Checklist of Myrmica species

The following species and subspecies belong to the genus Myrmica. Synonyms are listed under their senior names. For valid names only see Myrmica species and for distribution information see Myrmica species by Country.

A
ademonia Bolton, 1995 (Democratic Peoples Republic of Korea, Republic of Korea, Russian Federation)

• Myrmica aspersa Kupyanskaya, 1990

adrijae Bharti, 2012 (India)

afghanica Radchenko & Elmes, 2003 (Afghanistan, India)

aimonissabaudiae Menozzi, 1939 (Afghanistan, Bhutan, India, Nepal, Pakistan)

• Myrmica dicaporiaccoi Menozzi, 1939

alaskensis Wheeler, W.M., 1917 (Canada, United States)

• Myrmica brevinodis kuschei Wheeler, W.M., 1917

aloba Forel, 1909 (Balearic Islands, China, France, Iberian Peninsula, Portugal, Spain)

• Myrmica albuferensis Lomnicki, 1925

• Myrmica rolandi Bondroit, 1918

alperti Elmes & Radchenko, 2009 (Nepal)

americana Weber, 1939 (Canada, United States)

anatolica Elmes, Radchenko & Aktaç, 2002 (Turkey)

angulata Radchenko, Zhou & Elmes, 2001 (China, Vietnam)

angulinodis Ruzsky, 1905 (China, Democratic Peoples Republic of Korea, Mongolia, Republic of Korea, Russian Federation)

• Myrmica incurvata Collingwood, 1976

• Myrmica saposhnikovi baikalensis Pisarski, 1969

arisana Wheeler, W.M., 1930 (Taiwan)

• Myrmica kurokii tipuna Santschi, 1937

arnoldii Dlussky, 1963 (Mongolia, Russian Federation)

assimilis Spinola, 1851 (Brazil)

B
bactriana Ruzsky, 1915 (China, Nepal, Tibet)

• Myrmica ruzskyana Radchenko & Elmes, 2010

• Myrmica smythiesi exigua Ruzsky, 1915

• Myrmica tibetana furva Ruzsky, 1915

bakurianica Arnoldi, 1970 (Georgia, Russian Federation)

bergi Ruzsky, 1902 (Austria, Azerbaijan, Balearic Islands, Belgium, Finland, Germany, Gibraltar, Iran, Kazakhstan, Kyrgyzstan, Romania, Russian Federation, Switzerland, Turkey)

• Myrmica bergi barchanica Ruzsky, 1905

• Myrmica bergi fortior Crawley, 1920

• Myrmica bergi persiana Weber, 1947

• Myrmica kamyschiensis Arnoldi, 1934

• Myrmica rugosa kirgisica Ruzsky, 1903

bessarabicus Nasonov, 1889 (Armenia, Austria, Germany, Luxembourg, Netherlands, Poland, Republic of Moldova, Romania, Russian Federation, Slovakia, Sweden, Turkmenistan)

bibikoffi Kutter, 1963 (Germany, Spain, Switzerland)

boltoni Radchenko & Elmes, 1998 (India, Nepal)

brancuccii Radchenko, Elmes & Collingwood, 1999 (India, Nepal, Pakistan)

breviceps Smith, F., 1878

brevispinosa Wheeler, W.M., 1917 (Canada, United States)

• Myrmica rubra decedens Wheeler, W.M., 1917

C
cachmiriensis Forel, 1904 (India, Pakistan)

• Myrmica smythiesi lutescens Menozzi, 1939

caeca Jerdon, 1851

cagnianti Espadaler, 1996 (Algeria, Morocco)

caucasicola Arnoldi, 1934 (Armenia, Azerbaijan, Georgia, Turkey)

colax (Cole, 1957) (United States)

collingwoodi Radchenko & Elmes, 1998 (Bhutan, India)

commarginata Ruzsky, 1905 (Mongolia, Russian Federation)

constricta Karavaiev, 1934 (Austria, Bulgaria, Czech Republic, Finland, Germany, Greece, Hungary, Iran, Italy, Poland, Romania, Serbia, Switzerland, Ukraine)

• Myrmica rugulosa minuta Karavaiev, 1929

• Myrmica rugulosa slobodensis Arnoldi, 1934

corrugata Say, 1836

crassirugis Francoeur, 2007 (Canada, United States)

curiosa Radchenko, Zhou & Elmes, 2008 (China)

cursor Smith, F., 1878

curvispinosa Bharti & Sharma, 2013 (India)

curvithorax Bondroit, 1920 (Austria, Belarus, Bulgaria, Czech Republic, Germany, Hungary, Kazakhstan, Poland, Romania, Russian Federation, Slovakia, Spain, Switzerland, Ukraine)

