Temnothorax interruptus

This species occurs mostly in xerothermic grasslands or other open habitats, especially in northern localities, and it can often be collected in overgrown limestone or gypsum rocks. It nests in soil, under stones, in moss or in rock rubble. Sometimes specimens can be found at the edges of dry, deciduous oak forests (Csosz et al., 2018).

Identification
Prebus (2017) - A member of the Palearctic clade.

Collingwood (1979) - Light bright yellow with dark areas at the side of the dorsum of the first gaster segment and frequently at the front corners of the head. The antennal club is distinctly dark. The dorsal outline of the alitrunk is smoothly curved without a break and the propodeal spines are long and curved. The petiole node is steeply peaked in profile. Length: 2.3-3.4 mm.

Csosz et al. (2018) - Morphological characteristics of Temnothorax interruptus are considered the most distinct amongst three closely related species. Its workers differ from Temnothorax morea and Temnothorax strymonensis in: 1) shorter antennal scape which never reaches occipital margin of head; 2) wide frontal lobes (FL / SL > 0.5 μm); and 3) low SL / CS ratio (0.800 ± 0.02 in T. interruptus vs. 0.894 ± 0.02 in T. morea vs. 0.856 ± 0.02 in T. strymonensis).

Gynes of Temnothorax interruptus differ from those of T. morea and T. strymonensis in: 1) darker body colour (brown to dark brown vs. orange to bright orange); 2) shorter antennal scape which never reaches occipital margin of head (in T. morea and T. strymonensis antennal scape reaches occipital margin of head); and 3) the whole surface of the scutum is conspicuously costulate (in T. morea and T. strymonensis the dorsal surface of scutum inconspicuously costulate or smooth).

Distribution
This is a Turano-European species. Its known distribution stretches from Spain to the Caucasus, from Central Europe to the Mediterranean peninsula. This species, by crossing the Caucasus, also gained a foothold in Eastern Turkey, where it co-occurs with Temnothorax strymonensis (Csosz et al., 2018).

Distribution based on Regional Taxon Lists
Palaearctic Region: Armenia, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Germany, Hungary, Iberian Peninsula, Montenegro, Netherlands, Norway, Poland, Republic of Macedonia, Romania, Russian Federation, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine , United Kingdom of Great Britain and Northern Ireland.

Biology
Collingwood (1979) - This is a rather uncommon ground nesting species. In Britain where it has been well studied by Donisthorpe (1927) it is found nesting in dry peat or among small stones and heather roots in small colonies of 50-100 workers and single queens. Alatae are present in the nests during July.

