Odontomachus rixosus

Odontomachus rixosus inhabits plantations, secondary and primary forests in lowlands, and nests under leaf litter, in the soil near the base of living trees, and under rotten logs and stumps. It is a A widely distributed forest species. This species has been collected nesting together with Pheidole tandjongensis. Details concerning the nature of the relationship between O. rixosus and Pheidole spp. are unknown. (Satria et al. 2015, Sorger & Zettel 2011)

Identification
A member of the Odontomachus rixosus group.

Satria et al. (2015) - In the syntypes of O. rixosus and O. rixosus conifer the pronotal disc is finely striate transversely, but finely concentric in O. rixosus obscurior.

Odontomachus rixosus is very similar in general appearance to Odontomachus pararixosus, but is distinguishable from the latter by the weakly notched posterior head margin (almost straight in the latter); the apical spine of the petiolar node is needle-shaped (acutely triangular in the latter); and the first gastral tergite lacks long suberect setae (present in the latter). The delimitation between O. rixosus and Odontomachus minangkabau is discussed under O. minangkabau.

Sorger & Zettel (2011) - There is considerable variation in O. rixosus, especially in size and colour, but it seems unconnected with certain geographical populations. This includes the var. obscurior, described from Myanmar and Thailand (Forel 1900), and later recorded from Palawan by Stitz (1925). In the Philippines, O. rixosus can be recognized by group characters, especially by the elongate and truncate subapical tooth of the mandible.

Distribution
Sorger & Zettel (2011) - Southeast Asian mainland (Myanmar, Thailand, Peninsular Malaysia, Singapore), Sumatra, Mentawai Islands, Java, Borneo, Philippines.

Distribution based on Regional Taxon Lists
Indo-Australian Region: Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore. Oriental Region: Bangladesh, Cambodia, India, Laos, Myanmar, Thailand, Vietnam. Palaearctic Region: China.

Biology
Polygynous (Ito et al. 1996) with multiple dealate queens.

Sorger & Zettel (2011) - The only encounter in the Philippines by the senior author was at a waterfall area in a slightly degraded dipterocarp forest. Observations from Borneo confirm that O. rixosus is a forest species.

Association with Other Organisms
Heraty et al. (2015) - Odontomachus rixosus is parasitized by the eucharitid wasp.

