Lasius meridionalis

This species is characteristic of lowland sandy heath in North Europe. Nests are in the ground, often with low earth mounds and carton lined chambers. Flight period August. Fertilised queens start colonies through adoption by established nests of, or. Males which have well toothed mandibles have been seen to pick up objects and to feed themselves (Collingwood, 1979).

Identification
Collingwood (1979) - Clear yellow; pubescence on head rather dilute but close and very fine on gaster. Funiculus segments distinctly longer than wide; scapes and tibiae elliptical in cross section with thin front edge. Petiole sides straight to weakly convex, dorsal margin flat to slightly emarginate. Body and appendage hairs numerous. Length: 3.5-5 mm.

Resembles Lasius umbratus but the different habits, flat appendages and rectangular scale in the queen, shining darker colour and fine sculpture in queen and male clearly distinguish the species from L. umbratus, although workers may be less easy to separate.

Distribution
Spain to Japan, ltaly to Scandinavia (Collingwood 1979).

Distribution based on Regional Taxon Lists
Palaearctic Region: Austria, Belgium, Bulgaria, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Japan, Netherlands, Norway, Poland, Republic of Korea, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, United Kingdom of Great Britain and Northern Ireland.

Nomenclature

 *  meridionalis. Formicina meridionalis Bondroit, 1920a: 143 (q.) FRANCE. Stärcke, 1937: 52 (w.m.). Combination in Lasius: Emery, 1922b: 13; in L. (Chthonolasius): Emery, 1925b: 234. Subspecies of umbratus: Emery, 1922b: 13; Menozzi, 1925d: 34; Stitz, 1939: 295; Novak & Sadil, 1941: 103. Status as species: Stärcke, 1937: 50. Junior synonym of rabaudi: Wilson, 1955a: 168; Bernard, 1967: 364. Revived from synonymy, status as species: Pisarski, 1975: 37; Collingwood, 1979: 102; Seifert, 1988: 154; Kupyanskaya, 1990: 226; Atanassov & Dlussky, 1992: 247.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antarea (at www.antarea.fr on June 11th 2017)
 * ArtDatabanken Bugs (via GBIG)
 * Assing V. 1989. Die Ameisenfauna (Hym.: Formicidae) nordwestdeutscher Calluna-Heiden. Drosera 89: 49-62.
 * Bees, Wasps and Ants Recording Society 243389481
 * Bees, Wasps and Ants Recording Society 243389482
 * Bees, Wasps and Ants Recording Society 243389483
 * Bees, Wasps and Ants Recording Society 243389484
 * Bees, Wasps and Ants Recording Society 243389485
 * Bees, Wasps and Ants Recording Society 243389486
 * Bees, Wasps and Ants Recording Society 243389487
 * Bees, Wasps and Ants Recording Society 243389488
 * Bees, Wasps and Ants Recording Society 243389489
 * Bees, Wasps and Ants Recording Society 243389490
 * Bees, Wasps and Ants Recording Society 243389491
 * Bees, Wasps and Ants Recording Society 243389492
 * Bees, Wasps and Ants Recording Society 243389493
 * Bees, Wasps and Ants Recording Society 243389494
 * Bees, Wasps and Ants Recording Society 243389495
 * Bees, Wasps and Ants Recording Society 243389496
 * Bees, Wasps and Ants Recording Society 243389497
 * Bees, Wasps and Ants Recording Society 243389498
 * Bees, Wasps and Ants Recording Society 243389499
 * Bees, Wasps and Ants Recording Society 243389500
 * Bees, Wasps and Ants Recording Society 243389501
 * Bees, Wasps and Ants Recording Society 243389502
 * Bees, Wasps and Ants Recording Society 243389503
 * Bees, Wasps and Ants Recording Society 243389504
 * Bees, Wasps and Ants Recording Society 243389505
 * Bees, Wasps and Ants Recording Society 243389506
 * Bees, Wasps and Ants Recording Society 243389507
