Lasius brunneus

This species is widespread in Europe. It lives only where broad-leaved trees occur (Rigato & Toni, 2011).

Identification
Bicoloured with gaster dark brown contrasting with testaceous or pale reddish brown head and alitrunk. Pubescence and body hairs sparse. Occipital hairs restricted to median area of back of head only. Scapes and tibiae never with erect hairs. Back of head flat or feebly concave. Frontal triangle and frontal furrow distinct, ocelli small but always clearly visible. Length: 3.2-4.5 mm (Collingwood 1979).

A distinct species characterized by large males with intermediate sitkaensis-niger type mandibles and workers with short scapes and sparse pilosity. (Wilson 1955)

Distribution
Spain to Crimea and West Himalayas, Italy to Sweden (Collingwood 1979).

Distribution based on Regional Taxon Lists
Oriental Region: India, Pakistan. Palaearctic Region: Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Israel, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Norway, Poland, Portugal, Republic of Korea, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, United Kingdom of Great Britain and Northern Ireland.

Biology
Collingwood (1979) - This species nests in the interior of old trees, chiefly oak, but has also been recorded from hedgerows. It is fugitive and non-aggressive, rapidly dispersing on disturbance and because of its cryptic habits may be somewhat under-recorded. In Norway and Sweden it has frequently occurred nesting in the timbers of old houses and farm buildings, where its populous colonies may be difficult to dislodge. It chiefly tends tree aphids including the large bark feeding Stomaphis. Single queens initiate colonies in the crevices of old trees but may also be accepted back into the mother nest after the mating flight which occurs in June and early July.

Wilson (1955) - Most European authors agree that brunneus is a timid species adapted to living under the bark and in the wood of tree trunks. Donisthorpe (1927) found a large population of this species in the Windsor Forest of England limited to living trees, which the ants penetrated from the trunk up into the main branches and down into the roots. Various trees were inhabited, including oaks, elm, ash, beech, poplar and maple. It is not clear whether the workers carried on much excavation in the living wood, but this seems unlikely due to the rather unspectacular mandibular apparatus of the species. Forsslund (1949) found brunneus in oaks in dense, undisturbed woodland in several localities in the vicinity of Stockholm. The nests were mostly in dead wood, but occasional galleries penetrated living wood. Scherdlin (1909) found the species in Alsace nesting in the trunks of trees and timber of houses. Clausen (1938) observed a swarm of reproductives inside a house in Zurich. Gosswald (1932) states that in Germany brunneus is found as often under stones as in dead wood; since this observation is divergent from those of other authors, the possibility must be considered that he was erroneously including some alienus in his concept of brunneus.

Donisthorpe (ibid.), who has undertaken the most careful study of this species to date, found workers transporting and tending aphids of the genus Stomaphis. He also observed them carrying psoeids and other small insects to the nests, presumable for use as animal food.

brunneus appears to hold its nuptial flights earlier in the day and season than other European members of the subgenus. Donisthorpe (ibid.) encountered winged queens and males swarming over the trunk of an oak at noon on June 25, and Forsslund (ibid.) saw the same thing from noon to 1:30 p.m. during the period June 10-16.

Other Insects
This species is a host for the temporary parasites, and.

This ant has been associated with the butterfly (Obregon et al. 2015).

Nematodes
This species is a host for the nematodes, , , and  (Kohler, 2012).

Nomenclature

 *  brunneus. Formica brunnea Latreille, 1798: 41 (w.q.) FRANCE. Mayr, 1855: 358 (m.); Lorite, Chica & Palomeque, 1998: 28 (k.). Combination in Lasius: Mayr, 1861: 50; in Donisthorpea: Donisthorpe, 1915d: 347; in Formicina: Emery, 1916b: 241; in Lasius: Müller, 1923: 127; in Acanthomyops: Kuznetsov-Ugamsky, 1927e: 188; in Lasius (Lasius): Wilson, 1955a: 47. Subspecies of niger: Forel, 1874: 47; Forel, 1892i: 307; Bondroit, 1910: 486; Kuznetsov-Ugamsky, 1927e: 188. Status as species: André, 1881b: 60; Dalla Torre, 1893: 182; Bondroit, 1912: 352; Forel, 1915d: 52; Bondroit, 1918: 26; Müller, 1923: 127; Karavaiev, 1927c: 279; Finzi, 1930d: 316; Menozzi, 1939a: 313; Wilson, 1955a: 47; Bernard, 1967: 358; Kutter, 1977c: 228; Collingwood, 1982: 285; Atanassov & Dlussky, 1992: 238; Seifert, 1992b: 6. Senior synonym of pallida: Latreille, 1802c: 169; Wilson, 1955a: 47; of timida: Smith, F. 1858b: 7; Seifert, 1992b: 6; of alienobrunneus: Stärcke, 1944a: 157; Wilson, 1955a: 47; Kutter, 1977c: 14; Seifert, 1992b: 6; of nigrobrunneus: Wilson, 1955a: 47; Seifert, 1992b: 6. Current subspecies: nominal plus emarginatobrunneus.
 * pallida. Formica pallida Latreille, 1798: 41 (w.q.) FRANCE. Junior synonym of brunneus: Latreille, 1802c: 169. Unrecognisable taxon, incertae sedis in Lasius: Seifert, 1992b: 48. [Note. As the taxa brunneus and pallida were both described by Latreille and the synonymy was also established by Latreille, it seems reasonable to let it stand.]
 * timida. Formica timida Foerster, 1850a: 35 (w.) GERMANY. Schenck, 1852: 54 (q.m.). Junior synonym of brunneus: Smith, F. 1858b: 7; Seifert, 1992b: 6.
 * alienobrunneus. Lasius niger var. alienobrunneus Forel, 1874: 47 (w.) SWITZERLAND . Emery, 1916b: 249 (q.). Junior synonym of alienus: Dalla Torre, 1893: 182. Revived from synonymy as subspecies of brunneus: Ruzsky, 1902d: 17. Raised to species: Bingham, 1903: 340. Subspecies of brunneus: Forel, 1915d: 53; Karavaiev, 1927c: 279. Junior synonym of brunneus: Stärcke, 1944a: 157; Wilson, 1955a: 47; Kutter, 1977c: 14; Seifert, 1992b: 6.
 * nigrobrunneus. Acanthomyops (Dendrolasius) brunneus var. nigrobrunneus Donisthorpe, 1926b: 18 (w.) ITALY. Junior synonym of brunneus: Wilson, 1955a: 47.