Nomenclature

 * . Odontomachus rixosus Smith, F. 1857a: 64 (w.) SINGAPORE.
 * Type-material: 2 syntype workers.
 * Type-locality: Singapore, “SING”, and “SING. 55/9 (A.R. Wallace).
 * [Note: one specimen in OXUM (“SING”), and one in BMNH (“SING. 55/9”). In the original description of rixosus, Smith states that the thorax is “tranversely striated.” The BMNH specimen more or less fits this description, whereas the OXUM specimen has weakly whorled, not transverse, striation on the pronotum.]
 * Type-depositories: BMNH, OXUM.
 * Crawley, 1924: 388 (q.); Wheeler, G.C. & Wheeler, J. 1952c: 651 (l.); Imai, et al. 1984: 67 (k.); Satria, et al. 2015: 27 (m.).
 * Status as species: Smith, F. 1858b: 79; Roger, 1861a: 27; Smith, F. 1861b: 44; Roger, 1863b: 21; Mayr, 1863: 437; Smith, F. 1871a: 319; Mayr, 1872: 148; Mayr, 1886c: 358; Emery, 1887b: 428; André, 1892b: 52; Emery, 1892d: 560 (in key); Dalla Torre, 1893: 52; Emery, 1893g: 261; Emery, 1895k: 463; Forel, 1900c: 58; Emery, 1900d: 672; Emery, 1901g: 566; Bingham, 1903: 48; Forel, 1905c: 8; Emery, 1911d: 114; Forel, 1911b: 193; Forel, 1911d: 382; Forel, 1912d: 97; Forel, 1913k: 19; Forel, 1915a: 22; Viehmeyer, 1916a: 116; Wheeler, W.M. 1919e: 60; Crawley, 1924: 388; Stitz, 1925: 115; Karavaiev, 1925c: 293; Wheeler, W.M. 1927h: 86; Santschi, 1928h: 124; Stitz, 1932b: 733; Donisthorpe, 1932c: 447; Karavaiev, 1935a: 75; Chapman & Capco, 1951: 45; Brown, 1976a: 105, 163; Bolton, 1995b: 297; Mathew & Tiwari, 2000: 290; Jaitrong & Nabhitabhata, 2005: 29; Zhou & Ran, 2010: 107; Pfeiffer, et al. 2011: 56; Sorger & Zettel, 2011: 157 (redescription); Guénard & Dunn, 2012: 60; Satria, et al. 2015: 21 (redescription); Bharti, Guénard, et al. 2016: 53; Jaitrong, Guénard, et al. 2016: 42; Chen, Shi & Zhou, 2018: 6 (in key).
 * Senior synonym of conifera: Brown, 1976a: 105; Bolton, 1995b: 297; Satria, et al. 2015: 21.
 * Senior synonym of obscurior: Brown, 1976a: 105; Bolton, 1995b: 297; Sorger & Zettel, 2011: 157; Satria, et al. 2015: 21.
 * Distribution: China, India, Indonesia (Bali, Java, Mentawei, Sumatra), Laos, Malaysia (Peninsula, Sabah, Sarawak), Myanmar, Philippines (Mindanao, Palawan), Singapore, Thailand.
 * conifera. Odontomachus rixosus var. conifera Forel, 1913k: 19 (w.q.) INDONESIA (Java).
 * Type-material: syntype workers, syntype queens (numbers not stated).
 * Type-locality: Indonesia: Java, Buitenzorg (= Bogor) u. Tjiogrek (v. Buttel-Reepen).
 * Type-depository: MHNG.
 * Subspecies of rixosus: Chapman & Capco, 1951: 45.
 * Junior synonym of rixosus: Brown, 1976a: 105; Bolton, 1995b: 295; Satria, et al. 2015: 21.
 * obscurior. Odontomachus rixosus var. obscurior Forel, 1900c: 58 (w.) THAILAND, MYANMAR.
 * Type-material: syntype workers (number not stated).
 * Type-localities: Thailand: frontier (Fulton), and Bangkok (Sigg), Myanmar (“Burma”) (no further data) (Bingham).
 * Type-depository: MHNG.
 * Viehmeyer, 1916a: 116 (q.).
 * Subspecies of rixosus: Bingham, 1903: 48; Emery, 1911d: 114; Viehmeyer, 1916a: 116; Stitz, 1925: 115; Chapman & Capco, 1951: 45.
 * Junior synonym of rixosus: Brown, 1976a: 105; Bolton, 1995b: 296; Sorger & Zettel, 2011: 157; Satria, et al. 2015: 21.

Smith Types
Odontomachus rixosus

The specimen at labelled as “type” by Donisthorpe may not be so. One cannot be positive as there is no datum number on the disc of this specimen, which merely states “SING” (= Singapore). However, a specimen in, labelled “SING. 55/9” has the following information in Acc. Reg.: “1859 no. 9. Singapore (Malacca), purchased of Stevens. Collected by Mr A.R. Wallace.”

I suspect that this specimen may be the true holotype but have no way of conclusively proving it. The only evidence in support of this view is that in the original description of rixosus, Smith states that the thorax is “tranversely striated.” The specimen more or less fits this description, whereas the  specimen has weakly whorled, not transverse, striation on the pronotum.

Worker
Sorger & Zettel (2011) - Worker with smallest HW: CI 71, HL 2.38, HW 1.68, MdI 57, MdL 1.37, MsL 3.33, PnW 1.05, PtH 0.78, PtL 0.77, PtW 0.42, SI 145, SL 2.43, TL 10.31; worker with largest HW: CI 76, HL 2.57, HW 1.95, MdI 56, MdL 1.43, MsL 3.62, PnW 1.17, PtH 0.84, PtL 0.92, PtW 0.48, SI 131, SL 2.63, TL 12.94.

