Tetramorium semilaeve

Tetramorium semilaeve prefers very warm and dry places. Most observed nests were located in the littoral zone on the flat, sandy areas covered with sparse vegetation or on stony pastures. The locality with the highest altitude in the examined material here is from Andalucia, Igualeja placed 720 m a.s.l. Sanetra et al. (1999) noted that in Italy most samples were collected below 1000 m a.s.l. and only occasionally above this altitude. All nests were located under stones, from small to very large size, and consisted of from several to several hundred workers. (Borowiec et al., 2015.)

Identification
Borowiec et al. (2015) - Tetramorium semilaeve belongs to the group of Mediterranean species with workers pale-coloured, from yellow to yellowish-brown but never dark brown or black, head sculpture partly reduced, at least with postocular area and sides of frons without distinct striation but with at least half of the surface of the anterior part of head striate, striae on top of head not diverging laterally, pronotum mostly striate, sides of mesosoma only microreticulate and microgranulate without carinae, short propodeal spines, petiole and postpetiole dorsally with polished area, and first abdominal tergite smooth and shiny, without microreticulation or striation; gyne with flat mesonotal plate, without a distinct striation or the striation is indistinct, short not extending behind half length of the plate, scutellum without striation or only on sides with short, indistinct striae, postpetiole not enlarged; male genitalia stout, in lateral view only slightly constricted before apex, ventral and dorsal margins of parameres shallowly incised, top of inner margin of paramere before apical denticle straight with dentiform plate distinctly clearly extending beyond the sharp edge of paramere (arrow indices this plate). The presence of dentiform plate on top of the inner margin of paramere is the best character distinguishing T. semilaeve from all other species related to this taxon.

Distribution
The concept of this taxon has change recently (Borowiec et al., 2015) and its distribution clarified. Previous occurrence records should be verified before being considered valid.

Distribution based on Regional Taxon Lists
Palaearctic Region: Azerbaijan, France, Iberian Peninsula, Italy, Malta, Portugal, Spain.

Predators
Pekár et al. (2018) - This ant is preyed upon by numerous spider species in the genus Zodarion (Araneae: Zodariidae). All members of this genus are specialized ant predators that exclusively prey on ants.

Nomenclature

 *  semilaeve. Tetramorium caespitum var. semilaeve André, 1883a: 286 (w.) Mediterranean region. Emery, 1891b: 2 (q.); Forel, 1902a: 148 (m.). Subspecies of caespitum: Forel, 1902a: 148; Forel, 1904b: 371; Emery, 1909d: 703; Emery, 1916b: 194; Santschi, 1921a: 111; Santschi, 1923a: 135; Menozzi, 1926b: 182; Emery, 1924a: 8. Status as species: Dalla Torre, 1893: 134; Bondroit, 1918: 109; Müller, 1923: 104; Santschi, 1927a: 54; Santschi, 1929e: 150; Finzi, 1930d: 315; Santschi, 1931a: 10; Bernard, 1967: 234; Arnol'di & Dlussky, 1978: 545. Senior synonym of tenuicornis: Radchenko, 1992b: 51; of siciliense: Sanetra, Güsten & Schulz, 1999: 327; of banyulense: Casevitz-Weulersse & Galkowski, 2009: 497. Current subspecies: atlante, galaticum, hoggarense, italicum, judas, jugurtha, kutteri. See also: Emery, 1925c: 179; López, 1991a: 36; López, 1991b: 76; López, et al. 1992: 169; Atanassov & Dlussky, 1992: 153; Radchenko, 1992b: 51.
 * tenuicornis. Tetramorium caespitum subsp. tenuicornis Emery, 1925c: 188, figs. B, D (w.m.) TRANSCASPIA. Junior synonym of semilaeve: Radchenko, 1992b: 51.
 * siciliense. Tetramorium semilaeve var. siciliense Santschi, 1927a: 56 (w.q.) ITALY (Sicily). Junior synonym of semilaeve: Sanetra, Güsten & Schulz, 1999: 327.
 * banyulense. Tetramorium banyulense Bernard, 1983: 98 (diagnosis in key) (w.m.) FRANCE. Junior synonym of semilaeve: Casevitz-Weulersse & Galkowski, 2009: 497.

Worker
Borowiec et al (2015) - (n=15): CL: 0.723 ± 0.034 (0.637-0.771); POC: 0.293 ± 0.021 (0.246-.324); CW: 0.693 ± 0.037 (0.606-0.749); FR: 0.253 ± 0.015 (0.234-0.279); FL: 0.262 ± 0.017 (0.235-0.291); SL: 0.534 ± 0.03 (0.503-0.626); OMD: 0.18 ± 0.021 (0.145-0.223); EL: 0.13 ± 0.01 (0.106-0.145); EH: 0.091 ± 0.006 (0.078-0.101); ML: 0.781 ± 0.043 (0.737-0.894); SPSP: 0.133 ± 0.015 (0.112-0.179); SPL: 0.099 ± 0.007 (0.089-0.112); PEL: 0.184 ± 0.01 (0.167-0.201); NOL: 0.144 ± 0.013 (0.128-0.168); PPL: 0.176 ± 0.009 (0.156- 0.19); PEH: 0.239 ± 0.018 (0.212-0.291); NOH: 0.158 ± 0.017 (0.14-0.218); PPH: 0.228 ± 0.02 (0.201-0.291); MW: 0.45 ± 0.027 (0.413-0.508); PEW: 0.22 ± 0.014 (0.201-0.246); PPW: 0.256 ± 0.018 (0.223-0.307); CS: 0.707 ± 0.036 (0.622-0.76); EYE: 0.155 ± 0.007 (0.143-0.166); CL/CW: 1.042 ± 0.013 (1.015-1.075); FR/CS: 0.358 ± 0.01 (0.345-0.378); FL/FR: 1.025 ± 0.02 (0.996-1.066); SL/CS: 0.756 ± 0.03 (0.732-0.841); MW/CS: 0.636 ± 0.018 (0.612-0.682); PEW/PPW: 0.861 ± 0.033 (0.801-0.918); NOH/NOL: 1.1 ± 0.113 (0.929-1.298); NOH/PEL: 0.86 ± 0.085 (0.819-1.085); NOL/PEL: 0.784 ± 0.045 (0.705-0.871); PEH/NOL: 1.66 ± 0.152 (1.399-1.922); PEW/PEH: 0.926 ± 0.049 (0.805-1.0); CS/PEW: 3.215 ± 0.089 (3.026-3.336); CS/PPW: 2.769 ± 0.154 (2.425-3.024); CW/MW: 1.541 ± 0.046 (1.435-1.602);

