Myrmica incompleta

This ant inhabits deciduous forests where it forms large colonies with many queens (polygynous) and consisting of numerous nests (polydomous) in moist soil and in mounds of moss. It is host for the xenobiotic ant.

Identification
The temperate M. incompleta is similar in appearance to the boreal Myrmica alaskensis; they are most easily separated geographically. Useful morphological characters to distinguish these two species are the clypeus, which bulges outward and has a concave margin in M. incompleta but is flattened and has a convex margin in M. alaskensis, and the rugae on the head, which are reticulate (net-like) in M. incompleta but parallel in M. alaskensis. (Ellison et.al. 2012)

Distribution based on Regional Taxon Lists
Nearctic Region: Canada, United States.

Biology
Ant of deciduous forests. This species feeds on honeydew from root-feeding aphids and coccids. It is a host to larvae of the the myrmecophilous syrphid fly Microdon albicomatus.

Nomenclature

 *  incompleta. Myrmica incompleta Provancher, 1881b: 359 (w.q.m.) CANADA.
 * [Also described as new by Provancher, 1883: 602.]
 * Wheeler, G.C. & Wheeler, J. 1953a: 116 (l.).
 * Junior synonym of laevinodis: Provancher, 1887: 247; Dalla Torre, 1893: 111; Donisthorpe, 1915d: 110; Smith, M.R. 1951a: 789.
 * Status as species: Francoeur & Béique, 1966: 141; Smith, D.R. 1979: 1349; Allred, 1982: 491; Wheeler, G.C. & Wheeler, J. 1986g: 23; Bolton, 1995b: 279; Mackay & Mackay, 2002: 161; Coovert, 2005: 36; Ellison, et al. 2012: 285.
 * Senior synonym of brevinodis (and its junior synonyms canadensis, subalpina): Francoeur & Béique, 1966: 141.
 * Senior synonym of frigida, sulcinodoides, whymperi: Shattuck & Cover, 2016: 18.
 * brevinodis. Myrmica rubra subsp. brevinodis Emery, 1895c: 312 (w.m.) U.S.A.
 * Wheeler, W.M. 1907b: 74 (q.); Wheeler, G.C. & Wheeler, J. 1953a: 116 (l.).
 * Subspecies of rubra: Forel, 1904a: 154; Wheeler, W.M. 1904e: 303; Wheeler, W.M. 1906b: 6; Wheeler, W.M. 1907b: 73 (redescription); Dennis, 1938: 288.
 * Status as species: Wheeler, W.M. 1914b: 53 (in text); Wheeler, W.M. 1916m: 587; Wheeler, W.M. 1917a: 502; Wheeler, W.M. 1917e: 16; Weber, 1939b: 150; Cole, 1942: 368; Weber, 1950b: 189 (redescription); Creighton, 1950a: 95; Smith, M.R. 1951a: 789; Cole, 1953d: 243.
 * Senior synonym of canadensis, subalpina: Weber, 1950b: 189.
 * Junior synonym of incompleta: Francoeur & Béique, 1966: 141.
 * canadensis. Myrmica brevinodis var. canadensis Wheeler, W.M. 1916m: 587 (w.q.m.) CANADA.
 * [First available use of Myrmica rubra subsp. brevinodis var. canadensis Wheeler, W.M. 1907b: 76 (w.q.m.) CANADA; unavailable (infrasubspecific) name.]
