Checklist of Temnothorax species

The following species and subspecies belong to the genus Temnothorax. Synonyms are listed under their senior names.

A
abeli (Fontenla Rizo, 1998) (Cuba)

adustus (Mackay, 2000) (Mexico)

affinis (Mayr, 1855) (Austria)

• Leptothorax tirolensis Gredler, 1858

• Leptothorax tuberoaffinis Bondroit, 1918

alayoi (Baroni Urbani, 1978) (Cuba)

alayorum (Fontenla Rizo, 1993) (Cuba)

albipennis (Curtis, 1854) (United Kingdom of Great Britain and Northern Ireland)

• Leptothorax tuberointerruptus Bondroit, 1918

albispinus (Wheeler, W.M., 1908) (Puerto Rico)

• Antillaemyrmex albispina pallipes Wheeler, W.M., 1931

algiricus (Forel, 1894) (Algeria)

• Leptothorax angustulus silvanus Forel, 1907

• Leptothorax convexus timida Santschi, 1912

• Leptothorax gazella Santschi, 1923

• Leptothorax gazella monticola Santschi, 1923

algiricus trabutii (Forel, 1894) (Algeria)

• Leptothorax lindbergi Santschi, 1931

alienus Schulz, Heinze & Pusch, 2007 (Italy)

alinae (Radchenko, 1994) (Russia)

allardycei (Mann, 1920) (Bahamas)

• Antillaemyrmex floridanus Wheeler, W.M., 1931

alpinus (Ruzsky, 1902) (Russia)

ambiguus (Emery, 1895) (United States)

ambiguus foveatus (Smith, M.R., 1934) (United States)

ambiguus pinetorum (Wesson, L.G. & Wesson, R.G., 1940) (United States)

anacanthus (Santschi, 1912)

• Leptothorax hesperius inermis Santschi, 1909

anacanthus maximus (Santschi, 1931) (Morocco)

andersoni (Mackay, 2000) (United States)

andrei (Emery, 1895) (United States)

• Leptothorax nitens heathii Wheeler, W.M., 1903

• Leptothorax nitens occidentalis Wheeler, W.M., 1903

• Leptothorax ocellatus Mackay, 2000

androsanus (Wheeler, W.M., 1905) (Bahamas)

angulohumerus Zhou, Huang, Yu & Liu, 2010 (China)

angustulus (Nylander, 1856) (France)

angustulus angustinodis (Stitz, 1917) (Libya)

angustulus bordagei (Bondroit, 1918) (France)

anira (Terayama & Onoyama, 1999) (Japan)

annexus (Baroni Urbani, 1978)

• Macromischa annectens Wheeler, W.M., 1931

annibalis (Santschi, 1909) (Tunisia)

annibalis numidicus (Santschi, 1929) (Tunisia)

anodonta (Arnoldi, 1977) (Armenia)

anodontoides (Dlussky & Zabelin, 1985) (Turkmenia)

antera (Terayama & Onoyama, 1999) (Japan)

antigoni (Forel, 1911) (Turkey)

arcanus (Kutter, 1973) (Switzerland)

archangelskiji (Kuznetsov-Ugamsky, 1926) (Kazakhstan)

arenarius (Santschi, 1908) (Tunisia)

arenarius fusciventris (Santschi, 1911) (Tunisia)

arenarius kruegeri (Kutter, 1928) (Libya)

argentipes (Wheeler, W.M., 1928) (China)

arimensis (Azuma, 1977) (Japan)

arpini (Tarbinsky, 1976) (Kyrgyzstan)

• Leptothorax narinicus Tarbinsky, 1976

artvinensis Seifert, 2006 (Turkey)

atlantis (Santschi, 1911) (Algeria)

• Leptothorax angustulus brunea Santschi, 1918

atomus (Cagniant & Espadaler, 1997) (Morocco)

augusti (Baroni Urbani, 1978)

• Macromischa foreli Aguayo, 1931

auresianus (Santschi, 1929) (Algeria)

aveli (Bondroit, 1918) (France)