Nomenclature

 *  interruptus. Myrmica interrupta Schenck, 1852: 106 (w.q.) GERMANY.
 * [Also described as new by Schenck, 1853: 188.]
 * Mayr, 1855: 446 (m.).
 * Combination in Leptothorax: Mayr, 1855: 446.
 * Combination in Temnothorax: Bolton, 2003: 271.
 * As unavailable (infrasubspecific) name: Lameere, 1892: 69.
 * Subspecies of tuberum: Forel, 1874: 85 (in key); Emery & Forel, 1879: 458; André, 1883a: 299; Emery, 1884a: 379; Forel, 1890a: lxxiv; Emery, 1891b: 6; Emery, in Dalla Torre, 1893: 127 (footnote); Emery, 1898c: 134; Forel, 1904f: 425; Bondroit, 1910: 497; Stitz, 1914: 63; Crawley, 1914a: 91 (in key); Forel, 1915d: 24 (in key); Emery, 1916b: 175; Escherich, 1917: 328 (in key); Bondroit, 1918: 132; Soudek, 1922: 55; Emery, 1924d: 256; Menozzi, 1925d: 29; Finzi, 1930d: 315; Stitz, 1939: 171.
 * Status as species: Mayr, 1855: 446 (redescription); Smith, F. 1858b: 120; Roger, 1859: 258; Mayr, 1861: 59 (in key); Roger, 1863b: 26; Mayr, 1863: 427; André, 1874: 191 (in key); Emery, 1878b: 51; Dalla Torre, 1893: 124; Bondroit, 1911: 12; Donisthorpe, 1915d: 166; Bondroit, 1918: 132; Müller, 1923b: 94; Donisthorpe, 1927b: 185; Lomnicki, 1928: 5; Zimmermann, 1935: 33; Novák & Sadil, 1941: 93 (in key); Novák, in Kratochvíl, et al. 1944: 119; Consani & Zangheri, 1952: 41; Bernard, 1956a: 163; Bernard, 1967: 212 (redescription); Baroni Urbani, 1971c: 109; Pisarski, 1975: 20; van Boven, 1977: 107; Kutter, 1977c: 131; Collingwood, 1978: 84 (in key); Arnol'di & Dlussky, 1978: 542 (in key); Collingwood, 1979: 75; Agosti & Collingwood, 1987b: 275 (in key); Atanassov & Dlussky, 1992: 138; Douwes, 1995: 90; Bolton, 1995b: 239; Czechowski, et al. 2002: 48; Seifert, 2007: 233; Casevitz-Weulersse & Galkowski, 2009: 495; Lapeva-Gjonova, et al. 2010: 22; Boer, 2010: 51; Legakis, 2011: 18; Borowiec, L. & Salata, 2012: 542; Czechowski, et al. 2012: 145; Borowiec, L. 2014: 179; Radchenko, 2016: 232; Csősz et al., 2018: 109.
 * Senior synonym of simpliciuscula: Roger, 1859: 258; Mayr, 1861: 59; Roger, 1863b: 26; Mayr, 1863: 427; André, 1874: 203 (in list); Forel, 1874: 102 (in list); Emery & Forel, 1879: 459; Dalla Torre, 1893: 124, Emery, 1924d: 256; Radchenko, 2016: 232.
 * Senior synonym of tuberoaffinis: Plateaux & Cagniant, 2013: 429.
 * Senior synonym of knipovitshi: Csősz et al., 2018: 109.
 * Senior synonym of junipereti: Csősz et al., 2018: 118.
 * Senior synonym of nikitae: Csősz et al., 2018: 118.
 * junipereti. Leptothorax junipereti Arnol'di, 1977a: 203 (w.) UKRAINE.
 * Status as species: Arnol'di & Dlussky, 1978: 543 (in key).
 * Junior synonym of knipovitshi: Radchenko, 1994d: 156 (in key); Radchenko, 1995c: 18.
 * Junior synonym of interruptus: Csősz et al., 2018: 118.
 * knipovitshi. Leptothorax tuberum subsp. knipovitshi Karavaiev, 1916: 499 (w.q.) UKRAINE.
 * Combination in Temnothorax: Bolton, 2003: 271.
 * Subspecies of tuberum: Karavaiev, 1927c: 266 (in key); Karavaiev, 1934: 139 (redescription).
 * Status as species: Radchenko, 1994d: 156 (in key); Bolton, 1995b: 240; Radchenko, 1995c: 17; Borowiec, L. 2014: 180; Radchenko, 2016: 222.
 * Junior synonym of interruptus: Csősz et al., 2018: 109.
 * Senior synonym of junipereti: Radchenko, 1994d: 156 (in key); Radchenko, 1995c: 18; Radchenko, 2016: 222.
 * Senior synonym of nikitae: Radchenko, 1994d: 156 (in key); Radchenko, 1995c: 18; Radchenko, 2016: 222.
 * nikitae. Leptothorax nikitae Arnol'di, 1977a: 202 (w.) UKRAINE.
 * Status as species: Arnol'di & Dlussky, 1978: 543 (in key); Arakelian, 1994: 55.
 * Junior synonym of knipovitshi: Radchenko, 1994d: 156 (in key); Radchenko, 1995c: 18.
 * Junior synonym of interruptus: Csősz et al., 2018: 118.
 * simpliciuscula. Myrmica simpliciuscula Nylander, 1856b: 92, pl. 3, fig. 40 (w.) FRANCE.
 * Status as species: Smith, F. 1858b: 120.
 * Junior synonym of interruptus: Roger, 1859: 258; Mayr, 1861: 59.
 * tuberoaffinis. Leptothorax tuberoaffinis Bondroit, 1918: 131 (w.q.) SWITZERLAND.
 * [First available use of Leptothorax tuberum r. affinis var. tuberoaffinis Forel, 1915d: 23 (in key) (w.q.) SWITZERLAND; unavailable (infrasubspecific) name.]
 * [Leptothorax tuberum var. tuberoaffinis Forel, 1874: 86; Dalla Torre, 1893: 128. Nomina nuda.]
 * Subspecies of affinis: Stitz, 1939: 174; Novák & Sadil, 1941: 92 (in key); Bolton, 1995b: 246 (error).
 * Junior synonym of affinis: Kutter, 1977c: 14; Radchenko, 2016: 217 (error).
 * Junior synonym of interruptus: Plateaux & Cagniant, 2013: 428.

