Brachymyrmex heeri

 Brachymyrmex heeri nests under stones and other objects, often in disturbed areas, as well as in bamboo (Fagundes et al., 2010).

Identification
Small workers (1.2 to 2mm) with nine segmented antenna. Color varies from yellow to brown.

Distribution
Central America, South America and the Caribbean. Also introduced in a few locations in Europe and in the Galapagos Islands.

Distribution based on Regional Taxon Lists
Neotropical Region: Argentina, Bolivia, Brazil , Colombia, Costa Rica, Ecuador, French Guiana, Galapagos Islands, Guatemala, Guyana, Haiti, Lesser Antilles, Mexico, Panama, Paraguay, Puerto Rico, Uruguay, Venezuela. Palaearctic Region: Switzerland, Ukraine.

Greater Antilles
In Haiti this species was "found to be very common beneath stones, boards etc. on the shores of the salty Lake Assuei (Haiti)" (Wheeler and Mann 1914).

Wheeler (1908) found "several small colonies under stones" in Puerto Rico and in the Bahamas (Wheeler 1905) discovered "a single colony, including several winged females, of the typical form of this species.under a stone on a key in the Southern Bight, Andros Island, May 23 (1904)."

Costa Rica
Longino 2004 - This is a common species of synanthropic habitats in Costa Rica. It can be found in small parks in the middle of San José, in hotel landscaping, along road edges, in scrubby second growth vegetation, and in pastures. It occurs in almost any bioclimatic region: dry Guanacaste lowlands, wet Atlantic slope lowlands, Central Valley urban areas, and roads and pastures near Monteverde cloud forest. Nests are often under stones on the ground but also occur in cavities in low vegetation. Colonies are polygynous, with multiple dealate queens often occurring together in nests. Workers are omnivorous and opportunistic foragers.

Nomenclature

 * . Brachymyrmex heeri Forel, 1874: 89, 91, figs. 16, 20 (w.) SWITZERLAND (hothouse in botanical garden, Zurich).
 * [Note: description repeated in Latin, p. 92.]
 * Forel, 1876: 52 (q.m.).
 * Status as species: Forel, 1876: 52; Emery, 1878b: 47; Emery & Forel, 1879: 466; Forel, 1881: 3; André, 1882b: 214 (in key); Mayr, 1886d: 431; Cresson, 1887: 257; Dalla Torre, 1893: 174; Forel, 1895b: 106; Forel, 1899c: 123; Wheeler, W.M. 1904e: 304; Wheeler, W.M. 1905b: 132; Wheeler, W.M. 1906e: 350; Forel, 1907e: 9; Wheeler, W.M. 1908a: 153; Wheeler, W.M. 1911a: 29; Forel, 1912i: 62; Wheeler, W.M. & Mann, 1914: 44; Forel, 1915d: 49 (in key); Wheeler, W.M. 1916m: 591; Mann, 1920: 431; Wheeler, W.M. 1921f: 166; Wheeler, W.M. 1922c: 15; Wheeler, W.M. 1923c: 5; Santschi, 1923b: 664; Emery, 1925b: 42; Stärcke, 1926: 118 (in key); Borgmeier, 1927c: 141; Menozzi, 1927c: 268; Menozzi & Russo, 1930: 166; Menozzi, 1931b: 274; Aguayo, 1932: 224; Santschi, 1933e: 122; Eidmann, 1936b: 92; Smith, M.R. 1937: 866; Santschi, 1939e: 319; Wheeler, W.M. 1942: 253; Kusnezov, 1953b: 339; Kempf, 1961b: 522; Bernard, 1967: 280 (redescription); Kempf, 1970b: 340; Kempf, 1972a: 39; Alayo, 1974: 26 (in key); Bolton, 1995b: 82; Wetterer & Wetterer, 2004: 215; Wild, 2007b: 27; Branstetter & Sáenz, 2012: 255; Borowiec, L. 2014: 24 (see note in bibliography); Radchenko, 2016: 348; Fernández & Ortiz-Sepúlveda, 2019: 728; Lubertazzi, 2019: 78; Ortiz-Sepúlveda, et al. 2019: 503 (redescription).
 * Senior synonym of cordobensis: Ortiz-Sepúlveda, et al. 2019: 503.
 * Senior synonym of goeldii: Ortiz-Sepúlveda, et al. 2019: 503.
 * Senior synonym of physogaster: Ortiz-Sepúlveda, et al. 2019: 503.
 * cordobensis. Brachymyrmex giardi var. cordobensis Santschi, 1929d: 309 (w.) ARGENTINA (Córdoba).
 * Subspecies of giardi: Kempf, 1972a: 39; Bolton, 1995b: 82.
 * Junior synonym of heeri: Ortiz-Sepúlveda, et al. 2019: 503.
 * goeldii. Brachymyrmex goeldii Forel, 1912i: 65 (w.) BRAZIL (São Paulo).
 * Status as species: Luederwaldt, 1918: 48; Santschi, 1923b: 663; Emery, 1925b: 42; Borgmeier, 1927c: 141; Kempf, 1972a: 39; Bolton, 1995b: 82.
 * Junior synonym of heeri: Ortiz-Sepúlveda, et al. 2019: 503.
 * physogaster. Brachymyrmex physogaster Kusnezov, 1960c: 382, figs. 1-4 (w.) ARGENTINA (Salta).
 * Status as species: Bolton, 1995b: 82.
 * Junior synonym of heeri: Ortiz-Sepúlveda, et al. 2019: 503.

