Lasius americanus

This North American species is commonly collected and widespread. They are primarily found in forest habitats but will occasionally occur in bogs and other wetlands. Nests are in soil, in and under rotten logs and stumps, and sometimes in deep leaf litter. (Ellison et al., 2012). This species has only recently (Schär et al. 2018) been recognized as distinct and separate from the morphologically similar Paleoarctic Lasius alienus. North American studies that refer to L. alienus, provided the authors correctly identified this ant, are records of L. americanus.

Identification
Mackay and Mackay (2002) - This is a small, dark brown or black species with relatively large eyes (12-14 ommatidia in maximum diameter). The scape has few (less than 5) or no erect hairs (except at apex). The penultimate tooth is about the same size as the adjacent teeth (other 2 basal teeth).

Ellison et al., (2012) - In New England, the large-eyed Lasius americanus can be confused with either of the other three species in the niger group: Lasius pallitarsis, Lasius neoniger, or Lasius niger. The key feature is that L. americanus lacks erect hairs on its scape and hind tibiae, and is more common in forested habitats. The other three species all have erect hairs on their scapes and hind tibiae, and are more common in open habitats.

Distribution
This North American species is found from southern British Columbia west to Nova Scotia, south to northern Florida and in the mountains of the southwestern United States and Mexico (Ellison et al., 2012).

Distribution based on Regional Taxon Lists
Nearctic Region: Canada, United States. Neotropical Region: Mexico.

Biology
Wilson (1955) - Over its entire range, this species shows a strong predilection for well shaded woodland, where it nests in rotting logs and stumps and under stones. Among the hundreds of colonies I have encountered in the field in the eastern United States, nearly all conformed to this ecological character. It may happen, however, that at high elevations or at the northern periphery of its range, the species occasionally nests in open situations. At the summit of Grandfather Mountain in North Carolina, for instance, I found a small but vigorous population living under stones in an open blueberry-and-heath "bald". The elevation was 5800 feet, higher by 700 feet than any other collection of the genus made in the course of several field trips in the southern Appalachians.

This species is replaced in nearly every available habitat except woodland by the equally successful and abundant members of the Lasius neoniger complex. Lasius pallitarsis occupies the same types of nesting sites as americanus and probably limits its northward spread. In general, one gains the impression that americanus has been squeezed into relatively narrow ecological ranges by its congeneric competitors, but is nevertheless eminently successful within those ranges.

Records of nuptial flights in this species are too sparse to allow a rigorous comparison with other species, with records ranging from May 30 (Decatur Co., Ga.) to December 4 (Alachua Co., Fla.). Both of these are very exceptional dates, however; the majority of the other records fall in August.



Wheeler 1906 - Lasius americanus occasionally builds detritus tents around the stems of plants. One of these which I found during the past summer at Colebrook, Connecticut, is represented in Fig. 3. A small colony of anericanus, nesting under a flat stone, was keeping aphides on a prostrate stem along which a broad and very irregular gallery had been excavated. Around the stem at the point where it emerged from under the stone, the ants had woven a cylindrical tube of fine vegetable detritus about iinches in length and closed at the outer end, as if to prevent the aphides from escaping from the nest.

New England
Ellison et al. (2012) - This omnivorous species collects elaiosomes from seeds, live insects and carcasses of dead ones, and tends a wide variety of aphids, scales, and treehoppers, and coccids that feed on plant roots. The colonies can be very large and have many queens, but individual queens disperse and found colonies independently.

Other Insects
This species is a host for the temporary parastites, and.

