Checklist of Tetramorium species

The following species and subspecies belong to the genus Tetramorium. Synonyms are listed under their senior names.

A
aculeatum (Mayr, 1866) (Ghana)

• Macromischa wasmanni Forel, 1901

• Macromischoides aculeatus inermis Bernard, 1953

• Macromischoides aculeatus melanogyne Santschi, 1923

• Macromischoides aculeatus militaris Santschi, 1924

• Macromischoides aculeatus pulchellus Santschi, 1924

• Macromischoides viridis Weber, 1943

• Macromischoides zumpti Santschi, 1937

• Tetramorium aculeatum andricum Emery, 1908

• Tetramorium aculeatum major Forel, 1915

• Tetramorium aculeatum rubroflava Forel, 1916

adamsi Hita Garcia & Fisher, 2012 (Madagascar)

adelphon Bolton, 1979 (Borneo)

adpressum (Bolton, 1976) (Indonesia (Sulawesi))

aegeum Radchenko, 1992 (Caucasus)

• Tetramorium inerme laevigatum Arnol'di, 1948

africanum (Mayr, 1866) (Ghana)

• Rhoptromyrmex tessmanni Forel, 1910

• Tetramorium lamottei Bernard, 1953

agile Arnold, 1960 (Zimbabwe)

agna (Santschi, 1935) (Democratic Republic of Congo)

aherni Hita Garcia & Fisher, 2012 (Madagascar)

akengense (Wheeler, W.M., 1922) (Democratic Republic of Congo)

akermani Arnold, 1926 (South Africa)

• Tetramorium akermani myersi Arnold, 1958

ala Hita Garcia & Fisher, 2012 (Madagascar)

alpestre Steiner, Schlick-Steiner & Seifert, 2010 (Austria)

alternans Santschi, 1929 (Morocco)

• Tetramorium biskrensis kahenae Menozzi, 1934

altivagans Santschi, 1914 (Kenya)

• Tetramorium simillimum isis Weber, 1943

amalae Sharaf & Aldawood, 2012 (Saudi Arabia)

amatongae Bolton, 1980 (Mozambique)

amaurum Bolton, 1980 (Zimbabwe)

ambanizana Hita Garcia & Fisher, 2012 (Madagascar)

ambatovy Hita Garcia & Fisher, 2012 (Madagascar)

amentete Bolton, 1980 (Ghana)

amissum Bolton, 1980 (Democratic Republic of Congo)

amium Forel, 1912 (Taiwan)

anatolicum Csösz & Schulz, 2007 (Turkey)

andohahela Hita Garcia & Fisher, 2012 (Madagascar)

andrei Forel, 1892 (Madagascar)

• Tetramorium andrei robustior Forel, 1892

andrynicum Bolton, 1977 (Australia)

angulinode Santschi, 1910 (Congo)

• Tetramorium humerosum Bernard, 1953

• Tetramorium nullispinum Bolton, 1980

• Xiphomyrmex angulinode daphnis Santschi, 1920

• Xiphomyrmex papyri Weber, 1943

ankarana Hita Garcia & Fisher, 2012 (Madagascar)

annectens Pisarski, 1969 (China)

anodontion Bolton, 1979 (Madagascar)

antennatum (Mann, 1919) (Solomon Is)

antipodum Wheeler, W.M., 1927 (Australia)

antrema (Bolton, 1976) (Cameroun)

anxium Santschi, 1914 (Guinea)

aptum Bolton, 1977 (Thailand)

argenteopilosum Arnold, 1926 (Zimbabwe)

argentirubrum Dietrich, 2004 (Jordan)

armatum Santschi, 1927 (Transcaspia)

• Tetramorium caespitum annauensis Karavaiev, 1931

• Tetramorium caespitum brevinodis Karavaiev, 1927

arnoldi (Forel, 1913) (Zimbabwe)

artemis Hita Garcia & Fisher, 2012 (Madagascar)

arzi Tohmé, G., 1969 (Lebanon)

asetyum Bolton, 1980 (Nigeria)

aspersum (Smith, F., 1865) (Indonesia (Morotai Island))

• Tetramorium bismarckii Forel, 1901

• Xiphomyrmex costatus willowsi Wheeler, W.M., 1934

ataxium Bolton, 1980 (Nigeria)

avaratra Hita Garcia & Fisher, 2012 (Madagascar)

avium Bolton, 1980 (South Africa)

azcatltontlium Marques, Vásquez-Bolaños & Quesada, 2011 (Mexico)

B
barbigerum Bolton, 1980 (Namibia)

barryi Mathew, 1981 (India)

basum Bolton, 1977 (New Guinea)

baufra (Bolton, 1976) (Zimbabwe)

beesoni (Mukerjee, 1934) (India)

belgaense Forel, 1902 (India)

bellicosum Bolton, 1980 (Nigeria)

bendai Hita Garcia, Fischer & Peters, 2010 (Burundi)

bequaerti Forel, 1913 (Democratic Republic of Congo)

• Tetramorium humile Santschi, 1914

berbiculum Bolton, 1980 (Zimbabwe)

bessonii Forel, 1891 (Madagascar)

bevisi Arnold, 1958 (Lesotho)

bicarinatum (Nylander, 1846) (United States)

• Myrmica cariniceps Guérin-Méneville, 1852

• Myrmica kollari Mayr, 1853

• Myrmica modesta Smith, F., 1860

• Myrmica reticulata Smith, F., 1862

bicolor Viehmeyer, 1914 (New Guinea)

bicolorum Vásquez-Bolaños, 2007 (Mexico)

binodis (Linnaeus, 1763) (Jamaica)

biskrense Forel, 1904 (Algeria)

