Camponotus nearcticus

A widespread North American species that is found in a wide range of habitats.

Identification
This is a small, bicolored carpenter ant. The majors of this species can be recognized by the shiny gaster, by the dull surface of the mandible, lack of erect hairs in the sides of the heads and cheeks, and by the convex propodeum (and mesosoma). There are few erect hairs, specifically 1 - 2 pair on the basal margin of the clypeus, none on the cheeks or malar area, few on the dorsum of the mesosoma, petiole and gaster. (Mackay and Mackay 2002)

Distribution
Some areas of southern Canada and widespread across the United States.

Distribution based on Regional Taxon Lists
Nearctic Region: Canada, United States.

Habitat
Prairies and woodlands, especially riparian habitats, ranging from deciduous forests, oaks, pinyon-juniper forests up to ponderosa pine forests (1490 - 1700 m). (Mackay and Mackay 2002)

Biology
Mackay and Mackay (2002) - This is an arboreal species, especially of oaks (Quercus arizonica in New Mexico), cottonwoods and even pines. In central and southern Texas, this species nests in galls of the cynipid Disholcaspis cinerosa in live oaks (Quercus virginiana). Specimens from the Everglades were collected in a dead branch hanging in the understory of a mature hammock forest. Brood and reproductives were found in nests in March. An typical nest contained 198 minors and 36 majors. The largest complete nest we collected contained 531 minors, 188 majors, 5 alate females, 1 dealate nest gyne, and 144 males. All nests contained a single gyne. The brood was not counted. Reproductives were found in nests in March to October. This is a timid species that hesitates to rescue brood when the nest is disturbed. Workers tend coccids and aphids and carry pieces of dead insects to the nest. This is a house-infesting ant.

Nomenclature

 * . Camponotus marginatus var. nearcticus Emery, 1893i: 675 (w.q.) U.S.A. (New York, District of Columbia, Pennsylvania, Florida, California).
 * [Note: type-locality designated as Pennsylvania by Creighton, 1950a: 388.]
 * Wheeler, W.M. 1910f: 223 (s.m.); Wheeler, G.C. & Wheeler, J. 1953e: 193 (l.).
 * Combination in C. (Myrmentoma): Emery, 1925b: 117.
 * As unavailable (infrasubspecific) name: Emery, 1896d: 373 (in list); Wheeler, W.M. 1910f: 222.
 * Junior synonym of caryae: Wheeler, W.M. 1917c: 28; Wheeler, W.M. 1917i: 466 (in text); Emery, 1925b: 117.
 * Subspecies of marginatus: Wheeler, W.M. 1905f: 402; Wheeler, W.M. 1906b: 24.
 * Subspecies of fallax: Wheeler, W.M. 1908g: 406; Wheeler, W.M. 1910d: 342; Wheeler, W.M. 1910f: 222 (in key); Wheeler, W.M. 1910g: 572; Wheeler, W.M. 1913c: 117; Wheeler, W.M. 1916m: 600; Wheeler, W.M. 1917a: 558; Cole, 1936a: 39.
 * Subspecies of caryae: Buren, 1944a: 294.
 * Status as species: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, M.R. 1958c: 145; Carter, 1962a: 7 (in list); Smith, M.R. 1967: 367; Hunt & Snelling, 1975: 22; Francoeur, 1977b: 207; Yensen, et al. 1977: 184; Smith, D.R. 1979: 1432; Allred, 1982: 456; Snelling, R.R. 1988: 70; DuBois & LaBerge, 1988: 146; Deyrup, et al. 1989: 100; Wheeler, G.C., et al. 1994: 305; Bolton, 1995b: 113; Mackay & Mackay, 2002: 296; Deyrup, 2003: 44; Coovert, 2005: 170; Hansen & Klotz, 2005: 91; MacGown & Forster, 2005: 66; MacGown, et al. 2007: 16; Ellison, et al. 2012: 122; Deyrup, 2017: 195.
 * Senior synonym of minutus: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Snelling, R.R. 1988: 70; Bolton, 1995b: 113.
 * Senior synonym of pardus: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Bolton, 1995b: 113; Coovert, 2005: 170.
 * Senior synonym of tanquaryi: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Snelling, R.R. 1988: 70; Bolton, 1995b: 113.
