Checklist of Polyrhachis species

The following species and subspecies belong to the genus Polyrhachis. Synonyms are listed under their senior names. For valid names only see Polyrhachis species and for distribution information see Polyrhachis species by Country.

A
abbreviata Kohout, 2006 (Australia)

abdita Kohout, 2007 (New Guinea)

abdominalis Smith, F., 1858 (Borneo, India, Indonesia, Krakatau Islands, Laos, Malaysia, Myanmar, Philippines, Thailand)

• Polyrhachis achilles confinis Forel, 1912

• Polyrhachis achilles discrepans Forel, 1912

• Polyrhachis phyllophilus Smith, F., 1860

aberrans Kohout, 2008 (Indonesia, Sulawesi)

abnormis Donisthorpe, 1948 (New Guinea)

abrupta Mayr, 1867 (Indonesia)

• Polyrhachis orsyllus halmaheirae Forel, 1886

achterbergi Kohout, 2006 (Borneo, Indonesia, Malaysia)

aculeata Mayr, 1879 (India, Philippines)

acuminata Kohout, 2007 (Borneo, Indonesia, Malaysia)

aedipus Forel, 1893 (India, Sri Lanka)

aenescens Stitz, 1910 (Cameroun)

aequalis Forel, 1910 (Borneo, Indonesia, Malaysia, Philippines)

aerope Wheeler, W.M., 1922 (Cameroun, Congo, Democratic Republic of Congo)

agesilas Forel, 1913 (Indonesia)

alata Forel, 1904 (Indonesia, Sulawesi)

alatisquamis Forel, 1893 (India, Myanmar, Thailand)

albertisi Emery, 1887 (New Guinea)

alexisi Forel, 1916 (Central African Republic, Democratic Republic of Congo)

alluaudi Emery, 1892 (Democratic Republic of Congo, Ivory Coast)

• Polyrhachis alluaudi anteplana Forel, 1916

alphea Smith, F., 1863 (Australia, New Guinea)

alphena Smith, F., 1860 (Indonesia)

amana Smith, F., 1861 (Indonesia, Sulawesi)

ammon (Fabricius, 1775) (Australia)

• Polyrhachis ammon angustata Forel, 1902

ammonoeides Roger, 1863 (Australia)

• Polyrhachis chalchas Forel, 1907

anderseni Kohout, 2013 (Australia)

andrei Emery, 1921 (Cameroun, Congo, Ghana, United Republic of Tanzania)

andromache Roger, 1863 (Australia, Indonesia, New Guinea, Solomon Islands)

• Polyrhachis chaonia Smith, F., 1861

• Polyrhachis connectens Emery, 1887

• Polyrhachis connectens australiae Emery, 1887

• Polyrhachis hector Smith, F. Smith, F., 1859

• Polyrhachis proxima semirufipes Donisthorpe, 1943

angusta Forel, 1902 (Australia)

annae Mann, 1919 (Solomon Islands)

annulata Kohout, 2007 (New Guinea)

antennata Viehmeyer, 1912 (New Guinea)

antennata reticulata Dorow, 1995 (New Guinea)

• Polyrhachis antennata aciculata Viehmeyer, 1912

antoniae Stitz, 1911 (New Guinea)

aporema Kohout, 2006 (New Guinea)

appendiculata Emery, 1893 (Australia)

arachne Emery, 1896 (Indonesia, Thailand)

• Polyrhachis uncinata André, 1896

arborea Kohout, 2008 (Indonesia, Sulawesi)

archeri Kohout, 2013 (Australia)

arcuata (Le Guillou, 1842) (Australia, Borneo, Indonesia, Krakatau Islands, Malaysia, Philippines, Sulawesi, Thailand)

• Polyrhachis arcuata continentis Forel, 1905

• Polyrhachis latifrons Roger, 1863

• Polyrhachis modiglianii Emery, 1888

arcuata acutinota Forel, 1901 (New Guinea, Solomon Islands)

argenteosignata Emery, 1900 (Australia, New Guinea)

argentosa Forel, 1902 (Australia)

armata (Le Guillou, 1842) (Borneo, Cambodia, China, India, Indonesia, Laos, Malaysia, Philippines, Thailand)

