Formica lemani

This is an abundant upland species in Europe not distinguished from Formica fusca until Bondroit (1917) and first clearly described by Yarrow (1954). It has similar habits to F. fusca but sometimes occurs in large multi-queened colonies in favourable sites such as stony banks. Colony founding is either by single queens or by nest fission. Alatae fly in July and August (Collingwood, 1979). In Japan this species nests in the soil under stones and near grass roots. Reproductive alates fly after mid August at Mt. Zao, Miyagi Prefecture (Japanese Ant Image Database).

Identification
Greyish to brownish black, legs paler. Short stout hairs present on promesonotum normally numerous but occasionally abraded or few. Underside of mid and hind femora normally with one or more hairs at mid length. Frons coarsely sculptured to that punctures readily seen under ordinary magnification. Length: 4.5-7.0 mm (Collingwood 1979).

Distribution
Mountains of Spain to Japan including Himalayas, Appenines to arctic Fennoscandia (Collingwood 1979).

The records of F. lemani from Korea are considered to be misidentications of Formica fusca by the following characteristics: 1. No more than 6 erect hairs (Radchenko, 2005) on the pronotum of worker. 2. Workers with less than 3 hairs in mid femur. However, according to Dlussky and Pisarski (1971), there are some variable individuals without any setae on the pronotum, which requires close examination (Dong & Kim, 2019).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Finland, France, Georgia, Germany, Greece, Iberian Peninsula, Italy, Japan, Kyrgyzstan, Mongolia, Montenegro, Netherlands, Norway, Poland, Republic of Korea, Republic of Macedonia, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland.

Biology
Kvifte and Soule (2017) - Lasius flavus and Formica lemani were found in a plesiobiotic association in a heathland in western Norway. The colonies were found in chambers under rocks, with both larvae and pupae of both species present.

This is the first confirmed case of F. lemani in a plesiobiontic relationship with another ant species, providing further evidence for Collingwood’s (1979) claim that the habits of F. lemani are similar to Formica fusca – the most frequently recorded plesiobiont in the Palearctic region (Kaniszai et al., 2013). Workers of Formica lemani and Lasius flavus differ markedly in size and foraging behaviour. Whereas F. lemani is a free-living and active predacious, aphidicolous and nectarivorous species, L. flavus is mostly subterranean and feeds on smaller arthropods and honeydew from root feeding aphids (Collingwood, 1979; Douwes et al., 2012). The resources exploited by each species thus show little overlap, permitting coexistence without competition. This follows the general pattern outlined for plesiobiontic relationships by Kanizsai et al. (2013). Colony sizes of the two species are listed in the literature as a few hundred to a few thousand for F. lemani and up to 100 000 workers for L. flavus (Douwes et al., 2012).

This species is a host for the temporary parasites, and.

Nomenclature

 *  lemani. Formica lemani Bondroit, 1917b: 186 (w.q.) FRANCE. Bondroit, 1920a: 145 (m.). Combination in F. (Serviformica): Emery, 1925b: 247. Subspecies of fusca: Müller, 1923: 139; Santschi, 1925g: 352; Novak & Sadil, 1941: 107. Revived status as species: Yarrow, 1954a: 231. Senior synonym of borealis: Dlussky, 1965a: 35. See also: Bernard, 1967: 295; Dlussky, 1967a: 60; Dlussky & Pisarski, 1971: 150; Kutter, 1977c: 251; Kupyanskaya, 1990: 184; Atanassov & Dlussky, 1992: 261; Seppä, et al. 2011: 31.
 * borealis. Formica fusca subsp. borealis Vashkevich, 1924b: 147 (w.) RUSSIA. Junior synonym of lemani: Dlussky, 1965a: 35.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Arnan X., M. Gracia, L. Comas, and J. Retana. 2009. Forest management conditioning ground ant community structure and composition in temperate conifer forests in the Pyrenees Mountains. Forest Ecology and Management 258(2): 5159.
 * ArtDatabanken Bugs (via GBIG)
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
 * Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Bernadou A., G. Latil, V. Fourcassié, and X. Espadaler. 2006. Etude des communautés de fourmis d'une vallée andorrane. Union International pour l'Etude des Insectes Sociaux, Colloque annuel de la section francaise, 4pp.