Description
Ouvriere: Longueur 1, 2 a 2mm

Tout le corps court, large, trapu. Mandibules munies de cinq dents (les autres especes en ont quatre). Pas trace d'ocelles (les autres especes ont trois ocelles). Yeux composes d'environ 35 facettes chacun ; ils sont situes legerement en avant du milieu des bords lateraux de la tete. Palpes maxillaires longs; le second et le troisieme article sont plus longs que les quatre autres. Les palpes labiaux sont assez longs aussi; leurs quatre articles sont egaux entre eux. Chaperon (c.) en forme de capuchon (comme chez le Brachymyrmex patagonicus), sans carene, recouvrant un peu les mandibules, fortement voute de droite a gauche, moins fortement d'avant en arriere: Il est arrondi posterienrement, parfois aussi un peu echancre au milieu de son bord posterieur ; il ne se prolonge pas ou presque pas entre les articulations des antennes (entre les aretes frontales). Aire frontale distincte, triangulaire, souvent un peu arrondie a son angle posterieur. Sillon frontal distinct. Aretes frontales courtes, divergentes, a peine recourbees. La tete, un peu aplatie en dessus, basse en arriere, est echancree a son bord posterieur. Premier article du fouet des antennes un peu plus long que les deux suivants ensemble ; dernier (8m.) article du fouet aussi long que les trois qui le precedent a. la fois (fg.) Thorax tres court, surtout le mesonotum et le pronotum qui sont voutesis et forment ensemble seulement les 2/5 de sa longueur totale. Un sillon transversal enfonce, assez etroit, separe le mesonotum du metanotum; ce sillon ne renferme pas de petite spirale (il en renferme une chez le Brachymyrmex tristis). La face basale du metanotum est extremement courte, convexe (mtb); sa face declive, longue, large et plate (mtd), va en pente douce et egale jusqu'au pedicule. Eperons des pattes anterieures tres forts. Ecaille encore plus fortement inclinee en avant que chez la Plagiolepis pygmaea (e). Premier segment de l'abdomen prolonge en avant et recouvrant l'ecaille ; abdomen grand. D'un jaune plus on moins brunatre, grisatre ou rougeatre. Dessus de la tete et de l'abdomen plus fonce. Tout le corps luisant; chaperon et face declive du metanotum lisses; quelques rugosites assez grossieres sur les joues; tout le reste tres finement rugueux ponctue. Pubescence grisatre, assez forte sur l'abdomen, un peu moins forte sur la tete, plus faible sur le thorax, les pattes et les antennes, nulle sur le chaperon et sur la face declive du metanotum. Poils epars, un peu partout, sauf sur les pattes et sur les antennes. Les deux stigmates posterieurs sont gros et tres distincts.

Type Locality
Forel (1874) reported "Jardin botanique de Zurich, dans la serre des Orchidees tropicales. (Botanical Garden of Zurich, in the greenhouse of tropical orchids.)" It is now clear that the type specimens must have been transported to the European botanical garden in tropical plant material. This has created the odd situation of Brachymyrmex heeri being a New World species with a type locality of Switzerland.

