Messor capitatus

A west-European ant, reported also from NW Africa (Rigato & Toni, 2011). However, Borowiec (2014) notes that this species needs taxonomic study as recently collected material showed that populations from the western part of the Mediterranean differ slightly from the eastern populations, suggesting that a complex of related species may be present.

Distribution based on Regional Taxon Lists
Palaearctic Region: Balearic Islands, Bulgaria, Croatia, France, Greece, Iberian Peninsula, Italy, Malta, Monaco, Montenegro, Portugal, North Macedonia, Romania, Slovenia, Spain.

Nomenclature

 * . Formica capitata Latreille, 1798: 46 (w.) FRANCE.
 * Type-material: syntype workers (number not stated).
 * Type-locality: France: vic. Bordeaux (Latreille?).
 * Type-depository: MNHN.
 * Latreille, 1802c: 235 (q.); Mayr, 1855: 463 (m.).
 * Combination in Manica: Jurine, 1807: 279;
 * combination in Myrmica: Losana, 1834: 325;
 * combination in Atta: Latreille, 1809: 130; Leach, 1815: 147; Brullé, 1833: 327; Lepeletier de Saint-Fargeau, 1835: 173; Lucas, H. 1849: 300;
 * combination in Aphaenogaster: Roger, 1863b: 29;
 * combination in Stenamma (Messor): Forel, 1902a: 148;
 * combination in Aphaenogaster (Messor): Ruzsky, 1905b: 744;
 * combination in Messor: Emery, 1908e: 449.
 * As unavailable (infrasubspecific) name: Emery, 1921f: 69; Emery, 1922c: 97; Emery, 1924b: 12; Kutter, 1927: 98.
 * Junior synonym of barbarus: Smith, F. 1858b: 162; Mayr, 1861: 66 (in key); Smith, F. 1861a: 35; Roger, 1863b: 29; Mayr, 1863: 395; Dours, 1873: 167; André, 1874: 203 (in list); Emery & Forel, 1879: 461; Nasonov, 1889: 37.
 * Subspecies of barbarus: Emery, 1891b: 12; Emery, in Dalla Torre, 1893: 100 (footnote); Emery, 1893c: 84; Forel, 1894d: 31; Forel, 1902a: 148; Forel, 1904b: 373; Forel, 1904c: 5; Forel, 1905b: 176; Ruzsky, 1905b: 744; Forel, 1906c: 190; Forel, 1907b: 204; Forel, 1907e: 15; Emery, 1908e: 449 (redescription); Forel, 1909c: 104; Forel, 1909e: 371; Karavaiev, 1910b: 62; Karavaiev, 1912a: 7; Emery, 1912f: 97; Emery, 1916b: 142; Stitz, 1917: 343; Santschi, 1919e: 244; Menozzi, 1922b: 326; Ruzsky, 1923: 4; Santschi, 1923f: 325; Finzi, 1924a: 13; Santschi, 1925g: 341; Santschi, 1926f: 286; Finzi, 1929: 80; Cori & Finzi, 1931: 238; Santschi, 1931a: 4; Santschi, 1932g: 3; Finzi, 1933: 163; Grandi, 1935: 100; Zimmermann, 1935: 17; Menozzi, 1936d: 276; Finzi, 1940: 159; Consani & Zangheri, 1952: 40; Ceballos, 1956: 301.