Structures: Mandibles long, with ca. 6 basal denticles (widely separated from each) and three apical teeth: proximate tooth truncated, intercalary tooth only slightly shorter than apical. Head rectangular, broadest at level of eyes. Striation on head until ocular ridge, some more striation between ocular and temporal ridge, rest of head smooth and shiny. Microsculpture on head with fine isodiametric reticulum. Mesosoma elongate, slender and low, broadest at level of pronotum. Pronotum rounded, metanotal groove in lateral view present. Coarse rounded sculpture on pronotum (closed circles visible in dorsal view), metanotum and propodeum with coarse transverse sculpture. Petiole short, smooth and shiny, almost conical, with very short petiolar spine, which is rarely absent in small specimens (for example, O. rixosus var. conifera Forel, 1913).

Pilosity: Fine loose semi-appressed white pubescence on head, mesosoma and petiole; distance between hairs approximately their length. Head with two standing setae, tergite 1 without setae, tergite 2 with a few setae, number of setae and length increasing towards apex of abdomen.

Colour: Medium brown, mesosoma slightly darker than head, petiole and gaster.

Satria et al. (2015) - Syntype (n=1, gaster missing): HW 2.10 mm, HL 2.98 mm, SL 2.93 mm, IFLW 0.64 mm, EL 0.36 mm, MDL 1.64 mm, WL 4.00 mm, PTL 0.72 mm, PTH 1.21 mm, CI 70, SI 139, MDI 55, PTHI 168.

Non-types (n=10): HW 1.77–2.19 mm, HL 2.56–3.03 mm, SL 2.56–3.03 mm, IFLW 0.45–0.56 mm, EL 0.31–0.42 mm, MDL 1.45–1.67 mm, WL 3.35–4.00 mm, PTL 0.57–0.69 mm, PTH 1.09–1.26 mm, CI 64–73, SI 131–150, MDI 55–57, PTHI 171–196.

Relatively small (HL 2.56–3.03 mm, WL 3.35–4.00 mm). Head in full-face view much longer than broad, with posterior margin weakly concave; median furrow on vertex present as dark line; each side of line not swollen; frontal lobes followed by strong frontal carinae which are nearly parallel; minimum distance between margin of ocular ridge and margin of compound eye less than half major axis of compound eye; mandible relatively slender; masticatory margin with 6–10 denticles (7 in a syntype of O. rixosus examined; 7 in a syntype of O. rixosus conifer [ANTWEB CASENT 0907432]; 6 in a syntype of O. rixosus obscurior [ANTWEB CASENT 0907431]); subapical tooth ca. 2.5 times as long as broad, with truncate apex; palp formula 4, 4. Mesosoma in lateral view relatively slender; pronotum including its anteromedian lobe long, in lateral view with anterodorsal slope gentle; mesopleuron without anteroventral ridge, with anterodorsal margin weakly carinate, clearly separated by distinct dorsal carina from mesonotum and metapleuron; propodeum in lateral view with dorsum almost straight and gradually sloping posteriad, and with posterior face steeply sloping; propodeal dorsum without median longitudinal depression. Petiolar node conical, with sharply pointed apical spine; node in lateral view, excluding apical spine with anterior and posterior faces weakly convex; apical spine needle-shaped, less than 1/4 as long as petiolar height, sometimes weakly curved posteriad (but shape variable within species); subpetiolar process anteroposteriorly as long as dorsoventrally high, lobate, directed ventrally. First gastral tergum in lateral view long, with anterior face relatively short and vertical.

Head in full-face view largely smooth and shiny, but with striate area bordered by frontal carinae and ocular ridges; frontal lobe with almost smooth and shiny; extraocular furrow faintly striate; median part of vertex along median furrow smooth and shiny or sometimes with rough texture; lateral face and venter of head smooth and shiny; median disc of clypeus smooth and shiny. Pronotum in dorsal view finely concentrically striate (but sculpture variable within species); mesonotum finely striate transversely; mesopleuron largely smooth and shiny, with anterior 1/3 and posteriormost parts finely striate; metapleuron and propodeum with transverse striation which is a little sparser and stronger than on pronotum and mesonotum. Petiolar node largely smooth and shiny, but with its basal area faintly striate.

