Monomorium bicolor

An Afrotropical species that appears to have a wide native range. It has also been found in the Dominican Republic. The latter is an unusual occurrence. While there are many Monomorium tramp species, most have been spread to many areas outside of their native range.

Identification
Bolton (1987) - A member of the M. bicolor complex in the M. salomonis species group.

Worker. TL 3.2-3.9, HL 0.70-0.93, HW 0.52-0.75, CI 73-83, SL 0.56-0.78, SI 104-115, PW 0.36-0.50, AL 0.82-1.20 (25 measured).

Third and fourth (basal) tooth of mandible approximately the same size or the fourth very slightly smaller than the third, but the basal tooth never reduced to a minute denticle. Median portion of clypeus with its anterior free margin usually indented, more rarely the margin approximately transverse but never with flanking sharp teeth. Eyes of moderate size, the maximum diameter 0.24-0.27 x HW. Ventral surface of head with curved simple hairs but lacking extremely long J-shaped ammochaete hairs. Dorsal alitrunk hairless, petiole with one pair and postpetiole with 1-2 pairs of posteriorly directed hairs. Discounting the apical transverse row the first gastral tergite usually with only 1-2 pairs of hairs, situated on the basal half; rarely 3-4 pairs of hairs present. Dorsum and sides of head and entirety of alitrunk densely and sharply reticulate-punctate. Gaster usually finely and densely shagreenate dorsally but the sculpture may fade posteriorly in some examples and is reduced in a few. Colour bright orange to red on the head and alitrunk, the gaster blackish brown to black, the two strongly contrasting. First gastral tergite frequently with an anteromedian paler area.

In the bicolor-complex Monomorium bicolor differs from its closest relatives in that it lacks the densely hairy alitrunk seen in Monomorium hirsutum, has such smaller eyes than Monomorium personatum (0.31-0.33 x HW), lacks the reduced basal mandibular tooth of Monomorium rufulum, lacks the pair of sharp clypeal teeth characteristic of Monomorium westi, and lacks the dense gastral pilosity of Monomorium dictator.

Distribution based on Regional Taxon Lists
Afrotropical Region: Burkina Faso, Cameroun, Democratic Republic of Congo, Djibouti, Eritrea, Ethiopia, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Liberia, Nigeria, Saudi Arabia, Senegal, Sudan, Togo, United Arab Emirates. Neotropical Region: Dominican Republic, Greater Antilles. Palaearctic Region: Egypt.

Biology
Bolton (1987) - In sub-Saharan Africa Monomorium bicolor is characteristically a species of open savannah or semi-arid zones, but it also occurs in forested areas where there is some direct insolation, often being found on forest paths in Nigeria and Ghana. Nests are constructed directly into the earth and the species appears to be a general scavenger in habits, quickly appearing in traps baited with crushed large insects.