 * Status as species: Latreille, 1802c: 234; Jurine, 1807: 279; Latreille, 1809: 130; Leach, 1815: 147; Brullé, 1833: 327; Losana, 1834: 325; Lepeletier de Saint-Fargeau, 1835: 173; Brullé, 1840: 84; Lucas, H. 1849: 300; Schenck, 1852: 135; Mayr, 1855: 462 (redescription); Nylander, 1856b: 83; Mayr, 1866b: 896; Dalla Torre, 1893: 101; Bondroit, 1918: 153; Lomnicki, 1925b: 2; Baroni Urbani, 1964b: 40; Baroni Urbani, 1964c: 150; Cagniant, 1966b: 279; Bernard, 1967: 147 (redescription); Baroni Urbani, 1968b: 427; Cagniant, 1968a: 143; Baroni Urbani, 1969a: 331; Collingwood & Yarrow, 1969: 62; Cagniant, 1970a: 416; Baroni Urbani, 1971c: 57; Baroni Urbani, 1974: 232; Collingwood, 1978: 81 (in key); Bernard, 1980: 266; Schembri & Collingwood, 1981: 425; Agosti & Collingwood, 1987a: 54; Agosti & Collingwood, 1987b: 271 (in key); Casevitz-Weulersse, 1990b: 156 (in key); De Haro & Collingwood, 1994: 99; Bolton, 1995b: 253; Mei, 1995: 761; Poldi, et al. 1995: 3; Espadaler, 1997b: 30; Cagniant & Espadaler, 1998: 423; Markó & Csösz, 2002: 116; Bračko, 2006: 137; Cagniant, 2006a: 197; Petrov, 2006: 93 (in key); Bračko, 2007: 17; Casevitz-Weulersse & Galkowski, 2009: 489; Lapeva-Gjonova, et al. 2010: 15; Karaman, M.G. 2011b: 36; Legakis, 2011: 9; Borowiec, L. & Salata, 2012: 512; Borowiec, L. 2014: 103; Bračko, et al. 2014: 14; Lebas, et al. 2016: 294.
 * Senior synonym of galbula: Emery, 1921f: 70; Bolton, 1995b: 253.
 * Senior synonym of huberiana: Emery, 1921f: 70; Bolton, 1995b: 253.
 * Senior synonym of nigra: Menozzi, 1921: 26; Emery, 1922c: 97; Finzi, 1924a: 13; Finzi, 1929: 80; Cori & Finzi, 1931: 238; Zimmermann, 1935: 17; Bernard, 1967: 147; Baroni Urbani, 1971c: 58; Collingwood, 1978: 68; Bolton, 1995b: 253; Casevitz-Weulersse & Galkowski, 2009: 489.
 * Distribution: Algeria, Bosnia, Bulgaria, Croatia, France (+ Corsica), Greece, Italy (+ Sardinia, Sicily), Macedonia, Malta, Morocco, Portugal, Serbia, Slovenia, Spain (+ Balearics), Tunisia.
 * Current subspecies: nominal plus nigricans Santschi, 1929f.
 * galbula. Myrmica galbula Losana, 1834: 326, pl. 36, fig. 5 (w.) ITALY.
 * Type-material: holotype (?) worker.
 * [Note: no indication of number of specimens is given.]
 * Type-locality: Italy: Piedmont (M. Losana).
 * Type-depository: unknown (no material is known to exist).
 * [Note: author not mentioned in Horn & Kahle, 1935: 159.]
 * Combination in Aphaenogaster: Roger, 1863b: 29.
 * Status as species: Dalla Torre, 1893: 109.
 * Junior synonym of barbarus: Roger, 1863b: 29; Emery & Forel, 1879: 461.
 * Junior synonym of capitatus: Emery, 1921f: 70; Bolton, 1995b: 254.
 * huberiana. Formica huberiana Leach, 1825: 290 (w.q.m.) FRANCE.
 * Type-material: syntype worker(s), syntype queen(s), syntype male(s) (numbers not stated).
 * Type-localities: Switzerland: northern Switzerland (“Habitat in Helvetia Boreali”) (W.E. Leach), France: Risso, nr Nice (W.E. Leach).
 * Type-depository: unknown (in BMNH, according to Horn & Kahle, 1935: 151).
 * Combination in Aphaenogaster: Roger, 1863b: 29.
 * Junior synonym of barbarus: Roger, 1863b: 29; Mayr, 1863: 395; Emery & Forel, 1879: 461; Dalla Torre, 1893: 100.
 * Junior synonym of capitatus: Emery, 1921f: 70; Bolton, 1995b: 254.
 * nigra. Aphaenogaster barbara var. nigra André, 1883b: 355 (w.q.) FRANCE (mainland, Corsica), ITALY (mainland, Sardinia, Sicily), ALGERIA.
 * Type-material: syntype workers, syntype queens (numbers not stated).
 * Type-localities: France: (+ Corsica) (no further data), Italy: (+ Sardinia, Sicily) (no further data), Algeria (no further data).
 * Type-depository: MNHN.
 * Emery, 1908e: 447 (m.).
 * Combination in Stenamma (Messor): Ruzsky, 1902d: 28;
 * combination in Messor: Emery, 1908e: 446.