Worker
Wilson (1955) - (1) Scape shorter relative to head width than in any other member of the subgenus; S1 82-91 in all European series measured; 94 in the himalayanus lectotype and in one specimen from Lahore, Pakistan, both small specimens.

(2) Small individuals (PW 0.50-0.57 mm.), when viewed in perfect full face, with the lateral margins of the eyes not reaching the lateral borders of the head; in Lasius niger and Lasius alienus they reach or exceed it.

(3) Mandibles proportionately shorter, more incurved, and inserted slightly closer to the midline, and head more massive relative to the alitrunk, than other Lasius s. s. (Pl. 1, Fig. 9). Occipital margin viewed in full face flat to feebly convex, as opposed to the typically concave outline of niger and alienus.

(4) Mandible with only two basal teeth in all of seven nest series examined for this character.

(5) Scapes and tibiae completely devoid of standing hairs and nearly devoid of hairs of any inclination. Body pilosity sparse; the curving portion of the occipital angles viewed in full face typically devoid of hairs, rarely with one or two; the latter condition occurs in other members of the subgenus but is highly exceptional.

(6) Alitrunk and petiole homogeneous light reddish brown, rarely medium reddish brown, contrasting against the dark brown gaster. The head usually the same color, occasionally darkening to medium or dark reddish brown to contrast against the alitrunk. (niger and alienus typically concolorous.)

Size range and dispersion probably about the same as in niger. In a sample of 29, with no more than 2 individuals per nest series, PW range 0.50-0.73 mm., mean with standard error 0.630 ± 0.012 mm., standard deviation 0.063 mm. ML less than EW. Anterior margin of median clypeal lobe and structure of the mandibular basal angle essentially as in the niger complex. The greater massiveness of the alitrunk in this species can perhaps best be expressed as a ratio of alitrunk length to the maximum head depth measured perpendicular to the long axis of the head. Several medium-sized brunneus gave such a ratio of 67-69, whereas alienus of comparable size ranged between 70 and 76. Viewed from the side the propodeal angle tends to be more acute and the declivitous face of the propodeum tends to be longer relative to the dorsal face than in other Lasius s. s. The dorsal margin of the petiole was invariably concave in all series examined; occasionally the concavity is so deep as to be nearly semicircular.

Queen
Wilson (1955) - (1) SI low; 68-71 in 9 individuals from 6 localities having HW 1.49-1.64 mm., and 76 in a smaller individual with HW 1.39 mm,

(2) Pilosity and mandibular dentition as in worker.

(3) Frontal suture well marked, set in the middle of a conspicuous broad, shallow trough.

(4) Color distinctive; body uniformly dark reddish brown, appendages a contrasting yellowish brown.

(5) Fore wings infumated in the inner and upper thirds.

Male
Wilson (1955) - (1) Larger than other members of the subgenus, HW 1.04-1.10 mm.

(2) Mandibles of a type intermediate between Lasius pallitarsis and Lasius niger: there is a well marked subapical cleft as in sitkaensis, but it is set more posteriorly than in this primitive species; the basal angle is broadly rounded and the masticatory border lacks denticles, both of which characters are associated with the more advanced niger type.

(3) Frontal suture conspicuous as in queen.

(4) The entire dorsal petiolar margin involved in a deep concavity. In a series from Windsor Forest, Berks, England, secondary lateral convexities are present within this concavity.

(5) Parameres shorter relative to HW than in other members of the subgenus.