• Leptothorax berlandi Bondroit, 1918

• Leptothorax curvithorax Bondroit, 1918

• Leptothorax rabaudi Bondroit, 1918

aveli pici (Bondroit, 1918) (France)

aztecus (Wheeler, W.M., 1931) (Mexico)

• Macromischa azteca maya Wheeler, W.M., 1937

B
baeticus (Emery, 1924) (Spain)

banao (Fontenla Rizo, 1998) (Cuba)

barbouri (Aguayo, 1931) (Cuba)

barroi (Aguayo, 1931) (Cuba)

barryi (Cagniant, 1967) (Algeria)

basara (Terayama & Onoyama, 1999) (Japan)

bermudezi (Wheeler, W.M., 1931) (Cuba)

• Croesomyrmex bermudezi mutabilis Wheeler, W.M., 1931

• Leptothorax anemicus Baroni Urbani, 1978

• Macromischa bermudezi gracilis Aguayo, 1932

bestelmeyeri (Mackay, 2000) (United States)

bicolor (Mackay, 2000) (Mexico)

bikara (Terayama & Onoyama, 1999) (Japan)

bimbache Espadaler, 2007 (Spain (Canary Is))

blascoi (Espadaler, 1997) (Spain)

bradleyi (Wheeler, W.M., 1913) (United States)

brauneri (Ruzsky, 1905) (Caucasus)

brevispinosus (Mackay, 2000) (Mexico)

brevispinus (Chang & He, 2001) (China)

bristoli (Mackay, 2000) (United States)

bruneri (Mann, 1924) (Cuba)

• Macromischa bruneri imitatrix Wheeler, W.M., 1937

• Macromischa bruneri ornatipes Wheeler, W.M., 1937

• Macromischa bruneri proxima Wheeler, W.M., 1937

bucheti (Santschi, 1909) (Morocco)

bucheti festai (Emery, 1924) (Libya)

bucheti mixtus (Santschi, 1929) (Morocco)

bucheti tuneticus (Santschi, 1918) (Tunisia)

bugnioni (Forel, 1894) (Algeria)

bugnioni pupseli (Santschi, 1909) (Tunisia)

bulgaricus (Forel, 1892) (Bulgaria)

bulgaricus aeolius (Forel, 1911) (Turkey)

bulgaricus cypridis (Santschi, 1930) (Cyprus)

bulgaricus smyrnensis (Forel, 1911) (Turkey)

C
cabrerae (Forel, 1893) (Spain (Canary Is))

• Leptothorax cabrerae stitzi Santschi, 1918

caesari (Espadaler, 1997) (Spain)

cagnianti (Tinaut, 1983) (Spain)

canescens (Santschi, 1908) (Spain (Canary Is))

carinatus (Cole, 1957) (United States)

cenatus (Bolton, 1982) (Kenya)

chandleri (Mackay, 2000) (United States)

ciferrii (Menozzi & Russo, 1930) (Dominican Republic)

clypeatus (Mayr, 1853) (Austria)

cokendolpheri (Mackay, 2000) (United States)

coleenae (Mackay, 2000) (United States)

confucii (Forel, 1912) (Taiwan)

congruus (Smith, F., 1874) (Japan)

convexus (Forel, 1894) (Algeria)

• Leptothorax submuticus Emery, 1915

cornibrevis (Collingwood, 1961) (Afghanistan)

corticalis (Schenck, 1852) (Germany)

crassispinus (Karavaiev, 1926) (Ukraine)

• Leptothorax nylanderi slavonicus Seifert, 1995

creightoni (Mann, 1929) (Cuba)

creolus (Baroni Urbani, 1978) (Dominican Republic (Santo Domingo))

crepuscularis (Tinaut, 1994) (Spain)

cristinae (Espadaler, 1997) (Spain)

cuneinodis Radchenko, 2004 (North Korea)

curtulus (Santschi, 1929) (Morocco)

curtulus gentilis (Santschi, 1923) (Tunisia)

curvispinosus (Mayr, 1866) (United States)