 * As unavailable (infrasubspecific) name: Emery, 1891b: 12; Emery, 1908e: 446; Emery, 1912f: 97; Emery, 1914d: 156; Emery, 1915a: 245; Emery, 1916b: 142; Menozzi, 1918: 82; Soudek, 1925b: 15.
 * Subspecies of barbarus: Dalla Torre, 1893: 100; Ruzsky, 1902d: 28; Emery, 1908d: 23; Forel, 1909c: 104; Forel, 1909e: 379; Forel, 1911d: 346; Krausse, 1912b: 164; Müller, 1923b: 63.
 * Junior synonym of capitatus: Menozzi, 1921: 26; Emery, 1922c: 97; Finzi, 1924a: 13; Finzi, 1929: 80; Cori & Finzi, 1931: 238; Zimmermann, 1935: 17; Bernard, 1967: 147; Baroni Urbani, 1971c: 58; Collingwood, 1978: 68; Bolton, 1995b: 255; Casevitz-Weulersse & Galkowski, 2009: 489.

References based on Global Ant Biodiversity Informatics

 * Abril S., and C. Gomez. 2013. Rapid assessment of ant assemblages in public pine forests of the central Iberian Peninsula. Forest Ecology and Management 293: 7984.
 * Acosta, F. J., M. D. Martínez Ibáñez, and M. A. Morales. "Contribución al conocimiento de la mirmecofauna del encinar peninsular. I." Boletín de la Asociación Española de Entomología 6 (2) (1983): 379-391.
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * Andoni V. 1977. Kontribut mbi Himenopteret e familjes Formicidae te vendit tone. Buletini I Shkencave te Natyres 31(2): 93-101.
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antarea (at www.antarea.fr on June 11th 2017)
 * Arnan, X., A. Rodrigo and J. Retana. 2006. Post-fire recovery of Mediterranean ground ant communities follows vegetation and dryness gradients. Journal of Biogeography 33(7):1246-1258
 * Arnan, X., J. Retana, A. Rodrigo and X. Cerda. 2010. Foraging behaviour of harvesting ants determines seed removal and dispersal. Insectes Sociaux 54:421-430
 * Asociacion Iberica de Mirmecologia. 2012. List of species collected during the Taxomara Tres Cantos 2012. Iberomyrmex 4: 17-18.
 * Asociacion Iberica de Mirmecologia. 2015. List of species collected during the Taxomara Lisboa 2015 Iberomyrmex 7: 44-45.
 * Asociacion Iberica de Mirmecologia. 2016. List of species collected during the Taxomara Murcia 2016. Iberomyrmex 8: 48-49.
 * Barech G., M. Khaldi, F. Zahra Boujelal, and X. Espadaler. 2018. Diversity and structure of the ant fauna of the El Ksob dam in Algeria: new record for Aphaenogaster rupestris Forel, 1909 (Hymenoptera: Formicidae). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.) 62: 253-258.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. IV. La fauna mirmecologica delle isole Maltesi ed il suo significato ecologico e biogeografico. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 77: 408-559.
 * Baroni Urbani C. 1968. Studi sulla mirmecofauna d'Italia. V. Aspetti ecologici della Riviera del M. Cònero. Boll. Zool. 35: 39-76.
 * Baroni Urbani C. 1969. Studi sulla mirmecofauna d'Italia. VIII. L'Isola di Giannutri ed alcuni scogli minori dell'arcipelago Toscano. Atti Soc. Toscana Sci. Nat. Mem. Ser. B 75: 325-338.
 * Baroni Urbani C. 1974. Studi sulla mirmecofauna d'Italia. XII. Le Isole Pontine. Fragm. Entomol. 9: 225-252.
 * Baroni Urbani, C.. "Formiche dell'Italia appenninica (Studi sulla mirmecofauna d'Italia, III)." Memorie del Museo Civico di Storia Naturale di Verona 12 (1964): 149-172.
 * Baroni Urbani, C.. "Studi sulla mirmecofauna d'Italia. II. Formiche di Sicilia." Atti dell'Accademia Gioenia di Scienze Naturali in Catania (6) 16 (1964): 25-66.
 * Barrett K. E. J. 1968. Ants in western France. Entomologist 101: 153-155.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Baugnee J. Y., and J. F. Godeau. 2000. Signalement de Monomorium monomorium Bolton et de quelques autres fourmis à Belle-Ile-en-Mer, en Bretagne (Hymenoptera, Formicidae). Bulletin de la Société entomologique de France 105(2): 205-208.