SI of several individuals measured 60-64, overlapping part of range of variation of niger (q. v.). ML 0.15-0.17 mm., overlapping part of ranges of niger and emarqinatus.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antonov I. A. 2013. Ant Assemblages (Hymenoptera: Formicidae) of Cities of the Temperate Zone of Eurasia. Russian Journal of Ecology 44(6): 523526.
 * Antonova V., and L. Penev. 2008. Classification of assemblages of ants in the green areas in Sofia City. Acta Zoologica Bulgarica 60(2): 103-110.
 * Arahou M. 2008. Catalogue de l'entomofaune du Chêne vert du Moyen Atlas (Maroc). Documents de lInstitut Scientifique, Rabat 22, 39 pages.
 * ArtDatabanken Bugs (via GBIG)
 * Asociacion Iberica de Mirmecologia. 2012. List of species collected during the Taxomara Tres Cantos 2012. Iberomyrmex 4: 17-18.
 * Asociacion Iberica de Mirmecologia. 2014. List of species collected during the Taxomara 2014 Oviedo. Iberomyrmex 6: 23-24.
 * Azuma M. 1938. A list of ants found in Osaka Prefecture, Japan. Entomol. World Tokyo 6: 238-243.
 * Azuma M. 1938. A list of ants found in Osaka Prefecture, Japan. Entomological World. Tokyo 6:238-243.
 * Azuma M. 1951. On the Myrmecological fauna of Osaka Prefecture, Japan with description of new species (Formicidae, Hymenoptera). Hyogo Biology 1(5): 1-5.
 * Azuma M. 1953. On the myrmecological fauna of Mt. Rokko, Hyogo Prefecture. Warera 2:1-7.
 * Azuma M. 1955. A list of ants (Formicidae) from Hokkaido Is. Hyogo Biology 3:79-80.
 * Azuma M. 1977. On the myrmecological-fauna of Mt. Rokko, Hyogo, with description of a new species (Formicidae, Hymenoptera). Hyogo Biology 7:112-118.
 * Babik H. 2011. Ants of Botanical and Zoological Gardens of Warsaw (Poland). Entomologica romanica 16: 53.
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
 * Barrett K. E. 1967. Ants in South Brittany. Entomologist's Record and Journal of Variation 79:112-116.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
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 * Bees, Wasps and Ants Recording Society 243383515
 * Bees, Wasps and Ants Recording Society 243383516
 * Bees, Wasps and Ants Recording Society 243383517
 * Bees, Wasps and Ants Recording Society 243383518
 * Bees, Wasps and Ants Recording Society 243383519
 * Bees, Wasps and Ants Recording Society 243383520
 * Bees, Wasps and Ants Recording Society 243383521
 * Bees, Wasps and Ants Recording Society 243383522
 * Bees, Wasps and Ants Recording Society 243383523
 * Bees, Wasps and Ants Recording Society 243383524
 * Bees, Wasps and Ants Recording Society 243383525
 * Bees, Wasps and Ants Recording Society 243383526
 * Bees, Wasps and Ants Recording Society 243383527
 * Bees, Wasps and Ants Recording Society 243383528
 * Bees, Wasps and Ants Recording Society 243383529
 * Bees, Wasps and Ants Recording Society 243383530
 * Bees, Wasps and Ants Recording Society 243383531
 * Bees, Wasps and Ants Recording Society 243383532
 * Bees, Wasps and Ants Recording Society 243383533
 * Bees, Wasps and Ants Recording Society 243383534
 * Bees, Wasps and Ants Recording Society 243383535
 * Bees, Wasps and Ants Recording Society 243383536
 * Bees, Wasps and Ants Recording Society 243383537
 * Bees, Wasps and Ants Recording Society 243383538
 * Bees, Wasps and Ants Recording Society 243383539
 * Bees, Wasps and Ants Recording Society 243383540
 * Bees, Wasps and Ants Recording Society 243383541
 * Bees, Wasps and Ants Recording Society 243383542
 * Bees, Wasps and Ants Recording Society 243383543
 * Bees, Wasps and Ants Recording Society 243383544
 * Bees, Wasps and Ants Recording Society 243383545
 * Bees, Wasps and Ants Recording Society 243383546
 * Bees, Wasps and Ants Recording Society 243383547
 * Bees, Wasps and Ants Recording Society 243383548
 * Bees, Wasps and Ants Recording Society 243383549
 * Bees, Wasps and Ants Recording Society 243383550
 * Bees, Wasps and Ants Recording Society 243383551
 * Bees, Wasps and Ants Recording Society 243383552
 * Bees, Wasps and Ants Recording Society 243383553
 * Bees, Wasps and Ants Recording Society 243383554
 * Bees, Wasps and Ants Recording Society 243383555
 * Bees, Wasps and Ants Recording Society 243383556
 * Bees, Wasps and Ants Recording Society 243383557
 * Bees, Wasps and Ants Recording Society 243383558
 * Bees, Wasps and Ants Recording Society 243383559
 * Bees, Wasps and Ants Recording Society 243383560
 * Bees, Wasps and Ants Recording Society 243383561