 * Subspecies of brevinodis: Wheeler, W.M. 1916m: 587; Emery, 1921f: 37; Menozzi, 1932b: 310.
 * Junior synonym of brevinodis: Weber, 1950b: 189.
 * frigida. Myrmica brevinodis var. frigida Wheeler, W.M. 1917a: 502 (w.) CANADA.
 * [First available use of Myrmica rubra subsp. brevinodis var. frigida Forel, 1902i: 699 (w.) CANADA; unavailable (infrasubspecific) name.]
 * Subspecies of brevinodis: Emery, 1921f: 37.
 * Junior synonym of sulcinodoides: Wheeler, W.M. 1917e: 16; Weber, 1950b: 195; Smith, M.R. 1951a: 789.
 * Junior synonym of incompleta: Shattuck & Cover, 2016: 18.
 * subalpina. Myrmica brevinodis var. subalpina Wheeler, W.M. 1917a: 503 (w.q.m.) U.S.A.
 * [First available use of Myrmica rubra subsp. brevinodis var. subalpina Wheeler, W.M. 1907b: 77 (w.q.m.) U.S.A.; unavailable (infrasubspecific) name.]
 * [Unresolved junior primary homonym of Myrmica kozlovi subsp. subalpina Ruzsky, 1915a: 438 (Bolton, 1995b: 283).]
 * Subspecies of brevinodis: Emery, 1921f: 37.
 * Junior synonym of brevinodis: Weber, 1950b: 189.
 * sulcinodoides. Myrmica brevinodis var. sulcinodoides Wheeler, W.M. 1917a: 502 (w.) U.S.A.
 * [First available use of Myrmica rubra subsp. brevinodis var. sulcinodoides Emery, 1895c: 313 (w.) U.S.A.; unavailable (infrasubspecific) name.]
 * Wheeler, W.M. 1907b: 76 (q.m.).
 * Subspecies of brevinodis: Wheeler, W.M. 1917a: 502; Wheeler, W.M. 1917e: 16; Emery, 1921f: 37; Cole, 1942: 368; Weber, 1950b: 195 (redescription); Smith, M.R. 1951a: 789.
 * Subspecies of incompleta: Smith, D.R. 1979: 1349.
 * Junior synonym of brevinodis: Creighton, 1950a: 96.
 * Status as species: Hunt & Snelling, 1975: 21; Mackay, Lowrie, et al. 1988: 92 (in key).
 * Senior synonym of frigida: Wheeler, W.M. 1917e: 16; Weber, 1950b: 195.
 * Junior synonym of whymperi: Wheeler, W.M. 1917a: 502.
 * [Note: Wheeler gives sulcinodoides as senior synonym, but whymperi has priority (Bolton, 1995b: 284).]
 * Junior synonym of incompleta: Shattuck & Cover, 2016: 18.
 * whymperi. Myrmica brevinodis var. whymperi Forel, 1913l: 215 (w.) U.S.A.
 * [First available use of Myrmica rubra r. brevinodis var. whymperi Forel, 1904a: 154 (w.) U.S.A.; unavailable (infrasubspecific) name.]
 * Synonym of sulcinodoides: Wheeler, W.M. 1917a: 502; Smith, M.R. 1951a: 789.
 * [Note: Wheeler, and Smith, M.R., give sulcinodoides as senior synonym, but whymperi has priority (Bolton, 1995b: 284).]
 * Status as species: Bolton, 1995b: 284; Mackay & Mackay, 2002: 168.
 * Junior synonym of incompleta: Shattuck & Cover, 2016: 18.