Nomenclature

 * . Monomorium bicolor Emery, 1877b: 368 (w.) ERITREA.
 * Type-material: syntype workers (number not stated, “a great number”).
 * Type-locality: Eritrea: Bogos, Sciotel, 1870 (O. Beccari).
 * Type-depositories: MRAC, MSNG.
 * [Misspelled as bicolour by Hita Garcia, et al. 2013: 212.]
 * Mayr, 1901b: 7 (q.).
 * Combination in M. (Xeromyrmex): Wheeler, W.M. 1922a: 868.
 * Subspecies of subopacum: Santschi, 1927d: 244; Santschi, 1935b: 268; Finzi, 1936: 178; Santschi, 1936a: 42 (in key); Finzi, 1939a: 158.
 * Status as species: Emery, 1881b: 531 (in key); André, 1883a: 334 (in key); André, 1884b: 540; Dalla Torre, 1893: 66; Forel, 1894b: 88; Mayr, 1901b: 7; Forel, 1903a: 687; Forel, 1903e: 564; Mayr, 1904b: 4; Forel, 1907a: 18; Forel, 1907e: 15; Emery, 1908h: 677; Santschi, 1908: 518; Forel, 1910c: 251; Karavaiev, 1911: 5; Santschi, 1912b: 163; Santschi, 1914d: 353; Arnold, 1916: 209; Emery, 1922e: 176; Wheeler, W.M. 1922a: 162, 868; Emery, 1922e: 176; Menozzi, 1926a: 41; Menozzi, 1930b: 108; Santschi, 1937d: 222; Donisthorpe, 1942a: 28; Weber, 1943c: 314; Donisthorpe, 1947e: 109; Menozzi & Consani, 1952: 66; Ettershank, 1966: 87; Collingwood, 1985: 269; Bolton, 1987: 338 (redescription); Agosti & Collingwood, 1987a: 55; Agosti & Collingwood, 1987b: 272 (in key); Bolton, 1995b: 259; Collingwood & Agosti, 1996: 342; Petrov, 2006: 94 (in key); Lush, 2008: 72 (in key); Collingwood, et al. 2011: 433; Hita Garcia, et al. 2013: 212; Borowiec, L. 2014: 116 (see note in bibliography); Salata & Borowiec, 2018c: 47; Lubertazzi, 2019: 128; Madl, 2019: 15.
 * Senior synonym of aequatoriale: Bolton, 1987: 338; Bolton, 1995b: 259.
 * Senior synonym of coerulescens: Santschi, 1914d: 353 (in key); Emery, 1922e: 176; Wheeler, W.M. 1922a: 869; Bolton, 1987: 338; Bolton, 1995b: 259.
 * Senior synonym of rufibasis: Santschi, 1926b: 240 (in text); Santschi, 1927d: 244; Bolton, 1987: 338; Bolton, 1995b: 259.
 * Senior synonym of tropicale: Bolton, 1987: 338; Bolton, 1995b: 259.
 * Senior synonym of uelense: Bolton, 1987: 338; Bolton, 1995b: 259.
 * Distribution: Burkina Faso, Cameroon, Cyprus, Democratic Republic of Congo, Djibouti, Dominican Republic (introduced), Egypt, Eritrea, Ethiopia, Ghana, Greece, Guinea, Iran, Israel, Kenya, Liberia, Libya, Nigeria, Oman, Saudi Arabia, Sudan, Togo, United Arab Emirates.
 * aequatoriale. Monomorium (Xeromyrmex) bicolor var. aequatoriale Santschi, 1926b: 240 (w.) CAMEROON, DEMOCRATIC REPUBLIC OF CONGO.
 * Type-material: 3 syntype workers.
 * Type-locality: Cameroon: Gr Batanga, 1991 (Schwab).
 * Type-depository: NHMB.
 * As unavailable (infrasubspecific) name: Santschi, 1927d: 244.
 * Subspecies of bicolor: Ettershank, 1966: 87.
 * Junior synonym of bicolor: Bolton, 1987: 338; Bolton, 1995b: 258.
 * coerulescens. Monomorium bicolor var. coerulescens Santschi, 1912c: 148 (w.) DJIBOUTI.
 * Type-material: holotype worker.
 * Type-locality: Djibouti: Obock, 1893 (Maindron).
 * Type-depository: MNHN.
