Checklist of Messor species

The following species and subspecies belong to the genus Messor. Synonyms are listed under their senior names. For valid names only see Messor species and for distribution information see Messor species by Country.

A
abdelazizi Santschi, 1921 (Morocco)

aciculatus (Smith, F., 1874) (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation)

• Messor barbarus lobulifera Emery, 1901

• Stenamma aciculatum brunneicorne Forel, 1901

aciculatus risianus Forel, 1922 (China)

aegyptiacus (Emery, 1878) (Egypt, Israel, Morocco)

aegyptiacus canaliculatus Donisthorpe, 1944 (Algeria)

aegyptiacus felah Santschi, 1923 (Egypt, Israel)

aegyptiacus phasianicus Bernard, 1953 (Algeria)

aegyptiacus tunetinus Santschi, 1923 (Algeria, Tunisia)

alexandri Tohmé, G. & Tohmé, H., 1981 (Greece, Iran, Israel, Syrian Arab Republic)

angularis Santschi, 1928 (Kenya)

antennatus Emery, 1908 (Algeria, Morocco)

antennatus fodorii Röszler, 1942

antennatus personatus Santschi, 1931 (Morocco)

aphaenogasteroides Pisarski, 1967 (Afghanistan)

aralocaspius (Ruzsky, 1902) (China, Iran, Iraq, Kuwait, Kyrgyzstan, Montenegro, Republic of Macedonia, Russian Federation, Saudi Arabia, Turkmenistan)

aralocaspius infumatus Kuznetsov-Ugamsky, 1929 (China, Kazakhstan, Kyrgyzstan)

arenarius (Fabricius, 1787) (Algeria, Israel, Kuwait, Saudi Arabia, Tunisia)

• Messor barbarus beduinus Emery, 1922

• Myrmica amaurocyclia Foerster, 1850

• Myrmica scalpturata Nylander, 1856

• Stenamma bugnioni Forel, 1904

arenarius alfierii Santschi, 1938 (Egypt)

arenarius diabolus Santschi, 1938 (Egypt)

arenarius ratus Menozzi, 1933 (Israel)

arenarius thebaicus Santschi, 1912 (Egypt)

atanassovii Atanassov, 1982 (Bulgaria)

B
barbarus (Linnaeus, 1767) (Algeria, Balearic Islands, Bulgaria, Egypt, France, Gibraltar, Iberian Peninsula, Monaco, Morocco, Portugal, Romania, Spain, Tunisia)

• Aphaenogaster barbara sordida Forel, 1892

• Formica binodis Fabricius, 1775

• Formica juvenilis Fabricius, 1804

• Formica megacephala Leach, 1825

• Messor barbarus ambiguus Santschi, 1925

• Messor barbarus nigriceps Santschi, 1925

• Myrmica rufitarsis Foerster, 1850

barbarus gallaoides Santschi, 1929 (Morocco)

• Messor barbarus nigricans Santschi, 1929

barbarus mediosanguineus Donisthorpe, 1946 (Turkey)

barbarus politus Karavaiev, 1912 (Algeria)

• Messor barbarus dentiscapus Finzi, 1930

barbarus sahlbergi Forel, 1913 (Egypt)

barbarus santschii Emery, 1908 (Tunisia)

• Messor carthaginensis Bernard, 1980

berbericus Bernard, 1955 (Algeria)

bernardi Cagniant, 1967 (Algeria)

bouvieri Bondroit, 1918 (Balearic Islands, France, Gibraltar, Iberian Peninsula, Malta, Portugal, Spain)

boyeri Cagniant, 2006 (Morocco)

buettikeri Collingwood, 1985 (Iraq, Kuwait, Saudi Arabia)

C
caducus (Victor, 1839) (Albania, Armenia, Bulgaria, Georgia, Greece, Iran, Malta, Republic of Macedonia, Romania, Russian Federation, Slovakia, Turkey)

• Messor caducus caucasicola Arnoldi, 1977

caducus glabrescens Arnoldi, 1977 (Russian Federation)

capensis (Mayr, 1862) (Botswana, Namibia, South Africa, Zimbabwe)

• Aphaenogaster pseudoaegyptiaca Emery, 1884

• Messor barbarus schencki Wheeler, W.M., 1922

• Messor braunsi Forel, 1913

• Messor donisthorpei Santschi, 1937

capitatus (Latreille, 1798) (Balearic Islands, Bulgaria, Croatia, France, Greece, Iberian Peninsula, Italy, Malta, Monaco, Montenegro, Portugal, Republic of Macedonia, Romania, Slovenia, Spain)