Small to medium size, CS: 0.707 [0.622-0.76]. In most specimens whole body pale yellowish-brown and appendages yellow, the palest specimens completely yellow, the darkest specimens yellowish-brown but never dark brown or black. Head nearly square CL/CW: 1.042 [1.015-1.075], with almost parallel sides, straight or slightly concave occipital margin and narrowly rounded occipital corners. Eyes small, EYE 0.155 [0.143-0.166]. Frons moderately wide, FR/CS 0.358 [0.345-0.378], frontal lobes as wide as frons, FL/FR: 1.025 [0.996-1.066]. Scape short, SL/CS 0.756 [0.732-0.841], without dorsal carina basally, surface smooth and shiny. Promesonotal dorsum slightly convex, metanotal groove shallow, but distinct. Propodeal teeth short, spiniform, apex of spine located approximately at 2/3 height of mesosoma. Dorsal surface of petiole flat to slightly convex, NOH/NOL 1.1 [0.929-1.298], petiole relatively high, PEH/ NOL 1.66 [1.399-1.922], postpetiole distinctly transverse. General appearance finely rugose, ground surface shiny. Head dorsum partly longitudinally rugose and shiny between rugae, rugae extend occipital margin of head, occiput mostly smooth and shiny, sides in anterior half longitudinally rugose and shiny between rugae. In most specimens between frontal rugose area and rugosities along ocular area on each side runs longitudinal band without rugosities but smooth area never exceeds 1/5 of the anterior surface of head; in extremely sculptured specimens almost whole frontal surface of head with long rugae with very small smooth patch between interrupted rugae and occiput with fine rugosities, dark coloured specimens usually have more distinct sculpture than pale coloured specimens but strongly sculptured and completely yellow specimens were also observed. Alitrunk dorsum rugose and microreticulate but never reticulate, only occasionally rugae on pronotum partly interrupted with indistinct microreticulation but pronotum never with smooth and shiny areas. Sides of pronotum and meso- and metapleuron usually coarsely microreticulate, sometimes reticulation tends to form transverse lines but surface never appears striate or rugose. Dorsum of petiolar node smooth and shiny with sides carinate, lateral surface microreticulate. Dorsum of postpetiole smooth and shiny, sides microreticulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with sparse setae, the longest on pronotum and the shortest on frons. Ventral surface of head with sparse short and 2-3 moderately long setae not forming a psammophore.

Queen
Borowiec et al (2015) - (n=11): CL: 0.999 ± 0.015 (0.983- 1.027); POC: 0.378 ± 0.019 (0.34-.413); CW: 1.09 ± 0.05 (0.978-1.161); FR: 0.397 ± 0.009 (0.38-0.412); FL: 0.385 ± 0.018 (0.357-0.413); SL: 0.715 ± 0.015 (0.693- 0.737); OMD: 0.223 ± 0.016 (0.212-0.257); EL: 0.274 ± 0.01 (0.257-0.291); EH: 0.218 ± 0.012 (0.193-0.235); ML: 1.699 ± 0.217 (1.053-1.813); SPSP: 0.267 ± 0.015 (0.24-0.291); SPL: 0.193 ± 0.01 (0.173-0.201); PEL: 0.298 ± 0.014 (0.279-0.324); NOL: 0.225 ± 0.022 (0.179-0.256); PPL: 0.301 ± 0.01 (0.285-0.313); PEH: 0.424 ± 0.011 (0.408-0.447); NOH: 0.283 ± 0.019 (0.257-0.313); PPH: 0.423 ± 0.017 (0.391-0.447); MW: 1.069 ± 0.04 (1.0-1.141); PEW: 0.393 ± 0.022 (0.348-0.419); PPW: 0.509 ± 0.019 (0.48-0.547); CS: 1.046 ± 0.032 (0.986-1.094); EYE: 0.235 ± 0.012 (0.219-0.251); CL/CW: 0.917 ± 0.038 (0.882-0.985); FR/CS: 0.377 ± 0.011(0.36-0.393); FL/FR: 0.978 ± 0.035 (0.913-1.027); SL/CS: 0.686 ± 0.019 (0.654- 0.717); MW/CS: 1.024 ± 0.043 (0.94-1.088); PEW/PPW: 0.773 ± 0.035 (0.692- 0.821); NOH/NOL: 1.266 ± 0.136 (1.094-1.587); NOH/PEL: 0.949 ± 0.053 (0.883-1.036); NOL/PEL: 0.755 ± 0.059 (0.63-0.847); PEH/NOL: 1.9 ± 0.175 (1.746-2.374); PEW/PEH: 0.931 ± 0.048 (0.818-0.971); CS/PEW: 2.62 ± 0.123 (2.475-2.879); CS/PPW: 2.052 ± 0.085 (1.947-2.185); CW/MW: 1.02 ± 0.057 (0.911-1.129); WAIST: 0.871 ± 0.035 (0.817-0.918).