 * Junior synonym of bicolor: Santschi, 1914d: 353 (in key); Emery, 1922e: 176; Wheeler, W.M. 1922a: 869; Bolton, 1987: 338; Bolton, 1995b: 260.
 * rufibasis. Monomorium bicolor var. rufibasis Santschi, 1914d: 353 (diagnosis in key) (w.) EGYPT.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Egypt: Upper Egypt (F. Silvestri).
 * Type-depository: not in NHMB (perhaps lost, or perhaps in DEUN).
 * [Misspelled as rufobasalis by Santschi, 1926b: 240, Santschi, 1927d: 244.]
 * As unavailable (infrasubspecific) name: Finzi, 1936: 178.
 * Subspecies of bicolor: Emery, 1922e: 176.
 * Junior synonym of bicolor: Santschi, 1926b: 240 (in text); Santschi, 1927d: 244; Bolton, 1987: 338; Bolton, 1995b: 266.
 * tropicale. Monomorium (Xeromyrmex) bicolor var. tropicale Santschi, 1926b: 240 (w.q.m.) DEMOCRATIC REPUBLIC OF CONGO.
 * Type-material: syntype workers, syntype queen(s), syntype male(s) (numbers not stated).
 * Type-locality: Democratic Republic of Congo (“Congo belge”): Stanleyville (= Kisangani) (Majella).
 * Type-depository: NHMB.
 * As unavailable (infrasubspecific) name: Santschi, 1927d: 244; Santschi, 1936a: 42 (in key).
 * Suubspecies of bicolor: Ettershank, 1966: 92.
 * Junior synonym of bicolor: Bolton, 1987: 338; Bolton, 1995b: 268.
 * ''Type-localities: “Habitat in Aegypto, Arabia, Palaestina” (no further data).
 * [Note: Linnaeus, 1764: 418, restricts the distribution to Egypt, which may be regarded as the type-locality.]
 * Type-depository: ZMLS, perhaps also in UUUS.
 * [Misspelled as salamonis by Arnold, 1916: 219, Arnold, 1944: 13; misspelled as solomonis by Wheeler, W.M. & Mann, 1916: 170; misspelled as solamonis by Menozzi, 1940: 268; misspelled as salmonis by Petrov, 2006: 94 (in key).]
 * André, 1883a: 342 (m.); Emery, 1908h: 674 (w.q.m.).
 * Combination in Monomorium: Roger, 1862c: 294;
 * combination in M. (Xeromyrmex): Emery, 1915i: 190.
 * Status as species: Linnaeus, 1764: 418; Linnaeus, 1767: 963; Forskål, 1775: 84; Gmelin, 1790: 2799; Christ, 1791: 507; Olivier, 1792; 503; Latreille, 1802c: 291; Roger, 1862c: 294; Roger, 1863b: 32; Mayr, 1863: 429; Mayr, 1877: 20 (in list); Emery, 1877b: 369; Emery, 1878b: 50; Emery & Forel, 1879: 456; Emery, 1880: 391; Emery, 1881b: 531; André, 1883a: 336 (in key); André, 1884b: 540; Emery, 1884a: 381; Saunders, E. 1890: 204; Emery, 1891b: 13; Scudder, 1891: 708; Dalla Torre, 1893: 69; Forel, 1894a: 162; Forel, 1894d: 19; Emery, 1895b: 66; Forel, 1895d: 227, 233; Emery, 1901e: 61; Ruzsky, 1902d: 25; Forel, 1902a: 152; Rothney, 1903: 97; Ruzsky, 1903b: 316; Mayr, 1904b: 4; Forel, 1904c: 5; Forel, 1904f: 422; Ruzsky, 1905b: 633; Wheeler, W.M. 1905b: 123; Forel, 1907a: 18; Forel, 1907e: 15; Emery, 1908h: 674; Santschi, 1908: 518; Forel, 1909e: 372; Forel, 1910c: 251; Karavaiev, 1910b: 59; Karavaiev, 1911: 4; Karavaiev, 1912a: 3; Wheeler, W.M. & Mann, 1914: 19; Arnold, 1916: 219; Wheeler, W.M. & Mann, 1916: 170; Emery, 1916b: 161; Stitz, 1917: 