 * Bernadou A., X. Espadaler, A. Le Goff, and V. Fourcassie. 2015. Ant community organization along elevational gradients in a temperate ecosystem. Insect. Soc. 62:5971
 * Bernadou, A., G. Latil, V. Fourcassié, and X. Espadaler. "Les formigues de la Vall del Madriu-Perafita-Claror : diversitat i distribució." Hàbitats, 13 (2006): 10-21.
 * Bernard F. 1960. Fourmis récoltées en Corse par J. Bonfils (1957). Compte Rendu Sommaire des Séances de la Société de Biogéographie 36: 108-114.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1967. Recherches sur les fourmis des Monts-Dore. Annales de la Station biologique de Besse-en-Chandesse : 1-11.
 * Bernard F. 1973. Etudes ecologiques sur les fourmis de Breuil-Cervinia (Val d"Aoste). Revue Ecol Biol Sol, 102: 237-269.
 * Bernard, F.. "Notes sur les fourmis de France. II. Peuplement des montagnes méridionales." Annales de la Société Entomologique de France 115 (1950): 1-36.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bezdecka P., and K. Bezdeckova. 2012. Updated list of the ants of the Czech Republic. Pp 7-12. Bezd??ka P. & Bezd??ková K. (eds) 2012: Blanok?ídlí v ?eských zemích a na Slovensku 8, Chaloupky, 1.-3. ?ervna 2012, sborník abstrakt? z konference.  MVJ Jihlava, 37 pp.
 * Blacker N. C. 1989. The ants (Hymenoptera, Formicidae) of the Gower Peninsula, West Glamorgan, South Wales. Entomologist's Record and Journal of Variation 101: 261-266.
 * Blacker N.C. 2007. Ants (Hym., Formicidae) in East Anglia-Additional Records from . Entomologist's Monthly Magazine 143: 69-90
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., and W. Dekoninck. 2006. First record of the permanent social parasitic, slavemaking ant Harpagoxenus sublaevis (Nylander, 1852) for Belgium (Hymenoptera, Formicidae). Bulletin S.R.B.E/K.B.V.E 142: 21-24.
 * Bondroit J. 1917. Diagnoses de trois nouveaux Formica d'Europe (Hym.). Bull. Soc. Entomol. Fr. 1917: 186-187.
 * Bondroit, J.. "Diagnoses de trois nouveaux Formica d'Europe (Hym.)." Bulletin de la Société Entomologique de France (1917): 186-187.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Boven J. van 1949. Notes sur la faune des Hautes-Fagnes en Belgique. Bulletin et Annales de la Société Entomologique de Belgique 85: 135-143.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Cherix D., and S. Higashi. 1979. Distribution verticale des fourmis dans le Jura vaudois et recensement prelimaire des bourdons (Hymenoptera, Formicidae et Apidae). Bull. Soc. Vaud. Sc. Nat. 356(74): 315-324.
 * Choi B.-M. 1987. Taxonomic study on ants (Formicidae) in Korea (1). On the genus Monomorium. Journal of the Institute of Science Education (Cheongju National Teachers' College) 11:17-30.
 * Choi B.M. 1986. Studies on the distribution of ants (Formicidae) in Korea. Journal of Chongju National Teacher College 23: 317-386.
 * Choi B.M., Bang, J.R. 1992. Studies on the distribution of ants (Formicidae) in Korea (9). Ant fauna in Mt. Togyusan. Korean Journal of Applied Entomology 31:101-112.
 * Choi B.M., K. Ogata, and M. Terayama. 1993. Comparative studies of ant faunas of Korea and Japan. 1. Faunal comparison  among islands of Southern Korean and northern Kyushu, Japan. Bull. Biogeogr. Soc. Japan 48(1): 37-49.
 * Choi B.M., Kim, C.H., Bang, J.R. 1993. Studies on the distribution of ants (Formicidae) in Korea (13). A checklist of ants from each province (Do), with taxonomic notes. Cheongju Sabom Taehakkyo Nonmunjip (Journal of Cheongju National University of Education) 30: 331-380.
 * Choi B.M.; Bang, J.R. 1992. Studies on the distribution of ants (Formicidae) in Korea (10). Ant distribution in Gangweon Do. Journal of the Institute of Science Education (Cheongju National Teachers' College) 14:12-30.