Vertex with a pair of long erect setae; frontal lobe with erect seta; pronotal disc without long erect setae; 1st gastral tergum without erect setae. Head, mesosoma, petiole and gaster with fine appressed pubescence which is sparse and very short on head and petiole and is sparse and extremely short on gaster.

Body orange-brown, with darker mesosoma and gaster; leg yellowish brown to orange-brown, with coxae and femora paler.

Queen
Satria et al. (2015) - (n=10): HW 1.87–2.17 mm, HL 2.53–2.96 mm, SL 2.49–2.76 mm, IFLW 0.42–0.57 mm, EL 0.39–0.50 mm, OL 0.12–0.18 mm, MDL 1.45–1.63 mm, WL 3.50–4.05 mm, FWL 6.46–8.33 mm, PTL 0.65–0.70 mm, PTH 1.40–1.56 mm, CI 72–76, SI 124–136, MDI 52–58, PTHI 214–229.

In general appearance queen is similar to worker. Vertex near ocelli not swollen; ocular ridge clearly developed; distance between lateral ocelli shorter than or as long as distance between lateral and median ocelli, and as long as major axis of median ocellus; ocelli in lateral view protruded dorsad. Mesosoma with main sclerites associated with wing function, in dorsal view long and slender; anterodorsal slope of pronotum in lateral view gentle; anterodorsal outline of mesoscutum in lateral view relatively gentle; mesoscutum without posteromedian depression; parapsidal furrow very weak, and slightly curved; anterior third of mesopleuron with fine, oblique furrow (sometimes without any furrow); propodeum in lateral view long, with dorsum almost straight and gradually sloping posteriad. Wing venation as in Figs. 12E and 12F. Petiolar node in lateral view, excluding apical spine with anterior and posterior faces weakly convex; apical spine short and stout, and sometimes weakly curved posteriad; subpetiolar process anteroposteriorly as long as dorsoventrally high. First gastral tergum in lateral view long, with anterior face relatively short and vertical.

Head in full-face view largely smooth and shiny, but with a striate area bordered by frontal carinae and ocular ridges; frontal lobe almost smooth and shiny; extraocular furrow faintly striate; median part of vertex along median furrow smooth and shiny or sometimes with rough texture; lateral face and venter of head smooth and shiny; median disc of clypeus smooth and shiny. Pronotum densely and weakly striate transversely (often dorsolateral part smooth and shiny); mesoscutum faintly longitudinally striate (but sometime largely smooth and shiny, and faintly striate along posterior margin of mesoscutum); mesopleuron largely smooth and shiny, with anterior third and posteriormost part faintly striate; mesoscutellum smooth and shiny; propodeum strongly and sparsely striate transversely. Petiolar node largely smooth and shiny, with its basal area faintly striate.

Pair of long erect setae present on vertex near lateral ocelli; frontal lobe with a erect seta; pronotum and 1st gastral tergum without erect setae. Head, mesosoma, and petiole with sparse subdecumbent to decumbent pubescence which is extremely short in head and gaster; mesopleuron very sparsely pubescent.

Male
Satria et al. (2015) - (n=10): HW 1.19–1.35 mm, HL 1.04–1.14 mm, SL 0.20–0.26 mm, EL 0.70–0.84 mm, EW 0.37–0.46 mm, OL 0.16–0.22 mm, OES 0.16–0.18 mm, WL 2.63–2.93 mm, FWL 4.52–5.58 mm, PTL 0.56–0.64 mm, PTH 0.78–0.90 mm, CI 107–120, SI 15–20, PTHI 135–152.