• Tetramorium biskrense jarbas Cagniant, 1970

• Tetramorium caespitum oxyomma Karavaiev, 1912

boehmei Hita Garcia & Fisher, 2010 (Kenya)

boltoni Hita Garcia, Fischer & Peters, 2010 (Kenya)

bonibony Hita Garcia & Fisher, 2012 (Madagascar)

bothae Forel, 1910 (South Africa)

• Tetramorium guillarmodi Arnold, 1960

brevicorne Bondroit, 1918 (France (Corsica))

brevidentatum (Kutter, 1932) (Indonesia (Java))

brevispinosum (Stitz, 1910) (Cameroun)

• Triglyphothrix nion Bernard, 1953

browni Bolton, 1980 (Ghana)

bulawayense Forel, 1913 (Zimbabwe)

• Tetramorium bequaerti bruni Santschi, 1917

buthrum Bolton, 1980 (Central African Republic)

C
caespitum (Linnaeus, 1758) (Europe)

• Formica fusca Leach, 1825

• Myrmica brevinodis transversinodis Enzmann, J., 1946

• Myrmica fuscula Nylander, 1846

• Myrmica modesta Foerster, 1850

• Tetramorium caespitum fusciclavum Consani & Zangheri, 1952

• Tetramorium caespitum hammi Donisthorpe, 1915

• Tetramorium caespitum himalayanum Viehmeyer, 1914

• Tetramorium caespitum immigrans Santschi, 1927

• Tetramorium caespitum indocile Santschi, 1927

• Tetramorium jiangxiense Wang & Xiao, 1988

• Tetramorium semilaeve transbaicalense Ruzsky, 1936

caespitum barabense Ruzsky, 1925 (Russia)

caespitum caespitomoravicum Kratochvíl, 1941 (Czechoslovakia)

caespitum flavidulum Emery, 1924 (Turkey)

caespitum japonicum Röszler, 1936 (Japan)

caespitum pallidum Stitz, 1934 (China)

caespitum typicum Ruzsky, 1902 (Kazakhstan)

caldarium (Roger, 1857) (Poland)

• Tetramorium minutum Donisthorpe, 1942

• Tetramorium pauper transformans Santschi, 1914

• Tetramorium pusillum hemisi Wheeler, W.M., 1922

calidum Forel, 1907 (Oman)

calinum Bolton, 1980 (Ghana)

camerunense Mayr, 1895 (Cameroun)

candidum Bolton, 1980 (Democratic Republic of Congo)

capense Mayr, 1865 (South Africa)

• Tetramorium braunsi Forel, 1913

• Tetramorium popovici Forel, 1914

capillosum Bolton, 1980 (Gabon)

capitale (McAreavey, 1949) (Australia)

cardiocarenum Xu & Zheng, 1994 (China)

carinatum (Smith, F., 1859) (Indonesia (Aru Island))

• Xiphomyrmex aruensis Karavaiev, 1935

• Xiphomyrmex costatus Emery, 1897

• Xiphomyrmex costatus deficiens Emery, 1897

• Xiphomyrmex costatus flavescens Emery, 1897

centum Bolton, 1977 (New Guinea)

chapmani Bolton, 1977 (Philippines)

chefketi Forel, 1911 (Turkey)

• Tetramorium caespitum sarkissiani Forel, 1911

• Tetramorium caespitum turcomanicum Santschi, 1921

• Tetramorium taurocaucasicum Arnol'di, 1968

chepocha (Bolton, 1976) (Borneo)

chloe (Santschi, 1920) (Zimbabwe)

christiei Forel, 1902 (India)

ciliatum Bolton, 1977 (Thailand)

clunum Forel, 1913 (South Africa)

cognatum Bolton, 1979 (Madagascar)

coillum Bolton, 1979 (Madagascar)

coloreum Mayr, 1901 (Cameroun)

• Tetramorium humerosum muscicola Bernard, 1953

concaviceps Bursakov, 1984 (Kazakhstan)

confusum Bolton, 1977 (Australia)

constanciae Arnold, 1917 (Zimbabwe)

• Triglyphothrix longispinosa Arnold, 1956

convexum Bolton, 1980 (Ghana)

coonoorense Forel, 1902 (India)

cordatum Sheela & Narendran, 1998 (India)

crepum Wang & Wu, 1988 (China)

cristatum Stitz, 1910 (Togo)

• Tetramorium guineense medje Wheeler, W.M., 1922

crypticum (Bolton, 1976) (Ghana)

cuneinode Bolton, 1977 (Thailand)

curtulum Emery, 1895 (Myanmar)

curvispinosum Mayr, 1897 (Sri Lanka)

cyclolobium Xu & Zheng, 1994 (China)

cynicum Bolton, 1977 (Philippines)

D
davidi Forel, 1911 (Israel)

decamerum (Forel, 1902) (India)

deceptum Bolton, 1977 (Australia)

dedefra (Bolton, 1976) (Angola)

degener Santschi, 1911 (Madagascar)

delagoense Forel, 1894 (Mozambique)

• Tetramorium simillimum madecassum Forel, 1895

densopilosum Radchenko & Arakelian, 1990 (Armenia)

depressiceps Menozzi, 1933 (Israel)

depressum Forel, 1892 (Spain (Canary Is))

