Checklist of Messor species

The following species and subspecies belong to the genus Messor. Synonyms are listed under their senior names. For valid names only see Messor species.

A
abdelazizi Santschi, 1921 (Morocco)

aciculatus (Smith, F., 1874) (China, Democratic People's Republic of Korea, Japan, Mongolia, Russian Federation)

• Messor barbarus lobulifera Emery, 1901

• Stenamma aciculatum brunneicorne Forel, 1901

aciculatus risianus Forel, 1922 (China)

aegyptiacus (Emery, 1878) (Egypt, Israel)

aegyptiacus canaliculatus Donisthorpe, 1944 (Algeria)

aegyptiacus felah Santschi, 1923 (Egypt, Israel)

aegyptiacus phasianicus Bernard, 1953 (Algeria)

aegyptiacus tunetinus Santschi, 1923 (Tunisia)

alexandri Tohmé, G. & Tohmé, H., 1981 (Greece, Iran, Israel, Syrian Arab Republic)

andrei (Mayr, 1886) (Mexico, United States)

• Veromessor andrei castaneus Wheeler, W.M. & Creighton, 1934

• Veromessor andrei flavus Wheeler, W.M. & Creighton, 1934

angularis Santschi, 1928 (Kenya)

antennatus Emery, 1908 (Morocco)

antennatus fodorii Röszler, 1942 (Greece)

antennatus personatus Santschi, 1931 (Morocco)

aphaenogasteroides Pisarski, 1967 (Afghanistan)

aralocaspius (Ruzsky, 1902) (China, Iran, Iraq, Kuwait, Kyrgyzstan, Russian Federation, Saudi Arabia, The former Yugoslav Republic of Macedonia, Turkmenistan)

aralocaspius infumatus Kuznetsov-Ugamsky, 1929 (China, Kazakhstan, Kyrgyzstan)

arenarius (Fabricius, 1787) (Algeria, Israel, Kuwait, Saudi Arabia)

• Myrmica amaurocyclia Foerster, 1850

• Myrmica scalpturata Nylander, 1856

• Stenamma bugnioni Forel, 1904

arenarius alfierii Santschi, 1938 (Egypt)

arenarius diabolus Santschi, 1938 (Egypt)

arenarius ratus Menozzi, 1933 (Israel)

arenarius thebaicus Santschi, 1912 (Egypt)

atanassovii Atanassov, 1982 (Bulgaria)

B
barbarus (Linnaeus, 1767) (Algeria, Balearic Islands, Bulgaria, Egypt, France, Greece, Iberian Peninsula, Morocco, Portugal, Spain, Tunisia)

• Aphaenogaster barbara sordida Forel, 1892

• Formica binodis Fabricius, 1775

• Formica juvenilis Fabricius, 1804

• Formica megacephala Leach, 1825

• Messor barbarus ambiguus Santschi, 1925

• Myrmica rufitarsis Foerster, 1850

• Messor barbarus nigricans Santschi, 1929

barbarus gallaoides Santschi, 1929 (Morocco)

• Messor barbarus nigriceps Santschi, 1925

barbarus mediosanguineus Donisthorpe, 1946 (Turkey)

barbarus politus Karavaiev, 1912 (Algeria)

• Messor barbarus dentiscapus Finzi, 1930

barbarus sahlbergi Forel, 1913 (Egypt)

barbarus santschii Emery, 1908 (Tunisia)

• Messor carthaginensis Bernard, 1980

beduinus Emery, 1922 (Tunisia)

berbericus Bernard, 1955 (Algeria)

bernardi Cagniant, 1967 (Algeria)

bouvieri Bondroit, 1918 (Balearic Islands, France, Greece, Iberian Peninsula, Malta, Portugal, Spain)

boyeri Cagniant, 2006 (Morocco)

buettikeri Collingwood, 1985 (Iraq, Kuwait, Saudi Arabia)

C
caducus (Victor, 1839) (Armenia, Bulgaria, Georgia, Greece, Iran, Malta, Russian Federation, The former Yugoslav Republic of Macedonia)

• Messor caducus caucasicola Arnol'di, 1977

caducus glabrescens Arnoldi, 1977 (Kazakhstan)

capensis (Mayr, 1862) (Botswana, Namibia, South Africa, Zimbabwe)

• Aphaenogaster pseudoaegyptiaca Emery, 1884

• Messor barbarus schencki Wheeler, W.M., 1922

• Messor braunsi Forel, 1913

• Messor donisthorpei Santschi, 1937

capitatus (Latreille, 1798) (Balearic Islands, Bulgaria, Croatia, France, Greece, Iberian Peninsula, Italy, Malta, Montenegro, Portugal, Slovenia, Spain, The former Yugoslav Republic of Macedonia)