 * Bees, Wasps and Ants Recording Society 243383562
 * Bees, Wasps and Ants Recording Society 243383563
 * Bees, Wasps and Ants Recording Society 243383564
 * Bees, Wasps and Ants Recording Society 243383565
 * Bees, Wasps and Ants Recording Society 243383566
 * Bees, Wasps and Ants Recording Society 243383567
 * Bees, Wasps and Ants Recording Society 243383568
 * Bees, Wasps and Ants Recording Society 243383569
 * Bees, Wasps and Ants Recording Society 243383570
 * Bees, Wasps and Ants Recording Society 243383571
 * Bees, Wasps and Ants Recording Society 243383572
 * Bees, Wasps and Ants Recording Society 243383573
 * Bees, Wasps and Ants Recording Society 243383574
 * Bees, Wasps and Ants Recording Society 243383575
 * Bees, Wasps and Ants Recording Society 243383576
 * Bees, Wasps and Ants Recording Society 243383577
 * Bees, Wasps and Ants Recording Society 243383578
 * Bees, Wasps and Ants Recording Society 243383579
 * Bees, Wasps and Ants Recording Society 243383580
 * Bees, Wasps and Ants Recording Society 243383581
 * Bees, Wasps and Ants Recording Society 243383582
 * Bees, Wasps and Ants Recording Society 243383583
 * Bees, Wasps and Ants Recording Society 243383584
 * Bees, Wasps and Ants Recording Society 243383585
 * Bees, Wasps and Ants Recording Society 243383586
 * Bees, Wasps and Ants Recording Society 243383587
 * Bees, Wasps and Ants Recording Society 243383588
 * Bees, Wasps and Ants Recording Society 243383589
 * Bees, Wasps and Ants Recording Society 243383590
 * Bees, Wasps and Ants Recording Society 243383591
 * Bees, Wasps and Ants Recording Society 243383592
 * Bees, Wasps and Ants Recording Society 243383593
 * Bees, Wasps and Ants Recording Society 243383594
 * Bees, Wasps and Ants Recording Society 243383595
 * Bees, Wasps and Ants Recording Society 243383596
 * Bees, Wasps and Ants Recording Society 243383597
 * Bees, Wasps and Ants Recording Society 243383598
 * Bees, Wasps and Ants Recording Society 243383599
 * Bees, Wasps and Ants Recording Society 243383600
 * Bees, Wasps and Ants Recording Society 243383601
 * Bees, Wasps and Ants Recording Society 243383602
 * Bees, Wasps and Ants Recording Society 243383603
 * Bees, Wasps and Ants Recording Society 243383604
 * Bees, Wasps and Ants Recording Society 243383605
 * Bees, Wasps and Ants Recording Society 243383606
 * Bees, Wasps and Ants Recording Society 243383607
 * Bees, Wasps and Ants Recording Society 243383608
 * Bees, Wasps and Ants Recording Society 243383609
 * Bees, Wasps and Ants Recording Society 243383610
 * Bees, Wasps and Ants Recording Society 243383611
 * Bees, Wasps and Ants Recording Society 243383612
 * Bees, Wasps and Ants Recording Society 243383613
 * Bees, Wasps and Ants Recording Society 243383614
 * Bees, Wasps and Ants Recording Society 243383615
 * Bees, Wasps and Ants Recording Society 243383616
 * Bees, Wasps and Ants Recording Society 243383617
 * Bees, Wasps and Ants Recording Society 243383618
 * Bees, Wasps and Ants Recording Society 243383619
 * Bees, Wasps and Ants Recording Society 243383620
 * Bees, Wasps and Ants Recording Society 243383621
 * Bees, Wasps and Ants Recording Society 243383622
 * Bees, Wasps and Ants Recording Society 243383623
 * Bees, Wasps and Ants Recording Society 243383624
 * Bees, Wasps and Ants Recording Society 243383625
 * Bees, Wasps and Ants Recording Society 243383626
 * Bees, Wasps and Ants Recording Society 243383627
 * Bees, Wasps and Ants Recording Society 243383628
 * Bees, Wasps and Ants Recording Society 243383629
 * Bees, Wasps and Ants Recording Society 243383630
 * Bees, Wasps and Ants Recording Society 243383631
 * Bees, Wasps and Ants Recording Society 243383632
 * Bees, Wasps and Ants Recording Society 243383633
 * Bees, Wasps and Ants Recording Society 243383634
 * Bees, Wasps and Ants Recording Society 243383635
 * Bees, Wasps and Ants Recording Society 243383636
 * Bees, Wasps and Ants Recording Society 243383637
 * Bees, Wasps and Ants Recording Society 243383638
 * Bees, Wasps and Ants Recording Society 243383639
 * Bees, Wasps and Ants Recording Society 243383640
 * Bees, Wasps and Ants Recording Society 243383641
 * Bees, Wasps and Ants Recording Society 243383642
 * Bees, Wasps and Ants Recording Society 243383643
 * Bees, Wasps and Ants Recording Society 243383644
 * Bees, Wasps and Ants Recording Society 243383645
 * Bees, Wasps and Ants Recording Society 243383646
