Checklist of Messor species

The following species and subspecies belong to the genus Messor. Synonyms are listed under their senior names.

A
abdelazizi Santschi, 1921 (Morocco)

aciculatus (Smith, F., 1874) (Japan)

• Messor barbarus lobulifera Emery, 1901

• Stenamma aciculatum brunneicorne Forel, 1901

aciculatus risianus Forel, 1922 (China)

aegyptiacus (Emery, 1878) (Egypt)

aegyptiacus canaliculatus Donisthorpe, 1944 (Algeria)

aegyptiacus felah Santschi, 1923 (Egypt)

aegyptiacus phasianicus Bernard, 1953 (Algeria)

aegyptiacus tunetinus Santschi, 1923 (Tunisia)

alexandri Tohmé, G. & Tohmé, H., 1981 (Syria)

andrei (Mayr, 1886) (United States)

• Veromessor andrei castaneus Wheeler, W.M. & Creighton, 1934

• Veromessor andrei flavus Wheeler, W.M. & Creighton, 1934

angularis Santschi, 1928 (Kenya)

antennatus Emery, 1908 (Morocco)

antennatus fodorii Röszler, 1942 (Greece)

antennatus personatus Santschi, 1931 (Morocco)

aphaenogasteroides Pisarski, 1967 (Afghanistan)

aralocaspius (Ruzsky, 1902) (Russian Federation)

aralocaspius infumatus Kuznetsov-Ugamsky, 1929 (Kazakhstan)

arenarius (Fabricius, 1787) (North Africa)

• Myrmica amaurocyclia Foerster, 1850

• Myrmica scalpturata Nylander, 1856

• Stenamma bugnioni Forel, 1904

arenarius alfierii Santschi, 1938 (Egypt)

arenarius diabolus Santschi, 1938 (Egypt)

arenarius ratus Menozzi, 1933 (Israel)

arenarius thebaicus Santschi, 1912 (Egypt)

atanassovii Atanassov, 1982 (Bulgaria)

B
barbarus (Linnaeus, 1767) (Morocco)

• Aphaenogaster barbara sordida Forel, 1892

• Formica binodis Fabricius, 1775

• Formica juvenilis Fabricius, 1804

• Formica megacephala Leach, 1825

• Messor barbarus ambiguus Santschi, 1925

• Myrmica rufitarsis Foerster, 1850

• Messor barbarus nigricans Santschi, 1929

barbarus gallaoides Santschi, 1929 (Morocco)

• Messor barbarus nigriceps Santschi, 1925

barbarus mediosanguineus Donisthorpe, 1946 (Turkey)

barbarus politus Karavaiev, 1912 (Algeria)

• Messor barbarus dentiscapus Finzi, 1930

barbarus sahlbergi Forel, 1913 (Egypt)

barbarus santschii Emery, 1908 (Tunisia)

• Messor carthaginensis Bernard, 1980

beduinus Emery, 1922 (Tunisia)

berbericus Bernard, 1955 (Algeria)

bernardi Cagniant, 1967 (Algeria)

bouvieri Bondroit, 1918 (France)

boyeri Cagniant, 2006 (Morocco)

buettikeri Collingwood, 1985 (Saudi Arabia)

C
caducus (Victor, 1839) (Russia)

• Messor caducus caucasicola Arnol'di, 1977

caducus glabrescens Arnoldi, 1977 (Kazakhstan)

capensis (Mayr, 1862) (South Africa)

• Aphaenogaster pseudoaegyptiaca Emery, 1884

• Messor barbarus schencki Wheeler, W.M., 1922

• Messor braunsi Forel, 1913

• Messor donisthorpei Santschi, 1937

capitatus (Latreille, 1798) (France)

• Aphaenogaster barbara nigra André, 1883

• Formica huberiana Leach, 1825

• Myrmica galbula Losana, 1834

caviceps (Forel, 1902) (Algeria)

• Messor planiceps Stitz, 1917

celiae Reyes, 1985 (Spain)

cephalotes (Emery, 1895) (Ethiopia)

• Messor plinii Santschi, 1912

ceresis Santschi, 1934 (Lebanon)

chamberlini Wheeler, W.M., 1915 (United States)

chicoensis (Smith, M.R., 1956) (United States)

clypeatus Kuznetsov-Ugamsky, 1927 (Kazakhstan)

collingwoodi Bolton, 1982 (Niger)

crawleyi Santschi, 1928 (Egypt)

D
decipiens Santschi, 1917 (South Africa)

• Messor arcistriatus Santschi, 1928

• Messor barbarus probus Wheeler, W.M., 1922

dentatus Santschi, 1927 (Israel)

denticornis Forel, 1910 (Namibia)

