Tetramorium ferox

Very little is known about the biology of T. ferox, other than it is a host for the inquilines and.

Identification
Csösz and Schulz (2010) - A member of the Tetramorium ferox species complex in the Tetramorium caespitum species group. The feebly rugulose general sculpture, the microreticulate ground surface, and the relatively sparse pubescence on the first gastral tergite make the T. ferox gynes unique within this group.

Tetramorium ferox workers cannot be confused with those of Tetramorium diomedeum or Tetramorium aegeum. The general appearance of the head and mesosoma of T. ferox workers is always rugulose, microreticulate and dull, but head and mesosoma of the latter two species are always shiny and usually at least partly smooth.

Tetramorium feroxoides workers usually bear much feebler (and often parallel) rugulae on head and mesosoma than T. ferox workers, but in a few cases T. feroxoides workers can be confused with the smallest T. ferox workers.

A discriminant function using various morphological measurements has also been defined and can be used to make determinations in this species group.

Distribution
Widely distributed over the western Palaearctic, from Central Europe to the Caucasus and Turkey. (Csösz and Schulz 2010)

Distribution based on Regional Taxon Lists
Palaearctic Region: Armenia, Azerbaijan, Bulgaria, China, Croatia, Czech Republic, Georgia, Greece, Hungary, Iran, Kyrgyzstan, Malta, Montenegro, Republic of Macedonia, Romania, Russian Federation, Turkmenistan.

Nomenclature

 *  ferox. Tetramorium caespitum var. ferox Ruzsky, 1903b: 309 (w.q.m.) RUSSIA. Subspecies of caespitum: Emery, 1909d: 703; Emery, 1916b: 194. Raised to species: Cori & Finzi, 1931: 239; Soudek, 1931: 10; Menozzi, 1933b: 75 (in key); Kratochvíl, 1944: 72; Tarbinsky, 1976: 112; Atanassov & Dlussky, 1992: 155. Senior synonym of silhavyi: Bernard, 1967: 235; Dlussky, Soyunov & Zabelin, 1990: 201; Radchenko, 1992b: 55; of confinis, perspicax: Csösz & Schulz, 2010: 20; material of the unavailable name striaticeps referred here by Radchenko, 1992b: 55.
 * silhavyi. Tetramorium (Lobomyrmex) ferox subsp. silhavyi Kratochvíl, in Novak & Sadil, 1941: 84, fig. 1 (w.q.) CZECHOSLOVAKIA. [Also described as new by Kratochvíl, in Kratochvíl, Novak & Snoflak, 1944: 72.] Junior synonym of ferox: Bernard, 1967: 235; Radchenko, 1992b: 55.
 * confinis. Tetramorium confinis Radchenko & Arakelian, 1990: 374, fig. 2 (w.q.m.) ARMENIA. Junior synonym of ferox: Csösz & Schulz, 2010: 20.
 * perspicax. Tetramorium perspicax Radchenko, 1992b: 56 (w.q.) TURKEY. [First available use of Tetramorium caespitum st. ferox var. perspicax Santschi, 1921a: 111; unavailable name.] Synonym of feroxoides: Radchenko, 1992b: 56. [Radchenko gives perspicax as senior name, but feroxoides has priority: Bolton, 1995b: 412.] Junior synonym of ferox: Csösz & Schulz, 2010: 21.

Type Material
Csösz and Schulz (2010) - Syntype workers, gynes, male (2w, 2q, 1m): Russia, Saratov (Ruzsky) [labels: ''Tetr. caespitum L., v. ferox'' Rusz, Ssaratov M.R.] (examined),.

Bezděčková et al. (2017) - Tetramorium ferox silhavyi: Five syntypes are deposited in (all dry-mounted):
 * SYNTYPES (3 males 1 alate queen each glued on a small rectangular label and all together on one larger card label ca. 40 × 16 mm): ‘Tetramorium / ferrox [sic] / Šilhavýi. Krat. [hw] // Tetramorium ferrox [sic] / worker šilhavýi Krat. / Mohelno, cotypy, 05062 [hw] // Syntypus / (Bezděčková et al. 2017) [p, red label]’ [IN: E26-T10–E26-T13].
 * SYNTYPE (1 dealate queen, only head with one antenna and thorax with one leg glued on a card label ca. 40 × 16 mm): ‘Tetramorium / ferrox [sic] / šilhavýi Krat. [hw] // Tetramorium ferrox [sic]  / šilhavýi Krat. / Mohelno – paratyp, 05063 [hw] // Syntypus / (Bezděčková et al. 2017) [p, red label]’ [IN: E26-T14].