• Stenamma gallarum Patton, 1879

cuyagueteje (Fontenla Rizo, 1998) (Cuba)

D
darlingtoni (Wheeler, W.M., 1937) (Cuba)

• Macromischa opalina Wheeler, W.M., 1937

delaparti (Forel, 1890) (Algeria)

• Leptothorax emeryi Santschi, 1923

desertorum Dlussky & Soyunov, 1988 (Russian Federation)

desioi (Menozzi, 1939) (India)

desioi melanicus (Menozzi, 1939) (India)

dessyi (Menozzi, 1936) (Greece)

discoloratus (Arnoldi, 1977) (Russia)

dissimilis (Aguayo, 1932) (Cuba)

• Macromischa chloana Wheeler, W.M., 1937

ditifet (Espadaler, 1997) (Morocco)

duloticus (Wesson, L.G., 1937) (United States)

E
eburneipes (Wheeler, W.M., 1927) (China)

emmae (Mackay, 2000) (United States)

exilis (Emery, 1869) (Italy)

• Leptothorax exilis boccaris Santschi, 1923

• Leptothorax exilis creticus Forel, 1910

• Leptothorax exilis darii Forel, 1911

• Leptothorax exilis nitidulus Santschi, 1923

• Leptothorax tuberum ruficornis Emery, 1914

F
finzii (Menozzi, 1925) (Italy)

flavicornis (Emery, 1870) (Italy)

flavidulus (Wheeler, W.M. & Mann, 1914)

flavispinus (André, 1883)

• Leptothorax nigrita curvispinosus André, 1881

flavispinus amilcaris (Santschi, 1939) (Tunisia)

flavispinus clivulus (Santschi, 1939) (Tunisia)

flavispinus impurus (Santschi, 1939) (Tunisia)

flavispinus rufescens (Forel, 1907) (Tunisia)

foreli (Santschi, 1907) (Tunisia)

• Leptothorax petiolatus Forel, 1901

formosus (Santschi, 1909) (Morocco)

fragosus (Mackay & Buschinger, 2002)

fuentei (Santschi, 1919) (Spain)

fultonii (Forel, 1902) (India)

fumosus (Ruzsky, 1923) (Transcaucasus)

furunculus (Wheeler, W.M., 1909)

fuscatus (Mann, 1920) (Guatemala)

G
gaetulus (Santschi, 1923) (Morocco)

gallae (Smith, M.R., 1949)

gibbifer (Baroni Urbani, 1978) (Cuba)

goniops (Baroni Urbani, 1978) (Mexico)

gracilicornis (Emery, 1882) (Spain (Canary Is))

gracilicornis nivarianus (Santschi, 1925) (Spain (Canary Is))

• Leptothorax nivarianus guancha Santschi, 1925

graecus (Forel, 1911) (Greece)

gredosi (Espadaler & Collingwood, 1982) (Spain)

grouvellei (Bondroit, 1918) (France)

gundlachi (Wheeler, W.M., 1913) (Cuba)

• Croesomyrmex aguayoi Wheeler, W.M., 1931

• Croesomyrmex aguayoi archeri Wheeler, W.M., 1931

• Macromischa aguayoi natenzoni Aguayo, 1931

• Macromischa bierigi Santschi, 1931

H
hadrumetensis (Santschi, 1918) (Tunisia)

haira (Terayama & Onoyama, 1999) (Japan)

hengshanensis (Huang, Chen & Zhou, 2004) (China)

hesperius (Santschi, 1909) (Spain (Canary Is))

himachalensis Bharti, Gul & Schulz, 2012 (India)

hispaniolae (Baroni Urbani, 1978) (Dominican Republic)

hispidus (Cole, 1957) (United States)

huatuo Terayama, 2009 (Taiwan)

huehuetenangoi (Baroni Urbani, 1978) (Guatemala)

I
ibericus (Menozzi, 1922) (Spain)

imias (Fontenla Rizo, 1998) (Cuba)

indra (Terayama & Onoyama, 1999) (Japan)

inermis (Forel, 1902) (India)

interruptus (Schenck, 1852) (Germany)