• Aphaenogaster barbara nigra André, 1883

• Formica huberiana Leach, 1825

• Myrmica galbula Losana, 1834

caviceps (Forel, 1902) (Algeria, Morocco)

• Messor planiceps Stitz, 1917

celiae Reyes, 1985 (Iberian Peninsula, Spain)

cephalotes (Emery, 1895) (Ethiopia, Kenya, United Republic of Tanzania)

• Messor plinii Santschi, 1912

ceresis Santschi, 1934 (Iran, Lebanon, Saudi Arabia)

clivorum Ruzsky, 1905 (Republic of Moldova, Russian Federation)

clivorum sevani Arnoldi, 1977 (Armenia)

clypeatus Kuznetsov-Ugamsky, 1927 (Kazakhstan)

collingwoodi Bolton, 1982 (Mali, Niger)

crawleyi Santschi, 1928 (Egypt, Saudi Arabia)

D
decipiens Santschi, 1917 (Botswana, Iraq, Lesotho, Saudi Arabia, South Africa, Zimbabwe)

• Messor arcistriatus Santschi, 1928

• Messor barbarus probus Wheeler, W.M., 1922

dentatus Santschi, 1927 (Iran, Israel, Turkey)

denticornis Forel, 1910 (Botswana, Namibia, South Africa)

• Messor denticornis brunni Forel, 1910

• Messor denticornis parvidens Forel, 1910

desertora He & Song, 2009 (China)

diabarensis Arnoldi, 1970 (Kazakhstan)

E
ebeninus Santschi, 1927 (Egypt, Iran, Iraq, Israel, Kuwait, Lebanon, Montenegro, Oman, Saudi Arabia, Turkey, United Arab Emirates)

eglalae Sharaf, 2007 (Egypt)

erectus Espadaler, 1998 (Morocco)

excursionis Ruzsky, 1905 (China, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Turkmenistan)

excursionis alaschanicus Ruzsky, 1915 (Mongolia)

excursionis fraternus Ruzsky, 1905 (China)

excursionis sillemi Stärcke, 1935 (Kazakhstan)

F
ferreri Collingwood, 1993 (Kenya)

foreli Santschi, 1923 (Algeria, Egypt, Morocco, Oman, Saudi Arabia, Tunisia, United Arab Emirates)

• Messor aegyptiacus fossulatus Santschi, 1923

foreli brevispinosus Santschi, 1923 (Morocco)

foreli surcoufi Santschi, 1923 (Algeria)

G
galla (Mayr, 1904) (Burkina Faso, Chad, Eritrea, Ethiopia, Ghana, Iran, Ivory Coast, Kenya, Mali, Niger, Nigeria, Oman, Saudi Arabia, Senegal, Somalia, Sudan)

• Messor barbarus latinoda Santschi, 1917

• Messor galla airensis Bernard, 1950

• Messor galla nobilis Santschi, 1928

• Messor galla obscurus Menozzi & Consani, 1952

• Messor galla rufula Finzi, 1939

grandinidus Emery, 1912 (Morocco, Tunisia)

H
hebraeus Santschi, 1927 (Iran, Israel)

hellenius Agosti & Collingwood, 1987 (Greece)

himalayanus (Forel, 1902) (Afghanistan, India, Turkey)

hismai Collingwood & Agosti, 1996 (Saudi Arabia, United Arab Emirates)

hispanicus Santschi, 1919 (Balearic Islands, Gibraltar, Iberian Peninsula, Morocco, Portugal, Spain)

hoggarensis Santschi, 1929 (Algeria, Morocco)

I
ibericus Santschi, 1931 (Bulgaria, Croatia, France, Germany, Greece, Italy, Romania, Slovenia, Spain, Switzerland)

• Messor structor novaki Finzi, 1929

incisus Stitz, 1923 (Namibia)

incorruptus Kuznetsov-Ugamsky, 1929 (Armenia, Georgia, Iran, Kazakhstan, Russian Federation)

• Messor barbarus hamulifera Karavaiev, 1926

inermis Kuznetsov-Ugamsky, 1929 (China, Kazakhstan, Kyrgyzstan)

instabilis (Smith, F., 1858) (India, Israel)

• Aphaenogaster barbara punctata Forel, 1886

intermedius Santschi, 1927 (Greece, Israel, Oman, Syrian Arab Republic, Turkey, Turkmenistan)

K
kasakorum Arnoldi, 1970 (Kazakhstan)

kasakorum nadezhdae Arnoldi, 1970 (Kazakhstan)

kisilkumensis Arnoldi, 1970 (Turkmenistan, Uzbekistan)