• Aphaenogaster barbara nigra André, 1883

• Formica huberiana Leach, 1825

• Myrmica galbula Losana, 1834

caviceps (Forel, 1902) (Algeria)

• Messor planiceps Stitz, 1917

celiae Reyes, 1985 (Iberian Peninsula, Spain)

cephalotes (Emery, 1895) (Ethiopia, Kenya, Tanzania)

• Messor plinii Santschi, 1912

ceresis Santschi, 1934 (Iran, Lebanon, Saudi Arabia)

chamberlini Wheeler, W.M., 1915 (United States)

chicoensis (Smith, M.R., 1956) (United States)

clypeatus Kuznetsov-Ugamsky, 1927 (Kazakhstan)

collingwoodi Bolton, 1982 (Mali, Niger)

crawleyi Santschi, 1928 (Egypt, Saudi Arabia)

D
decipiens Santschi, 1917 (Botswana, Iraq, Lesotho, Saudi Arabia, South Africa, Zimbabwe)

• Messor arcistriatus Santschi, 1928

• Messor barbarus probus Wheeler, W.M., 1922

dentatus Santschi, 1927 (Iran, Israel)

denticornis Forel, 1910 (Botswana, Namibia, South Africa)

• Messor denticornis brunni Forel, 1910

• Messor denticornis parvidens Forel, 1910

denticulatus Santschi, 1927 (China, Greece, Iran, Israel, Kazakhstan, Kyrgyzstan, Montenegro, The former Yugoslav Republic of Macedonia, Turkmenistan)

desertora He & Song, 2009 (China)

diabarensis Arnoldi, 1970 (Kazakhstan)

E
ebeninus Santschi, 1927 (Greece, Iran, Iraq, Israel, Kuwait, Lebanon, Montenegro, Oman, Saudi Arabia, United Arab Emirates)

eglalae Sharaf, 2007 (Egypt)

erectus Espadaler, 1998 (Morocco)

excursionis Ruzsky, 1905 (China, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Turkmenistan)

excursionis alaschanicus Ruzsky, 1915 (Mongolia)

excursionis sillemi Stärcke, 1935 (Kazakhstan)

F
ferreri Collingwood, 1993 (Kenya)

foreli Santschi, 1923 (Oman, Saudi Arabia, Tunisia, United Arab Emirates)

• Messor aegyptiacus fossulatus Santschi, 1923

foreli brevispinosus Santschi, 1923 (Morocco)

foreli surcoufi Santschi, 1923 (Algeria)

fraternus Ruzsky, 1905 (Russian Federation)

G
galla (Mayr, 1904) (Burkina Faso, Chad, Eritrea, Ethiopia, Ghana, Iran, Ivory Coast, Kenya, Mali, Niger, Nigeria, Oman, Saudi Arabia, Senegal, Somalia, Sudan)

• Messor barbarus latinoda Santschi, 1917

• Messor galla airensis Bernard, 1950

• Messor galla nobilis Santschi, 1928

• Messor galla rufula Finzi, 1939

galla obscurus Menozzi & Consani, 1952 (Ethiopia)

H
hebraeus Santschi, 1927 (Israel)

hellenius Agosti & Collingwood, 1987 (Greece)

himalayanus (Forel, 1902) (Afghanistan, India)

hismai Collingwood & Agosti, 1996 (Saudi Arabia, United Arab Emirates)

hispanicus Santschi, 1919 (Balearic Islands, Iberian Peninsula, Portugal, Spain)

hoggarensis Santschi, 1929 (Algeria)

I
ibericus Santschi, 1931 (Spain)

incisus Stitz, 1923 (Namibia)

incorruptus Kuznetsov-Ugamsky, 1929 (Armenia, Georgia, Iran, Kazakhstan, Russian Federation)

• Messor barbarus hamulifera Karavaiev, 1926

inermis Kuznetsov-Ugamsky, 1929 (China, Kazakhstan, Kyrgyzstan)

instabilis (Smith, F., 1858) (India, Israel)

• Aphaenogaster barbara punctata Forel, 1886

intermedius Santschi, 1927 (Greece, Israel, Oman, Syrian Arab Republic, Turkmenistan)

J
julianus (Pergande, 1894) (Mexico)

julianus clarior (Wheeler, W.M. & Creighton, 1934) (Mexico)

julianus manni (Wheeler, W.M. & Creighton, 1934) (Mexico, United States)