• Tetramorium semilaeve guancha Santschi, 1929

desertorum (Forel, 1910) (Botswana)

dichroum Santschi, 1932 (South Africa)

difficile Bolton, 1977 (Nepal)

diligens (Smith, F., 1865) (New Guinea)

• Pheidole diligentissima Dalla Torre, 1892

diomandei Bolton, 1980 (Ivory Coast)

diomedeum Emery, 1908 (Italy)

• Tetramorium caespitum bariensis Forel, 1911

• Tetramorium caespitum laevior Menozzi, 1936

distinctum (Bolton, 1976) (Ghana)

do Forel, 1914 (Zimbabwe)

dogieli Karavaiev, 1931 (Kenya)

dolichosum Bolton, 1980 (Democratic Republic of Congo)

dominum Bolton, 1980 (South Africa)

doriae Emery, 1881 (Ethiopia)

dumezi Menozzi, 1942 (Democratic Republic of Congo)

• Tetramorium dumezi Bolton, 1980

dunhuangense Chang & He, 2001 (China)

dysalum Bolton, 1979 (Madagascar)

dysderke Bolton, 1980 (Nigeria)

E
edouardi Forel, 1894 (Ethiopia)

• Tetramorium kivuense Stitz, 1911

• Tetramorium tersum Santschi, 1911

• Xiphomyrmex kivuense atrinodis Santschi, 1928

eleates Forel, 1913 (Indonesia (Sumatra))

electrum Bolton, 1979 (Madagascar)

elf Hita Garcia & Fisher, 2012 (Madagascar)

elidisum Bolton, 1980 (Ghana)

elisabethae Forel, 1904 (India)

emeryi Mayr, 1901 (South Africa)

• Tetramorium emeryi cristulatum Forel, 1913

eminii (Forel, 1894) (Ethiopia)

• Tetramorium marleyi akermani Arnold, 1926

• Triglyphothrix cinereus Weber, 1943

• Triglyphothrix guillodi Santschi, 1937

• Triglyphothrix guillodi mus Santschi, 1937

• Triglyphothrix marleyi Forel, 1914

• Triglyphothrix marthae Forel, 1911

erectum Emery, 1895 (South Africa)

• Tetramorium bacchus Forel, 1910

ericae Arnold, 1917 (Zimbabwe)

• Triglyphothrix pauper Santschi, 1917

etiolatum Bolton, 1977 (New Guinea)

exasperatum Emery, 1891 (Tunisia)

• Tetramorium zahrae Santschi, 1923

exasperatum acutiseta Santschi, 1921 (Morocco)

exile Csösz & Radchenko, 2007 (Afghanistan)

F
fergusoni Forel, 1902 (India)

ferox Ruzsky, 1903 (Russian Federation)

• Tetramorium confinis Radchenko & Arakelian, 1990

• Tetramorium ferox perspicax Radchenko, 1992

• Tetramorium ferox silhavyi Kratochvíl, 1941

feroxoides Dlussky & Zabelin, 1985 (Kazakhstan)

• Tetramorium bursakovi Radchenko, 1992

fezzanense Bernard, 1948 (Algeria)

flabellum Bolton, 1980 (Ghana)

flagellatum Bolton, 1977 (Borneo)

flaviceps Arnold, 1960 (Zimbabwe)

• Tetramorium do mus Arnold, 1960

flavipes Emery, 1893 (Thailand)

flavithorax (Santschi, 1914) (Ghana)

flavum Chang & He, 2001 (China)

forte Forel, 1904 (France)

• Tetramorium caespitum hispanicum Bondroit, 1918

• Tetramorium caespitum ruginode Stitz, 1917

• Tetramorium ferox marocanum De Haro & Collingwood, 1994

• Tetramorium maura tingitanum Santschi, 1929

• Tetramorium ruginode marocana Cagniant, 1997

frenchi Forel, 1914 (South Africa)

frigidum Arnold, 1926 (South Africa)

• Tetramorium akermani drakensbergensis Arnold, 1926

fulviceps (Emery, 1897) (New Guinea)

furtivum (Arnold, 1956) (Zimbabwe)

fuscipes (Viehmeyer, 1925) (Australia)

G
gabonense (André, 1892) (Gabon)

• Triglyphothrix areolata burgeoni Santschi, 1935

• Triglyphothrix areolatus Stitz, 1910

• Triglyphothrix gabonensis boulognei Forel, 1916

• Triglyphothrix gabonensis kamerunensis Santschi, 1937

• Triglyphothrix gabonensis soyauxi Forel, 1901

• Triglyphothrix mucidus Forel, 1909

galoasanum Santschi, 1910 (Congo)

gambogecum (Donisthorpe, 1941) (New Guinea)

• Xiphomyrmex gambogecus flavus Donisthorpe, 1941

gazense Arnold, 1958 (Zimbabwe)

gegaimi Forel, 1916 (Democratic Republic of Congo)

geminatum Bolton, 1980 (Gabon)

gestroi (Menozzi, 1933) (Uganda)

ghindanum Forel, 1910 (Ethiopia)

glabratum Stitz, 1923 (Namibia)

• Tetramorium rutilum Prins, 1973

gladstonei Forel, 1913 (Zimbabwe)

goniommoide Poldi, 1979 (Turkey)

gracile Forel, 1894 (Ethiopia)

grandinode Santschi, 1913 (South Africa)

• Tetramorium grandinode hopensis Forel, 1914

granulatum Bolton, 1980 (Nigeria)

grassii Emery, 1895 (South Africa)

