Dolichoderus quadripunctatus

This is a moderately common species in Greece, known from most provinces except Cyclades. It prefers shadow habitats, close to streams and rivers. In Achaia, it was observed in rural sites in urban areas and on a large plane tree (Borowiec & Salata, 2021). Pashaei Rad et al. (2018) found this species in Iran on the ground and on trees in a Caspian moist littoral area.

Distribution
This is an uncommon species although known from almost all Greek provinces except Cyclades (Borowiec et al., 2022).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Liechtenstein, Lithuania, Luxembourg, Monaco, Montenegro, Norway, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine.

Biology
Czechowski et al. (2019) observed an alate queen of Dolichoderus quadripunctatus visiting aphids in a Polish garden. The gyne walked along a foraging trail to a colony of the aphid Panaphis juglandis where it imbibed honeydew excreted on the leaf by the aphids, after which it returned to the trail. This recurred during two more days, always a single alate gyne at a time; hence the total number of gynes, one or more, remains an open question. This feeding phenomenon is practically unknown in ants.

Nomenclature

 *  quadripunctatus. Formica quadripunctata Linnaeus, 1771: 541 (w.) FRANCE.
 * [Note: type-locality is now France, after Fabricius, 1775: 392, “Habitat in Alsatia”; the latter confirmed by Latreille, 1802c: 181, “Linnée avoit reçu cette espèce de l’Alsace”.]
 * [Misspelled as quatuorpunctata by Christ, 1791: 513; misspelled as 4 punctatus by Forel, 1895d: 227, and others.]
 * Latreille, 1802c: 179 (q.); Mayr, 1855: 380 (m.); Karavaiev, 1916: 497 (m.).
 * Combination in Tapinoma: Schenck, 1852: 129.
 * Combination in Hypoclinea: Mayr, 1855: 379; Mayr, 1863: 424; Roger, 1863b: 14.
 * Combination in Formica (Hypoclinea): Roger, 1859: 242.
 * Combination in Dolichoderus: Emery & Forel, 1879: 455.
 * Combination in D. (Hypoclinea): Emery, 1894c: 229.
 * Status as species: Fabricius, 1775: 392; Fabricius, 1782: 490; Fabricius, 1787: 308; Gmelin, 1790: 2799; Christ, 1791: 513; Olivier, 1792: 494; Fabricius, 1793: 355; Latreille, 1802c: 179; Walckenaer, 1802: 163; Fabricius, 1804: 403; Losana, 1834: 322; Mayr, 1855: 379 (redescription); Nylander, 1856b: 73; Smith, F. 1858b: 57; Roger, 1859: 242; Mayr, 1861: 40 (in key); Mayr, 1863: 424; Roger, 1863b: 14; Emery, 1869b: 11; Mayr, 1870b: 957 (in key); Forel, 1874: 59 (in key); André, 1874: 186 (in key); Emery, 1878b: 49; Emery & Forel, 1879: 455; André, 1882b: 226 (in key); Mayr, 1886d: 435 (in key); Forel, 1886e: clxvii; Nasonov, 1889: 27; Forel, 1892i: 307; Dalla Torre, 1893: 160; Emery, 1894c: 229; Forel, 1895d: 227; Ruzsky, 1902d: 18; Ruzsky, 1903b: 308; Ruzsky, 1905b: 468; Wasmann, 1906: 115 (in key); Bondroit, 1910: 489; Forel, 1911d: 350; Emery, 1913a: 11; Stitz, 1914: 79; Forel, 1915d: 41 (in key); Emery, 1916b: 210; Karavaiev, 1916: 497; Escherich, 1917: 329; Bondroit, 1918: 87; Menozzi, 1921: 32; Soudek, 1922: 58; Müller, 1923b: 108; Soudek, 1925b: 15; Santschi, 1926f: 287; Karavaiev, 1927c: 272 (in key); Lomnicki, 1928: 6; Kuznetsov-Ugamsky, 1929b: 42; Zimmermann, 1935: 40; Karavaiev, 1936: 165 (redescription); Teranishi, 1940: 19; Novák & Sadil, 1941: 95 (in key); Schmitz, 1950: 14; Consani & Zangheri, 1952: 42; Ceballos, 1956: 309; Bernard, 1967: 250 (redescription); Baroni Urbani, 1971c: 157; van Pisarski, 1975: 27; Boven, 1977: 124; Kutter, 1977c: 172; Arnol’di & Dlussky, 1978: 546 (in key); Collingwood, 1978: 86 (in key); Agosti & Collingwood, 1987b: 278 (in key); Atanassov & Dlussky, 1992: 186; Arakelian, 1994: 71; Radchenko, 1994b: 112 (in key); Shattuck, 1994: 63; Bolton, 1995b: 176; Czechowski, et al. 2002: 13; Seifert, 2007: 252; Casevitz-Weulersse & Galkowsky, 2009: 478; Lapeva-Gjonova, et al. 2010: 31; Boer, 2010: 14; Legakis, 2011: 23; Borowiec, L. & Salata, 2012: 491; Czechowski, et al. 2012: 63; Borowiec, L. 2014: 70 (see note in bibliography); Radchenko, 2016: 82.