• Myrmica simpliciuscula Nylander, 1856

interruptus nitidiceps (Dalla Torre, 1893) (Algeria)

iranicus (Radchenko, 1994) (Iran)

iris (Roger, 1863) (Cuba)

• Croesomyrmex iris rufithorax Wheeler, W.M., 1931

• Croesomyrmex iris tristis Wheeler, W.M., 1931

• Macromischa iris nigripes Santschi, 1930

isabellae (Wheeler, W.M., 1908) (Puerto Rico)

• Leptothorax hyperisabellae Baroni Urbani, 1978

• Macromischa isabellae mutica Smith, M.R., 1937

italicus (Consani, 1952) (Italy)

ixili (Baroni Urbani, 1978) (Guatemala)

J
jailensis (Arnoldi, 1977) (Ukraine)

janushevi (Radchenko, 1994) (Tajikistan)

josephi (Mackay, 2000)

K
kashmirensis Bharti, Gul & Schulz, 2012 (India)

kaszabi (Pisarski, 1969) (Mongolia)

• Leptothorax tuberum sachalinensis Kupyanskaya, 1990

kemali (Santschi, 1934) (Turkey)

kinomurai (Terayama & Onoyama, 1999) (Japan)

kirghizicus (Tarbinsky, 1976) (Kyrgyzstan)

kiudiria (Espadaler, 1997) (Morocco)

knipovitshi (Karavaiev, 1916) (Russia)

• Leptothorax junipereti Arnol'di, 1977

• Leptothorax nikitae Arnol'di, 1977

korbi (Emery, 1924) (Transcaucasia)

koreanus (Teranishi, 1940) (Republic of Korea)

kraussei (Emery, 1916) (Italy (Sardinia))

kubira (Terayama & Onoyama, 1999) (Japan)

kuixing Terayama, 2009 (Taiwan)

kurilensis (Radchenko, 1994) (Russia (Kunashir Island))

L
laciniatus (Stitz, 1917) (Algeria)

laestrygon (Santschi, 1931) (Italy (Sicily))

laestrygon splendidiceps (Baroni Urbani, 1968) (Malta)

laetus (Wheeler, W.M., 1937) (Cuba)

• Leptothorax similis Baroni Urbani, 1978

• Macromischa affinis Mann, 1920

• Macromischa affinis maerens Wheeler, W.M., 1937

• Macromischa affinis umbratipes Wheeler, W.M., 1937

lagrecai (Baroni Urbani, 1964) (Italy)

laurae (Emery, 1884) (Tunisia)

laurae antoniae (Cagniant, 1997) (Morocco)

laurae colettae (Santschi, 1915) (Tunisia)

laurae rosae (Santschi, 1915) (Tunisia)

leigong Terayama, 2009 (Taiwan)

leimu Terayama, 2009 (Taiwan)

leoni (Arnoldi, 1971) (Kazakhstan)

lereddei (Bernard, 1953) (Algeria)

leucacanthus (Baroni Urbani, 1978) (Mexico)

leviceps (Emery, 1898) (Italy)

• Leptothorax exilis dichroa Emery, 1916

leyeensis Zhou, Huang, Yu & Liu, 2010 (China)

lichtensteini (Bondroit, 1918) (France)

liebi (Mackay, 2000)

liviae (Agosti & Collingwood, 2011) (United Arab Emirates)

longipilosus (Santschi, 1912) (Tunisia)

longispinosus (Roger, 1863) (United States)

• Leptothorax longispinosus iowensis Buren, 1945

• Leptothorax longispinosus laeviceps Buren, 1944

luteus (Forel, 1874) (Switzerland)

M
makora (Terayama & Onoyama, 1999) (Japan)

manni (Wheeler, W.M., 1914)

maoerensis Zhou, Huang, Yu & Liu, 2010 (China)

marocana (Santschi, 1909) (Morocco)

mauritanicus (Santschi, 1909) (Morocco)

maurus (Santschi, 1921) (Morocco)

megalops (Hamann & Klemm, 1967) (Sudan)

mekira Terayama & Kubota, 2011 (Japan)

melas (Espadaler, Plateaux & Casevitz-Weulersse, 1984) (France (Corsica))

melleus (Forel, 1904) (Uzbekistan)