• Tetramorium grassii laevigatum Mayr, 1901

• Tetramorium grassii simulans Santschi, 1914

• Tetramorium grazsii mayri Emery, 1924

• Tetramorium joffrei Forel, 1914

• Tetramorium joffrei algoa Arnold, 1917

guangxiense Zhou & Zheng, 1997 (China)

guineense (Bernard, 1953) (Guinea)

H
hapale Bolton, 1980 (Ivory Coast)

hasinae Yamane & Jaitrong, 2011 (Thailand)

hector Hita Garcia & Fisher, 2012 (Madagascar)

hippocratis Agosti & Collingwood, 1987 (Turkey)

hirsutum Collingwood & van Harten, 2005 (Yemen)

hispidum (Wheeler, W.M., 1915) (United States)

hortorum Arnold, 1958 (Zimbabwe)

humbloti Forel, 1891 (Comoros)

• Tetramorium humbloti pembensis Forel, 1907

• Tetramorium humblotii victoriensis Forel, 1913

hungaricum Röszler, 1935 (Hungary)

I
ibycterum Bolton, 1979 (Madagascar)

ictidum Bolton, 1980 (Cameroun)

imbelle (Emery, 1915) (Ethiopia)

impressum (Viehmeyer, 1925) (Australia)

impurum (Foerster, 1850) (Germany)

• Tetramorium caespitum penninum Santschi, 1927

• Tetramorium caespitum staerckei Kratochvíl, 1944

impurum gregori Kratochvíl, 1944 (Czechoslovakia)

incruentatum Arnold, 1926

• Rhoptromyrmex arnoldi Santschi, 1916

indicum Forel, 1913 (Indonesia (Sumatra))

indosinense Wheeler, W.M., 1927 (Vietnam)

inerme Mayr, 1877 (Kazakhstan)

• Tetramorium caespitum picta Karavaiev, 1912

• Tetramorium caespitum plana Karavaiev, 1927

• Tetramorium caespitum reticuliventre Ruzsky, 1902

inezulae (Forel, 1914) (South Africa)

• Tetramorium hepburni Arnold, 1917

• Triglyphothrix hepburni mashonana Arnold, 1949

infraspinosum Karavaiev, 1935 (Vietnam)

infraspinum Forel, 1905 (Indonesia (Java))

inglebyi Forel, 1902 (India)

insolens (Smith, F., 1861) (Indonesia (Sulawesi))

• Tetramorium guineense macra Emery, 1914

• Tetramorium melanogyna pallidiventre Wheeler, W.M., 1934

• Tetramorium pacificum wilsoni Mann, 1921

intermedium Hita Garcia, Fischer & Peters, 2010 (Ghana)

intextum Santschi, 1914 (Kenya)

• Tetramorium intextum cataractae Santschi, 1916

intonsum Bolton, 1980 (Ghana)

invictum Bolton, 1980 (Ivory Coast)

isectum Bolton, 1979 (Madagascar)

isipingense Forel, 1914 (South Africa)

isoelectrum Hita Garcia & Fisher, 2012 (Madagascar)

J
jauresi Forel, 1914 (South Africa)

• Tetramorium latens Arnold, 1948

jedi Hita Garcia & Fisher, 2012 (Madagascar)

jejunum Arnold, 1926 (Zimbabwe)

jizane Collingwood, 1985 (Saudi Arabia)

jordani Santschi, 1937 (Namibia)

• Tetramorium aspinatum Prins, 1973

juba Collingwood, 1985 (Algeria)

jugatum Bolton, 1980 (Ivory Coast)

K
kabulistanicum Pisarski, 1967 (Afghanistan)

kakamega Hita Garcia, Fischer & Peters, 2010 (Kenya)

kali Hita Garcia & Fisher, 2012 (Madagascar)

katypum (Bolton, 1976) (Philippines)

kelleri Forel, 1887 (Madagascar)

keralense Sheela & Narendran, 1998 (India)

kestrum Bolton, 1980 (Uganda)

kheperra (Bolton, 1976) (Indonesia (Java))

khnum Bolton, 1977 (Philippines)

khyarum Bolton, 1980 (Nigeria)

kieti Roncin, 2002 (Vietnam)

kisilkumense Dlussky, 1990 (Turkmenistan)

kraepelini Forel, 1905 (Indonesia (Java))

• Tetramorium tanakai Bolton, 1977

• Tetramorium yanoi Santschi, 1937

krynitum Bolton, 1980 (Namibia)

kydelphon Bolton, 1979 (New Guinea)

L
laevithorax Emery, 1895 (South Africa)

• Tetramorium jeanae Weber, 1943

lanuginosum Mayr, 1870 (Indonesia)

• Tetramorium obesum striatidens Emery, 1889

• Triglyphothrix ceramensis Stitz, 1912

• Triglyphothrix mauricei Donisthorpe, 1946

• Triglyphothrix striatidens australis Forel, 1902

• Triglyphothrix striatidens felix Forel, 1912

• Triglyphothrix striatidens flavescens Wheeler, W.M., 1929

• Triglyphothrix striatidens laevidens Forel, 1900

• Triglyphothrix striatidens orissana Forel, 1902

• Triglyphothrix tricolor Donisthorpe, 1948

laparum Bolton, 1977 (Philippines)

laticephalum Bolton, 1977 (Australia)

latinode Collingwood & Agosti, 1996 (Yemen)

latreillei Forel, 1895 (Madagascar)

legone Bolton, 1980 (Ghana)

lobulicorne Santschi, 1916 (Zimbabwe)

longicorne Forel, 1907 (Kenya)

longoi Forel, 1915 (South Africa)

lucayanum Wheeler, W.M., 1905 (Bahamas)