Moderate size, CS 1.046 [0.986-1.094]. Whole body brown, appendages yellowish. Head wider than long, CL/CW 0.917 [0.882-0.985], with straight subparallel sides, shallowly emarginate occipital margin and narrowly rounded occipital corners. Frons moderately wide, FR/CS 0.377 [0.36-0.393], frontal lobes as wide as frons, FL/ FR 0.978 [0.913-1.027]. Scape short, SL/CS 0.686 [0.654-0.717], without dorsal carina basally, smooth and shiny. Head as wide as scutum, MW/CS 1.024 [0.94-1.088]. Propodeal teeth very short. Dorsal crest of petiolar node in frontal view straight. Petiolar node dorsum steeply rounded backward. Petiole and postpetiole relatively narrow, WAIST 0.871 [0.817-0.918]. General appearance smooth and shiny. Head dorsum, occiput and sides rugulose, ground surface shiny or indistinctly microreticulate. Frons longitudinally rugulose. Mesosoma flat, pronotum visible from above. Scutum and scutellum punctate along sides, in most specimens smooth and shiny, only occasionally scutellum in corners with very short rugae and scutum in basal part with thin longitudinal rugae never extending to half length of scutum. Sides of pronotum ruguloso-reticulate and feebly microreticulate, anepisternum mostly smooth and shiny only in dorsal part with fine longitudinal rugae, katepisternum smooth and shiny. Dorsum of petiolar node distinctly reticulate and smooth, posterior surface granulate and more or less transversely rugose. Postpetiole distinctly transverse, sides narrowly rounded to subangulate, dorsum of postpetiole smooth, sides granulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with short, sparse setae. Ventral surface of head with several short setae, as long as to 1.5 times longer than frontal setae, arising posteriorly to buccal cavity.

Male
Borowiec et al (2015) - (n=10): CL: 0.667 ± 0.018 (0.637- 0.693); POC: 0.279 ± 0.019 (0.263-.313); CW: 0.807 ± 0.024 (0.771-0.827); FR: 0.238 ± 0.012 (0.221-0.257); FL: 0.292 ± 0.013 (0.277-0.307); SL: 0.333 ± 0.016 (0.307- 0.358); OMD: 0.068 ± 0.016 (0.056-0.089); EL: 0.312 ± 0.01 (0.302-0.324); EH: 0.251 ± 0.011 (0.235-0.263); ML: 1.716 ± 0.069 (1.626-1.785); SPSP: 0.226 ± 0.014 (0.212-0.246); SPL: 0.205 ± 0.013 (0.184-0.223); PEL: 0.275 ± 0.017 (0.257-0.302); NOL: 0.176 ± 0.018 (0.156-0.201); PPL: 0.287 ± 0.017 (0.263-0.302); PEH: 0.286 ± 0.022 (0.257-0.313); NOH: 0.163 ± 0.01 (0.156-0.179); PPH: 0.418 ± 0.015 (0.391- 0.425); MW: 1.125 ± 0.056 (1.056-1.223); PEW: 0.337 ± 0.023 (0.307-0.368); PPW: 0.478 ± 0.022 (0.453-0.503); CS: 0.737 ± 0.019 (0.704-0.755); EYE: 0.382 ± 0.009 (0.37-0.395); CL/CW: 0.826 ± 0.023 (0.797-0.857); FR/CS: 0.323 ± 0.012 (0.314- 0.345); FL/FR: 1.228 ± 0.06 (1.132-1.306); SL/CS: 0.452 ± 0.015 (0.436-0.478); MW/CS: 1.528 ± 0.093 (1.411-1.684); PEW/PPW: 0.705 ± 0.04 (0.668-0.767); NOH/NOL: 0.928 ± 0.076 (0.812-1.0); NOH/PEL: 0.595 ± 0.059 (0.517-0.668); NOL/PEL: 0.643 ± 0.069 (0.559-0.75); PEH/NOL: 1.625 ± 0.114 (1.502-1.788); PEW/PEH: 1.182 ± 0.061 (1.1-1.284); CS/PEW: 2.216 ± 0.165 (2.019-2.438); CS/ PPW: 1.558 ± 0.085 (1.443-1.652); CW/MW: 0.719 ± 0.045 (0.639-0.783)

Whole body dark brown, appendages yellowish. Head with feebly convex sides, slightly rounded occipital margin and widely rounded occipital corners. Scutum distinctly wider than head. Propodeum rounded in profile or with only indistinct angulation in position of propodeal teeth. Dorsal crest of petiolar node with obtuse transversal edge, slightly emarginated in frontal view. Head distinctly granulate, dull, scutum in anterior part and laterally smooth and shiny, rest microreticulate. Scutellum ostly smooth and shiny, at corners microreticulate and in anterior part with thin transverse rugosities. Sides of alitrunk smooth and shiny. Dorsum of petiolar node microgranulate and microreticulate, dull, postpetiole mostly microreticulate only top partly smooth and shiny. First gastral tergite smooth and shiny. Male genitalia stout, in lateral view only slightly constricted before apex with obtuse inner angle, top shortly and sparsely pubescent, ventral and dorsal margins of parameres shallowly incised, top of inner margin of paramere before apical denticle straight with dentiform plate distinctly clearly extending beyond the sharp edge of paramere (arrow indices this plate).