 * Bernard F. 1958. Les fourmis des iles Pelagie, comparaison avec d'autres faunes insulaires. Esttrato dalla Rivista di Biologia Coloniale XVI: 67-79.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1978. Fourmis et milieu dans le massif des Maures. Vie et Milieu. Série C. Biologie Terrestre 27: 83-118.
 * Bonaric J. C. 1971. Contribution a l'etude systematique et ecologique des formicides du Bas-Languedoc. PhD thesis Universite des sciences et techniques du Languedoc, 175 pages.
 * Bonaric J. C. 1971. Étude systématique et écologique des fourmis de lHérault. Ann. Soc. Hortic. Hist. Nat. Hérault 111: 81-87.
 * Bondroit, J.. "Les fourmis de France et de Belgique." Annales de la Société Entomologique de France 87 (1918): 1-174.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Boulay, R., J.M. Fedriani, A.J. Manzaneda and X. Cerda. 2005. Indirect Effects of Alternative Food Resources in an Ant-Plant Interaction. Oecologia 144(1):72-79
 * Bouzekri M. A. 2011. Bioecologie des Formicidae dans la region de Djelfa: nidification et relation avec les plantes. Mphil Ecole National Superieure agronomique. 153pp.
 * Bouzekri M., S. Daoudi-Hacini, H. Cagniant, and S. E. Doumandji. 2015. Etude comparative des associations (plantes-fourmis) dans une region steppique (cas de la region de Djelfa, Algerie). Lebanese Science Journal 16(1): 69-77.
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko G., and L. Cesnik. 2016. Fist records of six ant species (Hymenoptera: Formicidae) for Slovenia. Natura Sloveniae 18(2): 39-46.
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Cagniant, H., and X. Espadaler. "Le genre Messor au Maroc." Annales de la Société Entomologique de France (N.S.) 33 (4) (1998): 419-434.
 * Cagniant, H., and X. Espadaler. "Liste des espèces de fourmis du Maroc." Actes des Colloques Insectes Sociaux 8 (1993): 89-93.
 * Cagniant, H.. "Contribution à la connaissance des fourmis marocaines. Description des trois castes d'Aphaenogaster wilsoni n. sp. (Hymenoptera, Myrmicidae)." Nouvelle Revue d'Entomologie (n.s.)5 (1988): 49-55.
 * Cagniant, H.. "Deuxième liste de fourmis d'Algérie récoltées principalement en forêt. (1re partie)." Bulletin de la Société d'Histoire Naturelle de Toulouse 105 (1970): 405-430.
 * Cagniant, H.. "Liste actualisee des fourmis du Maroc (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 193-200.
 * Cagniant, H.. "Note sur le peuplement en fourmis d'une montagne de la région d'Alger, l'Atlas de Blida." Bulletin de la Société d'Histoire Naturelle de Toulouse 102 (1966): 278-284.
 * Cagniant, H.. Les peuplements de fourmis des forêts algériennes: écologie, biocénotique, essai biologique. Universite de Toulouse, 1973.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J. 1990. Étude systématique de la myrmécofaune corse (Hymenoptera, Formicidae) (Première partie). Bull. Mus. Natl. Hist. Nat. Sect. A Zool. Biol. Écol. Anim. (4) 12: 135-163.
 * Casevitz-Weulersse J. 2014. A propos des fourmis de la Corse: Plagiolepis delaugerrei n. sp., parasite de Plagiolepis taurica Santschi, 1920 (Insecta: Hymenoptera: Formicidae: Formicinae). Le bulletin d’Arthropoda 47: 27-33.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse, J.. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Castracani C., D. A. Grasso, A. Fanfani, and A. Mori. 2010. The ant fauna of Castelporziano Presidential Reserve (Rome, Italy) as a model for the analysis of ant community structure in relation to environmental variation in Mediterranean ecosystems. J Insect Conserv 14: 585594.
 * Cecconi G. 1908. Contributo alla fauna delle Isole Tremiti. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 23(583): 1-53.
 * Cerdá, X., J. Retana, and J. Bosch. "Hormigas de Port-Bou (Gerona): una aproximación a su estudio ecológico." Ecología (Madrid) 5 (1991): 413-425.