• Tetramorium camerunense waelbroeki Forel, 1909

• Tetramorium lucayanum sexdens Forel, 1915

• Tetramorium rectinodis Menozzi, 1942

lucidulum Menozzi, 1933 (Syria)

lucyae Sorger, 2011 (Borneo, Sarawak, Malaysia)

luteipes Santschi, 1910 (Congo)

luteolum Arnold, 1926 (Zimbabwe)

M
mackae Media:Hita Garcia & Fisher 2012a.pdf|Hita Garcia & Fisher, 2012 (Madagascar)

magnificum Bolton, 1980 (Ivory Coast)

mahafaly Hita Garcia & Fisher, 2011 (Madagascar)

mai Wang, M., 1993 (China)

malabarense Sheela & Narendran, 1998 (India)

malagasy Hita Garcia & Fisher, 2012 (Madagascar)

mallenseana Hita Garcia & Fisher, 2012 (Madagascar)

manni Bolton, 1985

• Tetramorium manni Dlussky, 1994

• Triglyphothrix pacifica Mann, 1921

manobo (Calilung, 2000) (Philippines (Mindanao))

marginatum Forel, 1895 (Madagascar)

marojejy Hita Garcia & Fisher, 2012 (Madagascar)

matopoense Arnold, 1926 (Zimbabwe)

maurum Santschi, 1918 (Tunisia)

mayri (Mann, 1919) (Solomon Is)

megalops Bolton, 1977 (Australia)

meghalayense Bharti, 2011 (India)

• Tetramorium browni Tiwari, 2000

melanogyna Mann, 1919 (Solomon Is)

menkaura (Bolton, 1976) (Ghana)

meressei Forel, 1916 (Democratic Republic of Congo)

meridionale Emery, 1870 (Italy)

meshena (Bolton, 1976) (West Malaysia)

metactum Bolton, 1980 (Kenya)

mexicanum Bolton, 1979 (Mexico)

microgyna Santschi, 1918 (South Africa)

microps (Mayr, 1901) (South Africa)

• Tetramorium auropunctatus bulawayensis Arnold, 1917

• Triglyphothrix auropunctata rhodesiana Forel, 1913

• Triglyphothrix auropunctatus Forel, 1910

• Triglyphothrix auropunctatus fusciventris Forel, 1913

• Triglyphothrix auropunctatus pallens Forel, 1910

minimum (Bolton, 1976) (Ghana)

minusculum (Santschi, 1914) (Cameroun)

• Xiphomyrmex minusculus amen Weber, 1943

miserabile Santschi, 1918 (Kenya)

mixtum Forel, 1902 (India)

mkomazi Hita Garcia, Fischer & Peters, 2010 (Tanzania)

monardi (Santschi, 1937) (Angola)

moravicum Kratochvíl, 1941 (Czechoslovakia)

• Tetramorium rhenanum Schulz, 1996

mossamedense Forel, 1901 (Angola)

• Tetramorium caespitum schultzei Forel, 1910

muralti Forel, 1910 (Cameroun)

• Xiphomyrmex muralti trilineata Santschi, 1919

muscorum Arnold, 1926 (Mozambique)

mutatum Bolton, 1985

• Triglyphothrix pulchella Mann, 1919

myops Bolton, 1977 (India)

N
nacta (Bolton, 1976) (Thailand)

naganum Bolton, 1979 (Madagascar)

nautarum Santschi, 1918 (Annobon I)

navum Bolton, 1977 (New Guinea)

nazgul Hita Garcia & Fisher, 2012 (Madagascar)

nefassitense Forel, 1910 (Ethiopia)

nify Hita Garcia & Fisher, 2012 (Madagascar)

nigrum Forel, 1907 (Kenya)

• Tetramorium brevis Weber, 1943

nipponense Wheeler, W.M., 1928 (Japan)

nitidissimum Pisarski, 1967 (Transcaspia)

nodiferum (Emery, 1901) (Cameroun)

noeli Hita Garcia & Fisher, 2012 (Madagascar)

noratum Bolton, 1977 (West Malaysia)

norvigi Hita Garcia & Fisher, 2012 (Madagascar)

nosybe Hita Garcia & Fisher, 2012 (Madagascar)

notiale Bolton, 1980 (Zimbabwe)

• Tetramorium guineense striatum Arnold, 1917

notomelanum Vásquez-Bolaños, Castaño-Meneses & Guzmán-Mendoza, 2011 (Mexico)

nube Weber, 1943 (Sudan)

nursei Bingham, 1903 (Pakistan)

O
obesum André, 1887 (India)

obtusidens Viehmeyer, 1916 (Singapore)

occidentale (Santschi, 1916) (Cameroon)

• Xiphomyrmex insularis Menozzi, 1924

ochrothorax Chang & He, 2001 (China)

ocothrum Bolton, 1979 (Borneo)

oculatum Forel, 1913 (Zimbabwe)

olana Hita Garcia & Fisher, 2012 (Madagascar)

orc Hita Garcia & Fisher, 2012 (Madagascar)

orientale Forel, 1895 (Madagascar)

ornatum Emery, 1897 (Papua New Guinea)

• Tetramorium ornatum obscurius Forel, 1901

osiris (Bolton, 1976) (Ghana)

P
pacificum Mayr, 1870 (Tonga)