 * minutus. Camponotus marginatus var. minutus Emery, 1893i: 676 (w.q.m.) U.S.A. (District of Columbia, Maryland, Missouri, New Jersey).
 * Wheeler, W.M. 1910f: 224 (s.w.q.m.).
 * Combination in C. (Myrmentoma): Emery, 1925b: 117.
 * As unavailable (infrasubspecific) name: Emery, 1896d: 373 (in list); Wheeler, W.M. 1910f: 224.
 * Subspecies of marginatus: Wheeler, W.M. 1905f: 403; Wheeler, W.M. 1906b: 24.
 * Subspecies of fallax: Wheeler, W.M. 1910d: 342; Wheeler, W.M. 1910f: 224; Wheeler, W.M. 1910g: 572; Wheeler, W.M. 1917a: 558.
 * Subspecies of caryae: Wheeler, W.M. 1917c: 28; Wheeler, W.M. 1917i: 466; Emery, 1925b: 117.
 * Junior synonym of nearcticus: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Snelling, R.R. 1988: 70; Bolton, 1995b: 111.
 * pardus. Camponotus caryae var. pardus Wheeler, W.M. 1917c: 28.
 * [First available use of Camponotus fallax subsp. fallax var. pardus Wheeler, W.M. 1910f: 225 (s.w.q.m.) U.S.A. (New York, New Jersey); unavailable (infrasubspecific) name.]
 * [Camponotus fallax var. pardus Wheeler, W.M. 1910d: 342. Nomen nudum.]
 * Subspecies of caryae: Emery, 1925b: 117; Wesson, L.G. & Wesson, R.G. 1940: 103.
 * Junior synonym of nearcticus: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Bolton, 1995b: 116; Coovert, 2005: 170.
 * tanquaryi. Camponotus caryae var. tanquaryi Wheeler, W.M. 1917c: 28.
 * [First available use of Camponotus fallax subsp. fallax var. tanquaryi Wheeler, W.M. 1910f: 226 (s.w.q.m.) U.S.A. (Illinois); unavailable (infrasubspecific) name.]
 * [Camponotus fallax var. tanquaryi Wheeler, W.M. 1910d: 342. Nomen nudum.]
 * Combination in C. (Myrmentoma): Emery, 1925b: 117.
 * Subspecies of caryae: Emery, 1925b: 117.
 * Junior synonym of nearcticus: Creighton, 1950a: 388; Smith, M.R. 1951a: 845; Smith, D.R. 1979: 1432; Snelling, R.R. 1988: 70; Bolton, 1995b: 126.

References based on Global Ant Biodiversity Informatics

 * Allred D. M. 1982. Ants of Utah. The Great Basin Naturalist 42: 415-511.
 * Allred, D.M. 1982. The ants of Utah. Great Basin Naturalist 42:415-511.
 * Andersen A. N. 1997. Functional Groups and Patterns of Organization in North American Ant Communities: A Comparison with Australia. Journal of Biogeography. 24: 433-460
 * Annotated Ant Species List Ordway-Swisher Biological Station. Downloaded at http://ordway-swisher.ufl.edu/species/os-hymenoptera.htm on 5th Oct 2010.
 * Beckmann R. L., and J. M. Stucky. 1981. Extrafloral Nectaries and Plant Guarding in Ipomoea pandurata (L.) G. F. W. Mey. (Convolvulaceae). American Journal of Botany 68(1): 72-79.
 * Belcher A. K., M. R. Berenbaum, and A. V. Suarez. 2016. Urbana House Ants 2.0.: revisiting M. R. Smith's 1926 survey of house-infesting ants in central Illinois after 87 years. American Entomologist 62(3): 182-193.
 * Bodner G. 2005. Invertebrates of the Peloncillo Region: Richness and Mystery. In Bodner, G.S., J. Atchley Montoya, R. Hanson, and W. Anderson, Editors. 2006. Natural heritage of the Peloncillo Mountain Region: a synthesis of science. World Wildlife Fund and Sky Island Alliance, Tucson, AZ. Available at skyislandalliance.org/media/Peloncillo%20Report.pdf
 * Buczkowski G., and D. S. Richmond. 2012. The Effect of Urbanization on Ant Abundance and Diversity: A Temporal Examination of Factors Affecting Biodiversity. PLoS ONE 7(8): e41729. doi:10.1371/journal.pone.0041729
 * Canadensys Database. Dowloaded on 5th February 2014 at http://www.canadensys.net/
 * Carroll T. M. 2011. The ants of Indiana (Hymenoptera: Formicidae). Master's Thesis Purdue university, 385 pages.
