Checklist of Pheidole species

The following species and subspecies belong to the genus Pheidole. Synonyms are listed under their senior names. For valid names only see Pheidole species and for distribution information see Pheidole species by Country.

A
aana Wilson & Taylor, 1967 (Samoa)

abakytan Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

abaticanga Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

aberrans Mayr, 1868 (Argentina, Bolivia, Brazil, Paraguay, Peru)

• Pheidole aberrans diversiceps Santschi, 1916

• Pheidole aberrans fartilia Forel, 1913

• Pheidole aberrans mutica Emery, 1906

absurda Forel, 1886 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, United States)

• Pheidole ridicula Wheeler, W.M., 1916

acamata Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

acantha Eguchi, 2001 (Borneo, Indonesia, Malaysia)

accinota Wheeler, W.M., 1925 (Peru)

aciculata Wilson, 2003 (Brazil, Peru)

aculifera Wilson, 2003 (Panama)

acutidens (Santschi, 1922) (Argentina, Brazil)

acutilobata Mackay, Sola & Josens, 2011 (Argentina)

ademonia Wilson, 2003 (Ecuador, Guyana, Peru)

adrianoi Naves, 1985 (United States)

aeberlii Forel, 1894 (Senegal)

aeberlii erythraea Emery, 1901 (Eritrea, Ethiopia)

aelloea Salata & Fisher, 2020 (Madagascar)

aenescens Wilson, 2003 (Brazil)

aequiseta Santschi, 1923 (Bolivia)

aglae Forel, 1913 (Borneo, Brunei Darussalam, Indonesia, Malaysia, New Guinea, Philippines, Singapore, Thailand)

• Pheidole exasperata concordia Santschi, 1916

• Pheidole exasperata fusiformis Viehmeyer, 1914

• Pheidole exasperata polita Viehmeyer, 1914

agricola Wilson, 2003 (Mexico)

ajax Forel, 1899 (Colombia, Costa Rica, Nicaragua)

ajaxigibba Longino, 2019 (Costa Rica)

akermani Arnold, 1920 (South Africa)

ala Salata & Fisher, 2020 (Madagascar)

alacris Santschi, 1923 (Argentina)

• Pheidole flavens mediorubra Santschi, 1933

alayoi Wilson, 2003 (Cuba, Greater Antilles)

albidula Santschi, 1928 (Democratic Republic of Congo)

albipes Wilson, 2003 (Guatemala, Honduras, Mexico)

alexeter Wilson, 2003 (Brazil)

alfaroi Emery, 1896 (Colombia, Costa Rica, Panama)

alienata Borgmeier, 1929 (Brazil)

alina Salata & Fisher, 2020 (Madagascar)

allani Bingham, 1903 (China, India, Myanmar)

allarmata Wilson, 2003 (Brazil, Colombia, Ecuador, Peru, Suriname)

alpestris Wilson, 2003 (Ecuador)

alpinensis Forel, 1912 (Brazil)

alticola Wilson, 2003 (Colombia)

amabilis Wilson, 2003 (Costa Rica, Mexico)

amazonica Wilson, 2003 (Ecuador, Peru)

amber Donisthorpe, 1941 (New Guinea)

ambigua Wilson, 2003 (Brazil)

ambohimanga Salata & Fisher, 2020 (Madagascar)

ambonensis Karavaiev, 1930 (Indonesia)

ampangabe Salata & Fisher, 2021 (Madagascar)

ampla Forel, 1893 (Australia)

• Pheidole variabilis latigena Forel, 1907

ampla mackayensis Forel, 1902 (Australia)

ampla parviceps Forel, 1915 (Australia)

amplificata Viehmeyer, 1914 (New Guinea)

analavelona Salata & Fisher, 2020 (Madagascar)

anastasii Emery, 1896 (Belize, Costa Rica, Denmark, Guatemala, Honduras, Mexico, Nicaragua, Panama)

andapa Salata & Fisher, 2020 (Madagascar)

andersoni Longino, 2019 (Guatemala, Mexico)

andohahela Salata & Fisher, 2020 (Madagascar)

andrieui Santschi, 1930 (Mali, Sudan)

androsana Wheeler, W.M., 1905 (Bahamas)

angulicollis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

angulifera Wilson, 2003 (Colombia, Costa Rica)

angusta Forel, 1908 (Brazil)

• Pheidole angusta idulis Forel, 1912

angusticeps Wilson, 2003 (Costa Rica, El Salvador, Honduras)

• Pheidole gradifer Wilson, 2003

angustinigra Longino, 2019 (Guatemala)

anima Wilson, 2003 (Honduras)

ankerana Salata & Fisher, 2020 (Madagascar)

annemariae Forel, 1918 (Madagascar)

annexa Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

anomala Salata & Fisher, 2020 (Madagascar)

anosyenne Salata & Fisher, 2020 (Madagascar)

anthracina Forel, 1902 (Australia)

anthracina grandii Emery, 1914 (Australia)

anthracina orba Forel, 1902 (Australia)

antillana Forel, 1893 (Dominica, Grenada, Guadeloupe, Lesser Antilles, St. Vincent)

antipodum (Smith, F., 1858) (Australia)

• Monomorium lippulum Wheeler, W.M., 1927

• Pheidole froggatti Forel, 1902

• Pheidole myops Forel, 1902

antranohofa Salata & Fisher, 2020 (Madagascar)

antsahabe Salata & Fisher, 2020 (Madagascar)

aper Forel, 1912 (Brazil)

aphrasta Zhou & Zheng, 1999 (China)

arachnion Wilson, 2003 (Costa Rica, Nicaragua)

• Pheidole iracunda Wilson, 2003

araneoides Wilson, 2003 (Brazil, Ecuador)

arboricola Wilson, 2003 (Costa Rica)

arcifera Santschi, 1925 (Brazil)

areniphila Forel, 1910 (Botswana)

• Pheidole arenicola Forel, 1910

areniphila aurora Santschi, 1928 (United Republic of Tanzania)

argentina (Bruch, 1932) (Argentina)

arhuaca Forel, 1901 (Brazil, Colombia)

• Pheidole arhuaca bimons Forel, 1912

ariel Wilson, 2003 (Mexico)

aripoensis Wilson, 2003 (Trinidad and Tobago)

aristotelis Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Singapore, Thailand)

arivo Salata & Fisher, 2021 (Madagascar)

arnoldi Forel, 1913 (Zimbabwe)

arnoldi ballaensis Arnold, 1920 (Zimbabwe)

arnoldi rufescens Arnold, 1920 (Zimbabwe)

aspera Mayr, 1862 (South Africa)

asperata Emery, 1895 (India, Myanmar)

asperithorax Emery, 1894 (Bolivia, Brazil, Peru)

aspidata Eguchi & Bui, 2005 (Vietnam)

astur Wilson, 2003 (Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname)

athertonensis Forel, 1915 (Australia)

athertonensis cedarensis Forel, 1915 (Australia)

athertonensis tambourinensis Forel, 1915 (Australia)

atitlana Longino, 2019 (Guatemala)

atsirakambiaty Salata & Fisher, 2020 (Madagascar)

atticola Forel, 1912 (Argentina)

atua Wilson & Taylor, 1967 (Samoa)

aurea Wilson, 2003 (United States)

aurivillii Mayr, 1896 (Cameroun, Guinea, Kenya, Uganda)

aurivillii attenuata Santschi, 1910 (Congo)

aurivillii kasaiensis Forel, 1911 (Democratic Republic of Congo)

aurivillii rubricalva Forel, 1915 (Democratic Republic of Congo)

auropilosa Mayr, 1887 (Argentina, Brazil, Paraguay)

avaratra Salata & Fisher, 2020 (Madagascar)

avia Forel, 1908 (Brazil)

azteca Wilson, 2003 (Mexico)

B
bahai (Forel, 1922)

bajaensis Wilson, 2003 (Mexico)

bakeri Forel, 1912 (Cuba, Greater Antilles)

balatro Longino, 2019 (Costa Rica, Guatemala, Honduras)

balcanica Seifert, 2016 (Bosnia and Herzegovina, Bulgaria, Croatia, Greece, Montenegro, Turkey)

balzani Emery, 1894 (Bolivia, Venezuela)

bambusarum Forel, 1908 (Argentina, Brazil)

• Pheidole ogloblini Kusnezov, 1952

bandata Bharti, 2004 (India)

barbata Wheeler, W.M., 1908 (Mexico, United States)

barreleti Forel, 1903 (Sri Lanka)

barumtaun Donisthorpe, 1938 (New Guinea)

batrachorum Wheeler, W.M., 1922 (Central African Republic, Democratic Republic of Congo, Gabon)

beanka Salata & Fisher, 2020 (Madagascar)

beauforti Emery, 1911 (New Guinea)

befotaka Salata & Fisher, 2020 (Madagascar)

bellatrix Wilson, 2003 (Colombia)

belli Mann, 1919 (Solomon Islands)

beloceps Wilson, 2003 (Costa Rica, Honduras, Mexico, Nicaragua)

belonorte Longino, 2019 (Guatemala, Honduras, Mexico, Nicaragua)

bemarahaensis Salata & Fisher, 2020 (Madagascar)

bemarivoensis Salata & Fisher, 2020 (Madagascar)

bequaerti Forel, 1913 (Democratic Republic of Congo)

bergi Mayr, 1887 (Argentina, Brazil, Paraguay, Uruguay)

• Pheidole bergi pulliventris Santschi, 1929

• Pheidole bergi subparallela Emery, 1906

• Pheidole goetschi Santschi, 1939

besalon Longino, 2019 (Mexico)

bessonii Forel, 1891 (Madagascar)

bicarinata Mayr, 1870 (United States)

• Pheidole bicarinata vinelandica Forel, 1886

• Pheidole hayesi Smith, M.R., 1924

• Pheidole vinelandica buccalis Wheeler, W.M., 1908

• Pheidole vinelandica laeviuscula Emery, 1895

• Pheidole vinelandica longula Emery, 1895

• Pheidole vinelandica nebrascensis Forel, 1922

biconstricta Mayr, 1870 (Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Trinidad and Tobago, Venezuela)

• Pheidole biconstricta bicolor Emery, 1890

• Pheidole biconstricta burtoni Mann, 1916

• Pheidole biconstricta hybrida Emery, 1894

• Pheidole biconstricta rubicunda Emery, 1890

• Pheidole biconstricta surda Forel, 1912

• Pheidole holmgreni Wheeler, W.M., 1925

• Pheidole holmgreni festata Wheeler, W.M., 1925

• Pheidole inermis Mayr, 1870

• Pheidole radoszkowskii lallemandi Forel, 1901

bicornis Forel, 1899 (Costa Rica, Panama)

bicornisculpta Longino, 2019 (Costa Rica)

bidens Wilson, 2003 (Brazil)

bifurca Donisthorpe, 1941 (New Guinea)

bigote Longino, 2009 (Guatemala, Mexico)

bilimeki Mayr, 1870 (Bahamas, Belize, Colombia, Costa Rica, Cuba, Greater Antilles, Guatemala, Honduras, Mexico, Nicaragua, Panama, Venezuela)

• Pheidole anastasii cellarum Forel, 1908

• Pheidole anastasii johnsoni Wheeler, W.M., 1907

• Pheidole anastasii venezuelana Forel, 1905

• Pheidole floridana antoniensis Forel, 1901

• Pheidole floridana ares Forel, 1908

• Pheidole floridana deplanata Pergande, 1896

• Pheidole punctatissima annectens Wheeler, W.M., 1905

• Pheidole punctatissima insulana Wheeler, W.M., 1905

• Pheidole rectiluma Wilson, 2003

biloba (Karavaiev, 1935) (Cambodia)

binara Salata & Fisher, 2020 (Madagascar)

binasifera Wilson, 2003 (Colombia)

binghamii Forel, 1902 (China, India, Pakistan, Singapore, Vietnam)

biolleyi Forel, 1908 (Costa Rica, Mexico)

• Pheidole biolleyi tristani Forel, 1908

bison Wilson, 2003 (Bolivia)

blumenauensis Kempf, 1964 (Brazil)

• Pheidole diligens Santschi Santschi, 1923

bluntschlii Forel, 1911 (Indonesia, Thailand)

boliviana Wilson, 2003 (Bolivia, Colombia, Costa Rica, Panama)

• Pheidole mincana Wilson, 2003

• Pheidole scitula Wilson, 2003

boltoni Wilson, 2003 (Costa Rica)

• Pheidole humida Wilson, 2003

borgmeieri Kempf, 1972 (Brazil)

boribora Salata & Fisher, 2020 (Madagascar)

boruca Wilson, 2003 (Colombia, Costa Rica, Honduras)

bos Forel, 1893 (Australia)

• Pheidole ampla perthensis Crawley, 1922

• Pheidole pyriformis Clark, 1938

bos baucis Forel, 1910 (Australia)

bos eubos Forel, 1915 (Australia)

brachyops Wilson, 2003 (Costa Rica)

brandaoi Wilson, 2003 (Colombia)

branstetteri Longino, 2009 (Belize, Guatemala, Honduras, Mexico)

braueri Forel, 1897 (Seychelles)

brevicona Mayr, 1887 (Brazil)

