) (Mexico)

gracilicornis (Emery, 1882) (Canary Islands)

gracilicornis nivarianus (Santschi, 1925)

• Leptothorax nivarianus guancha Santschi, 1925

graecus (Forel, 1911) (Bulgaria, Croatia, Greece)

gredosi (Espadaler & Collingwood, 1982) (Andorra, Iberian Peninsula, Spain)

grouvellei (Bondroit, 1918) (France, Iberian Peninsula, Spain)

gundlachi (Wheeler, W.M., 1913) (Cuba, Greater Antilles)

• Croesomyrmex aguayoi Wheeler, W.M., 1931

• Croesomyrmex aguayoi archeri Wheeler, W.M., 1931

• Macromischa aguayoi natenzoni Aguayo, 1931

• Macromischa bierigi Santschi, 1931

H
hadrumetensis (Santschi, 1918) (Tunisia)

haira (Terayama & Onoyama, 1999) (Japan)

hengshanensis (Huang, Chen & Zhou, 2004) (China)

hesperius (Santschi, 1909) (Canary Islands)

himachalensis Bharti, Gul & Schulz, 2012 (India)

hispaniolae (Baroni Urbani, 1978) (Dominican Republic, Greater Antilles)

hispidus (Cole, 1957) (Mexico, United States)

huatuo Terayama, 2009 (Taiwan)

huehuetenangoi (Baroni Urbani, 1978) (Guatemala)

I
ibericus (Menozzi, 1922) (Iberian Peninsula, Spain)

imias (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

indra (Terayama & Onoyama, 1999) (Japan)

inermis (Forel, 1902) (India)

interruptus (Schenck, 1852) (Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France, Germany, Greece, Iberian Peninsula, Montenegro, Poland, Slovenia, Spain, United Kingdom of Great Britain and Northern Ireland)

• Myrmica simpliciuscula Nylander, 1856

interruptus nitidiceps (Dalla Torre, 1893) (Algeria)

iranicus (Radchenko, 1994) (Iran)

iris (Roger, 1863) (Cuba, Greater Antilles)

• Croesomyrmex iris rufithorax Wheeler, W.M., 1931

• Croesomyrmex iris tristis Wheeler, W.M., 1931

• Macromischa iris nigripes Santschi, 1930

isabellae (Wheeler, W.M., 1908) (Greater Antilles, Puerto Rico)

• Leptothorax hyperisabellae Baroni Urbani, 1978

• Macromischa isabellae mutica Smith, M.R., 1937

italicus (Consani, 1952) (Croatia, Italy, Slovenia)

ixili (Baroni Urbani, 1978) (Guatemala)

J
jailensis (Arnoldi, 1977) (Ukraine)

janushevi (Radchenko, 1994) (Tajikistan)

josephi (Mackay, 2000) (United States)

K
kashmirensis Bharti, Gul & Schulz, 2012 (India)

kaszabi (Pisarski, 1969) (Democratic People's Republic of Korea, Mongolia, Russian Federation)

• Leptothorax tuberum sachalinensis Kupyanskaya, 1990

kemali (Santschi, 1934) (Turkey)

kinomurai (Terayama & Onoyama, 1999) (Japan)

kirghizicus (Tarbinsky, 1976) (Kyrgyzstan)

kiudiria (Espadaler, 1997) (Morocco)

knipovitshi (Karavaiev, 1916) (Armenia, Russian Federation, Ukraine)

• Leptothorax junipereti Arnol'di, 1977

• Leptothorax nikitae Arnol'di, 1977

korbi (Emery, 1924) (Bulgaria, Georgia)

koreanus (Teranishi, 1940) (China, Democratic People's Republic of Korea, Japan, Republic of Korea)

kraussei (Emery, 1916) (Andorra, Iberian Peninsula, Italy, Malta, Montenegro, Spain)

kubira (Terayama & Onoyama, 1999) (Japan)

kuixing Terayama, 2009 (Taiwan)

kurilensis (Radchenko, 1994) (Russian Federation)