Remarks. The description was originally prepared by Josef Kratochvíl for publication in the proceedings on the fauna of the Mohelno serpentine steppe, which was, however, released later (KRATOCHVÍL 1944) than the identification key by NOVÁK & SADIL (1941) including a short description based on Kratochvíl’s manuscript and citing Kratochvíl as the author of the new name. Therefore, the text in NOVÁK & SADIL (1941) has priority for nomenclature.

Neither NOVÁK & SADIL (1941) nor KRATOCHVÍL (1944) provided any information on the number of the type specimens or the type locality; they only stated that T. ferox silhavyi was known from Moravia, Slovakia and Hungary. Based on the fact that the taxon was listed among new forms of ants discovered in the Mohelno serpentine steppe (KRATOCHVÍL 1944) and based on the locality data of the specimens in, at least a part of the type series was collected in Mohelno (Czech Republic, Moravia). Four specimens deposited in were originally labelled as cotypes (“cotypy”) and one specimen as a paratype, but no specimen designated as a holotype is available. Therefore, we consider all these specimens syntypes.

Worker
Csösz and Schulz (2010) - Medium to large size, CS 825 [693, 918]. Whole body and appendages greyish brown to dark brown. Head slightly longer than broad, CL/CW 1.03 [0.99, 1.09], with feebly convex sides, straight posterior margin with widely rounded corners. Eyes small, EYE 0.16 [0.15, 0.18]. Frons moderately wide, FR/CS 0.39 [0.36, 0.41], frontal lobes usually as wide as the frons, rarely slightly broader, FL/FR 1.01 [1.00, 1.03]. Scape long, SL/CS 0.79 [0.74, 0.83], with a feeble dorsal carina basally, its surface very finely microreticulate. Promesonotal dorsum slightly convex, metanotal groove shallow. Propodeal denticles moderately long. Petiolar node moderately high, NOH/NOL 1.05 [0.88, 1.29] trapezoid to cubic seen in profile.

Head longitudinally rugulose and microreticulate, posterior surface ruguloso-reticulate, ground surface microreticulate, appears dull. Mesosoma dorsum and mesopleura ruguloso-reticulate, ground surface coarsely microreticulate. Dorsum of petiolar node ruguloso-reticulate, ground surface microreticulate, median part smooth. Dorsum of postpetiole rugulose and microreticulate. First gastral tergite smooth, appears shiny. Ventral surface of head with a row of short setae only (without long C-shape hairs).

Queen
Csösz and Schulz (2010) - Medium to large size, CS 999 [955, 1095]. Whole body and appendages black. Head broader than long, CL/CW 0.92 [0.90, 0.94] with sides and posterior margin straight, with widely rounded corners. Head trapezoid in full face view, narrower at genae. Frons moderately wide, FR/CS 0.40 [0.39, 0.42]. Scape long, SL/CS 0.75 [0.70, 0.81], with a feeble dorsal carina basally, its surface smooth and shiny. Head slightly narrower than scutum, MW/CS 1.03 [0.99, 1.08]. Propodeal teeth long. Petiole and postpetiole wide, CS/PEW 1.65 [1.56, 1.78], CS/PPW 1.29 [1.11, 1.43].

Head dorsum, posterior surface and sides ruguloso-reticulate, ground surface microreticulate. Frons longitudinally rugulose and microreticulate. Scutum and scutellum longitudinally rugulose. Sides of mesosoma, rugoso-reticulate and microreticulate, ventral part of katepisternum always rugulose, or microreticulate. Dorsum of petiolar node and postpetiole coarsely reticulate and microreticulate. First gastral tergite smooth, appears shiny. Ventral surface of head with a row of short setae only (without long C-shape hairs).

Male
Csösz and Schulz (2010) - Head sides feebly convex, posterior margin widely rounded. Scutum much broader than head. Propodeal denticles reduced, propodeum nearly rounded in profile. Dorsum of petiolar node blunt, dorsolaterally sharp and angulate. Whole body and appendages black. Head, scutum, scutellum, anepi- and katepisternum as well as petiole and postpetiole finely rugulo-reticulate, between main sculpturation microreticulate. First gastral tergite smooth, appears shiny.

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