• Tetramorium pacificum subscabrum Emery, 1893

palaense Bolton, 1979 (Borneo)

parasiticum Bolton, 1980 (South Africa)

parvioculum Guillem & Bensusan, 2009 (Gibraltar)

parvispinum (Emery, 1893) (Borneo)

• Triglyphothrix parvispina formosae Forel, 1912

parvum Bolton, 1977 (Indonesia (Java))

pauper Forel, 1907 (Kenya)

pelagium Poldi, 1995 (Italy)

peringueyi Arnold, 1926 (South Africa)

perlongum Santschi, 1923 (Angola)

persignatum Bolton, 1995

• Tetramorium signatum Menozzi Menozzi, 1933

petersi Forel, 1910 (Namibia)

petiolatum Sheela & Narendran, 1998 (India)

peutli Forel, 1916 (Democratic Republic of Congo)

phasias Forel, 1914 (South Africa)

• Tetramorium guineense hertigi Santschi, 1937

philippwagneri Hita Garcia, Fischer & Peters, 2010 (Zambia)

pialtum Bolton, 1980 (Ghana)

pilosum Emery, 1893 (Sri Lanka)

pinnipilum Bolton, 1980 (Angola)

placidum Bolton, 1979 (Mexico)

platynode Bolton, 1980 (South Africa)

pleganon Bolton, 1979 (Madagascar)

plesiarum Bolton, 1979 (Madagascar)

plumosum Bolton, 1980 (Swaziland)

pnyxis (Bolton, 1976) (West Malaysia)

pogonion Bolton, 1980 (Namibia)

politum Emery, 1897 (New Guinea)

polymorphum Yamane & Jaitrong, 2011 (Laos)

popell Hita Garcia & Fisher, 2012 (Madagascar)

postpetiolatum Santschi, 1919 (Democratic Republic of Congo)

poweri Forel, 1914 (South Africa)

praetextum Bolton, 1980 (Namibia)

proximum Bolton, 1979 (Madagascar)

psymanum Bolton, 1980 (Ghana)

pulchellum Emery, 1897 (New Guinea)

pulcherrimum (Donisthorpe, 1945) (Uganda)

pullulum Santschi, 1924 (Democratic Republic of Congo)

• Tetramorium fernandensis Menozzi, 1942

• Xiphomyrmex uelensis Santschi, 1935

punctatum Santschi, 1927 (Italy (Sicily))

• Tetramorium semilaeve liparaeum Santschi, 1927

punctiventre Emery, 1887 (New Guinea)

punicum (Smith, F., 1861) (Israel)

punicum cyrenaicum Menozzi, 1927 (Libya)

pusillum Emery, 1895 (South Africa)

• Tetramorium caespitum ladismithensis Forel, 1913

• Tetramorium pusillum tablensis Forel, 1914

pylacum Bolton, 1980 (Ivory Coast)

pyrenaeicum Röszler, 1937 (Andorra)

Q
quadridentatum Stitz, 1910 (Cameroun)

• Tetramorium commodum Santschi, 1924

qualarum Bolton, 1980 (Ghana)

quasirum Bolton, 1979 (Madagascar)

R
ranarum Forel, 1895 (Madagascar)

regulare Bolton, 1980 (South Africa)

rekhefe Bolton, 1979 (Indonesia (Timor Island))

renae Hita Garcia, Fischer & Peters, 2010 (São Tomé & Principe)

repentinum Arnold, 1926 (Zimbabwe)

repletum Wang & Xiao, 1988 (China)

reptana (Bolton, 1976) (Uganda)

reticuligerum Bursakov, 1984 (Kazakhstan)

rhetidum Bolton, 1980 (Ghana)

rhodium Emery, 1924 (Greece)

rigidum Bolton, 1977 (New Guinea)

rimytyum Bolton, 1980 (Ghana)

rinatum Bolton, 1977 (Philippines)

robertsoni Hita Garcia, Fischer & Peters, 2010 (Tanzania)

robitika Hita Garcia & Fisher, 2012 (Madagascar)

rogatum Bolton, 1980 (Angola)

rossi (Bolton, 1976) (India)

rothschildi (Forel, 1907) (Ethiopia)

rotundatum (Santschi, 1924) (Democratic Republic of Congo)

rubrum Hita Garcia, Fischer & Peters, 2010 (Cameroun)

rufescens Stitz, 1923 (Namibia)

rugigaster Bolton, 1977 (India)

ryanphelanae Hita Garcia & Fisher, 2012 (Madagascar)

S
sabatra Hita Garcia & Fisher, 2012 (Madagascar)

sada Hita Garcia & Fisher, 2012 (Madagascar)

saginatum Bolton, 1980 (Angola)

sahlbergi Finzi, 1936 (Egypt)

salomo Mann, 1919 (Solomon Is)

salvatum Forel, 1902 (India)

salwae Mohamed, Zalat, Fadl, Gadalla & Sharaf, 2001 (Egypt)

sanetrai Schulz & Csösz, 2007 (Italy)

sargina Hita Garcia & Fisher, 2012 (Madagascar)

scabrosum (Smith, F., 1859) (Indonesia (Aru Island))

• Tetramorium papuanum Emery, 1887

scabrum Mayr, 1879 (Borneo)

schaufussii Forel, 1891 (Madagascar)

• Tetramorium nassonowii Forel, 1892

schmidti Forel, 1904 (Israel)

schneideri Emery, 1898 (Kazakhstan)

schoutedeni Santschi, 1924 (Democratic Republic of Congo)

sculptatum Bolton, 1977 (New Guinea)

scytalum Bolton, 1979 (Madagascar)

securis Roncin, 2002 (Vietnam)

semilaeve André, 1883 (Mediterranean region)