 * Bees, Wasps and Ants Recording Society 243383647
 * Bees, Wasps and Ants Recording Society 243383648
 * Bees, Wasps and Ants Recording Society 243383649
 * Bees, Wasps and Ants Recording Society 243383650
 * Bees, Wasps and Ants Recording Society 243383651
 * Bees, Wasps and Ants Recording Society 243383652
 * Bees, Wasps and Ants Recording Society 243383653
 * Bees, Wasps and Ants Recording Society 243383654
 * Bees, Wasps and Ants Recording Society 243383655
 * Bees, Wasps and Ants Recording Society 243383656
 * Bees, Wasps and Ants Recording Society 243383657
 * Bees, Wasps and Ants Recording Society 243383658
 * Bees, Wasps and Ants Recording Society 243383659
 * Bees, Wasps and Ants Recording Society 243383660
 * Bees, Wasps and Ants Recording Society 243383661
 * Bees, Wasps and Ants Recording Society 243383662
 * Bees, Wasps and Ants Recording Society 243383663
 * Bees, Wasps and Ants Recording Society 243383664
 * Bees, Wasps and Ants Recording Society 243383665
 * Bees, Wasps and Ants Recording Society 243383666
 * Bees, Wasps and Ants Recording Society 243383667
 * Bees, Wasps and Ants Recording Society 243383668
 * Bees, Wasps and Ants Recording Society 243383669
 * Bees, Wasps and Ants Recording Society 243383670
 * Bees, Wasps and Ants Recording Society 243383671
 * Bees, Wasps and Ants Recording Society 243383672
 * Bees, Wasps and Ants Recording Society 243383673
 * Bees, Wasps and Ants Recording Society 243383674
 * Bees, Wasps and Ants Recording Society 243383675
 * Bees, Wasps and Ants Recording Society 243383676
 * Bees, Wasps and Ants Recording Society 243383677
 * Bees, Wasps and Ants Recording Society 243383678
 * Bees, Wasps and Ants Recording Society 243383679
 * Bees, Wasps and Ants Recording Society 243383680
 * Bees, Wasps and Ants Recording Society 243383681
 * Bees, Wasps and Ants Recording Society 243383682
 * Bees, Wasps and Ants Recording Society 243383683
 * Bees, Wasps and Ants Recording Society 243383684
 * Bees, Wasps and Ants Recording Society 243383685
 * Bees, Wasps and Ants Recording Society 243383686
 * Bees, Wasps and Ants Recording Society 243383687
 * Bees, Wasps and Ants Recording Society 243383688
 * Bees, Wasps and Ants Recording Society 243383689
 * Bees, Wasps and Ants Recording Society 243383690
 * Bees, Wasps and Ants Recording Society 243383691
 * Bees, Wasps and Ants Recording Society 243383692
 * Bees, Wasps and Ants Recording Society 243383693
 * Bees, Wasps and Ants Recording Society 243383694
 * Bees, Wasps and Ants Recording Society 243383695
 * Bees, Wasps and Ants Recording Society 243383696
 * Bees, Wasps and Ants Recording Society 243383697
 * Bees, Wasps and Ants Recording Society 243383698
 * Bees, Wasps and Ants Recording Society 243383699
 * Bees, Wasps and Ants Recording Society 243383700
 * Bees, Wasps and Ants Recording Society 243383701
 * Bees, Wasps and Ants Recording Society 243383702
 * Bees, Wasps and Ants Recording Society 243383703
 * Bees, Wasps and Ants Recording Society 243383704
 * Bees, Wasps and Ants Recording Society 243383705
 * Bees, Wasps and Ants Recording Society 243383706
 * Bees, Wasps and Ants Recording Society 243383707
 * Bees, Wasps and Ants Recording Society 243383708
 * Bees, Wasps and Ants Recording Society 243383709
 * Bees, Wasps and Ants Recording Society 243383710
 * Bees, Wasps and Ants Recording Society 243383711
 * Bees, Wasps and Ants Recording Society 243383712
 * Bees, Wasps and Ants Recording Society 243383713
 * Bees, Wasps and Ants Recording Society 243383714
 * Bees, Wasps and Ants Recording Society 243383715
 * Bees, Wasps and Ants Recording Society 243383716
 * Bees, Wasps and Ants Recording Society 243383717
 * Bees, Wasps and Ants Recording Society 243383718
 * Bees, Wasps and Ants Recording Society 243383719
 * Bees, Wasps and Ants Recording Society 243383720
 * Bees, Wasps and Ants Recording Society 243383721
 * Bees, Wasps and Ants Recording Society 243383722
 * Bees, Wasps and Ants Recording Society 243383723
 * Bees, Wasps and Ants Recording Society 243383724
 * Bees, Wasps and Ants Recording Society 243383725
 * Bees, Wasps and Ants Recording Society 243383726
 * Bees, Wasps and Ants Recording Society 243383727
 * Bees, Wasps and Ants Recording Society 243383728
 * Bees, Wasps and Ants Recording Society 243383729
 * Bees, Wasps and Ants Recording Society 243383730
 * Bees, Wasps and Ants Recording Society 243383731
 * Bees, Wasps and Ants Recording Society 243383732