• Myrmica slovaca Sadil, 1952

D
deplanata Emery, 1921 (Austria, Azerbaijan, Bulgaria, China, Czech Republic, Georgia, Hungary, Iran, Kazakhstan, Poland, Romania, Russian Federation, Slovakia, Turkey, Turkmenistan, Ukraine)

• Myrmica lobicornis plana Karavaiev, 1927

• Myrmica moravica Soudek, 1922

• Myrmica moravica dagmarae Sadil, 1939

detritinodis Emery, 1921 (Canada, United States)

dimidiata Say, 1836

discontinua Weber, 1939 (United States)

displicentia Bolton, 1995 (Russian Federation)

• Myrmica bicolor Kupyanskaya Kupyanskaya, 1986

divergens Karavaiev, 1931 (Mongolia, Russian Federation)

diversa Buckley, 1867

dongi Chen, Zhou & Huang, 2016 (China)

draco Radchenko, Zhou & Elmes, 2001 (China)

dshungarica Ruzsky, 1905 (Kazakhstan, Kyrgyzstan, Tajikistan, Uzbekistan)

• Leptothorax svartshevskii Karavaiev, 1916

• Myrmica laevinodis minuta Ruzsky, 1905

• Myrmica minuta iskanderi Arnoldi, 1976

• Myrmica minuta tarbinskii Arnoldi, 1976

• Myrmica rugulosa ruginodiformis Karavaiev, 1929

E
eidmanni Menozzi, 1930 (China, Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

• Myrmica lobicornis littoralis Weber, 1948

elbrusi Radchenko & Yusupov, 2012 (Russian Federation)

elmesi Bharti & Sharma, 2011 (India)

emeryi Radchenko & Elmes, 2009 (Borneo, Indonesia, Malaysia)

ereptrix Bolton, 1988 (India)

excelsa Kupyanskaya, 1990 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Myrmica cadusa Kim, Park & Kim, 1997

• Myrmica sinica Wu, J. & Wang, C., 1995

F
ferganensis Finzi, 1926 (Kyrgyzstan)

forcipata Karavaiev, 1931 (China, Mongolia, Russian Federation)

foreliana Radchenko & Elmes, 2001 (India)

• Myrmica smythiesii carbonaria Forel, 1902

fortior Forel, 1904 (India)

fracticornis Forel, 1901 (Canada, United States)

G
gallienii Bondroit, 1920 (Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iran, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Sweden, Switzerland, Turkey, Ukraine)

• Myrmica jacobsoni Kutter, 1963

• Myrmica limanica obensis Arnoldi, 1970

• Myrmica rugulosa limanica Karavaiev, 1934

gebaueri Seifert, Schultz, Ritz & Ritz, 2018 (Tibet)

gigantea (Collingwood, 1962) (Myanmar)

glacialis Emery, 1921 (Canada, United States)

granulinodis Nylander, 1846

H
hamulata Weber, 1939 (United States)

hecate Weber, 1947 (India, Nepal)

hellenica Finzi, 1926 (Armenia, Austria, Azerbaijan, Bulgaria, Croatia, Finland, Georgia, Germany, Greece, Iran, Italy, Montenegro, Poland, Romania, Russian Federation, Serbia, Slovenia, Switzerland, Turkey, Ukraine)

• Myrmica rugulosa caucasica Arnoldi, 1934

• Myrmica rugulosa rugulososcabrinodis Karavaiev, 1929

heterorhytida Radchenko & Elmes, 2009 (China)

hirsuta Elmes, 1978 (Austria, Belgium, Croatia, Czech Republic, Denmark, Finland, France, Germany, Greece, Hungary, Netherlands, Poland, Russian Federation, Serbia, Slovakia, Sweden, United Kingdom of Great Britain and Northern Ireland)

hlavaci Radchenko & Elmes, 2009 (China)

huaii Chen, Zhou & Huang, 2016 (China)

I
incompleta Provancher, 1881 (Canada, United States)