 * Bees, Wasps and Ants Recording Society 243389508
 * Bees, Wasps and Ants Recording Society 243389509
 * Bees, Wasps and Ants Recording Society 243389510
 * Bees, Wasps and Ants Recording Society 243389511
 * Bees, Wasps and Ants Recording Society 243389512
 * Bees, Wasps and Ants Recording Society 243389513
 * Bees, Wasps and Ants Recording Society 243389514
 * Bees, Wasps and Ants Recording Society 243389515
 * Bees, Wasps and Ants Recording Society 243389516
 * Bees, Wasps and Ants Recording Society 243389517
 * Bees, Wasps and Ants Recording Society 243389518
 * Bees, Wasps and Ants Recording Society 243389519
 * Bees, Wasps and Ants Recording Society 243389520
 * Bees, Wasps and Ants Recording Society 243389521
 * Bees, Wasps and Ants Recording Society 243389522
 * Bees, Wasps and Ants Recording Society 243389523
 * Bees, Wasps and Ants Recording Society 243389524
 * Bees, Wasps and Ants Recording Society 243389525
 * Bees, Wasps and Ants Recording Society 243389526
 * Bees, Wasps and Ants Recording Society 243389527
 * Bees, Wasps and Ants Recording Society 243389528
 * Bees, Wasps and Ants Recording Society 243389529
 * Bees, Wasps and Ants Recording Society 243389530
 * Bees, Wasps and Ants Recording Society 243389531
 * Bees, Wasps and Ants Recording Society 243389532
 * Bees, Wasps and Ants Recording Society 243389533
 * Bees, Wasps and Ants Recording Society 243389534
 * Bees, Wasps and Ants Recording Society 243389535
 * Bees, Wasps and Ants Recording Society 243389536
 * Bees, Wasps and Ants Recording Society 243389537
 * Bees, Wasps and Ants Recording Society 243389538
 * Bees, Wasps and Ants Recording Society 243389539
 * Bees, Wasps and Ants Recording Society 243389540
 * Bees, Wasps and Ants Recording Society 243389541
 * Bees, Wasps and Ants Recording Society 243389542
 * Bees, Wasps and Ants Recording Society 243389543
 * Bees, Wasps and Ants Recording Society 243389544
 * Bees, Wasps and Ants Recording Society 243389545
 * Bees, Wasps and Ants Recording Society 243389546
 * Bees, Wasps and Ants Recording Society 243389547
 * Bees, Wasps and Ants Recording Society 243389548
 * Bees, Wasps and Ants Recording Society 243389549
 * Bees, Wasps and Ants Recording Society 243389550
 * Bees, Wasps and Ants Recording Society 243389551
 * Bees, Wasps and Ants Recording Society 243389552
 * Bees, Wasps and Ants Recording Society 243389553
 * Bees, Wasps and Ants Recording Society 243389554
 * Bees, Wasps and Ants Recording Society 243389555
 * Bees, Wasps and Ants Recording Society 243389556
 * Bees, Wasps and Ants Recording Society 243389557
 * Bees, Wasps and Ants Recording Society 243389558
 * Bees, Wasps and Ants Recording Society 243389559
 * Bees, Wasps and Ants Recording Society 243389560
 * Bees, Wasps and Ants Recording Society 243389561
 * Bees, Wasps and Ants Recording Society 243389562
 * Bees, Wasps and Ants Recording Society 243389563
 * Bees, Wasps and Ants Recording Society 243389564
 * Bees, Wasps and Ants Recording Society 243389565
 * Bees, Wasps and Ants Recording Society 243389566
 * Bees, Wasps and Ants Recording Society 243389567
 * Bees, Wasps and Ants Recording Society 243389568
 * Bees, Wasps and Ants Recording Society 243389569
 * Bees, Wasps and Ants Recording Society 243389570
 * Bees, Wasps and Ants Recording Society 243389571
 * Bees, Wasps and Ants Recording Society 243389572
 * Bees, Wasps and Ants Recording Society 243389573
 * Bees, Wasps and Ants Recording Society 243389574
 * Bees, Wasps and Ants Recording Society 243389575
 * Bees, Wasps and Ants Recording Society 243389576
 * Bees, Wasps and Ants Recording Society 243389577