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Comín del Río, P., and X. Espadaler. "Ants of the Pityusic islands." In Biogeography of the Pityusic Islands, edited by H. Kuhbler, J. A. Alcover and C. Guerau, 278-231. The Hague: Junk, 1984.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Cros S., X. Cerda, and J. Retana. 1997. Spatial and temporal variations in the activity patterns of Mediterranean ant communities. Ecoscience 4(3): 269-278.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Du Merle P. 1978. Les peuplements de fourmis et les peuplements d'acridiens du Mont Ventoux II. - Les peuplements de fourmis. Terre Vie 32(1): 161-218.
 * Emery C. 1914. Wissenschaftliche Ergebnisse der Bearbeitung von O. Leonhard's Sammlungen. 5. Südeuropäische Ameisen (Hym.). Entomologische Mitteilungen. Berlin-Dahlem 3: 156-159.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Emery, C.. "Alcune formiche di Malta." Bollettino della Società Entomologica Italiana 56 (1924): 11-12.
 * Emery, C.. "Beiträge zur Monographie der Formiciden des paläarktischen Faunengebietes. (Hym.) Teil III. Messor, Goniomma, Oxyopomyrmex." Deutsche Entomologische Zeitschrift 1908 (1908): 437-465.
 * Emery, C.. "Fauna entomologica italiana. I. Hymenoptera.-Formicidae." Bollettino della Societa Entomologica Italiana 47 (1916): 79-275.
 * Espadaler X. F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2009. Ants (Hymenoptera, Formicidae) from Mont-Rebei gorge (Pallars Jussà). Boletín Sociedad Entomológica Aragonesa 44: 393399.
 * Espadaler X., and J. I. López-Colón. 2011. Hormigas (Hymenoptera, Formicidae) de una zona de yesos de la Comunidad de Madrid (España) Boletín de la Sociedad Entomológica Aragonesa 49: 261-264.
 * Espadaler, X., and J. Reyes-López. "Dos nuevas localidades para una (¿rara?) hormiga endémica ibérica,Temnothorax blascoi (Espadaler, 1997) (Hymenoptera, Formicidae)." Boletín de la Sociedad Entomológica Aragonesa 42 (2008): 449-450.
 * Espadaler, X.. "Formicidos de las sierras de Cazorla, del Pozo y Segura (Jaén, España)." Ecología 11 (1997): 489-499.
 * Espadaler, X.. "Lasius neglectus Van Loon, Boosma & Andrásfalvy, 1990 a potential pest ant in Spain." Orsis 14 (1999): 43-46.
 * Finzi B. 1929. Le forme italiane del genere Messor. Bollettino della Società Entomologica Italiana 61: 75-94.
 * Finzi, B.. "Formiche dell'isola d'Elba e Monte Argentario." Bollettino della Società Entomologica Italiana 56 (1924): 12-15.
 * Finzi, B.. "Formiche della Libia." Memorie della Società Entomologica Italiana 18 (1940): 155-166.
 * Finzi, B.. "Raccolte entomologiche nell'Isola di Capraia fatte da C. Mancini e F. Capra (1927-1931). II. Formicidae." Memorie della Società Entomologica Italiana 11 (1933): 162-165.
 * Forel A. 1904. Miscellanea myrmécologiques. Rev. Suisse Zool. 12: 1-52.
 * Forel A. 1904. Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Ezheg. Zool. Muz. 8: 368-388.
 * Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel, A.. "Fourmis d'Espagne récoltées par M. O. Vogt et Mme Cécile Vogt, Docteurs en médecine." Annales de la Société Entomologique de Belgique 53 (1909): 103-106.
 * Forel, A.. "Miscellanea myrmécologiques." Revue Suisse de Zoologie 12 (1904): 1-52.
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Galkowski C. 2011. Une liste des fourmis (Hymenoptera, Formicidae) récoltées dans la région de Grasse, avec la mention dune nouvelle espèce de la faune de France. Bulletin de la Société linnéenne de Provence, 62 : 41-44.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * García J. C., and X. Espadaler. 2011. Segunda cita de Lasius bicornis (Förster, 1850) en España (Hymenoptera, Formicidae) Iberomyrmex 3: 21-22
 * Gomez C., D. Casellas, J. Oliveras, and J. M. Bas. 2003. Structure of ground-foraging ant assemblages in relation to land-use change in the northwestern Mediterranean region. Biodiversity and Conservation 12: 21352146.