Worker
Csosz et al. (2018) - Head orange to dark orange with brown to dark brown posterior part of gena or darker orange to brown posterior part of head. Scapes same colouration as head. Funicles same colouration as scapes or darker brown to dark brown. Mesosoma, legs, petiole and postpetiole orange to dark orange. Sometimes femora darker. Gaster orange to dark orange with complete brown to brownish-black, transverse band on the apical part of the first tergite. In most specimens the band is broad, occupies apical ⅓ to ½ length of first tergite, its anterior margin straight or on sides only slightly protruding forward. In rare aberrations first tergite with transverse band narrower than ⅓ length of the tergite, narrowly interrupted along the middle and laterally distinctly protruding forward.

Head quadratic (CL / CWb: 1.228 [1.161, 1.276]). Eyes small, oval, (EL / CS: 0.259 [0.231, 0.278]). Antennal scape short (SL / CS: 0.80 [0.748, 0.843]), not reaching occipital margin of head. Surface of the scape with very fine microsculpture, shiny, covered with short, moderately dense, suberect or adpressed setae. Mesosoma elongate (ML / CS: 1.223 [1.167, 1.303]), dorsal contour line in profile slightly rounded. Propodeal spines long (SPST / CS: 0.356 [0.314, 0.392]), wide at base, curved downwards with pointed apex. Frontal carinae short, extending to ⅓ length of eye; antennal fossa shallow, rugulose with costae. Frontal lobes distinctly wider than frons (FL / FR: 1.168 [1.111, 1.236]), and shiny. Frons longitudinally costate and sometimes rugose, interstices shiny. Postocular area of head rugulose, sometimes longitudinally costulate. Genae sometimes with sparser rugosity. Interstices feebly microreticulate, shiny. Entire head bearing suberect to erect, pale and thin setae.

Dorsum of mesosoma densely rugose. Lateral surface of promesonotum longitudinally costulate. Lateral surface of propodeum rugulose. Interstices shiny with microreticulation or micropunctation. Area between and below propodeal spines shiny and punctate. Dorsal surface of mesosoma with sparse, erect, long, thick and pale setae. Petiole punctate to rugulose, interstices shiny.

Queen
Csosz et al. (2018) - Head dark brown, sometimes temples orange. Antennal club brown to bright brown. Mesosoma, petiole and postpetiole dark brown to brown, sometimes with brighter spots on the pronotum. Legs orange. First gastral tergite mostly brown with orange spot basally but apical margin always pale, remaining tergites yellow-orange basally and brown apically but the light goes dark gradually.

Eyes big, oval [EL / CS: 0.34 ± 0.004]. Antennal scape short [SL / CS: 0.76 ± 0.03], not reaching occipital margin of head. Propodeal spines medium length [SPST / CS: 0.38 ± 0.01], wide at base, triangular, straight, with acute apex. Clypeus shiny with diffuse, longitudinally carinulae, interstices smooth. Antennal fossa deep, rugulose with concentric carinae. Frontal lobes wide [FLS / CS: 0.46 ± 0.01], rugulose with thick longitudinal costae, interstices shiny. Frons shiny, entire surface longitudinally costate and rugose, interstices smooth and shiny. Area above eyes and sides of head rugulose and sometimes longitudinally costate, interstices shiny. Entire head bearing suberect to erect, pale and thin setae.