• Pheidole kraepelini Forel, 1901

brevicornis Mayr, 1876 (Australia)

brevipilosa Salata & Fisher, 2020 (Madagascar)

breviseta Santschi, 1919 (Argentina)

brownampla Longino, 2019 (Costa Rica)

browni Wilson, 2003 (Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

bruchella Forel, 1915 (Argentina)

bruchi Forel, 1914 (Argentina, Paraguay)

bruesi Wheeler, W.M., 1911 (Grenada, Lesser Antilles)

brunnescens Santschi, 1929 (Brazil)

bucculenta Forel, 1908 (Brazil)

buchholzi Mayr, 1901 (Cameroun, Guinea, Ivory Coast)

buckleyi Smith, M.R., 1951

• Atta pennsylvanica Buckley, 1867

bufo Wilson, 2003 (Brazil, Colombia, Ecuador)

bula Sarnat, 2008 (Fiji)

bulliceps Wilson, 2003 (Colombia, Ecuador)

bureni Wilson, 2003 (United States)

butteli Forel, 1913 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Thailand)

C
caffra Emery, 1895 (South Africa, United Republic of Tanzania)

caffra abyssinica Forel, 1910 (Eritrea, Ethiopia)

caffra amoena Forel, 1911 (Eritrea, Ethiopia)

caffra bayeri Forel, 1916 (Democratic Republic of Congo)

caffra montivaga Santschi, 1939 (Zimbabwe)

caffra senilifrons Wheeler, W.M., 1922 (Democratic Republic of Congo)

cahui Longino, 2019 (Guatemala)

cairnsiana Forel, 1902 (Australia)

caldwelli Mann, 1921 (Fiji)

calens Forel, 1901 (Brazil, Colombia, Costa Rica, Honduras, Mexico, Panama, Peru, Venezuela)

californica Mayr, 1870 (Mexico, United States)

• Pheidole californica hagermani Cole, 1936

• Pheidole californica incenata Wheeler, W.M., 1915

• Pheidole californica nevadensis Wheeler, W.M., 1915

• Pheidole californica pyramidensis Emery, 1922

• Pheidole californica satura Wheeler, W.M., 1915

• Pheidole californica shoshoni Cole, 1933

• Pheidole oregonica Emery, 1895

caliginosa Longino, 2019 (Costa Rica)

calimana Wilson, 2003 (Colombia)

caltrop Wilson, 2003 (Costa Rica, Honduras, Panama)

cameroni Mayr, 1887 (Ecuador)

• Pheidole monticola Cameron, 1891

camilla Wilson, 2003 (Ecuador)

camptostela Kempf, 1972 (Brazil)

cangussu Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

capellinii Emery, 1887 (China, India, Indonesia, Thailand, Vietnam)

• Pheidole attila Forel, 1913

• Pheidole lighti Wheeler, W.M., 1927

capensis Mayr, 1862 (Mozambique, South Africa)

capensis dregei Emery, 1895 (South Africa)

capensis modestior Santschi, 1919 (South Africa)

capensis reddersbergensis Forel, 1913 (South Africa)

capillata Emery, 1906 (Brazil, Peru)

caracalla Wilson, 2003 (Ecuador, Peru)

carapuna Mann, 1916 (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Trinidad and Tobago, Venezuela)

• Pheidole carapuna chaquimayensis Wheeler, W.M., 1925

• Pheidole tristicula Wilson, 2003

carapunco Kusnezov, 1952 (Argentina)

cardiella Wilson, 2003 (Ecuador, Peru)

cardinalis Wilson, 2003 (Brazil)

caribbaea Wheeler, W.M., 1911 (Jamaica)

• Pheidole caribbaea sloanei Wheeler, W.M., 1917

carinata Wilson, 2003 (Costa Rica)

cariniceps Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore, Thailand)

carinitida Longino, 2019 (Costa Rica)

carinote Longino, 2009 (Costa Rica)

carrolli Naves, 1985 (United States)

casta Wheeler, W.M., 1908 (United States)

castanea (Smith, F., 1858) (Brazil)

cataphracta Wilson, 2003 (Colombia)

cataractae Wheeler, W.M., 1916 (Brazil, Guyana, Peru)

caulicola Wilson, 2003 (Costa Rica)

cavifrons Emery, 1906 (Argentina, Brazil, Uruguay)

• Pheidole arciruga Forel, 1908

• Pheidole cavifrons fuscipunctis Santschi, 1916

cavigenis Wheeler, W.M., 1915 (Mexico, United States)

ceibana Wilson, 2003 (Honduras)

celaena Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

centeotl Wheeler, W.M., 1914 (Mexico)

cerebrosior Wheeler, W.M., 1915 (Mexico, United States)

ceres Wheeler, W.M., 1904 (Mexico, United States)

• Pheidole ceres tepaneca Wheeler, W.M., 1914

cerina Wilson, 2003 (Costa Rica, Panama)

cervicornis Emery, 1900 (New Guinea)

ceylonica (Motschoulsky, 1863) (Sri Lanka)

chalca Wheeler, W.M., 1914 (Mexico)

charazana Wilson, 2003 (Colombia)

cheesmanae Donisthorpe, 1941 (Indonesia, New Guinea)

chilensis Mayr, 1862 (Chile, Peru)

chocoensis Wilson, 2003 (Colombia, Costa Rica, Honduras, Nicaragua, Panama)

christinae Fischer, Hita Garcia & Peters, 2012 (Uganda)

christopherseni Forel, 1912 (Colombia, Costa Rica, Honduras, Mexico, Panama)

chrysops Wilson, 2003 (Brazil, Paraguay)

cicatricosa Stitz, 1917 (Egypt, Iran, Israel, Mali)

• Pheidole pallidula recticeps Menozzi, 1932

cingulata (Smith, F., 1857) (Borneo, Indonesia, Malaysia)

citrina Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

clara Salata & Fisher, 2020 (Madagascar)

clavata (Emery, 1877) (Eritrea, Ethiopia)

claviscapa Santschi, 1925 (Bolivia, Brazil)

clementensis Gregg, 1969 (Mexico, United States)

clydei Gregg, 1950 (Mexico, United States)

clypeocornis Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore, Thailand)

cockerelli Wheeler, W.M., 1908 (Mexico, United States)

coffeicola Borgmeier, 1934 (Brazil, Suriname)

colaensis Mann, 1921 (Fiji)

colobopsis Mann, 1916 (Brazil, Colombia, Guatemala, Honduras, Mexico, Peru, Venezuela)

colpigaleata Eguchi, 2006 (Vietnam)

comata Smith, F., 1858 (Borneo, Indonesia, Malaysia, Thailand)

• Pheidole multicoma Eguchi, 1999

comosa Salata & Fisher, 2021 (Madagascar)

concentrica Forel, 1902 (Australia)

concentrica recurva Forel, 1910 (Australia)

concinna Santschi, 1910 (Congo)

conficta Forel, 1902 (Australia)

confoedusta Wheeler, W.M., 1909 (Mexico)

constanciae Forel, 1902 (China, India)

constanciae nigra Forel, 1902 (India)

constipata Wheeler, W.M., 1908 (United States)

coonoorensis Forel, 1902 (India)

coracina Wilson, 2003 (Mexico)

cordiceps Mayr, 1868 (Argentina)

• Pheidole cordiceps lilianae Forel, 1912

corniclypeus Longino, 2019 (Costa Rica, Honduras, Nicaragua)

cornicula Wilson, 2003 (Bolivia, Paraguay)

corticicola Santschi, 1910 (Congo, Guinea, Ivory Coast)

costaricensis Longino, 2019 (Costa Rica)

coveri Wilson, 2003 (Trinidad and Tobago, Venezuela)

cramptoni Wheeler, W.M., 1916 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

• Pheidole cramptoni petiolicola Wheeler, W.M., 1921

crassicornis Emery, 1895 (Mexico, United States)

crassinoda Emery, 1895 (Botswana, Kenya, Namibia, South Africa, Uganda)

crassinoda pluto Arnold, 1920 (Mozambique)

crassinoda ruspolii Emery, 1897 (Ethiopia, Kenya, Zimbabwe)

crassinoda sordidula Santschi, 1937 (United Republic of Tanzania)

creightoni Gregg, 1955 (United States)

crinita Wilson, 2003 (Ecuador)

crozieri Wilson, 2003 (Peru)

cryptocera Emery, 1900 (New Guinea)

cubaensis Mayr, 1862 (Cuba, Greater Antilles)

• Pheidole cubaensis grayi Mann, 1920

cuevasi Wilson, 2003 (Brazil, Peru)

cuitensis Forel, 1910 (Angola)

cuprina Wilson, 2003 (Peru)

cursor Wilson, 2003 (Brazil, Ecuador, Guyana, Peru)

curupira Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

curvistriata Salata & Fisher, 2020 (Madagascar)

cusuco Longino, 2019 (Guatemala, Honduras)

cyrtostela Wilson, 2003 (Brazil, Paraguay)

D
dammermani Wheeler, W.M., 1924 (Indonesia)

daphne Wilson, 2003 (Costa Rica)

darlingtoni Wheeler, W.M., 1936 (Greater Antilles, Haiti)

darwini Fischer, Hita Garcia & Peters, 2012 (Gabon)

dasos Salata & Fisher, 2020 (Madagascar)

dasypyx Wilson, 2003 (Costa Rica, Panama)

davidsonae Wilson, 2003 (Peru)

davisi Wheeler, W.M., 1905 (Mexico, United States)

dea Santschi, 1921 (Democratic Republic of Congo)

debilis Longino, 2009 (Costa Rica)

decarinata Santschi, 1929 (Mali, Sudan)

deceptrix Forel, 1899 (Belize, Guatemala, Honduras, Mexico)

• Pheidole chiapasana Wilson, 2003

• Pheidole variceps Wilson, 2003

decollata Forel, 1892 (Madagascar)

defecta Santschi, 1923 (Guatemala)

deima Wilson, 2003 (Brazil, Ecuador, Peru)

delecta Forel, 1899 (Mexico)

delicata Wilson, 2003 (Bolivia)

deltea Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines)

demeter Wilson, 2003 (Peru)

dentata Mayr, 1886 (Mexico, United States)

• Leptothorax tennesseensis Cole, 1938

• Pheidole commutata Mayr, 1886

• Pheidole dentata faisonsica Forel, 1901

dentigula Smith, M.R., 1927 (Mexico, United States)

depressinoda Longino, 2019 (Guatemala)

descolei Kusnezov, 1952 (Argentina, Brazil)

deserticola Forel, 1910 (Australia)

deserticola foveifrons Viehmeyer, 1924 (Australia)

desertorum Wheeler, W.M., 1906 (Mexico, United States)

• Pheidole desertorum comanche Wheeler, W.M., 1906

• Pheidole desertorum maricopa Wheeler, W.M., 1906

diabolus Wilson, 2003 (Colombia, Costa Rica)

diakritos Salata & Fisher, 2020 (Madagascar)

diana Forel, 1908 (Costa Rica)

diffidens (Walker, 1859) (Sri Lanka)

diffusa (Jerdon, 1851) (India)

diligens (Smith, F., 1858) (Brazil, Peru)

dinophila Wilson, 2003 (Brazil)

dione Forel, 1913 (Argentina)

dispar (Forel, 1895) (Australia)

distincta Donisthorpe, 1943 (New Guinea)

distorta Forel, 1899 (Colombia)

diversipilosa Wheeler, W.M., 1908 (Mexico, United States)

dodo Fischer & Fisher, 2013 (Mauritius)

dolon Wilson, 2003 (Bolivia, Peru)

dorsata Wilson, 2003 (Colombia, Guyana)

dossena Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

drepanon Wilson, 2003 (Dominican Republic, Greater Antilles)

drogon Sarnat, Fischer & Economo, 2016 (New Guinea)

dryas Wilson, 2003 (Costa Rica)

dugasi Forel, 1911 (China, Laos, Thailand, Vietnam)

dumicola Wilson, 2003 (Mexico)

duneraensis Bharti, 2001 (India)

durionei Santschi, 1923 (Argentina)

dwyeri Gregg, 1969 (Mexico)

dyctiota Kempf, 1972 (Brazil)

E
ectatommoides Wilson, 2003 (Colombia, Costa Rica, Ecuador, Panama)

ecuadorana Wilson, 2003 (Ecuador)

ehazoara Salata & Fisher, 2020 (Madagascar)

eidmanni Menozzi, 1926 (Brazil)

elecebra (Wheeler, W.M., 1904) (United States)

elegans Donisthorpe, 1938 (New Guinea)

elisae Emery, 1900 (Borneo, Indonesia, Malaysia, Philippines, Singapore, Thailand)

• Pheidole elisae nenia Forel, 1913

elongicephala Eguchi, 2008 (China, Vietnam)

embolopyx Brown, 1968 (Brazil, Ecuador, Peru)

emmae Forel, 1905 (Indonesia)

ensifera Forel, 1897 (Madagascar)

eosimilis Longino, 2019 (Costa Rica, Nicaragua)

eowilsoni Longino, 2009 (Costa Rica, Honduras, Nicaragua)

eparmata Wilson, 2003 (Venezuela)

epetrion Wilson, 2003 (Ecuador)

epiphyta Longino, 2009 (Costa Rica)

erato Mann, 1919 (Solomon Islands)

erethizon Wilson, 2003 (Mexico)

eriophora Wilson, 2003 (Colombia)

ernsti Forel, 1912 (Taiwan)

erratilis Wilson, 2003 (Costa Rica, Nicaragua)

