Checklist of Myrmica species

The following species and subspecies belong to the genus Myrmica. Synonyms are listed under their senior names.

A
ademonia Bolton, 1995 (Russian Federation)

• Myrmica aspersa Kupyanskaya, 1990

adrijae Bharti, 2012 (India)

afghanica Radchenko & Elmes, 2003 (Afghanistan)

aimonissabaudiae Menozzi, 1939 (Pakistan)

• Myrmica dicaporiaccoi Menozzi, 1939

alaskensis Wheeler, W.M., 1917 (United States)

• Myrmica brevinodis kuschei Wheeler, W.M., 1917

aloba Forel, 1909 (Spain)

• Myrmica albuferensis Lomnicki, 1925

• Myrmica rolandi Bondroit, 1918

alperti Elmes & Radchenko, 2009 (Nepal)

americana Weber, 1939 (United States)

anatolica Elmes, Radchenko & Aktaç, 2002 (Turkey)

angulata Radchenko, Zhou & Elmes, 2001 (China)

angulinodis Ruzsky, 1905 (Russian Federation)

• Myrmica incurvata Collingwood, 1976

• Myrmica saposhnikovi baikalensis Pisarski, 1969

arisana Wheeler, W.M., 1930 (Taiwan)

• Myrmica kurokii tipuna Santschi, 1937

arnoldii Dlussky, 1963 (Russian Federation)

assimilis Spinola, 1851 (Brazil)

B
bactriana Ruzsky, 1915 (Tibet)

• Myrmica tibetana furva Ruzsky, 1915

bakurianica Arnoldi, 1970 (Georgia)

bergi Ruzsky, 1902 (Kazakhstan)

• Myrmica bergi barchanica Ruzsky, 1905

• Myrmica bergi fortior Crawley, 1920

• Myrmica bergi persiana Weber, 1947

• Myrmica kamyschiensis Arnol'di, 1934

• Myrmica rugosa kirgisica Ruzsky, 1903

bessarabicus Nasonov, 1889 (Moldova)

bibikoffi Kutter, 1963 (Switzerland)

boltoni Radchenko & Elmes, 1998 (Nepal)

brancuccii Radchenko, Elmes & Collingwood, 1999 (Nepal)

breviceps Smith, F., 1878 (India)

brevispinosa Wheeler, W.M., 1917 (United States)

• Myrmica rubra decedens Wheeler, W.M., 1917

C
cachmiriensis Forel, 1904 (India)

• Myrmica smythiesi lutescens Menozzi, 1939

caeca Jerdon, 1851 (India)

cagnianti Espadaler, 1996 (Morocco)

caucasicola Arnoldi, 1934 (Armenia, Azerbaijan)

colax (Cole, 1957) (United States)

collingwoodi Radchenko & Elmes, 1998 (Bhutan)

commarginata Ruzsky, 1905 (Russian Federation)

constricta Karavaiev, 1934 (Ukraine)

• Myrmica rugulosa minuta Karavaiev, 1929

• Myrmica rugulosa slobodensis Arnol'di, 1934

corrugata Say, 1836 (United States)

crassirugis Francoeur, 2007 (United States)

curiosa Radchenko, Zhou & Elmes, 2008 (China)

cursor Smith, F., 1878 (India)

curvispinosa Bharti & Sharma, 2013 (India)

curvithorax Bondroit, 1920 (Spain)

D
deplanata Emery, 1921 (Ukraine, Russian Federation, Georgia)

• Myrmica lobicornis plana Karavaiev, 1927

• Myrmica moravica Soudek, 1922

• Myrmica moravica dagmarae Sadil, 1939

detritinodis Emery, 1921 (United States)

discontinua Weber, 1939 (United States)

displicentia Bolton, 1995 (Russian Federation)

• Myrmica bicolor Kupyanskaya, 1986

divergens Karavaiev, 1931 (Russian Federation)

diversa Buckley, 1867 (United States)

draco Radchenko, Zhou & Elmes, 2001 (China)

dshungarica Ruzsky, 1905 (Kazakhstan)

• Leptothorax svartshevskii Karavaiev, 1916

• Myrmica laevinodis minuta Ruzsky, 1905

• Myrmica minuta iskanderi Arnol'di, 1976

• Myrmica minuta tarbinskii Arnol'di, 1976

• Myrmica rugulosa ruginodiformis Karavaiev, 1929

E
eidmanni Menozzi, 1930 (Russian Federation)