Nomenclature

 *  americanus. Lasius niger var. americanus Emery, 1893i: 639 (w.q.m.) U.S.A.
 * Wheeler, G.C. & Wheeler, J. 1953c: 147 (l.).
 * As unavailable (infrasubspecific) name: Wheeler, W.M. 1916k: 172; Wheeler, W.M. 1917a: 525; Wheeler, W.M. 1917i: 463; Emery, 1925b: 230; Essig, 1926: 866; Smith, M.R. 1930a: 5; Menozzi, 1932b: 311; Smith, M.R. 1951a: 850.
 * Subspecies of niger: Forel, 1900e: 285; Wheeler, W.M. 1904e: 305; Wheeler, W.M. 1905f: 393; Wheeler, W.M. 1906b: 12; Wheeler, W.M. 1906d: 343; Wheeler, W.M. 1908f: 623; Wheeler, W.M. 1910g: 569; Santschi, 1911d: 7; Wheeler, W.M. 1913c: 116; Wheeler, W.M. 1916m: 592; Cole, 1936a: 37; Dennis, 1938: 295; Wing, 1939: 164; Cole, 1942: 374; Buren, 1944a: 296; Wheeler, G.C. & Wheeler, E.W. 1944: 253.
 * Subspecies of alienus: Creighton, 1950a: 419.
 * Junior synonym of alienus: Wilson, 1955a: 77; Smith, D.R. 1979: 1435.
 * Status as species: Gregg, E.V. 1945: 530; Schar et al., 2018: 6.

Taxonomic Notes
Schar et al. (2018) - Lasius americanus Emery, 1893 was originally described as a subspecies of Lasius niger, and was then raised to species in 1945 by Gregg. Creighton (1950) treated it as a subspecies of Lasius alienus, and then Wilson (1955) finally placed it in synonymy with Lasius alienus. Our results support the species-level status of this taxon as suggested by Gregg (1945). Therefore Lasius americanus is removed from synonymy with Lasius alienus and treated as a full species.