L
laboriosus Santschi, 1927 (China, Greece, Iran, Israel, Kazakhstan, Kyrgyzstan, Montenegro, Republic of Macedonia, Russian Federation, Turkey, Turkmenistan)

• Messor minor denticulatus Santschi, 1927

lamellicornis Arnoldi, 1968 (Kazakhstan, Turkmenistan)

lobicornis Forel, 1894 (Algeria, Gibraltar, Iberian Peninsula, Morocco, Portugal, Spain)

lobicornis batnensis Forel, 1909 (Algeria)

lobicornis laurenti Santschi, 1939 (Algeria)

lobicornis normandi Santschi, 1918 (Tunisia)

lobicornis rugulosus Santschi, 1926 (Algeria)

• Messor lobicornis rugosa Karavaiev, 1912

lobicornis submuticus Emery, 1908 (Tunisia)

luebberti Forel, 1910 (Angola, Botswana, Namibia, South Africa, United Republic of Tanzania, Zimbabwe)

luridus Santschi, 1927 (Turkey)

luridus virginalis Santschi, 1927 (Syrian Arab Republic)

lusitanicus Tinaut, 1985 (Gibraltar, Iberian Peninsula, Portugal, Spain)

M
maculifrons Santschi, 1927 (Israel, Syrian Arab Republic)

marikovskii Arnoldi, 1970 (Kazakhstan, Kyrgyzstan)

marocanus Santschi, 1927 (Gibraltar, Iberian Peninsula, Morocco, Portugal, Spain)

mcarthuri Steiner, Csősz, Markó, Gamisch, Rinnhofer, Folterbauer, Hammerle, Stauffer, Arthofer & Schlick-Steiner, 2018 (Greece, Turkey)

medioruber Santschi, 1910 (Algeria, Iran, Iraq, Kuwait, Morocco, Saudi Arabia, Tunisia, United Arab Emirates)

• Messor meridionalis sublaeviceps Santschi, 1927

medioruber maurus Santschi, 1927 (Tunisia)

medioruber montanus Santschi, 1927 (Algeria)

medioruber postpetiolatus Santschi, 1917 (Tunisia)

medioruber postquadratus Santschi, 1932 (Algeria)

melancholicus Arnoldi, 1977 (Armenia, Georgia, Iran, Kazakhstan)

• Messor melancholicus schelkovnikovi Arnoldi, 1977

meridionalis (André, 1883) (Afghanistan, Albania, Georgia, Greece, Iran, Iraq, Israel, Kuwait, Montenegro, Republic of Macedonia, Saudi Arabia, Turkey, United Arab Emirates)

minor (André, 1883) (Canary Islands, India, Iran, Iraq, Israel, Italy, Kuwait, Montenegro, Morocco, Saudi Arabia)

• Messor minor pompeianus Santschi, 1927

minor calabricus Santschi, 1927 (Italy)

minor capreensis Santschi, 1927 (Italy)

minor hesperius Santschi, 1927 (Canary Islands)

minor maurus Barquin, 1981 (Canary Islands)

muraywahus Collingwood & Agosti, 1996 (Saudi Arabia)

muscatus Collingwood & Agosti, 1996 (Oman)

muticus (Nylander, 1849) (Armenia, Kazakhstan, Kyrgyzstan, Romania, Russian Federation, Ukraine)

N
nahali Tohmé, G. & Tohmé, H., 1981 (Iran, Syrian Arab Republic)

niloticus Santschi, 1938 (Egypt)

nondentatus He & Song, 2009 (China)

O
oertzeni Forel, 1910 (Bulgaria, Greece, Republic of Macedonia, Turkey)

• Messor oertzeni amphigea Forel, 1911

oertzeni amasiensis Emery, 1921 (Turkey)

oertzeni carpathous Menozzi, 1936

olegianus Arnoldi, 1970 (Kyrgyzstan, Tajikistan)

orientalis (Emery, 1898) (Afghanistan, China, Greece, Iran, Iraq, Israel, Republic of Macedonia, Turkey)

P
perantennatus Arnoldi, 1970 (China, Kazakhstan, Turkmenistan)

piceus Stitz, 1923 (Botswana, Namibia, South Africa)

picturatus Santschi, 1927 (Algeria, Iran, Iraq, Kuwait, Morocco, Saudi Arabia)

ponticus Steiner, Csősz, Markó, Gamisch, Rinnhofer, Folterbauer, Hammerle, Stauffer, Arthofer & Schlick-Steiner, 2018 (Bulgaria, Romania, Turkey, Ukraine)

