Camponotus maculatus

Well known, common and widely distributed throughout Africa. A common ground nesting and foraging ant which often ascends trees to tend aphids, but does not build tents (Taylor 1977). It was not among the commoner species in the 76-farm survey of Taylor & Adedoyin (1978). Known to be prey of Oecophylla longinoda (Taylor et al., 2018). While this species has been suggested as being invasive in may parts of the world, these reports appear to be largely the result of misidentifications or are presented with little or no justification; additional details are required to confirm its occurrence outside its native range.

Distribution based on Regional Taxon Lists
Afrotropical Region: Angola, Benin, Cameroun, Central African Republic, Chad, Comoros, Eritrea, Ethiopia, Gambia, Guinea, Ivory Coast, Kenya, Lesotho, Liberia, Mali, Mozambique, Namibia, Saudi Arabia, Senegal, Sierra Leone, South Africa, Sudan, São Tomé & Principe, Uganda, United Republic of Tanzania, Zimbabwe. Indo-Australian Region: Micronesia (Federated States of), Palau. Malagasy Region: Madagascar, Mauritius, Mayotte, Réunion. Palaearctic Region: Algeria, Iran, Oman, Turkey.

Nomenclature

 * . Formica maculata Fabricius, 1782: 491 (w.) (no state data,”Habitat in Africa aequinoctiali”).
 * Smith, F. 1858b: 29 (s.w.); Mayr, 1862: 654 (q.m.); Forel, 1891b: 29 (s.w.q.m.).
 * Combination in Camponotus: Mayr, 1862: 654;
 * combination in C. (Myrmoturba): Forel, 1912i: 91;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 86.
 * Subspecies of sylvaticus: Forel, 1879a: 64; Emery & Forel, 1879: 448; Mayr, 1880: 21; Emery, 1881b: 526; André, 1882a: 145; André, 1884b: 536; Emery, 1884a: 375; Emery, 1887a: 212 (footnote).
 * Subspecies of rubripes: Forel, 1886f: 143, 149; Emery, 1887a: 212 (footnote).
 * Status as species: Fabricius, 1787: 308; Gmelin, 1790: 2800; Olivier, 1792: 495; Fabricius, 1793: 356; Latreille, 1802c: 283; Fabricius, 1804: 403; Lepeletier de Saint-Fargeau, 1835: 214; Haliday, 1836: 329; Smith, F. 1858b: 28 (redescription); Gerstäcker, 1859: 262; Mayr, 1862: 654 (redescription); Gerstäcker, 1862: 509; Mayr, 1863: 400; Roger, 1863b: 2; Mayr, 1865: 27; Gerstäcker, 1871: 355; Mayr, 1877: 20 (in list); Emery, 1877b: 364; Emery, 1878b: 44; Forel, 1891b: 23, 29 (redescription); Emery, 1891b: 17; Forel, 1892e: 349; Emery, 1892a: 119; Emery, 1892d: 566; Dalla Torre, 1893: 241; Emery, 1893i: 671; Mayr, 1893: 195; Mayr, 1895: 150; Emery, 1895a: 180; Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1896i: 158; Mayr, 1896: 249; Mayr, 1897: 432; Emery, 1897e: 604; Emery, 1898a: 225; Forel, 1899c: 134; Emery, 1899a: 501; Mayr, 1901b: 28; Mayr, 1904b: 11; Emery, 1905d: 37; Ruzsky, 1905b: 191; Mayr, 1907a: 389; Mayr, 1907b: 19; Forel, 1907e: 16; Forel, 1907g: 88; Forel, 1908a: 16; Emery, 1908a: 188 (infraspecific taxa key); Santschi, 1908: 518; Bondroit, 1910: 487; Forel, 1910c: 266; Forel, 1910f: 26; Karavaiev, 1910b: 9; Stitz, 1910: 148; Wheeler, W.M. 1910d: 301; Stitz, 1911b: 392; Forel, 1911d: 371; Viehmeyer, 1912: 8; Forel, 1913a: 145; Forel, 1913b: 340; Santschi, 1913h: 42; Santschi, 1914b: 129; Santschi, 1914c: 288; Santschi, 1914d: 379; Forel, 1914d: 249; Wheeler, W.M. & Mann, 1914: 47; Donisthorpe, 1915a: 221 (redescription of holotype); Wheeler, W.M. 1915g: 814; Forel, 1915d: 70 (in key); Emery, 1915f: 79; Emery, 1916b: 224; Stitz, 1916: 403; Mann, 1916: 474; Stitz, 1917: 351; Luederwaldt, 1918: 49; Mann, 1920: 435; Emery, 1920c: 5, 12; Arnold, 1922: 621; Wheeler, W.M. 1922a: 234, 949, 1040; Soudek, 1922: 91; Viehmeyer, 1923: 93; Stitz, 1923: 164; Wheeler, W.M. 1924b: 253; Wheeler, W.M. 1925b: 3; Emery, 1925b: 86; Santschi, 1925h: 166; Crawley, 1926: 393 (redescription); Menozzi, 1926a: 41; Karavaiev, 1926e: 189; Arnold, 1926: 292; Karavaiev, 1927a: 294; Santschi, 1932f: 519; Finzi, 1936: 