Stictoponera coxalis

Identification
Lattke (2004) - Clypeal lamella projecting anterad as broad triangular lobe; occipital lobes poorly developed. Pronotal dorsum punctate, with rough transverse strigae that arch anterad, becoming longitudinal on posterior pronotum and mesonotum; propodeum unarmed and transversely strigose.

Distribution based on Regional Taxon Lists
Indo-Australian Region: Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore. Oriental Region: India, Myanmar, Nicobar Island, Sri Lanka , Thailand, Vietnam. Palaearctic Region: China.

Nomenclature

 * . Ponera coxalis Roger, 1860: 308.
 * Replacement name for Ponera rugosa Smith, F. 1857a: 66. [Junior primary homonym of Ponera rugosa Le Guillou, 1842: 318.]
 * Combination in Ectatomma (Rhytidoponera): Mayr, 1862: 732;
 * combination in Ectatomma: Emery, 1887b: 444;
 * combination in E. (Stictoponera): Mayr, 1887: 539;
 * combination in Gnamptogenys: Brown, 1958g: 228;
 * combination in Stictoponera: Emery, 1900d: 662; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Status as species: Mayr, 1862: 732; Mayr, 1863: 410; Roger, 1863b: 17; Mayr, 1867a: 89 (redescription); Mayr, 1872: 150; Emery, 1887b: 444; Emery, 1889b: 494 (footnote, in key); Dalla Torre, 1893: 24; Forel, 1899d: 316; Bingham, 1903: 84; Emery, 1911d: 48; Wheeler, W.M. 1919e: 51; Karavaiev, 1925a: 79; Wheeler, W.M. & Chapman, 1925: 59; Chapman & Capco, 1951: 29; Brown, 1954h: 9; Brown, 1958g: 228, 300; Baltazar, 1966: 235; Bolton, 1995b: 209; Lattke, 2004: 103 (redescription); Mohanraj, et al. 2010: 6; Pfeiffer, et al. 2011: 35; Bharti, Guénard, et al. 2016: 23; Lattke & Delsinne, 2016: 144; ; Dias, R.K.S. et al. 2020: 28; Khachonpisitsak, et al. 2020: 38; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of costatum: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of parva: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of pinealis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of simalurensis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of unicolor: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Senior synonym of wallacei: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * Distribution: Brunei, China, India (+ Andaman Is, Nicobar Is), Indonesia (Java, Kalimantan, Sulawesi, Sumatra), Malaysia (Peninsula, Sabah, Sarawak), Myanmar, Philippines (Luzon, Mindanao), Singapore, Sri Lanka, Thailand, Vietnam.
 * costatum. Ectatomma (Stictoponera) costatum Emery, 1889b: 494 (w.) MYANMAR.
 * Type-material: holotype worker.
 * Type-locality: Myanmar (“Burma”): Tenasserim, Thagata (L. Fea).
 * Type-depository: MSNG.
 * Wheeler, G.C. & Wheeler, J. 1976a: 44 (l.); Lattke, 2004: 100 (q.m.).
 * Combination in Stictoponera: Emery, 1900d: 662;
 * combination in Gnamptogenys: Brown, 1958g: 227.
 * Status as species: Emery, 1889b: 494 (footnote, in key); Dalla Torre, 1893: 24; Emery, 1895k: 458; Forel, 1899d: 316; Emery, 1900d: 662; Forel, 1901f: 335; Emery, 1911d: 48; Forel, 1912d: 98; Forel, 1912n: 52; Crawley, 1924: 383; Wheeler, W.M. 1929g: 31; Chapman & Capco, 1951: 29; Brown, 1954h: 7; Brown, 1958g: 227; Baltazar, 1966: 235; Radchenko, 1993a: 78; Bolton, 1995b: 208; Lattke, 2004: 97 (redescription); Pfeiffer, et al. 2011: 35; Liu, C., Guénard, et al. 2015: 38.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * parva. Stictoponera rugosa r. parva Forel, 1913k: 6 (w.) INDONESIA (Sumatra).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Sumatra, Lau-Boentoe, 1912 (von Buttel-Reepen).
 * Type-depository: MHNG.
 * Subspecies of rugosa: Chapman & Capco, 1951: 30.
 * Junior synonym of costatum: Brown, 1954h: 7; Lattke, 2004: 98.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * pinealis. Stictoponera costata var. pinealis Wheeler, W.M. 1929g: 31 (q.) WEST MALAYSIA (Penang I.).
 * Type-material: holotype worker.
 * Type-locality: Malaysia: Penang I. (F. Silvestri).
 * Type-depository: MCZC.
 * Subspecies of costatum: Chapman & Capco, 1951: 29.
 * Junior synonym of costatum: Brown, 1954h: 7; Lattke, 2004: 98.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * rugosa. Ponera rugosa Smith, F. 1857a: 66 (q.) BORNEO (East Malaysia: Sarawak).
 * Type-material: holotype queen.
 * Type-locality: Malaysia (“Borneo”): Sararwak, “SAR. 20” (A.R. Wallace).
 * Type-depository: OXUM.
 * [Junior primary homonym of Ponera rugosa Le Guillou, 1842: 318.]
 * Combination in Stictoponera: Emery, 1911d: 48.
 * Status as species: Smith, F. 1858b: 88; Smith, F. 1861b: 44; Emery, 1911d: 48; Wheeler, W.M. 1919e: 51; Chapman & Capco, 1951: 30.
 * Replacement name: Ponera coxalis Roger, 1860: 308.
 * simalurensis. Stictoponera costata st. simalurensis Forel, 1915a: 23 (w.) INDONESIA (Sumatra: Simalur I.).
 * Type-material: holotype worker.
 * Type-locality: Indonesia: Sumatra, Simalur I. (E. Jacobson).
 * Type-depository: MHNG.
 * Subspecies of costatum: Chapman & Capco, 1951: 29.
 * Junior synonym of costatum: Brown, 1954h: 7; Lattke, 2004: 98.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * unicolor. Stictoponera costata var. unicolor Forel, 1901f: 335 (w.m.) BORNEO (East Malaysia: Sarawak).
 * Type-material: syntype worker(s), syntype males(s) (numbers not stated).
 * Type-locality: Malaysia: Sarawak (Haviland).
 * Type-depository: MHNG.
 * Subspecies of costatum: Emery, 1911d: 48; Wheeler, W.M. 1919e: 51; Chapman & Capco, 1951: 29.
 * Junior synonym of costatum: Brown, 1954h: 7; Lattke, 2004: 98.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144; Camacho, Franco, Branstetter, et al. 2022: 12.
 * wallacei. Stictoponera wallacei Donisthorpe, 1932c: 447.
 * Unnecessary (second) replacement name for Ponera rugosa Smith, F. 1857a: 66; hence automatic junior synonym of coxalis Roger, 1860: 308 (first replacement name).
 * Junior synonym of costatum: Brown, 1954h: 7; Lattke, 2004: 98.
 * Junior synonym of coxalis: Lattke & Delsinne, 2016: 144.