 * Collingwood C. A. 1955. Ants in S.W. Scotland. Entomologist's Record and Journal of Variation 67: 11-12.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1961. Ants in the Scottish Highlands. Scotish Naturalist 70: 12-21.
 * Collingwood C. A. 1962. Some ants (Hym. Formicidae) from north-east Asia. Entomologisk Tidskrift 83: 215-230.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood C. A. 1976. Ants (Hymenoptera: Formicidae) from North Korea. Annales Historico-Naturales Musei Nationalis Hungarici 68:
 * Collingwood C. A., and J. E. Satchell. 1956. The ants of the South Lake District. Journal of the Society for British Entomology 5: 159-164.
 * Collingwood C. A., and J. Hughes. 1987. Ant species in Yorkshire, England. Naturalist (Leeds) 112: 95-101.
 * Collingwood C.A. 1955. Ants in S.W. Scotland. Entomol.Rec. 67: 11-12
 * Collingwood C.A. 1959. Ants in the Scottish Highlands. The Scottish Naturalist. 70: 12-21
 * Collingwood C.A. 1961. Ants in Finland. Entomol. Rec. 73: 190-195
 * Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
 * Collingwood C.A. and Satchell J.E. 1956. The Ants of the South Lake District. Journal of the Society for British Entomology. 5: 159-164
 * Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
 * Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
 * Collingwood, C. A. 1974. A revised list of Norwegian ants (Hymenoptera: Formicidae). Norsk Entomologisk Tidsskrift 21: 31-35.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Collingwood, C. A.. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Collingwood, C.A. 1958. A survey of Irish Formicidae. Proceedings of the Royal Irish Academy 59B:213-219
 * Csősz S. and Markó, B. 2005. European ant species (Hymenoptera: Formicidae) in the ant collection of the Natural History Museum of Sibiu (Hermannstadt/Nagyszeben), Romania II. Subfamily Formicinae. Annales Historico-Naturales Musei Nationalis Hungarici 97: 225-240.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Dlussky G. M., and B. Pisarski. 1970. Formicidae aus der Mongolei. Ergebnisse der Mongolisch-Deutschen Biologischen Expeditionen seit 1962, Nr. 46. Mitteilungen aus dem Zoologischen Museum in Berlin 46: 85-90.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Espadaler X., X. Roig, K. Gómez, and F. García. 2011. Formigues de les Planes de Son i mata de València (Hymenoptera, Formicidae) Treballs de la Institució Catalana d'Història Natural 16: 609-627.
 * Espadaler, X., and D. Cuesta. "Teleutomyrmex schneideri Kutter, 1950 en España (Hymenoptera, Formicidae)." Graellsia 62 (2) (2006): 261-262.
 * Espadaler, X.. "Contribución al conocimiento de los formícidos (Hymenoptera, Formicidae) del Pirineo catalán." Tesis Universida (1979): 285 pp.
 * Fiedler, K., F. Kuhlmann, B. C. Schlick-Steiner, F. M. Steiner and G. Gebauer. 2007. Stable N-isotope signatures of central European ants  assessing positions in a trophic gradient. Insectes Sociaux 54(4):393-402.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Formidabel Database
 * Fowles, A.P. 1996. A provisional checklist of the invertebrates recorded from Wales. 2. Aculeate wasps, bees and ants (Hymenoptera: Aculeata). Countryside Council for Wales
 * Galle L. 1997. Contribution to the ant fauna of Slovenia with special reference to the submediterranean and eudinaric regions. Annals for Istrian and Mediterranean studies 11: 209-214.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gardner, M.G., K. Schonrogge, G.W. Elmes and J.A. Thomas. 2007. Increased Genetic Diversity as a Defence against Parasites Is Undermined by Social Parasites: Microdon mutabilis Hoverflies Infesting Formica lemani Ant Colonies. Proceedings: Biological Sciences 274(1606):103-110
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Gibb H., and T. Johansson. 2011. Field tests of interspecific competition in ant assemblages: revisiting the dominant red wood ants. Journal of Animal Ecology 80: 548-557.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Glaser, F., M. J. Lush, and B. Seifert. "Rediscovered after 140 years at two localities: Myrmica myrmicoxena Forel, 1895 (Hymenoptera: Formicidae)." Myrmecological News 14 (2010): 107-111.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Hayashida K. 1971. Vertical distribution of ants in the southern part of the Hidaka mountains. [In Japanese.]. Memoirs of the National Science Museum (Tokyo) 4:29-38.