• Myrmica lobicornis littoralis Weber, 1948

elmesi Bharti & Sharma, 2011 (India)

emeryi Radchenko & Elmes, 2009 (Borneo (Pulo Laut))

ereptrix Bolton, 1988 (India)

excelsa Kupyanskaya, 1990 (Russian Federation)

• Myrmica cadusa Kim, Park & Kim, 1997

• Myrmica sinica Wu & Wang, 1995

F
ferganensis Finzi, 1926 (Kyrgyzstan)

forcipata Karavaiev, 1931 (Russian Federation)

foreliana Radchenko & Elmes, 2001 (India)

• Myrmica smythiesii carbonaria Forel, 1902

fortior Forel, 1904 (India)

fracticornis Forel, 1901 (United States)

G
gallienii Bondroit, 1920 (France)

• Myrmica jacobsoni Kutter, 1963

• Myrmica limanica obensis Arnol'di, 1970

• Myrmica rugulosa limanica Karavaiev, 1934

gigantea (Collingwood, 1962) (Myanmar)

glacialis Emery, 1921 (Canada)

granulinodis Nylander, 1846 (Russia)

H
hamulata Weber, 1939 (United States)

hecate Weber, 1947 (India)

hellenica Finzi, 1926 (Greece)

• Myrmica rugulosa caucasica Arnol'di, 1934

• Myrmica rugulosa rugulososcabrinodis Karavaiev, 1929

heterorhytida Radchenko & Elmes, 2009 (China)

hirsuta Elmes, 1978 (United Kingdom of Great Britain and Northern Ireland)

hlavaci Radchenko & Elmes, 2009 (China)

I
incompleta Provancher, 1881 (Canada)

• Myrmica rubra brevinodis Emery, 1895

• Myrmica rubra canadensis Wheeler, W.M., 1916

• Myrmica rubra subalpina Wheeler, W.M., 1917

indica Weber, 1950 (India)

inezae Forel, 1902 (India)

inflecta Say, 1836 (United States)

inucta Radchenko & Elmes, 2006 (Kazakhstan)

J
jennyae Elmes, Radchenko & Aktaç, 2002 (Turkey)

jessensis Forel, 1901 (Japan)

• Myrmica hyungokae Elmes, Radchenko & Kim, 2001

jucunda Smith, F., 1861 (Lebanon)

juglandeti Arnoldi, 1976 (Kyrgyzstan)

• Myrmica tianshanica Arnol'di, 1976

• Myrmica tianshanica alajensis Arnol'di, 1976

K
kabylica (Cagniant, 1970) (Algeria)

kamtschatica Kupyanskaya, 1986 (Russian Federation)

• Myrmica aborigenica Zhigul'skaya, 1991

karavajevi (Arnoldi, 1930) (Ukraine)

• Myrmica faniensis Boven, 1970

• Sifolinia pechi Samsinak, 1957

• Sifolinia winterae Kutter, 1973

kasczenkoi Ruzsky, 1905 (Russian Federation)

khamensis Ruzsky, 1915 (Tibet)

kirghisorum Arnoldi, 1976 (Kyrgyzstan)

• Myrmica lobicornis foreliella Arnol'di, 1976

koreana Elmes, Radchenko & Kim, 2001 (Republic of Korea)

kothiensis Bharti & Sharma, 2013 (India)

kotokui Forel, 1911 (Japan)

• Myrmica ruginodis orientalis Karavaiev, 1926

kozlovi Ruzsky, 1915 (Tibet)

• Myrmica kozlovi mekongi Ruzsky, 1915

• Myrmica kozlovi ruzskyi Weber, 1947

• Myrmica kozlovi subalpina Ruzsky, 1915

• Myrmica kozlovi subbrevispinosa Ruzsky, 1915

• Myrmica specularis Donisthorpe, 1929

kryzhanovskii Arnoldi, 1976 (Tajikistan)

kurokii Forel, 1907 (Japan)

• Myrmica chinensis Viehmeyer, 1922

• Myrmica helleri Viehmeyer, 1922

• Myrmica kozlovi ruzskyi Kiseleva, 1925

• Myrmica longiscapus Curtis, 1854

• Myrmica rubra champlaini Forel, 1901

L
lampra Francoeur, 1968 (Canada)

latifrons Stärcke, 1927 (United States)