L
laciniatus (Stitz, 1917) (Algeria)

laestrygon (Santschi, 1931) (Italy)

laestrygon splendidiceps (Baroni Urbani, 1968) (Malta)

laetus (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

• Leptothorax similis Baroni Urbani, 1978

• Macromischa affinis Mann, 1920

• Macromischa affinis maerens Wheeler, W.M., 1937

• Macromischa affinis umbratipes Wheeler, W.M., 1937

lagrecai (Baroni Urbani, 1964) (Italy)

laurae (Emery, 1884) (Tunisia)

laurae antoniae (Cagniant, 1997) (Morocco)

laurae colettae (Santschi, 1915) (Tunisia)

laurae rosae (Santschi, 1915) (Tunisia)

leigong Terayama, 2009 (Taiwan)

leimu Terayama, 2009 (Taiwan)

leoni (Arnoldi, 1971) (Kazakhstan)

lereddei (Bernard, 1953) (Algeria)

leucacanthus (Baroni Urbani, 1978) (Mexico)

leviceps (Emery, 1898) (Italy, Spain)

• Leptothorax exilis dichroa Emery, 1916

leyeensis Zhou, Huang, Yu & Liu, 2010 (China)

lichtensteini (Bondroit, 1918) (Balearic Islands, Croatia, France, Iberian Peninsula, Montenegro, Slovenia, Spain)

liebi (Mackay, 2000) (United States)

liviae (Agosti & Collingwood, 2011) (United Arab Emirates)

longipilosus (Santschi, 1912) (Tunisia)

longispinosus (Roger, 1863) (Canada, Iberian Peninsula, Spain, United States)

• Leptothorax longispinosus iowensis Buren, 1945

• Leptothorax longispinosus laeviceps Buren, 1944

luteus (Forel, 1874) (Israel, Spain, Switzerland, Syrian Arab Republic)

M
makora (Terayama & Onoyama, 1999) (Japan)

manni (Wheeler, W.M., 1914) (Mexico)

maoerensis Zhou, Huang, Yu & Liu, 2010 (China)

marocana (Santschi, 1909) (Morocco)

mauritanicus (Santschi, 1909) (Morocco)

maurus (Santschi, 1921) (Morocco)

megalops (Hamann & Klemm, 1967) (Sudan, United Arab Emirates)

mekira Terayama & Kubota, 2011 (Japan)

melas (Espadaler, Plateaux & Casevitz-Weulersse, 1984) (France)

melleus (Forel, 1904) (Kazakhstan, Kyrgyzstan, Mongolia, Turkmenistan, Uzbekistan)

• Leptothorax balchashensis Arnol'di, 1971

• Leptothorax melleus csikii Pisarski, 1969

melnikovi (Ruzsky, 1905) (Georgia)

mexicanus (Mackay, 2000) (Mexico)

michali Radchenko, 2004 (Democratic People's Republic of Korea)

microreticulatus Bharti, Gul & Schulz, 2012 (India)

mimeuri (Cagniant, 1997) (Morocco)

minozzii (Santschi, 1922) (Italy)

• Leptothorax silvestrii Menozzi, 1921

minutissimus (Smith, M.R., 1942) (United States)

mirabilis (Espadaler & Cagniant, 1996) (Morocco)

miserabilis (Santschi, 1918) (Tunisia)

mongolicus (Pisarski, 1969) (China, Democratic People's Republic of Korea, Mongolia, Russian Federation)

monjauzei (Cagniant, 1968) (Algeria)

mortoni (Aguayo, 1937) (Cuba, Greater Antilles)

myersi (Wheeler, W.M., 1931) (Cuba, Greater Antilles)

N
nadigi (Kutter, 1925) (Andorra, Armenia, Bulgaria, Georgia, Iberian Peninsula, Iran, Poland, Spain, Switzerland, The former Yugoslav Republic of Macedonia)

• Leptothorax caucasicus Arnol'di, 1977

• Leptothorax hasardaghi Dlussky & Zabelin, 1985

naeviventris (Santschi, 1910) (Iberian Peninsula, Spain, Tunisia)

naeviventris kefensis (Santschi, 1929) (Tunisia)

nassanovi (Ruzsky, 1895) (Democratic People's Republic of Korea, Kyrgyzstan, Mongolia, Republic of Korea, Russian Federation, Ukraine)

• Leptothorax galeatus Wheeler, W.M., 1927

• Leptothorax nassonovi firssovi Kuznetsov-Ugamsky, 1928

• Leptothorax nassonovi subnudus Ruzsky, 1905

neminan (Espadaler, 1997) (Morocco)

neomexicanus (Wheeler, W.M., 1903) (United States)

nevadensis (Wheeler, W.M., 1903) (United States)

• Leptothorax eldoradensis Wheeler, W.M., 1915

• Leptothorax lindae Mackay, 2000

• Leptothorax maryanae Mackay, 2000

• Leptothorax melanderi Wheeler, W.M., 1909

niger (Forel, 1894) (France, Greece, Iberian Peninsula, Israel, Spain)