 * Gomez K. 2017. Two species of exotic ants (Hymenoptera: Formicidae) new to Malta. Boletin de la Sociedad Entomologica Aragonesa (S.E.A.) 61: 233-235.
 * Gomez K., D. Martinez, and X. Espadaler. 2018. Phylogeny of the ant genus Aphaenogaster (Hymenoptera: Formicidae) in the Iberian Peninsula, with the description of a new species. Sociobiology 65: 215-224.
 * González Martín J. D., and X. Espadaler. 2011. Formicidos del País Vasco (Hymenoptera: Formicidae): Nuevas aportaciones. Heteropterus Revista de Entomologia 11(1): 109-122.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Grasso, D.A., M.F. Sledge, F. Le Moli, A. Mori and S. Turillazzi. 2005. Nest-area marking with faeces: a chemical signature that allows colony-level recognition in seed harvesting ants (Hymenoptera, Formicidae). Insectes Sociaux 52:36-44
 * Gómez C., and S. Abril. 2011. Selective logging in public pine forests of the central Iberian Peninsula: Effects of the recovery process on ant assemblages Forest Ecology and Management 262: 1061-1066.
 * Hernandez Cuba O., C. Perez Banon, and M. A. Marcos Garcia. 2008. Los formicidos (Hymenoptera, Formicidae) del Parque natural de la Font Roja. Iberis 6: 9-20.
 * Hernandez Cuba O., J. L. Perez-Bote, and J. M. Garcia Jimenez. 2006. Los formicidos (Hymenoptera: Formicidae) del Parque Natural de Cornaldo (suroeste de la Peninsula Iberica). Boletin Sociedad Entomologica Aragonesa 38: 356-358.
 * Jiménez, J., and A. Tinaut. "Mirmecofauna de la Sierra de Loja (Granada)." Orsis 7 (1992): 97-111.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karavaiev V. 1912. Ameisen aus Tunesien und Algerien, nebst einigen unterwegs in Italien gesammelten Arten. Rus. Entomol. Obozr. 12: 1-22.
 * Korlevic, A.. "Prilozi fauni hrvatskih opnokrilaca." Glasn. Hrv. Narav. Dr. 5 (1890): 189-250.
 * Krausse, A. H.. "Ueber sardische Ameisen." Archiv für Naturgeschichte, Berlin A 78(7) (1912): 162-166.
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis A. 2001. Ants (Hymenoptera, Formicidae) collected in the eastern Pyrenees and surrounding area, September 1999. Proceedings of the International Colloqium of the European Invertebrate Survey: Cartography and Conservation of Biodiversity Marcevol Priory, Arboussols, france 30.8-4.9.1999. OPIE.
 * Legakis Collection Database
 * Lomnicki, J.. "Une contribution à la connaissance de la faune des fourmis des îles Baléares." Polskie Pismo Entomologiczne - Bulletin Entomologique de Pologne 4 (1925): 1-3.
 * López, F.. "Descripción de un nido de Lasius umbratus Nylander 1864 encontrado en el interior de una cueva y algunas consideraciones sobre el caraacter cavernícola de las hormigas (Hymenoptera, Formicidae)." Memories Bioespelologiques 15 (1988): 107-115.
 * Markó B., and S. Csősz. 2002. Die europäischen Ameisenarten (Hymenoptera: Formicidae) des Hermannstädter (Sibiu, Rumänien) Naturkundemuseums I.: Unterfamilien Ponerinae, Myrmicinae und Dolichoderinae. Annales Historico-Naturales Musei Nationalis Hungarici 94: 109-121.
 * Martínez Ibáñez, M. D., and J. M. Serrano Talavera. "Contribución al conocimiento de las hormigas (Hym. Formicidae) del sabinar español." Boletim da Sociedade Portuguesa de Entomologia Suplemento Actas Do I (1985): 33-41.
 * Martínez Ibáñez, M. D., and X. Espadaler. "Revisión de las hormigas ibéricas de la colección M. Medina y nuevos datos de distribución." Actas de las VIII Jornadas de la Asociación Española de Entomología (1986): 1022-1034.
 * Mayr, G.. "Formicina austriaca. Beschreibung der bisher im österreichischen Kaiserstaate aufgefundenen Ameisen, nebst Hinzufügung jener in Deutschland, in der Schweiz und in Italien vorkommenden Arten." Verhandlungen der Zoologisch-Botanischen Vereins in Wien 5 (1855): 273-478.