 * Clark A. T., J. J. Rykken, and B. D. Farrell. 2011. The Effects of Biogeography on Ant Diversity and Activity on the Boston Harbor Islands, Massachusetts, U.S.A. PloS One 6(11): 1-13.
 * Clark Adam. Personal communication on November 25th 2013.
 * Cokendolpher J.C., Reddell J.R., Taylor S.J, Krejca J.K., Suarez A.V. and Pekins C.E. 2009. Further ants (Hymenoptera: Formicidae) from caves of Texas [Hormigas (Hymenoptera: Formicdae) adicionales de cuevas de Texas]. Texas Memorial Museum Speleological Monographs, 7. Studies on the cave and endogean fauna of North America, V. Pp. 151-168
 * Colby, D. and D. Prowell. 2006. Ants (Hymenoptera: Formicidae) in Wet Longleaf Pine Savannas in Louisiana. Florida Entomologist 89(2):266-269
 * Cole A. C., Jr. 1953. A checklist of the ants (Hymenoptera: Formicidae) of the Great Smoky Mountains National Park, Tennessee. Journal of the Tennessee Academy of Science 28: 34-35.
 * Coovert G. A. 2005. The Ants of Ohio (Hymenoptera: Formicidae). Ohio Biological Survey, Inc. 15(2): 1-207.
 * Coovert, G.A. 2005. The Ants of Ohio (Hymenoptera: Formicidae) Ohio Biological Survey Bulletin New Series Volume 15(2):1-196
 * Dash S. T. and L. M. Hooper-Bui. 2008. Species diversity of ants (Hymenoptera: Formicidae) in Louisiana. Conservation Biology and Biodiversity. 101: 1056-1066
 * Davis W. T., and J. Bequaert. 1922. An annoted list of the ants of Staten Island and Long Island, N. Y. Bulletin of the Brooklyn Entomological Society 17(1): 1-25.
 * Del Toro, I. 2010. PERSONAL COMMUNICATION. MUSEUM RECORDS COLLATED BY ISRAEL DEL TORO
 * Deyrup M. 2016. Ants of Florida: identification and natural history. CRC Press, 423 pages.
 * Deyrup M., C. Johnson, G. C. Wheeler, J. Wheeler. 1989. A preliminary list of the ants of Florida. Florida Entomologist 72: 91-101
 * Deyrup, M. 2003. An updated list of Florida ants (Hymenoptera: Formicidae). Florida Entomologist 86(1):43-48.
 * Deyrup, M. and J. Trager. 1986. Ants of the Archbold Biological Station, Highlands County, Florida (Hymenoptera: Formicidae). Florida Entomologist 69(1):206-228
 * DuBois M. B. 1981. New records of ants in Kansas, III. State Biological Survey of Kansas. Technical Publications 10: 32-44
 * Dubois, M.B. and W.E. Laberge. 1988. An Annotated list of the ants of Illionois. pages 133-156 in Advances in Myrmecology, J. Trager
 * Ellison A. M., E. J. Farnsworth, and N. J. Gotelli. 2002. Ant diversity in pitcher-plant bogs of Massachussetts. Northeastern Naturalist 9(3): 267-284.
 * Ellison A. M., and E. J. Farnsworth. 2014. Targeted sampling increases knowledge and improves estimates of ant species richness in Rhode Island. Northeastern Naturalist 21(1): NENHC-13NENHC-24.
 * Emery C. 1893. Beiträge zur Kenntniss der nordamerikanischen Ameisenfauna. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 7: 633-682.
 * Forster J.A. 2005. The Ants (hymenoptera: Formicidae) of Alabama. Master of Science, Auburn University. 242 pages.
 * Frye J. A., T. Frye, and T. W. Suman. 2014. The ant fauna of inland sand dune communities in Worcester County, Maryland. Northeastern Naturalist, 21(3): 446-471.