346; Crawley, 1920a: 165; Wheeler, W.M. 1922a: 870, 1027; Emery, 1922e: 177; Stitz, 1923: 156; Wheeler, W.M. 1923c: 4; Wheeler, W.M. 1926: 3; Menozzi, 1927e: 117; Wheeler, W.M. 1927g: 107; Menozzi & Russo, 1930: 157; Menozzi, 1933b: 66 (in key); Menozzi, 1934: 160; Grandi, 1935: 100; Finzi, 1936: 176; Santschi, 1936a: 49 (redescription); Finzi, 1939a: 158; Finzi, 1940: 160; Donisthorpe, 1942a: 28; Arnold, 1944: 13; Donisthorpe, 1947e: 109; Bernard, 1948: 145; Bernard, 1950b: 290; Bernard, 1953a: 159; Wellenius, 1955: 8; Bernard, 1956b: 259; Ceballos, 1956: 305; Kempf, 1960f: 436; Cagniant, 1964: 89; Cagniant, 1966b: 279; Ettershank, 1966: 92; Bernard, 1967: 169 (redescription); Hamann & Klemm, 1967: 413; Cagniant, 1968a: 143; Collingwood & Yarrow, 1969: 66; Cagniant, 1970a: 418; Bernard, 1971: 7; Baroni Urbani, 1971c: 91; Kempf, 1972a: 144; Alayo, 1974: 14 (in key); Báez & Ortega, 1978: 190; Collingwood, 1978: 82 (in key); Barquin Diez, 1981: 183; Collingwood, 1985: 272; Agosti & Collingwood, 1987b: 273 (in key); Hohmann, et al. 1993: 155; De Haro & Collingwood, 1994: 100; Bolton, 1995b: 266; Collingwood & Agosti, 1996: 354; Espadaler, 1997b: 30; Fernández, 2007b: 137; Gratiashvili & Barjadze, 2008: 140; Paknia, et al. 2008: 155; Vonshak, et al. 2009: 43; Borowiec, L. 2014: 125 (see note in bibliography); Lebas, et al. 2016: 304; Fernández & Serna, 2019: 808; Madl, 2019: 15; Borowiec, L. & Salata, 2020: 14.
 * Senior synonym of debilis: Emery & Forel, 1879: 456; Bolton, 1995b: 266.
 * Senior synonym of obscuratum: Collingwood, 1978: 69; Bolton, 1995b: 266.
 * Senior synonym of thorense: Emery & Forel, 1879: 456; Dalla Torre, 1893: 69; Ruzsky, 1905b: 633; Bolton, 1995b: 266.
 * Distribution: Algeria, Colombia (introducton), Egypt, Eritrea, France, Georgia, Iran, Israel, Italy (+ Sicily), Jordan, Kuwait, Libya, Malta, Morocco, Portugal (Madeira), Saudi Arabia, Spain (Balearics), Tunisia.
 * [Note: earlier Neotropical records (Kempf, 1972a: 144) are probably best regarded as misidentifications; note that specimens from Dominican Republic and Haiti, earlier identified as salomonis, are identified as bicolor by Lubertazzi, 2019: 128.]
 * Current subspecies: nominal plus bedui, didonis, molestum, obscuriceps, scipionis, subnitidum, targui, transversale, volubile, zanoni.
 * uelense. Monomorium (Xeromyrmex) bicolor var. uelense Santschi, 1926b: 239 (w.) DEMOCRATIC REPUBLIC OF CONGO.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Democratic Republic of Congo (“Congo belge”): Haut Uelé, Moto, 1920 (L. Burgeon).
 * Type-depositories: MRAC, NHMB.
 * [Misspelled as uluense by Santschi, 1926b: 240.]
 * [Note: Bolton, 1987: 338, records that the spelling uluense occurs on the syntype data labels, but that uelense was the published form of the name.]
 * As unavailable (infrasubspecific) name: Santschi, 1927d: 245.
 * Subspecies of bicolor: Ettershank, 1966: 93.
 * Junior synonym of bicolor: Bolton, 1987: 338; Bolton, 1995b: 268.