• Myrmica brevinodis sulcinodoides Wheeler, W.M., 1917

• Myrmica rubra brevinodis Emery, 1895

• Myrmica rubra canadensis Wheeler, W.M., 1916

• Myrmica rubra frigida Wheeler, W.M., 1917

• Myrmica rubra subalpina Wheeler, W.M., 1917

• Myrmica whymperi Forel, 1913

indica Weber, 1950 (Bhutan, India, Nepal)

inezae Forel, 1902 (India)

inflecta Say, 1836

inucta Radchenko & Elmes, 2006 (Kazakhstan)

J
jennyae Elmes, Radchenko & Aktaç, 2002 (Russian Federation, Turkey)

jessensis Forel, 1901 (Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Myrmica hyungokae Elmes, Radchenko & Kim, 2001

jucunda Smith, F., 1861

juglandeti Arnoldi, 1976 (Kyrgyzstan)

• Myrmica tianshanica Arnoldi, 1976

• Myrmica tianshanica alajensis Arnoldi, 1976

K
kabylica (Cagniant, 1970) (Algeria)

kamtschatica Kupyanskaya, 1986 (Democratic Peoples Republic of Korea, Mongolia, Russian Federation)

• Myrmica aborigenica Zhigulskaya, 1991

karavajevi (Arnoldi, 1930) (Austria, Belarus, Belgium, Czech Republic, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Italy, Norway, Poland, Romania, Russian Federation, Spain, Sweden, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Myrmica faniensis Boven, 1970

• Sifolinia pechi Samsinak, 1957

• Sifolinia winterae Kutter, 1973

kasczenkoi Ruzsky, 1905 (Democratic Peoples Republic of Korea, Mongolia, Republic of Korea, Russian Federation)

khamensis Ruzsky, 1915

kirghisorum Arnoldi, 1976 (Kyrgyzstan, Mongolia, Russian Federation, Tajikistan)

• Myrmica lobicornis foreliella Arnoldi, 1976

koreana Elmes, Radchenko & Kim, 2001 (China, Democratic Peoples Republic of Korea, Kazakhstan, Mongolia, Republic of Korea, Russian Federation)

kothiensis Bharti & Sharma, 2013 (India)

kotokui Forel, 1911 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Myrmica ruginodis orientalis Karavaiev, 1926

kozakorum Radchenko & Elmes, 2010 (Russian Federation, Ukraine)

kozlovi Ruzsky, 1915 (China, India, Nepal, Tibet)

• Myrmica kozlovi mekongi Ruzsky, 1915

• Myrmica kozlovi ruzskyi Weber Weber, 1947

• Myrmica kozlovi subalpina Ruzsky, 1915

• Myrmica kozlovi subbrevispinosa Ruzsky, 1915

• Myrmica specularis Donisthorpe, 1929

kryzhanovskii Arnoldi, 1976 (Kyrgyzstan, Russian Federation, Tajikistan)

kurokii Forel, 1907 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)

• Myrmica chinensis Viehmeyer, 1922

• Myrmica helleri Viehmeyer, 1922

• Myrmica kozlovi ruzskyi Kiseleva Kiseleva, 1925

L
lampra Francoeur, 1968 (Canada, United States)

latifrons Stärcke, 1927 (Canada, United States)

• Myrmica schencki emeryana Cole, 1942

latra Bharti, Radchenko & Sasi, 2016 (India)

laurae (Emery, 1907) (Italy)

• Myrmica samnitica Mei, 1987

lemasnei Bernard, 1967 (France, Iberian Peninsula, Spain)

lineolata Buckley, 1867

liui Chen, Zhou & Huang, 2016 (China)

lobicornis Nylander, 1846 (Albania, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Canary Islands, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Poland, Republic of Korea, Romania, Russian Federation, Slovakia, Slovenia, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Myrmica denticornis Curtis, 1854

• Myrmica lobicornis alpestris Arnoldi, 1934

• Myrmica lobicornis angustifrons Stärcke, 1927

• Myrmica lobicornis arduennae Bondroit, 1911

• Myrmica lobicornis foreli Santschi, 1931

• Myrmica lobicornis kieviensis Karavaiev, 1934

• Myrmica lobicornis lissahorensis Stitz, 1939

• Myrmica schencki brunescens Karavaiev, 1929

• Myrmica schencki burtshakabramovitshi Karavaiev, 1929

• Myrmica schencki starki Karavaiev, 1929

lobifrons Pergande, 1900 (Canada, United States)

lobulicornis Nylander, 1857 (Andorra, Austria, Canary Islands, France, Iberian Peninsula, Italy, Slovenia, Spain, Switzerland)

