Checklist of Camponotus species

The following species and subspecies belong to the genus Camponotus. Synonyms are listed under their senior names. For valid names only see Camponotus species and for distribution information see Camponotus species by Country.

A
aberrans Mayr, 1895 (Ghana, Nigeria, Uganda)

abjectus Santschi, 1937 (Angola, Senegal)

abrahami Forel, 1913 (Lebanon, Turkey)

abscisus Roger, 1863 (Brazil, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

absquatulator Snelling, R.R., 2006 (Mexico, United States)

abunanus Mann, 1916 (Brazil)

acutirostris Wheeler, W.M., 1910 (Mexico, United States)

• Camponotus acutirostris clarigaster Wheeler, W.M., 1915

acvapimensis Mayr, 1862 (Benin, Cameroun, Eritrea, Ghana, Ivory Coast, Kenya, Liberia, Nigeria, Senegal, Socotra Archipelago, Togo, United Arab Emirates, Yemen, Zimbabwe)

• Camponotus acwapimensis poultoni Forel, 1913

• Camponotus flavosetosus Donisthorpe, 1945

adami Forel, 1910 (Australia)

adenensis Emery, 1893 (United Arab Emirates, Yemen)

adustus Viehmeyer, 1916 (Singapore)

aegaeus Emery, 1915 (Bulgaria, Greece, North Macedonia, Turkey)

aegyptiacus Emery, 1915 (Benin, Egypt, Eritrea, Guinea, Kenya, Kuwait, Oman, Senegal, Yemen)

• Camponotus maculatus infrasquameus Santschi, 1926

aeneopilosus Mayr, 1862 (Australia)

aeneopilosus flavidopubescens Forel, 1902 (Australia)

aequatorialis Roger, 1863 (Gabon, Guinea)

aequatorialis kohli Forel, 1915 (Democratic Republic of Congo)

aequitas Santschi, 1920 (South Africa, Zimbabwe)

aethiops (Latreille, 1798) (Albania, Andorra, Armenia, Austria, Azerbaijan, Balearic Islands, Bulgaria, China, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Malta, Montenegro, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Saudi Arabia, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Camponotus aethiops concava Dalla Torre, 1893

• Camponotus aethiops sylvaticoides Dalla Torre, 1893

• Camponotus maculatus glaber Emery, 1925

• Camponotus marginatus hyalinipennis Costa, 1884

• Camponotus sylvaticus sylvaticoaethiops Forel, 1874

• Formica angustata Latreille, 1798

• Formica marginata Latreille, 1798

• Formica nigrata Nylander, 1849

aethiops cachmiriensis Emery, 1925 (India)

aethiops escherichi Emery, 1925 (Turkey)

afflatus Viehmeyer, 1925 (Australia)

ager (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Peru, Suriname, Trinidad and Tobago, Venezuela)

agonius Santschi, 1915 (Democratic Republic of Congo, Guinea)

aguilerai Kusnezov, 1952 (Argentina)

aina Rakotonirina & Fisher, 2022 (Madagascar)

aktaci Karaman, C., 2013 (Turkey)

alacer Forel, 1912 (Brazil)

alamaina Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

albicoxis Forel, 1899 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

albipes Emery, 1893 (Sri Lanka)

albistramineus Wheeler, W.M., 1936 (Dominican Republic, Greater Antilles, Haiti)

albivillosus Zhou, 2001 (China)

alboannulatus Mayr, 1887 (Bolivia, Brazil, Venezuela)

alboannulatus montanus Emery, 1894 (Bolivia)

alboannulatus nessus Forel, 1912 (Brazil)

albocinctus (Ashmead, 1905) (New Guinea, Philippines)

albosparsus Bingham, 1903 (China, India, Japan, Singapore)

alii Forel, 1890 (Algeria, United Arab Emirates)

alii auresi Emery, 1920 (Algeria)

alii dallatorrei Özdikmen, 2010 (Algeria)

• Camponotus alii concolor Dalla Torre, 1893

alii hoggarensis Santschi, 1929 (Algeria)

altivagans Wheeler, W.M., 1936 (Greater Antilles, Haiti)

amamianus Terayama, 1991 (Japan, Republic of Korea)

amaurus Espadaler, 1997 (Iberian Peninsula, Spain)

americana (Buckley, 1866)

americanus Mayr, 1862 (United States)

• Camponotus castaneus rufinasis Santschi, 1936

amoris Forel, 1904 (Brazil, Colombia, Peru)

amphidus Santschi, 1926 (Zimbabwe)

anatolicus Karaman, C. & Aktac, 2013 (Turkey)

andrei Forel, 1885 (Mexico)

andrei cholericus Wheeler, W.M., 1914 (Mexico)

andrewsi Donisthorpe, 1936 (Christmas Island)

andrianjaka Rasoamanana & Fisher, 2022 (Madagascar)

androy Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

andyyoungi McArthur, 2008 (Australia)

anezkae Klimes & McArthur, 2014 (New Guinea)

anguliceps Stitz, 1938 (Borneo, Indonesia, Malaysia)

angusticeps Emery, 1886 (South Africa)

angusticollis (Jerdon, 1851) (Bangladesh, Borneo, India, Indonesia, Malaysia, Nepal, Sri Lanka)

• Camponotus prismaticus Mayr, 1862

• Formica ardens Smith, F., 1858

• Formica impetuosa Smith, F., 1858

angusticollis sanguinolentus Forel, 1895 (Borneo, India, Indonesia, Malaysia)

anningensis Wu, J. & Wang, C., 1989 (China)

annulatus Karavaiev, 1929 (Indonesia)

anthrax Wheeler, W.M., 1911 (Mexico, United States)

antsaraingy Rasoamanana & Fisher, 2022 (Madagascar)

apicalis (Mann, 1916) (Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, Trinidad and Tobago, Venezuela)

• Dendromyrmex apicalis filiae Weber, 1943

• Dendromyrmex apicalis guyanensis Santschi, 1921

• Dendromyrmex apicalis opaciceps Wheeler, W.M., 1923

• Dendromyrmex wheeleri Donisthorpe, 1937

apostolus Forel, 1901 (Costa Rica)

arabicus Collingwood, 1985 (Oman, Saudi Arabia)

arboreus (Smith, F., 1858) (Argentina, Brazil, Ecuador, Panama, Paraguay)

• Camponotus adpressisetosus Forel, 1879

arcuatus Mayr, 1876 (Australia)

arcuatus aesopus Forel, 1907 (Australia)

arenatus Shattuck & McArthur, 2002 (Australia)

argus Santschi, 1935 (Democratic Republic of Congo)

arhuacus Forel, 1902 (Colombia)

armeniacus Arnoldi, 1967 (Armenia, Iran, Turkey)

arminius Forel, 1910 (Mozambique, Yemen)

arminius bicontractus Wheeler, W.M., 1922 (South Africa)

armstrongi McAreavey, 1949 (Australia)

arnoldinus Forel, 1914 (South Africa)

aro Rakotonirina & Fisher, 2022 (Madagascar)

arrogans (Smith, F., 1858) (Bangladesh, Borneo, India, Indonesia, Malaysia, Singapore, Thailand)

• Camponotus maculatus junctus Forel, 1892

asara Rakotonirina & Fisher, 2022 (Madagascar)

ashokai Karmaly & Narendran, 2006 (India)

asli Dumpert, 1989 (Malaysia)

aterrimus Emery, 1895 (China, Kazakhstan, Mongolia, Pakistan, Russian Federation)

atimo Rakotonirina & Fisher, 2022 (Madagascar)

atlantis Forel, 1890 (Canary Islands, Iran, Oman, Socotra Archipelago, Tunisia, United Arab Emirates, Yemen)

atlantis atrioris Santschi, 1922 (Algeria)

atlantis maroccanus Emery, 1925 (Morocco)

atlantis nigrovarius Forel, 1894 (Algeria)

atra (Buckley, 1866)

atriceps (Smith, F., 1858) (Argentina, Barbados, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States, Uruguay, Venezuela)

• Camponotus abdominalis costaricensis Forel, 1899

• Camponotus abdominalis cupiens Forel, 1908

• Camponotus abdominalis depressidens Santschi, 1922

• Camponotus abdominalis epistomatus Santschi, 1936

• Camponotus abdominalis fuchsae Forel, 1908

• Camponotus abdominalis fumata Stitz, 1933

• Camponotus abdominalis laevilata Santschi, 1922

• Camponotus abdominalis rabidus Santschi, 1922

• Camponotus abdominalis romani Wheeler, W.M., 1923

• Camponotus abdominalis transvectus Wheeler, W.M., 1910

• Camponotus atriceps atricipitoesuriens Forel, 1879

• Camponotus atriceps stercorarius Forel, 1885

• Camponotus atriceps ustulatus Forel, 1885

• Camponotus taeniatus Roger, 1863

• Camponotus ustulatus mediopallidus Forel, 1901

• Camponotus vulpinus Mayr, 1862

• Formica abdominalis Fabricius Fabricius, 1804

• Formica esuriens Smith, F., 1858

• Formica fulvacea Norton, 1868

atriceps nocens Wheeler, W.M., 1911 (Grenada)

atricolor (Nylander, 1849) (Austria, Azerbaijan, Bulgaria, Greece, Hungary, Romania, Russian Federation, Turkey)

• Camponotus lateralis rectus Forel, 1892

atriscapus Santschi, 1926 (Democratic Republic of Congo)

atrox Emery, 1925 (Democratic Peoples Republic of Korea, Republic of Korea)

• Camponotus herculeanus jeholensis Teranishi, 1936

• Camponotus japonicus cruentata Emery, 1925

augustei Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

auratiacus Zhou, 2001 (China)

aurelianus Forel, 1912 (Indonesia)

aureopilus Viehmeyer, 1914 (New Guinea)

• Camponotus aureopilus velutinus Stitz, 1938

aureus Dumpert, 2006 (Thailand)

auricomus Roger, 1862 (Colombia, French Guiana, Greater Antilles, Guyana, Lesser Antilles, Mexico, Saint Vincent and the Grenadines, Suriname, Trinidad and Tobago)

auricomus lucianus Forel, 1899 (Lesser Antilles)

auricomus vincentensis Forel, 1899 (Lesser Antilles)

auriculatus Mayr, 1897 (Sri Lanka)

auriventris Emery, 1889 (Borneo, Indonesia, Malaysia, Myanmar, Thailand, Vietnam)

aurocinctus (Smith, F., 1858) (Australia)

• Camponotus midas Froggatt, 1896

aurofasciatus Santschi, 1915 (São Tomé & Principe)

auropubens Forel, 1894 (Madagascar, Mozambique, Seychelles, Zimbabwe)

• Camponotus auropubens aldabrensis Forel, 1897

• Camponotus olivieri freyeri Santschi, 1915

auropubens absalon Santschi, 1915 (South Africa)

auropubens argentopubens Santschi, 1915 (Angola)

auropubens jacob Santschi, 1915 (Mozambique)

aurosus Roger, 1863 (Madagascar, Mauritius, Réunion)

autrani Forel, 1886 (Borneo, Indonesia, Malaysia)

avius Santschi, 1926 (Zimbabwe)

avius hertigi Santschi, 1937 (Angola)

B
bakeri Wheeler, W.M., 1904 (United States)

bakhtiariensis Salata, Khalili-Moghadam & Borowiec, 2020 (Iran)

baldaccii Emery, 1908 (Greece, Israel, Saudi Arabia, Turkey)

balzani Emery, 1894 (Argentina, Bolivia, Brazil, Ecuador, Paraguay)

banghaasi Emery, 1903 (Ecuador, Peru)

barbaricus Emery, 1905 (Algeria, Gibraltar, Iberian Peninsula, Malta, Portugal, Spain)

• Camponotus barbaricus baetica Emery, 1924

barbaricus eubarbaricus Cagniant, 1970 (Algeria)

barbaricus xanthomelas Emery, 1905 (Algeria)

barbarossa Emery, 1920 (Cameroun)

barbarossa micipsa Wheeler, W.M., 1922 (Democratic Republic of Congo)

barbarossa sulcatinasis Santschi, 1926 (Democratic Republic of Congo)

barbatus Roger, 1863 (Borneo, India, Indonesia, Malaysia, Philippines, Sri Lanka)

barbatus infuscoides Bingham, 1903 (Sri Lanka)

barbatus samarus Santschi, 1932 (Borneo, Indonesia, Malaysia)

barbatus taylori Forel, 1892 (Bangladesh, China, India, Sri Lanka)

barbosus Baroni Urbani, 1971 (Philippines)

• Camponotus barbatus Menozzi Menozzi, 1926

basuto Arnold, 1958 (Lesotho)

batesii Forel, 1895 (Madagascar)

bayeri Forel, 1913 (Democratic Republic of Congo, Kenya)

baynei Arnold, 1922 (South Africa)

beccarii Emery, 1887 (Indonesia)

becki Santschi, 1923 (Madagascar)

bedoti Emery, 1893 (Borneo, Brunei Darussalam, China, Indonesia, Krakatau Islands, Malaysia, New Guinea, Singapore, Solomon Islands)

beebei Wheeler, W.M., 1918 (Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)

bellacolor Mackay, 2019 (Mexico)

belligerus Santschi, 1920 (Namibia)

bellus Forel, 1908 (Indonesia, Singapore)

• Camponotus ornatus Forel, 1907

belumensis Dumpert, 1995 (Malaysia)

bemaheva Rakotonirina & Fisher, 2022 (Madagascar)

benguelensis Santschi, 1911 (Angola)

benguetensis Zettel & Balàka, 2018 (Philippines)

bermudezi Aguayo, 1932 (Cuba, Greater Antilles)

berthoudi Forel, 1879 (South Africa)

bertolonii Emery, 1895 (Kenya, South Africa)

bevohitra Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

bianconii Emery, 1895 (South Africa)

bidens Mayr, 1870 (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Peru, Suriname, Trinidad and Tobago)

bidens repressus Forel, 1912 (Brazil)

bifossus Santschi, 1917 (South Africa)

bigenus Santschi, 1919 (Australia)

binghamii Forel, 1894 (India, Myanmar)

biolleyi Forel, 1902 (Costa Rica)

bishamon Terayama, 1999 (Japan)

bispinosus Mayr, 1870 (Colombia, Ecuador)

bituberculatus André, 1889 (Guinea, Senegal, Sierra Leone)

blandus (Smith, F., 1858) (Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Guyana, Honduras, Mexico, Paraguay, Peru, Suriname, Uruguay, Venezuela)

blandus aequinotus Forel, 1902 (Venezuela)

blandus denudatus Emery, 1903 (Bolivia)

blandus pronotalis Santschi, 1936 (Panama)

blandus semiceps Emery, 1925 (Costa Rica)

bocki Forel, 1907 (Bolivia)

boghossiani Forel, 1911 (Greece, Turkey)

• Camponotus boghossiani stenoticus Emery, 1915

• Camponotus kiesenwetteri angustatus Forel, 1889

boivini Forel, 1891 (Madagascar, Seychelles)

• Camponotus maculatus fairmairei Santschi, 1911

bonariensis Mayr, 1868 (Argentina, Brazil, Paraguay, Uruguay)

bonariensis curtulus Santschi, 1936 (Brazil)

bonariensis garbei Santschi, 1922 (Brazil)

bonariensis luteolus Emery, 1906 (Argentina)

• Camponotus maculatus carolinus Forel, 1916

bonariensis parvulus Emery, 1894 (Brazil)

bonariensis tucumanus Santschi, 1922 (Argentina)

bonariensis weiseri Santschi, 1922 (Argentina)

borellii Emery, 1894 (Argentina)

borellii saltensis Emery, 1925 (Argentina)

bottegoi Emery, 1895 (Kenya, Somalia)

bozaka Rakotonirina & Fisher, 2022 (Madagascar)

branneri (Mann, 1916) (Brazil, Ecuador)

brasiliensis Mayr, 1862 (Brazil, Paraguay)

brasiliensis chacoensis Santschi, 1919 (Argentina)

brasiliensis clivius Emery, 1925 (Brazil)

brasiliensis costaricanus Özdikmen, 2010 (Costa Rica)