Type Material

 * Monomorium bicolor var. coerulescens: Holotype worker, DJIBOUTI: Obock, 1893 (M. Maindron).
 * Monomorium bicolor var. rufibasis: Syntype workers, EGYPT: Upper Egypt (diagnosis in key) (not in, presumed lost).
 * Monomorium (Xeromyrmex) bicolor var. uelense: Syntype workers, ZAIRE: Haut Uele, Moto, 1920 (L. Burgeon) (NHMB; MRAC).
 * Monomorium (Xeromyrmex) bicolor var. aequatoriale: Syntype workers, CAMEROUN: Gr. Batanga, 1911 (Schwab), (Wasmann) (NHMB).
 * Monomorium (Xeromyrmex) bicolor var. tropicale: Syntype workers, female, ZAIRE: Stanleyville (Majella) (NHMB).

Worker
Bolton (1987) - TL 3.2-3.9, HL 0.70-0.93, HW 0.52-0.75, CI 73-83, SL 0.56-0.78, SI 104-115, PW 0.36-0.50, AL 0.82-1.20 (25 measured).

Third and fourth (basal) tooth of mandible approximately the same size or the fourth very slightly smaller than the third, but the basal tooth never reduced to a minute denticle. Median portion of clypeus with its anterior free margin usually indented, more rarely the margin approximately transverse but never withftanking sharpteeth. Eyes of moderate size, the maximum diameter 0.24-0.27 x HW. Ventral surface of head with curved simple hairs but lacking extremely long J-shaped ammochaete hairs. Dorsal alitrunk hairless, petiole with one pair and postpetiole with 1-2 pairs of posteriorly directed hairs. Discounting the apical transverse row the first gastral tergite usually with only 1-2 pairs of hairs, situated on the basal half; rarely 3-4 pairs of hairs present. Dorsum and sides of head and entirety of alitrunk densely and sharply reticulate-punctate. Gaster usually finely and densely shagreenate dorsally but the sculpture may fade posteriorly in some examples and is reduced in a few. Colour bright orange to red on the head and alitrunk, the gaster blackish brown to black, the two strongly contrasting. First gastral tergite frequently with an anteromedian paler area.