References based on Global Ant Biodiversity Informatics

 * Aguayo C. G. 1932. Notes on West Indian ants. Bulletin of the Brooklyn Entomological Society 27: 215-227.
 * Ahuatzin D. A., E. J. Corro, A. Aguirre Jaimes, J. E. Valenzuela Gonzalez, R. Machado Feitosa, M. Cezar Ribeiro, J. Carlos Lopez Acosta, R. Coates, W. Dattilo. 2019. Forest cover drives leaf litter ant diversity in primary rainforest remnants within human-modified tropical landscapes. Biodiversity and Conservation 28(5): 1091-1107.
 * Alayo D. P. 1974. Introduccion al estudio de los Himenopteros de Cuba. Superfamilia Formicoidea. Academia de Ciencias de Cuba. Instituto de Zoologia. Serie Biologica no.53: 58 pp. La Habana.
 * Alonso L. E., J. Persaud, and A. Williams. 2016. Biodiversity assessment survey of the south Rupununi Savannah, Guyana. BAT Survey Report No.1, 306 pages.
 * Alvarez, G., I. Armbrecht, E. Jimenez, H. Armbrecht and P. Ulloa-Chacon. 2001. Ant-plant association in two Tococa species from a primary rain forest of Colombia Choco (Hymenoptera: Formicidae) Sociobiology 38(3B):585-602
 * Armbrecht I., E. Jimenez, G. Alvarez, P. Ulloa-Chacon, and H. Armbrecht. 2001. An ant mosaic in the Colombian rain forest of Choco (Hymenoptera: formicidae0. Sociobiology 37(3B): 491-509.
 * Arruda F. V., M. A. Pesquero, D. G. Marcelino, G. A. Leiter, J. H. C. Delabie, and R. Fagundes. 2015. Size and condition of bamboo as structural factors behind the vertical stratification of the bamboo-nesting ant community. Insectes Sociaux DOI 10.1007/s00040-015-0440-4
 * Badano, E.I., H.A. Regidor, H.A. Nunez, R. Acosta and E. Gianoli. 2005. Species richness and structure of ant communities in a dynamic archipelago: effects of island area and age. Journal of Biogeography
 * Castano-Meneses G., R. De Jesus Santos, J. R. Mala Dos Santos, J. H. C. Delabie, L. L. Lopes, and C. F. Mariano. 2019. Invertebrates associated to Ponerine ants nests in two cocoa farming systems in the southeast of the state of Bahia, Brazil. Tropical Ecology 60: 52–61.
 * Castano-Meneses, G., M. Vasquez-Bolanos, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha, and I. Alcala-Martinez. 2015. Avances de Formicidae de Mexico. Universidad Nacional Autonoma de Mexico.
 * Cuezzo, F. 1998. Formicidae. Chapter 42 in Morrone J.J., and S. Coscaron (dirs) Biodiversidad de artropodos argentinos: una perspectiva biotaxonomica Ediciones Sur, La Plata. Pages 452-462.
 * Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
 * Del Toro, I., M. Vázquez, W.P. Mackay, P. Rojas and R. Zapata-Mata. Hormigas (Hymenoptera: Formicidae) de Tabasco: explorando la diversidad de la mirmecofauna en las selvas tropicales de baja altitud. Dugesiana 16(1):1-14.
 * Diehl-Fleig E. 2014. Termites and Ants from Rio Grande do Sul, Brazil. Sociobiology (in Press).
 * Fagundes R., G. Terra, S. P. Ribeiro, and J. D. Majer. 2010. The Bamboo Merostachys fi scheriana (Bambusoideae: Bambuseae) as a Canopy Habitat for Ants of Neotropical Montane Forest. Neotropical Entomology 39(6):906-911
 * Fernandes T. T., R. R. Silva, D. Rodrigues de Souza-Campana, O. Guilherme Morais da Silva, and M. Santina de Castro Morini. 2019. Winged ants (Hymenoptera: Formicidae) presence in twigs on the leaf litter of Atlantic Forest. Biota Neotropica 19(3): http://dx.doi.org/10.1590/1676-0611-bn-2018-0694
 * Fernandez Triana J. L., H. Sariol, M. A. Vega Castillo, S. Ricardo, M. Gonzalez, and E. P. Ferrer. 2002. Datos preliminares dobre la biodiversidad del orden Hymenoptera en la provincia Granma, Cuba. Bol. S.E.A. 31: 43-48.
 * Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
 * Fichaux M., B. Bechade, J. Donald, A. Weyna, J. H. C. Delabie, J. Murienne, C. Baraloto, and J. Orivel. 2019. Habitats shape taxonomic and functional composition of Neotropical ant assemblages. Oecologia 189(2): 501-513.
 * Figueiredo C. J. de, R. R. da Silva, C. de Bortoli Munhae, and M. S. de Castro Morini. 2013. Ant fauna (Hymenoptera: Formicidae) attracted to underground traps in Atlantic Forest. Biota Neotrop 13(1):  176-182
 * Fleck M. D., E. Bisognin Cantarelli, and F. Granzotto. 2015. Register of new species of ants (Hymenoptera: Formicidae) in Rio Grande do Sul state. Ciencia Florestal, Santa Maria 25(2): 491-499.
 * Fontanla Rizo J.L. 1997. Lista preliminar de las hormigas de Cuba. Cocuyo 6: 18-21.
 * Fontenla J. L. 2005. Species of ants (Formicidae) recorded in the Sierra de Cubitas and adjacent areas, Camagüey Province, 16-19 September 2002. In: Díaz, L., M., W. S. Alverson, A. Barreto V., y / and T. Wachter. 2006. Cuba: Camagüey, Sierra de Cubitas. Rapid Biological Inventories Report 08. The Field Museum, Chicago
 * Fontenla J. L., and J. Alfonso-Simonetti. 2018. Classification of Cuban ants (Hymenoptera: Formicidae) into functional groups. Poeyana Revista Cubana de Zoologia 506: 21-30.
 * Fontenla Rizo J. L. 1993. Composición y estructura de comunidades de hormigas en un sistema de formaciones vegetales costeras. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 441: 1-19.
 * Fontenla Rizo J. L. 1993. Mirmecofauna de Isla de la Juventud y de algunos cayos del archipielago cubano. Poeyana. Instituto de Ecologia y Sistematica, Academia de Ciencias de Cuba 444:1-7.
 * Fontenla Rizo J. L. 1997. Lista preliminar de las hormigas de Cuba (Hymenoptera: Formicidae). Cocuyo 6: 18-21.
 * Fontenla Rizo J. L., and L. M. Hernández. 1993. Relaciones de coexistencia en comunidades de hormigas en un agroecosistema de caña de azúcar. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 438: 1-16.
 * Forel A. 1912. Formicides néotropiques. Part VI. 5me sous-famille Camponotinae Forel. Mémoires de la Société Entomologique de Belgique. 20: 59-92.
 * Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
 * Garcia M. A. The vulnerability of leaflitter ants to forest disturbances in the islands of Puerto Rico, Greater Antilles. Novitates Caribaea 13: 74-91.
 * Kempf W. W. 1961. A survey of the ants of the soil fauna in Surinam (Hymenoptera: Formicidae). Studia Entomologica 4: 481-524.
 * Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
 * Kusnezov N. 1953. La fauna mirmecológica de Bolivia. Folia Universitaria. Cochabamba 6: 211-229.
 * Kusnezov N. 1978. Hormigas argentinas: clave para su identificación. Miscelánea. Instituto Miguel Lillo 61:1-147 + 28 pl.
 * Leponce, M., L. Theunis, J.H.C. Delabie and Y. Roisin. 2004. Scale dependence of diversity measures in a leaf-litter ant assemblage. Ecography. 27:253-267.
 * Longino J. T., J. Coddington, and R. K. Colwell. 2002. The ant fauna of a tropical rain forest: estimating species richness three different ways. Ecology 83: 689-702.
 * Longino J. et al. ADMAC project. Accessed on March 24th 2017 at https://sites.google.com/site/admacsite/
 * Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
 * Menozzi C, Russo G. 1930. Contributo alla conoscenza della mirmecofauna della Repubblica Dominicana (Antille). Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici. 24: 148-173.