• Leptothorax glaber Santschi, 1925

nigricans (Baroni Urbani, 1978) (Cuba, Greater Antilles)

• Macromischa melanocephala Wheeler, W.M., 1931

nigriceps (Mayr, 1855) (Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Georgia, Greece, Hungary, Iberian Peninsula, Poland, Slovenia, Spain)

nigriceps pyrenaeus (Bondroit, 1918) (France)

nigritus (Emery, 1878) (Algeria, Georgia, Greece)

nigritus rupestris (Forel, 1909) (Algeria)

nigritus saharensis (Santschi, 1929) (Algeria)

nigritus salambo (Santschi, 1912) (Tunisia)

nipensis (Fontenla Rizo, 1998) (Cuba, Greater Antilles)

nitens (Emery, 1895) (Mexico, United States)

• Leptothorax melinus Mackay, 2000

• Leptothorax nitens mariposa Wheeler, W.M., 1917

nordmeyeri (Schulz, 1997) (India)

normandi (Santschi, 1912) (Tunisia)

nylanderi (Foerster, 1850) (Andorra, Armenia, Belgium, Bulgaria, Denmark, France, Georgia, Germany, Greece, Iberian Peninsula, Montenegro, Poland, Portugal, Russian Federation, Spain, Sweden, Switzerland, The former Yugoslav Republic of Macedonia, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Leptothorax nylanderi nylanderotuberum Ruzsky, 1902

• Leptothorax tuberum nylanderocorticalis Forel, 1874

• Myrmica cingulata Schenck, 1852

nylanderi nylanderonigriceps (Stitz, 1939) (Germany)

O
obliquicanthus (Cole, 1953) (Mexico, United States)

obscurior (Dalla Torre, 1893) (Algeria, Morocco)

• Leptothorax exilis retifer Santschi, 1929

obturator (Wheeler, W.M., 1903) (United States)

ocarinae (Baroni Urbani, 1978) (Mexico)

opaciabdomin (Chang & He, 2001) (China)

oraniensis (Forel, 1894) (Algeria, Tunisia)

• Leptothorax rusticus Santschi, 1918

• Leptothorax rusticus chobauti Santschi, 1923

• Leptothorax rusticus sergensis Santschi, 1929

orchidus Zhou, Huang, Yu & Liu, 2010 (China)

oxianus (Ruzsky, 1905) (Kyrgyzstan, Uzbekistan)

• Leptothorax oxianus juglandeti Arnol'di, 1976

• Leptothorax talassicus Tarbinsky, 1976

oxynodis (Mackay, 2000) (United States)

P
pallidipes (Santschi, 1910) (Tunisia)

pallidus (Collingwood, 1961) (Afghanistan)

palustris (Deyrup & Cover, 2004) (United States)

pamiricus (Ruzsky, 1902) (Russian Federation)

pan (Santschi, 1936) (Morocco)

pardoi (Tinaut, 1987) (Iberian Peninsula, Portugal, Spain)

parvulus (Schenck, 1852) (Andorra, Armenia, Belgium, Bulgaria, Croatia, Czech Republic, Georgia, Germany, Iberian Peninsula, Iran, Montenegro, Poland, Portugal, Slovenia, Spain, Turkmenistan)

pastinifer (Emery, 1894) (Bahamas, Cuba, Greater Antilles, Haiti)

• Leptothorax pastoris Baroni Urbani, 1978

• Macromischa lucayensis Forel, 1901

• Macromischa pastinifera opacipes Wheeler, W.M., 1905

pelagosanus (Müller, 1923) (Croatia, Italy)

peninsularis (Wheeler, W.M., 1934) (Mexico)

pergandei (Emery, 1895) (Mexico, United States)

• Leptothorax floridanus Emery, 1895

• Leptothorax manni Wesson, L.G., 1935

• Leptothorax pergandei flavus Smith, M.R., 1929

personatus (Cagniant, 1987) (Morocco)

peyerimhoffi (Santschi, 1929) (Algeria)

pisarskii Radchenko, 2004 (China, Democratic People's Republic of Korea)

platycephalus (Espadaler, 1997) (Iberian Peninsula, Spain)

platycnemis (Wheeler, W.M., 1937) (Cuba, Greater Antilles)

poeyi (Wheeler, W.M., 1913) (Cuba, Greater Antilles)

