Leptogenys pilaka

Worker specimens were found foraging on the forest floor, and the colony nests were located in the rotten logs.

Identification
A member of the alluaudi complex of the incisa species group. Rakotonirina and Fisher (2014) - Worker. Small, peg-like setae absent from near anteromedian margin of clypeus, above anteriorly fringing semi-translucent lamella; in full-face view, eye extending beyond lateral cephalic margin; head short (CI: 94–97); antennal scape relatively short; mandible narrow, inner margin not convex and slightly curved near bases; blades mostly smooth with faint longitudinal striation.

Leptogenys pilaka is very similar to Leptogenys incisa andLeptogenys alluaudi, but is separable from L. alluaudi by the location of its eyes, which split the lateral outline of the head. In L. alluaudi, the eyes do not break the lateral cephalic margin. The species can be separated from L. incisa by its broader head and by the narrower mandibular blades which are not strongly curved at the base and have a largely smooth and shiny surface.

Key to Malagasy Leptogenys

Distribution based on Regional Taxon Lists
Malagasy Region: Madagascar.

Habitat
Leptogenys pilaka occurs in northern Madagascar, in the transitional forest of the RS Ankarana at 80 m and in the lower montane rainforest of PN Montagne d’Ambre at 885 m.

Nomenclature

 *  pilaka. Leptogenys pilaka Rakotonirina & Fisher, 2014: 95, figs. 46B, 47B, 106, 122 (w.) MADAGASCAR.

Worker
(5 specimens). HW: 1.98–2.17, HL: 2.09–2.23, CI: 94–97, SL: 2.38–2.52, SI: 116–123, PW: 1.37–1.47, WL: 3.45–3.66, PNH: 1.14–1.22, PNL: 1.04–1.11, PNW: 0.97–1.09, DNI: 90–99, LNI: 106–112.

Head more or less elongate and broader anteriorly than posteriorly; lateral border weakly convex and diverging from back to front. Eye large, jutting from head surface and splitting lateral cephalic margin. Antennal scape relatively short. Lateral margin of clypeus convex and rounding to short, narrowly triangular anteromedian lobe; edge of clypeus fringed with narrow and obtusely angulate semi-translucent lamella. Mandible long and slender, weakly curved near base; inner margin concave; apical portions not strongly intersecting and not closing tightly against clypeus. Hypostomal teeth not visible when head in full-face view. In lateral view, mesosoma relatively high and short; with bluntly angulate junction between propodeal dorsum and declivity; propodeal lobe with blunt angle. In dorsal view, mesosoma with three visible segments, metanotal groove impressed. Constriction between third and fourth abdominal segments fairly visible. Mandibles are mostly smooth and shining, though sparse and short striation may be present. Dorsum of head finely longitudinally rugulose from front to level of anterior margin of eye, becoming densely and finely reticulate-rugose toward mid-length of head, rest of head dorsum with fine microreticulate rugulae superimposed with large and shallow elongate punctures. Anterior portion of pronotum finely microreticulate, superimposed with coarse punctures; rest of mesosomal dorsum and petiolar node microreticulate, overlaid by a mixture of fine and coarse transverse rugae and shallow, large elongate punctures. Declivitous surface transversely rugose. Third abdominal tergite with elongate small punctures. Standing hairs present, pubescence abundant on head dorsum but very sparse on rest of body dorsum. Coloration black, basal portion of appendages brown, apical portion and tip of gaster light brown.

Type Material
Holotype worker: Madagascar, Antsiranana, RS Ankarana, 22.9 km 224° SW Anivorano Nord, -12.9089, 49.1098, 80 m, tropical dry forest, ex rotten log, 10–16 Feb 2001 (Fisher, Griswold et al.) collection code: BLF02867, specimen code: CASENT0428200. Paratypes: 4 workers with same data as holotype but with the following specimen codes: CASENT0428201, CASENT0428202, CASENT0428116, CASENT0247223 (CASC,, ).