R
regalis (Emery, 1892) (Benin, Congo, Ivory Coast, Nigeria)

• Cratomyrmex regalis rubea Santschi, 1913

• Cratomyrmex sculpturatus Stitz, 1916

reticuliventris Karavaiev, 1910 (Afghanistan, Kazakhstan, Turkmenistan)

rufotestaceus (Foerster, 1850) (Algeria, Iran, Israel, Oman, Saudi Arabia, Syrian Arab Republic, United Arab Emirates)

• Aphaenogaster gracilinodis Emery, 1878

• Atta thoracica Mayr, 1862

rufus Santschi, 1923 (Kazakhstan, Kyrgyzstan, Russian Federation)

• Messor clivorum karawajevi Tarbinsky, 1976

• Messor similis Kuznetsov-Ugamsky, 1927

rufus striolatus Kuznetsov-Ugamsky, 1927 (Kyrgyzstan)

ruginodis Stitz, 1916 (Congo)

rugosus (André, 1881) (Israel)

rugosus bodenheimeri Menozzi, 1933 (Israel)

S
sanctus Emery, 1921 (Algeria, Balearic Islands, Libya, Morocco, Tunisia)

sanganus Collingwood & Agosti, 1996 (Yemen)

semirufus (André, 1883) (Afghanistan, India, Iran, Iraq, Israel, Italy, Turkey)

• Messor semirufus kutteri Santschi, 1934

• Messor semirufus nigricans Santschi, 1932

• Messor semirufus nigriceps Santschi, 1929

semirufus emeryi Santschi, 1927 (Israel)

semoni (Forel, 1906) (Algeria)

striatellus Arnoldi, 1970 (China, Tajikistan)

striaticeps (André, 1883) (Iraq, Kuwait, Saudi Arabia)

striatifrons Stitz, 1923 (Namibia, South Africa)

striativentris Emery, 1908 (Algeria, Morocco)

striatulus (Emery, 1891) (Algeria, Morocco, Tunisia)

structor (Latreille, 1798) (Austria, Bulgaria, Czech Republic, France, Hungary, Romania, Slovenia)

• Formica aedificator Schilling, 1839

• Formica lapidum Fabricius, 1804

• Formica rufitarsis Fabricius, 1804

• Messor platyceras rubella Crawley, 1920

• Messor rufitarsis darianus Pisarski, 1967

• Messor rufitarsis jakowlevi Arnoldi, 1977

• Messor structor subpolitus Kuznetsov-Ugamsky, 1927

• Messor structor turanicus Kuznetsov-Ugamsky, 1927

structor platyceras Crawley, 1920 (Iran)

structor tyrrhenus Emery, 1898 (Gibraltar, Italy, Spain)

subgracilinodis Arnoldi, 1970 (China, Russian Federation, Turkmenistan)

sultanus Santschi, 1917 (Iran, Israel, Turkey)

syriacus Tohmé, G., 1969 (Armenia, Iran, Iraq, Israel, Saudi Arabia, Syrian Arab Republic, Turkey)

T
tadzhikorum Arnoldi, 1970 (Tajikistan)

tataricus Ruzsky, 1905 (Russian Federation)

testaceus Donisthorpe, 1950 (Iran, Syrian Arab Republic, Turkey)

testaceus Bernard Bernard, 1980

timidus Espadaler, 1997 (Iberian Peninsula, Spain)

tropicorum Wheeler, W.M., 1922 (Angola, Namibia)

• Messor braunsi nigriventris Stitz, 1923

• Messor denticornis laevifrons Stitz, 1923

turcmenochorassanicus Arnoldi, 1977 (Iran, Kazakhstan, Turkmenistan)

V
valentinae Arnoldi, 1970 (China, Kyrgyzstan, Tajikistan)

variabilis Kuznetsov-Ugamsky, 1927 (Kazakhstan, Turkmenistan)

• Messor excursionis macrops Santschi, 1927

varrialei Emery, 1921 (Republic of Macedonia, Turkey)

• Messor structor aegaeus Santschi, 1926

vaucheri Emery, 1908 (Morocco)

vicinus Kuznetsov-Ugamsky, 1927 (Kazakhstan, Turkmenistan)

W
wasmanni Krausse, 1910 (Bulgaria, Croatia, Greece, Iran, Israel, Italy, Lebanon, Malta, Montenegro, Oman, Republic of Macedonia, Romania, Slovenia, Syrian Arab Republic, Turkey)

• Messor semirufus concolor Santschi, 1927

=Fossil Species=

S
sculpturatus Carpenter, 1930