• Pheidole petersoni Wilson, 2003

escherichii Forel, 1910 (Eritrea, Ethiopia)

euryscopa Wilson, 2003 (Colombia)

exarata Emery, 1896 (Colombia, Costa Rica, Honduras, Nicaragua, Venezuela)

• Pheidole crassipes grantae Forel, 1908

exasperata (Mayr, 1866) (China)

excellens Mayr, 1862 (Ghana, Guinea, Ivory Coast)

excellens fulvobasalis Santschi, 1921 (Benin)

excellens weissi Santschi, 1910 (Congo)

excubitor Wilson, 2003 (Costa Rica, Mexico)

exigua Mayr, 1884 (Barbados, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Greater Antilles, Guadeloupe, Guyana, Mexico, Netherlands Antilles, Panama, Peru, Puerto Rico, Trinidad and Tobago, Venezuela)

exquisita Wilson, 2003 (Ecuador)

F
fabricator (Smith, F., 1858) (Brazil, Colombia, Venezuela)

• Atta nigriventris Smith, F., 1858

fadli Sharaf, 2007 (Egypt)

fallax Mayr, 1870 (Colombia, Costa Rica, Cuba, Grenada, Guadeloupe, Guatemala, Guyana, Honduras, Jamaica, Netherlands Antilles, Nicaragua, Puerto Rico, Trinidad and Tobago, Venezuela)

• Pheidole fallax britoi Forel, 1912

• Pheidole fallax columbica Forel, 1886

• Pheidole fallax ovalis Forel, 1912

• Pheidole fallax rubens Forel, 1899

• Pheidole jelskii fallacior Forel, 1901

familiaparra Longino, 2019 (Costa Rica)

fantasia Chapman, 1963 (Borneo, Indonesia, Malaysia, Philippines)

fatigata Bolton, 1995 (New Guinea)

• Pheidole granulatus Donisthorpe, 1949

feae Emery, 1895 (China, India, Myanmar)

fera Santschi, 1925 (Brazil)

fergusoni Forel, 1902 (India, Pakistan, Sri Lanka)

ferruginea Salata & Fisher, 2020 (Madagascar)

fervens Smith, F., 1858 (Borneo, China, Cook Islands, Fiji, French Polynesia, Guam, India, Indonesia, Iran, Japan, Kiribati, Krakatau Islands, Malaysia, Marshall Islands, Mauritius, Micronesia (Federated States of), New Caledonia, Northern Mariana Islands, Palau, Philippines, Pitcairn, Samoa, Singapore, Sri Lanka, Taiwan, Thailand, Tokelau, Tonga, United States, Vanuatu, Vietnam)

• Pheidole amia Forel, 1912

• Pheidole cavannae Emery, 1887

• Pheidole javana Mayr, 1867

• Pheidole javana desucta Wheeler, W.M., 1929

• Pheidole javana dharmsalana Forel, 1902

• Pheidole javana dolenda Forel, 1912

• Pheidole javana soror Santschi, 1937

• Pheidole nodus azumai Santschi, 1941

• Pheidole oceanica nigriscapa Santschi, 1928

• Solenopsis pungens Smith, F., 1861

fervens pectinata Stitz, 1912 (Indonesia)

fervida Smith, F., 1874 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation, Vietnam)

fervida kwazana Teranishi, 1940 (Republic of Korea)

fimbriata Roger, 1863 (Argentina, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole diversa Smith, F., 1860

• Pheidole fimbriata tucumana Forel, 1913

• Pheidole smithii Dalla Torre, 1892

fincanaranjo Longino, 2019 (Costa Rica)

fiorii Emery, 1890 (Costa Rica, Nicaragua, Panama)

fisaka Salata & Fisher, 2020 (Madagascar)

fissiceps Wilson, 2003 (Peru)

fitarata Salata & Fisher, 2020 (Madagascar)

flammea Salata & Fisher, 2020 (Madagascar)

flavens Roger, 1863 (Argentina, Bahamas, Barbados, Belize, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, Galapagos Islands, Greater Antilles, Grenada, Guadeloupe, Guatemala, Haiti, Honduras, Lesser Antilles, Madagascar, Mexico, Nicaragua, Panama, Paraguay, Peru, Trinidad and Tobago, United States)

• Pheidole exigua tuberculata Mayr, 1887

• Pheidole flavens gracilior Forel, 1901

• Pheidole flavens haytiana Forel, 1907

• Pheidole flavens spei Santschi, 1930

• Pheidole flavens vincentensis Forel, 1893

• Pheidole floridana aechmeae Wheeler, W.M., 1934

• Pheidole greggi Naves, 1985

flavens thomensis Emery, 1894

flaveria Zhou & Zheng, 1999 (China)

flavida Mayr, 1887 (Brazil)

flavifrons Wilson, 2003 (Brazil, Peru)

flavigaster Zhong, 2021 (China)

flavodepressa Salata & Fisher, 2020 (Madagascar)

flavominuta Salata & Fisher, 2020 (Madagascar)

flavothoracica Viehmeyer, 1914 (New Guinea)

floricola Wilson, 2003 (Costa Rica, Ecuador, Honduras, Nicaragua)

floridana Emery, 1895 (Mexico, United States)

• Pheidole lauta Wheeler, W.M., 1908

foederalis Borgmeier, 1928

foreli Mayr, 1901 (South Africa)

foreli pubens Forel, 1910 (South Africa)

fortis Eguchi, 2006 (Thailand, Vietnam)

fossimandibula Longino, 2009 (Costa Rica)

foveolata Eguchi, 2006 (Vietnam)

fowleri Wilson, 2003 (Brazil)

fracticeps Wilson, 2003 (Brazil, Ecuador, Paraguay, Peru, Trinidad and Tobago)

fullerae Wilson, 2003 (Ecuador)

funki LaPolla, 2005 (Guyana)

funkikoensis Wheeler, W.M., 1929 (China, Taiwan)

furcata Sarnat, 2008 (Fiji)

furtiva Wilson, 2003 (United States)

fuscula Emery, 1900 (New Guinea)

G
gagates Wilson, 2003 (Ecuador, Peru)

gaigei Forel, 1914 (Colombia)

galba Wilson, 2003 (Peru)

gambogia Donisthorpe, 1948 (New Guinea)

gatesi (Wheeler, W.M., 1927) (China, Myanmar, Thailand, Vietnam)

gauthieri Forel, 1901 (Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama)

• Pheidole gauthieri oxymora Forel, 1912

gellibrandi Clark, 1934 (Australia)

geminata Wilson, 2003 (Ecuador)

gemmula Wilson, 2003 (Peru)

geraesensis Santschi, 1929 (Brazil)

germaini Emery, 1896 (Brazil)

gertrudae Forel, 1886 (Argentina, Bolivia, Brazil, Colombia, Paraguay)

• Megalomyrmex humilis Borgmeier, 1930

• Pheidole gertrudae leonhardi Forel, 1901

• Pheidole gertrudae loretensis Santschi, 1933

ghatica Forel, 1902 (India)

ghigii Emery, 1900 (Borneo, Indonesia, Malaysia)

gibba Mayr, 1887 (Brazil)

• Pheidole gibba lopesi Borgmeier, 1937

gibbata Borgmeier, 1934 (Colombia, Suriname)

gigaflavens Wilson, 2003 (Argentina, Bolivia, Paraguay)

gigas Wilson, 2003 (Brazil)

gigliolii Menozzi, 1935

gilva Wilson, 2003 (Ecuador, Peru)

gilvescens Creighton & Gregg, 1955 (Mexico, United States)

glabra Salata & Fisher, 2020 (Madagascar)

glabrella Fischer, Hita Garcia & Peters, 2012 (Cameroun)

globularia Wilson, 2003 (Ecuador)

glomericeps Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

gnomus Wilson, 2003 (Ecuador)

goavana Salata & Fisher, 2020 (Madagascar)

godmani Forel, 1893 (Lesser Antilles, St. Vincent)

goeldii Forel, 1895 (Brazil)

gombakensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

gouldi Forel, 1886 (Costa Rica, Guatemala, Honduras, Mexico)

gracilescens (Smith, F., 1860) (Indonesia, Sulawesi)

gracilipes (Motschoulsky, 1863) (Sri Lanka)

gracilis Salata & Fisher, 2020 (Madagascar)

grallatrix Emery, 1899 (Madagascar)

grandinodus Wilson, 2003 (Brazil)

granulata Pergande, 1896 (Mexico)

gravida Wilson, 2003 (Venezuela)

grayi Forel, 1902 (India)

grex Wilson, 2003 (Colombia)

grundmanni Smith, M.R., 1953 (United States)

guajirana Wilson, 2003 (Colombia)

guayasana Wilson, 2003 (Ecuador)

guerrerana Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

guilelmimuelleri Forel, 1886 (Argentina, Brazil, Colombia)

• Pheidole guilelmimuelleri gustavi Forel, 1911

guineensis (Fabricius, 1793) (Borneo, Guinea, Indonesia, Malaysia)

gulo Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

gymnoceras Longino, 2009 (Costa Rica, Nicaragua)

H
haboka Salata & Fisher, 2020 (Madagascar)

hainanensis Chen, Ye, Lu & Zhou, 2011 (China)

hamtoni Wilson, 2003 (Mexico)

hansoni Longino, 2019 (Costa Rica)

harlequina Wilson, 2003 (Dominican Republic, Greater Antilles)

harrisonfordi Wilson, 2003 (Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Pheidole prolixa Wilson, 2003

• Pheidole ruida Wilson, 2003

• Pheidole tenebra Wilson, 2003

hartmeyeri Forel, 1907 (Australia)

haskinsorum Wilson, 2003 (Bolivia, Ecuador)

hasticeps Wilson, 2003 (Colombia, Costa Rica, Mexico, Panama)

havilandi Forel, 1911 (Borneo, Indonesia, Malaysia)

• Pheidole havilandi sapuana Forel, 1911

• Pheidole havilandi selangorensis Forel, 1913

havoana Salata & Fisher, 2020 (Madagascar)

haywardi Kusnezov, 1952 (Argentina, Paraguay)

hazenae Wilson, 2003 (Colombia, Ecuador)

hazo Salata & Fisher, 2020 (Madagascar)

hecate Wheeler, W.M., 1911 (Jamaica)

• Pheidole hecate bruesi Wheeler, W.M., 1917

• Pheidole hecate malevola Wheeler, W.M., 1911

hector Wilson, 2003 (Costa Rica)

hectornitida Longino, 2019 (Costa Rica)

hedlundorum Wilson, 2003 (Venezuela)

heliosa Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

hercules Donisthorpe, 1941 (New Guinea)

heterothrix Santschi, 1923 (Argentina)

• Pheidole gavrilovi Kusnezov, 1952

hetschkoi Emery, 1896 (Brazil)

• Pheidole hetschkoi frivola Santschi, 1923

hewitti Santschi, 1932 (South Africa)

• Pheidole capensis hewitti Arnold, 1944

heyeri Forel, 1899 (Brazil)

• Pheidole guilelmimuelleri ultrix Forel, 1912

hierax Wilson, 2003 (Panama)

hirsuta Emery, 1896 (Costa Rica, Guatemala, Panama)

hispaniolae Wilson, 2003 (Dominican Republic, Greater Antilles)

hitoy Longino, 2019 (Costa Rica)

hizemops Wilson, 2003 (Costa Rica)

hoelldobleri Wilson, 2003 (Costa Rica)

hongkongensis Wheeler, W.M., 1928 (China, Laos, Thailand, Vietnam)

hoplitica Wilson, 2003 (Mexico, United States)

horni Emery, 1901 (Sri Lanka)

horribilis Wilson, 2003 (Ecuador, Peru)

hortensis Forel, 1913 (Borneo, Indonesia, Malaysia, Philippines, Singapore, Thailand)

hortonae Wilson, 2003 (Venezuela)

hospes Smith, F., 1865 (New Guinea)

hospita Bingham, 1903 (India)

huacana Wilson, 2003 (Ecuador, Peru)

huarache Longino, 2019 (Guatemala, Mexico)

huberi Forel, 1911 (Borneo, Indonesia, Malaysia)

• Pheidole huberi perakensis Forel, 1911

huilana Wilson, 2003 (Colombia)

humeralis Wheeler, W.M., 1908 (United States)

humeridens Wilson, 2003 (Argentina)

hyatti Emery, 1895 (Mexico, United States)

• Pheidole hyatti ecitonodora Wheeler, W.M., 1908

• Pheidole hyatti solitanea Wheeler, W.M., 1915

• Pheidole vaslitii Pergande, 1896

I
iceni Fernández, 2011 (Colombia)

idiota Santschi, 1923 (Argentina, Brazil, Costa Rica)