189; Santschi, 1936e: 81; Finzi, 1939a: 159; Finzi, 1940: 163; Weber, 1943c: 385; Donisthorpe, 1946e: 33; Chapman & Capco, 1951: 248; Bernard, 1953b: 260; Prins, 1963: 106; Baroni Urbani, 1972: 125; Hamann & Klemm, 1976: 676; Collingwood, 1985: 282; Taylor & Brown, 1985: 116; Marsh, 1986: 342; Bolton, 1995b: 109; Collingwood & Agosti, 1996: 374; Blard, et al. 2003: 129; Paknia, et al. 2008: 153; Mackay & Delsinne, 2009: 496 (in key); Legakis, 2011: 31; Borowiec, L. & Salata, 2012: 480; Hita Garcia, et al. 2013: 205; Borowiec, L. 2014: 36 (see note in bibliography); Madl, 2019: 14.
 * Senior synonym of atramentarius: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of ballioni: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of boera: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of cavallus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of cluisoides: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of cognata: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
 * Senior synonym of conakryensis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of erythraea: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of flavifemur: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of flavominor: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of hannae: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of hieroglyphicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of intonsus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of lacteipennis: Baroni Urbani, 1972: 127; Bolton, 1995b: 110.
 * Senior synonym of liengmei: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of liocnemis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of lividior: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of lohieri: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of manzer: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of mathildae: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of melanocnemis: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of miserabilis: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of nubis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of proletaria: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of radamoides: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of sarmentus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of schereri: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of schultzei: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of semispicatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * Senior synonym of sudanicus: Baroni Urbani, 1972: 126; Bolton, 1995b: 110.
 * Senior synonym of thomensis: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of tuckeri: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Senior synonym of zumpti: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * Material of the unavailable names calceatus, cataractae, citinus, cluis, contaminatus, diffusus, hansingi, importunoides, madecassa, pessimus referred here by Baroni Urbani, 1972: 124-127; Bolton, 1995b: 110.
 * Current subspecies: nominal plus foveolatus (unresolved junior homonym), miserabilis (unresolved junior homonym), obfuscatus, subnudus, sylvaticomaculatus, ugandensis.
 * atramentarius. Camponotus maculatus var. atramentarius Forel, 1904b: 379 (w.) ETHIOPIA.
 * Forel, 1910f: 26 (q.).
 * Combination in C. (Myrmoturba): Forel, 1913b: 340;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Forel, 1905b: 180; Forel, 1907e: 14; Forel, 1907g: 88; Forel, 1909b: 54, 67; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1911f: 280; Forel, 1913b: 340; Santschi, 1914b: 131; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 952; Stitz, 1923: 164; Emery, 1925b: 88; Santschi, 1930b: 75; Santschi, 1937d: 240; Bernard, 1953b: 261.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 86.