References based on Global Ant Biodiversity Informatics

 * Abril S., C. Gómez. 2012. Lista actualizada de las especies de hormigas de Menorca (Islas Baleares, España) y primera cita de Monomorium andrei Saunders, 1890 (Hymenoptera, Formicidae) de la isla Boletín de la Sociedad Entomológica Aragonesa 50: 403-407.
 * Acosta, F. J., M. D. Martínez Ibáñez, and M. A. Morales. "Contribución al conocimiento de la mirmecofauna del encinar peninsular. I." Boletín de la Asociación Española de Entomología 6 (2) (1983): 379-391.
 * Acosta, F. J., and M. D. Martínez Ibáñez. "Consideraciones sobre la dulosis en el género Strongylonathus Mayr, 1853." Boletín de la Asociación Española de Entomología 6(1) (1982): 121-124.
 * Aktaç, N. "Studies on the myrmecofauna of Turkey I. Ants of Siirt, Bodrum and Trabzon." Istanbul Universitesi Fen Fakultesi Mecmuasi. Seri B 41 (1977): 115-135.
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2010. Iberomyrmex 3: 32-33.
 * Asociacion Iberica de Mirmecologia. 2011. List of species collected during the Taxomara Lisboa 2011. Iberomyrmex 3: 30-31.
 * Asociacion Iberica de Mirmecologia. 2015. List of species collected during the Taxomara Lisboa 2015 Iberomyrmex 7: 44-45.
 * Asociacion Iberica de Mirmecologia. 2016. List of species collected during the Taxomara Murcia 2016. Iberomyrmex 8: 48-49.
 * Azcarate F. M., and B. Peco. 2012. Abandonment of grazing in a mediterranean grassland area: consequences for ant assemblages. Insect Conservation and Diversity 5: 279288.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. IV. La fauna mirmecologica delle isole Maltesi ed il suo significato ecologico e biogeografico. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 77: 408-559.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. V. Aspetti ecologici della Riviera del M. Cònero. Boll. Zool. 35: 39-76.
 * Baroni Urbani C. 1974. Studi sulla mirmecofauna d'Italia. XII. Le Isole Pontine. Fragm. Entomol. 9: 225-252.
 * Baroni Urbani C. 1976. Le formiche dell'arcipelago della Galita (Tunisia). Redia 59: 207-223.
 * Baroni Urbani, C. "Formiche dell'Italia appenninica (Studi sulla mirmecofauna d'Italia, III)." Memorie del Museo Civico di Storia Naturale di Verona 12 (1964): 149-172.
 * Baroni Urbani, C. "Studi sulla mirmecofauna d'Italia. II. Formiche di Sicilia." Atti dell'Accademia Gioenia di Scienze Naturali in Catania (6) 16 (1964): 25-66.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Bernard F. 1958. Les fourmis des iles Pelagie, comparaison avec d'autres faunes insulaires. Esttrato dalla Rivista di Biologia Coloniale XVI: 67-79.
 * Bernard F. 1959. Les fourmis de l'île de Port-Cros. Contribution à l'écologie des anciennes forêts méditerranéennes. Vie Milieu 9: 340-360.
 * Bernard F. 1960. Fourmis récoltées en Corse par J. Bonfils (1957). Compte Rendu Sommaire des Séances de la Société de Biogéographie 36: 108-114.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1973. Tendances calcicoles ou silicicoles chez les fourmis méditerranéennes. Pp. 16-21 in: International Union for the Study of Social Insects. Congress 1973. Proceedings IUSSI VIIth International Congress, London, 10-15 September, 1973. Southampton: University of Southampton, vi + 418 pp.
 * Bernard F. 1975. Rapports entre fourmis et vegetation pres des Gorges du Verdon. Annales du Muséum d'Histoire Naturelle de Nice 2: 57-79.
 * Bernard F. 1978. Fourmis et milieu dans le massif des Maures. Vie et Milieu. Série C. Biologie Terrestre 27: 83-118.
 * Bernard, F. "Remarques sur le peuplement des Baléares en fourmis." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 47 (1956): 254-266.
 * Berville L., M. Renucci, and E. Provost. 2012. Mise en place de protocoles de contrôle de la fourmi dArgentine (Linepithema humile) sur les îles de Port-Cros et de Porquerolles (Var, France). Sci. Rep. Port-Cros natl. Park, 26: 91-108.
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Bonaric J. C. 1971. Contribution a l'etude systematique et ecologique des formicides du Bas-Languedoc. PhD thesis Universite des sciences et techniques du Languedoc, 175 pages.
 * Bonaric J. C. 1971. Étude systématique et ecologique des fourmis de lHérault (suite). Ann. Soc. Hortic. Hist. Nat. Hérault 111: 119-126.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., C. Galkowski, and S. Salata. 2015. What is Tetramorium semilaeve André, 1883? (Hymenoptera, Formicidae). ZooKeys 512:39-62.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Borowiec L., and S. Salata. 2017. New records of ants (Hymenoptera: Formicidae) from southern Portugal. Acta Entomologica Silesiana 25(3): 1-10.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Cagniant H. 1997. Le genre Tetramorium au Maroc (Hymenoptera: Formicidae): clé et catalogue des espèces. Ann. Soc. Entomol. Fr. (N.S.) 33: 89-100.
 * Cagniant H., and C. Galkowski. 2013. Aphaenogaster koniari n. sp. du Maroc (Hymenoptera, Formicidae). Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 175-185.
 * Cagniant, H. 2006. Liste actualisée des fourmis du Maroc. Myrmecologische Nachrichten 8: 193-200.
 * Cagniant, H. "Aphaenogaster nadigi Santschi, bona especies (n. status)." EOS (Revista española de entomología) LXII (1987): 31-43.
 * Cagniant, H. "Contribution à la connaissance des fourmis marocaines: description des sexués et compléments à la définition de l'espèce Aphaenogaster theryi Santschi 1923 (Hyménoptères, Formicoidea, Myrmicidae)." Bulletin de la Société d'Histoire Naturelle de Toulouse 122 (1986): 139-144.
 * Cagniant, H. "Liste actualisee des fourmis du Maroc (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 193-200.