• Macromischa poeyi rugiceps Aguayo, 1932

politus (Smith, M.R., 1939) (United States)

porphyritis (Roger, 1863) (Cuba, Greater Antilles)

• Macromischa jaumei Santschi, 1930

• Macromischa porphyritis latispina Wheeler, W.M., 1931

productus (Santschi, 1918) (Canary Islands, Morocco)

pulchellus (Emery, 1894) (Lesser Antilles)

punctaticeps (Mackay, 2000) (Mexico)

punctatissimus (Mackay, 2000) (Mexico)

punctithorax (Mackay, 2000) (Mexico)

punicans (Roger, 1863) (Cuba, Greater Antilles)

purpuratus (Roger, 1863) (Cuba, Greater Antilles)

R
racovitzai (Bondroit, 1918) (Balearic Islands, France, Iberian Peninsula, Montenegro, Portugal, Spain)

• Leptothorax massiliensis Bondroit, 1918

recedens (Nylander, 1856) (Armenia, Balearic Islands, Bulgaria, Croatia, France, Georgia, Greece, Iberian Peninsula, Israel, Malta, Montenegro, Portugal, Slovenia, Spain, Tunisia, Turkmenistan)

• Leptothorax ergatogyna Bernard, 1950

• Leptothorax mordax Santschi, 1919

• Leptothorax pictus Emery, 1924

• Leptothorax recedens barbarus Santschi, 1939

• Temnothorax rogeri Emery, 1869

reduncus (Wang & Wu, 1988) (China)

reticulatus (Chang & He, 2001) (China)

risii (Forel, 1892) (Canary Islands)

• Leptothorax elongatus Santschi, 1919

• Leptothorax risii armouri Wheeler, W.M., 1927

• Leptothorax risii globulinodis Santschi, 1909

rothneyi (Forel, 1902) (India)

rothneyi simlensis (Forel, 1904) (India)

rottenbergii (Emery, 1870) (Croatia, Italy)

rottenbergii ernesti (Santschi, 1909) (Algeria)

rottenbergii jesus (Forel, 1913) (Israel)

rottenbergii scabriosus (Santschi, 1909) (Algeria)

rudis (Wheeler, W.M., 1917) (United States)

rugatulus (Emery, 1895) (Mexico, United States)

• Leptothorax curvispinosus annectens Wheeler, W.M., 1903

• Leptothorax rugatulus brunnescens Wheeler, W.M., 1917

• Leptothorax rugatulus cockerelli Wheeler, W.M., 1917

• Leptothorax rugatulus dakotensis Wheeler, G.C. & Wheeler, E.W., 1944

• Leptothorax rugatulus mediorufus Wheeler, W.M., 1917

ruginosus Zhou, Huang, Yu & Liu, 2010 (China)

rugithorax (Mackay, 2000) (Mexico)

rugosus (Mackay, 2000) (Mexico)

rugulosus (Mackay, 2000) (Mexico)

S
sallei (Guérin-Méneville, 1852) (Dominican Republic, Greater Antilles, Haiti)

• Macromischa sallei haytiana Wheeler, W.M. & Mann, 1914

• Macromischa sallei opacinoda Wheeler, W.M., 1931

• Myrmica russula Nylander, 1856

salvini (Forel, 1899) (Costa Rica, Guatemala, Honduras, Mexico, Panama)

salvini obscurior (Forel, 1899)

santra (Terayama & Onoyama, 1999) (Japan)

santschii (Forel, 1905) (Algeria, Morocco, Tunisia)

• Leptothorax flavispinus boiteli Santschi, 1929

• Leptothorax flavispinus facetus Santschi, 1939

sardous (Santschi, 1909) (Italy)

satunini (Ruzsky, 1902) (Armenia, Georgia, Kyrgyzstan)

saudiae (Collingwood & Agosti, 1996) (Saudi Arabia)

saxatilis Schulz, Heinze & Pusch, 2007 (Italy)

saxonicus (Seifert, 1995) (Bulgaria, Czech Republic, Germany, Hungary, Poland, Slovenia)

schaufussi (Forel, 1879) (Iberian Peninsula, Spain)

schaumii (Roger, 1863) (United States)