 * General D., and L. Thompson. 2008. New distributional records of ants in Arkansas. Journal of the Arkansas Academy of Science 62: 148-150.
 * General D.M. & Thompson L.C. 2007. Ants (Hymenoptera: Formicidae) of Arkansas Post National Memorial. Journal of the Arkansas Acaedemy of Science. 61: 59-64
 * Glasier J. R. N., S. E. Nielsen, J. Acorn, and J. Pinzon. 2019. Boreal sand hills are areas of high diversity for Boreal ants (Hymenoptera: Formicidae). Diversity 11, 22; doi:10.3390/d11020022.
 * Gotelli, N.J. and A.M. Ellison. 2002. Biogeography at a Regional Scale: Determinants of Ant Species Density in New England Bogs and Forests. Ecology 83(6):1604-1609
 * Gregg R. E. 1945 (1944). The ants of the Chicago region. Annals of the Entomological Society of America 37: 447-480
 * Gregg, R.T. 1963. The Ants of Colorado.
 * Guénard B., K. A. Mccaffrey, A. Lucky, and R. R. Dunn. 2012. Ants of North Carolina: an updated list (Hymenoptera: Formicidae). Zootaxa 3552: 1-36.
 * Heithaus R. E., and M. Humes. 2003. Variation in Communities of Seed-Dispersing Ants in Habitats with Different Disturbance in Knox County, Ohio. OHIO J. SCI. 103 (4): 89-97.
 * Ipser R. M. 2004. Native and exotic ants (Hymenoptera: Formicidae) of Georgia: Ecological Relationships with implications for development of biologically-based management strategies. Doctor of Philosophy thesis, University of Georgia. 165 pages.
 * Ipser, R.M., M.A. Brinkman, W.A. Gardner and H.B. Peeler. 2004. A Survey of Ground-Dwelling Ants (Hymenoptera: Formicidae) in Georgia. The Florida Entomologist 87(3) 253-260.
 * Ivanov, K. 2019. The ants of Ohio (Hymenoptera, Formicidae): an updated checklist. Journal of Hymenoptera Research 70: 65–87.
 * Ivanov K., L. Hightower, S. T. Dash, and J. B. Keiper. 2019. 150 years in the making: first comprehensive list of the ants (Hymenoptera: Formicidae) of Virginia, USA. Zootaxa 4554 (2): 532–560.
 * Jeanne R. J. 1979. A latitudinal gradient in rates of ant predation. Ecology 60(6): 1211-1224.
 * Johnson C. 1986. A north Florida ant fauna (Hymenoptera: Formicidae). Insecta Mundi 1: 243-246
 * Kjar D. 2009. The ant community of a riparian forest in the Dyke Marsh Preserve, Fairfax County, Virginiam and a checklist of Mid-Atlantic Formicidae. Banisteria 33: 3-17.
 * Kjar D., and E. M. Barrows. 2004. Arthropod community heterogeneity in a mid-Atlantic forest highly invaded by alien organisms. Banisteria 23: 26-37.
 * Kjar D., and Z. Park. 2016. Increased ant (Hymenoptera: Formicidae) incidence and richness are associated with alien plant cover in a small mid-Atlantic riparian forest. Myrmecological News 22: 109-117.
 * Knowlton G. F. 1970. Ants of Curlew Valley. Proceedings of the Utah Academy of Sciences, Arts and Letters 47(1): 208-212.
 * Longino, J.T. 2010. Personal Communication. Longino Collection Database
 * Lubertazi, D. Personal Communication. Specimen Data from Museum of Comparative Zoology at Harvard
 * Lubertazzi D. and Tschinkel WR. 2003. Ant community change across a ground vegetation gradient in north Floridas longleaf pine flatwoods. 17pp. Journal of Insect Science. 3:21
 * Lynch J. F. 1988. An annotated checklist and key to the species of ants (Hymenoptera: Formicidae) of the Chesapeake Bay region. The Maryland Naturalist 31: 61-106
 * MacGown J. A., J. G. Hill, R. L. Brown, T. L. Schiefer, J. G. Lewis. 2012. Ant diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi. Mississippi Agricultural and Forestry Experiment Station Bulletin 1197: 1-30
 * MacGown J. A., J. G. Hill, and R. L. Brown. 2010.  Native and exotic ant in Mississippi state parks.  Proceedings:  Imported Fire Ant Conference, Charleston, South Carolina, March 24-26, 2008: 74-80.