• Camponotus brasiliensis antennatus Santschi, 1925

braunsi Mayr, 1895 (Kenya, Mozambique, Zimbabwe)

braunsi candidus Santschi, 1926 (United Republic of Tanzania)

braunsi epinotalis Santschi, 1916 (Zimbabwe)

braunsi erythromelus Emery, 1896 (Kenya, Somalia)

brettesi Forel, 1899 (Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Trinidad and Tobago)

brettesi canalis Forel, 1912 (Panama)

brevicollis Stitz, 1916 (Congo)

brevis Forel, 1899 (Colombia, Costa Rica, Ecuador, Panama)

brevis obscurifrons Santschi, 1936 (Panama)

brevis pauli Forel, 1908 (Costa Rica)

breviscapus Zhou, 2001 (China)

brevisetosus Forel, 1910 (South Africa)

britteni Donisthorpe, 1931 (United Kingdom of Great Britain and Northern Ireland)

brookei Forel, 1914 (South Africa)

bruchi Forel, 1912 (Argentina, Bolivia)

• Camponotus distinguendus boliviana Stitz, 1933

bruchi lysistrata Santschi, 1919 (Argentina)

bruchi titicacensis Emery, 1920 (Peru)

brullei (Smith, F., 1858) (Canary Islands)

• Formica carinata Brullé, 1840

bruneiensis Viehmeyer, 1922 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

brunni Forel, 1901 (Japan)

brutus Forel, 1886 (Angola, Cameroun, Ivory Coast, Kenya, Sierra Leone)

buchholzi Mayr, 1902 (Cameroun)

buchneri Forel, 1886 (Angola, Cameroun)

• Phasmomyrmex sericeus Stitz, 1910

buchneri camerounensis Ward, Blaimer & Fisher, 2016 (Cameroun)

• Phasmomyrmex buchneri griseus Santschi, 1937

buddhae Forel, 1892 (India, Iran, Kyrgyzstan, Pakistan, Russian Federation, Turkey, Turkmenistan)

• Camponotus fedtschenkoi obliquipilosa Emery, 1920

bugnioni Forel, 1899 (Colombia)

burgeoni Santschi, 1926 (Democratic Republic of Congo)

burtoni Mann, 1916 (Brazil)

butteli Forel, 1905 (Madagascar)

buttikeri Arnold, 1958 (South Africa)

C
cacicus Emery, 1903 (Brazil, Ecuador, Peru)

caesar Forel, 1886 (Angola, Guinea)

caesar imperator Emery, 1899 (Cameroun)

caffer Emery, 1895 (South Africa)

callistus Emery, 1911 (Bolivia, Ecuador)

callistus bradleyi Wheeler, W.M., 1934 (Ecuador, Peru)

callmorphus Stitz, 1923 (Namibia)

camelinus (Smith, F., 1857) (Borneo, India, Indonesia, Laos, Malaysia, Singapore, Thailand)

• Camponotus senilis Mayr, 1862

cameranoi Emery, 1894 (Argentina, Brazil, Paraguay, Peru)

cameranoi eugaster Santschi, 1922 (Brazil)

cameranoi inlex Forel, 1901 (Brazil)

cameratus Viehmeyer, 1925 (Australia)

cameroni Forel, 1892 (China, Ecuador)

• Camponotus mayri Cameron Cameron, 1891

candiotes Emery, 1894 (Azerbaijan, Greece, Turkey)

canescens Mayr, 1870 (Bolivia, Brazil, Colombia, Costa Rica, Panama, Trinidad and Tobago)

canescens antennatus Santschi, 1920 (Bolivia)

canescens stomatus Santschi, 1925 (Panama)

capito Mayr, 1876 (Australia)

capito ebeninithorax Forel, 1915 (Australia)

capperi Forel, 1899 (Greater Antilles, Jamaica)

capperi corticalis Forel, 1899 (Jamaica)

capperi formosulus Wheeler, W.M., 1917 (Jamaica)

capperi subdepilis Wheeler, W.M., 1917 (Jamaica)

capperi unctulus Wheeler, W.M., 1917 (Jamaica)

caracalla Forel, 1912 (Brazil)

carbo Emery, 1877 (Benin, Eritrea, Ethiopia, Oman)

carbo honorus Forel, 1910 (Ethiopia)

carbo occidentalis Mayr, 1902 (Ghana)

carbonarius (Latreille, 1802) (Greater Antilles)

carin Emery, 1889 (Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Sri Lanka)

carin babiensis Emery, 1925 (Indonesia)

carin tenuisquamis Forel, 1901 (Indonesia)

carin tipunus Forel, 1913 (Taiwan)

carinifer Viehmeyer, 1916 (Singapore)

caryae (Fitch, 1855) (China, Mexico, United States)

casicus Santschi, 1920 (French Guiana)

castaneus (Latreille, 1802) (United States)

• Camponotus clarus Mayr, 1862

• Formica mellea Say Say, 1836

castanicola Donisthorpe, 1943 (Philippines)

• Camponotus castanea Stitz, 1925

cecconii Emery, 1908 (Cyprus, Greece, Iran)

cemeryi Özdikmen, 2010 (Madagascar)

• Camponotus hova luteolus Emery, 1925

ceriseipes Clark, 1938 (Australia)

cervicalis Roger, 1863 (Madagascar)

• Camponotus gaullei Santschi, 1911

• Camponotus gerberti Donisthorpe, 1949

• Camponotus perroti Forel, 1897

• Camponotus perroti aeschylus Forel, 1913

chalceoides Clark, 1938 (Australia)

chalceus Crawley, 1915 (Australia)

championi Forel, 1899 (Costa Rica, Mexico, Panama)

chapini Wheeler, W.M., 1922 (Democratic Republic of Congo)

chapini ganzii Weber, 1943 (Sudan)

chartifex (Smith, F., 1860) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Honduras, Nicaragua, Panama, Suriname)

• Dendromyrmex madeirensis melinoni Santschi, 1921

chazaliei Forel, 1899 (Lesser Antilles)

cheesmanae Donisthorpe, 1932 (Colombia)

chilensis (Spinola, 1851) (Argentina, Chile, Peru)

chloroticus Emery, 1897 (Australia, Cook Islands, Fiji, New Caledonia, New Guinea, Niue, Samoa, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Camponotus irritans chlorogaster Emery, 1920

• Camponotus maculatus sanctaecrucis Mann, 1919

chongqingensis Wu, J. & Wang, C., 1989 (China)

chrislaini Rasoamanana & Fisher, 2022 (Madagascar)

christi Forel, 1886 (Madagascar)

christophei Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

christopherseni Forel, 1912 (Guatemala, Panama)

chromaiodes Bolton, 1995 (United States)

• Formica ferruginea Fabricius Fabricius, 1798

chrysurus Gerstäcker, 1871 (Kenya)

chrysurus acutisquamis Mayr, 1902 (Cameroun)

chrysurus apellis Wheeler, W.M., 1922 (Angola)

chrysurus securifer Emery, 1920 (Cameroun)

chrysurus yvonnae Forel, 1920 (Democratic Republic of Congo)

cilicicus Emery, 1908 (Iraq, Israel, Turkey)

cillae Forel, 1912 (Brazil)

cinctellus (Gerstäcker, 1859) (Eritrea, Kenya, Mali, Mozambique, Rwanda, United Republic of Tanzania, Zimbabwe)

• Camponotus venustus Mayr, 1867

cinctellus belliceps Santschi, 1939 (Democratic Republic of Congo)

cinerascens (Fabricius, 1787) (India, Philippines)

cinereus Mayr, 1876 (Australia)

cinereus amperei Forel, 1913 (Australia)

cinereus notterae Forel, 1907 (Australia)

cingulatus Mayr, 1862 (Bolivia, Brazil, Paraguay, Uruguay)

• Camponotus cingulatus bambusarum Forel, 1902

• Camponotus cingulatus brunneiventris Santschi, 1920

• Camponotus cingulatus damocles Forel, 1909

• Camponotus cingulatus fissa Stitz, 1933

• Camponotus cingulatus myster Santschi, 1922

• Camponotus rufipes lessonai Emery, 1894

circularis Mayr, 1870 (Colombia)

circularis rufitibia Emery, 1920 (Peru)

circumspectus (Smith, F., 1861) (Indonesia, Sulawesi)

clarior Forel, 1902 (Australia)

claripes Mayr, 1876 (Australia)

• Camponotus claripes piperatus Wheeler, W.M., 1933

• Camponotus walkeri Forel, 1893

claripes nudimalis Forel, 1913 (Australia)

claripes orbiculatopunctatus Viehmeyer, 1925 (Australia)

clarithorax Creighton, 1950 (Mexico, United States)

claveri Rasoamanana & Fisher, 2022 (Madagascar)

claviscapus Forel, 1899 (Belize, Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Netherlands Antilles, Nicaragua, Panama, Trinidad and Tobago, Venezuela)

claviscapus lehmanni Forel, 1911 (Nicaragua)

claviscapus occultus Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

claviscapus subcarinatus Forel, 1899 (Guatemala)

cleobulus Santschi, 1919 (South Africa)

clypeatus Mayr, 1866 (Brazil)

cocosensis Wheeler, W.M., 1919 (Cocos (Keeling) Islands)

cognatocompressus Forel, 1904 (Iran)

coloratus Forel, 1904 (Argentina, Colombia, Costa Rica, Mexico, Paraguay)

compositor Santschi, 1922 (Bolivia, Brazil)

compressiscapus André, 1889 (Senegal, Sierra Leone)

compressus (Fabricius, 1787) (Bangladesh, Borneo, Canary Islands, China, India, Indonesia, Liechtenstein, Malaysia, Maldives, Nepal, Pakistan, Philippines, Sri Lanka, Turkey, United Arab Emirates)

• Camponotus quadrilaterus Roger Roger, 1863

• Formica callida Smith, F., 1858

• Formica indefessa Sykes, 1835

compressus bernardi Özdikmen, 2010 (Algeria)

• Camponotus compressus nitens Bernard, 1953

compressus martensi Forel, 1907 (Algeria)

compressus occipitalis Stitz, 1917 (Algeria)

compressus probativus Santschi, 1921 (Mozambique)

compressus rectinotus Santschi, 1921 (Libya)

concavus Kim & Kim, 1994 (Greece, Republic of Korea)

concilians Forel, 1915 (Indonesia)

• Camponotus jacobsoni Forel, 1915

concolor Forel, 1891 (Madagascar)

• Camponotus gallienii Forel, 1916

concurrens Zettel & Laciny, 2018 (Borneo, Brunei Darussalam, Malaysia, Thailand)

confluens Forel, 1913 (Democratic Republic of Congo)

confluens bequaerti Forel, 1913 (Democratic Republic of Congo)

confluens trematogaster Santschi, 1915 (Democratic Republic of Congo)

confucii Forel, 1894 (China, India, Pakistan)

confusus Emery, 1887 (Australia, New Guinea)

congolensis Emery, 1899 (Democratic Republic of Congo, Guinea, Ivory Coast, Liberia, Sierra Leone, Zimbabwe)

congolensis weissi Santschi, 1911 (Congo)

coniceps Santschi, 1926 (Congo)

conradti Forel, 1914 (Cameroun)

conradti fimbriatipes Santschi, 1920 (Cameroun)

consanguineus (Smith, F., 1861) (Indonesia, Sulawesi)

consectator (Smith, F., 1858) (Australia)

• Camponotus discors angustinodus Emery, 1925

consobrinus (Erichson, 1842) (Australia)

• Camponotus dimidiatus Roger, 1863

• Camponotus nigriceps obniger Forel, 1902

conspicuus (Smith, F., 1858) (Argentina, Bahamas, Brazil, Costa Rica, Cuba, Greater Antilles, Guatemala, Jamaica, Lesser Antilles, Mexico, Panama, Uruguay)

conspicuus sharpi Forel, 1893 (Grenada, Mexico)

conspicuus williamsi Forel, 1916 (Guadeloupe, Lesser Antilles)

constructor Forel, 1899 (Costa Rica, Ecuador, Panama)

contractus Mayr, 1872 (Borneo, Indonesia, Malaysia)

contractus buttesi Forel, 1902 (Malaysia)

contractus scortechinii Emery, 1888 (Malaysia)

conulus Mayr, 1870 (Colombia)

convexiclypeus Mackay, W.P., 1997 (Peru)

coptobregma Kempf, 1968 (Brazil)

cordiceps Santschi, 1939 (Argentina)

cordincola Wheeler, W.M., 1934 (Panama)

coriolanus Forel, 1912 (Brazil)

• Camponotus scrobifer Borgmeier, 1928

corniculatus Wheeler, W.M., 1934 (Guyana)

cornis Wang, C. & Wu, J., 1994 (China)

coruscus (Smith, F., 1862) (Colombia, Costa Rica, Guatemala, Honduras, Mexico, Panama, Venezuela)

coruscus astilbus Stitz, 1938 (Suriname)

coruscus fulgens Forel, 1885 (Guatemala)

cosmicus (Smith, F., 1858) (Mozambique, South Africa, Zimbabwe)

coxalis (Smith, F., 1859) (Indonesia, New Guinea)

crassicornis Emery, 1920 (Bolivia, Brazil)

crassisquamis Forel, 1902 (Bangladesh, India)

crassus Mayr, 1862 (Argentina, Brazil, Paraguay, Trinidad and Tobago)

• Camponotus crassus picticornis Santschi, 1936

• Camponotus flexus Mayr, 1862

• Camponotus vezenyii Forel, 1907

crassus amazonicus Santschi, 1922 (Brazil)

crassus delabiatus Santschi, 1925 (Brazil)

crawleyi Emery, 1920 (Cameroun)

crenatus Mayr, 1876 (Australia)

crepusculi Arnold, 1922 (Zimbabwe)

cressoni André, 1887 (Bolivia, Brazil, Mexico)

cressoni purensis Forel, 1912 (Brazil)

criniticeps Menozzi, 1939 (Ethiopia)

crispulus Santschi, 1922 (Argentina)

crozieri McArthur & Leys, 2006 (Australia)

crucheti Santschi, 1911 (Angola)

cruentatus (Latreille, 1802) (Algeria, Andorra, France, Gibraltar, Iberian Peninsula, Italy, Monaco, Morocco, Portugal, Spain, Tunisia)

• Camponotus cruentatus cruentior Santschi, 1931

• Camponotus cruentatus lindbergi Santschi, 1931

• Formica gigas Leach, 1825

• Formica opaca Nylander, 1856

cruentatus asper Menozzi, 1925 (Australia)

cuauhtemoc Snelling, R.R., 1988 (Mexico, Mexico, United States)

cubangensis Forel, 1901 (Angola)

cubangensis dofleini Forel, 1911 (South Africa)

• Camponotus mayri ledieui Forel, 1916

cuneidorsus Emery, 1920 (Costa Rica, Guatemala, Honduras, Mexico, Peru)

cuneiscapus Forel, 1910 (Namibia, South Africa)

curviscapus Emery, 1896 (Costa Rica, Mexico)

cyrtomyrmodes Donisthorpe, 1941 (New Guinea)

D
daitoensis Terayama, 1999 (Japan)

dalmasi Forel, 1899 (Colombia)

dalmaticus (Nylander, 1849) (Bulgaria, Croatia, Greece, Israel, Italy, Lebanon, Montenegro, North Macedonia, Slovenia, Switzerland, Syria, Turkey)

dalmaticus rhodius Santschi, 1934 (Greece)

daraina Rakotonirina & Fisher, 2022 (Madagascar)

darlingtoni Wheeler, W.M., 1934 (Australia)