• Myrmica schencki emeryana Cole, 1942

laurae (Emery, 1907) (Italy)

• Myrmica samnitica Mei, 1987

lemasnei Bernard, 1967 (France)

lineolata Buckley, 1867 (United States)

lobicornis Nylander, 1846 (Finland)

• Myrmica denticornis Curtis, 1854

• Myrmica lobicornis alpestris Arnol'di, 1934

• Myrmica lobicornis angustifrons Stärcke, 1927

• Myrmica lobicornis arduennae Bondroit, 1911

• Myrmica lobicornis foreli Santschi, 1931

• Myrmica lobicornis kieviensis Karavaiev, 1934

• Myrmica lobicornis lissahorensis Stitz, 1939

• Myrmica schencki brunescens Karavaiev, 1929

• Myrmica schencki burtshakabramovitshi Karavaiev, 1929

• Myrmica schencki starki Karavaiev, 1929

lobifrons Pergande, 1900 (United States)

lobulicornis Nylander, 1857 (France)

• Myrmica arduennae pyrenaea Bondroit, 1918

• Myrmica lobicornis alpina Stärcke, 1927

• Myrmica lobicornis apennina Stärcke, 1927

lonae Finzi, 1926 (Finland)

longisculpta Bharti & Sharma, 2011 (India)

luctuosa Smith, F., 1878 (India)

luteola Kupyanskaya, 1990 (Russian Federation)

M
margaritae Emery, 1889 (Myanmar)

martensi Radchenko & Elmes, 1998 (Nepal)

mexicana Wheeler, W.M., 1914 (Mexico)

mirabilis Elmes & Radchenko, 1998 (Taiwan)

mixta Radchenko & Elmes, 2008 (China)

montana Buckley, 1867 (United States)

monticola Creighton, 1950 (United States)

multiplex Radchenko & Elmes, 2009 (China)

myrmicoxena Forel, 1895 (Switzerland)

N
nearctica Weber, 1939 (United States)

nefaria Bharti, 2012 (India)

nitida Radchenko & Elmes, 1999 (India)

O
obscura Finzi, 1926 (Italy)

obscurata Motschoulsky, 1863 (Sri Lanka)

onoyamai Radchenko & Elmes, 2006 (Japan)

ordinaria Radchenko & Elmes, 1999 (India)

orthostyla Arnoldi, 1976 (Kyrgyzstan)

P
pachei Forel, 1906 (Nepal)

pararitae Radchenko & Elmes, 2008 (China)

pelops Seifert, 2003 (Greece)

petita Radchenko & Elmes, 1999 (India)

phalacra Radchenko & Elmes, 2009 (China)

pilinodis Motschoulsky, 1863 (Sri Lanka)

pinetorum Wheeler, W.M., 1905 (United States)

pisarskii Radchenko, 1994 (Russia, Mongolia)

pleiorhytida Radchenko & Elmes, 2009 (China)

poldii Radchenko & Rigato, 2008 (China)

polyglypta Radchenko & Rigato, 2008 (China)

pseudorugosa Bharti, 2012 (Pakistan)

pulchella Santschi, 1937 (Taiwan)

• Myrmica margaritae formosae Wheeler, W.M., 1929

punctinops Francoeur, 2007 (United States)

punctiventris Roger, 1863 (United States)

• Myrmica punctiventris isfahani Forel, 1922

Q
quebecensis Francoeur, 1981 (Canada)

R
radchenkoi Bharti & Sharma, 2011 (India)

ravasinii Finzi, 1923 (Albania (Tomor))

religiosa Media:Three new species of Myrmica-2013-Journal of Asia Pacific Entomology.pdf|Bharti & Sharma, 2013 (India)

rhynchophora Foerster, 1850 (Algeria)

rhytida Radchenko & Elmes, 1999 (India)

rigatoi Radchenko & Elmes, 1998 (Pakistan)

ritae Emery, 1889 (Myanmar)

rubra (Linnaeus, 1758) (Europe)