 * ballioni. Camponotus maculatus r. ballioni Forel, 1904d: 176 (w.q.) SOUTH AFRICA.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
 * As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 954.
 * Status as species: Santschi, 1925h: 166.
 * Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 87.
 * boera. Camponotus (Myrmoturba) ballioni var. boera Santschi, 1925h: 166.
 * [First available use of Camponotus maculatus subsp. cognatus var. boera Forel, 1910f: 27 (s.w.q.) SOUTH AFRICA; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Emery, 1920c: 5; Arnold, 1922: 632; Wheeler, W.M. 1922a: 954; Emery, 1925b: 87.
 * Subspecies of ballioni: Santschi, 1930b: 57.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 89.
 * cavallus. Camponotus maculatus st. cavallus Santschi, 1911g: 211 (s.w.q.) ANGOLA.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Stitz, 1916: 403; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Emery, 1925b: 88; Santschi, 1930b: 57; Santschi, 1937d: 241.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 91.
 * cluisoides. Camponotus (Myrmoturba) maculatus var. cluisoides Forel, 1913h: 354 (s.w.) KENYA.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 87.
 * As unavailable (infrasubspecific) name: Emery, 1920c: 5; Wheeler, W.M. 1922a: 952.
 * Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1927f: 361.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
 * cognata. Formica cognata Smith, F. 1858b: 35 (s.w.) SOUTH AFRICA.
 * Mayr, 1862: 655 (q.m.).
 * Combination in Camponotus: Mayr, 1862: 655;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * As unavailable (infrasubspecific) name: Emery, 1894i: 46; Emery, 1895h: 46.
 * Status as species: Mayr, 1877: 20 (in list); Dalla Torre, 1893: 225.
 * Subspecies of sylvaticus: Forel, 1879a: 65; Emery & Forel, 1879: 448; Mayr, 1880: 22; Emery, 1881b: 526; André, 1882a: 146; Emery, 1882: 448; Emery, 1884a: 375; André, 1884b: 536; Forel, 1885b: 176; Forel, 1886d: 239; Forel, 1886e: clxvii; Medina, 1893: 104.
 * Subspecies of rubripes: Forel, 1886f: 143, 151; Emery, 1887a: 212 (footnote); Mayr, 1889: 278; Forel, 1889: 255; Forel, 1890a: lxv; Saunders, E. 1890: 202.
 * Subspecies of maculatus: Mayr, 1862: 655; Emery, 1891b: 19; Emery, 1892d: 566; Emery, in Dalla Torre, 1893: 225, 241 (footnote); Mayr, 1893: 195; André, 1895a: 1; Mayr, 1896: 249; Forel, 1904b: 378; Forel, 1904c: 12; Ruzsky, 1905b: 205; Stitz, 1910: 148; Forel, 1910f: 25; Emery, 1920c: 5; Wheeler, W.M. 1922a: 953; Arnold, 1922: 631; Emery, 1925b: 88; Menozzi, 1942: 180; Eidmann, 1944: 469.
 * Junior synonym of maculatus: Roger, 1863b: 2; Mayr, 1863: 400; Mayr, 1865: 27; Gerstäcker, 1871: 355; Baroni Urbani, 1972: 127; Bolton, 1995b: 93.
 * conakryensis. Camponotus (Myrmoturba) maculatus var. conakryensis Emery, 1920c: 13 (s.w.) GUINEA.
 * Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1930a: 73.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 93.
 * erythraea. Camponotus (Myrmoturba) maculatus var. erythraea Emery, 1920c: 14 (s.) ERITREA.
 * Subspecies of maculatus: Emery, 1925b: 87; Santschi, 1935b: 275.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 98.