• Polyrhachis pandarus Smith, F., 1857

armata defensa Smith, F., 1857 (India, Philippines, Singapore)

• Polyrhachis armata minor Forel, 1886

arnoldi Forel, 1914 (Malawi, South Africa)

aruensis Viehmeyer, 1912 (New Guinea)

asomaningi Bolton, 1973 (Central African Republic, Gabon, Ghana)

aspasia Forel, 1911 (Borneo, Indonesia, Malaysia)

atossa Forel, 1913 (Indonesia, New Guinea)

atossa aequicuspis Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

• Polyrhachis constructor Smith, F. Smith, F., 1858

atropos Smith, F., 1860 (Australia, Indonesia, New Guinea)

• Polyrhachis eucharis Karavaiev, 1927

atrovirens Emery, 1900 (Borneo, Indonesia, Malaysia)

aurea Mayr, 1876 (Australia)

aurea fiorii Emery, 1914 (Vanuatu)

aureovestita Donisthorpe, 1937 (New Guinea, Philippines)

auriformis Donisthorpe, 1943 (New Guinea)

aurita Emery, 1911 (New Guinea)

• Polyrhachis schlaginhaufeni Viehmeyer, 1912

• Polyrhachis schlaginhaufeni longispina Viehmeyer, 1912

aurora Kohout, 2013 (Australia)

australis Mayr, 1870 (Australia)

• Polyrhachis doddi Donisthorpe, 1938

• Polyrhachis nox Donisthorpe, 1938

• Polyrhachis townsvillei Donisthorpe, 1938

B
baca Sorger & Zettel, 2010 (Philippines)

bakana Xu, Z., 1998 (China)

bakeri Viehmeyer, 1916 (Philippines)

bamaga Kohout, 1990 (Australia)

banghaasi Viehmeyer, 1922 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines)

barretti Clark, 1928 (Australia)

barryi Kohout, 2006 (New Guinea)

basirufa Emery, 1900 (Borneo, Indonesia, Malaysia)

batesi Forel, 1911 (Indonesia)

• Polyrhachis distincta Karavaiev, 1927

beauforti Emery, 1911 (New Guinea)

beauforti punctinota Viehmeyer, 1914 (New Guinea)

beccarii Mayr, 1872 (Borneo, Indonesia, Malaysia)

bedeloweryi Kohout, 2007 (New Guinea)

bedoti Forel, 1902 (Australia, New Guinea)

bellicosa Smith, F., 1859 (Australia, Borneo, Indonesia, Laos, Malaysia, New Guinea, Philippines)

• Polyrhachis bellicosa crudelis Emery, 1887

bicolor Smith, F., 1858 (Australia, Bangladesh, Borneo, Brunei Darussalam, Cambodia, China, India, Indonesia, Krakatau Islands, Laos, Malaysia, Myanmar, New Guinea, Philippines, Singapore, Thailand, Vietnam)

• Polyrhachis bicolor concolor Forel, 1910

bicolor atrocastanea Karavaiev, 1927 (Indonesia)

bicolor aurata Karavaiev, 1935 (Indonesia)

bicolor aurinasis Forel, 1901 (Borneo, India, Indonesia, Malaysia, Philippines)

bicolor brachyacantha Karavaiev, 1935 (Cambodia)

bicolor comata Emery, 1911 (New Guinea)

bicolor erecta Karavaiev, 1935 (Indonesia)

bicolor exflavicornis Baltazar, 1966 (Philippines)

• Polyrhachis bicolor flavicornis Stitz, 1925

bicolor fumata Stitz, 1925 (Philippines)

bicolor weyeri Karavaiev, 1930 (Indonesia)

bihamata (Drury, 1773) (Borneo, Brunei Darussalam, Cambodia, China, India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Thailand, Vietnam)