• Messor denticornis brunni Forel, 1910

• Messor denticornis parvidens Forel, 1910

denticulatus Santschi, 1927 (Kazakhstan)

desertora He & Song, 2009 (China)

diabarensis Arnoldi, 1970 (Kazakhstan)

E
ebeninus Santschi, 1927 (Lebanon)

eglalae Sharaf, 2007 (Egypt)

erectus Espadaler, 1998 (Morocco)

excursionis Ruzsky, 1905 (Kazakhstan)

excursionis alaschanicus Ruzsky, 1915 (Mongolia)

excursionis sillemi Stärcke, 1935 (Kazakhstan)

F
ferreri Collingwood, 1993 (Kenya)

foreli Santschi, 1923 (Tunisia)

• Messor aegyptiacus fossulatus Santschi, 1923

foreli brevispinosus Santschi, 1923 (Morocco)

foreli surcoufi Santschi, 1923 (Algeria)

fraternus Ruzsky, 1905 (Russia)

G
galla (Mayr, 1904) (Ethiopia)

• Messor barbarus latinoda Santschi, 1917

• Messor galla airensis Bernard, 1950

• Messor galla nobilis Santschi, 1928

• Messor galla rufula Finzi, 1939

galla obscurus Menozzi & Consani, 1952 (Ethiopia)

H
hebraeus Santschi, 1927 (Israel)

hellenius Agosti & Collingwood, 1987 (Greece)

himalayanus (Forel, 1902) (India)

hismai Collingwood & Agosti, 1996 (Saudi Arabia)

hispanicus Santschi, 1919 (Spain)

hoggarensis Santschi, 1929 (Algeria)

I
ibericus Santschi, 1931 (Spain)

incisus Stitz, 1923 (Namibia)

incorruptus Kuznetsov-Ugamsky, 1929 (Kazakhstan)

• Messor barbarus hamulifera Karavaiev, 1926

inermis Kuznetsov-Ugamsky, 1929 (Kazakhstan)

instabilis (Smith, F., 1858) (India)

• Aphaenogaster barbara punctata Forel, 1886

intermedius Santschi, 1927 (Syria)

J
julianus (Pergande, 1894) (Mexico)

julianus clarior (Wheeler, W.M. & Creighton, 1934) (Mexico)

julianus manni (Wheeler, W.M. & Creighton, 1934) (United States)

K
kasakorum Arnoldi, 1970 (Kazakhstan)

kasakorum nadezhdae Arnoldi, 1970 (Kazakhstan)

kisilkumensis Arnoldi, 1970 (Uzbekistan)

L
lamellicornis Arnoldi, 1968 (Kazakhstan)

lariversi (Smith, M.R., 1951) (United States)

lobicornis Forel, 1894 (Algeria)

lobicornis batnensis Forel, 1909 (Algeria)

lobicornis laurenti Santschi, 1939 (Algeria)

lobicornis normandi Santschi, 1918 (Tunisia)

lobicornis rugulosus Santschi, 1926 (Algeria)

• Messor lobicornis rugosa Karavaiev, 1912

lobicornis submuticus Emery, 1908 (Tunisia)

lobognathus Andrews, 1916 (United States)

luebberti Forel, 1910 (Namibia)

luridus Santschi, 1927 (Turkey)

luridus virginalis Santschi, 1927 (Syria)

lusitanicus Tinaut, 1985 (Portugal)

M
maculifrons Santschi, 1927 (Syria)

marikovskii Arnoldi, 1970 (Kazakhstan)

marocanus Santschi, 1927 (Morocco)

medioruber Santschi, 1910 (Tunisia)

• Messor meridionalis sublaeviceps Santschi, 1927

medioruber maurus Santschi, 1927 (Tunisia)

medioruber montanus Santschi, 1927 (Algeria)

medioruber postpetiolatus Santschi, 1917 (Tunisia)

medioruber postquadratus Santschi, 1932 (Algeria)

melancholicus Arnoldi, 1977 (Caucasus)

• Messor melancholicus schelkovnikovi Arnol'di, 1977

meridionalis (André, 1883) (Greece)

minor (André, 1883) (Italy)

• Messor minor pompeianus Santschi, 1927

minor calabricus Santschi, 1927 (Italy)

minor capreensis Santschi, 1927 (Italy)

minor hesperius Santschi, 1927 (Spain (Canary Is))

minor laboriosus Santschi, 1927 (Kazakhstan)

minor maurus Barquin, 1981 (Spain (Canary Is))

muraywahus Collingwood & Agosti, 1996 (Saudi Arabia)

muscatus Collingwood & Agosti, 1996 (Muscat)