• Camponotus christmasensis McArthur, 2008

• Camponotus rottnesti Donisthorpe, 1941

darwinii Forel, 1886 (Madagascar, Seychelles)

• Camponotus darwinii rubropilosus Forel, 1891

debellator Santschi, 1926 (Zimbabwe)

decessor Forel, 1908 (Brazil)

decipiens Emery, 1893 (Mexico, United States)

• Camponotus fallax rasilis Wheeler, W.M., 1910

declivus Santschi, 1922 (Mexico)

dedalus Forel, 1911 (Indonesia)

dedalus moeschiellus Forel, 1912 (Indonesia)

deletangi Santschi, 1920 (Bolivia)

densopilus Shattuck, 2005 (Indonesia, New Guinea)

depressiceps Forel, 1879 (Brazil)

depressus Mayr, 1866 (Argentina, Brazil, Paraguay)

desantii Santschi, 1915 (Benin)

descarpentriesi Santschi, 1926 (Madagascar)

detritus Emery, 1886 (Namibia, South Africa)

devestivus Wheeler, W.M., 1928 (China, Japan)

dewitzii Forel, 1886 (Democratic Republic of Congo)

dicksoni Arnold, 1948 (South Africa)

difformis Stitz, 1938 (New Guinea)

dimorphus Emery, 1894 (Argentina, Bolivia, Brazil, Paraguay)

diplopunctatus Emery, 1915 (Eritrea, Ethiopia, Oman)

diplopunctatus subconvexus Viehmeyer, 1923 (Sudan)

discolor (Buckley, 1866) (United States)

discors Forel, 1902 (Australia)

discors laetus Emery, 1925 (Australia)

discors yarrabahensis Forel, 1915 (Australia)

distinguendus (Spinola, 1851) (Argentina, Chile, Peru)

• Camponotus distinguendus denudata Emery, 1906

• Camponotus distinguendus santacruziana Emery, 1906

• Camponotus distinguendus tenuipubens Santschi, 1916

divergens Mayr, 1887 (Brazil)

diversipalpus Santschi, 1922 (Brazil)

dolabratus Menozzi, 1927 (Costa Rica)

dolendus Forel, 1892 (China, India, Laos, Nepal)

dolichoderoides Forel, 1911 (Borneo, Indonesia, Malaysia, Thailand)

donisthorpei Emery, 1920 (Cameroun)

donnellani Shattuck & McArthur, 2002 (Australia)

dorycus (Smith, F., 1860) (China, Indonesia, New Guinea)

dorycus recticeps Stitz, 1938 (New Guinea)

dromas Santschi, 1919 (Australia)

dromedarius Forel, 1891 (Madagascar)

druryi Forel, 1886 (United Republic of Tanzania)

dryandrae McArthur, 1996 (Australia)

dufouri Forel, 1891 (Madagascar)

• Camponotus dufouri imerinensis Forel, 1891

dumetorum Wheeler, W.M., 1910 (Mexico, United States)

E
eastwoodi McArthur, 1996 (Australia)

ebneri Finzi, 1930 (Lebanon)

echinoploides Forel, 1891 (Madagascar)

edmondi André, 1887 (Comoros, Madagascar, Mayotte)

• Camponotus edmondi ernesti Forel, 1891

efitra Rakotonirina, Csősz & Fisher, 2017 (Madagascar)

egregius (Smith, F., 1858) (Brazil)

elegans Forel, 1902 (Australia)

elevatus Forel, 1899 (Costa Rica, Mexico)

ellioti Forel, 1891 (Madagascar)

ellioti relucens Santschi, 1911 (Madagascar)

emarginatus Emery, 1886 (South Africa)

emeryodicatus Forel, 1901 (Argentina, Brazil, Ecuador, Paraguay)

• Camponotus emeryi Forel, 1901 Forel, 1901

empedocles Emery, 1920 (Saudi Arabia, United Arab Emirates, Zimbabwe)

enigmaticus Mackay, W.P., Mackay, M. & Mackay, E.E., 2004 (Nicaragua)

eperiamorum Clouse, 2007 (Micronesia (Federated States of))

ephippiatus Viehmeyer, 1916 (Singapore)

ephippium (Smith, F., 1858) (Australia)

ephippium narses Forel, 1915 (Australia)

eremicus Wheeler, W.M., 1915 (Australia)

erigens Forel, 1894 (Algeria)

erigens subconcolor Emery, 1920 (Algeria)

erinaceus Gerstäcker, 1871 (Kenya, United Republic of Tanzania)

errabundus Arnold, 1949 (Zimbabwe)

erythrocephalus Clouse, 2007 (Micronesia (Federated States of))

erythrostoma Emery, 1920 (Brazil)

esau Forel, 1915 (Australia)

essigi Smith, M.R., 1923 (Mexico, United States)

• Camponotus nevadensis Gregg, 1973

• Camponotus pertusus Mackay, W.P., 1997

ethicus Forel, 1897 (Madagascar)

etiolipes Bolton, 1995 (Kenya, Mozambique, Zimbabwe)

• Formica longipes Gerstäcker Gerstäcker, 1859

eugeniae Forel, 1879 (Lesotho, Mozambique, South Africa, Zimbabwe)

eugeniae amplior Forel, 1913 (Democratic Republic of Congo)

eurynotus Forel, 1907 (Bolivia, Brazil, Colombia, Ecuador)

evae Forel, 1910 (Australia)

evae zeuxis Forel, 1915 (Australia)

evansi Crawley, 1920 (Iran, Iraq)

excisus Mayr, 1870 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Panama, Trinidad and Tobago)

exiguoguttatus Forel, 1886 (China, India)

exsectus Emery, 1900 (Borneo, Indonesia, Malaysia)

extensus Mayr, 1876 (Australia)

ezotus Bolton, 1995 (Australia)

• Camponotus erythropus Viehmeyer Viehmeyer, 1925

F
fabricator (Smith, F., 1858) (Saint Helena)

falco Forel, 1902 (Colombia)

fallatus Bolton, 1995 (Brazil)

• Camponotus escherichi Menozzi, 1926

fallax (Nylander, 1856) (Albania, Andorra, Armenia, Austria, Azerbaijan, Balearic Islands, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Liechtenstein, Lithuania, Luxembourg, Malta, Montenegro, Netherlands, Poland, Portugal, Republic of Korea, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)

• Camponotus fallax pageti Hamann & Klemm, 1976

• Camponotus marginatus kamensis Ruzsky, 1903

• Camponotus marginatus ruzskyi Emery, 1898

fasciatellus Dalla Torre, 1892 (Bolivia, Colombia, French Guiana, Guyana, Mexico, Suriname)

• Camponotus fasciatus Mayr Mayr, 1870

fasciatus (Mayr, 1867) (Borneo, Indonesia, Malaysia)

fastigatus Roger, 1863 (Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Suriname, Trinidad and Tobago)

fastigatus barbiculus Santschi, 1922 (Bolivia)

fastigatus schmalzi Emery, 1894 (Brazil)

favorabilis Santschi, 1919 (Zimbabwe)

fayfaensis Collingwood, 1985 (Saudi Arabia, United Arab Emirates)

feae Emery, 1882 (Canary Islands)

• Camponotus feae erythropus Stitz, 1917

• Camponotus feae excelsus Stitz, 1917

• Formica pubescens Brullé Brullé, 1840

fedtschenkoi Mayr, 1877 (Afghanistan, Armenia, Azerbaijan, China, Iran, Kazakhstan, Kyrgyzstan, Russian Federation, Turkey, Turkmenistan)

• Camponotus fedtschenkoi adi Stitz, 1930

• Camponotus tichomirovi Ruzsky, 1905

fellah Dalla Torre, 1893 (Egypt, Eritrea, Iran, Iraq, Israel, Jordan, Kuwait, Oman, Saudi Arabia, Syria, Turkey, United Arab Emirates)

femoratus (Fabricius, 1804) (Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname, Trinidad and Tobago, Venezuela)

fergusoni McArthur, 2003 (Australia)

ferreri Forel, 1913 (Cameroun)

ferreri akka Forel, 1916 (Democratic Republic of Congo)

ferreri cavisquamis Santschi, 1926 (Kenya)

fervidus Donisthorpe, 1943 (Borneo, Indonesia, Malaysia)

• Formica fervens Smith, F. Smith, F., 1857

festai Emery, 1894 (Greece, Israel, Lebanon, Syria, Turkey)

• Camponotus maculatus caeciliae Forel, 1906

festinatus (Buckley, 1866) (Mexico, United States)

• Camponotus fumidus pubicornis Emery, 1893

• Camponotus fumidus spurcus Wheeler, W.M., 1910

festinus (Smith, F., 1857) (Borneo, India, Indonesia, Malaysia)

festinus cetegus Forel, 1911 (Borneo, Indonesia, Malaysia)

festinus diligens (Smith, F., 1857) (Malaysia)

festinus eximius Emery, 1900 (Borneo, Indonesia, Malaysia, Singapore)

festinus inezae Forel, 1913 (Malaysia)

festinus simaluranus Forel, 1915 (Indonesia, Singapore)

fiebrigi Forel, 1906 (Argentina, Paraguay)

fieldeae Forel, 1902 (Australia)

• Camponotus nigroaeneus xuthus Emery, 1925

• Camponotus spenceri Clark, 1930

fieldellus Forel, 1910 (Australia)

flavescens (Fabricius, 1793) (French Guiana)

flavicomans Clouse, 2007 (Palau)

flavocassis Donisthorpe, 1941 (New Guinea)

flavocrines Donisthorpe, 1941 (New Guinea)

flavomarginatus Mayr, 1862 (Benin, Eritrea, Ghana, Guinea, Kenya, Oman, Rwanda, Saudi Arabia, Somalia, South Africa, United Arab Emirates)

• Camponotus jizani Collingwood, 1985

• Camponotus micans albisectus Emery, 1892

fletcheri Donisthorpe, 1942 (Sri Lanka)

floridanus (Buckley, 1866) (United States)

• Camponotus atriceps yankee Forel, 1885

florius Santschi, 1926 (Congo)

foersteri Forel, 1886 (Madagascar)

• Camponotus cambouei Forel, 1891

• Camponotus christi ambustus Forel, 1892

• Camponotus christi ferrugineus Emery, 1899

• Camponotus pictipes Forel, 1891

foleyi Santschi, 1939 (Algeria)

foleyi ajjer Bernard, 1953 (Algeria)

foleyi fezzanensis Bernard, 1953 (Algeria)

foleyi grasi Bernard, 1953 (Algeria)

foleyi lelubrei Bernard, 1953 (Algeria)

foleyi pseudocompressus Özdikmen, 2010 (Algeria)

• Camponotus foleyi hoggarensis Bernard, 1953

foleyi rufescens Bernard, 1953 (Algeria)

folicola Forel, 1904 (Brazil)

foraminosus Forel, 1879 (Benin, Cape Verde, Guinea, Kenya, Mozambique, Oman, Saudi Arabia, Senegal)

foraminosus chrysogaster Emery, 1895 (Ethiopia, Kenya)

foraminosus cuitensis Forel, 1901 (Angola)

foraminosus deductus Santschi, 1915 (Nigeria)

foraminosus dorsalis Santschi, 1926 (United Republic of Tanzania)

foraminosus flavus Stitz, 1916 (Congo)

foreli Emery, 1881 (Algeria, Greece, Iberian Peninsula, Portugal, Spain)

foreli tingitanus Santschi, 1921 (Morocco)

formiciformis Forel, 1885 (Belize, Ecuador, Guatemala, Mexico)

formosensis Wheeler, W.M., 1927 (Taiwan)

fornasinii Emery, 1895 (Mozambique)

fragilis Pergande, 1893 (Mexico, United States)

fraseri McArthur, 2008 (Australia)

friedae Forel, 1912 (China, Japan, Taiwan)

• Camponotus friedae amia Forel, 1912

froggatti Forel, 1902 (Australia)

frontalis Pergande, 1896 (Mexico)

fryi Mann, 1916 (Brazil)

fugax Forel, 1902 (Jamaica)

fulvopilosus (De Geer, 1778) (Botswana, India, Lesotho, Namibia, South Africa)

• Camponotus fulvopilosus detritoides Forel, 1910

• Camponotus fulvopilosus flavopilosus Emery, 1895

• Formica pilosa Olivier, 1792

• Formica rufiventris Fabricius Fabricius, 1804

fumidus Roger, 1863 (Argentina, Bahamas, Brazil, Lesser Antilles, Mexico, Venezuela)

fumidus dominicensis Wheeler, W.M., 1913 (Dominica)

fumidus fraterculus Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

fumidus haytianus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus illitus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus imbecillus Wheeler, W.M. & Mann, 1914 (Haiti)

fumidus soulouquei Forel, 1901 (Dominican Republic, Haiti)

fumidus toltecus Forel, 1899 (Mexico)

furvus Santschi, 1911 (Congo)

fuscivillosus Xiao & Wang, C., 1989 (China)

fuscocinctus Emery, 1888 (Brazil)

fuscus Kim & Kim, 1994 (Republic of Korea)

G
gabonensis Santschi, 1926 (Gabon)

galla Forel, 1894 (Eritrea, Ethiopia, Kenya)

• Camponotus foraminosus latinotus Forel, 1907

gallagheri Collingwood & Agosti, 1996 (Oman)

galoko Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

gambeyi Emery, 1883 (New Caledonia)

gambeyi marthae Forel, 1894 (New Caledonia)

geayi Santschi, 1922 (French Guiana)

genatus Santschi, 1922 (Brazil)

gentingensis Dumpert, 1995 (Malaysia)

germaini Emery, 1903 (Argentina, Brazil, Paraguay)

• Camponotus germaini tacuruensis Emery, 1911

gestroi Emery, 1878 (Algeria, Armenia, Bulgaria, Cyprus, Gibraltar, Greece, Iberian Peninsula, Iran, Iraq, Israel, Italy, Jordan, Lebanon, Portugal, Spain, Turkey)

• Camponotus gestroi creticus Forel, 1886

gestroi ponariensis Santschi, 1921 (Algeria)

gibber Forel, 1891 (Madagascar)

gibbinotus Forel, 1902 (Australia)

gibbosus Karavaiev, 1929 (Indonesia)

gilviceps Roger, 1863 (Borneo, Indonesia, Malaysia)

• Camponotus smithi Donisthorpe, 1932

• Formica distinguenda Smith, F., 1871

• Formica ruficeps Smith, F. Smith, F., 1857

gilviventris Roger, 1863 (Cuba, Greater Antilles)

gilviventris refectus Wheeler, W.M., 1937 (Cuba)

gilviventris renormatus Wheeler, W.M., 1937 (Cuba)

glabrisquamis Emery, 1911 (New Guinea)

godmani Forel, 1899 (Brazil, Mexico, Panama, Trinidad and Tobago)

godmani palliolatus Wheeler, W.M., 1922 (Trinidad and Tobago)

goeldii Forel, 1894 (Brazil)

gombaki Dumpert, 1986 (Malaysia)

gouldi Forel, 1886 (Madagascar)

gouldianus Forel, 1922 (Australia)

grandidieri Forel, 1886 (Comoros, Kenya, Madagascar, Mauritius, Mayotte, Mozambique, Namibia, Seychelles, Zimbabwe)

• Camponotus grandidieri atrabilis Santschi, 1915

• Camponotus grandidieri comorensis Santschi, 1915

grandidieri eumendax Özdikmen, 2010 (South Africa)