K
kasakorum Arnoldi, 1970 (Kazakhstan)

kasakorum nadezhdae Arnoldi, 1970 (Kazakhstan)

kisilkumensis Arnoldi, 1970 (Turkmenistan, Uzbekistan)

L
lamellicornis Arnoldi, 1968 (Kazakhstan, Turkmenistan)

lariversi (Smith, M.R., 1951) (United States)

lobicornis Forel, 1894 (Algeria, Iberian Peninsula, Portugal, Spain)

lobicornis batnensis Forel, 1909 (Algeria)

lobicornis laurenti Santschi, 1939 (Algeria)

lobicornis normandi Santschi, 1918 (Tunisia)

lobicornis rugulosus Santschi, 1926 (Algeria)

• Messor lobicornis rugosa Karavaiev, 1912

lobicornis submuticus Emery, 1908 (Tunisia)

lobognathus Andrews, 1916 (United States)

luebberti Forel, 1910 (Angola, Botswana, Namibia, South Africa, Tanzania, Zimbabwe)

luridus Santschi, 1927 (Turkey)

luridus virginalis Santschi, 1927 (Syrian Arab Republic)

lusitanicus Tinaut, 1985 (Iberian Peninsula, Portugal, Spain)

M
maculifrons Santschi, 1927 (Israel, Syrian Arab Republic)

marikovskii Arnoldi, 1970 (Kazakhstan, Kyrgyzstan)

marocanus Santschi, 1927 (Iberian Peninsula, Morocco, Portugal, Spain)

medioruber Santschi, 1910 (Iran, Iraq, Kuwait, Saudi Arabia, Tunisia, United Arab Emirates)

• Messor meridionalis sublaeviceps Santschi, 1927

medioruber maurus Santschi, 1927 (Tunisia)

medioruber montanus Santschi, 1927 (Algeria)

medioruber postpetiolatus Santschi, 1917 (Tunisia)

medioruber postquadratus Santschi, 1932 (Algeria)

melancholicus Arnoldi, 1977 (Armenia, Georgia, Kazakhstan)

• Messor melancholicus schelkovnikovi Arnol'di, 1977

meridionalis (André, 1883) (Albania, Georgia, Greece, Iran, Iraq, Israel, Kuwait, Montenegro, Saudi Arabia, The former Yugoslav Republic of Macedonia, United Arab Emirates)

minor (André, 1883) (Iran, Iraq, Israel, Italy, Montenegro, Saudi Arabia)

• Messor minor pompeianus Santschi, 1927

minor calabricus Santschi, 1927 (Italy)

minor capreensis Santschi, 1927 (Italy)

minor hesperius Santschi, 1927 (Canary Islands)

minor laboriosus Santschi, 1927 (Kazakhstan)

minor maurus Barquin, 1981 (Canary Islands)

muraywahus Collingwood & Agosti, 1996 (Saudi Arabia)

muscatus Collingwood & Agosti, 1996 (Oman)

N
nahali Tohmé, G. & Tohmé, H., 1981 (Syrian Arab Republic)

niloticus Santschi, 1938 (Egypt)

nondentatus He & Song, 2009 (China)

O
oertzeni Forel, 1910 (Bulgaria, Greece, The former Yugoslav Republic of Macedonia)

• Messor oertzeni amphigea Forel, 1911

oertzeni amasiensis Emery, 1921 (Turkey)

oertzeni carpathous Menozzi, 1936 (Greece)

olegianus Arnoldi, 1970 (Kyrgyzstan, Tajikistan)

orientalis (Emery, 1898) (China, Greece, Iran, Iraq, Israel, Turkey)

P
perantennatus Arnoldi, 1970 (China, Kazakhstan, Turkmenistan)

pergandei (Mayr, 1886) (Mexico, United States)

piceus Stitz, 1923 (Botswana, Namibia, South Africa)

picturatus Santschi, 1927 (Algeria, Iran, Iraq, Kuwait, Saudi Arabia)

R
regalis (Emery, 1892) (Benin, Congo, Ivory Coast, Nigeria)

• Cratomyrmex regalis rubea Santschi, 1913

• Cratomyrmex sculpturatus Stitz, 1916

reticuliventris Karavaiev, 1910 (Kazakhstan, Turkmenistan)

rufotestaceus (Foerster, 1850) (Algeria, Iran, Israel, Oman, Saudi Arabia, Syrian Arab Republic, United Arab Emirates)