• Leptothorax balchashensis Arnol'di, 1971

• Leptothorax melleus csikii Pisarski, 1969

melnikovi (Ruzsky, 1905) (Kazan)

mexicanus (Mackay, 2000)

michali Radchenko, 2004 (North Korea)

microreticulatus Bharti, Gul & Schulz, 2012 (India)

mimeuri (Cagniant, 1997) (Morocco)

minozzii (Santschi, 1922)

• Leptothorax silvestrii Menozzi, 1921

minutissimus (Smith, M.R., 1942)

mirabilis (Espadaler & Cagniant, 1996) (Morocco)

miserabilis (Santschi, 1918) (Tunisia)

mongolicus (Pisarski, 1969) (Mongolia)

monjauzei (Cagniant, 1968) (Algeria)

mortoni (Aguayo, 1937) (Cuba)

myersi (Wheeler, W.M., 1931) (Cuba)

N
nadigi (Kutter, 1925) (Switzerland)

• Leptothorax caucasicus Arnol'di, 1977

• Leptothorax hasardaghi Dlussky & Zabelin, 1985

naeviventris (Santschi, 1910) (Tunisia)

naeviventris kefensis (Santschi, 1929) (Tunisia)

nassanovi (Ruzsky, 1895) (Russian Federation)

• Leptothorax galeatus Wheeler, W.M., 1927

• Leptothorax nassonovi firssovi Kuznetsov-Ugamsky, 1928

• Leptothorax nassonovi subnudus Ruzsky, 1905

neminan (Espadaler, 1997) (Morocco)

neomexicanus (Wheeler, W.M., 1903)

nevadensis (Wheeler, W.M., 1903)

• Leptothorax eldoradensis Wheeler, W.M., 1915

• Leptothorax lindae Mackay, 2000

• Leptothorax maryanae Mackay, 2000

• Leptothorax melanderi Wheeler, W.M., 1909

niger (Forel, 1894) (France)

• Leptothorax glaber Santschi, 1925

nigricans (Baroni Urbani, 1978)

• Macromischa melanocephala Wheeler, W.M., 1931

nigriceps (Mayr, 1855) (Austria)

nigriceps pyrenaeus (Bondroit, 1918) (France)

nigritus (Emery, 1878) (Algeria)

nigritus rupestris (Forel, 1909) (Algeria)

nigritus saharensis (Santschi, 1929) (Algeria)

nigritus salambo (Santschi, 1912) (Tunisia)

nipensis (Fontenla Rizo, 1998) (Cuba)

nitens (Emery, 1895)

• Leptothorax melinus Mackay, 2000

• Leptothorax nitens mariposa Wheeler, W.M., 1917

nordmeyeri (Schulz, 1997) (India)

normandi (Santschi, 1912) (Tunisia)

nylanderi (Foerster, 1850) (Germany)

• Leptothorax nylanderi nylanderotuberum Ruzsky, 1902

• Leptothorax tuberum nylanderocorticalis Forel, 1874

• Myrmica cingulata Schenck, 1852

nylanderi nylanderonigriceps (Stitz, 1939) (Germany)

O
obliquicanthus (Cole, 1953)

obscurior (Dalla Torre, 1893) (Algeria)

• Leptothorax exilis retifer Santschi, 1929

obturator (Wheeler, W.M., 1903)

ocarinae (Baroni Urbani, 1978) (Mexico)

opaciabdomin (Chang & He, 2001) (China)

oraniensis (Forel, 1894) (Algeria)

• Leptothorax rusticus Santschi, 1918

• Leptothorax rusticus chobauti Santschi, 1923

• Leptothorax rusticus sergensis Santschi, 1929

orchidus Zhou, Huang, Yu & Liu, 2010 (China)

oxianus (Ruzsky, 1905) (Uzbekistan)