 * Hayashida K. 1972. Ecological survey on ants in Nakagawa Experiment Forest of Hokkaido University. Res. Bull. Exper. Forests, Coll. Agr., Hokkaido Univ. 29: 25-36.
 * Hågvar S. 2005. Altitudinal zonation of ants (Formicidae) in a steep fjord landscape in Sogndal, Western Norway. Norw. J. Entomol. 52: 3-12.
 * Ichinose K. 1990. The Ant Fauna of the Tomakomai Experiment Forest, Hokkaido University (Hymenoptera: Formicidae) with Notes on the Nuptial Season. Research Bulletins of College Experiment Forests 47(1) 137-144.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Kim B.J. 1996. Synonymic list and distribution of Formicidae (Hymenoptera) in Korea. Entomological Research Bulletin Supplement 169-196.
 * Kim B.J. and C.W; Kim. 1983. A systematic revision of the genus Formica in Korea on the basis of external fine features (Hym.: Formicidae). Ent. Res. Bulletin 9: 57-67.
 * Kim et al. 1993. Systematic study of ants from Chejudo Province. Koran Journal of Entomology 23(3): 117-141.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Krugova T. M. 2017. Ant's population in the chern taiga of Tigireksky mountain range (North-Western Altai). Proceedings of the Tigirek Reserve 9: 68-77.
 * Kupianskaya A. N., Lelej, A.S., and Urbain, B. K. 2000. The Ants (Hymenoptera: Formicidae) of the Kuril Islands. Far Eastern Entomologist. 92:1-21.
 * Kupianskaya A. N., and A. S. Lelej. 2000. The ants (Hymenoptera, Formicidae) collected in the Habomai and Shikotan (Kuril Islands) in 1998.  Far Eastern entomologist 92: 22-24.
 * Kupianskaya, A. N., Lelej, A.S., and Urbain, B. K. 2000. The Ants (Hymenoptera: Formicidae) of the Kuril Islands. Far Eastern Entomologist. 92:1-21.
 * Kupianskaya, A.N. and A.S. Lelej. 2000. The ants (Hymenoptera, Formicidae) collected in the Habomai and Shikotan (Kuril Islands) in 1998. Far Eastern Entomologist 92:22-24
 * Kupianskaya, A.N., A.S. Lelej and B.K. Urbain. 2000. The ants (Hymenoptera: Formicidae) of the Kuril Islands. Far Eastern Entomologist 92:1-21
 * Kvamme T. 1982. Atlas of the Formicidae of Norway (Hymenoptera: Aculeata). Insecta Norvegiae 2: 1-56.
 * Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur  Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Le Moli F., and A. Zaccone. 1995. Ricerche sulla mirmecofauna del Cansiglio (Prealpi Carniche). Soc. Ven. Sc. Nat. 20: 33-52.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Lebas C., and C. Galkowski. 2016. Myrmica hirsuta Elmes, 1978, a new species from France (Hymenoptera, Formicidae). Bull. Soc. Linn. Bordeaux 151, 44(2/3): 239-244.
 * Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
 * Liu X. 2012. Taxonomy, diversity and spatial distribution characters of the ant family Formicidae (Insecta: Hymenoptera) in southeastern Tibet. PhD Thesis 139 pages
 * Liu X., Z. H. Xu, X. Y. Zhou, N. N. Yu, and C. L. Zhang. 2011. Ant communities of West slope of Mount Demula and Bomi Valley in Southeastern Tibet. Forest Research 24(4): 458-463.
 * Liu X., Z. Xu, C. Zhang, N. Yu, and G. Xu. 2012. Distribution patterns of ants from West slope of Mount Demula and Bomi Valley in Southeastern Tibet. Journal of Northwest Forestry University 27(4): 77-82.
 * Liu X., Z. Xu, X. Zhou, N. Yu, and C. Zhang. 2011. Ant Communities of West Slope of Mount Demula and Bomi Valley in Southeastern Tibet. Forest Research 24(4): 458-463.