 * Cagniant, H. "Liste préliminaire de fourmis forestières d'Algérie. Résultats obtenus de 1963 à 1966." Bulletin de la Societe d'Histoire Naturelle de Toulouse 104 (1968): 138-147.
 * Cagniant, H. "Étude de quelques fourmis marocaines. Statistique provisoire des Formicidae du Maroc." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 53 (1964): 83-118.
 * Cagniant, H. Les peuplements de fourmis des forêts algériennes: écologie, biocénotique, essai biologique. Universite de Toulouse, 1973.
 * Cammell, M. E., M. J. Way, and M. R. Paiva. "Diversity and structure of ant communities associated with oak, pine, eucalyptus and arable habitats in Portugal." Insectes Sociaux 43 (1996): 37-46.
 * Carpintero S., J. Reyes-Lopes, and G. M. Luque. 2011. Ant community structure under Retama sphaerocarpa shrubs in a semi-arid environment. Entomological Science 14: 147153.
 * Carpintero, S., A. Tinaut, J. Reyes, and L. Arias. "Estudio faunístico de los formícidos (Hymenoptera: Formicidae) del Parque Nacional de Doñana." Boletín de la Asociación Española de Entomología 25 (1-2) (2001): 133-152.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J. 1992. La myrmecofaune de la reserve naturelle de Scandola, inventaire spécifique (1984/85-1991). Trav. Sci. Parc nat. Res. Nat. Corse, Fr, 36: 85-108.
 * Casevitz-Weulersse J. 2014. A propos des fourmis de la Corse: Plagiolepis delaugerrei n. sp., parasite de Plagiolepis taurica Santschi, 1920 (Insecta: Hymenoptera: Formicidae: Formicinae). Le bulletin d’Arthropoda 47: 27-33.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Casevitz-Weulersse, J. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Castillo-Miralbes M. 2001. Artropodos presentes en carrona de cerdos en la comarca de la litera (Huesca). Bol. S.E.A. 28: 133-140.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Catarineu C., G. G. Barbera, and J. L. Reyes-Lopez. 2017. A new ant species, Temnothorax ansei sp. N. (Hymenoptera: Formicidae) from the arid environments of South-eastern Spain. Sociobiology 64(2): 138-145.
 * Cerda, X., E. Angulo, R. Boulay and A. Lenoir. 2009. Individual and collective foraging decisions: a field study of worker recruitment in the gypsy ant Aphaenogaster senilis. Behavioral Ecology and Sociobiology 63:551-562
 * Cerdá, X., J. Retana, and J. Bosch. "Hormigas de Port-Bou (Gerona): una aproximación a su estudio ecológico." Ecología (Madrid) 5 (1991): 413-425.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Comín del Río, P., and Andrés de Haro. "Datos iniciales para un estudio ecológico de las hormigas de Menorca." Bolletí de la Societat d´Historia Natural de les Balears 24 (1980): 23-48.
 * Comín del Río, P., and X. Espadaler. "Ants of the Pityusic islands." In Biogeography of the Pityusic Islands, edited by H. Kuhbler, J. A. Alcover and C. Guerau, 278-231. The Hague: Junk, 1984.
 * Comín del Río, P. "Los Formícidos de Menorca. Contribución al estudio taxonómico, geográfico y biológico." Tesina de licenciatura Universida (1977): 135 pp.
 * Comín del Río, P. Estudio de los formícidos de Baleares: Contribución al estudio taxonómico, geográfìco y biológico. Palma de Mallorca: Universidad de las Islas Baleares, 1988.
 * Cros S., X. Cerda, and J. Retana. 1997. Spatial and temporal variations in the activity patterns of Mediterranean ant communities. Ecoscience 4(3): 269-278.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * De Stefani T. 1889. Miscellanea imenotterologica sicula. Nat. Sicil. 8: 140-145.
 * Delye, G., and J. L. Bonaric. "Fourmis du sud Marocain." Etude de certains milieux du Maroc et de leux evolution recente RCP 249 (1973).
 * Donisthorpe, H. "A first instalment of the ants of Turkey." Annals and Magazine of Natural History (12)3 (1950): 1057-1067.
 * Eidmann, H. "Die Ameisenfauna der Balearen." Zeitschrift für Morphologie und Ökologie der Tiere 6 (1926): 694-742.
 * Eidmann, H. "Zur Kenntnis der Insektekfauna der Balearischen Inseln." Entomologische Mitteilungen 16 (1927): 24-37.
 * Emery C. 1914. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Südeuropäische Ameisen (Hym.). Entomologische Mitteilungen. Berlin-Dahlem 3: 156-159.
 * Emery C. 1925. Notes critiques de myrmécologie. Annales de la Société Entomologique de Belgique 64: 177-191.
 * Emery, C. "Beiträge zur Kenntniss der palaearktischen Ameisen." Öfversigt af Finska Vetenskaps-Societetens Förhandlingar (Helsinki) 20 (1898): 124-151.
 * Emery, C. "Escursioni zoologiche del Dr. Enrico Festa nell'Isola di Rodi. XII. Formiche." Bollettino del Museo di Zoologia ed Anatomia Comparatadella Reale Università di Torino 30 (1915): 1-7.
 * Emery, C. "Exploration scientifique de la Tunisie. Zoologie. - Hyménoptères. Révision critique des fourmis de la Tunisie." Explor. Scient. De la Tunisie Zoll. Hym. (Folleto) Paris. Imp (1891): iii + 21 pp.
 * Espadaler X. F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2009. Ants (Hymenoptera, Formicidae) from Mont-Rebei gorge (Pallars Jussà). Boletín Sociedad Entomológica Aragonesa 44: 393399.
 * Espadaler, X. and Cagniant, H. Plagiolepis Xene Starcke, The First Inquiline Ant From The Balearic Islands (Spain). Psyche. 98:351-354.
 * Espadaler, X. and H. Cagniant. 1991. Psyche. 98: 351-354.
 * Espadaler, X., and H. Cagniant. "Plagiolepis xene Starckë, the first inquiline ant from the Balearic Islands (Spain)." Psyche 98 (4) (1992): 351-354.
 * Espadaler, X., and J. Reyes-López. "Dos nuevas localidades para una (¿rara?) hormiga endémica ibérica,Temnothorax blascoi (Espadaler, 1997) (Hymenoptera, Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 449-450.