• Myrmica laevinodis Nylander, 1846

• Myrmica microrubra Seifert, 1993

• Myrmica rubra bruesi Weber, 1947

• Myrmica rubra europaea Finzi, 1926

rubra neolaevinodis Forel, 1901 (United States)

ruginodis Nylander, 1846 (Finland)

• Myrmica diluta Nylander, 1849

• Myrmica dimidiata Say, 1836

• Myrmica kurokii sontica Santschi, 1937

• Myrmica rubra macrogyna Brian, M.V. & Brian, A.D., 1949

• Myrmica rubra microgyna Brian, M.V. & Brian, A.D., 1949

• Myrmica rubra mutata Sadil, 1952

• Myrmica rubra ruginodolaevinodis Forel, 1874

• Myrmica rubra yoshiokai Weber, 1947

• Myrmica ruginodis silvestrii Wheeler, W.M., 1928

rugiventris (Smith, M.R., 1943) (United States)

rugosa Mayr, 1865 (Himalaya (India?))

rugulosa Nylander, 1849 (Finland)

• Myrmica clandestina Foerster, 1850

• Myrmica rugulosa sulcinodorugulosa Nasonov, 1889

rupestris Forel, 1902 (India)

• Myrmica everesti Donisthorpe, 1929

• Myrmica rugosa debilior Forel, 1902

• Myrmica rugosa rugososmythiesi Forel, 1902

ruzskyana Radchenko & Elmes, 2010 (Tibet)

• Myrmica smythiesi exigua Ruzsky, 1915

S
sabuleti Meinert, 1861 (Denmark)

• Myrmica rubra scabrinodolobicornis Forel, 1874

sabuleti trullicornis Buren, 1944 (United States)

salina Ruzsky, 1905 (Russia Kazakhstan)

• Myrmica scabrinodis ahngeri Karavaiev, 1926

• Myrmica georgica Seifert, 1987

• Myrmica tobiasi Radchenko & Elmes, 2004

saposhnikovi Ruzsky, 1904 (Kazakhstan)

• Myrmica saposhnikovi bergiana Tarbinsky, 1976

• Myrmica scabrinodis intermedia Kuznetsov-Ugamsky, 1927

scabrata Buckley, 1867 (United States)

scabrinodis Nylander, 1846 (Finland)

• Myrmica pilosiscapus Bondroit, 1920

• Myrmica rolandi reticulata Stärcke, 1942

• Myrmica scabrinodis rugulosoides Forel, 1915

• Myrmica scabrinodis scabrinodosabuleti Sadil, 1952

• Sommimyrma symbiotica Menozzi, 1925

schencki Viereck, 1903 (Europe)

• Myrmica betuliana Ruzsky, 1946

• Myrmica schencki kutteri Finzi, 1926

• Myrmica schenckioides Boer & Noordijk, 2005

• Myrmica subopaca Arnol'di, 1934

schoedli Radchenko, Elmes & Bui, 2006 (Viet Nam)

schulzi Radchenko & Elmes, 2009 (China)

sculptiventris Radchenko & Elmes, 2009 (China)

semiparasitica Francoeur, 2007 (United States)

serica Wheeler, W.M., 1928 (China)

siciliana Radchenko, Elmes & Alicata, 2006 (Italy (Sicily))

sinensis Radchenko, Zhou & Elmes, 2001 (China)

sinoschencki Radchenko & Elmes, 2008 (China)

slovaca Sadil, 1952 (Czechia, Slovakia, Ukraine, Bulgaria)

smythiesii Forel, 1902 (India)

• Myrmica smythiesi himalayana Weber, 1947

spatulata Smith, M.R., 1930 (United States)

specioides Bondroit, 1918 (Belgium)

• Leptothorax nevodovskii Karavaiev, 1926

• Myrmica balcanica Sadil, 1952

• Myrmica balcanica scabrinodoides Sadil, 1952

• Myrmica kozakorum Radchenko & Elmes, 2010

• Myrmica puerilis Stärcke, 1942

• Myrmica puerilis dolens Stärcke, 1942

• Myrmica rugulosoides striata Finzi, 1926

• Myrmica sancta tshuliensis Arnol'di, 1976

• Myrmica scabrinodis sancta Karavaiev, 1926

• Myrmica scabrinodis turcica Santschi, 1931

• Tetramorium silvestrianum Emery, 1924

spinosior Santschi, 1931 (France)

stangeana Ruzsky, 1902 (Russia, Kazakhstan)