 * Bees, Wasps and Ants Recording Society 243389578
 * Bees, Wasps and Ants Recording Society 243389579
 * Bees, Wasps and Ants Recording Society 243389580
 * Bees, Wasps and Ants Recording Society 243389581
 * Bees, Wasps and Ants Recording Society 243389582
 * Bees, Wasps and Ants Recording Society 243389583
 * Bees, Wasps and Ants Recording Society 243389584
 * Bees, Wasps and Ants Recording Society 243389585
 * Bees, Wasps and Ants Recording Society 243389586
 * Bees, Wasps and Ants Recording Society 243389587
 * Bees, Wasps and Ants Recording Society 243389588
 * Bees, Wasps and Ants Recording Society 243389589
 * Bees, Wasps and Ants Recording Society 243389590
 * Bees, Wasps and Ants Recording Society 243389591
 * Bees, Wasps and Ants Recording Society 243389592
 * Bees, Wasps and Ants Recording Society 243389593
 * Bees, Wasps and Ants Recording Society 243389594
 * Bees, Wasps and Ants Recording Society 243389595
 * Bees, Wasps and Ants Recording Society 243389596
 * Bees, Wasps and Ants Recording Society 243389597
 * Bees, Wasps and Ants Recording Society 243389598
 * Bees, Wasps and Ants Recording Society 243389599
 * Bees, Wasps and Ants Recording Society 243389600
 * Bees, Wasps and Ants Recording Society 243389601
 * Bees, Wasps and Ants Recording Society 243389602
 * Bees, Wasps and Ants Recording Society 243389603
 * Bees, Wasps and Ants Recording Society 243389604
 * Bees, Wasps and Ants Recording Society 243389605
 * Bees, Wasps and Ants Recording Society 243389606
 * Bees, Wasps and Ants Recording Society 243389607
 * Bees, Wasps and Ants Recording Society 243389608
 * Bees, Wasps and Ants Recording Society 243389609
 * Bees, Wasps and Ants Recording Society 243389610
 * Bees, Wasps and Ants Recording Society 243389611
 * Bees, Wasps and Ants Recording Society 243389612
 * Bees, Wasps and Ants Recording Society 243389613
 * Bees, Wasps and Ants Recording Society 243389614
 * Bees, Wasps and Ants Recording Society 243389615
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Blacker N. C. 1989. The ants (Hymenoptera, Formicidae) of the Gower Peninsula, West Glamorgan, South Wales. Entomologist's Record and Journal of Variation 101: 261-266.
 * Blacker N. C. and C. A. Collingwood. 2002. Some significant new records of ants (Hymenoptera: Formicidae) from the Salisbury area, south Wiltshire, England, with a key to the British species of Lasius. British Journal of Entomology and Natural History 15: 25-46
 * Blacker N.C. 2007. Ants (Hym., Formicidae) in East Anglia-Additional Records from . Entomologist's Monthly Magazine 143: 69-90
 * Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
 * Boer P., and J. Noordijk. 2004. De ruige gaststeekmier Myrmica hirsuta nieuw voor Nederland (Hymenoptera: Formicidae). Ned. Faun. Meded. 20: 25-32.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Bra?ko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Choi B.M. 1985. Study on distribution of ants (Formicidae) from Korea (1). Formic fauna in Mt. Songni. Cheongju Sabom Taehak Nonmunjip (Journal of Cheongju National Teachers' College) 22:401-437.
 * Choi B.M. 1986. Studies on the distribution of ants (Formicidae) in Korea. Journal of Chongju National Teacher College 23: 317-386.
 * Choi B.M. 1996. Studies on the distribution of ants (Formicidae) in Korea (15) -Ant fauna islands Ullungdo and Dokdo. Journal of Chongju National University of Education 33: 201-219.
 * Choi B.M., K. Ogata, and M. Terayama. 1993. Comparative studies of ant faunas of Korea and Japan. 1. Faunal comparison  among islands of Southern Korean and northern Kyushu, Japan. Bull. Biogeogr. Soc. Japan 48(1): 37-49.