• Pheidole laticornis Wilson, 2003

imbrilis Longino, 2019 (Costa Rica)

impressa Mayr, 1870 (Brazil)

impressa detrita Emery, 1890 (Venezuela)

impressiceps Mayr, 1876 (Australia, New Guinea)

impressiceps commista Forel, 1901 (New Guinea)

inca Wilson, 2003 (Peru)

incerta (Smith, F., 1863) (Indonesia)

incisa Mayr, 1870 (Colombia)

incurvata Viehmeyer, 1924 (Australia)

indagarama Longino, 2019 (Costa Rica)

indagatrix Wilson, 2003 (Costa Rica, Honduras, Nicaragua, Panama)

indica Mayr, 1879 (Antigua and Barbuda, Balearic Islands, Bangladesh, Barbados, Canary Islands, China, Colombia, Comoros, Cuba, Dominican Republic, Egypt, Eritrea, Greater Antilles, Greece, Guadeloupe, Iberian Peninsula, India, Iran, Israel, Japan, Kenya, Kuwait, Madagascar, Malta, Mauritius, Mayotte, Nepal, Oman, Pakistan, Peru, Republic of Korea, Saint Helena, Saudi Arabia, Seychelles, Socotra Archipelago, Spain, Sri Lanka, United Arab Emirates, United States, Yemen)

• Pheidole indica himalayana Forel, 1902

• Pheidole indica rotschana Forel, 1902

• Pheidole striativentris Mayr, 1879

• Pheidole teneriffana Forel, 1893

• Pheidole teneriffana taina Aguayo, 1932

indosinensis Wheeler, W.M., 1928 (China, Vietnam)

industa Santschi, 1939 (Uruguay)

infernalis Wilson, 2003 (Bolivia)

inflexa Santschi, 1923

innotata Mayr, 1866

innupta Menozzi, 1931 (Costa Rica, Peru)

inornata Eguchi, 2001 (Borneo, Indonesia, Malaysia, Thailand)

inquilina (Wheeler, W.M., 1903) (United States)

inscrobiculata Viehmeyer, 1916 (Malaysia, Philippines, Thailand)

insipida Forel, 1899 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole fariasana Wilson, 2003

• Pheidole mooreorum Wilson, 2003

inversa Forel, 1901 (Colombia)

irritans (Smith, F., 1858) (South Africa)

isis Mann, 1919 (Solomon Islands)

isis taki Mann, 1919 (Solomon Islands)

itremo Salata & Fisher, 2020 (Madagascar)

J
jacobsoni Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

• Pheidole orophila Eguchi, 2001

jaculifera Wilson, 2003 (Brazil)

jamaicensis Wheeler, W.M., 1908 (Cuba, Dominican Republic, Haiti, Jamaica)

janzeni Longino, 2009 (Costa Rica, El Salvador, Guatemala, Honduras, Mexico)

jeannei Wilson, 2003 (Brazil)

jelskii Mayr, 1884 (Argentina, Barbados, Brazil, Dominican Republic, French Guiana, Greater Antilles, Grenada, Haiti, Netherlands Antilles, Paraguay, Saint Lucia, Trinidad and Tobago)

• Pheidole fallax emiliae Forel, 1901

• Pheidole jelskii antillensis Forel, 1901

• Pheidole jelskii arenicola Emery, 1894

jivaro Wilson, 2003 (Ecuador, Panama)

joffreville Salata & Fisher, 2020 (Madagascar)

jonas Forel, 1907 (Comoros, Madagascar, Mayotte)

jordanica Saulcy, 1874 (Egypt, Israel, Saudi Arabia)

• Pheidole megacephala obtusa Stitz, 1917

• Pheidole schmitzi Forel, 1911

• Pheidole sinaitica laticeps Mayr, 1904

jubilans Forel, 1911 (Indonesia)

jucunda Forel, 1885 (China, India, Sri Lanka)

jucunda fossulata Forel, 1902 (China, India)

jujuyensis Forel, 1913 (Argentina)

juniperae Wilson, 2003 (United States)

K
karolmorae Longino, 2009 (Costa Rica, Honduras, Nicaragua)

karolsetosa Longino, 2009 (Costa Rica)

kasparii Longino, 2019 (Costa Rica)

katonae Forel, 1907 (Saudi Arabia, United Republic of Tanzania)

kava Fischer, Sarnat & Economo, 2016 (Fiji)

kelainos Longino, 2019 (Costa Rica)

kely Salata & Fisher, 2020 (Madagascar)

kikutai Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

kitschneri Forel, 1910 (Kenya, South Africa)

klaman Gómez, Kouakou, Fischer, Hita-Garcia, Katzke & Economo, 2022 (Ivory Coast)

knowlesi Mann, 1921 (Fiji)

• Pheidole knowlesi extensa Mann, 1921

kochi (Emery, 1911) (New Guinea)

kohli Mayr, 1901 (Kenya, South Africa)

komori Fischer & Fisher, 2013 (Comoros, Mozambique)

koshewnikovi Ruzsky, 1905 (Azerbaijan, Cyprus, Greece, Iran, Kazakhstan, Kyrgyzstan, Montenegro, Russian Federation, Turkey, Uzbekistan)

• Pheidole pallidula arenarum Ruzsky, 1905

• Pheidole pallidula orientalis Müller, 1923

kugleri Wilson, 2003 (Peru)

kukrana Wilson, 2003 (Costa Rica, Guadeloupe, Guatemala, Nicaragua, Venezuela)

kuna Wilson, 2003 (Panama)

kusnezovi Wilson, 2003 (Argentina)

• Eriopheidole symbiotica Kusnezov, 1952

L
laelaps Wilson, 2003 (Colombia, Ecuador)

laevicolor Eguchi, 2006 (Vietnam)

laevifrons Mayr, 1887 (Brazil)

laevinota Forel, 1908 (Argentina, Brazil)

laevithorax Eguchi, 2008 (Vietnam)

laeviventris Mayr, 1870 (Colombia)

laevivertex Forel, 1901 (Mexico)

lagunculiminor Longino, 2019 (Guatemala, Honduras)

lagunculinoda Longino, 2009 (Guatemala, Mexico)

laidlowi Mann, 1916 (Brazil, Ecuador, Peru)

lamancha Longino, 2019 (Mexico)

lamellinoda Forel, 1902 (India, Socotra Archipelago, Yemen)

lamia Wheeler, W.M., 1901 (United States)

laminata Emery, 1900 (New Guinea)

lamperos Salata & Fisher, 2020 (Madagascar)

lancifera Wilson, 2003 (Brazil, Peru)

lanigera Wilson, 2003 (Peru, Venezuela)

lanuginosa Wilson, 1984 (India)

laselva Wilson, 2003 (Colombia, Costa Rica, Ecuador, Honduras, Nicaragua, Panama, Venezuela)

• Pheidole ebenina Wilson, 2003

laselvoides Longino, 2019 (Costa Rica, Nicaragua, Panama)

laticrista Santschi, 1916 (Argentina)

latinoda Roger, 1863 (Iran, Pakistan, Sri Lanka)

latinoda angustior Forel, 1902 (India)

latinoda major Forel, 1885 (India)

latinoda peradeniyae Forel, 1911 (Sri Lanka)

lattkei Wilson, 2003 (Venezuela)

laudatana Wilson, 2003 (Dominica, Guadeloupe)

lavasoa Salata & Fisher, 2020 (Madagascar)

leloi Eguchi & Bui, 2016 (Vietnam)

lemnisca Wilson, 2003 (Colombia, Ecuador)

lemur Forel, 1912 (Brazil)

leoncortesi Longino, 2009 (Mexico)

leonina Wilson, 2003 (Brazil, Peru)

leptina Wilson, 2003 (Brazil, Peru)

liengmei Forel, 1894 (Mozambique)

liengmei malindana Forel, 1913 (Zimbabwe)

liengmei micrartifex Forel, 1913 (Zimbabwe)

liengmei shinsendensis Forel, 1913 (Democratic Republic of Congo)

lignicola Mayr, 1887 (Argentina, Brazil, Paraguay)

• Pheidole lignicola levocciput Forel, 1908

lilloi (Kusnezov, 1952) (Argentina)

lineafrons Longino, 2019 (Costa Rica)

liteae Forel, 1910 (Australia)

litigiosa Forel, 1892 (Madagascar, Madagascar)

littoralis Cole, 1952 (United States)

lobulata Emery, 1900 (New Guinea)

loki Fischer & Fisher, 2013 (Mayotte)

lokitae Forel, 1913 (Borneo, Indonesia, Malaysia, Philippines)

longiceps Mayr, 1876 (Australia)

longiceps doddi Forel, 1910 (Australia)

longiceps frontalis Forel, 1902 (Australia)

longicornis Emery, 1888 (Indonesia)

longinoi Wilson, 2003 (Costa Rica)

longior Santschi, 1933 (Argentina)

longipes (Latreille, 1802) (Borneo, French Guiana, India, Indonesia, Malaysia, Nicobar Island, Thailand)

longipilosa Salata & Fisher, 2020 (Madagascar)

longiscapa Forel, 1901 (Brazil, Colombia, Costa Rica, Greater Antilles, Guyana, Nicaragua, Puerto Rico, Suriname, Venezuela)

• Pheidole cocciphaga Borgmeier, 1934

• Pheidole longiscapa martensis Forel, 1914

longiseta Wilson, 2003 (Brazil, Peru)

longispinosa Forel, 1891 (Madagascar)

lourothi Wilson, 2003 (Costa Rica)

lovejoyi Wilson, 2003 (Brazil)

lucaris Wilson, 2003 (Brazil, Costa Rica)

lucida Forel, 1895 (Madagascar)

lucioccipitalis Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

lucretii Santschi, 1923 (Brazil)

lupus Wilson, 2003 (Ecuador, Peru)

lustrata Wilson, 2003 (Costa Rica)

lutea Salata & Fisher, 2020 (Madagascar)

luteagossamer Longino, 2019 (Honduras, Nicaragua)

luteipes Emery, 1914 (New Caledonia)

luteipes obvia Emery, 1914 (New Caledonia)

lutzi Forel, 1905 (Brazil)

• Pheidole lutzi heinzi Forel, 1913

M
macclendoni Wheeler, W.M., 1908 (United States)

machaquila Longino, 2019 (Guatemala)

machetula Wilson, 2003 (Peru)

mackayi Wilson, 2003 (Mexico)

macracantha Wilson, 2003 (Panama)

macromischoides Wilson, 2003 (Cuba)

macrops Wilson, 2003 (Mexico)

maculifrons Wheeler, W.M., 1929 (Philippines)

madecassa Forel, 1892 (Madagascar)

madinika Salata & Fisher, 2020 (Madagascar)

madrensis Wilson, 2003 (Mexico)

magna Eguchi, 2006 (Vietnam)

magrettii Emery, 1887 (Indonesia, Malaysia, Singapore, Thailand)

• Pheidole nodgii maxwellensis Forel, 1913

mahaboensis Salata & Fisher, 2020 (Madagascar)

mahamavo Salata & Fisher, 2020 (Madagascar)

mainty Salata & Fisher, 2020 (Madagascar)

maizina Salata & Fisher, 2020 (Madagascar)

maja Forel, 1886 (Guatemala)

makaensis Salata & Fisher, 2020 (Madagascar)

makilingi Viehmeyer, 1916 (Philippines)

makirovana Salata & Fisher, 2020 (Madagascar)

malabarica (Jerdon, 1851) (India)

malinsii Forel, 1902 (China, India, Sri Lanka)

mallota Wilson, 2003 (Panama)

mamirapiratra Salata & Fisher, 2021 (Madagascar)

mamiratra Salata & Fisher, 2020 (Madagascar)

mamore Mann, 1916 (Brazil, Guyana, Peru, Suriname)

manantenensis Salata & Fisher, 2020 (Madagascar)

manantenina Salata & Fisher, 2020 (Madagascar)

mantadia Salata & Fisher, 2020 (Madagascar)

mantadioflava Salata & Fisher, 2020 (Madagascar)

manteroi Emery, 1897 (New Guinea)

mantilla Wilson, 2003 (Costa Rica, Panama)

manuana Wilson, 2003 (Peru)

manukana Eguchi, 2001 (Borneo, Indonesia, Malaysia)

mapinguari Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

marcidula Wheeler, W.M., 1908 (Mexico, United States)

marieannae Salata & Fisher, 2020 (Madagascar)

marmor Longino, 2019 (Guatemala)

maro Salata & Fisher, 2020 (Madagascar)

masoala Salata & Fisher, 2020 (Madagascar)

masoandro Salata & Fisher, 2020 (Madagascar)

maufei Arnold, 1920 (Zimbabwe)

mavesatra Salata & Fisher, 2020 (Madagascar)

mavohavoana Salata & Fisher, 2020 (Madagascar)

mayri Forel, 1894 (Senegal)

medioflava Donisthorpe, 1941 (New Guinea)

megacephala (Fabricius, 1793) (Aruba, Australia, Bahamas, Barbados, Bermuda, Borneo, Brazil, Cameroun, Canary Islands, Cape Verde, China, Colombia, Cook Islands, Costa Rica, Cuba, Dominican Republic, Ecuador, Egypt, Fiji, French Guiana, Gambia, Greater Antilles, Greece, Guadeloupe, Guam, Guatemala, Guinea, Guyana, Haiti, Hawaii, Honduras, India, Indonesia, Iran, Italy, Ivory Coast, Japan, Kenya, Kiribati, Lesser Antilles, Madagascar, Malaysia, Mali, Marshall Islands, Mauritius, Mayotte, Mexico, Micronesia (Federated States of), Montenegro, Mozambique, Netherlands Antilles, Netherlands Antilles, New Caledonia, New Guinea, New Zealand, Nicaragua, Nicobar Island, Niue, Northern Mariana Islands, Oman, Palau, Panama, Peru, Philippines, Puerto Rico, Republic of Macedonia, Romania, Rwanda, Réunion, Saint Lucia, Saint Martin (French part), Samoa, Saudi Arabia, Senegal, Seychelles, Sierra Leone, Singapore, Solomon Islands, Sri Lanka, Suriname, Thailand, Tokelau, Tonga, Trinidad and Tobago, Uganda, United Arab Emirates, United Kingdom of Great Britain and Northern Ireland, United States, Vanuatu, Venezuela, Vietnam, Wallis and Futuna Islands, Zimbabwe)