 * Mei M. 1995. Arthropoda di Lampedusa, Linosa e Pantelleria (Canale di Sicilia, Mar Mediterraneo). Hymenoptera Formicidae (con diagnosi di due nuove specie). Il Naturalista Siciliano (4)19(suppl.): 753-772.
 * Menozzi C. 1918. Primo contributo alla conoscenza della fauna mirmecologica del Modenese. Atti della Società dei Naturalisti e Matematici di Modena. (5)4: 81-88.
 * Menozzi C. 1921. Formiche dei dintorni di Sambiase di Calabria. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici. 15: 24-32.
 * Morillo C. 1981. Inventario y fenologia de los Formicidae de un encinar de Espana central. Rev. Ecol. Biol. Sol. 18(2): 243-251.
 * Müller, G.. "Faunisticna istrazivanja sjeverodalmatinskih otoka: Dugi otok i Kornati (1925-1927): Orthopteroidea, Coleoptera i Formicidae." Acta biol. 1 (1957): 187-218.
 * Müller, G.. "Le formiche della Venezia Guilia e della Dalmazia." Bollettino della Società Adriatica di Scienze Naturali in Trieste 28 (1923): 11-180.
 * Nylander, W.. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * Obregon R., J. Lopez, J. L. Reyes-Lopez. 2014. A catalogue of the ant fauna (Hymenoptera, Formicidae) of Sierra Mágina (Jaén, southern Spain). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.), 54: 370374.
 * Ortiz, F. J.. "Formícidos del litoral granadino." Memoria de Licenciatura Universida (1985): 206 pp.
 * Passera L. 1967. Peuplement en fourmis terricoles du rebord Meridional des Causses jurassiques du Quercy: la lande calcaire a buis. Vie et milieu. Serie C, Biologie terrestre 18(1): 189-205.
 * Passera L. 1977. Peuplement myrmecologique du cordon littoral du Languedoc-Roussillon modifications anthropiques. Vie et Milieu 27: 249-265.
 * Petrov I. Z. 2004. A list of currently known ant species (Formicidae, Hymenoptera) of Serbia. Arch. Biol. Sci., Belgrade 56(3-4): 121-125.
 * Petrov I. Z. 2007. Contribution to knowledge of the myrmecofauna (Formicidae, Hymenoptera) of Belgrade, Serbia. Archives of Biological Sciences 59: 11-12.
 * Petrov I. Z., and A. Legakis. 1996. Contribution to the Myrmecofauna (Formicidae, Hymenoptera) of Greece. Archives of biological sciences, Belgrade, 48 (3-4): 31-32.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Poldi B., M. Mei, and F. Rigato. 1995. Hymenoptera, Formicidae. Vol 102. Checklist delle specie della fauna Italiana: 1-10.
 * Retana J., and X. Cerdá X. 2000. Patterns of diversity and composition of Mediterranean ground ant communites tracking spatial and temporal variability in the thermal environment. Oecologia 123: 436-444.
 * Rigato S., and I. Toni. 2011. Short notes 21. Hymenoptera, Formicidae. Pp. 873-882 in: Nardi, G.; Whitmore, D.; Bardiani, M.; Birtele, D.; Mason, F.; Spada, L.; Cerretti, P. (eds.) 2011. Biodiversity of Marganai and Montimannu (Sardinia). Research in the framework of the ICP Forests network. Conservazione Habitat Invertebrati, 5. Sommacampagna, Verona: Cierre Edizioni, 896 pp.
 * Rodríguez, A.. "Contribución al conocimiento de las hormigas (Hymenoptera, Formicidae) de Sierra Morena Central." Boletín de la Asociación Española de Entomología 5 (1982): 181-188.
 * Roig X., X. Espadaler, R. Cusco, F. Garcia, K. Gomez, S. Serrano, and R. Vila. 2008. Ants in gypsum areas. First record from Catalonia (Iberian Peninsula) of Cardiocondyla batesii Forel (Hymenoptera, Formicidae) with an updated list of Catalan Formicidae. Boletin Sociedad Entomologica Aragonesa 42: 189-192.
 * Sanchez-Gil Jimeno R., and J. L. Reyes-Lopez. 2016. Study of ants species of the Sierra de San Carlos del Valle (Ciudad Real) and updating the provincial check list (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent. 40 (1-2): 93-109.