References based on Global Ant Biodiversity Informatics

 * Allred D. M. 1982. Ants of Utah. The Great Basin Naturalist 42: 415-511.
 * Allred, D.M. 1982. The ants of Utah. Great Basin Naturalist 42:415-511.
 * Blacker, N.C. 1992. Some Ants from Southern Vancouver Island, British Columbia. J. Entomol. Soc. Bri. Columbia 89:3-12.
 * Blacker, N.C. 1992. Some ants (Hymenoptera: Formicidae) from Southern Vancouver Island, British Columbia. Journal of the Entomological Society of British Columbia 89:3-12
 * Borchert, H.F. and N.L. Anderson. 1973. The Ants of the Bearpaw Mountains of Montana (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 46(2):200-224
 * Burrill A. C., and M. R. Smith. 1919. A key to the species of Wisconsin ants, with notes on their habits. Ohio Journal of Science 19: 279-292.
 * Choate B., and F. A. Drummond. 2012. Ant Diversity and Distribution (Hymenoptera: Formicidae) Throughout Maine Lowbush Blueberry Fields in Hancock and Washington Counties. Environ. Entomol. 41(2): 222-232.
 * Choate B., and F. A. Drummond. 2013. The influence of insecticides and vegetation in structuring Formica Mound ant communities (Hymenoptera: Formicidae) in Maine lowbush blueberry. Environ. Entomol. 41(2): 222-232.
 * Cole A. C., Jr. 1936. An annotated list of the ants of Idaho (Hymenoptera: Formicidae). Canadian Entomologist 68: 34-39.
 * Cole A. C., Jr. 1942. The ants of Utah. American Midland Naturalist 28: 358-388.
 * Cole, A.C. 1936. An annotated list of the ants of Idaho (Hymenoptera; Formicidae). Canadian Entomologist 68(2):34-39
 * Del Toro, I. 2010. PERSONAL COMMUNICATION. MUSEUM RECORDS COLLATED BY ISRAEL DEL TORO
 * Drummond F. A., A. M. llison, E. Groden, and G. D. Ouellette. 2012. The ants (Formicidae). In Biodiversity of the Schoodic Peninsula: Results of the Insect and Arachnid Bioblitzes at the Schoodic District of Acadia National Park, Maine. Maine Agricultural and forest experiment station, The University of Maine, Technical Bulletin 206. 217 pages
 * Ellison A. M., E. J. Farnsworth, and N. J. Gotelli. 2002. Ant diversity in pitcher-plant bogs of Massachussetts. Northeastern Naturalist 9(3): 267-284.
 * Ellison A. M., and E. J. Farnsworth. 2014. Targeted sampling increases knowledge and improves estimates of ant species richness in Rhode Island. Northeastern Naturalist 21(1): NENHC-13NENHC-24.
 * Forel A. 1904. Fourmis de British Columbia récoltées par M. Ed. Whymper. Annales de la Société Entomologique de Belgique 48: 152-155.
 * Francoeur, A. 1997. Ants (Hymenoptera: Formicidae) of the Yukon. Pages 901 910 in H.V. Danks and J.A. Downes (Eds.), Insects of the Yukon. Biological Survey of Canada (Terrestrial Arthropods), Ottawa.
 * Glasier J. R. N., S. E. Nielsen, J. Acorn, and J. Pinzon. 2019. Boreal sand hills are areas of high diversity for Boreal ants (Hymenoptera: Formicidae). Diversity 11, 22; doi:10.3390/d11020022.
 * Glasier, J. Alberta Ants. AntWeb.
 * Gregg R. E. 1946. The ants of northeastern Minnesota. American Midland Naturalist 35: 747-755.
 * Gregg, R.T. 1963. The Ants of Colorado.
 * Hoey-Chamberlain R. V., L. D. Hansen, J. H. Klotz and C. McNeeley. 2010. A survey of the ants of Washington and Surrounding areas in Idaho and Oregon focusing on disturbed sites (Hymenoptera: Formicidae). Sociobiology. 56: 195-207
 * Ivanov, K. 2019. The ants of Ohio (Hymenoptera, Formicidae): an updated checklist. Journal of Hymenoptera Research 70: 65–87.
 * Johnson R. Personnal Database. Accessed on February 5th 2014 at http://www.asu.edu/clas/sirgtools/resources.htm
 * Kannowski P. B. 1956. The ants of Ramsey County, North Dakota. American Midland Naturalist 56(1): 168-185.
 * Knowlton G. F. 1970. Ants of Curlew Valley. Proceedings of the Utah Academy of Sciences, Arts and Letters 47(1): 208-212.
 * La Rivers I. 1968. A first listing of the ants of Nevada. Biological Society of Nevada, Occasional Papers 17: 1-12.
 * Lavigne R., and T. J. Tepedino. 1976. Checklist of the insects in Wyoming. I. Hymenoptera. Agric. Exp. Sta., Univ. Wyoming Res. J. 106: 24-26.
 * Lesica P., and P. B. Kannowski. 1998. Ants Create Hummocks and Alter Structure and Vegetation of a Montana Fen. Am. Midl. Nat. 139: 5868
 * Lidgren, B.S. and A.M. MacIsaac. 2002. A Preliminary Study of Ant Diversity and of Ant Dependence on Dead Wood in Central Interior British Columbia. USDA Forest Service Gen. Tech. Rep. PSW-GTR-181.
 * Lindgren, B.S. and A.M. MacIsaac. 2002. Ant dependence on dead wood in Central Interior British Columbia. USDA Forest Service Gen. Tech. Rep.PSW-GTR-181
 * Mackay W. P., and E. E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