 * MacGown J. A., R. L. Brown, J. G. Hill, and B. Layton. 2007. Carpenter ants of Mississippi. Mississippi Agricultural and Forestry Experiment Station Bulletin 1158: 1-35.
 * MacGown J. A., and R. L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown J. A., and R. Whitehouse. 2015. A preliminary report of the ants of West Ship Island. A report submitted to the Gulf Islands National Seashore. Mississippi Entomological Museum Report #2015-02. 9 pp.
 * MacGown J.A., Hill J.G. and Skvarla M. 2011. New Records of Ants (Hymenoptera: Formicidae) for Arkansas with a Synopsis of Previous Records. Midsouth Entomologist. 4: 29-38
 * MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
 * MacGown, J.A. and JV.G. Hill. Ants of the Great Smoky Mountains National Park (Tennessee and North Carolina).
 * MacGown, J.A. and R.L. Brown. 2006. Observations on the High Diversity of Native Ant Species Coexisting with Imported Fire Ants at a Microspatial Scale in Mississippi. Southeastern Naturalist 5(4):573-586
 * MacGown, J.A. and R.L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A. and T. Lockley. Ants of Horn Island, Jackson County, Mississippi
 * MacGown, J.A., J.G. Hill, R.L. Brown and T.L. 2009. Ant Diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi Report #2009-01. Schiefer. 2009.
 * MacGown. J. 2011. Ants collected during the 25th Annual Cross Expedition at Tims Ford State Park, Franklin County, Tennessee
 * Macgown J. A., S. Y. Wang, J. G. Hill, and R. J. Whitehouse. 2017. A List of Ants (Hymenoptera: Formicidae) Collected During the 2017 William H. Cross Expedition to the Ouachita Mountains of Arkansas with New State Records. Transactions of the American Entomological Society, 143(4): 735-740.
 * Mackay W. P., and E. E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
 * Mahon M. B., K. U. Campbell, and T. O. Crist. 2017. Effectiveness of Winkler litter extraction and pitfall traps in sampling ant communities and functional groups in a temperate forest. Environmental Entomology 46(3): 470–479.
 * Mann H. R., E. Rowe, J. Selfridge, and D. L. Price. 2018. Leaf litter and arboreal ants (Hymenoptera: Formicidae) in a Mid-Atlantic Forest. Northeastern Naturalist 25(2): 341-354.
 * McClelland L. A. 1978. The Nebraska distribution of the ant genus Camponotus Mayr (Hymenoptera: Formicidae). Master's Thesis, Department of Biology and the faculty of the Graduate of Nebraska at Omaha, 72 pages.
 * Moser J. C. and M. S. Blum. 1960. The Formicidae of Louisiana. Insect Conditions in Louisiana 3: 48-50
 * Nuhn, T.P. and C.G. Wright. 1979. An Ecological Survey of Ants (Hymenoptera: Formicidae) in a Landscaped Suburban Habitat. American Midland Naturalist 102(2):353-362
 * O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
 * O'Neill J.C. and Dowling A.P.G. 2011. A Survey of the Ants (hymenoptera: Formicidae) of Arkansas and the Ozark Mountains. An Undergraduate Honors, University of Arkansas. 18pages.
 * Reddell J. R., and J. C. Cokendolpher. 2001. Ants (Hymenoptera: Formicidae) from caves of Belize, Mexico, and California and Texas (U.S.A.) Texas. Texas Memorial Museum Speleological Monographs 5: 129-154.
 * Roeder K. A., and D. V. Roeder. 2016. A checklist and assemblage comparison of ants (Hymenoptera: Formicidae) from the Wichita Mountains Wildlife Refuge in Oklahoma. Check List 12(4): 1935.