References based on Global Ant Biodiversity Informatics

 * Allred D. M. 1982. Ants of Utah. The Great Basin Naturalist 42: 415-511.
 * Allred, D.M. 1982. The ants of Utah. Great Basin Naturalist 42:415-511.
 * Amstutz M. E. 1943. The ants of the Kildeer plain area of Ohio (Hymenoptera, Formicidae). The Ohio Journal of Science 43(4): 165-173.
 * Banschbach V. S., and E. Ogilvy. 2014. Long-term Impacts of Controlled Burns on the Ant Community (Hymenoptera: Formicidae) of a Sandplain Forest in Vermont. Northeastern Naturalist 21(1): 1-12.
 * Beck D. E., D. M. Allred, W. J. Despain. 1967. Predaceous-scavenger ants in Utah. Great Basin Naturalist 27: 67-78
 * Belcher A. K., M. R. Berenbaum, and A. V. Suarez. 2016. Urbana House Ants 2.0.: revisiting M. R. Smith's 1926 survey of house-infesting ants in central Illinois after 87 years. American Entomologist 62(3): 182-193.
 * Blacker, N.C. 1992. Some Ants from Southern Vancouver Island, British Columbia. J. Entomol. Soc. Bri. Columbia 89:3-12.
 * Blacker, N.C. 1992. Some ants (Hymenoptera: Formicidae) from Southern Vancouver Island, British Columbia. Journal of the Entomological Society of British Columbia 89:3-12
 * Blades, D.C.A. and S.A. Marshall. Terrestrial arthropods of Canadian Peatlands: Synopsis of pan trap collections at four southern Ontario peatlands. Memoirs of the Entomological Society of Canada 169:221-284
 * Borchert, H.F. and N.L. Anderson. 1973. The Ants of the Bearpaw Mountains of Montana (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 46(2):200-224
 * Brandt, M. and S. Foitzik. 2004. Community Context and Specialization Influence Coevolution between a Slavemaking Ant and Its Hosts. Ecology 85(11):2997-3009
 * Campbell K. U., and T. O. Crist. 2017. Ant species assembly in constructed grasslands isstructured at patch and landscape levels. Insect Conservation and Diversity doi: 10.1111/icad.12215
 * Choate B., and F. A. Drummond. 2012. Ant Diversity and Distribution (Hymenoptera: Formicidae) Throughout Maine Lowbush Blueberry Fields in Hancock and Washington Counties. Environ. Entomol. 41(2): 222-232.
 * Choate B., and F. A. Drummond. 2013. The influence of insecticides and vegetation in structuring Formica Mound ant communities (Hymenoptera: Formicidae) in Maine lowbush blueberry. Environ. Entomol. 41(2): 222-232.
 * Clark Adam. Personal communication on November 25th 2013.
 * Colby, D. and D. Prowell. 2006. Ants (Hymenoptera: Formicidae) in Wet Longleaf Pine Savannas in Louisiana. Florida Entomologist 89(2):266-269
 * Cole A. C. 1940. A Guide to the Ants of the Great Smoky Mountains National Park, Tennessee. American Midland Naturalist 24(1): 1-88.
 * Cole A. C., Jr. 1936. An annotated list of the ants of Idaho (Hymenoptera: Formicidae). Canadian Entomologist 68: 34-39.
 * Cole A. C., Jr. 1942. The ants of Utah. American Midland Naturalist 28: 358-388.
 * Cole A. C., Jr. 1949. The ants of Mountain Lake, Virginia. Journal of the Tennessee Academy of Science 24: 155-156.
 * Cole, A.C. 1936. An annotated list of the ants of Idaho (Hymenoptera; Formicidae). Canadian Entomologist 68(2):34-39
 * Coovert G. A. 2005. The Ants of Ohio (Hymenoptera: Formicidae). Ohio Biological Survey, Inc. 15(2): 1-207.
 * Coovert, G.A. 2005. The Ants of Ohio (Hymenoptera: Formicidae) Ohio Biological Survey Bulletin New Series Volume 15(2):1-196
 * Cover S. P., and R. A. Johnson. 20011. Checklist of Arizona Ants. Downloaded on January 7th at http://www.asu.edu/clas/sirgtools/AZants-2011%20updatev2.pdf
 * Del Toro I., K. Towle, D. N. Morrison, and S. L. Pelini. 2013. Community Structure, Ecological and Behavioral Traits of Ants (Hymenoptera: Formicidae) in Massachusetts Open and Forested Habitats. Northeastern Naturalist 20: 1-12.
 * Del Toro, I. 2010. PERSONAL COMMUNICATION. MUSEUM RECORDS COLLATED BY ISRAEL DEL TORO
 * Des Lauriers J., and D. Ikeda. 2017. The ants (Hymenoptera: Formicidae) of the San Gabriel Mountains of Southern California, USA with an annotated list. In: Reynolds R. E. (Ed.) Desert Studies Symposium. California State University Desert Studies Consortium, 342 pp. Pages 264-277.