 * Bees, Wasps and Ants Recording Society 243383733
 * Bees, Wasps and Ants Recording Society 243383734
 * Bees, Wasps and Ants Recording Society 243383735
 * Bees, Wasps and Ants Recording Society 243383736
 * Bees, Wasps and Ants Recording Society 243383737
 * Bees, Wasps and Ants Recording Society 243383738
 * Bees, Wasps and Ants Recording Society 243383739
 * Bees, Wasps and Ants Recording Society 243383740
 * Bees, Wasps and Ants Recording Society 243383741
 * Bees, Wasps and Ants Recording Society 243383742
 * Bees, Wasps and Ants Recording Society 243383743
 * Bees, Wasps and Ants Recording Society 243383744
 * Bees, Wasps and Ants Recording Society 243383745
 * Bees, Wasps and Ants Recording Society 243383746
 * Bees, Wasps and Ants Recording Society 243383747
 * Bees, Wasps and Ants Recording Society 243383748
 * Bees, Wasps and Ants Recording Society 243383749
 * Bees, Wasps and Ants Recording Society 243383750
 * Bees, Wasps and Ants Recording Society 243383751
 * Bees, Wasps and Ants Recording Society 243383752
 * Bees, Wasps and Ants Recording Society 243383753
 * Bees, Wasps and Ants Recording Society 243383754
 * Bees, Wasps and Ants Recording Society 243383755
 * Bees, Wasps and Ants Recording Society 243383756
 * Bees, Wasps and Ants Recording Society 243383757
 * Bees, Wasps and Ants Recording Society 243383758
 * Bees, Wasps and Ants Recording Society 243383759
 * Bees, Wasps and Ants Recording Society 243383760
 * Bees, Wasps and Ants Recording Society 243383761
 * Bees, Wasps and Ants Recording Society 243383762
 * Bees, Wasps and Ants Recording Society 243383763
 * Bees, Wasps and Ants Recording Society 243383764
 * Bees, Wasps and Ants Recording Society 243383765
 * Bees, Wasps and Ants Recording Society 243383766
 * Bees, Wasps and Ants Recording Society 243383767
 * Bees, Wasps and Ants Recording Society 243383768
 * Bees, Wasps and Ants Recording Society 243383769
 * Bees, Wasps and Ants Recording Society 243383770
 * Bees, Wasps and Ants Recording Society 243383771
 * Bees, Wasps and Ants Recording Society 243383772
 * Bees, Wasps and Ants Recording Society 243383773
 * Bees, Wasps and Ants Recording Society 243383774
 * Bees, Wasps and Ants Recording Society 243383775
 * Bees, Wasps and Ants Recording Society 243383776
 * Bees, Wasps and Ants Recording Society 243383777
 * Bees, Wasps and Ants Recording Society 243383778
 * Bees, Wasps and Ants Recording Society 243383779
 * Bees, Wasps and Ants Recording Society 243383780
 * Bees, Wasps and Ants Recording Society 243383781
 * Bees, Wasps and Ants Recording Society 243383782
 * Bees, Wasps and Ants Recording Society 243383783
 * Bees, Wasps and Ants Recording Society 243383784
 * Bees, Wasps and Ants Recording Society 243383785
 * Bees, Wasps and Ants Recording Society 243383786
 * Bees, Wasps and Ants Recording Society 243383787
 * Bees, Wasps and Ants Recording Society 243383788
 * Bees, Wasps and Ants Recording Society 243383789
 * Bees, Wasps and Ants Recording Society 243383790
 * Bees, Wasps and Ants Recording Society 243383791
 * Bees, Wasps and Ants Recording Society 243383792
 * Bees, Wasps and Ants Recording Society 243383793
 * Bees, Wasps and Ants Recording Society 243383794
 * Bees, Wasps and Ants Recording Society 243383795
 * Bees, Wasps and Ants Recording Society 243383796
 * Bees, Wasps and Ants Recording Society 243383797
 * Bees, Wasps and Ants Recording Society 243383798
 * Bees, Wasps and Ants Recording Society 243383799
 * Bees, Wasps and Ants Recording Society 243383800
 * Bees, Wasps and Ants Recording Society 243383801
 * Bees, Wasps and Ants Recording Society 243383802
 * Bees, Wasps and Ants Recording Society 243383803
 * Bees, Wasps and Ants Recording Society 243383804
 * Bees, Wasps and Ants Recording Society 243383805
 * Bees, Wasps and Ants Recording Society 243383806
 * Bees, Wasps and Ants Recording Society 243383807
 * Bees, Wasps and Ants Recording Society 243383808
 * Bees, Wasps and Ants Recording Society 243383809
 * Bees, Wasps and Ants Recording Society 243383810
 * Bees, Wasps and Ants Recording Society 243383811
 * Bees, Wasps and Ants Recording Society 243383812
 * Bees, Wasps and Ants Recording Society 243383813
 * Bees, Wasps and Ants Recording Society 243383814
 * Bees, Wasps and Ants Recording Society 243383815
 * Bees, Wasps and Ants Recording Society 243383816
 * Bees, Wasps and Ants Recording Society 243383817
 * Bees, Wasps and Ants Recording Society 243383818