• Camponotus foraminosus mendax Emery, 1895

grandidieri ruspolii Forel, 1892 (Somalia)

greeni Forel, 1911 (Sri Lanka)

gretae Forel, 1902 (India, Myanmar, Singapore)

guanchus Santschi, 1908 (Canary Islands)

guatemalensis Forel, 1885 (Guatemala)

guayapa Kusnezov, 1952 (Argentina)

guidae McArthur, 2007 (Australia)

guizhouensis Wang, M., 1992 (China)

gundlachi Mann, 1920 (Cuba, Greater Antilles)

guttatus Emery, 1899 (Benin, Cameroun, Guinea, Niger)

guttatus minusculus Viehmeyer, 1914 (United Republic of Tanzania)

H
habereri Forel, 1911 (Japan, Taiwan)

haematocephalus Emery, 1903 (Peru)

haereticus Santschi, 1914 (Benin, Nigeria)

hagensii Forel, 1886 (Madagascar)

hannani Forel, 1899 (Greater Antilles, Jamaica)

• Camponotus hannani willardi Forel, 1899

hapi Weber, 1943 (Sudan)

harenarum Rakotonirina & Fisher, 2022 (Madagascar)

haroi Espadaler, 1997 (Iberian Peninsula, Spain)

hartogi Forel, 1902 (Australia)

• Camponotus ferruginipes Crawley, 1922

hastifer Emery, 1911 (New Guinea)

havilandi Arnold, 1922 (South Africa)

heathi Mann, 1916 (Brazil, Ecuador)

heathi gilvigaster Wheeler, W.M., 1923 (Brazil)

hedwigae Forel, 1912 (Brazil)

heidrunvogtae Seifert, 2019 (Bosnia and Herzegovina, Croatia, Greece, Montenegro)

helleri Emery, 1903 (Argentina, Ecuador, Paraguay, Peru)

hellmichi Menozzi, 1935 (Chile)

helvus Xiao & Wang, C., 1989 (China)

hemichlaena Yasumatsu & Brown, 1951 (Japan)

herculeanus (Linnaeus, 1758) (Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Canada, China, Croatia, Czech Republic, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kazakhstan, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Netherlands, Norway, Poland, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United States)

• Camponotus herculeanus altaica Ruzsky, 1915

• Camponotus herculeanus caucasicus Arnoldi, 1967

• Camponotus herculeanus eudokiae Ruzsky, 1926

• Camponotus herculeanus jacuticus Karavaiev, 1929

• Camponotus herculeanus montanus Ruzsky, 1904

• Camponotus herculeanus nadigi Menozzi, 1922

• Camponotus herculeanus sachalinensis Forel, 1904

• Camponotus herculeanus shitkowi Ruzsky, 1904

• Camponotus herculeanus whymperi Forel, 1902

• Formica atra Zetterstedt, 1838

• Formica castanea Lepeletier de Saint-Fargeau, 1835

• Formica intermedia Zetterstedt, 1838

hermanni Emery, 1911 (Brazil)

heros Santschi, 1926 (United Republic of Tanzania)

hesperius Emery, 1893 (Canary Islands)

heteroclitus Forel, 1895 (Madagascar)

hildebrandti Forel, 1886 (Madagascar)

hildebrandti dichromothrix Emery, 1920 (Madagascar)

himalayanus Forel, 1893 (India, Nepal)

hippocrepis Emery, 1920 (Bolivia, Ecuador)

hirtus Karaman, C. & Aktac, 2013 (Turkey)

hispidus Emery, 1906 (Argentina, Paraguay)

hoelldobleri Cagniant, 1991 (Morocco)

holosericeus Emery, 1889 (China, India, Myanmar)

holzi Forel, 1921 (Ecuador)

honaziensis Karaman, C. & Aktac, 2013 (Croatia, Cyprus, Greece, North Macedonia, Spain, Turkey)

hoplites Emery, 1914 (New Caledonia)

horni Clark, 1930 (Australia)

horrens Forel, 1910 (Philippines)

horseshoetus Datta & Raychaudhuri, 1985 (India)

hospes (Emery, 1884) (Borneo, Brunei Darussalam, Indonesia, Malaysia)

hospes adultus Viehmeyer, 1916 (Malaysia)

hova Forel, 1891 (Comoros, Madagascar, Mayotte, Mozambique, Oman, Seychelles, Socotra Archipelago, Yemen)

• Camponotus hova obscuratus Emery, 1925

hova fulvus Emery, 1894 (Comoros, Seychelles)

hova pictiventris Mayr, 1901 (South Africa)

hovahovoides Forel, 1892 (Madagascar)

• Camponotus radamae hovoides Dalla Torre, 1893

humeralis Emery, 1920 (Peru)

humerus Wang, C. & Wu, J., 1994 (China)

humilior Forel, 1902 (Australia)

husseini Dietrich, 2004 (Jordan)

hyatti Emery, 1893 (Mexico, United States)

• Camponotus sayi bicolor Pergande, 1894

• Camponotus sayi californica Emery, 1925

hypoclineoides Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

I
icarus Forel, 1912 (Indonesia)

ignestii Menozzi, 1935 (Guyana)

ihazofotsy Rasoamanana & Fisher, 2022 (Madagascar)

iheringi Forel, 1908 (Argentina, Brazil, Paraguay, Peru)

• Camponotus iheringi bajulus Emery, 1925

• Camponotus iheringi latinota Forel, 1909

ilgii Forel, 1894 (Ethiopia, Oman, Senegal)

imitator Forel, 1891 (Madagascar)

• Camponotus imitator resinicola Santschi, 1911

immaculatus Forel, 1892 (Madagascar)

• Camponotus quadrimaculatus opaca Emery, 1899

• Camponotus quadrimaculatus opacata Emery, 1925

immigrans Santschi, 1914 (Cameroun)

importunus Forel, 1911 (Mozambique, South Africa)

impressilabris Stitz, 1938 (Malaysia)

improprius (Forel, 1879) (Colombia)

inaequalis Roger, 1863 (Cuba, Mexico, United States)

• Camponotus maculatus tortuganus Emery, 1895

inca Emery, 1903 (Brazil, Peru)

inca rector Santschi, 1923 (Brazil)

incensus Wheeler, W.M., 1932 (United States)

inconspicuus Mayr, 1872 (Borneo, Indonesia, Malaysia)

indefinitus Karavaiev, 1929 (Indonesia)

indeflexus (Walker, 1859) (India, Sri Lanka)

indianus Forel, 1879 (Colombia, Panama, Peru)

indicatus Santschi, 1922 (French Guiana)

inflatus Lubbock, 1880 (Australia)

• Camponotus aurofasciatus Wheeler, W.M. Wheeler, W.M., 1915

innexus Forel, 1902 (Australia)

innocens Forel, 1909 (Argentina, Paraguay)

inquilinus Zettel & Laciny, 2018 (Borneo, Brunei Darussalam)

insipidus Forel, 1893 (Australia)

• Camponotus claripes minima Crawley, 1922

integellus Forel, 1899 (Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Peru, Venezuela)

• Camponotus rudigenis Emery, 1903

interjectus Mayr, 1877 (Afghanistan, Azerbaijan, China, Iran, Iraq, Israel, Jordan, Kazakhstan, Kyrgyzstan, Russian Federation, Turkmenistan)

intrepidus (Kirby, W., 1819) (Australia)

• Camponotus magnus Mayr, 1862

• Formica agilis Smith, F., 1858

intrepidus bellicosus Forel, 1902 (Australia)

inverallensis Forel, 1910 (Australia)

• Camponotus churchetti McArthur, 2008

invidus Forel, 1892 (India, Pakistan)

ionius Emery, 1920 (Bulgaria, Greece, North Macedonia, Serbia, Turkey)

iridis Santschi, 1922 (Brazil)

irritabilis (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

irritabilis winkleri Forel, 1911 (Borneo, Indonesia, Malaysia)

irritans (Smith, F., 1857) (Borneo, Brunei Darussalam, China, India, Malaysia, Nicobar Island, Singapore, Sri Lanka, Vietnam)

• Camponotus agnatus Roger, 1863

irritans carensis Emery, 1920 (India, Nicobar Island)

irritans cliens Forel, 1911 (Indonesia)

irritans croceomaculatus Emery, 1914 (New Caledonia)

irritans curtus Emery, 1900 (Indonesia)

irritans fatuus Forel, 1886 (Indonesia, New Guinea)

irritans hongkongensis Forel, 1912 (China)

irritans inferior Emery, 1925 (Borneo, Indonesia, Malaysia)

irritans melanogaster Stitz, 1938 (Micronesia (Federated States of))

irritans pallidus (Smith, F., 1857) (Bangladesh, Borneo, India, Indonesia, Malaysia, Philippines)

irritans procax Santschi, 1919 (Vietnam)

irritans sumatranus Özdikmen, 2010 (Indonesia)

• Camponotus irritans puberula Emery, 1920

irritans volkensi Wheeler, W.M., 1935 (Micronesia (Federated States of))

isabellae Forel, 1909 (Sri Lanka)

itoi Forel, 1912 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea)

itoi kwansienensis Viehmeyer, 1922 (China)

ivadia Rasoamanana & Fisher, 2022 (Madagascar)

iwoensis Terayama & Kubota, 2011 (Japan)

J
jaliensis Dalla Torre, 1893 (Cyprus, Greece, Israel, Turkey)

janeti Forel, 1895 (Australia)

janforrestae McArthur & Shattuck, 2001 (Australia)

japonicus Mayr, 1866 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Pakistan, Philippines, Republic of Korea, Russian Federation)

• Camponotus herculeanus manczshuricus Emery, 1925

• Camponotus japonicus miltotus Wheeler, W.M., 1929

• Camponotus japonicus sanguinea Karavaiev, 1929

• Camponotus japonicus wui Wheeler, W.M., 1929

javaensis Ward, Blaimer & Fisher, 2016 (Borneo, Cambodia, Indonesia, Malaysia, Philippines)

• Forelophilus overbecki Kutter, 1931

jeanneli Santschi, 1914 (Kenya)

jejuensis Kim & Kim, 1986 (Republic of Korea)

jianghuaensis Xiao & Wang, C., 1989 (China)

jjacquia Rasoamanana & Fisher, 2022 (Madagascar)

joany Rakotonirina & Fisher, 2022 (Madagascar)

jodina Rasoamanana, Csősz & Fisher, 2017 (Madagascar)

johnclarki Taylor, 1992 (Australia)

• Notostigma sanguinea Clark, 1930

johnsoni Mackay, 2019 (Mexico)

judithmorrisae McArthur, 2008 (Australia)

juliae Emery, 1903 (Costa Rica)

K
kaguya Terayama, 1999 (Japan)

karaha Rasoamanana, Csősz & Fisher, 2017 (Madagascar)

karsti Rakotonirina & Fisher, 2022 (Madagascar)

kattensis Bingham, 1903 (India)

kaura Snelling, R.R. & Torres, 1998 (Greater Antilles, Netherlands Antilles, Puerto Rico)

kefir Ionescu-Hirsch, 2010 (Israel)

keiferi Wheeler, W.M., 1934 (Mexico)

keihitoi Forel, 1913 (China, Japan)

• Camponotus caryae teranishii Wheeler, W.M., 1928

• Camponotus tokyoensis Teranishi, 1915

kelimaso Rakotonirina & Fisher, 2022 (Madagascar)

keralensis Karmaly & Narendran, 2006 (India)

kersteni Gerstäcker, 1871 (Kenya, Saudi Arabia)

khaosokensis Dumpert, 2006 (Thailand)

• Camponotus hoelldobleri Dumpert Dumpert, 2006

kiesenwetteri (Roger, 1859) (Cyprus, Greece, Iran, Turkey)

• Camponotus kiesenwetteri cypria Emery, 1920

kiusiuensis Santschi, 1937 (Japan, Republic of Korea)

klaesii (Forel, 1886) (Indonesia)

klugii Emery, 1895 (South Africa)

knysnae Arnold, 1922 (South Africa)

kollbrunneri Forel, 1910 (Ethiopia)

kolthoffi Stitz, 1934 (China)

kopetdaghensis Dlussky & Zabelin, 1985 (Azerbaijan, Iran, Russian Federation, Turkmenistan)

korthalsiae Emery, 1887 (Indonesia)

koseritzi Emery, 1888 (Bolivia, Brazil, Peru)

kraepelini Forel, 1901 (Bolivia)

kubaryi Mayr, 1876 (Palau)

kugleri Ionescu-Hirsch, 2010 (Israel)

kunigamiensis Terayama, 2013 (Japan)

kurdistanicus Emery, 1898 (Azerbaijan, China, Iran, Iraq, Israel, Russian Federation, Turkey)

kutteri Forel, 1915 (Indonesia)

kutterianus Baroni Urbani, 1972 (Cuba, Greater Antilles)

L
laconicus Emery, 1920 (Greece)

laevigatus (Smith, F., 1858) (Mexico, United States, United States)

• Camponotus quercicola Smith, M.R., 1954

laevissimus Mackay, 2019 (Canada, Mexico, United States)

lamarckii Forel, 1892 (India, Nepal)

lamborni Donisthorpe, 1933 (Malawi)

lameerei Emery, 1898 (Kyrgyzstan, Russian Federation, Turkmenistan)

• Camponotus fallax barbatus Vashkevich, 1926

• Camponotus fallax barbiger Donisthorpe, 1941

lamosy Rakotonirina & Fisher, 2018 (Madagascar)

lancifer Emery, 1894 (Brazil)

landolti Forel, 1879 (Brazil, Colombia, Costa Rica, Mexico, Panama)

langi Wheeler, W.M., 1922 (Democratic Republic of Congo)

langi jejunus Santschi, 1926 (Democratic Republic of Congo)

largiceps Wu, J. & Wang, C., 1989 (China)

lasiselene Wang, C. & Wu, J., 1994 (China, Thailand)

latangulus Roger, 1863 (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Suriname, Trinidad and Tobago)

latebrosus (Walker, 1859) (Sri Lanka)

lateralis (Olivier, 1792) (Albania, Algeria, Andorra, Armenia, Azerbaijan, Balearic Islands, Bulgaria, Croatia, Cyprus, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Malta, Monaco, Montenegro, Morocco, Portugal, North Macedonia, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine)

• Camponotus kosswigi Donisthorpe, 1950

• Camponotus lateralis armouri Wheeler, W.M., 1926

• Camponotus lateralis balearis Santschi, 1929

• Camponotus lateralis purius Santschi, 1929

• Formica axillaris Spinola, 1808

• Formica bicolor Latreille Latreille, 1798

• Formica hemipsila Foerster, 1850

• Formica melanogastes Latreille, 1802

• Formica merula Losana, 1834

• Formica pallidinervis Brullé, 1833

laurenti Santschi, 1939 (Algeria)

lautus (Say, 1836)

leae Wheeler, W.M., 1915 (Australia)

lenkoi Kempf, 1960 (Brazil)

leptocephalus Emery, 1923 (Brazil)

lespesii Forel, 1886 (Argentina, Brazil, Colombia, Mexico, Panama, Paraguay)

• Camponotus lespesi melancholicus Emery, 1894

leucodiscus Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia, Philippines)

leucophaeus (Smith, F., 1861) (Indonesia, Sulawesi)

leveillei Emery, 1895 (Madagascar)

leydigi Forel, 1886 (Argentina, Brazil, French Guiana, Guyana, Paraguay, Suriname)

• Camponotus nobilis Santschi, 1916

liandia Rakotonirina & Fisher, 2018 (Madagascar)

libanicus André, 1881 (Cyprus, Greece, Iran, Israel, Lebanon, Turkey)

• Camponotus libanicus sahlbergi Forel, 1913

• Camponotus nadimi Tohmé, G., 1969

ligea Donisthorpe, 1931 (Ethiopia)

lighti Wheeler, W.M., 1927 (China)

ligniperda (Latreille, 1802) (Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)