• Myrmica scabrinodis lacustris Ruzsky, 1905

• Myrmica stangeana medvedevi Arnol'di, 1970

striolagaster Cole, 1953 (United States)

sublanuginosa Buckley, 1867 (United States)

sulcinodis Nylander, 1846 (Finland)

• Myrmica myrmecophila Wasmann, 1910

• Myrmica perelegans Curtis, 1854

• Myrmica ruginodis sulcinodoruginodis Donisthorpe, 1915

• Myrmica sulcinodis derzhavini Ruzsky, 1920

• Myrmica sulcinodis eximia Kupyanskaya, 1990

• Myrmica sulcinodis nigripes Ruzsky, 1895

• Myrmica sulcinodis sulcinodoscabrinodis Forel, 1915

• Myrmica sulcinodis vicaria Kuznetsov-Ugamsky, 1928

• Myrmica sulcinodoscabrinodis Ruzsky, 1895

T
tahoensis Weber, 1948 (United States)

taibaiensis Wei, Zhou & Liu, 2001 (China)

tamarae Elmes, Radchenko & Aktaç, 2002 (Turkey)

tenuispina Ruzsky, 1905 (Uzbekistan)

tibetana Mayr, 1889 (Tibet)

titanica Radchenko & Elmes, 2001 (Viet Nam)

transsibirica Radchenko, 1994 (Russia)

• Myrmica carinata Kupyanskaya, 1990

• Myrmica taediosa Bolton, 1995

tschekanovskii Radchenko, 1994 (Russia)

tulinae Elmes, Radchenko & Aktaç, 2002 (Turkey)

U
urbanii Radchenko & Elmes, 1998 (India)

ussuriensis Kuznetsov-Ugamsky, 1928 (Russia)

V
vandeli Bondroit, 1920 (France)

varisculpta Radchenko & Rigato, 2009 (China)

villosa Radchenko & Elmes, 1999 (Bhutan)

vittata Radchenko & Elmes, 1999 (Pakistan)

W
wardi Radchenko & Elmes, 1999 (Pakistan)

weberi Elmes & Radchenko, 2009 (Nepal)

weii Radchenko & Zhou, 2008 (China)

wesmaeli Bondroit, 1918 (France)

wheeleri Weber, 1939 (United States)

wheelerorum Francoeur, 2007 (United States)

whymperi Forel, 1913 (United States)

• Myrmica brevinodis sulcinodoides Wheeler, W.M., 1917

• Myrmica rubra frigida Wheeler, W.M., 1917

williamsi Radchenko & Elmes, 1999 (India)

wittmeri Radchenko & Elmes, 1999 (Pakistan)

X
xavieri Radchenko, Elmes & Savolainen, 2008 (Spain)

Y
yamanei Radchenko & Elmes, 2001 (Viet Nam)

yunnanensis Radchenko & Elmes, 2009 (China)

Z
zhengi Ma & Xu, 2011 (China)

zojae Radchenko, 1994 (Russia)

A
aemula Heer, 1849 (Switzerland (Miocene))

angusticollis Heer, 1849 (Switzerland (Miocene))

B
bremii Heer, 1849 (Croatia (Miocene))

C
concinna Heer, 1867 (Croatia (Miocene))

E
eocenica Radchenko, Dlussky & Elmes, 2007 (Baltic Amber (Eocene))

I
intermedia (Wheeler, W.M., 1915) (Baltic Amber (Eocene))

J
jurinei Heer, 1849 (Croatia (Miocene))

L
longispinosa Mayr, 1868 (Baltic Amber (Eocene))

M
macrocephala Heer, 1849 (Switzerland (Miocene))

molassica Heer, 1849 (Switzerland (Miocene))

N
nebulosa Novak, O., 1878 (Germany (Oligocene))

O
obsoleta Heer, 1849 (Switzerland (Miocene))

P
paradoxa Radchenko, Dlussky & Elmes, 2007 (Germany (Saxonian amber, Eocene))

R
rudis (Mayr, 1868) (Baltic Amber (Eocene))

T
tertiaria Heer, 1849 (Croatia (Miocene))

• Myrmica bicolor Heer, 1867

V
venusta Heer, 1867 (Croatia (Miocene))