 * Choi B.M., and J. R. Bang. Studies on the distribution of ants (Formicidae) in Korea (12): the analysis of ant communities in 23 islands. Journal of Cheongju National University of Education 30:317-330.
 * Collingwood C., and A. Prince. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5: 1-49.
 * Collingwood, C. A.. "Especies raras de hormigas del género Lasius en España." Boletín de la Asociación Española de Entomología 15 (1991): 215-219.
 * Collingwood, C. A.. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Mem. Soc. Entomol. Ital. 31: 38-48.
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Dekoninck W., P. Boer, and J. P. Maelfait. 2004. Lasius platythorax Seifert, 1991 as a host of several Chthonolasius species, with remarks on the colony foundation of the parasites (Hymenoptera: Formicidae). Myrmecologische Nachrichten 6: 5-8.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Formidabel Database
 * Fowles, A.P. 1996. A provisional checklist of the invertebrates recorded from Wales. 2. Aculeate wasps, bees and ants (Hymenoptera: Aculeata). Countryside Council for Wales
 * Galkowski C. 2008. Quelques fourmis nouvelles ou intéressantes pour la faune de France (Hymenoptera, Formicidae). Bulletin de la Société Linnéenne de Bordeaux, 143, N.S. 36, 4 : 423-433.
 * Galle L. 1997. Contribution to the ant fauna of Slovenia with special reference to the submediterranean and eudinaric regions. Annals for Istrian and Mediterranean studies 11: 209-214.
 * Gallé L. 1991. Structure and succession of ant assemblages in a north European sand dune area. Holarctic Ecology 14: 31-37.
 * Giacalone I., and M. Moretti. 2001. Contributo alla conoscenza della mirmecofauna (Hymenoptera: Formicidae) dei castagneti al Sud delle Alpi (ticino, Svizzera). Bollettino della Societa ticinese di Scienze naturali 89(1-2): 51-60.
 * Glaser F., A. Freitag, and H. Martz. 2012. Ants (Hymenoptera: Formicidae) in the Münstertal (Val Müstair)  a hot spot of regional species richness between Italy and Switzerland. Gredleriana 12: 273 - 284.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Ha S.J, S.J. Park, and B.J. Kim. 2002. Comparative ant faunas between Seonyudo and seven other islands of West Sea in Korea. Korean Journal of Entomology 32(2): 75-79.
 * Kim B.J. 1996. Synonymic list and distribution of Formicidae (Hymenoptera) in Korea. Entomological Research Bulletin Supplement 169-196.
 * Kim B.J., K.G. Kim, D.P. Ryu, J.H. Kim. 1995. Ants of Chindo island in Korea (Hymenoptera; Formicidae). The Korean Journal of Systematic Zoology 11(1): 101-113.
 * Kim B.J.; Kim, K.G.; Lim, K.H. 1993. Systematic study of ants from Chejudo province. Korean Journal of Entomology 23: 117-116
 * Kim et al. 1993. Systematic study of ants from Chejudo Province. Koran Journal of Entomology 23(3): 117-141.
 * Kim, Byung-Jin, Ky-Gyong Kim, Dong-Pyo Ryu and Joong-Hyon Kim. 1995. The Korean Journal of Systematic Zoology. 11(1):101-113.
 * Kupianskaia A.N. 1990. Murav'I (Hymenoptera, Formicidae) Dal'nego Vostoka SSSR (1989). Vladivostok. 258 pages.
 * Kvamme T. 1982. Atlas of the Formicidae of Norway (Hymenoptera: Aculeata). Insecta Norvegiae 2: 1-56.
 * Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur  Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
 * Kwon T. S. 2015. Ant assemblages along the Baekdudaegan Mountain Range in South Korea: Human roads and temperature niche. Journal of Asia-Pacific Biodiversity 8: 152-157.