• Camponotus herculeanus herculeanoligniperdus Dalla Torre, 1893

• Formica obsoleta Christ Christ, 1791

ligniperda afer Stärcke, 1942 (Czech Republic)

ligniperda nigrescens Gösswald, 1932 (Germany)

lilianae Forel, 1913 (Democratic Republic of Congo, Kenya)

lilianae cornutus Forel, 1913 (Democratic Republic of Congo)

limbiventris Santschi, 1911 (United Republic of Tanzania)

lindigi Mayr, 1870 (Belize, Colombia, Costa Rica, Greater Antilles, Honduras, Nicaragua, Panama, Trinidad and Tobago, Venezuela)

linnaei Forel, 1886 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Peru)

• Camponotus angulatus Mayr, 1870

• Camponotus angulicollis Emery, 1887

linnaei comoedus Wheeler, W.M., 1934 (Mexico)

linnaei maccus Wheeler, W.M., 1934 (French Guiana, Guyana)

linnaei muticus Forel, 1899 (Costa Rica)

liogaster Santschi, 1932 (South Africa)

lividicoxis Viehmeyer, 1925 (Australia)

lividipes Emery, 1887 (Australia)

lokobe Rakotonirina & Fisher, 2022 (Madagascar)

longiceps (Smith, F., 1863) (China, Indonesia)

longicollis Rasoamanana, Csősz & Fisher, 2017 (Madagascar)

longideclivis McArthur, 1996 (Australia)

longifacies McArthur, 2003 (Australia)

longipalpis Santschi, 1926 (Democratic Republic of Congo)

longipilis Emery, 1911 (Bolivia, Peru)

loweryi McArthur, 1996 (Australia)

lownei Forel, 1895 (Australia)

• Formica nitida Lowne Lowne, 1865

lubbocki Forel, 1886 (Madagascar)

• Camponotus lubbocki christoides Forel, 1891

• Camponotus lubbocki rectus Forel, 1891

lucayanus Wheeler, W.M., 1905 (Bahamas, Dominican Republic, Haiti)

luctuosus (Smith, F., 1858) (Indonesia)

luteiventris Emery, 1897 (New Guinea)

luteus (Smith, F., 1858) (India)

lutzi Forel, 1905 (Brazil)

M
macareaveyi Taylor, 1992 (Australia)

• Camponotus sanguinea McAreavey, 1949

maccooki Forel, 1879 (Mexico, United States)

macilentus Smith, F., 1877 (Ecuador, Galapagos Islands)

macilentus albemarlensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

macilentus altinotus Stitz, 1932 (Galapagos Islands, Mexico)

macilentus barringtonensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus bindloensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus castellanus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus duncanensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus hoodensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus jacobensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus narboroensis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus pervicus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus sapphirinus Wheeler, W.M., 1924 (Galapagos Islands)

macilentus vulcanalis Wheeler, W.M., 1919 (Galapagos Islands)

macilentus wollebaeki Stitz, 1932 (Galapagos Islands)

mackayensis Forel, 1902 (Australia)

macrocephalus Emery, 1894 (Argentina, Bolivia, Brazil, Paraguay, Peru)

• Camponotus geralensis Emery, 1920

• Camponotus luederwaldti Santschi, 1922

macrochaeta Emery, 1903 (Brazil, Ecuador)

macromischoides Fontenla Rizo, 1997 (Cuba)

maculatus (Fabricius, 1782) (Algeria, Angola, Benin, Cameroun, Central African Republic, Chad, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Iran, Ivory Coast, Kenya, Lesotho, Liberia, Madagascar, Mali, Mauritius, Mayotte, Micronesia (Federated States of), Mozambique, Namibia, Oman, Palau, Réunion, Saudi Arabia, Senegal, Sierra Leone, South Africa, Sudan, São Tomé & Principe, Turkey, Uganda, United Republic of Tanzania, Zimbabwe)

• Camponotus maculatus atramentarius Forel, 1904

• Camponotus maculatus ballioni Forel, 1904

• Camponotus maculatus boera Santschi, 1925

• Camponotus maculatus cavallus Santschi, 1911

• Camponotus maculatus cluisoides Forel, 1913

• Camponotus maculatus conakryensis Emery, 1920

• Camponotus maculatus erythraea Emery, 1920

• Camponotus maculatus flavifemur Santschi, 1937

• Camponotus maculatus flavominor Emery, 1925

• Camponotus maculatus hannae Santschi, 1919

• Camponotus maculatus hieroglyphicus Santschi, 1917

• Camponotus maculatus intonsus Emery, 1905

• Camponotus maculatus liocnemis Emery, 1905

• Camponotus maculatus lividior Santschi, 1911

• Camponotus maculatus lohieri Emery, 1915

• Camponotus maculatus manzer Forel, 1910

• Camponotus maculatus mathildae Forel, 1910

• Camponotus maculatus melanocnemis Santschi, 1911

• Camponotus maculatus miserabilis Santschi, 1914

• Camponotus maculatus nubis Weber, 1943

• Camponotus maculatus proletaria Baroni Urbani, 1971

• Camponotus maculatus radamoide Forel, 1891

• Camponotus maculatus sarmentus Emery, 1920

• Camponotus maculatus schereri Forel, 1911

• Camponotus maculatus schultzei Forel, 1912

• Camponotus maculatus semispicatus Emery, 1920

• Camponotus maculatus sudanicus Weber, 1943

• Camponotus maculatus thomensis Santschi, 1920

• Camponotus maculatus tuckeri Santschi, 1932

• Camponotus maculatus zumpti Santschi, 1937

• Camponotus sexpunctatus liengmei Forel, 1894

• Formica cognata Smith, F., 1858

• Formica lacteipennis Smith, F., 1858

maculatus foveolatus Stitz, 1925 (Philippines)

maculatus obfuscatus Viehmeyer, 1916 (Brunei Darussalam, Singapore)

maculatus subnudus Emery, 1889 (Myanmar, Philippines, Singapore)

maculatus sylvaticomaculatus Dalla Torre, 1893 (Turkey)

maculatus ugandensis Santschi, 1923 (Uganda)

maculiventris Emery, 1895 (Madagascar)

madagascarensis Forel, 1886 (Madagascar)

magister Santschi, 1925 (Morocco)

magister tibestiensis Santschi, 1932 (Libya)

maguassa Wheeler, W.M., 1922 (Democratic Republic of Congo)

mahafaly Rakotonirina & Fisher, 2022 (Madagascar)

maintikibo Rakotonirina, Csősz & Fisher, 2017 (Madagascar)

maintilany Rasoamanana & Fisher, 2022 (Madagascar)

mainty Rakotonirina & Fisher, 2018 (Madagascar)

malleensis McArthur, 2007 (Australia)

manabo Rakotonirina & Fisher, 2018 (Madagascar)

marcens Forel, 1907 (Australia)

marianensis Clouse, 2007 (Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands)

maritimus Ward, 2005 (United States)

maschwitzi Dumpert, 2006 (Borneo, Indonesia, Malaysia)

massinissa Wheeler, W.M., 1922 (Democratic Republic of Congo)

matsilo Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

maxwellensis Forel, 1913 (Malaysia, Singapore)

maynei Forel, 1916 (Democratic Republic of Congo)

mayri Forel, 1879 (Kenya, Mozambique, South Africa, Zimbabwe)

mayri chimporensis Santschi, 1930 (Angola)

medeus Emery, 1920 (Peru)

medeus fulvulus Emery, 1920 (Peru)

megalonyx Wheeler, W.M., 1919 (Borneo, India, Indonesia, Malaysia)

• Camponotus chalaconotus Menozzi, 1929

• Camponotus dracocephalus Stitz, 1938

melanocephalus Roger, 1863 (Venezuela)

melanoticus Emery, 1894 (Argentina, Bolivia, Brazil, Ecuador, Mexico, Paraguay)

melanoticus catharinae Santschi, 1939 (Brazil)

melanoticus flavopubens Emery, 1925 (Mexico)

melanoticus kempfi Özdikmen, 2010 (Brazil)

• Camponotus melanoticus nigrescens Santschi, 1939

melanoticus paranaensis Santschi, 1939 (Brazil)

melanoticus publicola Forel, 1912 (Brazil)

melanoticus valerius Santschi, 1922 (Brazil)

melanus Dumpert, 1995 (Borneo, Indonesia, Malaysia)

melichloros Kirby, W.F., 1888 (Christmas Island)

melinus Mackay, W.P., 1997 (Mexico)

mendax Forel, 1895 (India, Sri Lanka)

• Camponotus pseudolus Forel, 1907

mendax integer Forel, 1895 (Sri Lanka)

micans (Nylander, 1856) (Algeria, Iberian Peninsula, Monaco, Portugal, Spain)

micans asniensis Santschi, 1939 (Morocco)

michaelseni Forel, 1907 (Australia)

• Camponotus walkeri bardus Forel, 1910

micragyne Dumpert, 1995 (Malaysia)

micronesicus Blanchard & Clouse, 2016 (Guam, Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands)

microps Snelling, R.R., 2006 (Mexico, United States)

micrositus Wheeler, W.M., 1937 (Cuba, Greater Antilles)

mifaka Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

mina Forel, 1879 (Mexico, United States)

• Camponotus erythropus Pergande Pergande, 1893

• Camponotus mina zuni Wheeler, W.M., 1910

minozzii Emery, 1920 (Peru)

minus Wang, C. & Wu, J., 1994 (China)

mirabilis Emery, 1903 (Bolivia, Brazil, Peru)

misturus (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

• Formica exasperata Smith, F., 1857

misturus fornaronis Forel, 1892 (Bangladesh, Borneo, India, Indonesia, Malaysia)

mita Rakotonirina, Csősz & Fisher, 2017 (Madagascar)

mitis (Smith, F., 1858) (China, India, Singapore, Sri Lanka)

mixtellus Dalla Torre, 1893 (Comoros, Madagascar, Mayotte, Seychelles)

mocquerysi Emery, 1899 (Madagascar)

mocsaryi Forel, 1902 (Brazil, Ecuador, French Guiana, Guyana, Peru, Suriname)

moderatus Santschi, 1930 (Ethiopia)

modoc Wheeler, W.M., 1910 (Canada, Mexico, United States)

moelleri Forel, 1912 (Brazil)

molossus Forel, 1907 (Australia)

mombassae Forel, 1886 (Kenya, United Republic of Tanzania)

monardi Santschi, 1930 (Angola)

monju Terayama, 1999 (Japan)

montivagus Forel, 1885 (Guatemala, Mexico)

• Camponotus nitidus nuperus Wheeler, W.M., 1914

morosus (Smith, F., 1858) (Argentina, Chile)

mozabensis Emery, 1899 (Algeria)

• Camponotus foreli impolitus Forel, 1902

mucronatus Emery, 1890 (Costa Rica, Guatemala, Honduras, Mexico)

• Dolichoderus tridentanodus Ortega-De Santiago & Vásquez-Bolaños, 2012

mucronatus formaster Wheeler, W.M., 1934 (Guatemala, Mexico)

mucronatus hirsutinasus Wheeler, W.M., 1934 (Guatemala)

mus Roger, 1863 (Argentina, Bolivia, Brazil, Colombia, Ecuador, Mexico, Paraguay, Trinidad and Tobago, Uruguay)

• Camponotus ogloblini Santschi, 1934

mus mendozanus Santschi, 1922 (Argentina)

mus mustelus Emery, 1920 (Bolivia)

mussolinii Donisthorpe, 1936 (New Guinea)

mutilarius Emery, 1893 (India, Myanmar)

mystaceus Emery, 1886 (Botswana, Namibia, South Africa, Zimbabwe)

mystaceus exsanguis Forel, 1910 (Namibia)

mystaceus kamae Forel, 1910 (Namibia)

N
nacerdus Norton, 1868 (Mexico)

naegelii Forel, 1879 (Brazil)

namacola Prins, 1973 (Namibia)

nasicus Forel, 1891 (Madagascar)

nasutus Emery, 1895 (Namibia, South Africa, Zimbabwe)

nasutus fenestralis Santschi, 1937 (Namibia)

nasutus pretiosus Arnold, 1922 (Zimbabwe)

nasutus quadridentatus Santschi, 1937 (Namibia)

nasutus quinquedentatus Forel, 1910 (South Africa)

nasutus subnasutus Arnold, 1922 (South Africa)

natalensis (Smith, F., 1858) (Lesotho, South Africa, Zimbabwe)

natalensis corvus Forel, 1879 (South Africa)

natalensis diabolus Forel, 1879 (South Africa)

natalensis fulvipes Forel, 1914 (South Africa)

natalensis politiceps Santschi, 1914 (South Africa)

navigator Wilson & Taylor, 1967 (Samoa)

nawai Ito, 1914 (Japan, Republic of Korea)

nearcticus Emery, 1893 (Canada, United States)

• Camponotus fallax pardus Wheeler, W.M., 1917

• Camponotus fallax tanquaryi Wheeler, W.M., 1917

• Camponotus marginatus minutus Emery, 1893

nepos Forel, 1912 (Brazil)

niavo Rakotonirina & Fisher, 2022 (Madagascar)

nicobarensis Mayr, 1865 (Bangladesh, China, India, Laos, Nicobar Island, Thailand, Vietnam)

nicobarensis monticola Bingham, 1903 (Myanmar, Philippines)

nicobarensis rabbani Emery, 1925 (Myanmar)

nidulans (Smith, F., 1860) (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Suriname, Venezuela)

• Camponotus fabricii Roger, 1862

• Camponotus fabricii acoma Forel, 1908

• Dendromyrmex chartifex felis Wheeler, W.M., 1916

• Dendromyrmex chartifex mamoreensis Mann, 1916

• Dendromyrmex fabricii io Weber, 1943

• Dendromyrmex fabricii niger Weber, 1943

• Dendromyrmex fabricii pictus Wheeler, W.M., 1923

• Dendromyrmex fabricii rufescens Wheeler, W.M., 1916

• Dendromyrmex mamoreensis vestita Santschi, 1921

• Dendromyrmex nidulans nigripes Wheeler, W.M., 1916

nigricans Roger, 1863 (Philippines)

nigricans enganensis Forel, 1916 (Indonesia, Singapore)

• Camponotus nigricans nitidiceps Viehmeyer, 1916

nigricans pantiensis Santschi, 1932 (Indonesia)

nigriceps (Smith, F., 1858) (Australia)

• Camponotus consobrinus perthianus Wheeler, W.M., 1933

nigripes Dumpert, 1995 (Malaysia)

nigroaeneus (Smith, F., 1858) (Australia)

• Camponotus nigroaeneus divus Forel, 1907

nigronitidus Azuma, 1951 (Japan)

nipponensis Santschi, 1937 (Democratic Peoples Republic of Korea, Japan, Republic of Korea)

nirvanae Forel, 1893 (India)

nitens Mayr, 1870 (Brazil, Colombia)

nitens fuhrmanni Forel, 1921 (Colombia)

nitidescens Forel, 1889 (Greece)

nitidior (Santschi, 1921) (Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Dendromyrmex fabricii isthmicus Weber, 1943

niveosetosus Mayr, 1862 (Botswana, Kenya, Namibia, South Africa)

niveosetosus irredux Forel, 1910 (South Africa)

normatus Forel, 1899 (Costa Rica)

norvigi Rasoamanana & Fisher, 2022 (Madagascar)

nossibeensis André, 1887 (Madagascar)

novaeboracensis (Fitch, 1855) (Canada, United States)

• Camponotus herculeanus rubens Emery, 1925

• Camponotus ligniperdus pictus Forel, 1886

novaehollandiae Mayr, 1870 (Australia, New Caledonia, Solomon Islands)