 * Bees, Wasps and Ants Recording Society 243383819
 * Bees, Wasps and Ants Recording Society 243383820
 * Bees, Wasps and Ants Recording Society 243383821
 * Bees, Wasps and Ants Recording Society 243383822
 * Bees, Wasps and Ants Recording Society 243383823
 * Bees, Wasps and Ants Recording Society 243383824
 * Bees, Wasps and Ants Recording Society 243383825
 * Bees, Wasps and Ants Recording Society 243383826
 * Bees, Wasps and Ants Recording Society 243383827
 * Bees, Wasps and Ants Recording Society 243383828
 * Bees, Wasps and Ants Recording Society 243383829
 * Bees, Wasps and Ants Recording Society 243383830
 * Bees, Wasps and Ants Recording Society 243383831
 * Bees, Wasps and Ants Recording Society 243383832
 * Bees, Wasps and Ants Recording Society 243383833
 * Bees, Wasps and Ants Recording Society 243383834
 * Bees, Wasps and Ants Recording Society 243383835
 * Bees, Wasps and Ants Recording Society 243383836
 * Bees, Wasps and Ants Recording Society 243383837
 * Bees, Wasps and Ants Recording Society 243383838
 * Bees, Wasps and Ants Recording Society 243383839
 * Bees, Wasps and Ants Recording Society 243383840
 * Bees, Wasps and Ants Recording Society 243383841
 * Bees, Wasps and Ants Recording Society 243383842
 * Bees, Wasps and Ants Recording Society 243383843
 * Bees, Wasps and Ants Recording Society 243383844
 * Bees, Wasps and Ants Recording Society 243383845
 * Bees, Wasps and Ants Recording Society 243383846
 * Bees, Wasps and Ants Recording Society 243383847
 * Bees, Wasps and Ants Recording Society 243383848
 * Bees, Wasps and Ants Recording Society 243383849
 * Bees, Wasps and Ants Recording Society 243383850
 * Bees, Wasps and Ants Recording Society 243383851
 * Bees, Wasps and Ants Recording Society 243383852
 * Bees, Wasps and Ants Recording Society 243383853
 * Bees, Wasps and Ants Recording Society 243383854
 * Bees, Wasps and Ants Recording Society 243383855
 * Bees, Wasps and Ants Recording Society 243383856
 * Bees, Wasps and Ants Recording Society 243383857
 * Bees, Wasps and Ants Recording Society 243383858
 * Bees, Wasps and Ants Recording Society 243383859
 * Bees, Wasps and Ants Recording Society 243383860
 * Bees, Wasps and Ants Recording Society 243383861
 * Bees, Wasps and Ants Recording Society 243383862
 * Bees, Wasps and Ants Recording Society 243383863
 * Bees, Wasps and Ants Recording Society 243383864
 * Bees, Wasps and Ants Recording Society 243383865
 * Bees, Wasps and Ants Recording Society 243383866
 * Bees, Wasps and Ants Recording Society 243383867
 * Bees, Wasps and Ants Recording Society 243383868
 * Bees, Wasps and Ants Recording Society 243383869
 * Bees, Wasps and Ants Recording Society 243383870
 * Bees, Wasps and Ants Recording Society 243383871
 * Bees, Wasps and Ants Recording Society 243383872
 * Bees, Wasps and Ants Recording Society 243383873
 * Bees, Wasps and Ants Recording Society 243383874
 * Bees, Wasps and Ants Recording Society 243383875
 * Bees, Wasps and Ants Recording Society 243383876
 * Bees, Wasps and Ants Recording Society 243383877
 * Bees, Wasps and Ants Recording Society 243383878
 * Bees, Wasps and Ants Recording Society 243383879
 * Bees, Wasps and Ants Recording Society 243383880
 * Bees, Wasps and Ants Recording Society 243383881
 * Bees, Wasps and Ants Recording Society 243383882
 * Bees, Wasps and Ants Recording Society 243383883
 * Bees, Wasps and Ants Recording Society 243383884
 * Bees, Wasps and Ants Recording Society 243383885
 * Bees, Wasps and Ants Recording Society 243383886
 * Bees, Wasps and Ants Recording Society 243383887
 * Bees, Wasps and Ants Recording Society 243383888
 * Bees, Wasps and Ants Recording Society 243383889
 * Bees, Wasps and Ants Recording Society 243383890
 * Bees, Wasps and Ants Recording Society 243383891
 * Bees, Wasps and Ants Recording Society 243383892
 * Bees, Wasps and Ants Recording Society 243383893
 * Bees, Wasps and Ants Recording Society 243383894
 * Bees, Wasps and Ants Recording Society 243383895
 * Bees, Wasps and Ants Recording Society 243383896
 * Bees, Wasps and Ants Recording Society 243383897
 * Bees, Wasps and Ants Recording Society 243383898
 * Bees, Wasps and Ants Recording Society 243383899
 * Bees, Wasps and Ants Recording Society 243383900
 * Bees, Wasps and Ants Recording Society 243383901
 * Bees, Wasps and Ants Recording Society 243383902
 * Bees, Wasps and Ants Recording Society 243383903