 * Menozzi C. 1927. Formiche raccolte dal Sig. H. Schmidt nei dintorni di San José di Costa Rica. Entomologische Mitteilungen. Berlin-Dahlem. 16: 266-277.
 * Mentone T. O., E. A. Diniz, C. B. Munhae, O. C. Bueno, and M. S. C. Morini. 2011. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop. 11(2): http://www.biotaneotropica.org.br/v11n2/en/abstract?inventory+bn00511022011.
 * Mentone T.O., M. S. C. Morini, L. Souza, and S. M. P. Braga. 2009. Hymenoptera communities in an agroecosyste using direct seedling in Southeastern Brazil. Sociobiology 53(2B): 473-486.
 * Mirmecofauna de la reserva ecologica de San Felipe Bacalar
 * Morrison L. W. 1998. A review of Bahamian ant (Hymenoptera: Formicidae) biogeography. Journal of Biogeography 25: 561-571.
 * Munhae C. B., Z. A. F. N. Bueno, M. S. C. Morini, and R. R. Silva. 2009. Composition of the Ant Fauna (Hymenoptera: Formicidae) in Public Squares in Southern Brazil. Sociobiology 53(2B): 455-472.
 * Nascimento Santos M., J. H. C. Delabie, and J. M. Queiroz. 2019. Biodiversity conservation in urban parks: a study of ground-dwelling ants (Hymenoptera: Formicidae) in Rio de Janeiro City. Urban Ecosystems https://doi.org/10.1007/s11252-019-00872-8
 * Neves F. S., K. S. Queiroz-Dantas, W. D. da Rocha, and J. H. C. Delabie. 2013. Ants of Three Adjacent Habitats of a Transition Region Between the Cerrado and Caatinga Biomes: The Effects of Heterogeneity and Variation in Canopy Cover. Neotrop Entomol 42: 258268.
 * Oliveira Mentone T. de, E. A. Diniz, C. de Bortoli Munhae, O. Correa Bueno and M. S. de Castro Morini. 2012. Composition of ant fauna (Hymenoptera: Formicidae) at litter in areas of semi-deciduous forest and Eucalyptus spp., in Southeastern Brazil. Biota Neotrop 11(2): 237-246.
 * Ortiz-Sepuvelda C. M., B. Van Bocxlaer, A. D. Meneses, and F. Fernandez. 2019. Molecular and morphological recognition of species boundaries in the neglected ant genus Brachymyrmex (Hymenoptera: Formicidae): toward a taxonomic revision. Organisms Diversity & Evolution https://doi.org/10.1007/s13127-019-00406-2
 * Osorio-Perez K., M. F. Barberena-Arias, and T. M. Aide. 2007. Changes in Ant Species Richness and Composition During Plant Secondary Succession in Puerto Rico. Caribbean Journal of Science 43(2): 244-253.
 * Ottonetti L., L. Tucci, F. Frizzi, G. Chelazzi, and G. Santini. 2010. Changes in ground-foraging ant assemblages along a disturbance gradient in a tropical agricultural landscape. Ethology Ecology & Evolution 22: 7386.
 * Pacheco, R., R.R. Silva, M.S. de C. Morini, C.R.F. Brandao. 2009. A Comparison of the Leaf-Litter Ant Fauna in a Secondary Atlantic Forest with an Adjacent Pine Plantation in Southeastern Brazil. Neotropical Entomology 38(1):055-065
 * Perez-Gelabert D. E. 2008. Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography. Zootaxa 1831:1-530.
 * Philpott S. M., I. Perfecto, and J. Vandermeer. 2006. Effects of management intensity and season on arboreal ant diversity and abundance in coffee agroecosystems. 15: 139-155.
 * Philpott, S.M. and P.F. Foster. 2005. Nest-site limitation in coffee agroecosytems: Artificial nests maintain diversity of arboreal ants. Ecological Applications 15(4):1478-1485
 * Pinheiro E. R. S., L. da S. Duarte, E. Diehl, and S. M. Hartz. 2010. Edge effects on epigeic ant assemblages in a grasslandeforest mosaic in southern Brazil. Acta Oecolo. doi:10.1016/j.actao.2010.03.004
 * Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