 * Deyrup M., C. Johnson, G. C. Wheeler, J. Wheeler. 1989. A preliminary list of the ants of Florida. Florida Entomologist 72: 91-101
 * Deyrup, M. 2003. An updated list of Florida ants (Hymenoptera: Formicidae). Florida Entomologist 86(1):43-48.
 * Drummond F. A., A. M. llison, E. Groden, and G. D. Ouellette. 2012. The ants (Formicidae). In Biodiversity of the Schoodic Peninsula: Results of the Insect and Arachnid Bioblitzes at the Schoodic District of Acadia National Park, Maine. Maine Agricultural and forest experiment station, The University of Maine, Technical Bulletin 206. 217 pages
 * Dubois, M.B. and W.E. Laberge. 1988. An Annotated list of the ants of Illionois. pages 133-156 in Advances in Myrmecology, J. Trager
 * Ellison A. M., E. J. Farnsworth, and N. J. Gotelli. 2002. Ant diversity in pitcher-plant bogs of Massachussetts. Northeastern Naturalist 9(3): 267-284.
 * Ellison A. M., S. Record, A. Arguello, and N. J. Gotelli. 2007. Rapid Inventory of the Ant Assemblage in a Temperate Hardwood Forest: Species Composition and Assessment of Sampling Methods. Environ. Entomol. 36(4): 766-775.
 * Ellison A. M., and E. J. Farnsworth. 2014. Targeted sampling increases knowledge and improves estimates of ant species richness in Rhode Island. Northeastern Naturalist 21(1): NENHC-13NENHC-24.
 * Feener D.H.,Jr. 1987. Response of Pheidole morrisi to two species of enemy ants, and a general model of defense behavior in Pheidole. Journal of the Kansas Entomological Society 60: 569-575.
 * Forster J.A. 2005. The Ants (hymenoptera: Formicidae) of Alabama. Master of Science, Auburn University. 242 pages.
 * Frye J. A., T. Frye, and T. W. Suman. 2014. The ant fauna of inland sand dune communities in Worcester County, Maryland. Northeastern Naturalist, 21(3): 446-471.
 * General D. M., and L. C. Thompson. 2011. New Distributional Records of Ants in Arkansas for 2009 and 2010 with Comments on Previous Records. Journal of the Arkansas Academy of Science 65: 166-168.
 * General D., and L. Thompson. 2008. Ants of Arkansas Post National Memorial: How and Where Collected. Journal of the Arkansas Academy of Science 62: 52-60.
 * General D.M. & Thompson L.C. 2007. Ants (Hymenoptera: Formicidae) of Arkansas Post National Memorial. Journal of the Arkansas Acaedemy of Science. 61: 59-64
 * Glasier J. R. N., S. E. Nielsen, J. Acorn, and J. Pinzon. 2019. Boreal sand hills are areas of high diversity for Boreal ants (Hymenoptera: Formicidae). Diversity 11, 22; doi:10.3390/d11020022.
 * Gregg E. V. 1945. A statistical study of taxonomic categories in ants (Formicidae: Lasius neoniger and Lasius americanus). Annals of the Entomological Society of America 38: 529-548.
 * Gregg R. E. 1946. The ants of northeastern Minnesota. American Midland Naturalist 35: 747-755.
 * Gregg, R.T. 1963. The Ants of Colorado.
 * Guénard B., K. A. Mccaffrey, A. Lucky, and R. R. Dunn. 2012. Ants of North Carolina: an updated list (Hymenoptera: Formicidae). Zootaxa 3552: 1-36.
 * Headley A. E. 1943. The ants of Ashtabula County, Ohio (Hymenoptera, Formicidae). The Ohio Journal of Science 43(1): 22-31.
 * Heithaus R. E., and M. Humes. 2003. Variation in Communities of Seed-Dispersing Ants in Habitats with Different Disturbance in Knox County, Ohio. OHIO J. SCI. 103 (4): 89-97.
 * Herbers J. M. 2011. Nineteen years of field data on ant communities (Hymenoptera: Formicidae): what can we learn. Myrmecological News 15: 43-52.
 * Herbers J. N. 1989. Community structure in north temperate ants: temporal and spatial variation. Oecologia 81: 201-211.
 * Hoey-Chamberlain R. V., L. D. Hansen, J. H. Klotz and C. McNeeley. 2010. A survey of the ants of Washington and Surrounding areas in Idaho and Oregon focusing on disturbed sites (Hymenoptera: Formicidae). Sociobiology. 56: 195-207
 * Holway D.A. 1998. Effect of Argentine ant invasions on ground-dwelling arthropods in northern California riparian woodlands. Oecologia. 116: 252-258