 * Liu X., Z.-H. Xu, C.-L. Zhang, N.-N., and G.-L. 2011. Distribution patterns of ants from West Slope of Mount Demula and Bomi Valley in southeastern Tibet. Journal of Northwest Forestry University 27(4): 77-82.
 * Malozemova L. A. 1972. Ants of steppe forests, their distribution by habitats, and perspectives of their utilization for protection of forests (north Kazakhstan). [In Russian.]. Zoologicheskii Zhurnal 51: 57-68.
 * Marko B., K. Kiss, and L. Galle. 2004. Mosaic structure of ant communities (Hymenoptera: Formicidae) in Eastern Carpathian marshes: regional versus local scales. Acta Zoologica Academiae Scientiarum Hungaricae 50(2): 7795.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Mizutani A. 1979. A myrmecofaunal survey at Hiyama Experiment Forest, Hokkaido University. Research Bulletin of the College Experiment Forests, Hokkaido University 36:509-516.
 * Müller, G.. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Núñez-Pérez, E.. "Bases para el desarrollo del control integrado de los pulgones (Hom. Aphididae) de los cultivos de la provincia de León." Universidad de León, Secretariado de Publicaciones Tesis doct (1992): 7 páginas y 4 microfichas. León.
 * Paik W.H. 1984. A checklist of Formicidae (Hymenoptera) of Korea. Korean J. Plant Prot. 23(3): 193-195.
 * Palanca, A., A. Urbieta, C. Castán, and X. Espadaler. "Ants in the supraforestal pastures of the Tendeñera massif." Écologie des milieux montagnards et de haute altitude. Documents d'Ecologie Pyrénéenne III-IV (1984): 223-224.
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Park S.J., and B.J. Kim. 2002. Faunal comparison of ants among Cheongsando and other islands of South Sea in Korea. Korean Journal of Entomology 32(1): 7-12.
 * Park, Seong, Joon and Byung, and Kim, Jin. 2002. Faunal Comparison of Ants among Cheongsando and Other Islands of South Sea in Korea. Korean Jornal of Entomology. 32(1):7-12.
 * Pascual, M. R.. "Estudio taxonómico y ecológico de los Formícidos de las Sierras de Alfacar, La Yedra, Huétor y Harana." Tesis Doctoral Universida (1986): 264 pp.
 * Paukkunen J., and M. V. Kozlov. 2015. Stinging wasps, ants and bees (Hy menoptera: Aculeata) of the Murmansk region, Northwest Russia. — Entomol. Fennica. 26: 53–73.
 * Petal J. 1994. Reaction of ant communities to degradations of forest habitats in the Karkonosze Mountains. Polish Academy of Sciences, Museum and Institute of Zoology, Memorabilia Zoologica 48: 171-179.
 * Petal J. M. 1967. Contribution a la connaissance des fourmis (Formicidae, Hymenoptera) de la region de Lublin. VII. Associations des fourmis des milieux de tourbieres, de forets et de dunes aux environs de Libiszow (dist. De Parczew). Annales Universitatis Mariae Curie-Sklodowska Lublin-Polonia 22(9): 117-130.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Pisarski B. 1969. Fourmis (Hymenoptera: Formicidae) de la Mongolie. Fragmenta Faunistica (Warsaw). 15: 221-236.
 * Punttila P., Y. Haila, and H. Tukia. 1996. Ant communities in taiga clearcuts: habitat effects and species interactions. Ecography 19: 16-28.
 * Punttila P., and Y. Haila. 1996. Colonisation of a burned forest by ants in the southern Finnish Boreal forest. Silva Fennica 30(4): 421-435.
 * Pusvaskyte O. 1979. Myrmecofauna of the Lituanian SSR. Acta Entomologica Lituanica 4: 99-105.
 * Radchenko, A. 2005. Monographic revision of the ants (Hymenoptera, Formicidae) of North Korea. Annales Zoologici 55(2): 127-221.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Rigato F., and R. Sciaky. 1991. The myrmecofauna of the Gesso Valley (Maritime Alps) (Hymenoptera Formicidae). Ethology Ecology and Evolution Special Issue 1: 87-89.