• Leptothorax oxianus juglandeti Arnol'di, 1976

• Leptothorax talassicus Tarbinsky, 1976

oxynodis (Mackay, 2000)

P
pallidipes (Santschi, 1910) (Tunisia)

pallidus (Collingwood, 1961) (Afghanistan)

palustris (Deyrup & Cover, 2004)

pamiricus (Ruzsky, 1902) (Russia)

pan (Santschi, 1936) (Morocco)

pardoi (Tinaut, 1987) (Spain)

parvulus (Schenck, 1852) (Germany)

pastinifer (Emery, 1894) (Bahamas)

• Leptothorax pastoris Baroni Urbani, 1978

• Macromischa lucayensis Forel, 1901

• Macromischa pastinifera opacipes Wheeler, W.M., 1905

pelagosanus (Müller, 1923) (Italy)

peninsularis (Wheeler, W.M., 1934)

pergandei (Emery, 1895)

• Leptothorax floridanus Emery, 1895

• Leptothorax manni Wesson, L.G., 1935

• Leptothorax pergandei flavus Smith, M.R., 1929

personatus (Cagniant, 1987) (Morocco)

peyerimhoffi (Santschi, 1929) (Algeria)

pisarskii Radchenko, 2004 (North Korea)

platycephalus (Espadaler, 1997) (Spain)

platycnemis (Wheeler, W.M., 1937) (Cuba)

poeyi (Wheeler, W.M., 1913) (Cuba)

• Macromischa poeyi rugiceps Aguayo, 1932

politus (Smith, M.R., 1939)

porphyritis (Roger, 1863) (Cuba)

• Macromischa jaumei Santschi, 1930

• Macromischa porphyritis latispina Wheeler, W.M., 1931

productus (Santschi, 1918) (Morocco)

pulchellus (Emery, 1894) (St)

punctaticeps (Mackay, 2000)

punctatissimus (Mackay, 2000)

punctithorax (Mackay, 2000)

punicans (Roger, 1863) (Cuba)

purpuratus (Roger, 1863) (Cuba)

R
racovitzai (Bondroit, 1918) (France)

• Leptothorax massiliensis Bondroit, 1918

recedens (Nylander, 1856) (France)

• Leptothorax ergatogyna Bernard, 1950

• Leptothorax mordax Santschi, 1919

• Leptothorax pictus Emery, 1924

• Leptothorax recedens barbarus Santschi, 1939

• Temnothorax rogeri Emery, 1869

reduncus (Wang & Wu, 1988) (China)

reticulatus (Chang & He, 2001) (China)

risii (Forel, 1892) (Spain (Canary Is))

• Leptothorax elongatus Santschi, 1919

• Leptothorax risii armouri Wheeler, W.M., 1927

• Leptothorax risii globulinodis Santschi, 1909

rothneyi (Forel, 1902) (India)

rothneyi simlensis (Forel, 1904) (India)

rottenbergii (Emery, 1870) (Italy (Sicily))

rottenbergii ernesti (Santschi, 1909) (Algeria)

rottenbergii jesus (Forel, 1913) (Israel)

rottenbergii scabriosus (Santschi, 1909) (Algeria)

rudis (Wheeler, W.M., 1917)

rugatulus (Emery, 1895)

• Leptothorax curvispinosus annectens Wheeler, W.M., 1903

• Leptothorax rugatulus brunnescens Wheeler, W.M., 1917

• Leptothorax rugatulus cockerelli Wheeler, W.M., 1917

• Leptothorax rugatulus dakotensis Wheeler, G.C. & Wheeler, E.W., 1944

• Leptothorax rugatulus mediorufus Wheeler, W.M., 1917

ruginosus Zhou, Huang, Yu & Liu, 2010 (China)

rugithorax (Mackay, 2000)

rugosus (Mackay, 2000)

rugulosus (Mackay, 2000)

S
sallei (Guérin-Méneville, 1852) (Haiti)