References based on Global Ant Biodiversity Informatics

 * Bakr R. F. A., H. H. Fadl, R. M. Badawy, and M. R. Sharaf. 2007. Myrmecophile insects associated with some ant species (Hymenoptera : Formicidae )in Egypt. The second international conference of economic entomology (Entomological Society of Egypt) ,Cairo, Egypt, 8-11 December, (1): 205-233.
 * Belshaw R., and B. Bolton. 1994. A survey of the leaf litter ant fauna in Ghana, West Africa (Hymenoptera: Formicidae). Journal of Hymenoptera Research 3: 5-16.
 * Bernard F. 1953. La réserve naturelle intégrale du Mt Nimba. XI. Hyménoptères Formicidae. Mémoires de l'Institut Français d'Afrique Noire 19: 165-270.
 * Bolton B. 1987. A review of the Solenopsis genus-group and revision of Afrotropical Monomorium Mayr (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History). Entomology 54: 263-452.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2018. Notes on ants (Hymenoptera: Formicidae) from Gambia (Western Africa). Annals of the Upper Silesian Museum in Bytom Entomology 26: 1-13.
 * Collingwood C. A., D. Agosti, M. R. Sharaf, A. Van Harten, 2011. Order Hymenoptera, family Formicidae. Arthropod Fauna of the UAE 4: 405-474
 * Collingwood C.A., D.Agosti, M.R. Sharaf, and A. van Harten. 2011. Order Hymenoptera, family Formicidae. Arthropod fauna of the UAE, 4: 405474
 * Collingwood, C. A., and Donat Agosti. "Formicidae (Insecta: Hymenoptera) of Saudi Arabia (Part 2)." Fauna of Saudi Arabia 15 (1996): 300-385.
 * Diame L., B. Taylor, R. Blatrix, J. F. Vayssieres, J. Y. Rey, I. Grechi, and K. Diarra. 2017. A preliminary checklist of the ant (Hymenoptera, Formicidae) fauna of Senegal. Journal of Insect Biodiversity 5(15): 1-16.
 * Dieng M. M., A. B. Ndiaye, C. T. Ba, and B. Taylor. 2016. Les fourmis (Hymenoptera, Formicidae) de l’enclos d’acclimatation de Katane de la reserve de faune du Ferlo nord (Senegal). Int. J. Biol. Chem. Sci. 10(4): 1626-1636.
 * Donisthorpe H. 1942. The Formicidae (Hym.) of the Armstrong College Expedition to the Siwa Oasis. Annals and Magazine of Natural History (11)9: 26-33.
 * Donisthorpe H. 1947. Results of the Armstrong College Expedition to Siwa Oasis (Libyan Desert), 1935: A second list of the Formicidae (Hymenoptera). Bulletin de la Société Fouad Ier d'Entomologie 31: 109-111.
 * Dufour B. 1991. Place et importance des différentes espèces d'insectes dans l'écologie du CSSV (Cocoa Swollen Shoot Virus) au Togo). Café Cacao Thé 35(3): 197-204.
 * Emery C. 1915. Formiche raccolte nell'Eritrea dal Prof. F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 10: 3-26.
 * Emery, C.. "Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) Teil V. Monomorium." Deutsche Entomologische Zeitschrift 1908 (1908): 663-686.
 * Emery, C.. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte prima. Formiche provenienti dal Viaggio dei signori Antinori, Beccari e Issel nel Mar Rosso e nel paese dei Bogos. [concl.]." Annali del Museo Civico di Storia Naturale 9 (1877): 363-381.
 * Ettershank G. 1966. A generic revision of the world Myrmicinae related to Solenopsis and Pheidologeton (Hymenoptera: Formicidae). Aust. J. Zool. 14: 73-171.
 * Finzi B. 1939. Materiali zoologici dell'Eritrea raccolti da G. Müller durante la spedizione dell'Istituto Sieroterapico Milanese e conservati al Museo di Trieste. Parte III. Hymenoptera: Formicidae. Atti del Museo Civico di Storia Naturale di Trieste 14: 153-168.
 * Forel A. 1901. Einige neue Ameisen aus Südbrasilien, Java, Natal und Mossamedes. Mitteilungen der Schweizerischen Entomologischen Gesellschaft. 10: 297-311.
 * Forel A. 1903. Einige neue Ameisen aus Sud-Angola. Pp 559-564, in: Baum, H. Kunene-Sambesi-Expedition, 1903. Berlin: Verlag des Kolonial-Wirtschaftlichen Komitees, 593pp.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1910. Ameisen aus der Kolonie Erythräa. Gesammelt von Prof. Dr. K. Escherich (nebst einigen in West-Abessinien von Herrn A. Ilg gesammelten Ameisen). Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 29: 243-274.
 * IZIKO South Africa Museum Collection
 * Kolo Y. 2006. Evaluation rapide des fourmis de la région de Boké, Guinée. In Wright, H.E. and J. McCullough et M.S. Diallo. (eds). 2006. A Rapid Biological Assessment of the Boké Préfecture, Northwestern Guinea. RAP Bulletin of Biological Assessment 41. Conservation International, Washington, DC.