 * Sackett T. E., S. Record, S. Bewick, B. Baiser, N. J. Sanders, and A. M. Ellison. 2011. Response of macroarthropod assemblages to the loss of hemlock (Tsuga canadensis), a foundation species. Ecosphere 2(7):art74. doi:10.1890/ES11-00155.1
 * Sargent J. M., Benson. E.P., Zungoli. P. A. and Bridges. W. C. 2002. Carpenter Ant (Hymenoptera: Formicidae) Fauna of South Carolina. J. Agric. Urban Entomol. 18: 227-236
 * Shik, J., A. Francoeur and C. Buddle. 2005. The effect of human activity on ant species (Hymenoptera: Formicidae) richness at the Mont St. Hilaire Biosphere Reserve, Quebec. Canadian Field-Naturalist 119(1): 38-42.
 * Smith M. R. 1924. An annotated list of the ants of Mississippi (Hym.) (continued from page 85). Entomological News 35: 121-127.
 * Snelling, R.R. 1988. Taxonomic notes on Nearctic species of Camponotus, subgenus Myrmentoma (Hymenoptera: Formicidae). pages 55-XX in J. Trager, editor, Advances in Myrmecology
 * Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
 * Tanquary M. C. 1912. A preliminary list of ants from Illinois. Trans. Ill. Acad. Sci. 4: 137-142.
 * Tynes J. S., and R. E. Hutchins. 1964. Studies of plant-nesting ants in east central Mississippi. American Midland Naturalist 72(1): 152-156.
 * Van Pelt A. F. 1948. A Preliminary Key to the Worker Ants of Alachua County, Florida. The Florida Entomologist 30(4): 57-67
 * Van Pelt A. F. 1956. The ecology of the ants of the Welaka Reserve, Florida (Hymenoptera: Formicidae). American Midland Naturalist 56: 358-387
 * Van Pelt A., and J. B. Gentry. 1985. The ants (Hymenoptera: Formicidae) of the Savannah River Plant, South Carolina. Dept. Energy, Savannah River Ecology Lab., Aiken, SC., Report SRO-NERP-14, 56 p.
 * Varady-Szabo, H. 2004. Spiders and Ants Associated with Fallen Logs in Forillon National Park of Canada, Quebec. Masters Thesis, MgGill University, Montreal, Canada
 * Wang C., J. Strazanac and L. Butler. 2000. Abundance, diversity and activity of ants (Hymenoptera: Formicidae) in oak-dominated mixed Appalachian forests treated with microbial pesticides. Environmental Entomology. 29: 579-586
 * Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
 * Wesson L. G., and R. G. Wesson. 1940. A collection of ants from southcentral Ohio. American Midland Naturalist 24: 89-103.
 * Wheeler G. C., J. N. Wheeler, and P. B. Kannowski. 1994. Checklist of the ants of Michigan (Hymenoptera: Formicidae). The Great Lakes Entomologist 26(4): 297-310
 * Wheeler G. C., and J. Wheeler J. 1989. A checklist of the ants of Oklahoma. Prairie Naturalist 21: 203-210.
 * Wheeler G. C., and J. Wheeler. 1987. A Checklist of the Ants of South Dakota. Prairie Nat. 19(3): 199-208.
 * Wheeler W. M. 1906. Fauna of New England. 7. List of the Formicidae. Occasional Papers of the Boston Society of Natural History 7: 1-24.
 * Wheeler W. M. 1910. The North American ants of the genus Camponotus Mayr. Annals of the New York Academy of Sciences 20: 295-354.
 * Wheeler W. M. 1910. The North American forms of Camponotus fallax Nylander. Journal of the New York Entomological Society 18: 216-232.
 * Wheeler W. M. 1913. Ants collected in Georgia by Dr. J. C. Bradley and Mr. W. T. Davis. Psyche (Cambridge) 20: 112-117.
 * Wheeler W. M. 1917. The mountain ants of western North America. Proceedings of the American Academy of Arts and Sciences 52: 457-569.
 * Wheeler, G.C. and J. Wheeler. 1985. A checklist of Texas ants. Prairie Naturalist 17:49-64.
 * Wheeler, G.C., J. Wheeler and P.B. Kannowski. 1994. CHECKLIST OF THE ANTS OF MICHIGAN (HYMENOPTERA: FORMICIDAE). Great Lakes Entomologist 26:1:297-310
 * Wheeler, G.C., J. Wheeler, T.D. Galloway and G.L. Ayre. 1989. A list of the ants of Manitoba. Proceedings of the Entomological Society of Manitoba 45:34-49
 * Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
 * Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71