Pronotum with thick rugosity on whole dorsal surface. Sides with thick rugosity or gentle, dense longitudinal costae. Surface between rugosity smooth and shiny. Scutum with dense, thick longitudinal costae, shiny. Sometimes rugosity weaker laterally. Scutellum smooth and shiny at the centre, sides smooth or with diffuse, thick, longitudinal costae. Metanotum with slight sculpture, rugulose or punctate. Sometimes with a few thick wrinkles. Propodeum with variable sculpture. Area above propodeal spines with very sparse, transverse and gentle costae or sparse irregular rugosity, interstices microreticulate, shiny. Area between and below propodeal spines with dense punctation. Sides of propodeum punctate or with slight, dense rugosity. Sometimes with a few longitudinal costae, always shiny. Anepisternum and katepisternum shiny, with gentle, dense longitudinal costae. Metaepisternum and metakatepisternum, shiny, with dense, longitudinal rugosity. Surface between rugosity punctate. Dorsal surface of mesosoma with sparse, erect, long, thick and pale setae. Petiole and postpetiole shiny, the entire surface punctate to rugulose, dorsal surface longitudinally costulate. Gaster smooth and shiny, bearing sparse, long, suberect to erect setae.

Type Material
Types studied by Salata et al. (2018): Temnothorax interruptus: Syntype (w.) (FOCOL2008): [Nassau | Leptothorax interruptus Schenck || interruptus Schenck || Type || GBIF-D/FoCol 2010 | specimen + label | data documented] (ZMHB).