• Myrmica arduennae pyrenaea Bondroit, 1918

• Myrmica lobicornis alpina Stärcke, 1927

• Myrmica lobicornis apennina Stärcke, 1927

lonae Finzi, 1926 (Austria, Belgium, Bulgaria, Croatia, Czech Republic, Finland, Germany, Greece, Hungary, Netherlands, Norway, Poland, Romania, Russian Federation, Slovakia, Slovenia, Turkey, United Kingdom of Great Britain and Northern Ireland)

longisculpta Bharti & Sharma, 2011 (India)

luctuosa Smith, F., 1878

luteola Kupyanskaya, 1990 (China, Democratic Peoples Republic of Korea, Republic of Korea, Russian Federation, Taiwan)

• Myrmica zhengi Ma & Xu, S., 2011

M
margaritae Emery, 1889 (India, Myanmar)

martensi Radchenko & Elmes, 1998 (India, Nepal)

martini Seifert, Yazdi & Schultz, 2014 (Andorra, France, Spain)

mexicana Wheeler, W.M., 1914 (Mexico)

mifui Chen, Zhou & Huang, 2016 (China)

mirabilis Elmes & Radchenko, 1998 (Taiwan)

mixta Radchenko & Elmes, 2008 (China)

montana Buckley, 1867

monticola Creighton, 1950 (United States)

multiplex Radchenko & Elmes, 2009 (China)

myrmicoxena Forel, 1895 (Switzerland)

N
nearctica Weber, 1939 (Canada, United States)

nefaria Bharti, 2012 (India)

nitida Radchenko & Elmes, 1999 (India)

O
obscura Finzi, 1926 (Italy)

obscurata Motschoulsky, 1863

onoyamai Radchenko & Elmes, 2006 (Japan)

ordinaria Radchenko & Elmes, 1999 (India, Pakistan)

orthostyla Arnoldi, 1976 (Kyrgyzstan, Russian Federation)

oui Chen, Zhou & Huang, 2016 (China)

P
pachei Forel, 1906 (India, Nepal)

pararitae Radchenko & Elmes, 2008 (China)

pelops Seifert, 2003 (Greece)

petita Radchenko & Elmes, 1999 (India)

phalacra Radchenko & Elmes, 2009 (China)

pilinodis Motschoulsky, 1863

pinetorum Wheeler, W.M., 1905 (United States)

pisarskii Radchenko, 1994 (Mongolia, Russian Federation)

pleiorhytida Radchenko & Elmes, 2009 (China)

poldii Radchenko & Rigato, 2008 (China)

polyglypta Radchenko & Rigato, 2008 (China)

pseudorugosa Bharti, 2012 (India, Pakistan)

pulchella Santschi, 1937 (Taiwan)

• Myrmica margaritae formosae Wheeler, W.M., 1929

punctinops Francoeur, 2007 (United States)

punctiventris Roger, 1863 (United States)

• Myrmica punctiventris isfahani Forel, 1922

Q
quebecensis Francoeur, 1981 (Canada)

R
radchenkoi Bharti & Sharma, 2011 (India)

ravasinii Finzi, 1923 (Albania, Armenia, Georgia, Greece, Russian Federation, Turkey)

religiosa Bharti & Sharma, 2013 (India)

rhynchophora Foerster, 1850

rhytida Radchenko & Elmes, 1999 (India)

rigatoi Radchenko & Elmes, 1998 (India, Pakistan)

ritae Emery, 1889 (China, India, Myanmar, Nepal, Thailand)

rubra (Linnaeus, 1758) (Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Canada, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Japan, Jersey, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Mongolia, Montenegro, Netherlands, Norway, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland, United States)

• Myrmica laevinodis Nylander, 1846

• Myrmica longiscapus Curtis, 1854

• Myrmica microrubra Seifert, 1993

• Myrmica rubra bruesi Weber, 1947

• Myrmica rubra champlaini Forel, 1901

• Myrmica rubra europaea Finzi, 1926

rubra neolaevinodis Forel, 1901 (United States)

ruginodis Nylander, 1846 (Albania, Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Latvia, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Poland, Portugal, Republic of Korea, North Macedonia, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, United Kingdom of Great Britain and Northern Ireland)