• Macromischa sallei haytiana Wheeler, W.M. & Mann, 1914

• Macromischa sallei opacinoda Wheeler, W.M., 1931

• Myrmica russula Nylander, 1856

salvini (Forel, 1899) (Panama)

salvini obscurior (Forel, 1899) (Panama)

santra (Terayama & Onoyama, 1999) (Japan)

santschii (Forel, 1905) (Tunisia)

• Leptothorax flavispinus boiteli Santschi, 1929

• Leptothorax flavispinus facetus Santschi, 1939

sardous (Santschi, 1909) (Italy (Sardinia))

satunini (Ruzsky, 1902) (Caucasus)

saudiae (Collingwood & Agosti, 1996) (Saudi Arabia)

saxatilis Schulz, Heinze & Pusch, 2007 (Italy)

saxonicus (Seifert, 1995) (Germany)

schaufussi (Forel, 1879) (Spain)

schaumii (Roger, 1863)

• Leptothorax fortinodis Mayr, 1886

• Leptothorax fortinodis gilvus Wheeler, W.M., 1903

• Leptothorax fortinodis melanoticus Wheeler, W.M., 1903

schmittii (Wheeler, W.M., 1903)

schoedli Seifert, 2006 (Turkey)

schurri (Forel, 1902) (India)

schwarzi (Mann, 1920) (Cuba)

semenovi (Ruzsky, 1903) (Transcaspia)

semiruber (André, 1881) (Israel)

• Leptothorax rottenbergi balcanica Santschi, 1909

senectutis (Baroni Urbani, 1978) (Cuba)

serviculus (Ruzsky, 1902) (Russia)

sevanensis (Arnoldi, 1977) (Armenia)

shannxiensis Zhou, Huang, Yu & Liu, 2010 (China)

shelkovnikovi (Karavaiev, 1926) (Caucasus)

silvestrii (Santschi, 1911)

simesno (Espadaler, 1997) (Morocco)

singularis (Radchenko, 1994) (Tajikistan)

skwarrae (Wheeler, W.M., 1931) (Mexico)

smithi (Baroni Urbani, 1978)

• Leptothorax wheeleri Smith, M.R., 1929

solerii (Menozzi, 1936) (Greece)

sordidulus (Müller, 1923) (Italy)

• Leptothorax carinthiacus Bernard, 1957

• Leptothorax mulleri Santschi, 1926

• Leptothorax sordidulus tergestina Finzi, 1928

specularis (Emery, 1916) (Italy)

spinosior (Forel, 1901) (Japan)

spinosus (Forel, 1909) (Algeria)

splendens (Wheeler, W.M., 1905) (Bahamas)

• Macromischa scabripes Mann, 1920

• Macromischa scabripes brunneipes Wheeler, W.M., 1937

squamifer (Roger, 1863) (Cuba)

• Macromischa squamifera atrinodis Mann, 1920

steinbergi (Arnoldi, 1971) (Kazakhstan)

stenotyle (Cole, 1956)

• Leptothorax angustinodus Cole, 1956

stollii (Forel, 1885)

• Leptothorax denticulatus Stitz, 1917

striatulus (Stitz, 1937)

striatus Zhou, Huang, Yu & Liu, 2010 (China)

subcingulatus (Emery, 1924) (Spain)

subditivus (Wheeler, W.M., 1903)

• Macromischa flavitarsis Mann, 1920

• Macromischa laevissima Wheeler, W.M., 1911

• Macromischa luciliae Mann, 1935

suberis (Forel, 1894) (Algeria)

susamyri (Dlussky, 1965) (Kyrgyzstan)

T
taivanensis (Wheeler, W.M., 1929) (Taiwan)

tamarae (Radchenko, 1994) (Georgia)

tauricus (Ruzsky, 1902) (Ukraine)

• Leptothorax pithyusae Arnol'di, 1977

• Leptothorax pithyusae babuganicus Arnol'di, 1977

tebessae (Forel, 1890) (Algeria)

tebessae gapsensis (Santschi, 1929) (Tunisia)

tenuisculptus (Baroni Urbani, 1978) (Mexico)

tenuispinus (Santschi, 1911) (Tunisia)

terricola (Mann, 1920) (Cuba)

terrigena (Wheeler, W.M., 1903)

tesquorum (Arnoldi, 1977) (Georgia)