 * flavifemur. Camponotus (Myrmoturba) maculatus var. flavifemur Santschi, 1937g: 84 (w.q.) TANZANIA.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 99.
 * flavominor. Camponotus (Tanaemyrmex) maculatus var. flavominor Emery, 1925b: 87.
 * [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. flavominor Santschi, 1920i: 4 (s.w.) BENIN (footnote); unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 956.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 100.
 * hannae. Camponotus (Myrmoturba) maculatus st. hannae Santschi, 1919a: 349, fig. 4 (s.w.) ZIMBABWE.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88; Arnold, 1926: 294.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 102.
 * hieroglyphicus. Camponotus maculatus st. hieroglyphicus Santschi, 1917b: 290 (s.w.q.) ANGOLA.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Emery, 1920c: 5; Wheeler, W.M. 1922a: 955; Emery, 1925b: 88.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 103.
 * intonsus. Camponotus maculatus subsp. intonsus Emery, 1905d: 29 (footnote) (s.w.) SOUTH AFRICA.
 * As unavailable (infrasubspecific) name: Emery, 1920c: 5; Emery, 1925b: 87; Santschi, 1926b: 249.
 * Subspecies of maculatus: Arnold, 1922: 632; Wheeler, W.M. 1922a: 955.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 105.
 * lacteipennis. Formica lacteipennis Smith, F. 1858b: 34 (s.w.q.m.) SOUTH AFRICA.
 * Combination in Camponotus: Roger, 1863b: 2;
 * combination in C. (Myrmoturba): Forel, 1914a: 267;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 90.
 * Status as species: Mayr, 1863: 416; Roger, 1863b: 2; Dalla Torre, 1893: 237.
 * Subspecies of sylvaticus: Mayr, 1886c: 355.
 * Subspecies of maculatus: Emery, 1895h: 46; Emery, 1896d: 370 (in list); Emery, 1920c: 5; Arnold, 1922: 626; Wheeler, W.M. 1922a: 955; Emery, 1925b: 89.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 127; Bolton, 1995b: 107.
 * liengmei. Camponotus sexpunctatus var. liengmei Forel, 1894b: 67 (s.w.) MOZAMBIQUE.
 * Forel, 1907g: 88 (m.).
 * Combination in C. (Myrmoturba): Forel, 1914a: 267;
 * combination in C. (Dinomyrmex): Santschi, 1921f: 312;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 87.
 * As unavailable (infrasubspecific) name: Emery, 1895h: 46; Emery, 1896d: 371 (in list.
 * Subspecies of maculatus: Forel, 1901h: 69; Forel, 1907g: 88; Forel, 1910e: 452; Forel, 1911e: 288; Forel, 1914d: 249; Emery, 1920c: 5; Arnold, 1922: 636; Wheeler, W.M. 1922a: 955; Emery, 1925b: 87; Prins, 1963: 107.
 * Status as species: Santschi, 1923e: 292.
 * Junior synonym of maculatus Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
 * liocnemis. Camponotus maculatus subsp. liocnemis Emery, 1905d: 30 (footnote) (s.w.) SOUTH AFRICA.
 * [Misspelled as lioculmis by Forel, 1905b: 180 (in text); misspelled as liochnemis by Santschi, 1914e: 38.]
 * Santschi, 1914e: 38 (q.m.).
 * Combination in C. (Myrmoturba): Santschi, 1914d: 379.
 * As unavailable (infrasubspecific) name: Forel, 1907e: 16; Forel, 1909b: 54; Forel, 1910f: 26; Forel, 1911d: 371; Forel, 1913b: 340; Santschi, 1917b: 293; Emery, 1920c: 5; Arnold, 1922: 625; Wheeler, W.M. 1922a: 958; Emery, 1925b: 88; Santschi, 1925h: 166; Santschi, 1930a: 73; Santschi, 1930b: 76.
 * Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1914e: 38.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 108.