 * Kwon T. S., S. S. Kim, and J. H. Chun. 2014. Pattern of ant diversity in Korea: An empirical test of Rapoport's altitudinal rule. Journal of Asia-Pacific Entomology 17: 161167.
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Lehouck V., D. Bonte, W. Dekoninck, and J. P. Maelfait. 2004. The distribution of ant nests (Hymenoptera, Formicidae) in coastal grey dunes of Flanders (Belgium) and their relationship to myrmecochorous plants. Belg. J. Zool. 134 (2/1) : 89-96.
 * Nielsen M. G. 2011. A check list of Danish ants and proposed common names. Ent. Meddr. 79: 13-18.
 * Noordijk, J., R. Morssinkhof, P. Boer, A. P. Schaffers, Th. Heijerman and K. V. Sýkora. 2008. How ants find each other; temporal and spatial patterns in nuptial flights. Insectes Sociaux 55(3):266-273.
 * Odegaard F. 2013. New and little known ants (Hymenoptera, Formicidae) in Norway. Norwegian Journal of Entomology 60, 172175.
 * Park S.J., and B.J. Kim. 2002. Faunal comparison of ants among Cheongsando and other islands of South Sea in Korea. Korean Journal of Entomology 32(1): 7-12.
 * Park, Seong, Joon and Byung, and Kim, Jin. 2002. Faunal Comparison of Ants among Cheongsando and Other Islands of South Sea in Korea. Korean Jornal of Entomology. 32(1):7-12.
 * Petrov I. Z. 2002. Contribution to the myrmecofauna (Formicidae, Hymenoptera) of the Banat Province (Serbia). Archives of Biological Sciences, Belgrade, 54(12): 57-64.
 * Petrov I. Z. 2012. Preliminary data on ants (Formicidae, Hymenoptera) on Mountain Avala (Belgrade, Serbia). Bulletin of the Natural History Museum 5: 95-99.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Radchenko A., W. Czechowska, W. Czechowski, and E. Siedlar. 1999. Four species of the ant genus Lasius F. new to Poland, with additions to the records for previously reported species (Hymenoptera, Formicidae). Fragmenta Faunistica 42(11): 115-121.
 * Radchenko, A. 2005. Monographic revision of the ants (Hymenoptera, Formicidae) of North Korea. Annales Zoologici 55(2): 127-221.
 * Sato T., N. Tsurusaki, K. Hamaguchi, and K. Kinomura. 2010. Ant fauna of Tottori prefecture, Honshu, Japan. Bulletin of the Tottori Prefectural Museum 47: 27-44.
 * Schar S., G Talavera, X. Espadaler, J. D. Rana, A. A. Andersen, S. P. Cover, and R. Vila. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 00: 1-12.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
 * Seifert B. 1998. Rote Liste der Ameisen. - in: M. Binot, R. Bless, P. Boye, H. Gruttke und P. Pretscher: Rote Liste gefährdeter Tiere Deutschlands. Bonn-Bad Godesberg 1998: 130-133.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Stärcke A. 1937. Retouches sur quelques fourmis d'Europe. II. Lasius groupe umbratus Nylander. Tijdschr. Entomol. 80: 38-72.
 * Terayama M. 1992. Structure of ant communities in East Asia. A. Regional differences and species richness. Bulletin of the Bio-geographical Society of Japan 47: 1-31.
 * Terayama M., Choi, B.M., Kim, C.H. 1992. A check list of ants from Korea, with taxonomic notes. Bulletin of the Toho Gakuen 7:19-54.
 * Terayama M., K. Ogata, and B.M. Choi. 1994. Distribution records of ants in 47 prefectures of Japan. Ari (report of the Myrmecologists Society of Japan) 18: 5-17.
 * Terayama M., S. Kubota, and K. Eguchi. 2014. Encyclopedia of Japanese ants. Asakura Shoten: Tokyo, 278 pp.
 * Tinaut A. 2016. Ants of the Tejeda, Almijara and Alhama Mountains Natural Park (Andalusia, Spain) (Hymenoptera, Formicidae). Boln. Asoc. esp. Ent., 40 (1-2): 125-159.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.