• Atta testacea Smith, F., 1858

• Formica edax Forskål, 1775

• Myrmica agilis Smith, F., 1857

• Myrmica laevigata Smith, F., 1855

• Myrmica suspiciosa Smith, F., 1859

• Myrmica trinodis Losana, 1834

• Oecophthora perniciosa Gerstäcker, 1859

• Oecophthora pusilla Heer, 1852

• Pheidole janus Smith, F., 1858

• Pheidole laevigata Mayr, 1862

megacephala atrox Forel, 1913 (Democratic Republic of Congo)

• Pheidole inquilina Forel Forel, 1914

megacephala costauriensis Santschi, 1914 (Ghana, Guinea)

megacephala duplex Santschi, 1937 (Angola)

megacephala ilgii Forel, 1907 (Ethiopia)

megacephala impressifrons Wasmann, 1905 (Eritrea, South Africa)

• Pheidole megacephala impressiceps Wasmann, 1904

megacephala melancholica Santschi, 1912 (Guinea, Ivory Coast)

megacephala nkomoana Forel, 1916 (Democratic Republic of Congo)

megacephala rotundata Forel, 1894 (Kenya, Mozambique, Zimbabwe)

megacephala speculifrons Stitz, 1911 (United Republic of Tanzania)

megacephala talpa Gerstäcker, 1871 (Kenya)

megatron Fischer & Fisher, 2013 (Comoros)

meihuashanensis Li & Chen, 1992 (China)

meinerti Forel, 1905 (Brazil, Ecuador, Guyana, Peru, Venezuela)

• Pheidole mimula Wheeler, W.M., 1916

meinertopsis Wilson, 2003 (Guyana, Peru)

melanogaster Donisthorpe, 1943 (New Guinea)

melastomae Wilson, 2003 (Colombia, Ecuador, Panama)

mena Salata & Fisher, 2021 (Madagascar)

mendanai Mann, 1919 (Solomon Islands)

mendicula Wheeler, W.M., 1925 (Brazil, Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Guadeloupe, Panama, Peru, Trinidad and Tobago)

mentita Santschi, 1914 (Guinea)

mentita pullata Santschi, 1914 (Guinea)

mera Wilson, 2003 (Guatemala, Mexico, United States)

merimbun Eguchi, 2001 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines)

mesomontana Longino, 2009 (Costa Rica)

metallescens Emery, 1895 (United States)

• Pheidole metallescens splendidula Wheeler, W.M., 1908

metana Wilson, 2003 (Colombia, Ecuador)

micon Wilson, 2003 (Colombia, Venezuela)

micridris Wilson, 2003 (Ecuador, Peru)

microgyna Wheeler, W.M., 1928 (Guyana)

microps Wilson, 2003 (Ecuador, Peru)

micula Wheeler, W.M., 1915 (Mexico, United States)

midas Wilson, 2003 (Ecuador, Peru, Trinidad and Tobago, Venezuela)

midongy Salata & Fisher, 2020 (Madagascar)

mikros Salata & Fisher, 2020 (Madagascar)

militicida Wheeler, W.M., 1915 (Mexico, United States)

minax Wilson, 2003 (Peru)

minensis Santschi, 1923 (Brazil)

minima Mayr, 1901 (Cameroun, Guinea, Mali)

minima catella Santschi, 1914 (Nigeria)

minima faurei Santschi, 1920 (Gabon)

minima malelana Wheeler, W.M., 1922 (Democratic Republic of Congo)

minor (Jerdon, 1851) (India)

minuscula Bernard, 1953 (Guinea, Iraq, Israel, Ivory Coast, Saudi Arabia)

minutula Mayr, 1878 (Brazil, Ecuador, French Guiana, Guyana, Panama, Peru)

• Pheidole minutula folicola Forel, 1904

• Pheidole minutula personata Santschi, 1929

mirabilis Wilson, 2003 (Colombia)

miramila Salata & Fisher, 2020 (Madagascar)

miseranda Wheeler, W.M., 1924 (Indonesia, Krakatau Islands)

mittermeieri Wilson, 2003 (Bolivia)

mivory Salata & Fisher, 2020 (Madagascar)

mixteca Wilson, 2003 (Mexico)

mjobergi Forel, 1915 (Australia)

modiglianii Emery, 1900 (Borneo, Indonesia, Malaysia)

moerens Wheeler, W.M., 1908 (Dominican Republic, Greater Antilles, Haiti, Puerto Rico, United States)

• Pheidole moerens creola Wheeler, W.M. & Mann, 1914

• Pheidole moerens dominicensis Wheeler, W.M., 1913

moffetti Wilson, 2003 (French Guiana)

monstrosa Wilson, 2003 (Brazil)

montana Eguchi, 1999 (Borneo, Indonesia, Malaysia)

monteverdensis Wilson, 2003 (Costa Rica)

moramanaensis Salata & Fisher, 2020 (Madagascar)

morelosana Wilson, 2003 (Mexico)

morrisii Forel, 1886 (United States)

• Pheidole morrisi impexa Wheeler, W.M., 1908

• Pheidole morrisii vanceae Forel, 1901

moseni Wheeler, W.M., 1925 (Brazil)

mosenopsis Wilson, 2003 (Argentina, Bolivia, Paraguay)

moskitia Longino, 2019 (Honduras)

multidens Forel, 1902 (China, India)

multispina Wilson, 2003 (Costa Rica, Panama)

muralla Longino, 2019 (Guatemala, Honduras)

mus Forel, 1902 (India, Niue, Pakistan)

musacolor Longino, 2019 (Costa Rica)

musinermis Longino, 2019 (Costa Rica, Panama)

mutisi Fernández & Wilson, 2008 (Colombia)

mylognatha Wheeler, W.M., 1922 (Democratic Republic of Congo)

N
nana Emery, 1894 (Brazil)

naoroji Forel, 1902 (India)

napoensis Wilson, 2003 (Ecuador)

nasifera Wilson, 2003 (Peru)

nasutoides Hölldobler & Wilson, 1992 (Costa Rica)

natalie Longino, 2019 (Guatemala, Honduras, Mexico, Nicaragua)

navigans Forel, 1901 (Mexico)

navoatrensis Salata & Fisher, 2020 (Madagascar)

naylae Wilson, 2003 (Cuba, Greater Antilles)

nebulosa Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole scabriventris Wilson, 2003

nemoralis Forel, 1892 (Madagascar)

neokohli Wilson, 1984 (Democratic Republic of Congo)

• Anergatides kohli Wasmann, 1915

neolongiceps Brown, 1950 (Cuba, Greater Antilles)

• Pheidole longiceps Aguayo Aguayo, 1932

neolongiscapa Özdikmen, 2010 (China)

• Pheidole longiscapa Zhou & Zheng Zhou & Zheng, 1999

neoschultzi LaPolla, 2006 (Guyana)

• Pheidole schultzi LaPolla, 2005

nephele Longino, 2019 (Honduras, Nicaragua)

nesiota Wilson, 2003 (Brazil)

nietneri Emery, 1901 (China, Pakistan, Sri Lanka)

nigella Emery, 1894 (Bolivia, Peru)

• Pheidole dimidiata Emery, 1894

nigeriensis Santschi, 1914 (Kenya, Nigeria, Uganda)

nigricula Wilson, 2003 (Costa Rica, Honduras, Nicaragua)

nigritella Bernard, 1953 (Guinea)

nimba Bernard, 1953 (Cameroun, Guinea)

nindi Mann, 1919 (Guam, Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands, Solomon Islands)

nitella Wilson, 2003 (Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Peru)

nitidicollis Emery, 1896 (Belize, Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Jamaica, Mexico, Nicaragua)

• Pheidole chalcoides Wilson, 2003

• Pheidole sagana Wheeler, W.M., 1934

nitidobruna Salata & Fisher, 2020 (Madagascar)

nitidula Emery, 1888 (Argentina, Brazil)

• Pheidole strobeli silvicola Borgmeier, 1927

njassae Viehmeyer, 1914 (United Republic of Tanzania)

njassae legitima Santschi, 1916 (United Republic of Tanzania)

• Pheidole njassae sculptior Viehmeyer, 1914

noar Wilson, 2003 (Dominican Republic, Greater Antilles)

nodgii Forel, 1905 (Indonesia, Singapore)

nodgii verlatenensis Wheeler, W.M., 1937 (Indonesia, Krakatau Islands, Singapore)

nodifera (Smith, F., 1858) (China, Thailand, Vietnam)

nodus Smith, F., 1874 (Bangladesh, India, Indonesia, Japan, Republic of Korea, Sri Lanka, Taiwan, Thailand, Vietnam)

• Pheidole nodus flebilis Santschi, 1937

• Pheidole nodus praevexata Wheeler, W.M., 1929

• Pheidole rhombinoda Mayr, 1879

• Pheidole rhombinoda formosensis Forel, 1913

• Pheidole rhombinoda micantiventris Mayr, 1897

• Pheidole rhombinoda stella Forel, 1911

• Pheidole rhombinoda taprobanae Forel, 1902

• Pheidole treubi Forel, 1905

nubicola Wilson, 2003 (Mexico)

• Pheidole cielana Wilson, 2003

• Pheidole petrensis Wilson, 2003

nubila Emery, 1906 (Argentina)

nuculiceps Wheeler, W.M., 1908 (Mexico, United States)

O
oaxacana Wilson, 2003 (Guatemala, Mexico)

obapara Casadei-Ferreira, Economo & Feitosa, 2020 (Brazil)

obnixa Forel, 1912 (Argentina)

obrima Wilson, 2003 (Venezuela)

obscurifrons Santschi, 1925 (Argentina)

obscurior Forel, 1886 (Argentina, Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Lesser Antilles, Paraguay, Trinidad and Tobago)

• Pheidole incisa evoluta Borgmeier, 1929

• Pheidole partita Mayr, 1887

obscurithorax Naves, 1985 (Argentina, Brazil, Paraguay, United States)

obturaculum Longino, 2019 (Mexico)

obtusopilosa Mayr, 1887 (Argentina, Brazil, Uruguay)

obtusospinosa Pergande, 1896 (Mexico, United States)

• Pheidole arizonica Santschi, 1911

• Pheidole subdentata Pergande, 1896

• Pheidole vasliti acolhua Wheeler, W.M., 1914

• Pheidole vasliti hirtula Forel, 1899

occipitalis André, 1890 (Guinea, Senegal, Sierra Leone)

occipitalis adami Santschi, 1939 (Guinea)

occipitalis neutralis Santschi, 1914 (Guinea)

oceanica Mayr, 1866 (Australia, Cook Islands, Fiji, French Polynesia, Guam, Kiribati, Marshall Islands, Micronesia (Federated States of), New Caledonia, New Guinea, Niue, Northern Mariana Islands, Samoa, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Pheidole oceanica boraborensis Wheeler, W.M., 1932

• Pheidole oceanica pattesoni Mann, 1919

• Pheidole oceanica upoluana Santschi, 1928

ocellata Zhou, 2001 (China)

ochracea Eguchi, 2008 (China, Vietnam)

oculata (Emery, 1899) (Madagascar)

ocypodea Salata & Fisher, 2020 (Madagascar)

oliveirai Wilson, 2003 (Colombia)

olsoni Wilson, 2003 (Costa Rica, Guatemala, Honduras, Panama)

onifera Mann, 1921 (Fiji)

onyx Wilson, 2003 (Costa Rica)

opaciventris Mayr, 1876 (Australia)

optiva Forel, 1901 (Mexico)

orbica Forel, 1893 (Guadeloupe, Puerto Rico, St. Vincent)

oswaldi Forel, 1891 (Madagascar)

otisi Wilson, 2003 (Costa Rica, Honduras)

ovalinoda Salata & Fisher, 2020 (Madagascar)

oxyops Forel, 1908 (Argentina, Brazil, Paraguay)

• Pheidole genalis Borgmeier, 1929

• Pheidole oxyops regia Forel, 1908

P
paiute Gregg, 1959 (United States)

palenquensis Wilson, 2003 (Ecuador)

pallidula (Nylander, 1849) (Afghanistan, Albania, Algeria, Andorra, Armenia, Austria, Balearic Islands, Bulgaria, Canary Islands, Croatia, France, Georgia, Gibraltar, Greece, Iberian Peninsula, Iran, Israel, Italy, Kazakhstan, Kyrgyzstan, Malta, Monaco, Montenegro, Portugal, Republic of Macedonia, Romania, Russian Federation, Slovenia, Socotra Archipelago, Spain, Turkey, Turkmenistan, Yemen)