 * lividior. Camponotus maculatus var. lividior Santschi, 1911e: 128 (s.w.q.m.) COMORO IS.
 * Subspecies of maculatus: Emery, 1920c: 5; Emery, 1925b: 88.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 108.
 * lohieri. Camponotus (Myrmoturba) maculatus var. lohieri Emery, 1915g: 22.
 * [First available use of Camponotus (Myrmoturba) maculatus st. melanocnemis var. lohieri Santschi, 1913c: 313 (w.) IVORY COAST; unavailable (infrasubspecific) name.]
 * As unavailable (infrasubspecific) name: Santschi, 1914d: 379; Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
 * Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1942: 180; Eidmann, 1944: 463.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 109.
 * manzer. Camponotus maculatus subsp. manzer Forel, 1910e: 452 (s.) ZIMBABWE.
 * Forel, 1911f: 280 (w.); Santschi, 1916b: 509 (m.).
 * Combination in C. (Myrmoturba): Santschi, 1916b: 509;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Forel, 1911f: 280; Santschi, 1916b: 509; Santschi, 1917b: 291; Emery, 1920c: 5; Arnold, 1922: 624; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88; Santschi, 1930b: 76.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * mathildae. Camponotus maculatus subsp. mathildae Forel, 1910c: 266 (s.w.q.) SOUTH AFRICA.
 * Combination in C. (Myrmoturba): Santschi, 1914b: 130;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * As unavailable (infrasubspecific) name: Arnold, 1922: 649.
 * Subspecies of maculatus: Santschi, 1914b: 130; Emery, 1920c: 5; Wheeler, W.M. 1922a: 956; Emery, 1925b: 88.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 110.
 * melanocnemis. Camponotus maculatus var. melanocnemis Santschi, 1911c: 368 (s.) DEMOCRATIC REPUBLIC OF CONGO.
 * Santschi, 1915c: 278 (w.).
 * As unavailable (infrasubspecific) name: Emery, 1920c: 5.
 * Subspecies of maculatus: Forel, 1913h: 355; Santschi, 1913c: 313; Forel, 1914d: 250; Santschi, 1914d: 379; Santschi, 1915c: 278; Santschi, 1920i: 4 (footnote); Wheeler, W.M. 1922a: 234, 956; Emery, 1925b: 87; Weber, 1943c: 333; Bernard, 1953b: 260.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 111.
 * miserabilis. Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379 (s.w.) GUINEA.
 * [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
 * Subspecies of maculatus: Santschi, 1915c: 281; Emery, 1920c: 5; Wheeler, W.M. 1922a: 219, 957; Emery, 1925b: 88.
 * Replacement name: Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126.
 * nubis. Camponotus (Myrmoturba) maculatus subsp. nubis Weber, 1943c: 385, pl. 16, fig. 30 (s.w.) SOUTH SUDAN.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 114.
 * proletaria. Camponotus maculatus subsp. proletaria Baroni Urbani, 1971b: 362.
 * Replacement name for Camponotus (Myrmoturba) maculatus st. miserabilis Santschi, 1914d: 379. [Junior primary homonym of *Camponotus miserabilis Förster, 1891: 430.]
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 118.
 * radamoides Camponotus maculatus var. radamoides'' Forel, 1891b: 213 (s.w.) MADAGASCAR.
 * Combination in C. (Myrmoturba): Santschi, 1917b: 292;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * As unavailable (infrasubspecific) name: Santschi, 1911e: 131 (caption of fig. 3d).
 * Subspecies of maculatus: Santschi, 1917b: 292; Santschi, 1919b: 239; Emery, 1920c: 5; Santschi, 1920g: 180; Arnold, 1922: 626; Wheeler, W.M. 1922a: 958, 1042; Emery, 1925b: 88; Santschi, 1925h: 166; Arnold, 1926: 292; Santschi, 1930a: 73; Santschi, 1930b: 76; Santschi, 1935a: 279; Santschi, 1937d: 240.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 119.