• Camponotus villosus Crawley, 1915

novogranadensis Mayr, 1870 (Argentina, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States)

novogranadensis modestior Forel, 1904 (Costa Rica)

nylanderi Emery, 1921 (United Arab Emirates)

• Formica pallens Nylander, 1849

nywet Bolton, 1995 (Brazil)

• Formica nana Smith, F. Smith, F., 1858

O
oasium Forel, 1890 (Afghanistan, Algeria, Egypt, Iran, Iraq, Kazakhstan, Mali, Oman, Saudi Arabia, Turkey, Turkmenistan, United Arab Emirates)

• Camponotus bactrianus Pisarski, 1967

oasium ninivae Pisarski, 1971 (Iran, Iraq)

oblongus (Smith, F., 1858) (Bangladesh, India, Myanmar, Pakistan, Sri Lanka)

oblongus binominatus Forel, 1916 (India)

• Camponotus oblongus opaca Forel, 1907

obreptivus Forel, 1899 (Costa Rica)

obscuripes Mayr, 1879 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

obscuriventris Cagniant, 1991 (Morocco)

obtritus Emery, 1911 (Peru)

occasus Emery, 1920 (Cape Verde)

ocreatus Emery, 1893 (Mexico, United States)

• Camponotus acutirostris primipilaris Wheeler, W.M., 1910

oculatior Santschi, 1935 (Democratic Republic of Congo)

odiosus Forel, 1886 (Indonesia)

oertzeni Forel, 1889 (Canary Islands, Greece, Iran, Montenegro, Russian Federation, Turkey)

• Camponotus karawajewi Radchenko, 1996

• Camponotus oertzeni andrius Dalla Torre, 1893

• Camponotus oertzeni kappariensis Dalla Torre, 1893

oetkeri Forel, 1910 (Australia)

oetkeri voltai Forel, 1913 (Australia)

ogasawarensis Terayama & Satoh, 1990 (Japan)

olivieri Forel, 1886 (Angola)

olivieri concordius Santschi, 1915 (Angola)

olivieri delagoensis Forel, 1894 (Mozambique)

olivieri infelix Santschi, 1915 (South Africa)

olivieri lemma Forel, 1886 (Angola)

olivieri moshianus Santschi, 1915 (United Republic of Tanzania)

olivieri osiris Forel, 1911 (Democratic Republic of Congo)

olivieri patersoni Arnold, 1959 (Botswana)

olivieri pax Menozzi, 1942 (Cameroun)

olivieri sorptus Santschi, 1915 (Democratic Republic of Congo)

olivieri tenuipilis Santschi, 1915 (South Africa)

olivieri zaireicus Özdikmen, 2010 (Democratic Republic of Congo)

• Camponotus olivieri nitidior Santschi, 1926

ominosus Forel, 1911 (Sri Lanka)

opaciceps Roger, 1863 (Brazil)

opaciventris Mayr, 1879 (India, Sri Lanka)

orinobates Santschi, 1919 (Kenya, Rwanda)

orinus Dumpert, 1995 (Malaysia)

• Camponotus montanus Dumpert, 1989

orites Santschi, 1919 (South Africa)

• Camponotus orinodromus Santschi, 1919

orombe Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

orthocephalus Emery, 1894 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Peru)

orthodoxus Santschi, 1914 (Guinea, Ivory Coast)

ostiarius Forel, 1914 (South Africa)

ovaticeps (Spinola, 1851) (Chile)

overbecki Viehmeyer, 1916 (Singapore)

owensae Shattuck & McArthur, 2002 (Australia)

oxleyi Forel, 1902 (Australia)

P
pachylepis Emery, 1920 (Bolivia, Colombia, Peru)

palkura McArthur, 2007 (Australia)

pallens (Le Guillou, 1842) (French Polynesia)

pallescens Mayr, 1887 (Brazil)

pallidiceps Emery, 1887 (Australia)

palmyrensis Tohmé, G. & Tohmé, H., 2000 (Syria)

palpatus Emery, 1897 (New Guinea)

panamensis Fernández, 2002 (Panama)

parabarbatus Bharti & Wachkoo, 2014 (India)

paracolobopsis Zettel & Yamane, 2018 (Borneo, Brunei Darussalam)

paradoxus (Mayr, 1866) (Brazil)

• Camponotus paradoxus janitor Forel, 1908

paraleonardi Zettel & Yamane, 2018 (Myanmar, Thailand)

parius Emery, 1889 (Bangladesh, China, India, Laos, Myanmar, Singapore, Sri Lanka)

patimae Wheeler, W.M., 1942 (Guyana)

patimae dolentulus Wheeler, W.M., 1942 (Guyana)

pavidus (Smith, F., 1860) (Indonesia)

pawseyi McArthur, 2003 (Australia)

peleliuensis Clouse, 2007 (Palau)

pellarius Wheeler, W.M., 1914 (Mexico)

pellax Santschi, 1919 (Australia)

pellitus Mayr, 1862 (Argentina, Brazil, Paraguay)

• Camponotus pellitus scintillans Forel, 1901

pennsylvanicus (De Geer, 1773) (Bermuda, Canada, United States)

• Camponotus herculeanus herculeanopennsylvanicus Forel, 1879

• Camponotus herculeanus mahican Emery, 1925

peperi Forel, 1913 (Guatemala)

perjurus Shattuck & McArthur, 2002 (Australia)

perrisii Forel, 1886 (Angola, Kenya)

perrisii densipunctatus Stitz, 1916 (Democratic Republic of Congo)

perrisii insularis Stitz, 1916 (Equatorial Guinea)

• Camponotus foraminosus annobonensis Santschi, 1920

perrisii jucundus Santschi, 1911 (Congo)

perrisii nigeriensis Santschi, 1914 (Nigeria)

personatus Emery, 1894 (Argentina, Brazil, Paraguay)

• Camponotus personatus aloysii Emery, 1925

peseshus Bolton, 1995 (Australia)

• Camponotus nitidiceps Viehmeyer, 1925

petersii Emery, 1895 (Botswana, Lesotho, Mozambique, South Africa, Zimbabwe)

petersii janus Forel, 1911 (South Africa)

pexus Santschi, 1929 (Morocco)

philippinensis (Zettel & Zimmermann, 2007) (Philippines)

philwardi McArthur, 2008 (Australia)

phytophilus Wheeler, W.M., 1934 (Mexico)

piceatus Norton, 1868 (Mexico)

piceus (Leach, 1825) (Albania, Algeria, Andorra, Armenia, Austria, Azerbaijan, Balearic Islands, Belgium, Bulgaria, China, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Italy, Lebanon, Malta, Montenegro, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine)

• Camponotus ebeninus Emery, 1869

• Camponotus figaro Collingwood & Yarrow, 1969

• Formica foveolata Mayr, 1853

picipes (Olivier, 1792) (Brazil, Ecuador, French Guiana, Guyana, Mexico, Suriname)

picipes jamaicensis Wheeler, W.M., 1917 (Jamaica)

picipes jason Forel, 1912 (Bolivia)

picipes pilosulus Emery, 1925 (Mexico)

picipes plombyi Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

picipes spengleri Forel, 1908 (Brazil)

pictostriatus Karavaiev, 1933 (Indonesia)

pilicornis (Roger, 1859) (France, Gibraltar, Iberian Peninsula, Portugal, Spain)

• Camponotus lichtensteini Bondroit, 1918

• Camponotus pilicornis massiliensis Schmitz, 1950

pilicornis siculus Grandi, 1935 (Italy, North Macedonia)

piliventris (Smith, F., 1858) (Australia)

pitjantjatarae McArthur, 2003 (Australia)

pittieri Forel, 1899 (Brazil, Costa Rica, Guadeloupe, Lesser Antilles, Netherlands Antilles)

pittieri fuscogaster Emery, 1920 (Brazil, Peru)

pittieri poenalis Wheeler, W.M., 1921 (Guyana)

placidus (Smith, F., 1858) (Indonesia)

planatus Roger, 1863 (Colombia, Costa Rica, Cuba, Ecuador, El Salvador, Guadeloupe, Guatemala, Honduras, Mexico, Nicaragua, Panama, United States, Venezuela)

planatus acaciae Emery, 1920 (Costa Rica)

planatus colombicus Forel, 1899 (Colombia)

planatus continentis Forel, 1901 (Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico)

planatus esdras Forel, 1916 (Mexico)

planitae Santschi, 1929 (Algeria, Tunisia)

planus Smith, F., 1877 (Ecuador, Galapagos Islands)

planus fernandinensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus fidelis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus hephaestus Wheeler, W.M., 1933 (Ecuador, Galapagos Islands)

planus indefessus Wheeler, W.M., 1919 (Galapagos Islands)

planus isabelensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus peregrinus Emery, 1893 (Ecuador, Galapagos Islands)

planus pinzonensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

planus sansalvadorensis Wheeler, W.M., 1924 (Ecuador, Galapagos Islands)

planus santacruzensis Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

platypus Roger, 1863 (Philippines)

platytarsus Roger, 1863 (French Guiana, Guyana, Mexico, Suriname)

plutus Santschi, 1922 (French Guiana)

poecilus Emery, 1893 (Indonesia)

politae Wu, J. & Wang, C., 1994 (China)

polymorphicus Mackay, W.P., López-Castro & Fernández, 2002 (Colombia)

polyrhachioides Emery, 1898 (Central African Republic, Gabon, Liberia, Sierra Leone)

• Polyrhachis paradoxa (André, 1892)

polyrhachioides cupreus (Santschi, 1923) (Congo)

pompeius Forel, 1886 (Gabon, Guinea, Kenya)

pompeius cassius Wheeler, W.M., 1922 (Democratic Republic of Congo)

pompeius iota Santschi, 1926 (Congo)

pompeius marius Emery, 1899 (Cameroun)

postangulatus Emery, 1911 (Peru)

postcornutus Clark, 1930 (Australia)

posteropilus Shattuck, 2005 (New Guinea)

posticus Santschi, 1926 (Democratic Republic of Congo, Kenya)

postoculatus Forel, 1914 (South Africa)

praegracilis Karaman & Kiran, 2017 (Turkey)

pressipes Emery, 1893 (Borneo, Indonesia, Malaysia, Philippines)

pressipes errans Forel, 1913 (Indonesia)

propinquellus Emery, 1920 (Peru)

propinquus Mayr, 1887 (Brazil, Colombia)

• Camponotus paris Forel, 1912

propinquus baretoi Santschi, 1922 (Argentina)

prosseri Shattuck & McArthur, 2002 (Australia)

prostans Forel, 1910 (Australia)

prosulcatus Santschi, 1935 (Democratic Republic of Congo)

pseudoirritans Wu, J. & Wang, C., 1989 (China)

pseudolendus Wu, J. & Wang, C., 1989 (China)

puberulus Emery, 1897 (Eritrea, Somalia)

pudorosus Emery, 1925 (Mexico, United States)

pulchellus Forel, 1894 (New Caledonia)

pulcher Forel, 1892 (Madagascar)

pullatus Mayr, 1866 (Mexico)

pulvinatus Mayr, 1907 (United Republic of Tanzania)

punctaticeps (Mayr, 1867) (Indonesia)

punctatissimus Forel, 1907 (Myanmar)

punctatus Forel, 1912 (Argentina, Brazil)

punctiventris Emery, 1920 (Australia)

punctulatus Mayr, 1868 (Argentina, Bolivia, Brazil, Mexico, Paraguay)

punctulatus andigenus Emery, 1903 (Peru)

punctulatus brevibarbis Santschi, 1939 (Bolivia)

punctulatus chubutensis Forel, 1913 (Argentina, Brazil)

punctulatus cruentus Santschi, 1922 (Argentina)

punctulatus hybridus Forel, 1901 (Argentina, Brazil)

punctulatus imberbis Emery, 1906 (Argentina)

punctulatus lillii Forel, 1908 (Brazil)

punctulatus lizeri Santschi, 1922 (Bolivia)

punctulatus minutior Forel, 1886 (Argentina)

punctulatus pergandei Emery, 1925 (Mexico)

• Camponotus punctulatus ruficornis Pergande, 1896

punctulatus tenuibarbis Santschi, 1939 (Argentina)

puniceps Donisthorpe, 1942 (India)

pupillus Santschi, 1939 (Algeria, Morocco)

putatus Forel, 1892 (Madagascar)

Q
quadrimaculatus Forel, 1886 (Comoros, Madagascar, Mayotte)

• Camponotus kelleri Forel, 1886

• Camponotus kelleri invalidus Forel, 1897

• Camponotus quadrimaculatus sellaris Emery, 1895

quadrinotatus Forel, 1886 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Camponotus marginatus nigricolor Ruzsky, 1926

• Camponotus marginatus rubicolor Ruzsky, 1925

quadrisectus (Smith, F., 1858) (Borneo, Indonesia, Malaysia, Philippines)

quadrisectus hians Forel, 1907 (Singapore)

quadrisectus margaritae Forel, 1907 (Singapore)

R
radamae Forel, 1891 (Madagascar, Mayotte)

radiatus Forel, 1892 (India)

radovae Forel, 1886 (Madagascar)

• Camponotus radovae radovaedarwinii Forel, 1891

raina Rakotonirina & Fisher, 2018 (Madagascar)

ramulorum Wheeler, W.M., 1905 (Bahamas, Cuba)

ramulorum marcidus Wheeler, W.M., 1905 (Bahamas)

ramulorum mestrei Wheeler, W.M., 1913 (Cuba, Greater Antilles)

ramulorum vernulus Wheeler, W.M., 1936 (Haiti)

rapax (Fabricius, 1804) (Brazil, Ecuador, French Guiana, Guyana, Suriname)

raphaelis Forel, 1899 (Colombia, Costa Rica, Guatemala, Honduras, Mexico)

reaumuri Forel, 1892 (Madagascar)

rebeccae Forel, 1913 (Cyprus, Greece, Iran, Israel, Jordan, Lebanon, Syria, Turkey)

• Camponotus lateralis cypridis Santschi, 1939

reburrus Mackay, W.P., 2013 (Ecuador)

rectangularis Emery, 1890 (Belize, Brazil, Costa Rica, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Suriname)

rectangularis aulicus Wheeler, W.M., 1933 (Guatemala, Mexico)

rectangularis rubroniger Forel, 1899 (El Salvador, Mexico)

rectangularis setipes Forel, 1904 (Trinidad and Tobago)

rectangularis sordidatus Wheeler, W.M., 1934 (Peru)

rectangularis willowsi Wheeler, W.M., 1933 (Mexico)

rectithorax Forel, 1895 (Guatemala, Mexico, Nicaragua)

• Camponotus nitida Norton, 1868

reevei Arnold, 1922 (South Africa)

reichardti Arnoldi, 1967 (China, Kyrgyzstan, Uzbekistan)

reinaldi Kempf, 1960 (Brazil)

renggeri Emery, 1894 (Argentina, Bolivia, Brazil, Ecuador, French Guiana, Guyana, Paraguay, Peru, Suriname, Uruguay)

repens Forel, 1897 (Madagascar)

• Mayria madagascarensis Forel, 1886

reticulatus Roger, 1863 (Borneo, Indonesia, Malaysia, Micronesia (Federated States of), Northern Mariana Islands, Palau, Seychelles, Singapore, Sri Lanka, Wallis and Futuna Islands)

reticulatus fullawayi Wheeler, W.M., 1912 (Guam)

reticulatus gestiens Forel, 1915 (Indonesia)

reticulatus imparilis Forel, 1915 (Indonesia)

reticulatus jagori Stitz, 1925 (Philippines)

reticulatus latitans Forel, 1893 (India, Sri Lanka)

reticulatus sericellus Viehmeyer, 1916 (Singapore)

reticulatus yerburyi Forel, 1893 (Sri Lanka)