 * Espadaler, X., and L. Lopez Soria. "Rareness of certain Mediterranean ant species: fact or artifact?" Insectes Sociaux 38 (1991): 365-377.
 * Espadaler, X., and X. Roig. "Ants from the Montnegre-Corredor Natural Park with description of the male lasius cinereus Seifert." Miscellanea Zoologica 23 (2) (2001): 45-53. Abstract
 * Espadaler, X. "Formicidos de las sierras de Cazorla, del Pozo y Segura (Jaén, España)." Ecología 11 (1997): 489-499.
 * Espadaler, X. "Formícidos de los alrededores de la laguna de Sariñena (Huesca). Descripción del macho de Camponotus foreli Emery." Colección Estudios Altoaragoneses 6 (1986): 109-126.
 * Finzi, B. "Quarto contributo alla conoscenza della fauna mirmecologica della Venezia Giulia." Bollettino della Società Entomologica Italiana 60 (1928): 128-130.
 * Finzi, B. "Raccolte entomologiche nell'Isola di Capraia fatte da C. Mancini e F. Capra (1927-1931). II. Formicidae." Memorie della Società Entomologica Italiana 11 (1933): 162-165.
 * Forel A. 1904. Miscellanea myrmécologiques. Rev. Suisse Zool. 12: 1-52.
 * Forel A. 1904. Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Ezheg. Zool. Muz. 8: 368-388.
 * Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
 * Forel, A. "Les fourmis du Sahara algérien récoltées par M. le Professeur A. Lameere et le Dr. A. Diehl." Annales de la Société Entomologique de Belgique 46 (1902): 147-158.
 * Forel, A. "Miscellanea myrmécologiques." Revue Suisse de Zoologie 12 (1904): 1-52.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gaspar C. 1968. Les fourmis de la Drome et des Basses-Alpes, en France (Hymenoptera, Formicidae). Naturaliste can. 95: 747-766.
 * Gomez C., D. Casellas, J. Oliveras, and J. M. Bas. 2003. Structure of ground-foraging ant assemblages in relation to land-use change in the northwestern Mediterranean region. Biodiversity and Conservation 12: 21352146.
 * Gomez K. 2017. Two species of exotic ants (Hymenoptera: Formicidae) new to Malta. Boletin de la Sociedad Entomologica Aragonesa (S.E.A.) 61: 233-235.
 * Goncalves C., X. Espadaler, J. A. Pereira, S. Santos, and M. I. Patanita. 2014. First records of Strongylognathus caeciliaeForel, 1897 and Temnothorax tyndalei(Forel, 1909) from mainland Portugal (Hymenoptera, Formicidae). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 54: 402.
 * González Martín J. D., and X. Espadaler. 2011. Formicidos del País Vasco (Hymenoptera: Formicidae): Nuevas aportaciones. Heteropterus Revista de Entomologia 11(1): 109-122.
 * Gonçalves M. F., and J. A. Pereira. 2012. Abundance and diversity of soil arthropods in the olive grove ecosystem. Journal of Insect Science 12:20 available online: insectscience.org/12.20
 * Gonçalves, C., I. Patanita, and X. Espadaler. "Substantial, and significant, expansion of ant hosts range for Myrmicinosporidium Hölldobler, 1933 (Fungi)." Insectes Sociaux 59 (2012): 395-399.
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Hamann H. H. F., and W. Klemm. 1976. Ergebnisse der von Dr. O. Paget und Dr. E. Kritscher auf Rhodos durchgeführten zoologischen Exkursionen. XVI. Formicidae. Annalen des Naturhistorischen Museums in Wien 80: 669-679.
 * Hohmann H., F. La Roche, G. Ortega, and J. Barquín. 1993. Bienen, Wespen und Ameisen der Kanarischen Inseln. Veröff. Überseemus. Bremen Naturwiss. 12: 14.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karavaiev V. 1912. Ameisen aus Tunesien und Algerien, nebst einigen unterwegs in Italien gesammelten Arten. Rus. Entomol. Obozr. 12: 1-22.
 * Karavaiev V. 1926. Beiträge zur Ameisenfauna des Kaukasus, nebst einigen Bemerkungen über andere palaearktische Formen. (Fortsetzung). Konowia 5: 161-169.
 * Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Krausse, A. H. "Ueber sardische Ameisen." Archiv für Naturgeschichte, Berlin A 78(7) (1912): 162-166.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis Collection Database
 * Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
 * Livory A. 2003. Les fourmis de la Manche. L'Argiope 39: 25-49.
 * Livory A. Un Nouvel Inventaire: les Fourmis. Argiope 29: 27-34.
 * Lopez, F., J. M. Zorrilla, F. J. Acosta Salmerón, and J. M. Serrano. "Comparative morphological study of Tetramorium caespitum (Linné, 1758) and Tetramorium semilaeve André, 1881 (Hym., Formicidae)." Miscellania Zoologica (Barcelona) 15 (1992): 169-178.
 * Lorite P., J. A. Carrillo, M. F. García, and T. Palomeque. 2000. Chromosome numbers in Spanish Formicidae. III. Subfamily Myrmicinae (Hymenoptera). Sociobiology 36: 555-570.
 * Luque-García, G., J. L. Reyes-López, and J. Fernández-Haeger. "Estudio faunístico de las hormigas (Hymenoptera: Formicidae) de la cuenca del río Guadiamar: Primeras aportaciones." Boletín de la Sociedad Entomológica Aragonesa 30 (2002): 153-159.
 * López, F., J. M. Serrano, and F. J. Acosta. "Compared Iberian distribution of Tetramorium caespitum (Linné, 1758) and Tetramorium semilaeve André, 1881 (Hym., Formicidae)." Anales de Biología 16 (1990): 53-61.
 * López, F. "Estudio morfológico y taxonómico de los grupos de especies ibéricas del género Tetramorium Mayr, 1855." Boletín de la Asociación Española de Entomología 15 (1991): 29-52.
 * Martínez Ibáñez, M. D., and J. M. Serrano Talavera. "Contribución al conocimiento de las hormigas (Hym. Formicidae) del sabinar español." Boletim da Sociedade Portuguesa de Entomologia Suplemento Actas Do I (1985): 33-41.
 * Martínez Ibáñez, M. D., and X. Espadaler. "Revisión de las hormigas ibéricas de la colección M. Medina y nuevos datos de distribución." Actas de las VIII Jornadas de la Asociación Española de Entomología (1986): 1022-1034.