 * Senior synonym of kratochvili: Radchenko, 2016: 82.
 * Senior synonym of unicolor: Arakelian, 1994: 72; Radchenko, 2016: 82.
 * kratochvili. Dolichoderus (Hypoclinea) quadripunctatus var. kratochvili Novak, V. 1941: 47, figs. 2-4 (w.) CZECHOSLOVAKIA.
 * Subspecies of quadripunctatus: Novák & Sadil, 1941: 95 (in key) (date of publication (31.xii).1941); Shattuck, 1994: 63.
 * Junior synonym of quadripunctatus: Radchenko, 2016: 82.
 * unicolor. Dolichoderus quadripunctatus var. unicolor Ruzsky, 1905b: 471 (w.q.) CAUCASUS.
 * Senior synonym of unicolor: Arakelian, 1994: 72; Radchenko, 2016: 82.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Alvarado M., and L. Galle. 2000. Ant assemblages associated with lowland forests in the southern part of the great Hungarian plain. Acta Zoologica Academiae Scientarum Hungaricae 46(2): 79-102.
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Antonova V., and L. Penev. 2008. Classification of assemblages of ants in the green areas in Sofia City. Acta Zoologica Bulgarica 60(2): 103-110.
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * Banert P, and B. Pisarski. 1972. Mrówki (Formicidae) Sudetów. Fragmenta Faunistica (Warsaw) 18: 345-359.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Bernadou A., V. Fourcassié, and X. Espadaler. 2013. A preliminary checklist of the ants (Hymenoptera, Formicidae) of Andorra. Zookeys 277: 13-23.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Blatrix R., P. Wednez, T. Colin, and C. Galkowski. 2017. Neuf nouvelles espèces de fourmis pour la Corse (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 26(2): 60-64.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. Ants of the Peloponnese, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology 86: 193-236.
 * Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) of Samos Island, Greece. Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-13.
 * Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko G., K. Kiran, C. Karaman, S. Salata, and L. Borowiec. 2016. Survey of the ants (Hymenoptera: Formicidae) of the Greek Thrace. Biodiversity Data Journal 4: e7945. doi: 10.3897/BDJ.4.e7945
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood, C. A.. "A provisional list of Iberian Formicidae with a key to the worker caste." EOS (Revista española de entomología) Nº LVII (1978): 65-95.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S., B. Markó, and L. Gallé. 2001. Ants (Hymenoptera: Formicidae) of Stana Valley (Romania): Evaluation of the effectiveness of a myrmecological survey.  Entomologica Romanica 6 : 121-126.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Diniz, M. A.. "Estado actual do conhecimento dos himenópteros de Portugal." Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 259 (1959): 1-42.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Dvorak, L., P. BOGUSCH, I. MALENOVSKÝ, P. BEZDÌÈKA, K. BEZDÌÈKOVÁ, K. HOLÝ, P. LIKA, J. MACEK, L. ROLLER, M. RÍHA et al. "Hymenoptera of Hády Hill, near the city of Brno (Czech Republic), collected during the Third Czech-Slovak Hymenoptera meeting." Acta Musei Moraviae, Scientiae biologicae (Brno) 93 (2008): 53-92.
 * Emery C. 1869. Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli con descrizioni di specie nuove o meno conosciute. Ann. Accad. Aspir. Nat. Secunda Era 2: 1-26.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Espadaler X., F. Garcia, X. Roig, and R. Vila. 2013. Ants (Hymenoptera, Formicidae) from the Castell de Montesquiu park (Osona, north-east of the Iberian Peninsula). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 53: 223-227.