N
nahali Tohmé, G. & Tohmé, H., 1981 (Syria)

niloticus Santschi, 1938 (Egypt)

nondentatus He & Song, 2009 (China)

O
oertzeni Forel, 1910 (Turkey)

• Messor oertzeni amphigea Forel, 1911

oertzeni amasiensis Emery, 1921 (Turkey)

oertzeni carpathous Menozzi, 1936 (Greece)

olegianus Arnoldi, 1970 (Tajikistan)

orientalis (Emery, 1898) (Turkey)

P
perantennatus Arnoldi, 1970 (Kazakhstan)

pergandei (Mayr, 1886) (United States)

piceus Stitz, 1923 (South Africa)

picturatus Santschi, 1927 (Algeria)

R
regalis (Emery, 1892) (Nigeria)

• Cratomyrmex regalis rubea Santschi, 1913

• Cratomyrmex sculpturatus Stitz, 1916

reticuliventris Karavaiev, 1910 (Kazakhstan)

rufotestaceus (Foerster, 1850) (Algeria)

• Aphaenogaster gracilinodis Emery, 1878

• Atta thoracica Mayr, 1862

rufus Santschi, 1923 (Russian Federation)

• Messor clivorum karawajevi Tarbinsky, 1976

• Messor similis Kuznetsov-Ugamsky, 1927

rufus striolatus Kuznetsov-Ugamsky, 1927 (Kyrgyzstan)

ruginodis Stitz, 1916 (Congo)

rugosus (André, 1881) (Israel)

rugosus bodenheimeri Menozzi, 1933 (Israel)

S
sanctus Emery, 1921 (Tunisia)

sanganus Collingwood & Agosti, 1996 (Yemen)

semirufus (André, 1883) (vicinity of Caspian Sea)

• Messor semirufus kutteri Santschi, 1934

• Messor semirufus nigricans Santschi, 1932

• Messor semirufus nigriceps Santschi, 1929

semirufus emeryi Santschi, 1927 (Israel)

semirufus grandinidus Emery, 1912 (Tunisia)

semoni (Forel, 1906) (Algeria)

smithi (Cole, 1963) (United States)

stoddardi (Emery, 1895) (United States)

striatellus Arnoldi, 1970 (Tajikistan)

striaticeps (André, 1883) (Caucasus)

striatifrons Stitz, 1923 (Namibia)

striativentris Emery, 1908 (Algeria)

striatulus (Emery, 1891) (Tunisia)

structor (Latreille, 1798) (France)

• Formica aedificator Schilling, 1839

• Formica lapidum Fabricius, 1804

• Formica rufitarsis Fabricius, 1804

• Messor barbarus varrialei Emery, 1921

• Messor clivorum sevani Arnol'di, 1977

• Messor platyceras rubella Crawley, 1920

• Messor rufitarsis darianus Pisarski, 1967

• Messor rufitarsis jakowlevi Arnol'di, 1977

• Messor structor clivorum Ruzsky, 1905

• Messor structor novaki Finzi, 1929

• Messor structor subpolitus Kuznetsov-Ugamsky, 1927

• Messor structor turanicus Kuznetsov-Ugamsky, 1927

• Messor tataricus Ruzsky, 1905

• Myrmica mutica Nylander, 1849

• Stenamma structor tyrrhena Emery, 1898

structor aegaeus Santschi, 1926 (Turkey)

structor platyceras Crawley, 1920 (Iran)

structor tadzhikorum Arnoldi, 1970 (Tajikistan)

subgracilinodis Arnoldi, 1970 (Russian Federation)

sultanus Santschi, 1917 (Israel)

syriacus Tohmé, G., 1969 (Syria)

T
testaceus Donisthorpe, 1950 (Turkey)

timidus Espadaler, 1997 (Spain)

tropicorum Wheeler, W.M., 1922 (Angola)

• Messor braunsi nigriventris Stitz, 1923

• Messor denticornis laevifrons Stitz, 1923

turcmenochorassanicus Arnoldi, 1977 (Kazakhstan)

V
valentinae Arnoldi, 1970 (Tajikistan)

variabilis Kuznetsov-Ugamsky, 1927 (Kazakhstan)

• Messor excursionis macrops Santschi, 1927

vaucheri Emery, 1908 (Morocco)

vicinus Kuznetsov-Ugamsky, 1927 (Kazakhstan)

W
wasmanni Krausse, 1910 (Italy (Sardinia))

• Messor barbarus concolor Santschi, 1927

S
sculpturatus Carpenter, 1930 (United States)