 * Kouakou L. M. 2015. Evaluation de la diversite des especes de fourmis anthropophiles, natives, exotiques et potentielles invasives en Cote d'Ivoire. Royal Belgian Institute of Natural Sciences
 * Kouakou L. M. M., K. Yeo, K. Ouattara, W. Dekoninck, T. Delsinne, and S. Konate. 2018. Investigating urban ant community (Hymenoptera: Formicidae) in port cities and in major towns along the border in Côte d’Ivoire: a rapid assessment to detect potential introduced invasive ant species. Journal of Animal and Plant Sciences 36(1): 5793-5811.
 * Kouakou L. M. M., W. Dekoninck, M. Kone, T. Delsinne, K. Yeo, K. Ouattara, and S. Konate. 2018. Diversity and distribution of introduced and potentially invasive ant species from the three main ecoregions of Côte d’Ivoire (West Africa). Belgian Journal of Zoology 148 (1): 83–103.
 * Madl M. 2019. Notes on the ant fauna of Eritrea (Insecta: Hymenoptera: Formicidae): type specimens deposited in the Natural History Museum Vienna (Austria) and a preliminary checklist. Ann. Naturhist. Mus. Wien, B 121: 9-18.
 * Medler J. T. 1980: Insects of Nigeria - Check list and bibliography. Mem. Amer. Ent. Inst. 30: i-vii, 1-919.
 * Menozzi C. 1930. Formiche della Somalia italiana meridionale. Memorie della Società Entomologica Italiana. 9: 76-130.
 * Menozzi C., M. Consani. 1952. Missione biologica Sagan-Omo diretta dal Prof. E. Zavattari. Hymenoptera Formicidae. Rivista di Biologia Coloniale 11: 57-71.
 * Santschi F. 1912. Fourmis d'Afrique et de Madagascar. Annales de la Société Entomologique de Belgique 56: 150-167.
 * Santschi F. 1914. Formicides de l'Afrique occidentale et australe du voyage de Mr. le Professeur F. Silvestri. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 8: 309-385.
 * Santschi F. 1926. Description de nouveaux Formicides éthiopiens (IIIme partie). Revue Zoologique Africaine (Brussels) 13: 207-267.
 * Santschi F. 1935. Hymenoptera. I. Formicidae. Mission Scientifique de l'Omo 2: 255-277.
 * Santschi, F.. "Quelques nouvelles variétés de fourmis africaines." Bulletin de la Société d' Histoire Naturelle de l' Afrique du Nord 3 (1912): 147-149.
 * Santschi, F. 1908. Nouvelles fourmis de l'Afrique du Nord (Égypte, Canaries, Tunisie). Annales de la Société Entomologique de France 77: 517-534.
 * Sharaf M. R., B. L. Fisher, H. M. Al Dhafer, A. Polaszek, and A. S. Aldawood. 2018. Additions to the ant fauna (Hymenoptera: Formicidae) of Oman: an updated list, new records and a description of two new species. Asian Myrmecology 10: e010004
 * Stephens S. S., P. B. Bosu, and M. R. Wager. 2016. Effect of overstory tree species diversity and composition on ground foraging ants (Hymenoptera: Formicidae) in timber plantations in Ghana. International Journal of Biodiversity Science, Ecosystem Services & management 12(1-2): 96-107.
 * Taylor B. 1980. Ants of the Nigerian Forest Zone (Hymenoptera: Formicidae). IV. Myrmicinae (Myrmecinini to Tetramoriini). Cocoa Research Institute of Nigeria Research Bulletin 7: 1-63.
 * Taylor B., N. Agoinon, A. Sinzogan, A. Adandonon, Y. N'Da Kouagou, S. Bello, R. Wargui, F. Anato, I. Ouagoussounon, H. Houngbo, S. Tchibozo, R. Todjhounde, and J. F. Vayssieres. 2018. Records of ants (Hymenoptera: Formicidae) from the Republic of Benin, with particular reference to the mango farm ecosystem. Journal of Insect Biodiversity 8(1): 006–029.
 * Weber N. A. 1943. The ants of the Imatong Mountains, Anglo-Egyptian Sudan. Bulletin of the Museum of Comparative Zoology 93: 263-389.
 * Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. II. The ants collected by the American Museum Congo Expedition. Bulletin of the American Museum of Natural History 45: 39-269.
 * Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bulletin of the American Museum of Natural History 45: 711-1004
 * Yeo K., and A. Hormenyo. 2007. A Rapid Survey of Ants in Ajenjua Bepo and Mamang River Forest Reserves, Eastern Region of Ghana. Pp 27-29. In McCullough, J., P. Hoke, P. Naskrecki, and Y. Osei-Owusu (eds.). 2008. A Rapid Biological Assessment of the Ajenjua Bepo and Mamang River Forest Reserves, Ghana. RAP Bulletin of Biological Assessment 50. Conservation International, Arlington, VA, USA.