References based on Global Ant Biodiversity Informatics

 * Alvarado M., and L. Galle. 2000. Ant assemblages associated with lowland forests in the southern part of the great Hungarian plain. Acta Zoologica Academiae Scientarum Hungaricae 46(2): 79-102.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnol'di K. V. 1977. New and little known species of the genus Leptothorax Mayr (Hymenoptera, Formicidae) in the European regions of the USSR and the Caucasus. Entomol. Rev. (Wash.) 56: 148-153.
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
 * Bernard, F. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bigot L. 1959. Complement a l'inventaire de la faune entomologique de la Camargue 93eme note). La Terre et la Vie 106 : 149-157.
 * Boer P., W. Dekoninck, A. J. Van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten (Amsterdam) 63: 54-58.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec M. L. 2009. New data on the occurrence of formicoxenine ants (Hymenoptera: Formicidae) in Poland. Wiadomo?ci Entomologiczne 28: 237-246.
 * Borowiec M. L. 2009. Nowe dane o rozmieszczeniu mrówek (Hymenoptera: Formicidae) z plemienia Formicoxenini w Polsce. Wiadomosci Entomologiczne 28: 237-246.
 * Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Børgesen L. W. 2000. Ecological notes on Leptothorax interruptus: an ant new to Denmark (Hymenoptera, Formicidae). Entomol. Medd. 68: 67-77.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Cherix D., and S. Higashi. 1979. Distribution verticale des fourmis dans le Jura vaudois et recensement prelimaire des bourdons (Hymenoptera, Formicidae et Apidae). Bull. Soc. Vaud. Sc. Nat. 356(74): 315-324.
 * Colindre L. 2015. Les fourmis en Picardie: bilan 2014 (Hymenoptera/ Formicidae). Entomologiste Picard 26, 15 pages.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
 * Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
 * Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
 * Collingwood, C. A. 1974. A revised list of Norwegian ants (Hymenoptera: Formicidae). Norsk Entomologisk Tidsskrift 21: 31-35.
 * Collingwood, C. A. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Csosz S., S. Salata, and L. Borowiec. 2018. Three Turano-European species of the Temnothorax interruptus group (Hymenoptera: Formicidae) demonstrated by quantitative morphology. Myrmecological News 26: 101-119.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
 * Dekoninck W., H. De Koninck, J. Y. Baugnee, and J. P. Maelfait. 2007. Ant biodiversity conservation in Belgian calcareous grasslands: active management is vital. Belg. J. Zool. 137 (2): 137-146.
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Forel A. 1890. Fourmis de Tunisie et de l'Algérie orientale. Annales de la Société Entomologique de Belgique 34: lxi-lxxvi.
 * Formidabel Database
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galle L., and G. Szonyi. 1988. A check list of ants (Hymenoptera: Formicoidea) of a sandy grassland in Kiskunsag National Park (Hungary). Acta Biol. Szeged 34: 167-168.
 * Gallé L. 1991. Structure and succession of ant assemblages in a north European sand dune area. Holarctic Ecology 14: 31-37.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Hauschteck-Jungen E., and H. Jungen. 1983. Ant chromosomes. II. Karyotypes of western palearctic species. Insectes Soc. 30: 149-164.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-203.
 * Kanizsai O., R. Gallé, and L. Gallé. 2009. Perception of spatial patchiness by ant assemblages (Hymenoptera: Formicidae). Tiscia 37: 3-7.
 * Karaman C., K. Kiran, and V. Aksoy. 2014. New records of the genus Strumigenys Smith, 1860 (Hymenoptera, Formicidae) from Black Sea region of Turkey. Trakya University Journal of Natural Sciences, 15(2): 59-63.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
 * Legakis Collection Database
 * Lomnicki J. 1928. Spis mrówek Lwowa i okolicy. Ksiegi Pamiatkowej (Lecia Gimn. IV Jana Dlugosza Lwowie) 50: 1-10.
 * Lorite P., J. A. Carrillo, M. F. García, and T. Palomeque. 2000. Chromosome numbers in Spanish Formicidae. III. Subfamily Myrmicinae (Hymenoptera). Sociobiology 36: 555-570.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Müller, G. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Nielsen M. G. 2011. A check list of Danish ants and proposed common names. Ent. Meddr. 79: 13-18.
 * Nylander, W. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * Odegaard F., K. M. Olsen, A. Staverlokk, and J. O. Gjershaug. 2015. Towards a new era for the knowledge of ants (Hymenoptera, Formicidae) in Norway? Nine species new to the country. Nowegian Journal of Entomology 62: 80–99.
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Radchenko A. G., T.N. Bazhenova, and E. I.  Simutnik. 2012. Records  of  previously  non-recorded  and  little-known  ant  species  and  genus (Hymenoptera: Formicidae) in the Ukrainian fauna. Ukrainian entomofaunistyka 3 (2): 1-9.
 * Santschi F. 1910. Contributions à la faune entomologique de la Roumanie. Formicides capturées par Mr. A. L. Montandon. Buletinul Societatii Române de Stiinte 19: 648-652.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
 * Somogyi A. A., G. Lorinczi, J. Kovacs, and I. E. Maak. 2017. Structure of ant assemblages in planted poplar (Populus alba) forests and the effect of the common milkweek (Asclepias syriaca). Acta Zoologica Academiae Scientiarum Hungaricae 63(4): 443–457.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Stumper R. 1953. Etudes myrmecologiques. XI. Fourmis luxembourgeoises. Bulletin Soc. Nat. luxemb. 57: 122-135.
 * Tausan I. 2014. Temnothorax parvulus (Schenck, 1852)(Hymenoptera: Formicidae) in Romania. Brukenthal, Acta Musei 9: 511-514.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
 * Wiezick M. 2008. Ant communities (Hymenoptera: Formicidae) of forest-steppe habitats at southern and South-Western slopes of Plesivecka a Planina plateau at Slovensky Kras Karst. Natura Carpatica 49: 85-94.
 * Wiezik M., A. Wiezikova, and M. Svitok. 2011. Vegetation structure, ecological stability, and low-disturbance regime of abandoned dry grasslands support specific ant assemblages in Central Slovakia. Tuexenia 31: 301315.
 * Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
 * de Biseau, J.-C., Y. Quinet, L. Deffernez and J.M. Pasteels. 1997. Explosive food recruitment as a competitive strategy in the ant Myrmica sabuleti (Hymenoptera, Formicidae). Insectes Sociaux 44:59-73