 * Espadaler X., M. Boieiro, R. Azedo, C. Bernardes, D. Figueiredo, and A. R. M. Serrano. 2008. Additions to the ant fauna of Portugal (Hymenoptera, Formicidae). Boletin Sociedad Entomologica Aragonesa 42: 349-351.
 * Espadaler, X., A. R. Azedo, C. Bernardes, D. Figueiredo, A. R. M. Serrano, and M. Boeiro. "Additions to the Ant fauna of Portugal (Hymenoptera: Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 349-351.
 * Espadaler, X., and J. L. Nieves. "Hormigas (Hymenoptera, Formicidae) pobladoras de agallas abandonadas de cinípidos (Hymenoptera, Cynipidae) sobre Quercus sp. en la Península Ibérica." Boletín de la Estación Central de Ecología 12 (1983): 89-93.
 * Espadaler, X., and X. Roig. "Ants from the Montnegre-Corredor Natural Park with description of the male lasius cinereus Seifert." Miscellanea Zoologica 23 (2) (2001): 45-53. Abstract
 * Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants  assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
 * Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel, A.. "Nouvelles fourmis de Grèce récoltées par M. E. von Oertzen." Annales de la Société Entomologique de Belgique 30 (1886): clix-clxviii.
 * Forel, A.. "Südpalaearctische Ameisen." Mitteilungen der Schweizerischen Entomologischen Gessellschaft 9 (1895): 227-234.
 * Francois J. 1958. Contribution a l'etude ecologique des Formicides (Insectes, Hymenopteres) de la region Dijonnaise. Travaux du laboratoire de Zoologie et de la Station Aquicole Grimaldi de la Faculte des Sciences de Dijon 25, 39 pages.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
 * Galle L., and G. Szonyi. 1988. A check list of ants (Hymenoptera: Formicoidea) of a sandy grassland in Kiskunsag National Park (Hungary). Acta Biol. Szeged 34: 167-168.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Glaser F., T. Kopf, and K. H. Steiberger. 2003. Ameisen (Hymenoptera: Formicidae) im Frastanzer Ried und den Illauen (Vorarlberg, Österreich) Artenspektrum, Gefährdung und Schutzempfehlungen. Vorarlberger Naturschau 13: 287-310.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Hosoishi, S., S. Yamane and K. Ogata. 2010. Subterranean species of the ant genus Crematogaster in Asia (Hymenoptera: Formicidae). Entomological Science 13:345-350.
 * IZIKO South Africa Museum Collection
 * Jaffe K., P. Horchler, M. Verghaagh, C. Gomez, R. Sievert, R. Jaffe, and W. Morawetz. 2007. Comparing the ant fauna in a tropical and a temperate forest canopy. Ecotropicos 20(2): 74-81.
 * Jaffe, K., et al. 2007. Comparing the ant fauna in a tropical and a temperat forest canopy. Ecotropicos 20(2):74-81
 * Karaman M. G. 2001. Contribution to the knowledge of the ants (Hymenoptera: Formicidae) of Demir Kapija Crag (Vardar River, Macedonia), second contribution. Protection of nature 53(2): 49-61.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karaman M. G., and G. S. Karaman. 2007. Contribution to the Knowledge of the ants (Hymenoptera, Formicidae) from Special nature reserve Zasavica, Serbia. Zbornik “Naucno-strucni skup Zasavica 2007, Sremska Mitrovica, 67-75.
 * Karavaiev V. 1926. Beiträge zur Ameisenfauna des Kaukasus, nebst einigen Bemerkungen über andere palaearktische Formen. (Schluss). Konowia 5: 187-199.
 * Karavaiev V. 1927. Übersicht der Ameisenfauna der Krim nebst einigen Neubeschreibungen. Konowia 5: 281-303.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Kiss K., and K. Fetyko. 2008. Notes about the ant fauna (Hymenoptera: Formicidae) of the Tur valley. In. Sike T., Márk-Nagy J. (eds), The flora and fauna of the Tur river natural reserve, Biharean Biologist, II, Oradea, pp.71-76.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Korlevic, A.. "Prilozi fauni hrvatskih opnokrilaca." Glasn. Hrv. Narav. Dr. 5 (1890): 189-250.
 * Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur  Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
 * Lapeva-Gjonova A., and K. Kiran. 2012. Ant fauna (Hymenoptera, Formicidae) of Strandzha (Istranca) Mountain and adjacent Black Sea coast. North-western journal of Zoology 8(1): 72-84.
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis Collection Database
 * Lenoir A. 1971. Les fourmis de Touraine, leur intérêt biogéographique. Cahiers des Naturalistes 27: 21-30.
 * Lomnicki J. 1928. Spis mrówek Lwowa i okolicy. Ksiegi Pamiatkowej (Lecia Gimn. IV Jana Dlugosza Lwowie) 50: 1-10.
 * Majzlan O., and P. Devan. 2009. Selected insect groups (Hymenoptera, Neuroptera, Mecoptera, Raphidioptera) of the Rokoš Massif (Strážovské vrchy Mts.). Rosalia (Nitra), 20, p. 63–70.
 * Marikovsky P. I. 1979. Ants of the Semireche Desert. [In Russian.]. Alma Ata: Nauka, 263 pp.
 * Marko B. 1999. Contribution to the knowledge of the myrmecofauna of the River Somes valley. In: Sárkány-Kiss, A., Hamar, J. (szerk.): The Somes/Szamos River Valley. A study of the geography, hydrobiology and ecology of the river system and its environment. Tiscia Monograph Series 3, Szolnok-Szeged-Tîrgu Mure?, pp. 297-302.
 * Marko B. 2008. Ants (Hymenoptera: Formicidae) of the Sand Dunes of Foieni Protected Area and Its Surroundings (Satu Mare County, Romania), and a New Species for the Romanian Fauna. Acta Scientiarum Transylvanica 16(3): 87-99.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Markó B., and S. Csősz. 2002. Die europäischen Ameisenarten (Hymenoptera: Formicidae) des Hermannstädter (Sibiu, Rumänien) Naturkundemuseums I.: Unterfamilien Ponerinae, Myrmicinae und Dolichoderinae. Annales Historico-Naturales Musei Nationalis Hungarici 94: 109-121.
 * Menke, S.B. Personal Communication. Field Museum Specimen Data
 * Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
 * Nemet E., Z. Czekes, I. Tausan, and B. Marko. 2012. Contribution to the knowledge of the myrmecofauna of the Cefa Nature Park (North-Western Romania). Acta Scientiarum Transylvanica 20(1): 61-72.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Nylander, W.. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * Odegaard F., K. M. Olsen, A. Staverlokk, and J. O. Gjershaug. 2015. Towards a new era for the knowledge of ants (Hymenoptera, Formicidae) in Norway? Nine species new to the country. Nowegian Journal of Entomology 62: 80–99.
 * Paknia, O., A. G. Radchenko, and M. Pfeiffer. "New records of ants (Hymenoptera: Formicidae) from Iran." Asian Myrmecology 3, no. 29-38 (2010).
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Pashaei Rad S., B. Taylor, R. Torabi, E. Aram, G. Abolfathi, R. Afshari, F. Borjali, M. Ghatei, F. Hediary, F. Jazini, V. Heidary Kiah, Z. Mahmoudi, F. Safariyan, and M. Seiri. 2018. Further records of ants (Hymenoptera: Formicidae) from Iran. Zoology in the Middle East 64(2): 145-159.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z. 2007. Contribution to knowledge of the myrmecofauna (Formicidae, Hymenoptera) of Belgrade, Serbia. Archives of Biological Sciences 59: 11-12.
 * Petrov I. Z. 2010. Contribution to the myrmecofauna (Hymenoptera: Formicidae) of Krusevac and its surroundings. Acta entomologica serbica, 2010, 15(1): 145-148.
 * Petrov I. Z., and A. Legakis. 1996. Contribution to the Myrmecofauna (Formicidae, Hymenoptera) of Greece. Archives of biological sciences, Belgrade, 48 (3-4): 31-32.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Poldi B. 1980. Imenotteri formicidi della Brughiera di Rovasenda (Piemonte). In: Poldi, B. 1980. Quaderni sulla "Struttura delle zoocenosi terrestri". 1. La brughiera pedemontana. III. Serie AQ/1/110. Collana del programma finalizzato "Promozione della qualità dell'ambiente". Roma: Consiglio Nazionale delle Ricerche, pp. 7-19.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
 * Salata S., P. Zurawlew, and J. K. Kowalczyk. 2018. New distribution records of several ant species (Hymenoptera: Formicidae) in Poland. Wiadomosci Entomologiczne 37(1): 46-53.