• Tetramorium banyulense Bernard, 1983

• Tetramorium caespitum tenuicornis Emery, 1925

• Tetramorium semilaeve siciliense Santschi, 1927

semilaeve atlante Cagniant, 1970 (Tunisia)

semilaeve gaetulum Santschi, 1936 (Morocco)

semilaeve galaticum Menozzi, 1936 (Turkey)

semilaeve hoggarense Santschi, 1929 (Algeria)

semilaeve italicum Menozzi, 1932 (Italy)

semilaeve judas Wheeler, W.M. & Mann, 1916 (Israel)

semilaeve jugurtha Menozzi, 1934 (Tunisia)

semilaeve kutteri Santschi, 1927 (Switzerland)

semireticulatum Arnold, 1917 (Zimbabwe)

• Tetramorium semireticulatum politum Arnold, 1948

seneb Bolton, 1977 (West Malaysia)

sentosum Sheela & Narendran, 1998 (India)

sepositum Santschi, 1918 (Zimbabwe)

sepultum Bolton, 1980 (Swaziland)

sericeiventre Emery, 1877 (Ethiopia)

• Atopula hortensis Bernard, 1948

• Tetramorium blochmannii Forel, 1887

• Tetramorium blochmanni calvum Stitz, 1923

• Tetramorium blochmanni continentis Forel, 1910

• Tetramorium blochmannii montanum Forel, 1891

• Tetramorium blochmanni nigriventre Stitz, 1910

• Tetramorium neuvillei Forel, 1907

• Tetramorium quadrispinosum Emery, 1886

• Tetramorium quadrispinosum angolense Santschi, 1930

• Tetramorium quadrispinosum beirae Arnold, 1926

• Tetramorium quadrispinosum elegans Santschi, 1918

• Tetramorium quadrispinosum eudoxia Santschi, 1918

• Tetramorium quadrispinosum otaviensis Arnold, 1926

• Tetramorium sericeiventre arenarium Santschi, 1918

• Tetramorium sericeiventre bipartita Santschi, 1918

• Tetramorium sericeiventre cinnamomeum Santschi, 1918

• Tetramorium sericeiventre debile Forel, 1894

• Tetramorium sericeiventre femoratum Emery, 1895

• Tetramorium sericeiventre gamaii Santschi, 1918

• Tetramorium sericeiventre hori Santschi, 1918

• Tetramorium sericeiventre inversa Santschi, 1910

• Tetramorium sericeiventre jasonis Santschi, 1918

• Tetramorium sericeiventre munda Santschi, 1918

• Tetramorium sericeiventre repertum Santschi, 1926

• Tetramorium sericeiventre vascoi Santschi, 1918

• Tetramorium sericeiventre vividum Santschi, 1926

sericeum Arnold, 1926 (Zimbabwe)

setigerum Mayr, 1901 (South Africa)

• Tetramorium setigerum anteversa Santschi, 1921

• Tetramorium setigerum quaerens Forel, 1914

setuliferum Emery, 1895 (South Africa)

• Tetramorium setuliferum cucalense Santschi, 1911

• Tetramorium setuliferum triptolemus Arnold, 1917

severini (Emery, 1895) (Madagascar)

shamshir Hita Garcia & Fisher, 2012 (Madagascar)

shensiense Bolton, 1977 (China)

shilohense Forel, 1913 (Zimbabwe)

shirlae Sharaf, 2007 (Saudi Arabia)

shivalikense Bharti & Kumar, 2012 (India)

sigillum Bolton, 1980 (Ivory Coast)

signatum Emery, 1895 (South Africa)

• Tetramorium solidum grootensis Forel, 1913

• Tetramorium solidum lugubre Forel, 1910

• Tetramorium solidum tuckeri Arnold, 1926

sikorae Forel, 1892 (Madagascar)

• Xiphomyrmex latior Santschi, 1926

silvicola Hita Garcia & Fisher, 2012 (Madagascar)

simillimum (Smith, F., 1851) (United Kingdom)

• Myrmica parallela Smith, F., 1859

• Tetramorium pusillum bantouana Santschi, 1910

• Tetramorium pusillum exoleta Santschi, 1914

• Tetramorium pygmaeum Emery, 1877

• Tetramorium simillimum denticulatum Forel, 1902

• Tetramorium simillimum insulare Santschi, 1928

• Tetramorium simillimum opacior Forel, 1913

• Wasmannia auropunctata brevispinosa Borgmeier, 1928

simulator Arnold, 1917 (Zimbabwe)

singletonae Hita Garcia & Fisher, 2012 (Madagascar)

sitefrum Bolton, 1980 (Ghana)

sjostedti Forel, 1915 (Australia)

smaug Hita Garcia & Fisher, 2012 (Madagascar)

smithi Mayr, 1879 (India)

• Tetramorium simillimum laevinode Forel, 1902

• Tetramorium smithi kanariense Forel, 1902

snellingi Hita Garcia, Fischer & Peters, 2010 (Kenya)

solidum Emery, 1886 (South Africa)

somniculosum Arnold, 1926 (Mozambique)

spininode Bolton, 1977 (Australia)

spinosum (Pergande, 1896) (Mexico)