 * Menozzi, C. "Zur Kenntnis der Ameisenfauna der Balearen." Zoologischer Anzeiger 66 (1926): 180-182.
 * Müller, G. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Neto-Silva A., R. J. Dinis-Oliveira, and C. Prado e Castro. 2018. Diversity of the Formicidae (Hymenoptera) carrion communities in Lisbon (Portugal): preliminary approach as seasonal and geographic indicators. Forensic Sciences Research 3(1): 65-73.
 * Obregon R., J. Lopez, J. L. Reyes-Lopez. 2014. A catalogue of the ant fauna (Hymenoptera, Formicidae) of Sierra Mágina (Jaén, southern Spain). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 370374.
 * Ordóñez-Urbano C., J. Reyes-López, and S. Carpintero-Ortega. 2007. Estudio faunísidos de los formícidos (Hymenoptera: Formicidae) asociados a los bosques de ribera en la Provincia de Córdoba (España). Primeras Aportaciones. Boletin Sociedad Entomológica Aragonesa 40: 367-375.
 * Ortiz, F. J., and A. Tinaut. "Formícidos del litoral granadino [1989]." Orsis 3 (1989): 145-163.
 * Ortiz, F. J. "Formícidos del litoral granadino." Memoria de Licenciatura Universida (1985): 206 pp.
 * Oussalah N., F. Marniche, X. Espadaler, and M. Biche. 2019. Exotic ants from the Maghreb (Hymenoptera, Formicidae) with first report of the Hairy Alien Ant Nylanderia jaegerskioeldi (Mayr) in Algeria. Arxius de Miscel·lania Zoologica 17: 45–58.
 * Ovazza M. 1950. Contribution à la connaissance des fourmis des Pyrénées-Orientales. Récoltes de J. Hamon Vie et Milieu 1: 93-94.
 * Ovazza M. 1953. Contributions a l'etude biologique de la Camargue, Formicides. Vie et Milieu 4(4) : 751-753.
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Pascual, M. R. "Estudio taxonómico y ecológico de los Formícidos de las Sierras de Alfacar, La Yedra, Huétor y Harana." Tesis Doctoral Universida (1986): 264 pp.
 * Pereira, J. A., A. Bento, D. Sousa, M. Campos, and L. Torres. "Estudo preliminar sobre as formigas (Hymenoptera: Formicidae) associadas ao olival da Terra Quente Transmontana (Nordeste de Portugal." Boletin de Sanidad Vegetal, Plagas 28 (2002): 357-365.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Pisarski B. 1964. Fauna Mrowek Afganistanu. Bibliogr. k. 160-166, Nieoprawiony maszynopis pracy, Praca doktorska. Instytut Zoologiczny PAN, 1964, Bibliogr. p. 160-166
 * Platner C., J. Pinol, D. Sanders, and X. Espadaler. 2012. Trophic diversity in a Mediterranean food webStable isotope analysis of an ant community of an organic citrus grove. Basic and Applied Ecology 13: 587596.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Prado e Castro C., M. D. Garcia, C. Palma, and M. D. Martinez-Ibanez. 2014. First report on sarcosaprophagous Formicidae from Portugal (Insecta: Hymenoptera). Annales de la Société entomologique de France 50(1): 51-58.
 * Redolfi I., A. Tinaut, F. Pascual, and M. Campos. 2004. Nest density of the nat assemblage (Formicidae) in three olive orchards under different agricultural management in Granada, Spain. Ecología Aplicada 3(1,2): 73-81.
 * Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
 * Reyes López, J. L., and G. Luque García. "Nuevas citas de Monomorium andrei Saunders, 1890 (Hymenoptera: Formicidae) para la Península Ibérica y Baleares." Boletín de la Asociación Española de Entomología 27 (1-4) (2003): 221-222.
 * Reyes, J. L. "Descripción de Messor celiae nov. sp." Boletín de la Asociación Española de Entomología 9 (1985): 255-261.
 * Reyes-Lopez J., S. Carpintero. 2014. Comparison of the exotic and native ant communities (Hymenoptera: Formicidae) in urban green areas at inland, coastal and insular sites in Spain. Eur. J. Entomol. 111(3): 421428.
 * Reyes-López J., S. Carpintero-Ortega, and E. Retamoza-Muñoz. 2010. Adiciones a la relación de especies de hormigas (Hym., Formicidae) del Parque Natural del Cabo de Gata- Níjar (Almería, España) Boletín de la Asociación Española de Entomología 34: 67-76.
 * Rigato S., and I. Toni. 2011. Short notes 21. Hymenoptera, Formicidae. Pp. 873-882 in: Nardi, G.; Whitmore, D.; Bardiani, M.; Birtele, D.; Mason, F.; Spada, L.; Cerretti, P. (eds.) 2011. Biodiversity of Marganai and Montimannu (Sardinia). Research in the framework of the ICP Forests network. Conservazione Habitat Invertebrati, 5. Sommacampagna, Verona: Cierre Edizioni, 896 pp.
 * Roig X., X. Espadaler, R. Cusco, F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2008. Ants in gypsum areas. First record from Catalonia (Iberian Peninsula) of Cardiocondyla batesii Forel (Hymenoptera, Formicidae) with an updated list of Catalan Formicidae. Boletin Sociedad Entomologica Aragonesa 42: 189-192.
 * Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
 * Sandor C., and M. Balint. 2004. Redescription of Tetramorium hungaricum RÖSZLER, 1935, a related species of T. caespitum (Linnaeus, 1758) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 6: 49-59.
 * Sanetra M., R. Güsten, and A. Schulz. 1999. On the taxonomy and distribution of Italian Tetramorium species and their social parasites (Hymenoptera Formicidae). Memorie della Societa Entomologica Italiana 77: 317-357.
 * Santos S. A. P., J. E. Cabanas, and J. A. Pereira. 2007. Abundance and diversity of soil arthropods in olive grove ecosystem (Portugal): Effect of pitfall trap type. European Journal of Soil Biology 43: 77-83.
 * Santschi F. 1910. Contributions à la faune entomologique de la Roumanie. Formicides capturées par Mr. A. L. Montandon. Buletinul Societatii Române de Stiinte 19: 648-652.
 * Santschi, F. 1921. Notes sur les fourmis paléarctiques. I. Quelques fourmis du nord de l'Afrique et des Canaries. Real Sociedad Española de Historia Natural, Memorias, Tomo del Cincuentenario : 424-436