 * Bees, Wasps and Ants Recording Society 243383904
 * Bees, Wasps and Ants Recording Society 243383905
 * Bees, Wasps and Ants Recording Society 243383906
 * Bees, Wasps and Ants Recording Society 243383907
 * Bees, Wasps and Ants Recording Society 243383908
 * Bees, Wasps and Ants Recording Society 243383909
 * Bees, Wasps and Ants Recording Society 243383910
 * Bees, Wasps and Ants Recording Society 243383911
 * Bees, Wasps and Ants Recording Society 243383912
 * Bees, Wasps and Ants Recording Society 243383913
 * Bees, Wasps and Ants Recording Society 243383914
 * Bees, Wasps and Ants Recording Society 243383915
 * Bees, Wasps and Ants Recording Society 243383916
 * Bees, Wasps and Ants Recording Society 243383917
 * Bees, Wasps and Ants Recording Society 243383918
 * Bees, Wasps and Ants Recording Society 243383919
 * Bees, Wasps and Ants Recording Society 243383920
 * Bees, Wasps and Ants Recording Society 243383921
 * Bees, Wasps and Ants Recording Society 243383922
 * Bees, Wasps and Ants Recording Society 243383923
 * Bees, Wasps and Ants Recording Society 243383924
 * Bees, Wasps and Ants Recording Society 243383925
 * Bees, Wasps and Ants Recording Society 243383926
 * Bees, Wasps and Ants Recording Society 243383927
 * Bees, Wasps and Ants Recording Society 243383928
 * Bees, Wasps and Ants Recording Society 243383929
 * Bees, Wasps and Ants Recording Society 243383930
 * Bees, Wasps and Ants Recording Society 243383931
 * Bees, Wasps and Ants Recording Society 243383932
 * Bees, Wasps and Ants Recording Society 243383933
 * Bees, Wasps and Ants Recording Society 243383934
 * Bees, Wasps and Ants Recording Society 243383935
 * Bees, Wasps and Ants Recording Society 243383936
 * Bees, Wasps and Ants Recording Society 243383937
 * Bees, Wasps and Ants Recording Society 243383938
 * Bees, Wasps and Ants Recording Society 243383939
 * Bees, Wasps and Ants Recording Society 243383940
 * Bees, Wasps and Ants Recording Society 243383941
 * Bees, Wasps and Ants Recording Society 243383942
 * Bees, Wasps and Ants Recording Society 243383943
 * Bees, Wasps and Ants Recording Society 243383944
 * Bees, Wasps and Ants Recording Society 243383945
 * Bees, Wasps and Ants Recording Society 243383946
 * Bees, Wasps and Ants Recording Society 243383947
 * Bees, Wasps and Ants Recording Society 243383948
 * Bees, Wasps and Ants Recording Society 243383949
 * Bees, Wasps and Ants Recording Society 243383950
 * Bees, Wasps and Ants Recording Society 243383951
 * Bees, Wasps and Ants Recording Society 243383952
 * Bees, Wasps and Ants Recording Society 243383953
 * Bees, Wasps and Ants Recording Society 243383954
 * Bees, Wasps and Ants Recording Society 243383955
 * Bees, Wasps and Ants Recording Society 243383956
 * Bees, Wasps and Ants Recording Society 243383957
 * Bees, Wasps and Ants Recording Society 243383958
 * Bees, Wasps and Ants Recording Society 243383959
 * Bees, Wasps and Ants Recording Society 243383960
 * Bees, Wasps and Ants Recording Society 243383961
 * Bees, Wasps and Ants Recording Society 243383962
 * Bees, Wasps and Ants Recording Society 243383963
 * Bees, Wasps and Ants Recording Society 243383964
 * Bees, Wasps and Ants Recording Society 243383965
 * Bees, Wasps and Ants Recording Society 243383966
 * Bees, Wasps and Ants Recording Society 243383967
 * Bees, Wasps and Ants Recording Society 243383968
 * Bees, Wasps and Ants Recording Society 243383969
 * Bees, Wasps and Ants Recording Society 243383970
 * Bees, Wasps and Ants Recording Society 243383971
 * Bees, Wasps and Ants Recording Society 243383972
 * Bees, Wasps and Ants Recording Society 243383973
 * Bees, Wasps and Ants Recording Society 243383974
 * Bees, Wasps and Ants Recording Society 243383975
 * Bees, Wasps and Ants Recording Society 243383976
 * Bees, Wasps and Ants Recording Society 243383977
 * Bees, Wasps and Ants Recording Society 243383978
 * Bees, Wasps and Ants Recording Society 243383979
 * Bees, Wasps and Ants Recording Society 243383980
 * Bees, Wasps and Ants Recording Society 243383981
 * Bees, Wasps and Ants Recording Society 243383982
 * Bees, Wasps and Ants Recording Society 243383983
 * Bees, Wasps and Ants Recording Society 243383984
 * Bees, Wasps and Ants Recording Society 243383985
 * Bees, Wasps and Ants Recording Society 243383986
 * Bees, Wasps and Ants Recording Society 243383987
 * Bees, Wasps and Ants Recording Society 243383988
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