 * Salata S. 2014. Ants (Hymenoptera: Formicidae) of the National Park of the Stołowe Mts. Przyroda Sudetow 17: 161-172.
 * Salata S. 2014. Mrówki (Hymenoptera: Formicidae) Parku Narodowego Gór Sto?owych. PRZYRODA SUDETÓW t. 17: 161-172.
 * Santschi F. 1925. Fourmis d'Espagne et autres espéces paléartiques EOS (Revista española de entomología) 1: 339-360.
 * Santschi, F.. "Fourmis d'Espagne et autres espéces paléartiques." EOS (Revista española de entomología) 1 (1925): 339-360.
 * Sato T., N. Tsurusaki, K. Hamaguchi, and K. Kinomura. 2010. Ant fauna of Tottori prefecture, Honshu, Japan. Bulletin of the Tottori Prefectural Museum 47: 27-44.
 * Schar S., G Talavera, X. Espadaler, J. D. Rana, A. A. Andersen, S. P. Cover, and R. Vila. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 00: 1-12.
 * Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
 * Seifert B. 2007. Die Ameisen Mittel- und Nordeuropas. Tauer: lutra Verlags- und Vertriebsgesellschaft, 368 pp.
 * Seppä P., H. Helanterä, A. Chernenko, K. Trontti, P. Punttila, L. Sundström. 2009. Population genetics of the black ant Formica lemani (Hymenoptera: Formicidae). Biological Journal of the Linnean Society 97:247-258.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Soulie J. 1962. Fourmis des Hautes-Pyrenees. Bull. Soc. Hist. Nat. Toulouse 97: 35-37.
 * Stärcke A. 1942. Ants, collected by Dr. C. F. Engelhard at Stockmarknes (Island Hadsekoy, Vesterale Archipelago circa 68°50' Lat. N. Norway) and between 64° and 66° Lat. N. on the western coast of Norway, July 1932. Entomologische Berichten (Amsterdam) 11: 21-23.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Tausan I. 2010. Notes on the ant fauna (Hymenoptera, Formicidae) of the Rodna Mountains National Park and it's surroundings (Transylvania-Maramures, Romania). Transylv. Rev. Syst. Ecol. Res. 9: 159-166.
 * Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
 * Terayama M. 1977. Checklist of the known ants of Saitama Prefecture. Insects and nature 12(4): 26-27
 * Terayama M. 1992. Structure of ant communities in East Asia. A. Regional differences and species richness. Bulletin of the Bio-geographical Society of Japan 47: 1-31.
 * Terayama M., Choi, B.M., Kim, C.H. 1992. A check list of ants from Korea, with taxonomic notes. Bulletin of the Toho Gakuen 7:19-54.
 * Terayama M., K. Murata, and N. Minakawa. 1998. Ants collected by the members of the International Kuril Islands Project, 1995. Ari 22: 10-12.
 * Terayama M., K. Ogata, and B.M. Choi. 1994. Distribution records of ants in 47 prefectures of Japan. Ari (report of the Myrmecologists Society of Japan) 18: 5-17.
 * Terayama M., S. Kubota, and K. Eguchi. 2014. Encyclopedia of Japanese ants. Asakura Shoten: Tokyo, 278 pp.
 * Terayama. M. 2004. Geological and ecological distribution of Japanese ants communities. (translated from Japanese) Reports of the Saitama Prefecture Animal Research Association. 48:24
 * Terayama. M. 2004. Geological and ecological distribution of Japanese ants communities. (translated from Japanese) Reports of the Saitama Prefecture Animal Research Association. 48:32
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Tizado Morales, E. J.. Estudio comparado de la fauna y la biología de pulgones (Hom.), afidíinos (Hym.) y otros insectos acompañantes en dos áreas de la provincia de León In Secretariado de Publicaciones, Tesis doctoral en microficha, nº 67. León: Universidad de León, 1991.
 * Urbieta Balado A., A. Palanca Soler, C. Castan Lanaspa, and X. Espadaler. 1984. Ants in the supraforestal pastures of the Tendenera massif. Documents dEcologie Pyrénéenne III-IV: 223-224.