• Tetramorium wheeleri Forel, 1901

• Xiphomyrmex spinosus insons Wheeler, W.M., 1915

splendens Ruzsky, 1902 (Caucasus)

splendidior (Viehmeyer, 1925) (Australia)

squaminode Santschi, 1911 (Tanzania)

steinheili Forel, 1892 (Madagascar)

striabdomen Chang & He, 2001 (China)

striativentre Mayr, 1877 (Kazakhstan)

strictum Bolton, 1977 (Australia)

striolatum Viehmeyer, 1914 (Australia)

subcoecum Forel, 1907 (Kenya)

• Tetramorium subcoecum inscia Forel, 1913

sudanense (Weber, 1943) (Sudan)

sulcinode Santschi, 1927 (Kazakhstan)

• Tetramorium karakalense Dlussky & Zabelin, 1985

surrogatum Bolton, 1985

• Triglyphothrix silvestrii Emery, 1915

susannae Hita Garcia, Fischer & Peters, 2010 (Ghana)

syriacum Emery, 1924 (Syria)

T
tabarum Bolton, 1980 (Democratic Republic of Congo)

talpa (Bolton, 1976) (Democratic Republic of Congo)

tanaense Hita Garcia, Fischer & Peters, 2010 (Kenya)

tantillum Bolton, 1979 (Madagascar)

taueret Bolton, 1995

• Tetramorium caespitum debilis Emery, 1924

taylori Bolton, 1985 (Australia)

• Tetramorium australe Bolton, 1977

tenebrosum Arnold, 1926 (Mozambique)

tenuicrine (Emery, 1914) (New Caledonia)

termitobium Emery, 1908 (Democratic Republic of Congo)

thalidum Bolton, 1977 (Australia)

thoth (Bolton, 1976) (South Africa)

titus Forel, 1910 (South Africa)

tonganum Mayr, 1870 (Tonga)

• Tetramorium magitae Forel, 1911

tortuosum Roger, 1863 (Sri Lanka)

• Tetramorium tortuosum bellii Forel, 1902

• Tetramorium tortuosum ethica Forel, 1911

tosii Emery, 1899 (Madagascar)

traegaordhi Santschi, 1914 (South Africa)

trafo Hita Garcia & Fisher, 2012 (Madagascar)

transversarium Roger, 1863 (Sri Lanka)

triangulatum Bharti & Kumar, 2012 (India)

tricarinatum Viehmeyer, 1914 (New Guinea)

• Xiphomyrmex tricolor Donisthorpe, 1949

trimeni (Emery, 1895) (South Africa)

trirugosum Hita Garcia, Fischer & Peters, 2010 (Cameroun)

tsingy Hita Garcia & Fisher, 2012 (Madagascar)

tsushimae Emery, 1925 (Japan)

• Tetramorium caespitum jacoti Wheeler, W.M., 1927

turneri Forel, 1902 (Australia)

tychadion Bolton, 1980 (Tanzania)

tylinum Bolton, 1977 (Borneo)

typhlops Bolton, 1980 (Ivory Coast)

tyrion Hita Garcia & Fisher, 2012 (Madagascar)

U
ubangense Santschi, 1937 (Democratic Republic of Congo)

umtaliense Arnold, 1926 (Zimbabwe)

undatium Chang & He, 2001 (China)

unicum Bolton, 1980 (Democratic Republic of Congo)

urbanii Bolton, 1977 (Bhutan)

V
validiusculum Emery, 1897 (Australia)

• Tetramorium longicarinum Donisthorpe, 1941

valky Hita Garcia & Fisher, 2012 (Madagascar)

vandalum Bolton, 1977 (New Guinea)

vernicosum Radchenko, 1992 (Tajikistan)

versiculum Bolton, 1980 (Ghana)

vertigum Bolton, 1977 (Indonesia (Sulawesi))

vexator Arnold, 1926 (South Africa)

viehmeyeri Forel, 1907 (Australia)

• Xiphomyrmex viehmeyeri venustus Wheeler, W.M., 1934

viticola Weber, 1943 (Sudan)

voasary Hita Garcia & Fisher, 2012 (Madagascar)

vohitra Hita Garcia & Fisher, 2012 (Madagascar)

vombis (Bolton, 1976) (Solomon Is)

vony Hita Garcia & Fisher, 2012 (Madagascar)

W
wadje Bolton, 1980 (Ghana)

wagneri Viehmeyer, 1914 (New Guinea)

walshi (Forel, 1890) (India)

• Triglyphothrix musculus Forel, 1902

• Triglyphothrix walshi spuria Forel, 1912

wardi Hita Garcia & Fisher, 2012 (Madagascar)

warreni Arnold, 1926 (South Africa)

weitzeckeri Emery, 1895 (South Africa)

• Tetramorium ebeninum Arnold, 1926

• Tetramorium escherichi Forel, 1910

• Xiphomyrmex weitzeckeri edithae Weber, 1943

• Xiphomyrmex weitzeckeri nigellus Santschi, 1932

X
xanthogaster Santschi, 1911 (Madagascar)

xuthum Bolton, 1980 (Ghana)

Y
yammer Hita Garcia & Fisher, 2012 (Madagascar)

yarthiellum (Bolton, 1976) (Zambia)

yemene Collingwood & Agosti, 1996 (Yemen)

yerburyi Forel, 1902 (Sri Lanka)

youngi Bolton, 1980 (Angola)

yulongense Xu & Zheng, 1994 (China)

Z
zambezium Santschi, 1939 (Zimbabwe)

zapyrum Bolton, 1980 (Ghana)

zenatum Bolton, 1979 (Madagascar)

zonacaciae (Weber, 1943) (Sudan)

zypidum Bolton, 1977 (Philippines)