• Oecophthora subdentata Mayr, 1853

• Pheidole pallidula emeryi Krausse, 1912

• Pheidole pallidula obscura Santschi, 1936

• Pheidole symbiotica Wasmann, 1909

• Xenoaphaenogaster inquilina Baroni Urbani, 1964

pallidula selenia Özdikmen, 2010 (Algeria)

• Pheidole pallidula inermis Stitz, 1917

pallidula tristior Schmitz, 1950 (Tunisia)

• Pheidole pallidula tasdelenia Özdikmen, 2010

• Pheidole pallidula tristis Forel, 1907

pampana Santschi, 1929 (Argentina)

paraensis Wilson, 2003 (Brazil)

paranana Santschi, 1925 (Brazil)

pararugiceps Longino, 2009 (Costa Rica)

parasitica Wilson, 1984 (India)

pariana Wilson, 2003 (Venezuela)

parva Mayr, 1865 (Borneo, China, India, Indonesia, Japan, Malaysia, Mauritius, Myanmar, Oman, Philippines, Saudi Arabia, Seychelles, Singapore, Sri Lanka, Taiwan, Thailand, United Arab Emirates, Vietnam)

• Pheidole bugi Wheeler, W.M., 1919

• Pheidole flavens farquharensis Forel, 1907

• Pheidole parva decanica Forel, 1902

• Pheidole rinae mala Forel, 1911

• Pheidole rinae tipuna Forel, 1912

• Pheidole sauteri Wheeler, W.M., 1909

• Pheidole tarda Donisthorpe, 1947

• Pheidole voeltzkowii Forel, 1894

parvicorpus Eguchi, 2001 (Borneo, Indonesia, Malaysia)

parviocula Salata & Fisher, 2020 (Madagascar)

parvula Salata & Fisher, 2020 (Madagascar)

parvulogibba Salata & Fisher, 2020 (Madagascar)

passivaeferox Longino, 2019 (Costa Rica, Nicaragua)

peckorum Wilson, 2003 (Ecuador)

pedana Wilson, 2003 (Brazil, Peru)

pegasus Sarnat, 2008 (Fiji)

peguensis Emery, 1895 (Myanmar)

pelor Wilson, 2003 (United States)

peltastes Wilson, 2003 (Brazil, Ecuador)

penetralis Smith, F., 1863 (Indonesia)

pepo Wilson, 2003 (Colombia)

peregrina Wheeler, W.M., 1916 (Brazil)

• Pheidole impariceps Santschi, 1923

• Pheidole impariceps fusciclava Santschi, 1923

perissothrix Longino, 2019 (Costa Rica, Guatemala, Honduras, Nicaragua)

perkinsi Wilson, 2003 (Dominica, Lesser Antilles)

perpilosa Wilson, 2003 (United States)

perpusilla Emery, 1894 (Brazil, Colombia, Costa Rica, Guatemala, Guyana, Mexico, Nicaragua, Panama, Trinidad and Tobago)

• Pheidole decem Forel, 1901

• Pheidole emersoni Wheeler, W.M., 1922

• Pheidole flavens breviscapa Forel, 1899

perryorum Wilson, 2003 (Brazil)

peruviana Wilson, 2003 (Ecuador, Peru)

petax Forel, 1895 (Madagascar)

phanigaster Longino, 2009 (Mexico)

philemon Forel, 1910 (Solomon Islands)

philippi Emery, 1915 (Eritrea, Ethiopia)

phipsoni Forel, 1902 (India)

pholeops Wilson, 2003 (Brazil, Ecuador, Peru)

picea (Buckley, 1867)

piceonigra Emery, 1922 (Costa Rica, Mexico)

• Pheidole picea Mayr Mayr, 1870

picobarva Longino, 2009 (Costa Rica)

pidax Wilson, 2003 (Colombia)

pieli Santschi, 1925 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Thailand, Vietnam)

• Pheidole rinae incensa Wheeler, W.M., 1928

pilifera (Roger, 1863) (Mexico, United States)

• Pheidole pennsylvanica Roger, 1863

• Pheidole pilifera artemisia Cole, 1933

• Pheidole pilifera coloradensis Emery, 1895

• Pheidole pilifera septentrionalis Wheeler, W.M., 1908

• Pheidole xerophila pacifica Wheeler, W.M., 1915

pilispina Wilson, 2003 (Panama)

piliventris (Smith, F., 1858) (Brazil)

pilosior Wilson, 2003 (United States)

pinealis Wheeler, W.M., 1908 (Mexico, United States)

pinicola Wilson, 2003 (Mexico)

plagiaria Smith, F., 1860 (Borneo, Brunei Darussalam, China, Indonesia, Krakatau Islands, Laos, Malaysia, Myanmar, Philippines, Singapore, Thailand, Vietnam)

• Pheidole divergens Mayr, 1867

• Pheidole peguensis yomensis Forel, 1903

plagiaria moica Forel, 1911 (Vietnam)

plagiaria palawanica Stitz, 1925 (Philippines)

plagiaria rectilineata Viehmeyer, 1916 (Indonesia, Sulawesi)

planidorsum Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines, Vietnam)

planifrons Santschi, 1920 (Thailand, Vietnam)

plato Wilson, 2003 (Ecuador)

platypus Crawley, 1915 (Australia)

platyscapa Longino, 2019 (Nicaragua)

plebecula Forel, 1899 (Costa Rica, Honduras, Mexico, Nicaragua, Panama)

• Pheidole perdiligens Wilson, 2003

• Pheidole texticeps Wilson, 2003

plinii Forel, 1911 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore)

podargea Salata & Fisher, 2020 (Madagascar)

polita Emery, 1894 (Bolivia, Brazil, Peru)

• Pheidole wheeleri Mann, 1916

polymorpha Wilson, 2003 (Mexico)

porcula Wheeler, W.M., 1908 (Mexico, United States)

poringensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

portalensis Wilson, 2003 (Mexico, United States)

potosiana Wilson, 2003 (Costa Rica, Mexico)

praegrandis Salata & Fisher, 2020 (Madagascar)

praeses Wilson, 2003 (Brazil)

praeusta Roger, 1863 (Colombia)

• Pheidole amata Forel, 1901

prattorum Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

prelli Forel, 1911 (Kenya)

prelli ingenita Santschi, 1928 (Ivory Coast)

prelli redbankensis Forel, 1913 (Mozambique, Zimbabwe)

probolonotum Longino, 2019 (Mexico)

pronotalis Forel, 1902 (India, Pakistan, Sri Lanka)

prostrata Wilson, 2003 (Costa Rica, Panama)

protaxi Oliveira & Lacau, 2015 (Brazil)

protea Forel, 1912 (Indonesia, Thailand)

protensa Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

providens (Sykes, 1835) (India, Iran)

proxima Mayr, 1876 (Australia, New Zealand)

proxima transversa Forel, 1902 (Australia)

psammophila Creighton & Gregg, 1955 (Mexico, United States)

psilogaster Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

pubiventris Mayr, 1887 (Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Mexico)

• Pheidole indistincta Forel, 1899

• Pheidole pubiventris cearensis Forel, 1901

• Pheidole pubiventris nevadensis Forel, 1901

• Pheidole pubiventris timmii Forel, 1901

• Pheidole pubiventris variegata Emery, 1896

pugnax Dalla Torre, 1892 (Costa Rica, El Salvador, Honduras, Panama)

• Pheidole radowszkowskii militaris Emery, 1890

pulchella Santschi, 1910 (Cameroun, Congo, Kenya)

• Pheidole niapuana Wheeler, W.M., 1922

• Pheidole pulchella achantella Santschi, 1939

pullula Santschi, 1911 (Argentina)

punctatissima Mayr, 1870 (Belize, Colombia, Costa Rica, Denmark, Guatemala, Honduras, Mexico, Nicaragua, Panama, United Kingdom of Great Britain and Northern Ireland)

• Pheidole punctatissima napaea Wheeler, W.M., 1934

punctithorax Borgmeier, 1929 (Brazil)

punctulata Mayr, 1866 (Comoros, Eritrea, Guinea, Ivory Coast, Kenya, Madagascar, Mayotte, Mozambique, Namibia, Seychelles, South Africa, Zimbabwe)

punctulata subatrox Santschi, 1937 (Democratic Republic of Congo)

purpurascens Emery, 1897 (New Guinea)

purpurea Longino, 2009 (Guatemala, Honduras, Mexico)

puttemansi Forel, 1911 (Brazil)

• Pheidole fallax janeirensis Forel, 1912

pygmaea Wilson, 2003 (Colombia)

Q
quadrensis Forel, 1900 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

quadriceps Wilson, 2003 (Colombia)

quadricuspis Emery, 1900 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Thailand)

quadriprojecta Smith, M.R., 1947 (New Guinea)

quadrispinosa (Smith, F., 1865) (Indonesia, New Guinea)

• Pheidolacanthinus armatus Smith, F., 1865

• Pheidole smithi Emery, 1887

quercicola Wilson, 2003 (Mexico)

quiaccana Wheeler, W.M., 1925 (Bolivia)

quinata Eguchi, 2000 (Borneo, Indonesia, Malaysia)

R
rabo Forel, 1913 (Borneo, China, Indonesia, Malaysia, Philippines, Thailand, Vietnam)

• Pheidole concinna Wheeler, W.M. Wheeler, W.M., 1928

• Pheidole tsailuni Wheeler, W.M., 1929

radoszkowskii Mayr, 1884 (Argentina, Bolivia, Brazil, Colombia, Costa Rica, French Guiana, Greater Antilles, Guatemala, Guyana, Honduras, Jamaica, Lesser Antilles, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Venezuela)

• Pheidole medialis Wilson, 2003

• Pheidole radoszkowskii luteola Forel, 1893

• Pheidole radoszkowskii opacissima Forel, 1901

• Pheidole radoszkowskii parvinoda Forel, 1912

• Pheidole radowszkowskii acuta Emery, 1894

• Pheidole radowszkowskii australis Emery, 1890

ragnax Fischer & Fisher, 2013 (Mayotte)

ranohirensis Salata & Fisher, 2020 (Madagascar)

rasnitsyni Dubovikoff, 2011

rebeccae Fischer, Hita Garcia & Peters, 2012 (Ivory Coast)

reclusi Forel, 1899 (Colombia)

recondita Clouse, 2007 (Guam, Marshall Islands, Micronesia (Federated States of), Northern Mariana Islands)

rectisentis Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua)

rectispina Wilson, 2003 (Belize, Costa Rica, Guatemala, Honduras, Mexico)

rectitrudis Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama)

reflexans Santschi, 1933 (Argentina, Paraguay)

reichenspergeri Santschi, 1923 (Brazil, Colombia)

renae Wilson, 2003 (Costa Rica)

renirano Salata & Fisher, 2020 (Madagascar)

retivertex Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

retronitens Santschi, 1930 (Democratic Republic of Congo)

rhea Wheeler, W.M., 1908 (Mexico, United States)

rhinoceros Forel, 1899 (Colombia, Costa Rica, Honduras, Mexico, Nicaragua, Panama)

rhinomontana Longino, 2009 (Costa Rica, Honduras, Nicaragua)

rhytifera Wilson, 2003 (Venezuela)

rima Longino, 2019 (Mexico)

rinae Emery, 1900 (Borneo, Indonesia, Malaysia, Singapore, Thailand, Vietnam)

risii Forel, 1892 (Argentina, Brazil)

riveti Santschi, 1911 (Colombia, Ecuador)

roberti Forel, 1902 (China, India, Pakistan)

rochai Forel, 1912 (Brazil)

rogeri Emery, 1896 (Costa Rica, Honduras)

rogeripolita Longino, 2019 (Costa Rica, Panama)

rogersi Forel, 1902 (Bangladesh, India, Sri Lanka)

rogersi taylori Forel, 1902 (India)

rohani Santschi, 1925 (Angola)

rohani monardi Santschi, 1930 (Angola)

rohani pellax Santschi, 1930 (Angola)

roosevelti Mann, 1921 (Fiji)

rosae Forel, 1901 (Argentina, Brazil)

• Pheidole silvestrii Emery, 1906

rosula Wilson, 2003 (Argentina)

rotundiceps Wilson, 2003 (Ecuador)

roushae Wilson, 2003 (Belize, Guatemala, Honduras, Mexico)

rubiceps Wilson, 2003 (Venezuela)

rudigenis Emery, 1906 (Argentina, Brazil, Paraguay)

• Pheidole bucolica Santschi, 1929

ruficeps (Smith, F., 1861) (Indonesia, Sulawesi)

• Pheidole mordax Smith, F., 1863

rufipilis Forel, 1908 (Brazil)