• Camponotus bedoti motschulskii Emery, 1893

rhamses Santschi, 1915 (Egypt)

rhamses completus Santschi, 1930 (Ethiopia)

riedeli Pisarski, 1971 (Iraq)

robecchii Emery, 1892 (Eritrea, Kenya, Namibia, Somalia, Yemen, Zimbabwe)

robecchii abyssinicus Santschi, 1913 (Ethiopia)

robecchii assumptionis Crawley, 1921 (Seychelles)

robecchii dispar Stitz, 1923 (Namibia)

robecchii rhodesianus Forel, 1913 (Zimbabwe)

robecchii troglodytes Forel, 1894 (Botswana, Kenya, Mozambique, Namibia)

robertae Santschi, 1926 (Ivory Coast)

robustus Roger, 1863 (Madagascar)

roeseli Forel, 1910 (Madagascar)

• Camponotus maculatus legionarium Santschi, 1911

rosariensis Forel, 1912 (Argentina, Paraguay)

rotrae Rakotonirina & Fisher, 2022 (Madagascar)

rotumanus Wilson & Taylor, 1967 (Fiji)

rotundinodis Santschi, 1935 (Democratic Republic of Congo)

roubaudi Santschi, 1911 (Congo)

royi Shattuck & Janda, 2009 (New Guinea)

ruber Emery, 1925 (Algeria, Balearic Islands, Portugal, Spain)

rubidus Xiao & Wang, C., 1989 (China)

rubiginosus Mayr, 1876 (Australia)

rubripes (Latreille, 1802) (Kenya, Sierra Leone)

rubrithorax Forel, 1899 (Mexico)

rubrithorax nigrithorax Stitz, 1937 (Mexico)

rudis McArthur, 2003 (Australia)

rufifemur Emery, 1900 (Borneo, India, Indonesia, Malaysia, Thailand)

rufifemur obscurus Stitz, 1925 (Philippines)

rufigaster Menozzi, 1928 (Indonesia)

rufipes (Fabricius, 1775) (Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Paraguay, Suriname, Uruguay, Venezuela)

• Camponotus cajurensis Luederwaldt, 1918

• Camponotus rufipes alpina Emery, 1906

• Camponotus rufipes magnifica Forel, 1913

• Camponotus rufipes subrufescens Santschi, 1929

• Formica herrichi Mayr, 1853

• Formica merdicola Lund, 1831

rufoglaucus (Jerdon, 1851) (Bangladesh, Borneo, Cambodia, Canary Islands, China, India, Indonesia, Kenya, Laos, Malaysia, Namibia, Nepal, Rwanda, Sri Lanka, Thailand, Vietnam, Zimbabwe)

• Camponotus redtenbacheri Mayr, 1862

rufoglaucus controversus Santschi, 1916 (Benin, United Republic of Tanzania)

• Camponotus rufoglaucus flavopilosus Viehmeyer, 1913

rufoglaucus latericius Stitz, 1923 (Namibia)

rufoglaucus syphax Wheeler, W.M., 1922 (Democratic Republic of Congo, Rwanda)

rufoglaucus tenuis Forel, 1907 (India)

rufoglaucus zanzibaricus Forel, 1911 (United Republic of Tanzania)

rufoglaucus zulu Emery, 1895 (South Africa)

rufonigrus Shattuck & McArthur, 2002 (Australia)

rufus Crawley, 1925 (Australia)

ruseni Karaman, C., 2012 (Turkey)

rusticus Santschi, 1916 (Argentina)

ruzskyellus Forel, 1922 (Iran, Iraq, Kazakhstan)

S
sacchii Emery, 1899 (Somalia)

sada Rakotonirina & Fisher, 2018 (Madagascar)

salvini Forel, 1899 (Costa Rica, Mexico, Panama)

sambiranoensis Rakotonirina & Fisher, 2022 (Madagascar)

samius Forel, 1889 (Bulgaria, Greece, Romania, Turkey)

• Camponotus maculatus spagnolinii Emery, 1920

sanctaefidei Dalla Torre, 1892 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Panama, Paraguay, Peru)

• Camponotus quadrilaterus Mayr Mayr, 1887

sanctaefidei convexinodis Wheeler, W.M., 1934 (Brazil)

sanctaefidei coronatus Santschi, 1922 (Brazil)

sanctaefidei hondurianus Mann, 1922 (Honduras)

sanctaefidei leonhardi Forel, 1901 (Bolivia)

sanctaefidei weberi Wheeler, W.M., 1935 (Trinidad and Tobago)

• Camponotus sanctaefidei darlingtoni Wheeler, W.M., 1934

sanctus Forel, 1904 (Afghanistan, Greece, Iran, Israel, Jordan, Lebanon, Syria, Turkey)

• Camponotus compressus cosensis Finzi, 1939

• Camponotus maculatus symiensis Forel, 1910

sankisianus Forel, 1913 (Democratic Republic of Congo)

sannini Tohmé, G. & Tohmé, H., 1999 (Greece, Israel, Lebanon, Syria)

sansabeanus (Buckley, 1866) (Mexico, United States)

• Camponotus maculatus bulimosus Wheeler, W.M., 1910

• Camponotus maculatus torrefactus Emery, 1925

santosi Forel, 1908 (Bahamas, Cuba, Greater Antilles)

santosi pazosi Santschi, 1922 (Cuba)

santschii Forel, 1899 (Colombia, Ecuador)

satan Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

saussurei Forel, 1879 (Dominican Republic, Haiti, Lesser Antilles)

saxatilis Ruzsky, 1895 (China, Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

sayi Emery, 1893 (Mexico, United States)

scabrinodis Arnold, 1924 (Zimbabwe)

scalaris Forel, 1901 (South Africa)

schaefferi Wheeler, W.M., 1909 (United States)

schneei Mayr, 1903 (Marshall Islands)

schoedli Dumpert, 2006 (Thailand)

schoutedeni Forel, 1911 (Benin, Benin, Democratic Republic of Congo, Kenya, Zimbabwe)

schulzi Salata, Loss, Karaman, Kiran, Borowiec, 2019 (Turkey)

schulzianus Zettel & Balàka, 2018 (Thailand)

scipio Forel, 1908 (Argentina, Paraguay)

scipio insignis Santschi, 1922 (Paraguay)

scissus Mayr, 1887 (Brazil)

scotti McArthur, 2003 (Australia)

scratius Forel, 1907 (Australia)

• Camponotus samueli McArthur, 2008

scratius nuntius Forel, 1907 (Australia)

sculptor Santschi, 1920 (Argentina)

sedulus (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

selene (Emery, 1889) (China, India, Myanmar)

selene obtusatus (Emery, 1895) (India, Myanmar)

sellidorsatus Prins, 1973 (South Africa)

semipunctata Kirby, W., 1837

semirufus Emery, 1925 (China, Kazakhstan, Kyrgyzstan, Turkmenistan)

semitestaceus Snelling, R.R., 1970 (Mexico, United States)

semoni Forel, 1905 (Venezuela)

senex (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

• Tapinoma tomentosa Norton, 1868

senkakuensis Terayama, 2013 (Japan)

septentrionalis (Buckley, 1866)

sericatus Mayr, 1887 (Brazil)

sericeiventris (Guérin-Méneville, 1838) (Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Suriname, Uruguay)

• Formica cuneata Perty, 1833

sericeiventris holmgreni Wheeler, W.M., 1931 (Bolivia)

sericeiventris otoquensis Wheeler, W.M., 1931 (Panama)

sericeiventris pontifex Santschi, 1936 (Mexico)

• Camponotus sericeiventris imperator Wheeler, W.M., 1934

sericeiventris rex Forel, 1907 (Costa Rica, Guatemala, Mexico)

sericeiventris satrapus Wheeler, W.M., 1931 (Brazil, Colombia, Costa Rica, Ecuador, Honduras, Panama, Peru)

sericeus (Fabricius, 1798) (Afghanistan, Algeria, Benin, Burkina Faso, Egypt, Eritrea, Gambia, India, Israel, Ivory Coast, Kenya, Kenya, Kuwait, Mali, Mauritius, Oman, Pakistan, Saudi Arabia, Senegal, Sri Lanka, Sudan, Thailand, United Arab Emirates, Zimbabwe)

• Formica aurulenta Latreille, 1802

• Formica pyrrhocephala Motschoulsky, 1863

sericeus euchrous Santschi, 1926 (Niger, Sudan)

sericeus obtusus (Smith, F., 1858) (Egypt)

sericeus peguensis Emery, 1895 (India, Myanmar)

sericeus sanguiniceps Donisthorpe, 1942 (Myanmar)

sericeus sulgeri Santschi, 1913 (Congo)

serotinus Cagniant, 1996 (Morocco)

sesquipedalis Roger, 1863 (Sri Lanka)

setitibia Forel, 1901 (Borneo, Indonesia, Malaysia)

setosus Shattuck & McArthur, 2002 (Australia)

seurati Emery, 1920 (Algeria)

sexguttatus (Fabricius, 1793) (Argentina, Barbados, Bolivia, Brazil, Colombia, Costa Rica, Dominican Republic, Ecuador, French Guiana, Greater Antilles, Grenada, Guadeloupe, Guyana, Honduras, Lesser Antilles, Martinique, Netherlands Antilles, Nicaragua, Panama, Paraguay, Peru, Puerto Rico, Saint Lucia, Suriname, United States, Uruguay)

• Camponotus sexguttatus fusciceps Emery, 1906

• Camponotus sexguttatus grenadensis Forel, 1897

• Formica albofasciata Smith, F., 1862

• Formica bimaculata Smith, F., 1858

• Formica ruficeps Fabricius Fabricius, 1804

sexguttatus albotaeniolatus Forel, 1921 (Ecuador)

sexguttatus antiguanus Wheeler, W.M., 1923 (Antigua and Barbuda, Guadeloupe)

sexguttatus basirectus Wheeler, W.M., 1923 (Brazil)

sexguttatus biguttatus Emery, 1894 (Bolivia)

sexguttatus decorus (Smith, F., 1858) (Brazil)

sexguttatus montserratensis Wheeler, W.M., 1923 (Lesser Antilles)

sexguttatus ornatus Emery, 1894 (Bolivia)

sexguttatus perturbans Kutter, 1931 (Colombia)

sexguttatus unitaeniatus Wheeler, W.M., 1923 (Peru)

sexpunctatus Forel, 1894 (Mozambique)

shaqualavensis Pisarski, 1971 (Iran, Iraq, Turkey)

• Camponotus arnoldi Pisarski, 1971

• Camponotus babau Bolton, 1995

siamensis Jaitrong & Jeenthong, 2022 (Thailand)

sibreei Forel, 1891 (Madagascar)

sicheli Mayr, 1866 (Algeria, Balearic Islands, Canary Islands, Gibraltar, Iberian Peninsula, Malta)

sicheli niger Emery, 1925 (Algeria)

siemsseni Forel, 1901 (China, India, Indonesia)

siemsseni dispar Forel, 1902 (India)

sikorae Emery, 1920 (Madagascar)

silvestrii Emery, 1906 (Brazil)

silvicola Forel, 1902 (Argentina, Argentina, Colombia, Ecuador, Paraguay)

simillimus (Smith, F., 1862) (Brazil, Colombia, Ecuador, French Guiana, Guyana, Panama, Suriname)

simillimus atratior Santschi, 1922 (Brazil)

simillimus impatibilis Forel, 1899 (Costa Rica)

simillimus riograndensis Emery, 1888 (Brazil)

simoni Emery, 1893 (Sri Lanka)

simpsoni McArthur, 2003 (Australia)

simulans Forel, 1910 (Namibia, Zimbabwe)

simulator Forel, 1915 (Australia)

simus Emery, 1908 (Democratic Republic of Congo)

simus manidis Forel, 1909 (Congo, Democratic Republic of Congo)

sinaiticus Ionescu-Hirsch, 2010 (Egypt, Israel, Jordan)

singularis (Smith, F., 1858) (Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Nepal, Thailand, Vietnam)

• Cataglyphis birmana Collingwood, 1962

singularis rufomaculatus Donisthorpe, 1941 (Borneo, Indonesia, Malaysia)

sklarus Bolton, 1995 (India, Myanmar, Sri Lanka)

• Camponotus similis Donisthorpe, 1933

snellingi Bolton, 1995 (United States)

• Camponotus fallax pavidus Snelling, R.R., 1988

socius Roger, 1863 (Brazil, United States)

• Camponotus socius osceola Wheeler, W.M., 1932

socorroensis Wheeler, W.M., 1934 (Mexico)

socrates Forel, 1904 (India, Pakistan)

solon Forel, 1886 (Angola, Benin, Guinea, Ivory Coast, Kenya, Senegal)

solon chiton Emery, 1925 (Senegal)

solon jugurtha Emery, 1925 (Democratic Republic of Congo)

somalinus André, 1887 (Eritrea, Kenya, Oman, Somalia)

somalinus curtior Forel, 1894 (Ethiopia)

somalinus pattensis Forel, 1907 (Kenya)

sophiae Zettel & Balàka, 2018 (Thailand)

spanis Xiao & Wang, C., 1989 (China)

sphaericus Roger, 1863 (Cuba)

sphaericus cardini Mann, 1920 (Cuba)

sphaericus rufipilis Aguayo, 1932 (Cuba, Greater Antilles)

sphaericus sphaeralis Roger, 1863 (Cuba)

sphenocephalus Emery, 1911 (Peru)

sphenoidalis Mayr, 1870 (Colombia)

spinitarsus Emery, 1920 (Australia)

spinolae Roger, 1863 (Chile)

• Camponotus chilensis ruficornis Emery, 1895

spissinodis Forel, 1909 (Algeria)

• Camponotus lateralis crassinodis Forel, 1894

sponsorum Forel, 1910 (Australia)

staryi Pisarski, 1971 (Iran, Iraq)

stefani McArthur, 2007 (Australia)

stefanschoedli (Zettel & Zimmermann, 2007) (Philippines)

storeatus Forel, 1910 (Namibia, South Africa)

strangulatus Santschi, 1911 (Comoros, Madagascar, Mayotte, Seychelles)

• Camponotus hova maculatoides Emery, 1920

striatipes Dumpert, 1995 (Borneo, Indonesia, Malaysia)

striatus (Smith, F., 1862) (Colombia, Costa Rica, Honduras, Mexico, Panama)

• Camponotus alfaroi Emery, 1890

• Dolichoderus granulatus Pergande, 1896

subbarbatus Emery, 1893 (United States)

subcircularis Emery, 1920 (Bolivia)

subnitidus Mayr, 1876 (Australia)

subnitidus famelicus Emery, 1887 (Australia)

subnitidus longinodis Forel, 1915 (Australia)

subpilus Shattuck, 2005 (Indonesia, New Guinea)

substitutus Emery, 1894 (Argentina, Brazil, Chile, Colombia, French Guiana, Guatemala, Guyana, Mexico, Panama, Paraguay, Suriname, Uruguay, Venezuela)

• Camponotus melanoticus multipilis Forel, 1907

substitutus clarus Stitz, 1938 (Argentina)

substitutus pullulus Emery, 1925 (Argentina)

subtilis (Smith, F., 1860) (Indonesia)

subtruncatus Borgmeier, 1929 (Brazil)

sucki Forel, 1901 (Borneo, Indonesia, Malaysia)

suffusus (Smith, F., 1858) (Australia)

• Camponotus schencki Mayr, 1862

suffusus bendigensis Forel, 1902 (Australia)

syaukanii Zettel & Yamane, 2018 (Borneo, Indonesia)

sylvaticus (Olivier, 1792) (Bulgaria, Canary Islands, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Luxembourg, Portugal, Spain, Turkey)

• Camponotus sylvaticus catalana Emery, 1924

• Formica castanipes Leach, 1825

sylvaticus basalis Smith, F., 1878 (India, Pakistan)