 * Santschi, F. "A propos du Tetramorium caespitum L." Folia Myrmecologica et Termitologica 1 (1927): 52-58.
 * Schembri, Stephen P. and Cedric A. Collingwood. A Revision of the Myrmecofauna of the Maltese Islands. 417-442.
 * Schlick-Steiner, B.C., F.M. Steiner, M. Sanetra, G. Heller, C. Stauffer, E. Christian and B. Seifert. 2005. Queen size dimorphism in the ant Tetramorium moravicum (Hymenoptera, Formicidae): Morphometric, molecular genetic and experimental evidence. Insects Sociaux 52:186-193
 * Schmitz, H. "Ein Verzeichnis portugiesischer Ameisen (Formicidae, Hymenoptera)." Brotéria Ser. Cienc. Nat. 24 (1955): 27-37.
 * Schulz A. 1991. Tetramorium semilaeve (Hym.: Formicidae, Myrmicinae) und Bothriomyrmex gibbus (Hym.: Formicidae, Dolichoderinae) neu für Österreich sowie über die Verbreitung von Leptothorax sordidulus (Hym.: Formicidae, Myrmicinae). Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 43: 120-122.
 * Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
 * Sommer F. 1984. Etude de la myrmécofaune de la réserve de la Massane. Laboratoire Arago, Banyuls sur Mer (66), 29 pages.
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Soyer B. 1951. Les fourmis des collines des environs de Marseille. Bull Soc Linn Provence 18: 3-6.
 * Steiner F. M., B. Seifert, K. Moder, and B. C. Schlick-Steiner. 2010. A multisource solution for a complex problem in biodiversity research: description of the cryptic ant species Tetramorium alpestre sp. n. (Hymenoptera: Formicidae). Zoologischer Anzeiger 249: 223-254.
 * Stitz, H. "Ameisen aus dem westlichen Mittelmeergebiet und von den Kanarischen Inseln." Mitteilungen aus den Zoologischen Museum in Berlin 8 (1917): 333-353.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Taheri A., J. Reyes-Lopez, and N. Bennas. 2014. Contribution a l'etude de la fauna myrmecologique du parc national de Talassemtane (nord du Maroc): biodiversite, biogeographie et especes indicatrices. Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 225236.
 * Taheri A., and J. Reyes-Lopez. 2017. First record in Morocco of the wooly ant Tetramorium lanuginosum (Hymenoptera: Formicidae), an Asian species now known from all southern Mediterranean countries. Transactions American Entomological Society 143: 515-516.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut A., O. Mira, J. M. Vidal, and A. Aguirre-Segura. 2009. The ants of Cabo de Gata (Almería). Faunistics aspects (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 33 (1-2): 227-251.
 * Tinaut A., and F. J. Ortiz. 1988. Introduccion al conocimiento de las hormigas de la provincia de Almeria (Hymenoptera: Formicidae). Publicaciones del Instituto de Estudios Almerienses. Boletín (Ciencias) 8: 223-231.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Tohme G. 1996. Formicidae. Etude de la diversité biologique n° 4 . Ministère de lAgriculture à Beyrouth (Eds.). P85-87.
 * Tohme G., and H. Tohme. 2014. Nouvelles liste des especes de fourmis du Liban (Hymenoptera, Formicoidea). Lebanese Science Journal 15(1): 133-141.
 * Trigos Peral G., and J. L. Reyes-Lopez. 2015. New record of Temnothorax algiricus (Hymenoptera, Formicidae) from the Iberian Peninsula and ten additional new records for the ant fauna from Málaga, with preliminary list of species from Natural Park Sierras de Tejeda, Almijara y Alhama (Andalusia, Spain). Boln. Asoc. esp. Ent. 39 (3-4): 411-416.
 * Trigos Peral G., and J. Reyes Lopez. 2016. Ants as bioindicators for monitoring of urban green in the south of the Iberian Peninsula. Proposal of functional groups. Iberomyrmex 8: 37-38.
 * Vicidomini S., and C. Pignataro. 2007. Contributo alla conoscenza della biodiversità del Parco Nazionale del Cilento e Vallo di Diano: gli Imenotteri veleniferi più comuni e check-list delle specie di Apoidea e Formicidae della provincia di Salerno. Il Naturalista Campano 34: 1-15.
 * Wheeler, W. M. "Ants of the Balearic Islands." Folia Myrmecologica et Termitologica 1 (1926): 1-6.
 * Zariquiey, R. "Formigues de Cadaqués (prov. De Gerona)." Butll. Inst. Catalana Hist. Nat. (1930): 92.
 * Zimmermann, S. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
 * Zina V., M. Branco, and J. C. Franco. 2017. Ant community associated to citrus canopy in Southern Portugal over a century after the invasion by the Argentine ant. Phytoparasitica 45:183–200.
 * de Haro, Andrés, C. A. Collingwood, and P. Comín del Río. "Prospección mirmecológica por Ibiza y Formentera (Baleares)." Orsis 2 (1986): 115-120.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Andalucia." Boletín de la Estación Central de Ecología 6 (12) (1977): 85-90.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por Extremadura (España) y Sao Brás-Almodovar, Alcácer do Sal, Serra da Estrela (Portugal)." Boletim da Sociedade Portuguesa de Entomologia Suplemento 3(1) (1992): 95-104.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por el litoral mediterráneo de Marruecos (Cabo Negro, Martil, Oued Lau) y su comparación con la zona meridional ibérica." Orsis 9 (1994): 97-104.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por la península Tingitana al norte del Rif (Marruecos)." Orsis (Organismes i Sistemes) 12 (1997): 93-99.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por las sierras de Aitana-Alfaro y los cabos de la Nao-San Antonio (Alicante) y su comparación con la fauna balear y de Córcega-Cerdeña." Orsis 3 (1988): 165-172.
 * de Haro, Andrés. "Contribución al conocimiento de los formícidos del cabo de Creus (Gerona)." Pirineos 117 (1982): 49-57.