• Aphaenogaster gracilinodis Emery, 1878

• Atta thoracica Mayr, 1862

rufus Santschi, 1923 (Kazakhstan, Kyrgyzstan, Russian Federation)

• Messor clivorum karawajevi Tarbinsky, 1976

• Messor similis Kuznetsov-Ugamsky, 1927

rufus striolatus Kuznetsov-Ugamsky, 1927 (Kyrgyzstan)

ruginodis Stitz, 1916 (Congo)

rugosus (André, 1881) (Israel)

rugosus bodenheimeri Menozzi, 1933 (Israel)

S
sanctus Emery, 1921 (Libya, Tunisia)

sanganus Collingwood & Agosti, 1996 (Yemen)

semirufus (André, 1883) (Iran, Iraq, Israel, Italy)

• Messor semirufus kutteri Santschi, 1934

• Messor semirufus nigricans Santschi, 1932

• Messor semirufus nigriceps Santschi, 1929

semirufus emeryi Santschi, 1927 (Israel)

semirufus grandinidus Emery, 1912 (Tunisia)

semoni (Forel, 1906) (Algeria)

smithi (Cole, 1963) (United States)

stoddardi (Emery, 1895) (Mexico, United States)

striatellus Arnoldi, 1970 (China, Tajikistan)

striaticeps (André, 1883) (Iraq, Kuwait, Saudi Arabia)

striatifrons Stitz, 1923 (Namibia, South Africa)

striativentris Emery, 1908 (Algeria)

striatulus (Emery, 1891) (Tunisia)

structor (Latreille, 1798) (Albania, Andorra, Armenia, Austria, Balearic Islands, Belgium, Bulgaria, Canary Islands, China, Croatia, Czech Republic, France, Georgia, Greece, Iberian Peninsula, Iran, Israel, Italy, Kazakhstan, Kyrgyzstan, Malta, Montenegro, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, The former Yugoslav Republic of Macedonia, Turkmenistan, Ukraine, Uzbekistan)

• Formica aedificator Schilling, 1839

• Formica lapidum Fabricius, 1804

• Formica rufitarsis Fabricius, 1804

• Messor barbarus varrialei Emery, 1921

• Messor clivorum sevani Arnol'di, 1977

• Messor platyceras rubella Crawley, 1920

• Messor rufitarsis darianus Pisarski, 1967

• Messor rufitarsis jakowlevi Arnol'di, 1977

• Messor structor clivorum Ruzsky, 1905

• Messor structor novaki Finzi, 1929

• Messor structor subpolitus Kuznetsov-Ugamsky, 1927

• Messor structor turanicus Kuznetsov-Ugamsky, 1927

• Messor tataricus Ruzsky, 1905

• Myrmica mutica Nylander, 1849

• Stenamma structor tyrrhena Emery, 1898

structor aegaeus Santschi, 1926 (The former Yugoslav Republic of Macedonia, Turkey)

structor platyceras Crawley, 1920 (Iran)

structor tadzhikorum Arnoldi, 1970 (Tajikistan)

subgracilinodis Arnoldi, 1970 (China, Russian Federation, Turkmenistan)

sultanus Santschi, 1917 (Greece, Iran, Israel)

syriacus Tohmé, G., 1969 (Armenia, Iraq, Israel, Saudi Arabia, Syrian Arab Republic)

T
testaceus Donisthorpe, 1950 (Turkey)

timidus Espadaler, 1997 (Iberian Peninsula, Spain)

tropicorum Wheeler, W.M., 1922 (Angola, Namibia)

• Messor braunsi nigriventris Stitz, 1923

• Messor denticornis laevifrons Stitz, 1923

turcmenochorassanicus Arnoldi, 1977 (Kazakhstan, Turkmenistan)

V
valentinae Arnoldi, 1970 (China, Kyrgyzstan, Tajikistan)

variabilis Kuznetsov-Ugamsky, 1927 (Kazakhstan, Turkmenistan)

• Messor excursionis macrops Santschi, 1927

vaucheri Emery, 1908 (Morocco)

vicinus Kuznetsov-Ugamsky, 1927 (Kazakhstan, Turkmenistan)

W
wasmanni Krausse, 1910 (Bulgaria, Croatia, Greece, Iran, Israel, Italy, Lebanon, Malta, Montenegro, Oman, Slovenia, Syrian Arab Republic, The former Yugoslav Republic of Macedonia)

• Messor barbarus concolor Santschi, 1927

=Fossil Species=

S
sculpturatus Carpenter, 1930 (United States)