• Camponotus maculatus lobinieri Forel, 1902

sylvaticus paradichrous Emery, 1925 (India, Pakistan)

T
tafo Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

tahatensis Santschi, 1929 (Algeria)

taino Fontenla Rizo Fontenla Rizo, 1997

taino Snelling & Torres Snelling, R.R. & Torres, 1998

taipingensis Forel, 1913 (Malaysia)

taniae Mackay, W.P. & Delsinne, 2009 (Argentina, Paraguay)

tanosy Rakotonirina & Fisher, 2018 (Madagascar)

tapia Rakotonirina & Fisher, 2022 (Madagascar)

tashcumiri Tarbinsky, 1976 (Kyrgyzstan, Mongolia)

tasmani Forel, 1902 (Australia)

tauricollis Forel, 1894 (Mozambique)

tejonia (Buckley, 1866)

tendryi Rakotonirina & Fisher, 2022 (Madagascar)

tenuipes (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

tenuiscapus Roger, 1863 (Argentina, Brazil, Uruguay)

tepicanus Pergande, 1896 (Mexico)

terbimaculatus Emery, 1920 (Brazil)

terebrans (Lowne, 1865) (Australia)

• Camponotus latrunculus victoriensis Santschi, 1928

• Camponotus myoporus Clark, 1938

• Formica testaceipes Smith, F. Smith, F., 1858

tergestinus Müller, 1921 (Bulgaria, Hungary, Italy, Romania, Slovenia, Turkey)

termitarius Emery, 1902 (Argentina, Brazil, Paraguay)

terricola Karavaiev, 1929 (Indonesia)

territus Santschi, 1939 (Argentina)

testaceus Emery, 1894 (Brazil)

texanus Wheeler, W.M., 1903 (United States)

texens Dumpert, 1986 (Malaysia)

textor Forel, 1899 (Costa Rica, Honduras, Panama)

thadeus Shattuck, 2005 (Australia)

thales Forel, 1910 (Lesotho)

themistocles Forel, 1910 (Madagascar)

thomasseti Forel, 1912 (Seychelles)

thoracicus (Fabricius, 1804) (Algeria, Iran, Kuwait, Oman, Saudi Arabia, Turkey, United Arab Emirates, Yemen)

• Camponotus sylvaticus dichrous André, 1882

thraso Forel, 1893 (India, Sri Lanka)

thraso agricola Forel, 1910 (South Africa)

thraso assabensis Emery, 1925 (Ethiopia)

thraso diogenes Forel, 1909 (Sri Lanka)

thraso montinanus Santschi, 1926 (Zimbabwe)

thraso nefasitensis Menozzi, 1931 (Ethiopia)

thraso negus Forel, 1907 (Ethiopia, Kenya, Zimbabwe)

thysanopus Wheeler, W.M., 1937 (Cuba, Greater Antilles)

tilhoi Santschi, 1926 (Chad)

timidus (Jerdon, 1851) (India)

tinctus (Smith, F., 1858) (Myanmar, Singapore)

tol Gibson & Clouse, 2016 (Micronesia (Federated States of))

tonduzi Forel, 1899 (Colombia, Costa Rica, Panama)

tonkinus Santschi, 1925 (China)

torrei Aguayo, 1932 (Cuba, Greater Antilles)

toussainti Wheeler, W.M. & Mann, 1914 (Greater Antilles, Haiti)

traegaordhi Santschi, 1914 (South Africa)

traegaordhi fumeus Santschi, 1926 (Zimbabwe)

traili Mayr, 1878 (Bolivia, Brazil, Colombia, Ecuador, Panama, Venezuela)

• Dendromyrmex graciliceps Viehmeyer, 1916

• Dendromyrmex madeirensis Mann, 1916

• Dendromyrmex madeirensis romani Wheeler, W.M., 1923

• Dendromyrmex traili rufogaster Mann, 1916

trajanus Forel, 1912 (Indonesia)

transvaalensis Arnold, 1948 (South Africa)

transvaalensis griqua Arnold, 1952 (South Africa)

transvaalensis prinsi Özdikmen, 2010 (Zimbabwe)

• Camponotus transvaalensis arnoldi Prins, 1965

trapeziceps Forel, 1908 (Argentina, Brazil, Paraguay)

trapeziceps prosaicus Santschi, 1922 (Brazil)

trapezoideus Mayr, 1870 (Brazil, Colombia)

tratra Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

trepidulus Creighton, 1965 (Mexico, United States)

triangulatus Klimes & McArthur, 2014 (New Guinea)

tricolor Weber, 1943 (Cameroun, New Guinea, Sudan)

• Camponotus tameri Bolton, 1995

tricoloratus Clark, 1941 (Australia)

trietericus Menozzi, 1926 (Philippines)

trifasciatus Santschi, 1926 (South Africa)

triodiae McArthur, 2009 (Australia)

tripartitus Mayr, 1887 (Brazil)

• Camponotus tripartitus collegianus Santschi, 1929

tristis Clark, 1930 (Australia)

truebi Forel, 1910 (Taiwan)

• Camponotus itoi genaiai Santschi, 1928

• Camponotus truebi arnoldi Forel, 1912

tsimelahy Rasoamanana & Fisher, 2022 (Madagascar)

tumidus Crawley, 1922 (Australia)

turkestanicus Emery, 1887 (Afghanistan, Azerbaijan, China, Georgia, Iran, Kazakhstan, Kyrgyzstan, Russian Federation, Turkmenistan)

• Camponotus attila Stärcke, 1935

• Camponotus maculatus kaschgariensis Forel, 1901

turkestanus André, 1882 (Afghanistan, Azerbaijan, China, Israel, Kazakhstan, Kyrgyzstan, Lebanon, Mongolia, Russian Federation, Turkey, Turkmenistan)

• Camponotus maculatus ferganensis Pisarski, 1967

• Camponotus maculatus flavonitidus Emery, 1925

• Camponotus maculatus ruzskyi Vashkevich, 1924

U
ulcerosus Wheeler, W.M., 1910 (Mexico, United States)

• Camponotus bruesi Wheeler, W.M., 1910

ulei Forel, 1904 (Peru)

ulvarum Forel, 1899 (Costa Rica)

universitatis Forel, 1890 (Albania, Bulgaria, France, Germany, Iberian Peninsula, Italy, Spain, Switzerland, Turkey)

urichi Forel, 1899 (Brazil, Trinidad and Tobago)

urichi sculnus Forel, 1904 (Brazil)

ursus Forel, 1886 (Madagascar)

ustus Forel, 1879 (Colombia, Dominican Republic, Greater Antilles, Grenada, Haiti, Puerto Rico)

• Camponotus larvigerus Wheeler, W.M. & Mann, 1914

• Camponotus larvigerus maculifrons Menozzi & Russo, 1930

• Camponotus ustus depolitus Wheeler, W.M., 1936

• Camponotus ustus furnissi Wheeler, W.M. & Mann, 1914

• Camponotus ustus sublautus Wheeler, W.M. & Mann, 1914

• Camponotus ustus ulysses Forel, 1907

V
vafer Wheeler, W.M., 1910 (Mexico, United States)

vagulus Forel, 1908 (Argentina, Brazil, Paraguay)

vagus (Scopoli, 1763) (Albania, Algeria, Andorra, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Turkey, Ukraine)

• Camponotus vagus kodorica Forel, 1913

• Formica fuscoptera Geoffroy, 1785

• Formica pubescens Fabricius Fabricius, 1775

vagus ifranensis Cagniant, 1987 (Morocco)

valdeziae Forel, 1879 (South Africa, Zimbabwe)

vanispinus Xia & Zheng, 1997 (China)

vano Rakotonirina & Fisher, 2022 (Madagascar)

varatra Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

varians Roger, 1863 (India)

variegatus (Smith, F., 1858) (Bangladesh, China, Hawaii, India, Micronesia (Federated States of), Pakistan, Singapore, Sri Lanka, Turkey)

• Camponotus variegatus hawaiensis Emery, 1920

variegatus ambonensis Karavaiev, 1930 (Indonesia)

variegatus bacchus (Smith, F., 1858) (India, Sri Lanka)

• Formica ventralis Smith, F., 1858

variegatus batta Menozzi, 1930 (Indonesia)

variegatus cleon Forel, 1913 (Indonesia)

variegatus comottoi Emery, 1887 (Myanmar)

variegatus crassinodis Forel, 1892 (Myanmar, Philippines, Sri Lanka)

variegatus dulcis Dalla Torre, 1893 (China, India, Myanmar, Pakistan, Philippines, Sri Lanka)

variegatus flavotestaceus Donisthorpe, 1948 (New Guinea)

variegatus fuscithorax Dalla Torre, 1893 (India, Laos, Singapore, Sri Lanka)

variegatus infuscus Forel, 1892 (India, Sri Lanka)

variegatus intrans Forel, 1911 (Sri Lanka)

variegatus papua Chapman & Capco, 1951 (New Guinea)

variegatus proles Emery, 1925 (Laos)

variegatus somnificus Forel, 1902 (India, Sri Lanka)

variegatus stenonotus Stitz, 1938 (Borneo, Indonesia, Malaysia)

varius Donisthorpe, 1943 (India)

varus Forel, 1910 (Angola)

velox (Jerdon, 1851) (India)

verae Forel, 1908 (Brazil)

versicolor Clark, 1930 (Australia)

vespertinus Arnold, 1960 (Zimbabwe)

vestitus (Smith, F., 1858) (Benin, Burkina Faso, Guinea-Bissau, Ivory Coast, Mali, Namibia, Senegal, South Africa, Zimbabwe)

vestitus anthracinus Santschi, 1930 (Angola)

vestitus bombycinus Santschi, 1930 (Angola)

vestitus comptus Santschi, 1926 (Ivory Coast)

vestitus intuens Santschi, 1926 (Angola, Eritrea)

vestitus lujai Santschi, 1930 (Democratic Republic of Congo)

vestitus pectitus Santschi, 1930 (Angola)

vestitus perpectitus Santschi, 1926 (Benin)

vestitus strophiatus Santschi, 1926 (South Africa)

vicinus Mayr, 1870 (Canada, Mexico, United States)

• Camponotus vicinus nitidiventris Wheeler, 1935

victoriae Arnold, 1959 (South Africa)

viehmeyeri Forel, 1911 (New Guinea)

vigilans (Smith, F., 1858) (Borneo, China, Indonesia, Malaysia)

vinosus (Smith, F., 1858) (Brazil)

viri Santschi, 1915 (Kenya)

virulens (Smith, F., 1861) (Indonesia, Sulawesi)

vitiosus Smith, F., 1874 (China, Japan, Republic of Korea)

• Camponotus itoi tokioensis Ito, 1912

• Camponotus tokioensis atrigenatus Santschi, 1937

• Camponotus tokioensis inconstans Santschi, 1937

vittatus Forel, 1904 (Brazil)

vividus (Smith, F., 1858) (Benin, Gambia, Guinea, Ivory Coast, Kenya, Liberia, Mozambique, Senegal, Sierra Leone, Uganda, Zimbabwe)

• Formica laboriosa Smith, F., 1858

vividus cato Forel, 1913 (Democratic Republic of Congo, Uganda)

• Camponotus vividus laevithorax Menozzi, 1924

vividus meinerti Forel, 1886 (Angola, Chad)

vividus reginae Forel, 1901 (Angola)

vividus semidepilis Wheeler, W.M., 1922 (Democratic Republic of Congo)

voeltzkowii Forel, 1894 (Madagascar)

vogti Forel, 1906 (Greece, Iran, Iraq, Israel, Turkey)

vulpus Santschi, 1926 (Democratic Republic of Congo)

W
wanangus Klimes & McArthur, 2014 (New Guinea)

wasmanni Emery, 1893 (Bangladesh, China, India)

wedda Forel, 1908 (Sri Lanka)

weiserti Zettel & Laciny, 2018 (Thailand)

weismanni Forel, 1901 (New Guinea)

weissflogi Dumpert, 2006 (Malaysia)

wellmani Forel, 1909 (Angola, Guinea)

wellmani gamma Santschi, 1926 (Democratic Republic of Congo)

wellmani rufipartis Forel, 1916 (Democratic Republic of Congo)

werthi Forel, 1908 (Kerguelen Islands)

werthi skaifei Arnold, 1959 (South Africa)

westermanni Mayr, 1862 (Argentina, Brazil, Paraguay)

westermanni fulvicornis Emery, 1903 (Brazil)

wheeleri Mann, 1916 (Brazil, Ecuador)

whitei Wheeler, W.M., 1915 (Australia)

• Camponotus scutellus Clark, 1930

wiederkehri Forel, 1894 (Australia)

• Camponotus denticulatus Kirby, W.F., 1896

• Camponotus latrunculus Wheeler, W.M., 1915

• Camponotus wiederkehri lucidior Forel, 1910

wolfi Emery, 1920 (Cameroun, Central African Republic, Kenya, Uganda)

woodroffeensis McArthur, 2008 (Australia)

wroughtonii Forel, 1893 (India, Nepal)

wytsmani Emery, 1920 (Bolivia, Ecuador)

X
xanthogaster Santschi, 1925 (Brazil)

xanthopilus Shattuck, 2005 (New Guinea)

xerxes Forel, 1904 (Afghanistan, Egypt, Iran, Iraq, Israel, Oman, Qatar, Saudi Arabia, Turkey, Turkmenistan, United Arab Emirates, Uzbekistan)

• Camponotus compressus irakensis Menozzi, 1927

xingdoushanensis Wang, W. & Chen, 2003 (China)

Y
yala Kusnezov, 1952 (Argentina)

yamaokai Terayama & Satoh, 1990 (Japan)

yambaru Terayama, 1999 (Japan)

yessensis Yasumatsu & Brown, 1951 (Japan)

yiningensis Wang, C. & Wu, J., 1994 (China)

yogi Wheeler, W.M., 1915 (Mexico, United States)

• Colobopsis apostemata Mackay, 2018

• Colobopsis cavibregma Mackay, 2018

Z
zavo Rakotonirina, Csősz & Fisher, 2016 (Madagascar)

zenon Forel, 1912 (Brazil)

zenon criton Emery, 1925 (Brazil)

zimmermanni Forel, 1894 (Ethiopia)

zimmermanni pansus Santschi, 1926 (Kenya)

zoc Forel, 1879 (Colombia, French Guiana, Guatemala, Guyana, Suriname, Trinidad and Tobago, Venezuela)

zonatus Emery, 1894 (Costa Rica, Ecuador, Galapagos Islands, Honduras, Mexico, Panama)

zoro Rasoamanana & Fisher, 2022 (Madagascar)

=Fossil Species=

A
ambon Zhang, J., 1989

ampullosus Zhang, J., 1989

C
cockerelli (Donisthorpe, 1920)

compactus Förster, 1891

crozei Riou, 1999

curviansatus Zhang, J., 1989

F
fuscipennis Carpenter, 1930

G
gracilis Zhang, J., 1989

H
heracleus (Heer, 1849)

I
induratus (Heer, 1849)

J
juliae Pierce & Gibron, 1962 (Pierce & Gibron, 1962)

L
lignitus (Germar, 1837)

longiventris Théobald, 1937

longus Zhang, J., 1989

M
mengei Mayr, 1868

microcephalus Carpenter, 1930

microthoracus Zhang, J., 1989

N
novotnyi Samsinak, 1967

O
obesus Piton, 1935

P
parvus Perfilieva, 2022

penninervis Théobald, 1937

petrifactus Carpenter, 1930

pictus Zhang, J., Sun & Zhang, X., 1994

plenus Zhang, J., 1989

S
shanwangensis Hong, 1984

T
theobaldi Özdikmen, 2010

tokunagai Naora, 1933

V
vehemens Förster, 1891

vetus Scudder, 1877