Size small (HL 1.04–1.14 mm, WL 2.63–2.93 mm). Major axis of median ocellus as long as minimum distance between lateral ocelli; antenna 13-merous; scape very short; 1/3 as long as 3rd antennomere; 2nd antennomere 1/2 as long as scape; 3rd to 13th antennomeres each extremely long; palp formula 6, 4; dorsal outline of clypeus in lateral view weakly convex. Mesosoma in lateral view relatively slender and long; dorsal outline of pronotum in lateral view almost straight; anterodorsal outline of mesoscutum in lateral view gently sloping; mesoscutum without median depression; parapsidal furrow very weak and almost straight; oblique mesopleural furrow relatively shallow and wide; ventrolateral part of katepisternum with very weak longitudinal furrow; propodeum in lateral view with its dorsal outline roundly convex; metapleuron separated from propodeum indistinctly by suture; wing venation similar to queen (see Figs. 12E and 12F for queen wings). Petiolar node in lateral view tapering gently to bluntly pointed apex; its anterior slopes weakly convex, and posterior slope almost straight; subpetiolar process in lateral view anteroposteriorly shorter than dorsoventrally high, lobate and slender; petiolar sternum without posteroventral process. First gastral tergum in lateral view short; posterior spine of 8th abdominal tergum long and slender, very weakly curved (Fig. 18B); pygostyle with long setae in its apical third; disc of 9th abdominal sternum much broader than long, with posterolateral corner expanding laterad; apical lobe much longer than disc and gently tapering to almost truncate apex; telomeral apex in lateral view longer than high; distiventral apex of valviceps strongly produced; basiventral corner of valviceps distinctly produced; ventral margin of valviceps with 21−22 denticles.

Head including area between lateral ocelli entirely smooth and shiny. Pronotum in dorsal view smooth and shiny; mesoscutum smooth and shiny; scuto-scutellar suture with sparse, strong, longitudinal rugae; mesopleural anepisternum and katepisternum smooth and shiny; metapleuron smooth and shiny; propodeum in dorsal view with rough texture and shiny. Petiole smooth and shiny.

Head, mesosoma, legs, petiole and gaster with fine sparse subdecumbent to decumbent pubescence; mandible, vertex near ocelli, and gaster except 1st gastral tergum with long erect setae.

Males from Thailand are always darker than those from the other localities.