 * Portuondo E. F., and J. L. Reyes. 2002. Mirmecofauna de los macizos montañosos de Sierra Maestra y Nipe-Sagua-Baracoa. Cocuyo 12: 10-13
 * Portuondo E. F., and J. L. Reyes. 2006. Species of Hymenoptera recorded in the Reserva Ecológica Pico Mogote, Santiago de Cuba Province, 20-25 September 2002. In: Maceira F., D., A. Fong G., y/and W. S. Alverson, eds. 2006. Cuba: Pico Mogote. Rapid Biological Inventories Report 09. The Field Museum, Chicago.
 * Portuondo E. F., and J. L. Reyes. 2006. Species of hymenopterans recorded in Siboney-Juticí Ecological Reserve, Santiago de Cuba Province, compiled from collections during the rapid inventory of 27-28 September 2002. Fong G., A., D. Maceira F., W. S. Alverson, y / and J. M. Shopland, eds. 2005. Cuba: Siboney-Juticí. Rapid Biological Inventories Report 10. The Field Museum, Chicago.
 * Portuondo Ferrer E., and J. L. Fernández Triana. 2005. Species of hymenopterans (bees, wasps, and ants) recorded in Alejandro de Humboldt National Park, from literature records, revision of the collection at BIOECO, and collections before and during the rapid inventory, 12-22 February 2004. In Fong G., A., D. Maceira F., W. S. Alverson, y/and T. Wachter, eds. 2005. Cuba: Parque Nacional Alejandro de Humboldt. Rapid Biological Inventories Report 14. The Field Museum, Chicago.
 * Portuondo Ferrer, E. and J. Fernandez Triana. Biodiversidad del orden Hymenoptera en Los Macizos Montanosos de Cuba Oriental. Boletin S.E.A. 35:121-136.
 * Reyes, J. L. "Inventario de la colección de hormigas (Hymenoptera: Formicidae) del Centro Oriental de Ecosistemas y Biodiversidad, Santiago de Cuba, Cuba." Boletín de la Sociedad Aragonesa 36 (2005): 279-283.
 * Reynoso-Campos J. J., J. A. Rodriguez-Garza, and M. Vasquez-Bolanos. 2015. Hormigas (Hymenoptera: Formicidae) de la Isla Cozumel, Quintana Roo, Mexico (pp. 27-39). En: Castaño Meneses G., M. Vásquez-Bolaños, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha e I. Alcalá-Martínez (Coords.). Avances de Formicidae de  México.  UNAM,  Universiad  de  Guadalajara, Guadalajara, Jalisco.
 * Rodrigues de Souza D., E. Stingel, L. C. de Almeida, M. A. Lazarini, C. de Bortoli Munhae, O. Correa Bueno, C. R. Archangelo, and M. Santina de C. Morini. 2010. Field methods for the study of ants in sugarcane plantations in Southeastern Brazil. Sci. Agric. (Piracicaba, Braz.) 67(6): 651-657.
 * Santschi F. 1923. Revue des fourmis du genre Brachymyrmex Mayr. Anales del Museo Nacional de Historia Natural de Buenos Aires 31: 650-678.
 * Santschi F. 1933. Fourmis de la République Argentine en particulier du territoire de Misiones. Anales de la Sociedad Cientifica Argentina. 116: 105-124.
 * Santschi F. 1939. Études et descriptions de fourmis néotropiques. Revista de Entomologia (Rio de Janeiro). 10: 312-330.
 * Siqueira de Castro F., A. B. Gontijo, P. de Tarso Amorim Castro, and S. Pontes Ribeiro. 2012. Annual and Seasonal Changes in the Structure of Litter-Dwelling Ant Assemblages (Hymenoptera: Formicidae) in Atlantic Semideciduous Forests. Psyche doi:10.1155/2012/959715
 * Smith M. A., W. Hallwachs, D. H. Janzen. 2014. Diversity and phylogenetic community structure of ants along a Costa Rican elevational gradient. Ecography 37(8): 720-731.
 * Smith M. R. 1937. The ants of Puerto Rico. Journal of Agriculture of the University of Puerto Rico 20: 819-875.
 * Smith M. R. 1942. The relationship of ants and other organisms to certain scale insects on coffee in Puerto Rico. Journal of Agriculture of the University of Puerto Rico 26: 21-27.