 * Ipser R. M. 2004. Native and exotic ants (Hymenoptera: Formicidae) of Georgia: Ecological Relationships with implications for development of biologically-based management strategies. Doctor of Philosophy thesis, University of Georgia. 165 pages.
 * Ipser, R.M., M.A. Brinkman, W.A. Gardner and H.B. Peeler. 2004. A Survey of Ground-Dwelling Ants (Hymenoptera: Formicidae) in Georgia. The Florida Entomologist 87(3) 253-260.
 * Ivanov, K. 2019. The ants of Ohio (Hymenoptera, Formicidae): an updated checklist. Journal of Hymenoptera Research 70: 65–87.
 * Ivanov K., L. Hightower, S. T. Dash, and J. B. Keiper. 2019. 150 years in the making: first comprehensive list of the ants (Hymenoptera: Formicidae) of Virginia, USA. Zootaxa 4554 (2): 532–560.
 * Jeanne R. J. 1979. A latitudinal gradient in rates of ant predation. Ecology 60(6): 1211-1224.
 * Johnson C. 1986. A north Florida ant fauna (Hymenoptera: Formicidae). Insecta Mundi 1: 243-246
 * Johnson R. Personnal Database. Accessed on February 5th 2014 at http://www.asu.edu/clas/sirgtools/resources.htm
 * Judd, W.W. 1962. Insects and other invertebrates from nest of the Cardinal, Richmondena cardinalis (L.) at London, Ontario. Canadian Entomologist 94(1):92-95
 * Kjar D. 2009. The ant community of a riparian forest in the Dyke Marsh Preserve, Fairfax County, Virginiam and a checklist of Mid-Atlantic Formicidae. Banisteria 33: 3-17.
 * Kjar D., and E. M. Barrows. 2004. Arthropod community heterogeneity in a mid-Atlantic forest highly invaded by alien organisms. Banisteria 23: 26-37.
 * Kjar D., and Z. Park. 2016. Increased ant (Hymenoptera: Formicidae) incidence and richness are associated with alien plant cover in a small mid-Atlantic riparian forest. Myrmecological News 22: 109-117.
 * Knowlton G. F. 1970. Ants of Curlew Valley. Proceedings of the Utah Academy of Sciences, Arts and Letters 47(1): 208-212.
 * La Rivers I. 1968. A first listing of the ants of Nevada. Biological Society of Nevada, Occasional Papers 17: 1-12.
 * Lavigne R., and T. J. Tepedino. 1976. Checklist of the insects in Wyoming. I. Hymenoptera. Agric. Exp. Sta., Univ. Wyoming Res. J. 106: 24-26.
 * Lesica P., and P. B. Kannowski. 1998. Ants Create Hummocks and Alter Structure and Vegetation of a Montana Fen. Am. Midl. Nat. 139: 5868
 * Lessard J. P., R. R. Dunn, C. R. Parker, and N. J. Sanders. 2007. Rarity and Diversity in Forest Ant Assemblages of Great Smoky Mountains National Park. Southeastern Naturalist 1: 215-228.
 * Lessard, J.-P., R. R. Dunn and N. J. Sanders. 2009. Temperature-mediated coexistence in temperate forest ant communities. Insectes Sociaux 56(2):149-456.
 * Longino, J.T. 2010. Personal Communication. Longino Collection Database
 * Lopez R., and D. A. Potter. 2003. Biodiversity of ants (Hymenoptera: Formicidae) in golf course and lawn turf habitats in Kentucky. Sociobiology 42(3): 701-713.
 * Lubertazi, D. Personal Communication. Specimen Data from Museum of Comparative Zoology at Harvard
 * Lynch J. F. 1981. Seasonal, successional, and vertical segregation in a Maryland ant community. Oikos 37: 183-198.
 * Lynch J. F. 1988. An annotated checklist and key to the species of ants (Hymenoptera: Formicidae) of the Chesapeake Bay region. The Maryland Naturalist 31: 61-106
 * Lynch J. F., and A. K. Johnson. 1988. Spatial and temporal variation in the abundance and diversity of ants (Hymenoptera: Formicidae) in the soild and litter layers of a Maryland forest. American Midland Naturalist 119(1): 31-44.
 * MacGown J. A., J. G. Hill, R. L. Brown, T. L. Schiefer, J. G. Lewis. 2012. Ant diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi. Mississippi Agricultural and Forestry Experiment Station Bulletin 1197: 1-30
 * MacGown J. A., J. G. Hill, and M. Deyrup. 2009. Ants (Hymenoptera: Formicidae) of the Little Ohoopee River Dunes, Emanuel County, Georgia. J. Entomol. Sci. 44(3): 193-197.
 * MacGown J. A., J. G. Hill, and R. L. Brown. 2010.  Native and exotic ant in Mississippi state parks.  Proceedings:  Imported Fire Ant Conference, Charleston, South Carolina, March 24-26, 2008: 74-80.
 * MacGown J. A., and R. L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