 * sarmentus. Camponotus (Myrmoturba) maculatus subsp. sarmentus Emery, 1920c: 14, fig. 2 (s.w.) TANZANIA.
 * Combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Emery, 1925b: 88.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
 * schereri. Camponotus maculatus subsp. schereri Forel, 1911e: 289 (w.q.) LIBERIA.
 * Combination in C. (Myrmoturba): Santschi, 1915c: 280;
 * combination in C. (Tanaemyrmex): Emery, 1925b: 88.
 * Subspecies of maculatus: Santschi, 1914d: 379; Santschi, 1915c: 280; Emery, 1920c: 5; Wheeler, W.M. 1922a: 959; Emery, 1925b: 88.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 122.
 * schultzei. Camponotus maculatus var. schultzei Forel, 1912j: 179 (s.w.m.) UGANDA.
 * Menozzi, 1933a: 110 (q.).
 * As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
 * Subspecies of maculatus: Santschi, 1914b: 130; Menozzi, 1933a: 110; Menozzi, 1939c: 106.
 * Junior synonym of melanocnemis: Forel, 1914d: 250; Emery, 1925b: 87.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 110.
 * schulzi. Camponotus schulzi Salata, Loss, et al. 2019: 95, figs. 1-10, 35 (s.w.) TURKEY.
 * [Note: subgenus C. (Myrmentoma).]
 * semispicatus. Camponotus (Tanaemyrmex) maculatus var. semispicatus Emery, 1925b: 87.
 * [First available use of Camponotus (Myrmoturba) maculatus st. mathildae var. semispicata Santschi, 1914b: 130 (w.) KENYA; unavailable (infrasubspecific) name.]
 * Menozzi, 1930b: 116 (q.).
 * As unavailable (infrasubspecific) name: Santschi, 1915c: 279; Emery, 1920c: 5; Wheeler, W.M. 1922a: 957.
 * Subspecies of maculatus: Emery, 1925b: 87; Menozzi, 1930b: 115; Menozzi, 1939c: 107; Menozzi & Consani, 1952: 69; Bernard, 1953b: 261.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 124; Bolton, 1995b: 123.
 * sudanicus. Camponotus (Myrmoturba) maculatus subsp. sudanicus Weber, 1943c: 385 (s.w.) SOUTH SUDAN.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 126; Bolton, 1995b: 125.
 * thomensis. Camponotus (Myrmoturba) maculatus var. thomensis Santschi, 1920i: 3 (s.w.m.) SÃO TOMÉ & PRÍNCIPE (São Tomé I.).
 * [Misspelled as thomeensis by Santschi, 1939f: 167.]
 * Subspecies of maculatus: Wheeler, W.M. 1922a: 951; Emery, 1925b: 87; Santschi, 1939f: 167.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 127.
 * tuckeri. Camponotus (Myrmoturba) maculatus var. tuckeri Santschi, 1932a: 391 (s.w.m.) NAMIBIA.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 128.
 * zumpti. Camponotus (Myrmoturba) maculatus var. zumpti Santschi, 1937b: 103 (w.) CAMEROON.
 * Junior synonym of maculatus: Baroni Urbani, 1972: 125; Bolton, 1995b: 130.

Taxonomic Notes
Taylor et al. (2018): Camponotus lohieri Santschi, 1913, new status (here, originally described as Camponotus (Myrmoturba) maculatus Fab. stirps melanocnemis Sants. v. lohieri n. var. Santschi, 1913; raised status to Camponotus maculatus F., var. lohieri Sant., Emery, 1915g: 22), CASENT0905268), Haut-Dahomey, DESANTI.

This treatment of C. lohieri as a valid species by Taylor et al. (2018) was proposed without comment or justification, has has not been accepted by subsequent authors.

Taylor et al. (2018) treat Camponotus proletaria as a full, distinct species but provide no justification for the change. Subsequent authors have not followed this treatment.

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