 * Vele A., J. Holisa, J. Tragnerova. 2015. Ant succession on burned areas in forested landscape: a case study from the Bohemian Switzerland National Forest. Zpravy Lesnickeho Vyzkumu 60(1): 47-52.
 * Vepsalainen K., H. Ikonene, and M. J. Koivula. 2008. The structure of ant assembalges in an urban area of Helsinki, southern Finland. Ann. Zool. Fennici 45: 109-127.
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vespalainen K., B. Pisarski, R. Kantorek, and K. J. Laine. 1984. Formicidae (Hymenoptera) of Inari Lapland. Kevo Notes 7: 115-116.
 * Wegnez P., D. Ignace, E. Lommelen, M. Hardy, J. Bogaert, and C. Nilsson. 2015. Redécouverte de Teleutomyrmex schneideriKutter, 1950 dans les Alpes françaises (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 52-57.
 * Wegnez P., and F. Mourey. 2016. Formica uralensis Ruzsky, 1895 une espèce encore présente en France mais pour combien de temps ? (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 152: 72-80.
 * Wiezik M. 2007. Ants (Hymenoptera: Formicidae) of mountain and alpine ecosystems at Southern part of Krá?ovoho?ské Tatry Mts. Naturae Tutela 11: 85-90.
 * Wiezik M. 2008. Vertical distribution of ants from alpine habitats at Nízke Tatry mts. Naturae Tutela 12: 61-67.
 * Wiezik M., A. Wiezikova, and J. Murin. 2008. Ant assemblages at horná orava peatbogs. Naturae Tutela 12: 55-60.
 * Wiezik M., M. Kozon, and A. Wiezikova. 2013. Ants of  selected  peat  bog  habitats  at  the  Horná Orava protected area. Naturae Tutela 17(1): 57-63.
 * Xu Z., J. Chu, C Zhang, and N. Yu. 2011. Ant species and distribution pattern in Gongbo Nature Reserve in Southeastern Tibet. Sichuan Journal of Zoology 30(1): 118-123.
 * Xu Z., J. Chu, C. Zhang, and N. Yu. 2011. Ant Species and D istribution Pattern in GongboNature Reserve in Southeastern T ibet. Sichuan Journal of Zoology 30(1): 118-123.
 * Yarrow I. H. H. 1954. The British ants allied to Formica fusca L. (Hym., Formicidae). Trans. Soc. Br. Entomol. 11: 229-244.
 * Yoshimura M. 1998. Ants from Islands in Hokkaido, Northern Japan (No. 1, Risiri Island). Rishiri Studies 17:33-38.
 * Yoshimura M. 1999. Ants in the island of Hokkaido (Part 2, Part Rebun) Ants from Islands in Hokkaido, Northern Japan (No. 2, Rebun-Island). Rishiri Studies 18: 49-54.
 * Yu N., Z. Xu, C. Zhang, J. Chu, B. Yang, and X. Liu. 2011. Distribution patterns of ant species from Mount Sejila, southeastern Tibet. Journal of Beijing Forestry University 33(5): 75-80.
 * Yu N., Z. Xu, C. Zhang, J. Chu, B. Yang, and X. Liu. 2011. Distribution patterns of ant species from mount Sejila, southeastern Tibet. Journal of Beijing Forestry University 33(5): 75-80
 * Yu N., Z. Xu, C. Zhang, X. Liu, and Q. Li. 2012. Ant Communities from East Slope of Mount Demola to Zayu Valley in Southeastern Tibet. Scientia Silvae Sinicae 48(10): 163-169.
 * Zhang C., Z. Xu, N. Yu, Q. He, and X. Liu. 2012. Distribution Patterns of Ant Species on East Slope of Mount Demola and Zayu Valley in Southeastern Tibet. Journal of Northeast Forestry University 40(3): 87-92
 * Zhigulskaya Z. A. 2009. The ants of the Chuya Depression and the Yustyd river basin in Southeastern Altai. Contemporary Problems of Ecology 2009 2(3): 210-215.
 * del Campo Gracia P., M. D. Martinez Ibanez, A. Tinaut, and S. Montagud Alario. 2014. Faunistic study of the Formicidae (Hymenoptera, Formicidae) from the Comunitat Valenciana (Spain). Boln. Asoc. esp. Ent. 38 (1-2): 33-65.