 * Smith, Marion R. 1954. American Museum Novitates. Ants of the Bimini Island Group, Bahamas, British West Indies (Hymenoptera, Formicidae). 1671:1-16
 * Smith, Marion R. 1954. Ants of the Bimini Island Group, Bahamas, British West Indies. American Museum of Natural History. 1671. 1-16.
 * Snelling R. 1993. Ants of Guana Island, British Virgin Islands. Notes From Underground 8: 11-12.
 * Suguituru S. S., D. R. de Souza, C. de Bortoli Munhae, R. Pacheco, and M. S. de Castro Morini. 2011. Diversidade e riqueza de formigas (Hymenoptera: Formicidae) em remanescentes de Mata Atlântica na Bacia Hidrográfica do Alto Tietê, SP. Biota Neotrop. 13(2): 141-152.
 * Suguituru S. S., M. Santina de Castro Morini, R. M. Feitosa, and R. Rosa da Silva. 2015. Formigas do Alto Tiete. Canal 6 Editora 458 pages
 * Suguituru S. S., R. Rosa Silva, D. R. de Souza, C. de Bortoli Munhae, and M. Santina de Castro Morini. Ant community richness and composition across a gradient from Eucalyptus plantations to secondary Atlantic Forest. Biota Neotrop. 11(1): 369-376.
 * Theunis, L., M. Gilbert, Y. Roisin and M. Leponce. 2005. Spatial structure of litter-dwelling ant distribution in a subtropical dry forest. Insectes Sociaux. 52:366377.
 * Torres J.A. 1984. Niches and Coexistence of Ant Communities in Puerto Rico: Repeated Patterns. Biotropica 16(4): 284-295.
 * Torres, Juan A. and Roy R. Snelling. 1997. Biogeography of Puerto Rican ants: a non-equilibrium case?. Biodiversity and Conservation 6:1103-1121.
 * Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
 * Wetterer J. K., and C. D. Lombard. 2010. Fire Ants (Hymenoptera: Formicidae) along an Important Sea Turtle Nesting Beach on St. Croix, USVI. Florida Entomologist 93(3): 449-450.
 * Wetterer J.K. and J.L.W. Keularts. 2008. Population explosion of the hairy crazy ant, Paratrechina pubens (Hymenoptera: Formicidae), on St. Croix, US Virgin Islands. Florida Entomologist 91(3): 423-427.
 * Wetterer, J.K. and J.L.W. Keularts. 2008. Population Explosion of the Hairy Crazy Ant, Paratrechina pubens (Hymenoptera: Formicidae), on St. Croix, US Virgin Islands. The Florida Entomologist 91(3):423-427
 * Wheeler W. M. 1905. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21: 79-135.
 * Wheeler W. M. 1908. The ants of Porto Rico and the Virgin Islands. Bulletin of the American Museum of Natural History 24: 117-158.
 * Wheeler W. M. 1922. The ants of Trinidad. American Museum Novitates 45: 1-16.
 * Wheeler W. M. 1942. Studies of Neotropical ant-plants and their ants. Bulletin of the Museum of Comparative Zoology 90: 1-262.
 * Wheeler W. M., and W. M. Mann. 1914. The ants of Haiti. Bulletin of the American Museum of Natural History 33: 1-61.
 * Wheeler, William Morton. 1911. Additions to the Ant-Fauna of Jamaica. Bulletin American Museum of Natural History. 30:21-29.
 * Wheeler, William Morton. 1911. Ants Collected in Grenada, W.I. by Mr. C. T. Brues. Bulletin of the Museum of Comparitive Zoology at Harvard College. 54(5):166-172.
 * Wheeler, William Morton. 1923. Report on the Ants. The University of Iowa Studies in Natural History. 10(3):3-9.
 * Wild, A. L. "A catalogue of the ants of Paraguay (Hymenoptera: Formicidae)." Zootaxa 1622 (2007): 1-55.
 * de Souza D. R., S. G. dos Santos, C. de B. Munhae, and M. S. de C. Morini. 2012. Diversity of Epigeal Ants (Hymenoptera: Formicidae) in Urban Areas of Alto Tietê. Sociobiology 59(3): 703-117.
 * do Nascimento, I.C. 2006. Fenologia dos Voos de Acasalamento em Formigas Tropicais