• Leptothorax fortinodis Mayr, 1886

• Leptothorax fortinodis gilvus Wheeler, W.M., 1903

• Leptothorax fortinodis melanoticus Wheeler, W.M., 1903

schmittii (Wheeler, W.M., 1903) (United States)

schoedli Seifert, 2006 (Turkey)

schurri (Forel, 1902) (India)

schwarzi (Mann, 1920) (Cuba, Greater Antilles)

semenovi (Ruzsky, 1903) (Kyrgyzstan, Turkmenistan)

semiruber (André, 1881) (Bulgaria, Greece, Israel)

• Leptothorax rottenbergi balcanica Santschi, 1909

senectutis (Baroni Urbani, 1978) (Cuba, Greater Antilles)

serviculus (Ruzsky, 1902) (Democratic People's Republic of Korea, Mongolia, Republic of Korea, Russian Federation)

sevanensis (Arnoldi, 1977) (Armenia)

shannxiensis Zhou, Huang, Yu & Liu, 2010 (China)

shelkovnikovi (Karavaiev, 1926) (Russian Federation)

silvestrii (Santschi, 1911) (United States)

simesno (Espadaler, 1997) (Morocco)

singularis (Radchenko, 1994) (Tajikistan)

skwarrae (Wheeler, W.M., 1931) (Mexico)

smithi (Baroni Urbani, 1978) (United States)

• Leptothorax wheeleri Smith, M.R., 1929

solerii (Menozzi, 1936) (Greece)

sordidulus (Müller, 1923) (Austria, Bulgaria, Croatia, Italy, Slovenia)

• Leptothorax carinthiacus Bernard, 1957

• Leptothorax mulleri Santschi, 1926

• Leptothorax sordidulus tergestina Finzi, 1928

specularis (Emery, 1916) (Balearic Islands, Iberian Peninsula, Italy, Portugal, Spain)

spinosior (Forel, 1901) (China, Japan, Republic of Korea)

spinosus (Forel, 1909) (Algeria)

splendens (Wheeler, W.M., 1905) (Bahamas, Cuba, Greater Antilles)

• Macromischa scabripes Mann, 1920

• Macromischa scabripes brunneipes Wheeler, W.M., 1937

squamifer (Roger, 1863) (Cuba, Greater Antilles)

• Macromischa squamifera atrinodis Mann, 1920

steinbergi (Arnoldi, 1971) (Kazakhstan)

stenotyle (Cole, 1956) (United States)

• Leptothorax angustinodus Cole, 1956

stollii (Forel, 1885) (Guatemala)

• Leptothorax denticulatus Stitz, 1917

striatulus (Stitz, 1937) (Guatemala, Honduras, Mexico)

striatus Zhou, Huang, Yu & Liu, 2010 (China)

subcingulatus (Emery, 1924) (Iberian Peninsula, Spain)

subditivus (Wheeler, W.M., 1903) (Costa Rica, Guatemala, Mexico, United States)

• Macromischa flavitarsis Mann, 1920

• Macromischa laevissima Wheeler, W.M., 1911

• Macromischa luciliae Mann, 1935

suberis (Forel, 1894) (Algeria)

susamyri (Dlussky, 1965) (Kyrgyzstan)

T
taivanensis (Wheeler, W.M., 1929) (China, Taiwan)

tamarae (Radchenko, 1994) (Armenia, Georgia)

tauricus (Ruzsky, 1902) (Armenia, Bulgaria, Russian Federation, Ukraine)

• Leptothorax pithyusae Arnol'di, 1977

• Leptothorax pithyusae babuganicus Arnol'di, 1977

tebessae (Forel, 1890) (Algeria)

tebessae gapsensis (Santschi, 1929) (Tunisia)

tenuisculptus (Baroni Urbani, 1978) (Mexico)

tenuispinus (Santschi, 1911) (Tunisia)

terricola (Mann, 1920) (Cuba, Greater Antilles)

terrigena (Wheeler, W.M., 1903) (United States)

tesquorum (Arnoldi, 1977) (Turkmenistan)

• Leptothorax elderensis Dlussky & Zabelin, 1985

texanus (Wheeler, W.M., 1903) (United States)

• Leptothorax texanus davisi Wheeler, W.M., 1905

theryi (Santschi, 1921) (Morocco)

tianpeng Terayama, 2009 (Taiwan)

tianschanicus (Tarbinsky, 1976) (Kyrgyzstan)

torrei (Aguayo, 1931) (Cuba, Greater Antilles, United States)

totonicapani (Baroni Urbani, 1978) (Guatemala)

tricarinatus (Emery, 1895) (Mexico, United States)

tricolor (Santschi, 1925) (Morocco)

tristis (Bondroit, 1918) (France, Iberian Peninsula, Spain)

tuberum (Fabricius, 1775) (Andorra, Armenia, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France, Georgia, Greece, Iberian Peninsula, Kyrgyzstan, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, Sweden, The former Yugoslav Republic of Macedonia)