 * Santschi F. 1925. Fourmis d'Espagne et autres espéces paléartiques EOS (Revista española de entomología) 1: 339-360.
 * Santschi F. 1926. Travaux scientifiques de l'Armée d'Orient (1916-1918). Fourmis. Bulletin du Muséum National d'Histoire Naturelle 32: 286-293.
 * Santschi, F.. "Fourmis du Portugal." Memórias e Estudos do Museu Zoológico da Universidade de Coimbra 59 (1932): 1-3.
 * Santschi, F.. "Inventa entomologica itineris Hispanici et Maroccani, quod a. 1926 fecerunt Harald et Håkan Lindberg. Fourmis du Bassin Méditerranéen occidental et du Maroc récoltées par MM. Lindberg." Societas Scientiarum Fennica (Helsingfors) 3 (14) (1931): 1-13.
 * Santschi, F.. "Messor et autres fourmis paléarctiques." Revue Suisse de Zoologie 30 (1923): 317-336.
 * Schembri, Stephen P. and Cedric A. Collingwood. A Revision of the Myrmecofauna of the Maltese Islands. 417-442.
 * Scupola A., and R. Poggi. 2014. Ricerche zoologiche della nave oceanografica "minerva" (C.N.R.) sulle isole circumsarde. XXXIII. Formiche. Estratto dagli Annali del Museo Civico di Storia Naturale “G. Doria” 106: 93-131.
 * Scupola I. 2009. Le formiche delle Isole Egadi (Sicilia) (Hymenoptera, Formicidae). Bollettino del Museo Civico di Storia Naturale di Verona Botanica Zoologia 33: 97-103
 * Sommer F. 1984. Etude de la myrmécofaune de la réserve de la Massane. Laboratoire Arago, Banyuls sur Mer (66), 29 pages.
 * Sommer F., and H. Cagniant. 1988. Peuplements de fourmis des Albères Orientales (Pyrénées-Orientales, France) (Première partie). Vie Milieu 38: 189-200.
 * Suñer i Escriche, David. "Contribució al coneixement mirmecologic de Gavarres, Montgrí, Guilleríes i la Serralada Transversal." Tesis Doctoral Universida (1991): 577 pp.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * Tinaut A. 2016. Wilhelm Gottlieb Rosenhauer (1813-1881), Maximilien Spinola (1780- 1857) and the first records of ants from Andalusia (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 161-174
 * Tinaut A., and F. J. Ortiz. 1988. Introduccion al conocimiento de las hormigas de la provincia de Almeria (Hymenoptera: Formicidae). Publicaciones del Instituto de Estudios Almerienses. Boletín (Ciencias) 8: 223-231.
 * Tinaut, A., J. Jiménez Rojas, and R. Pascual. "Estudio de la mirmecofauna de los bosques de Quercus Linneo 1753 de la provincia de Granada." Ecología 8 (1995): 429-438.
 * Tolg, F., and J. Fahringer. "Beitrag zur Dipteren- und Hymenopterenfauna Bosniens der Herzegowina und Dalmatiens, II." Mitt. Naturw. Ver. Univ. Wien 9 (1911): 23-28.
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vesnic A., and R. Skrijelj. 2014. Biodiversity and variation in species of the genus Messor Forel, 1890 (Hymenoptera, Formicidae) in the Mediterranean part of Bosnia and Herzegovina. Works of the Faculty of Forestry University of Sarajevo 1: 37-44.
 * Vogrin, V.. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Zimmermann, S.. "Beitrag zur Kenntnis der Ameisenfauna Süddalmatiens." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 84 (1935): 1-65.
 * Zimmermann, S.. "Beitrag zur Kenntnis der Ameisenfaunader Quarnerischen Inseln." Verhandlungen der Zoologisch-Botanisch Gesellschaft in Wien 80 (1930): 45-66.
 * de Haro, Andrés, C. A. Collingwood, and P. Comín del Río. "Prospección mirmecológica por Ibiza y Formentera (Baleares)." Orsis 2 (1986): 115-120.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica por la península Tingitana al norte del Rif (Marruecos)." Orsis (Organismes i Sistemes) 12 (1997): 93-99.
 * de Haro, Andrés. "Contribución al conocimiento de los formícidos del cabo de Creus (Gerona)." Pirineos 117 (1982): 49-57.