 * MacGown, J.A. and JV.G. Hill. Ants of the Great Smoky Mountains National Park (Tennessee and North Carolina).
 * MacGown, J.A. and R.L. Brown. 2006. Survey of the ants (Hymenoptera: Formicidae) of the Tombigbee National Forest in Mississippi. Journal of the Kansas Entomological Society 79(4):325-340.
 * MacGown, J.A., J.G. Hill, R.L. Brown and T.L. 2009. Ant Diversity at Noxubee National Wildlife Refuge in Oktibbeha, Noxubee, and Winston Counties, Mississippi Report #2009-01. Schiefer. 2009.
 * MacGown. J. 2011. Ants collected during the 25th Annual Cross Expedition at Tims Ford State Park, Franklin County, Tennessee
 * Macgown J. A., S. Y. Wang, J. G. Hill, and R. J. Whitehouse. 2017. A List of Ants (Hymenoptera: Formicidae) Collected During the 2017 William H. Cross Expedition to the Ouachita Mountains of Arkansas with New State Records. Transactions of the American Entomological Society, 143(4): 735-740.
 * Mackay, W. P., Lowrie, D., Fisher, A., Mackay, E. E., Barnes, F. 1988. The ants of Los Alamos County, New Mexico (Hymenoptera: Formicidae). Pp. 79-131 in: Trager, J. C. (ed.) Advances in myrmecology. Leiden: E. J. Brill, xxvii + 551 pp.
 * Mackay W. P. and Mackay, E. E. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
 * Mahon M. B., K. U. Campbell, and T. O. Crist. 2017. Effectiveness of Winkler litter extraction and pitfall traps in sampling ant communities and functional groups in a temperate forest. Environmental Entomology 46(3): 470–479.
 * Mann H. R., E. Rowe, J. Selfridge, and D. L. Price. 2018. Leaf litter and arboreal ants (Hymenoptera: Formicidae) in a Mid-Atlantic Forest. Northeastern Naturalist 25(2): 341-354.
 * Martelli, M.G., M.M. Ward and Ann M. Fraser. 2004. Ant Diversity Sampling on the Southern Cumberland Plateau: A Comparison of Litter Sifting and Pitfall Trapping. Southeastern Naturalist 3(1): 113-126
 * Matthewman, W.G. and D.P. Pielou. 1971. Arthropods inhabiting the sporophores of Fomes fomentarius (Polyporaceae) in Gatineau Park, Quebec. Canadian Entomologist 103(6):775-847
 * Menke S. B., E. Gaulke, A. Hamel, and N. Vachter. 2015. The effects of restoration age and prescribed burns on grassland ant community structure. Environmental Entomology http://dx.doi.org/10.1093/ee/nvv110
 * Menke S. B., and N. Vachter. 2014. A comparison of the effectiveness of pitfall traps and winkler litter samples for characterization of terrestrial ant (Formicidae) communities in temperate savannas. The Great Lakes Entomologist 47(3-4): 149-165.
 * Merle W. W. 1939. An Annotated List of the Ants of Maine (Hymenoptera: Formicidae). Entomological News. 50: 161-165
 * Nuhn, T.P. and C.G. Wright. 1979. An Ecological Survey of Ants (Hymenoptera: Formicidae) in a Landscaped Suburban Habitat. American Midland Naturalist 102(2):353-362
 * Ouellette G. D. and A. Francoeur. 2012. Formicidae [Hymenoptera] diversity from the Lower Kennebec Valley Region of Maine. Journal of the Acadian Entomological Society 8: 48-51
 * Ouellette G. D., F. A. Drummond, B. Choate and E. Groden. 2010. Ant diversity and distribution in Acadia National Park, Maine. Environmental Entomology 39: 1447-1556
 * Piers H. 1922. List of a small collection of ants (Formicidae) obtained in Queen's County, Nova Scotia, by the late Walter H. Prest. Proceedings and Transactions of the Nova Scotian Institute of Science, 15(4), 169-173.
 * Procter W. 1938. Biological survey of the Mount Desert Region. Part VI. The insect fauna. Philadelphia: Wistar Institute of Anatomy and Biology, 496 pp.
 * Raczkowski, J.M. and G.M. Luque. 2011. Colony founding and social parasitism in Lasius (Acanthomyops). Insectes Sociaux 58:237-244
 * Rees D. M., and A. W. Grundmann. 1940. A preliminary list of the ants of Utah. Bulletin of the University of Utah, 31(5): 1-12.
 * Sharplin, J. 1966. An annotated list of the Formicidae (Hymenoptera) of Central and Southern Alberta. Quaetiones Entomoligcae 2:243-253