• Formica tuberosa Latreille, 1802

• Leptothorax melanocephalus Emery, 1870

• Leptothorax pratostepposus Arnol'di, 1977

• Leptothorax tuberum acutinodis Arnol'di, 1977

• Leptothorax tuberum alaicus Tarbinsky, 1976

• Leptothorax tuberum ciscaucasicus Arnol'di, 1977

• Leptothorax tuberum nigricephala Karavaiev, 1930

• Leptothorax tuberum stipaceus Ruzsky, 1905

tuberum nylanderounifasciatus (Forel, 1874) (Switzerland)

tuberum tuberonigriceps (Forel, 1874) (Switzerland)

turcicus (Santschi, 1934) (Turkey)

turritellus (Espadaler, 1997) (Morocco)

tuscaloosae (Wilson, 1951) (United States)

tyndalei (Forel, 1909) (Algeria, Iberian Peninsula, Morocco, Spain)

• Leptothorax balli Santschi, 1939

• Leptothorax cervantesi Santschi, 1919

U
unifasciatus (Latreille, 1798) (Armenia, Austria, Belgium, Bulgaria, Channel Islands, Croatia, Czech Republic, Denmark, France, Georgia, Greece, Iberian Peninsula, Israel, Italy, Kazakhstan, Malta, Montenegro, Morocco, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Leptothorax anoplogynus Emery, 1869

• Leptothorax cordieri Bondroit, 1918

• Leptothorax tuberum brauneri Karavaiev, 1937

• Leptothorax tuberum brauniella Radchenko, 1995

• Leptothorax tuberum kirillovi Ruzsky, 1905

• Leptothorax tuberum salina Karavaiev, 1937

• Leptothorax tuberum unifasciatointerruptus Forel, 1874

• Leptothorax unifasciatus ucrainicus Arnol'di, 1977

• Leptothorax volubilis Santschi, 1929

unifasciatus obenbergeri (Sadil, 1939) (Czech Republic)

unifasciatus rougeti (Bondroit, 1918) (France)

unifasciatus staegeri (Bondroit, 1918) (Switzerland)

unifasciatus unifasciatonigriceps (Novak & Sadil, 1941) (Germany)

universitatis (Espadaler, 1997) (Iberian Peninsula, Spain)

usunkul (Ruzsky, 1924) (Russian Federation)

V
versicolor (Roger, 1863) (Cuba)

• Macromischa lugens Roger, 1863

• Macromischa lugens nigra Santschi, 1931

versicolor guanahacabibes (Fontenla Rizo, 1995) (Cuba, Greater Antilles)

villarensis (Aguayo, 1931) (Cuba, Greater Antilles)

• Macromischa manni Wheeler, W.M., 1931

• Leptothorax williami Baroni Urbani, 1978

violaceus (Mann, 1924) (Cuba, Greater Antilles)

volgensis (Ruzsky, 1905) (Russian Federation)

W
werneri (Radchenko, 1994) (Russian Federation)

wheeleri (Mann, 1920) (Cuba, Greater Antilles)

• Macromischa wheeleri petri Aguayo, 1931

whitfordi (Mackay, 2000) (Mexico, United States)

wollastoni (Donisthorpe, 1940) (Portugal)

wroughtonii (Forel, 1904) (India)

wui (Wheeler, W.M., 1929) (China)

X
xanthos Radchenko, 2004 (China, Democratic People's Republic of Korea)

Y
yanwan Terayama, 2009 (Taiwan)

Z
zhejiangensis Zhou, Huang, Yu & Liu, 2010 (China)

=Fossil Species=

G
glaesarius (Wheeler, W.M., 1915) (Baltic Amber (Eocene))

gracilis (Mayr, 1868) (Baltic Amber (Eocene))

H
hystriculus (Wheeler, W.M., 1915) (Baltic Amber (Eocene))

L
longaevus (Wheeler, W.M., 1915) (Baltic Amber (Eocene))

P
petiolata (Mayr, 1868) (Baltic Amber (Eocene))

placivus (Wheeler, W.M., 1915) (Baltic Amber (Eocene))

praecreolus (De Andrade, 1992) (Dominican Amber (Miocene))