Aenictus sagei

Jaitrong et al. (2010) - Aenictus sagei is known from the type series from Dharmasala (Forel, 1901), and additional material collected in Solon [?Solan] (1,380-1,400 m alt.), north-central India by L. Weatherill who observed workers preying on other small ants such as Paratrechina and Plagiolepis (Wilson, 1964). These prey ants are in size almost the same as the prey of A. camposi observed in Malay Peninsula reported by Rosciszewski and Maschwitz (1994).

Pisarski (1967) also notes that numerous specimens were collected by Dr. Knut Lindberg (1892 - 1962, Lund, Sweden) from Tour Kham [?Torkham], on the border with Pakistan, in the vicinity of Peshawar, Afghanistan.

Identification
A member of the wroughtonii group. This hairy species is closely related to Aenictus wroughtonii, but is slighty larger than the latter. The pronotal hairs are longer in A. sagei (0.20-0.25 mm) than in A.wroughtonii (0.10-0.13 mm). The antennal scape is short, shorter than head length with SI 100 in the former, while it reaches posterior margin of head with SI 111-122 in the latter. The subpetiolar process is weakly developed, with its anteroventral corner obtusely angulate in the former, while it is almost absent and unarmed in the latter. Aenictus sagei is also similar to Aenictus artipus, in which, however, the subpetiolar process is recognizable, with its anteroventral corner more clearly angulated. The antennal scape is longer in A. artipus (extending beyond posterolateral corner of head), while it does not exceed the posterior corner in A. sagei. (Jaitrong et al. 2010)

Distribution
North-central India

Distribution based on Regional Taxon Lists
Oriental Region: India, Nepal, Pakistan. Palaearctic Region: Afghanistan.

Castes
Known only from the worker caste.

Nomenclature

 * . Aenictus wroughtonii var. sagei Forel, 1901a: 469 (w.) INDIA (Himachal Pradesh).
 * Type-material: lectotype worker (by designation of Jaitrong, et al. 2010: 40), 5 paralectotype workers.
 * Type-locality: lectotype India: Dharmsala (Sage); paralectotypes with same data.
 * Type-depository: MHNG.
 * Subspecies of wroughtonii: Bingham, 1903: 17; Forel, 1906b: 90; Emery, 1910b: 30; Wheeler, W.M. 1927h: 83; Menozzi, 1939a: 327; Chapman & Capco, 1951: 12; Pisarski, 1967: 377 (redescription).
 * Status as species: Wilson, 1964a: 477; Dlussky, Soyunov & Zabelin, 1990: 180 (in key); Bolton, 1995b: 60; Jaitrong, et al. 2010: 40 (redescription); Bharti, Wachkoo & Kumar, 2012: 294 (in key); Sharaf, Aldawood & El Hawagry, 2012b: 47 (in key); Bharti, Guénard, et al. 2016: 21; Jaitrong & Ruangsittichai, 2018: 113 (in key); Rasheed, et al. 2019: 428.
 * Distribution: Afghanistan, India, Nepal, Pakistan.

Worker
Wilson (1964) - Syntypes: HW 0.51mm, HL 0.62mm, SL 0.54mm, Sl 106; HW 0.46mm, HL 0.60mm, SL 0.47 mm, Sl 102. HW of 7 other syntypes measured 0.46-0.53 mm. Close to wroughtoni, differing as follows: larger; with much shorter scapes; the petiolar node a little higher; a broad triangular area covering most of central occiput and vertex infuscate, contrasting with the yellow coloration of the remainder of head (wroughtoni is concolorous yellow). While sagei may in time prove to be a northern geographic variant of wroughtoni, I have considered the characters listed here adequate to justify provisional species rank for a form already bearing a name.

Jaitrong et al. (2010) - Worker lectotype and paralectotypes (n = 6): TL 3.1-3.2 mm; HL 0.63-0.65 mm; HW 0.50-0.53 mm: SL 0.50-0.53 mm: ML 1.0-1.1 mm; MTL 0.55-0.58 mm; PL 0.25 mm; CI 80-81; SI 100.

Head in full-face view subrectangular, distinctly longer than broad, with feebly convex sides and almost straight posterior margin; anterior portion slightly broader than posterior portion. Antennal scape short (shorter than head length: SI 100), not extending beyond the posterior margin of head; antennal segment II as long as III or less: II and III each clearly longer than each of IV-VII; VIII and IX combined as long as the last antennal segment (X). Frontal carina longer than in the other species of this group, extending posteriad beyond the posterior margin of antennal torulus. Clypeus short, with its anterior margin bearing 9-10 denticles. Mandible with the apical tooth large and curved, followed by 10-11 minute teeth on masticatory margin. Mesosoma elongate: in profile pronotum strongly convex dorsally; promesonotum sloping gradually to metanotal groove. Propodeum slightly lower than promesonotum; its dorsal outline almost straight propodeal junction rounded; declivity of propodeum flattened, laterally margined with a weak ridge. Petiole small, in profile rounded dorsally; subpetiolar process weakly developed, with its ventral outline nearly straight; its anteroventral corner obtusely angulate: postpetiole slightly smaller than petiole, with its node rounded and scarcely longer than broad.

Body extensively smooth and shiny. Anteriormost portion of pronotum minutely punctate; upper portion of mesopleuron with fine rugae.

Body with a few long standing hairs mixed with short decumbent hairs over the surface; length of the longest pronotal hair 0.20-0.25 mm. Head, mesosoma, waist and gaster deep yellow to pale brown; mandible pale yellow: antenna and legs extensively yellow.

Type Material
Jaitrong et al. (2010) - Six syntype workers (two pins, three on each pin) from Dharmsala. India (, examined). One worker (top on a pin) is selected as the lectotype, the others as paralectotypes.

References based on Global Ant Biodiversity Informatics

 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Dunford J. C., J. C. Turbyville, and J. M. Leavengood, Jr. 2014. Checklist of medically important Hymenoptera of Afghanistan. Insecta Mundi 0339: 1-13.
 * Emery C. 1910. Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum 102: 1-34.
 * Forel A. 1901. Les Formicides de l'Empire des Indes et de Ceylan. Part VIII. J. Bombay Nat. Hist. Soc. 13: 462-477
 * Forel A. 1906. Les fourmis de l'Himalaya. Bulletin de la Société Vaudoise des Sciences Naturelles 42: 79-94.
 * Jaitrong W.; Yamane, S.; Wiwatwitaya, D. 2010. The army ant Aenictus wroughtonii (Hymenoptera, Formicidae, Aenictinae) and related species in the Oriental region, with descriptions of two new species. Japanese Journal of Systematic Entomology 16:33-46.
 * Rasheed M. T., I. Bodlah, A. G. Fareen, A. A. Wachkoo, X. Huang, and S. A. Akbar. 2019. A checklist of ants (Hymenoptera: Formicidae) in Pakistan. Sociobiology 66(3): 426-439.
 * Wheeler W. M. 1927. Ants collected by Professor F. Silvestri in Indochina. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 20: 83-106.
 * Wilson E. O. 1964. The true army ants of the Indo-Australian area (Hymenoptera: Formicidae: Dorylinae). Pacific Insects 6: 427-483.