 * Mackay, W., D. Lowrie, A. Fisher, E. Mackay, F. Barnes and D. Lowrie. 1988. The ants of Los Alamos County, New Mexico (Hymenoptera: Formicidae). pages 79-131 in J.C. Trager, editor, Advances in Myrmecololgy.
 * Mallis A. 1941. A list of the ants of California with notes on their habits and distribution. Bulletin of the Southern California Academy of Sciences 40: 61-100.
 * Merle W. W. 1939. An Annotated List of the Ants of Maine (Hymenoptera: Formicidae). Entomological News. 50: 161-165
 * Munsee J. R., W. B. Jansma, and J. R. Schrock. 1986. Revision of the checklist of Indiana ants with the addition of five new species (Hymenoptera: Formicidae). Indiana Academy of Science 95: 265-274.
 * Ouellette G. D., F. A. Drummond, B. Choate and E. Groden. 2010. Ant diversity and distribution in Acadia National Park, Maine. Environmental Entomology 39: 1447-1556
 * Piers H. 1922. List of a small collection of ants (Formicidae) obtained in Queen's County, Nova Scotia, by the late Walter H. Prest. Proceedings and Transactions of the Nova Scotian Institute of Science, 15(4), 169-173.
 * Prest W. H., and H. Piers. 1922. List of a Small Collection of Ants (Formicidae) obtained in Queen's County, Nova Scotia. Nova Scotian Institute of Science 15(4): 169-173.
 * Procter W. 1938. Biological survey of the Mount Desert Region. Part VI. The insect fauna. Philadelphia: Wistar Institute of Anatomy and Biology, 496 pp.
 * Ratchford, J.S., S.E. Wittman, E.S. Jules, A.M. Ellison, N.J. Gotelli and N.J. Sanders. 2005. The effects of fire, local environment and time on ant assemblages in fens and forests. Diversity and Distributions 11:487-497.
 * Rees D. M., and A. W. Grundmann. 1940. A preliminary list of the ants of Utah. Bulletin of the University of Utah, 31(5): 1-12.
 * Sharplin, J. 1966. An annotated list of the Formicidae (Hymenoptera) of Central and Southern Alberta. Quaetiones Entomoligcae 2:243-253
 * Smith F. 1941. A list of the ants of Washington State. The Pan-Pacific Entomologist 17(1): 23-28.
 * Smith M. R. 1952. On the collection of ants made by Titus Ulke in the Black Hills of South Dakota in the early nineties. Journal of the New York Entomological Society 60: 55-63.
 * Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
 * Weber N. A. 1950. A revision of the North American ants of the genus Myrmica Latreille with a synopsis of the Palearctic species. III. Annals of the Entomological Society of America 43: 189-226.
 * Wheeler G. C., J. N. Wheeler, and P. B. Kannowski. 1994. Checklist of the ants of Michigan (Hymenoptera: Formicidae). The Great Lakes Entomologist 26(4): 297-310
 * Wheeler G. C., and E. W. Wheeler. 1944. Ants of North Dakota. North Dakota Historical Quarterly 11:231-271.
 * Wheeler G. C., and J. Wheeler. 1986. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp.
 * Wheeler G. C., and J. Wheeler. 1987. A Checklist of the Ants of South Dakota. Prairie Nat. 19(3): 199-208.
 * Wheeler W. M. 1905. Ants from the summit of Mount Washington. Psyche (Cambridge) 12: 111-114.
 * Wheeler W. M. 1906. Fauna of New England. 7. List of the Formicidae. Occasional Papers of the Boston Society of Natural History 7: 1-24.
 * Wheeler W. M. 1907. Notes on a new guest-ant, Leptothorax glacialis, and the varieties of Myrmica brevinodis Emery. Bulletin of the Wisconsin Natural History Society 5: 70-83.
 * Wheeler W. M. 1916. Formicoidea. Formicidae. Pp. 577-601 in: Viereck, H. L. 1916. Guide to the insects of Connecticut. Part III. The Hymenoptera, or wasp-like insects, of Connecticut. Connecticut State Geological and Natural History Survey. Bulletin 22: 1-824.
 * Wheeler W. M. 1917. The mountain ants of western North America. Proceedings of the American Academy of Arts and Sciences 52: 457-569.
 * Wheeler W. M. 1928. Ants of Nantucket Island, Mass. Psyche (Cambridge) 35: 10-11.
 * Wheeler, G.C. and J. Wheeler. 1978. Mountain ants of Nevada. Great Basin Naturalist 35(4):379-396
 * Wheeler, G.C. and J. Wheeler. 1988. A checklist of the ants of Montana. Psyche 95:101-114
 * Wheeler, G.C. and J. Wheeler. 1988. A checklist of the ants of Wyoming. Insecta Mundi 2(3&4):230-239
 * Wheeler, G.C., J. Wheeler and P.B. Kannowski. 1994. CHECKLIST OF THE ANTS OF MICHIGAN (HYMENOPTERA: FORMICIDAE). Great Lakes Entomologist 26:1:297-310
 * Wheeler, G.C., J. Wheeler, T.D. Galloway and G.L. Ayre. 1989. A list of the ants of Manitoba. Proceedings of the Entomological Society of Manitoba 45:34-49
 * Wing M. W. 1939. An annotated list of the ants of Maine (Hymenoptera: Formicidae). Entomological News 50:161-165.