• Pheidole pubiventris impia Santschi, 1923

rugaticeps Emery, 1877 (Burkina Faso, Eritrea, Ethiopia, Mali, Oman, Saudi Arabia, Senegal, South Africa)

rugaticeps arabs Emery, 1881 (Eritrea, Yemen)

rugatula Santschi, 1933 (Argentina, Paraguay)

rugiceps Wilson, 2003 (Belize, Brazil, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Peru, Trinidad and Tobago)

rugifera Eguchi, 2001 (Borneo, Indonesia, Malaysia, Singapore)

rugithorax Eguchi, 2008 (Vietnam)

rugocephala Salata & Fisher, 2020 (Madagascar)

rugofitarata Salata & Fisher, 2020 (Madagascar)

rugosa Smith, F., 1858 (Sri Lanka)

rugosula Forel, 1902 (Australia, New Zealand)

• Pheidole proxima bombalensis Forel, 1910

rugulosa Gregg, 1959 (Mexico, United States)

rutilana Wilson, 2003 (Brazil)

ryukyuensis Ogata, 1982 (Japan)

S
sabahna Eguchi, 2000 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

sabella Wilson, 2003 (Ecuador, Peru)

sabina Wilson, 2003 (Mexico)

sagax Wilson, 2003 (Colombia, Ecuador, Nicaragua, Venezuela)

sagei Forel, 1902 (China, India, Nepal, Pakistan)

sagittaria Wilson, 2003 (Costa Rica)

sarae (Sharaf, 2018) (Oman, Saudi Arabia)

sarawakana Forel, 1911 (Borneo, Indonesia, Malaysia, Philippines, Thailand)

sarcina Forel, 1912 (Brazil, Ecuador)

sarpedon Wilson, 2003 (Ecuador)

sauberi Forel, 1905 (Borneo, Indonesia, Krakatau Islands, Malaysia, New Guinea, Philippines, Thailand)

sava Salata & Fisher, 2020 (Madagascar)

savegre Longino, 2019 (Costa Rica)

saxicola Wheeler, W.M., 1922 (Democratic Republic of Congo)

sayapensis Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

scabrata Forel, 1895 (Madagascar)

scabriuscula Gerstäcker, 1871 (Kenya)

scapulata Santschi, 1923 (Argentina, Bolivia, Paraguay)

schmalzi Emery, 1894 (Bolivia, Brazil, Venezuela)

• Pheidole rata Borgmeier, 1929

• Pheidole rehi Forel, 1900

schoedli Eguchi, Hashimoto & Malsch, 2006 (Borneo, Indonesia, Malaysia)

schoutedeni Forel, 1913 (Democratic Republic of Congo)

schoutedeni platycephala Stitz, 1916 (Democratic Republic of Congo)

schultzei Forel, 1910 (Botswana)

schultzei ebangana Santschi, 1937 (Angola)

schultzei gwaaiensis Forel, 1913 (Zambia, Zimbabwe)

schultzei woodvalensis Arnold, 1960 (Zimbabwe)

schwarzmaieri Borgmeier, 1939 (Argentina, Brazil)

sciara Cole, 1955 (Mexico, United States)

scimitara Wilson, 2003 (Brazil, Peru)

sciophila Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole proserpina Wheeler, W.M., 1908

• Pheidole sciophila semilaevicephala Smith, M.R., 1934

scolioceps Wilson, 2003 (Brazil, Colombia, Ecuador, Guyana, Peru)

scrobifera Emery, 1896 (Costa Rica, Honduras, Nicaragua, Panama)

sculptior Forel, 1893 (Barbados, Colombia, Grenada, Guadeloupe, Netherlands Antilles, Puerto Rico, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole tayrona Wilson, 2003

sculpturata Mayr, 1866 (Guinea, Iran, Ivory Coast, Kenya, Namibia, Oman, Saudi Arabia, South Africa, United Arab Emirates)

sculpturata areolata Forel, 1911 (South Africa)

sculpturata berthoudi Forel, 1894 (South Africa)

sculpturata dignata Santschi, 1915 (Congo)

sculpturata particeps Santschi, 1921 (United Republic of Tanzania)

sculpturata rhodesiana Forel, 1913 (Zimbabwe)

sculpturata welgelegenensis Forel, 1913 (Democratic Republic of Congo)

sculpturata zambesiana Forel, 1913 (Zambia, Zimbabwe)

sebofila Longino, 2009 (Costa Rica)

securigera Wilson, 2003 (Colombia)

seeldrayersi Forel, 1910 (Argentina, Bolivia, Colombia)

selathorax Zhou, 2001 (China)

seligmanni Wilson, 2003 (Colombia)

semidea Fischer, Hita Garcia & Peters, 2012 (Nigeria)

semilaevis Forel, 1901 (Brazil)

senex Gregg, 1952 (Mexico, United States)

• Pheidole pilifera anfracta Cole, 1952

senilis Santschi, 1929 (Brazil)

sensipelada Longino, 2019 (Costa Rica)

sensitiva Borgmeier, 1959 (Brazil, Colombia, Costa Rica, Honduras, Nicaragua, Panama, Peru)

sepulchralis Bingham, 1903 (India)

sepultura Longino, 2019 (Mexico)

sericella Viehmeyer, 1914 (New Guinea)

servilia Wilson, 2003 (Colombia)

setosa Fischer, Hita Garcia & Peters, 2012 (Democratic Republic of Congo)

setsukoae Wilson, 2003 (Mexico)

severini Forel, 1904 (French Guiana, Suriname)

sexdentata Donisthorpe, 1948 (New Guinea)

sexspinosa Mayr, 1870 (Cook Islands, French Polynesia, Micronesia (Federated States of), New Guinea, Niue, Palau, Samoa, Singapore, Solomon Islands, Tokelau, Tuvalu, Vanuatu, Wallis and Futuna Islands)

• Pheidole sexspinosa adamsoni Wheeler, W.M., 1932

sexspinosa biroi Emery, 1900 (New Guinea)

sexspinosa fuscescens Emery, 1900 (New Guinea, Solomon Islands)

sharpi Forel, 1902 (India, Sri Lanka)

sharpi hoogwerfi Forel, 1902 (India)

sicaria Wilson, 2003 (Colombia, Costa Rica)

sigillata Wilson, 2003 (Brazil, Paraguay)

sikorae Forel, 1891 (Madagascar)

similigena Wheeler, W.M., 1937 (Cuba)

similis Salata & Fisher, 2020 (Madagascar)

simoni Emery, 1893 (Philippines)

simonsi Wilson, 2003 (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama)

• Pheidole arctos Wilson, 2003

• Pheidole gangamon Wilson, 2003

• Pheidole thrasys Wilson, 2003

simplex Wheeler, W.M., 1925 (Brazil, Peru)

simplispinosa Sarnat, 2008 (Fiji)

sinaitica Mayr, 1862 (Egypt, Eritrea, Iran, Iraq, Israel, Kenya, Saudi Arabia, United Arab Emirates)

sinaitica santschii Forel, 1907 (Tunisia)

singaporensis Özdikmen, 2010 (India, Pakistan, Philippines, Singapore)

• Myrmica longipes Smith, F., 1857

singaporensis conicollis (Emery, 1900) (Indonesia, Singapore)

singaporensis continentis (Forel, 1911) (Myanmar, Singapore)

singaporensis pseudola Forel, 1915 (Indonesia)

singularis Smith, F., 1863 (Indonesia, New Guinea)

sinica (Wu, J. & Wang, C., 1992) (China)

sitiens Wilson, 2003 (United States)

skwarrae Wheeler, W.M., 1934 (Mexico)

smythiesii Forel, 1902 (China, India, Nepal, Thailand, Vietnam)

• Pheidole bhavanae Bingham, 1903

• Pheidole smythiesii bengalensis Forel, 1902

socrates Forel, 1912 (Brazil, French Guiana, Guyana, Suriname)

soesilae Makhan, 2007 (Suriname)

sofia Salata & Fisher, 2020 (Madagascar)

soritis Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole sitarches Wheeler, W.M., 1908

• Pheidole sitarches campestris Creighton, 1950

• Pheidole sitarches rufescens Wheeler, W.M., 1908

• Pheidole sitarches transvarians Wheeler, W.M., 1908

sospes Forel, 1908 (Argentina, Brazil, Colombia, Ecuador, Peru)

• Pheidole minutissima Kusnezov, 1952

spadonia Wheeler, W.M., 1915 (Mexico, United States)

sparsa Salata & Fisher, 2020 (Madagascar)

sparsisculpta Longino, 2009 (Costa Rica)

spathicornis Wilson, 2003 (Mexico)

spathifera Forel, 1902 (Bangladesh, China, India, Sri Lanka, Thailand)

spathifera aspatha Forel, 1902 (India)

spathifera yerburyi Forel, 1902 (India, Myanmar, Sri Lanka)

spathipilosa Wilson, 2003 (Costa Rica, Nicaragua)

specularis Wilson, 2003 (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

speculifera Emery, 1877 (Cameroun, Eritrea, Ethiopia, Guinea, Kenya, Mali)

speculifera ascara Emery, 1901 (Eritrea, Ethiopia, Kenya)

speculifera bispecula Santschi, 1930 (Democratic Republic of Congo)

speculifera cubangensis Forel, 1901 (Angola)

sperata Forel, 1915 (Indonesia)

sphaerica Wilson, 2003 (Colombia)

spilota Wilson, 2003 (Colombia)

spinicornis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

spininodis Mayr, 1887 (Argentina, Brazil, Paraguay)

• Pheidole hohenlohei Emery, 1888

• Pheidole spielbergi Emery, 1888

• Pheidole spininodis bruta Santschi, 1934

• Pheidole spininodis lucifuga Santschi, 1923

• Pheidole spininodis pencosensis Forel, 1914

• Pheidole spininodis solaris Santschi, 1929

spinoda (Smith, F., 1858) (Australia)

spinosa Forel, 1891 (Comoros, Madagascar, Mayotte, Seychelles)

• Pheidole decepticon Fischer & Fisher, 2013

• Pheidole megacephala scabrior Forel, 1891

• Pheidole picata Forel, 1891

• Pheidole picata bernhardae Emery, 1915

• Pheidole picata gietleni Forel, 1905

spinulosa Forel, 1910 (Lesotho, Zimbabwe)

spinulosa conigera Forel, 1910 (South Africa)

spinulosa messalina Forel, 1910 (Lesotho)

squalida Santschi, 1910 (Congo)

steinheili Forel, 1901 (Colombia)

stigma Wilson, 2003 (Ecuador)

stomachosa Wheeler, W.M., 1917 (Jamaica)

strator Forel, 1910 (Eritrea, Ethiopia, Kenya)

strator fugax Arnold, 1920 (Mozambique)

strator tabida Menozzi, 1933 (Uganda)

striata Donisthorpe, 1947 (New Guinea)

striaticeps Mayr, 1870 (Costa Rica, Guatemala, Honduras, India, Mexico)

• Pheidole goeldii chloe Forel, 1908

strigosa Wilson, 2003 (Brazil)

strobeli Emery, 1906 (Argentina, Brazil, Uruguay)

• Pheidole nitidula daguerrei Santschi, 1931

• Pheidole perversa Forel, 1908

• Pheidole perversa richteri Forel, 1909

• Pheidole rufipilis divexa Forel, 1908

• Pheidole strobeli misera Santschi, 1916

stulta Forel, 1886 (Brazil, Costa Rica, Panama)

• Pheidole championi Forel, 1899

• Pheidole championii sima Forel, 1912

styrax Wilson, 2003 (Venezuela)

subaberrans (Kusnezov, 1952) (Argentina)

subarmata Mayr, 1884 (Argentina, Bahamas, Barbados, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, French Guiana, Greater Antilles, Grenada, Guadeloupe, Guatemala, Haiti, Honduras, Mexico, Panama, Paraguay, Puerto Rico, St. Vincent, Trinidad and Tobago)

• Pheidole cornutula Emery, 1890

• Pheidole cornutula dentimentum Santschi, 1929

• Pheidole cornutula imbecillis Emery, 1906

• Pheidole hondurensis Mann, 1922

• Pheidole subarmata borinquenensis Wheeler, W.M., 1908

• Pheidole subarmata elongatula Forel, 1893

• Pheidole subarmata nassavensis Wheeler, W.M., 1905

• Pheidole subarmata nefasta Santschi, 1929

submonticola Eguchi, 2001 (Borneo, Indonesia, Malaysia)

subnuda Wilson, 2003 (Colombia, Venezuela)

subreticulata Emery, 1894 (Brazil)

subsphaerica Wilson, 2003 (Colombia)

sulcaticeps Roger, 1863 (Bangladesh, China, India, Pakistan, Sri Lanka)

sulcaticeps punensis Forel, 1902 (India)

sulcaticeps vellicans Forel, 1911 (Sri Lanka)

superba Wilson, 2003 (Colombia, Panama)

susannae Forel, 1886 (Argentina, Barbados, Brazil, Colombia, Costa Rica, Dominican Republic, El Salvador, Guatemala, Honduras, Mexico, Netherlands Antilles, Nicaragua, Paraguay, Puerto Rico, Trinidad and Tobago, Venezuela)

• Pheidole susannae atricolor Forel, 1901

susanowo Onoyama & Terayama, 1999 (Japan)

sykesii Forel, 1902 (India)

synanthropica Longino, 2009 (Costa Rica, Honduras, Mexico, Nicaragua)

synarmata Wilson, 2003 (Brazil, Costa Rica, El Salvador, Guyana, Honduras, Nicaragua, Venezuela)