 * Salata S., and L Borowiec. 2017. Species of Tetramorium semilaeve complex from Balkans and western Turkey, with description of two new species of (Hymenoptera: Formicidae: Myrmicinae). Annales Zoologici (Warsaw) 62:279–313.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Salata S., and L. Borowiec. 2019. Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species. ZooKeys 877: 81-131.
 * Santschi F. 1910. Contributions à la faune entomologique de la Roumanie. Formicides capturées par Mr. A. L. Montandon. Buletinul Societatii Române de Stiinte 19: 648-652.
 * Santschi, F.. "Liste de fourmis d'Espagne recueilliés par Mr. J. M. Dusmet." Boletín de la Sociedad Entomológica de España 15 (1932): 69-74.
 * Schifani E., and A. Alicata. 2018. Exploring the myrmecofauna of Sicily: thirty-two new ant species recorded, including six new to Italy and many new aliens (Hymenoptera, Formicidae). Polish Journal of Entomology 87 (4): 323–348.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Schmitz H. 1950. Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit. Brotéria Ser. Cienc. Nat. 19: 12-16.
 * Schmitz, H.. "Formicidae quaedam a cl. A. Stärcke determinatae, quas in Lusitania collegit." Brotéria, Ciências Naturais 19 (1950): 12-16.
 * Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
 * Seifert B. 2007. Die Ameisen Mittel- und Nordeuropas. Tauer: lutra Verlags- und Vertriebsgesellschaft, 368 pp.
 * Sellier Y., C. Galkowski, C. Lebas, and P. Wegnez. 2016. Découverte de Temnothorax pardoi (Tinaut, 1987) dans la réserve naturelle nationale du Pinail (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 106-113.
 * Shattuck S. O. 1994. Taxonomic catalog of the ant subfamilies Aneuretinae and Dolichoderinae (Hymenoptera: Formicidae). University of California Publications in Entomology 112: i-xix, 1-241.
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Stukalyuk S. V. 2015. Structure of the ant assemblages (Hymenoptera, Formicidae) in the broad-leaved forests of Kiev. Entomological Review 95(3): 370–387.
 * Stukalyuk S. V. 2017. Stratification of the ant species (Hymenoptera, Formicidae) in the urban broadleaf woodlands of the city of Kiev. Entomological Review 97(3): 320-343.
 * Suvak M. 2007. The ants (Hymenoptera: Formicidae) in the stubble-fields of east Slovakia. Acta Zoologica Universitatis Comenianae. 47: 21-33
 * Tartally A. 2009. Data on the ant fauna of Gy?r?f? (SW-Hungary) (Hymenoptera: Formicidae). Natura Somogyiensis 13: 155-158.
 * Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
 * Torossian C. 1967. Recherches sur la biologie et l'ethologie de Dolichoderus quadripunctatus (L.) Hym. Formicoidea Dolichoderidae. Insectes Sociaux 14(2): 105-122.
 * Vagalinski B., and A. Lapeva-Gjonova. 2012. The ants (Hymenoptera, Formicidae) of Plana Mountain (Bulgaria). Historia naturalis bulgarica 20: 87-101.
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vesnic A., and S. Lelo. 2007. Members of subfamily Dolichoderinae Forel, 1878 (Insecta: Vespoidea, Formicidae) in Bosnia and Herzegovina. Prilozi fauni Bosne i Hercegovine 3: 23-28.
 * Vogrin, V.. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Yasumatsu, K. 1941a. On the ants of the genus Dolichoderus of Angaran element from the Far East (Hymenoptera, Formicidae). [In Japanese.]. Kontyû 14:179. [1941-01-25] PDF 130557
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Andalucia." Boletín de la Estación Central de Ecología 6 (12) (1977): 85-90.
 * del Campo Gracia P., M. D. Martinez Ibanez, A. Tinaut, and S. Montagud Alario. 2014. Faunistic study of the Formicidae (Hymenoptera, Formicidae) from the Comunitat Valenciana (Spain). Boln. Asoc. esp. Ent. 38 (1-2): 33-65.