Type Material
Lattke (2004) - Lectotype worker: Ceylon [Sri Lanka] (Nietner) [Not examined]. Designated by Brown, 1954b:9.

Lattke (2004) - Holotype worker by monotypy: Burma [Myanmar], Tenasserim (Fea) (MCSN) [Examined].

Stictoponera costata var. unicolor Forel, 1901b:335. Syntype workers, queen: Borneo (MHNG) [Not examined]. Synonymy by Brown, 1954b:7.

Stictoponera rugosa var. parva Forel, 1913:6. Holotype worker by monotypy: Sumatra (MHNG) [Not examined]. Synonymy by Brown, 1954b:7.

Stictoponera costata var. simalurensis Forel, 1915:23. Holotype worker by monotypy: Sumatra (MHNG) [Not examined]. Synonymy by Brown, 1954b:7.

Stictoponera costata var. pinealis Wheeler, W.M. 1929:31. Holotype queen by monotypy: Malaysia, Penang (Silvestri) (MCZC) [Examined]. Synonymy by Brown, 1954b:7.

Stictoponera wallacei Donisthorpe, 1932:447. Replacement name for Ponera rugosa Smith, F. 1857:66. Synonymy by Brown, 1954b:7.

Ponera rugosa

Holotype dealate queen in. Labelled “SAR. 20,” and “rugosa Sm.” There is also a Donisthorpe type-label and one that states “Stictoponera wallacei Donisthorpe,” his replacement name for the preoccupied P. rugosa F. Smith.

There is a specimen in that is incorrectly labelled as type of rugosa; it is actually a worker of Gnamptogenys menadensis (Mayr).

References based on Global Ant Biodiversity Informatics

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