• Myrmica diluta Nylander, 1849

• Myrmica kurokii sontica Santschi, 1937

• Myrmica rubra macrogyna Brian, M.V. & Brian, A.D., 1949

• Myrmica rubra microgyna Brian, M.V. & Brian, A.D., 1949

• Myrmica rubra mutata Sadil, 1952

• Myrmica rubra ruginodolaevinodis Forel, 1874

• Myrmica rubra yoshiokai Weber, 1947

• Myrmica ruginodis silvestrii Wheeler, W.M., 1928

rugiventris (Smith, M.R., 1943) (United States)

rugosa Mayr, 1865 (Bhutan, China, India, Nepal)

rugulosa Nylander, 1849 (Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Latvia, Lithuania, Luxembourg, Netherlands, Norway, Poland, Romania, Russian Federation, Slovakia, Slovenia, Sweden, Switzerland, Turkey, Ukraine)

• Myrmica clandestina Foerster, 1850

• Myrmica rugulosa sulcinodorugulosa Nasonov, 1889

rupestris Forel, 1902 (Afghanistan, Bhutan, China, India, Nepal)

• Myrmica everesti Donisthorpe, 1929

• Myrmica rugosa debilior Forel, 1902

• Myrmica rugosa rugososmythiesi Forel, 1902

S
sabuleti Meinert, 1861 (Albania, Andorra, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, Channel Islands, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Iran, Jersey, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Myrmica rubra scabrinodolobicornis Forel, 1874

salina Ruzsky, 1905 (Austria, Azerbaijan, Croatia, Czech Republic, Georgia, Kazakhstan, Kyrgyzstan, Romania, Russian Federation, Slovenia, Turkey)

• Myrmica georgica Seifert, 1987

• Myrmica scabrinodis ahngeri Karavaiev, 1926

• Myrmica tobiasi Radchenko & Elmes, 2004

saposhnikovi Ruzsky, 1904 (China, Kazakhstan, Kyrgyzstan, Republic of Korea, Russian Federation)

• Myrmica saposhnikovi bergiana Tarbinsky, 1976

scabrata Buckley, 1867

scabrinodis Nylander, 1846 (Albania, Andorra, Armenia, Austria, Azerbaijan, Balearic Islands, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Jersey, Kazakhstan, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Korea, North Macedonia, Republic of Moldova, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland, United States)

• Myrmica pilosiscapus Bondroit, 1920

• Myrmica rolandi reticulata Stärcke, 1942

• Myrmica scabrinodis rugulosoides Forel, 1915

• Myrmica scabrinodis scabrinodosabuleti Sadil, 1952

• Sommimyrma symbiotica Menozzi, 1925

scabrinodis intermedia Kuznetsov-Ugamsky, 1927

schencki Viereck, 1903 (Andorra, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Canary Islands, China, Croatia, Czech Republic, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kyrgyzstan, Latvia, Lithuania, Montenegro, Netherlands, Norway, Poland, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)

• Myrmica betuliana Ruzsky, 1946

• Myrmica schencki kutteri Finzi, 1926

• Myrmica schenckioides Boer & Noordijk, 2005

• Myrmica subopaca Arnoldi, 1934

schoedli Radchenko, Elmes & Bui, 2006 (Vietnam)

schulzi Radchenko & Elmes, 2009 (China)

sculptiventris Radchenko & Elmes, 2009 (China)

semiparasitica Francoeur, 2007 (United States)

serica Wheeler, W.M., 1928 (China, Taiwan, Vietnam)

siciliana Radchenko, Elmes & Alicata, 2006 (Italy)

sinensis Radchenko, Zhou & Elmes, 2001 (China)

sinoschencki Radchenko & Elmes, 2008 (China)

smythiesii Forel, 1902 (China, India, Nepal)

• Myrmica smythiesi himalayana Weber, 1947

spatulata Smith, M.R., 1930 (United States)

specioides Bondroit, 1918 (Andorra, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Italy, Montenegro, Netherlands, Norway, Poland, Portugal, Romania, Russian Federation, Slovenia, Spain, Turkey, Turkmenistan, Ukraine, United Kingdom of Great Britain and Northern Ireland, United States)

• Leptothorax nevodovskii Karavaiev, 1926

• Myrmica balcanica Sadil, 1952

• Myrmica balcanica scabrinodoides Sadil, 1952

• Myrmica puerilis Stärcke, 1942

• Myrmica puerilis dolens Stärcke, 1942

• Myrmica rugulosoides striata Finzi, 1926

• Myrmica sancta tshuliensis Arnoldi, 1976

• Myrmica scabrinodis sancta Karavaiev, 1926

• Myrmica scabrinodis turcica Santschi, 1931

• Tetramorium silvestrianum Emery, 1924

spinosior Santschi, 1931 (Andorra, France, Iberian Peninsula, Italy, Spain, Turkey)

stangeana Ruzsky, 1902 (Armenia, China, Czech Republic, Georgia, Kazakhstan, Kyrgyzstan, Romania, Russian Federation, Slovakia, Ukraine)