 * Shik, J., A. Francoeur and C. Buddle. 2005. The effect of human activity on ant species (Hymenoptera: Formicidae) richness at the Mont St. Hilaire Biosphere Reserve, Quebec. Canadian Field-Naturalist 119(1): 38-42.
 * Smith M. R. 1962. A new species of exotic Ponera from North Carolina (Hymenoptera, Formicidae). Acta Hymenopterologica 1: 377-382.
 * Sturtevant A. H. 1931. Ants collected on Cape Cod, Massachusetts. Psyche (Cambridge) 38: 73-79
 * Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
 * Toennisson T. A., N. J. Sanders, W. E. Klingeman, and K. M. Vail. 2011. Influences on the Structure of Suburban Ant (Hymenoptera: Formicidae) Communities and the Abundance of Tapinoma sessile. Environ. Entomol. 40(6): 1397-1404.
 * Van Pelt A. F. 1966. Activity and density of old-field ants of the Savannah River Plant, South Carolina. Journal of the Elisha Mitchell Scientific Society 82: 35-43.
 * Van Pelt A., and J. B. Gentry. 1985. The ants (Hymenoptera: Formicidae) of the Savannah River Plant, South Carolina. Dept. Energy, Savannah River Ecology Lab., Aiken, SC., Report SRO-NERP-14, 56 p.
 * Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
 * Wang C., J. Strazanac and L. Butler. 2000. Abundance, diversity and activity of ants (Hymenoptera: Formicidae) in oak-dominated mixed Appalachian forests treated with microbial pesticides. Environmental Entomology. 29: 579-586
 * Ward P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55: 1-16
 * Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
 * Wheeler G. C., J. N. Wheeler, and P. B. Kannowski. 1994. Checklist of the ants of Michigan (Hymenoptera: Formicidae). The Great Lakes Entomologist 26(4): 297-310
 * Wheeler G. C., and E. W. Wheeler. 1944. Ants of North Dakota. North Dakota Historical Quarterly 11:231-271.
 * Wheeler G. C., and J. Wheeler J. 1989. A checklist of the ants of Oklahoma. Prairie Naturalist 21: 203-210.
 * Wheeler G. C., and J. Wheeler. 1986. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp.
 * Wheeler G. C., and J. Wheeler. 1987. A Checklist of the Ants of South Dakota. Prairie Nat. 19(3): 199-208.
 * Wheeler J. N., G. C. Wheeler, R. J. Lavigne, T. A. Christiansen, and D. E. Wheeler. 2014. The ants of Yellowstone National Park. Lexington, Ky. : CreateSpace Independent Publishing Platform, 2013. 112 pages.
 * Wheeler W. M. 1905. Ants from the summit of Mount Washington. Psyche (Cambridge) 12: 111-114.
 * Wheeler W. M. 1906. Fauna of New England. 7. List of the Formicidae. Occasional Papers of the Boston Society of Natural History 7: 1-24
 * Wheeler W. M. 1906. Fauna of New England. 7. List of the Formicidae. Occasional Papers of the Boston Society of Natural History 7: 1-24.
 * Wheeler W. M. 1917. A list of Indiana ants. Proceedings of the Indiana Academy of Science 26: 460-466.
 * Wheeler W. M. 1917. The mountain ants of western North America. Proceedings of the American Academy of Arts and Sciences 52: 457-569.
 * Wheeler, G.C. and J. Wheeler. 1978. Mountain ants of Nevada. Great Basin Naturalist 35(4):379-396
 * Wheeler, G.C. and J. Wheeler. 1988. A checklist of the ants of Montana. Psyche 95:101-114
 * Wheeler, G.C. and J. Wheeler. 1988. A checklist of the ants of Wyoming. Insecta Mundi 2(3&4):230-239
 * Wheeler, G.C., J. Wheeler and P.B. Kannowski. 1994. CHECKLIST OF THE ANTS OF MICHIGAN (HYMENOPTERA: FORMICIDAE). Great Lakes Entomologist 26:1:297-310
 * Wheeler, G.C., J. Wheeler, T.D. Galloway and G.L. Ayre. 1989. A list of the ants of Manitoba. Proceedings of the Entomological Society of Manitoba 45:34-49
 * Wilson E. O. 1955. A monographic revision of the ant genus Lasius. Bulletin of the Museum of Comparative Zoology 113: 1-201
 * Wing M. W. 1939. An annotated list of the ants of Maine (Hymenoptera: Formicidae). Entomological News 50:161-165.
 * Yitbarek S., J. H. Vandermeer, and D. Allen. 2011. The Combined Effects of Exogenous and Endogenous Variability on the Spatial Distribution of Ant Communities in a Forested Ecosystem (Hymenoptera: Formicidae). Environ. Entomol. 40(5): 1067-1073.
 * Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
 * Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71