T
tachigaliae Wheeler, W.M., 1921 (French Guiana, Guyana)

tachirana Wilson, 2003 (Venezuela)

taipoana Wheeler, W.M., 1928 (China, Vietnam)

taivanensis Forel, 1912 (Taiwan, Thailand)

tambopatae Wilson, 2003 (Brazil, Colombia, Peru)

tampony Salata & Fisher, 2020 (Madagascar)

tandjongensis Forel, 1913 (China, Indonesia, Singapore, Thailand, Vietnam)

tanyscapa Wilson, 2003 (Costa Rica, Nicaragua, Panama)

tapanti Longino, 2019 (Costa Rica)

tarchon Wilson, 2003 (Argentina)

tasmaniensis Mayr, 1866 (Australia)

tasmaniensis continentis Forel, 1902 (Australia)

taurus Emery, 1906 (Argentina)

tawauensis Eguchi, 2001 (Borneo, Indonesia, Malaysia)

templaria Forel, 1902 (India)

templaria euscrobata Forel, 1913 (Sri Lanka)

tenebricosa Eguchi, 2001 (Borneo, Indonesia, Malaysia, Philippines)

tenebrovulgaris Salata & Fisher, 2020 (Madagascar)

tenerescens Wheeler, W.M., 1922 (Trinidad and Tobago)

tennantae Wilson, 2003 (Colombia, Costa Rica, Panama, Peru, Venezuela)

tenuicephala Longino, 2009 (Costa Rica)

tenuiclavata Donisthorpe, 1943 (New Guinea)

tenuinodis Mayr, 1901 (Cameroun, Namibia, South Africa)

tenuinodis bothae Forel, 1901 (South Africa)

tenuinodis robusta Stitz, 1923 (Namibia)

tenuinodis sipapomae Arnold, 1920 (Zimbabwe)

tenuis Wilson, 2003 (Colombia)

tepicana Pergande, 1896 (Mexico, United States)

• Pheidole carbonaria Pergande, 1896

• Pheidole kingi André, 1898

• Pheidole kingi instabilis Emery, 1901

• Pheidole kingi torpescens Wheeler, W.M., 1915

• Pheidole rugifrons Pergande, 1896

• Pheidole townsendi André, 1898

tepuicola Wilson, 2003 (Venezuela)

termitobia Forel, 1901 (Brazil)

termitophila Forel, 1904 (Guinea, Liberia, Mali, Senegal, Sudan)

termitophila liberiensis Forel, 1911 (Liberia)

terraceensis Bharti, 2001 (India)

terresi Wheeler, W.M. & Mann, 1914 (Dominican Republic, Greater Antilles, Haiti)

• Pheidole terresi illota Wheeler, W.M. & Mann, 1914

terribilis Wilson, 2003 (Bolivia, Brazil, Suriname)

tetra Creighton, 1950 (Mexico, United States)

tetracantha Emery, 1897 (New Guinea)

tetrica Forel, 1913 (Brazil)

tetroides Wilson, 2003 (Mexico)

texana Wheeler, W.M., 1903 (United States)

tigris Wilson, 2003 (Colombia, Panama)

tijucana Borgmeier, 1927 (Brazil)

tikal Longino, 2019 (Guatemala)

tillandsiarum Wheeler, W.M., 1934 (Mexico)

tinamu Longino, 2019 (Costa Rica)

tisiphone Wheeler, W.M., 1911 (Guatemala, Mexico)

titanis Wheeler, W.M., 1903 (Mexico, United States)

tjibodana Forel, 1905 (Borneo, Brunei Darussalam, Cambodia, China, Indonesia, Malaysia, Philippines, Singapore, Thailand, Vietnam)

tobini Wilson, 2003 (Ecuador, Peru)

tolteca Forel, 1901 (Mexico)

torosa Wilson, 2003 (Mexico)

trachyderma Emery, 1906 (Argentina)

trageri Wilson, 2003 (Brazil)

tragica Wheeler, W.M., 1934 (Mexico)

traini Wilson, 2003 (Costa Rica, Panama)

transfigens Forel, 1911 (New Guinea)

transversostriata Mayr, 1887 (Barbados, Brazil, Colombia, Costa Rica, Ecuador, Guyana, Nicaragua, Suriname, Trinidad and Tobago, Venezuela)

• Pheidole lacerta Wheeler, W.M., 1922

• Pheidole scalaris Wilson, 2003

• Pheidole transversostriata nigridens Forel, 1901

trapezoidea Viehmeyer, 1914 (Australia)

tricarinata Santschi, 1914 (Guinea)

trichotos Salata & Fisher, 2020 (Madagascar)

tricolor Donisthorpe, 1949 (New Guinea)

triconstricta Forel, 1886 (Argentina, Bolivia, Brazil, Paraguay, Uruguay)

• Pheidole radoszkowskii discursans Forel, 1912

• Pheidole radoszkowskii saviozae Forel, 1911

• Pheidole triconstricta ambulans Emery, 1906

• Pheidole triconstricta hebe Santschi, 1923

• Pheidole triconstricta rosariensis Forel, 1913

trinitatis Wilson, 2003 (Trinidad and Tobago)

triplex Wilson, 2003 (Ecuador, Trinidad and Tobago)

tristis (Smith, F., 1858) (Argentina, Brazil, Paraguay)

• Atta fumipennis Smith, F., 1858

• Atta rubra Smith, F., 1858

• Pheidole crassipes Mayr, 1887

• Pheidole emeryi Mayr, 1887

• Pheidole emeryi alsia Santschi, 1923

tristops Wilson, 2003 (Colombia)

truncula Wilson, 2003 (Costa Rica)

tsaravoniana Salata & Fisher, 2020 (Madagascar)

tschinkeli Wilson, 2003 (Guatemala, Honduras, Mexico, Nicaragua, Venezuela)

tsontekonwei Longino, 2019 (Mexico)

tuculutan Longino, 2019 (Guatemala)

tumida Eguchi, 2008 (China, Laos, Vietnam)

turneri Forel, 1902 (Australia)

tuxtlasana Wilson, 2003 (Guatemala, Honduras, Mexico)

typhlos Salata & Fisher, 2020 (Madagascar)

tysoni Forel, 1901 (Mexico, United States)

U
ulothrix Wilson, 2003 (Costa Rica, Panama)

ululevu Fischer, Sarnat & Economo, 2016 (Cook Islands, Fiji)

umbonata Mayr, 1870 (Fiji, French Polynesia, Guam, Kiribati, Marshall Islands, Micronesia (Federated States of), New Caledonia, Niue, Northern Mariana Islands, Palau, Samoa, Singapore, Solomon Islands, Tokelau, Tonga, Vanuatu, Wallis and Futuna Islands)

• Pheidole umbonata zimmermani Wheeler, W.M., 1936

umbonata fusciventris Emery, 1900 (New Guinea)

umphreyi Wilson, 2003 (Costa Rica, Panama)

uncagena Sarnat, 2008 (Fiji)

unicornis Wilson, 2003 (Colombia, Ecuador)

upeneci Forel, 1913 (Borneo, Indonesia, Malaysia)

uranus Salata & Fisher, 2020 (Madagascar)

urbana Camargo-Vanegas & Guerrero, 2020 (Colombia)

ursus Mayr, 1870 (Belize, Costa Rica, Guatemala, Honduras, Mexico)

• Macromischa cressoni André, 1887

• Pheidole ursus gracilinoda Forel, 1904

V
vadum Salata & Fisher, 2020 (Madagascar)

vafella Wheeler, W.M., 1925 (Bolivia)

vafra Santschi, 1923 (Argentina, Brazil, Colombia, Costa Rica, French Guiana, Nicaragua, Panama, Paraguay, Trinidad and Tobago, Venezuela)

valens Wilson, 2003 (Brazil)

vallicola Wheeler, W.M., 1915 (Mexico, United States)

vallifica Forel, 1901 (Brazil, Colombia, Costa Rica, Honduras, Paraguay)

vanderveldi Forel, 1913 (Democratic Republic of Congo)

variabilis Mayr, 1876 (Australia)

• Pheidole variabilis rugocciput Forel, 1902

variabilis aliena Emery, 1914 (New Caledonia)

variabilis croceithorax Emery, 1914 (New Caledonia)

variabilis mediofusca Forel, 1902 (Australia)

variabilis ocior Forel, 1915 (Australia)

variabilis ocyma Forel, 1915 (Australia)

variabilis parvispina Forel, 1902 (Australia)

variabilis praedo Forel, 1902 (Australia)

variabilis redunca Crawley, 1915 (Australia)

variolosa Emery, 1892 (Eritrea, Somalia)

vatovavensis Salata & Fisher, 2020 (Madagascar)

vatu Mann, 1921 (Fiji)

veletis Wilson, 2003 (Bolivia, Brazil, Panama)

velox Emery, 1887 (Indonesia)

venatrix Wilson, 2003 (Colombia)

verricula Wilson, 2003 (Costa Rica, Honduras)

vestita Wilson, 2003 (Costa Rica)

veteratrix Forel, 1891 (Madagascar)

• Pheidole veteratrix angustinoda Forel, 1892

victima Santschi, 1929 (Brazil)

victoris Forel, 1913 (Zimbabwe)

vieti Eguchi, 2008 (China, Vietnam)

vigilans (Smith, F., 1858) (Australia, New Zealand, Norfolk Island)

• Pheidole ampla norfolkensis Wheeler, W.M., 1927

• Pheidole ampla parallela Forel, 1902

• Pheidole ampla yarrensis Forel, 1902

• Pheidole dolichocephala André, 1896

violacea Wilson, 2003 (Costa Rica)

virago Wheeler, W.M., 1915 (United States)

viriosa Wilson, 2003 (Ecuador)

viserion Sarnat, Fischer & Economo, 2016 (New Guinea)

vistana Forel, 1914 (Mexico, United States)

• Pheidole grallipes Wheeler, W.M., 1916

• Pheidole susannae longipes Pergande, 1896

voasara Salata & Fisher, 2020 (Madagascar)

vohemarensis Salata & Fisher, 2020 (Madagascar)

volontany Salata & Fisher, 2020 (Madagascar)

vomer Wilson, 2003 (Colombia)

vony Salata & Fisher, 2020 (Madagascar)

vorax (Fabricius, 1804) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Trinidad and Tobago)

• Pheidole cephalica Smith, F., 1858

• Pheidole opaca Mayr, 1862

• Pheidole opaca apterostigmoides Weber, 1943

• Pheidole opaca incrustata Forel, 1908

• Pheidole opaca sarrita Forel, 1908

voreios Salata & Fisher, 2020 (Madagascar)

vulcan Fischer & Fisher, 2013 (Comoros)

vulgaris Eguchi, 2006 (China, India, Thailand, Vietnam)

W
walkeri Mann, 1922 (Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)

• Pheidole arietans Wilson, 2003

• Pheidole glyphoderma Wilson, 2003

• Pheidole triumbonata Wilson, 2003

wallacei Mann, 1916 (Brazil)

wardi Wilson, 2003 (Belize, Guatemala, Honduras, Mexico)

watsoni Forel, 1902 (Bangladesh, China, India, Myanmar, Nicobar Island, Sri Lanka)

weiseri Santschi, 1923 (Argentina)

wheelerorum Mackay, W.P., 1988 (United States)

wiesei Forel, 1910 (Australia)

williamsi Wheeler, W.M., 1919 (Ecuador, Galapagos Islands)

• Pheidole williamsi seymourensis Wheeler, W.M., 1924

wilsoni Mann, 1921 (Fiji)

wolfringi Forel, 1908 (Brazil)

woodmasoni Forel, 1885 (Bangladesh, India, Sri Lanka)

wroughtonii Forel, 1902 (India)

X
xanthocnemis Emery, 1914 (New Caledonia)

xanthogaster Wilson, 2003 (Colombia, Ecuador, Peru)

xerophila Wheeler, W.M., 1908 (Mexico, United States)

• Pheidole xerophila tucsonica Wheeler, W.M., 1908

xiloa Longino, 2019 (Nicaragua)

xocensis Forel, 1913 (Zimbabwe)

xocensis bulawayensis Forel, 1913 (Zimbabwe)

xyston Wilson, 2003 (Guatemala, Mexico)

Y
yaqui Creighton & Gregg, 1955 (Mexico, United States)

yeensis Forel, 1902 (China, Laos, Myanmar, Thailand, Vietnam)

yucatana Wilson, 2003 (Mexico)

Z
zannia Longino, 2019 (Guatemala, Honduras)

zavamanira Salata & Fisher, 2020 (Madagascar)

zelata Wilson, 2003 (Bolivia)

zeteki Smith, M.R., 1947 (Ecuador, Panama, Peru, Trinidad and Tobago)

zhoushanensis Li & Chen, 1992 (China)

zirafy Salata & Fisher, 2020 (Madagascar)

zoceana Santschi, 1925 (China, Thailand, Vietnam)

zoster Wilson, 2003 (Brazil, Peru)

=Fossil Species=

A
anticua Casadei-Ferreira, Chaul & Feitosa, 2019

C
cordata (Holl, 1829)

P
pauchil Varela-Hernández & Riquelme, 2021

primigenia Baroni Urbani, 1995

T
tertiaria Carpenter, 1930

tethepa Wilson, 1985