• Myrmica scabrinodis lacustris Ruzsky, 1905

• Myrmica stangeana medvedevi Arnoldi, 1970

striolagaster Cole, 1953 (Mexico, United States)

sublanuginosa Buckley, 1867

sulcinodis Nylander, 1846 (Albania, Andorra, Armenia, Austria, Belgium, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, Georgia, Germany, Greece, Iberian Peninsula, Latvia, Mongolia, Montenegro, Netherlands, Poland, Republic of Korea, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, United Kingdom of Great Britain and Northern Ireland)

• Myrmica myrmecophila Wasmann, 1910

• Myrmica perelegans Curtis, 1854

• Myrmica ruginodis sulcinodoruginodis Donisthorpe, 1915

• Myrmica sulcinodis derzhavini Ruzsky, 1920

• Myrmica sulcinodis eximia Kupyanskaya, 1990

• Myrmica sulcinodis nigripes Ruzsky, 1895

• Myrmica sulcinodis sulcinodoscabrinodis Forel, 1915

• Myrmica sulcinodis vicaria Kuznetsov-Ugamsky, 1928

• Myrmica sulcinodoscabrinodis Ruzsky, 1895

T
tahoensis Weber, 1948 (Mexico, United States)

taibaiensis Wei, Zhou & Liu, 2001 (China)

tamarae Elmes, Radchenko & Aktaç, 2002 (Turkey)

tenuispina Ruzsky, 1905 (Afghanistan, India, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan)

tibetana Mayr, 1889 (Afghanistan, China, Tibet)

titanica Radchenko & Elmes, 2001 (Vietnam)

transsibirica Radchenko, 1994 (Democratic Peoples Republic of Korea, Mongolia, Republic of Korea, Russian Federation)

• Myrmica carinata Kupyanskaya, 1990

• Myrmica taediosa Bolton, 1995

trullicornis Buren, 1944 (United States)

tschekanovskii Radchenko, 1994 (Russian Federation)

tulinae Elmes, Radchenko & Aktaç, 2002 (China, France, Georgia, Greece, Italy, Netherlands, Poland, Turkey)

U
urbanii Radchenko & Elmes, 1998 (China, India, Nepal)

ussuriensis Kuznetsov-Ugamsky, 1928

V
vandeli Bondroit, 1920 (Austria, Bulgaria, China, Czech Republic, France, Germany, Hungary, Iberian Peninsula, Poland, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)

varisculpta Radchenko & Rigato, 2009 (India)

villosa Radchenko & Elmes, 1999 (Bhutan, India)

vittata Radchenko & Elmes, 1999 (India, Pakistan)

W
wangi Chen, Zhou & Huang, 2016 (China)

wardi Radchenko & Elmes, 1999 (India, Pakistan)

weberi Elmes & Radchenko, 2009 (Bhutan, India, Nepal)

weii Radchenko & Zhou, 2008 (China)

wesmaeli Bondroit, 1918 (Andorra, China, France, Iberian Peninsula, Portugal, Spain)

wheeleri Weber, 1939 (United States)

wheelerorum Francoeur, 2007 (United States)

williamsi Radchenko & Elmes, 1999 (India)

wittmeri Radchenko & Elmes, 1999 (India, Pakistan)

X
xavieri Radchenko, Elmes & Savolainen, 2008 (Spain)

Y
yamanei Radchenko & Elmes, 2001 (Vietnam)

yani Chen, Zhou & Huang, 2016 (China)

yezomonticola Terayama, 2013 (Japan)

yunnanensis Radchenko & Elmes, 2009 (China)

Z
zojae Radchenko, 1994 (Russian Federation)

=Fossil Species=

A
andrannae Zharkov and Dubovikoff, 2022

angusticollis Heer, 1849

E
eocenica Radchenko, Dlussky & Elmes, 2007

I
intermedia (Wheeler, W.M., 1915)

L
longispinosa Mayr, 1868

M
minor Heer, 1849

molassica Heer, 1849

N
nebulosa Novak, O., 1878

nungesseri Jessen, 2020

O
obsoleta Heer, 1849

P
paradoxa Radchenko, Dlussky & Elmes, 2007

R
rudis (Mayr, 1868)

S
schaefersi Jessen, 2020

V
venusta Heer, 1867