References based on Global Ant Biodiversity Informatics

 * André E. 1892. Voyage de M. Chaper à Bornéo. Catalogue des fourmis et description des espèces nouvelles. Mém. Soc. Zool. Fr. 5: 46-55.
 * Berghoff S.M., U. Maschwitz, and K.E. Linsemair. 2003. Hypogaeic and epigaeic ant diversity on Borneo: evaluation of baited sieve buckets as a study method. Tropical Zoology 16: 153-163.
 * Bharti H., Y. P. Sharma, M. Bharti, and M. Pfeiffer. 2013. Ant species richness, endemicity and functional groups, along an elevational gradient in the Himalayas. Asian Myrmecology 5: 79-101.
 * Bolton, B. 1998. A preliminary analysis of the ants of the Pasoh Forest Reserve. Pp. 84-95 in: Lee, S. S.; Dan, Y. M.; Gauld, I. D.; Bishop, J. (eds.) Conservation, management and development of forest resources. Proceedings of the Malaysia-United Kingdom Programme Workshop 21-24 October 1996. Kuala Lumpur: International Institute for Environment and Development, 392 pp.: 84-95
 * Brown W. L., Jr. 1976. Contributions toward a reclassification of the Formicidae. Part VI. Ponerinae, tribe Ponerini, subtribe Odontomachiti. Section A. Introduction, subtribal characters. Genus Odontomachus. Stud. Entomol. 19: 67-171.
 * Bruhl C.A. 2001. Leaf litter ant communities in tropical lowland rain forests in Sabah, Malaysia: effects of forest disturbance and fragmentation. PHD thesis Wurzburg Universitat, 168 pp.
 * CSIRO Collection
 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Chung A.Y.C., and M. Mohamed. 1996. A comparative study of the ant fauna in a primary and secondary forest in Sabah, Malaysia. In Edwards, D.S. Booth, W.E. and S.C. Choy (eds). Tropical Rainforest Research. Kluwer Academic Publishers. Pp 357-366.
 * Crawley W.C. 1924. Ants from Sumatra, with biological notes by Edward Jacobson. Annals and Magazine of Natural History (9)13: 380-409
 * Eguchi K., B. T. Viet, and S. Yamane. 2014. Generic Synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part IICerapachyinae, Aenictinae, Dorylinae, Leptanillinae, Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae. Zootaxa 3860: 001-046.
 * Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). [concl.]. Ann. Mus. Civ. Stor. Nat. 25(5): 427-473.
 * Emery C. 1892. Voyage de M. Ch. Alluaud dans le territoire d'Assinie (Afrique occidentale) en juillet et août 1886. Formicides. Annales de la Société Entomologique de France 60: 553-574.
 * Emery C. 1901. Formiciden von Celebes. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 14:565-580.
 * Emery C. 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1-125.
 * Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine)." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 5, no. 25 (1887): 427-473.
 * Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
 * Fayle T. M., E. C. Turner, J. L. Snaddon, V. Khen Chey, A. Y. C. Chung, P. Eggleton, and W. A. Foster. 2010. Oil palm expansion into rain forest greatly reduces ant biodiversity in canopy, epiphytes and leaf-litter. Basic and Applied Ecology 11: 337345.
 * Forel A. 1900. Les Formicides de l'Empire des Indes et de Ceylan. Part VI. J. Bombay Nat. Hist. Soc. 13: 52-65.
 * Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
 * Forel A. 1911. Ameisen aus Java beobachtet und gesammelt von Herrn Edward Jacobson. II. Theil. Notes Leyden Mus. 33: 193-218.
 * Forel A. 1912. Ameisen aus Java beobachtet und gesammelt von Edward Jacobson. III. Theil. Notes Leyden Mus. 34: 97-112
 * Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
 * Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
 * Gillison A.N. 2000. Above ground biodiversity assesment working group summary report 1996-99: Impact of different land uses on biodiversity and social indicators. ASB Working Group Report, ICRAF, Nairobi, 160pp. http://www.asb.cgiar.org/PDFwebdocs/ASB Biodiversity Report.pdf
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Helms J. A., S. M. Helms, N. I. Fawzi, Tarjudin, F. Xaverius. 2017. Ant community of an Acacia mangium forest in Indonesian Borneo. Serangga 22(1): 147-159.
 * Herwina H., and K. Nakamura. 2007. Ant species diversity study using pitfall traps in a small yard in Bogor Botanic garden, West Java, Indonesia. Treubia 35: 99-116.
 * IZIKO South Africa Museum Collection
 * Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
 * Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
 * Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
 * Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
 * Karavaiev V. 1925. Ponerinen (Fam. Formicidae) aus dem Indo-Australischen Gebiet. (Schluss). Konowia 4: 276-296.
 * Karavaiev V. 1926. Ameisen aus dem Indo-Australischen Gebiet. Treubia 8: 413-445.