• Leptothorax elderensis Dlussky & Zabelin, 1985

texanus (Wheeler, W.M., 1903) (United States)

• Leptothorax texanus davisi Wheeler, W.M., 1905

theryi (Santschi, 1921) (Morocco)

tianpeng Terayama, 2009 (Taiwan)

tianschanicus (Tarbinsky, 1976) (Kyrgyzstan)

torrei (Aguayo, 1931) (Cuba)

totonicapani (Baroni Urbani, 1978) (Guatemala)

tricarinatus (Emery, 1895)

tricolor (Santschi, 1925) (Morocco)

tristis (Bondroit, 1918) (France)

tuberum (Fabricius, 1775) (Sweden)

• Formica tuberosa Latreille, 1802

• Leptothorax melanocephalus Emery, 1870

• Leptothorax pratostepposus Arnol'di, 1977

• Leptothorax tuberum acutinodis Arnol'di, 1977

• Leptothorax tuberum alaicus Tarbinsky, 1976

• Leptothorax tuberum ciscaucasicus Arnol'di, 1977

• Leptothorax tuberum nigricephala Karavaiev, 1930

• Leptothorax tuberum stipaceus Ruzsky, 1905

tuberum nylanderounifasciatus (Forel, 1874) (Switzerland)

tuberum tuberonigriceps (Forel, 1874) (Switzerland)

turcicus (Santschi, 1934) (Turkey)

turritellus (Espadaler, 1997) (Morocco)

tuscaloosae (Wilson, 1951)

tyndalei (Forel, 1909) (Algeria)

• Leptothorax balli Santschi, 1939

• Leptothorax cervantesi Santschi, 1919

U
unifasciatus (Latreille, 1798) (France)

• Leptothorax anoplogynus Emery, 1869

• Leptothorax cordieri Bondroit, 1918

• Leptothorax tuberum brauneri Karavaiev, 1937

• Leptothorax tuberum brauniella Radchenko, 1995

• Leptothorax tuberum kirillovi Ruzsky, 1905

• Leptothorax tuberum salina Karavaiev, 1937

• Leptothorax tuberum unifasciatointerruptus Forel, 1874

• Leptothorax unifasciatus ucrainicus Arnol'di, 1977

• Leptothorax volubilis Santschi, 1929

unifasciatus obenbergeri (Sadil, 1939) (Czechoslovakia)

unifasciatus rougeti (Bondroit, 1918) (France)

unifasciatus staegeri (Bondroit, 1918) (Switzerland)

unifasciatus unifasciatonigriceps (Novak & Sadil, 1941) (Germany)

universitatis (Espadaler, 1997) (Spain)

usunkul (Ruzsky, 1924) (Russia)

V
versicolor (Roger, 1863) (Cuba)

• Macromischa lugens Roger, 1863

• Macromischa lugens nigra Santschi, 1931

versicolor guanahacabibes (Fontenla Rizo, 1995) (Cuba)

villarensis (Aguayo, 1931) (Cuba)

• Macromischa manni Wheeler, W.M., 1931

• Leptothorax williami Baroni Urbani, 1978

violaceus (Mann, 1924) (Cuba)

volgensis (Ruzsky, 1905) (Russia)

W
werneri (Radchenko, 1994) (Caucasus)

wheeleri (Mann, 1920) (Cuba)

• Macromischa wheeleri petri Aguayo, 1931

whitfordi (Mackay, 2000)

wollastoni (Donisthorpe, 1940) (Portugal (Madeira Island))

wroughtonii (Forel, 1904) (India)

wui (Wheeler, W.M., 1929) (China)

X
xanthos Radchenko, 2004 (North Korea)

Y
yanwan Terayama, 2009 (Taiwan)

Z
zhejiangensis Zhou, Huang, Yu & Liu, 2010 (China)