 * Khoo Y.H. 1990. A note on the Formicidae (Hymenoptera) from pitfall traps at Ulu Kinchin, Pahang, Malaysia. Malayan Nature Journal 43: 290-293.
 * Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
 * Mohamed M. 1995. A preliminary list of ants (Hymenoptera: Formicidae) of the Tawau Hills Park, Sabah. In: Ghazally Ismail  et al.  (eds.),  Tawau  Hills Park,  Sabah,  Pelanduk Pub.  Pp.  205-213.
 * Pfeiffer M., D. Mezger, and J. Dyckmans. 2013. Trophic ecology of tropical leaf litter ants (Hymenoptera: Formicidae) - a stable isotope study in four types of Bornean rain forest. Myrmecological News 19: 31-41.
 * Pfeiffer M., and D. Mezger. 2012. Biodiversity Assessment in Incomplete Inventories: Leaf Litter Ant Communities in Several Types of Bornean Rain Forest. PLoS ONE 7(7): e40729. doi:10.1371/journal.pone.0040771
 * Pfeiffer M., and D. Mezger. 2012. Biodiversity Assessment in Incomplete Inventories: Leaf Litter Ant Communities in Several Types of Bornean Rain Forest. PLoS ONE 7(7): e40729. doi:10.1371/journal.pone.0040987
 * Pfeiffer M., and D. Mezger. 2012. Biodiversity Assessment in Incomplete Inventories: Leaf Litter Ant Communities in Several Types of Bornean Rain Forest. PLoS ONE 7(7): e40729. doi:10.1371/journal.pone.0041093
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
 * Philpott S.M., P. Bichier, R.A. Rice, and R. Greenberg. 2008. Biodiversity conservation, yield, and alternative products in coffee agroecosystems in Sumatra, Indonesia. Biodivers. Conserv. 17: 1805-1820. Data obtained from Stacy Philpott
 * Rizali A., M. M. Bos, D. Buchori, Sk. Yamane, and C. H. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 15(2): 77-84.
 * Rizali A., M.M. Bos, D. Buchori, Sk. Yamane, C. Hans, and J. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 77-84.
 * Sakchoowong W., W. Jaitrong, and K. Ogata. 2009. Comparison of ground-ant diversity between natural forests and disturbed forests along a natural gas pipeline transect in Thong Pha Phum National Park, Kanchanaburi province. Kasetsart J. (Nat. Sci) 43: 64-73.
 * Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
 * Satria R., H. Kurushima, H. Herwina, S. Yamane, and K. Eguchi. 2015. The trap-jaw ant genus Odontomachus Latreille (Hymenoptera: Formicidae) from Sumatra, with a new species description. Zootaxa 4048(1): 001-036.
 * Smith F. 1861. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Ceram, Celebes, Ternate, and Gilolo. [part]. Journal and Proceedings of the Linnean Society of London. Zoology 6: 36-48.
 * Song Y., Z. Xu, C. Li, N. Zhang, L. Zhang, H. Jiang, and F. Mo. 2013. An Analysis on the Ant Fauna of the Nangun river Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
 * Sorger, D.M. and H. Zettel. 2011. On the ants (Hymenoptera: Formicidae) of the Philippine Islands: V. The genus Odontomachus LATREILLE, 1804. Myrmecological News. 14:141-163.
 * Stitz H. 1925. Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1923: 110-136.
 * Stitz H. 1932. Formicidae der Deutschen Limnologischen Sunda-Expedition. Arch. Hydrobiol. Suppl.-Bd. 9(Tropische Binnengewässer 2 2: 733-737
 * Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
 * Watanasit S., Noon-anant N., and N. Binnima. 2005. Preliminary survey of ants at a reserve area of Prince of Songkla University, Songkhla Provinces, Southern Thailand. Songklanakarin J. Sci. Technol. 27(1): 39-46.
 * Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
 * Wheeler W. M. 1927. Ants collected by Professor F. Silvestri in Indochina. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 20: 83-106.
 * Woodcock P., D. P. Edwards, R. J. Newton, C. Vun Khen, S. H. Bottrell, and K. C. Hamer. 2013. Impacts of Intensive Logging on the Trophic Organisation of Ant Communities in a Biodiversity Hotspot. PLoS ONE 8(4): e60756. doi:10.1371/journal.pone.0060756
 * Woodcock P., D. P. Edwards, T. M. Fayle, R. J. Newton, C. Vun Khen, S. H. Bottrell, and K. C. Hamer. 2011. The conservation value of South East Asia's highly degraded forests: evidence from leaf-litter ants. Phil. Trans. R. Soc. B. 366: 3256-3264.
 * Wu W. 2010. The taxonomic and fauna study on the Formicidae of Liaoning Province (Insecta: Hymenoptera). Master's Thesis Northeast Normal University, 75 pages.
 * Zhang N. N., Y. Q. Chen, Z. X. Lu, W. Zhang, and K. L. Li. 2013. Species diversity, community structure difference and indicator species of leaf-litter ants in rubber plantations and secondary natural forests in Yunnan, southwestern China. Acta Entomologica Sinica 56(11): 1314-1323.
 * Zhang Z., M. Cao, X. Yang, X. Deng, and Y. She. 2000. A study on species diversity of ant in fragments of seasonal rain forest of Xishuangbanna, China. Zoological Research 21(1): 70-75.
 * Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.