• Formica affinis Le Guillou Le Guillou, 1842

• Polyrhachis bihamata minor Karavaiev, 1927

• Polyrhachis bihamata perplexa Santschi, 1925

• Polyrhachis bihamata tonsilis Santschi, 1928

binghamii Forel, 1893 (India, Myanmar)

bismarckensis Forel, 1901 (New Guinea)

bispinosa Kohout, 2013 (Australia)

bohemia Kohout, 2013 (Australia)

boltoni Dorow & Kohout, 1995 (Borneo, Indonesia, Malaysia)

bosi Kohout, 2008 (Indonesia, Sulawesi)

bouvieri Santschi, 1928 (Indonesia)

brachyspina Kohout, 2008 (Indonesia, Sulawesi)

braxa Bolton, 1973 (Ghana)

brendelli Kohout, 2008 (Indonesia, Sulawesi)

breviata Kohout, 2007 (New Guinea)

brevicorpa Xu, Z., 2002 (China)

brevinoda Kohout, 2006 (Australia, New Guinea)

breviorspinosa Donisthorpe, 1947 (New Guinea)

brevipilosa Rigato, 2016 (Congo, Gabon)

brisbanensis Kohout, 2013 (Australia)

browni Kohout, 2008 (Indonesia, Sulawesi)

bruehli Kohout, 2006 (Borneo, Indonesia, Malaysia)

brutella Kohout, 2013 (Australia)

bubastes Smith, F., 1863 (Indonesia, New Guinea)

• Polyrhachis hirta Donisthorpe Donisthorpe, 1949

• Polyrhachis kellyi Hung, 1967

• Polyrhachis rugousissima Donisthorpe, 1943

• Polyrhachis variolosa Emery, 1887

• Polyrhachis variolosa arcispina Santschi, 1916

• Polyrhachis variolosa curvispina Stitz, 1911

• Polyrhachis variolosa waigiouensis Forel, 1911

bugnioni Forel, 1908 (Sri Lanka)

burmanensis Donisthorpe, 1938 (China, Myanmar)

burwelli Kohout, 2013 (Australia)

busiris Smith, F., 1860 (New Guinea)

C
caeciliae Forel, 1912 (Borneo, Indonesia, Malaysia)

• Polyrhachis punctata Karavaiev, 1927

callima Kohout, 2013 (Australia)

calliope Emery, 1900 (New Guinea)

calypso Forel, 1911 (Borneo, India, Indonesia, Malaysia, Philippines, Thailand)

• Polyrhachis capra Karavaiev, 1927

• Polyrhachis sexspinosa malaccana Viehmeyer, 1916

campbelli Mann, 1919 (Solomon Islands)

capeyorkensis Kohout, 2013 (Australia)

capillata Kohout, 2013 (Australia)

captiva Kohout, 2013 (Australia)

carbonaria Smith, F., 1857 (Indonesia, Malaysia, Philippines)

• Polyrhachis orsyllus dentulata Stitz, 1925

• Polyrhachis orsyllus subcarinata Emery, 1900

carinata (Fabricius, 1804) (New Caledonia)

castaneiventris Smith, F., 1858 (Borneo, Indonesia, Malaysia)

caulomma Viehmeyer, 1914 (New Guinea)

• Polyrhachis caulomma parallela Viehmeyer, 1914

cedarensis Forel, 1915 (Australia)

celebensis Viehmeyer, 1913 (Indonesia, Sulawesi)

cephalotes Emery, 1893 (Borneo, Indonesia, Malaysia)

ceramensis Mayr, 1883 (Indonesia)

chaita Kohout, 2008 (Indonesia, Sulawesi)

chalybea Smith, F., 1857 (Borneo, Indonesia, Malaysia, Singapore)

• Polyrhachis sappho Forel, 1911

chapmani Kohout, 2006 (Philippines)

charaxa Smith, F., 1860 (Indonesia)

chartifex Emery, 1900 (New Guinea)

cheesmanae Donisthorpe, 1937 (New Guinea)

cincta Viehmeyer, 1913 (Indonesia, Sulawesi)

cingula Donisthorpe, 1947 (New Guinea)

circumdata Viehmeyer, 1913 (Indonesia, Sulawesi)

clarkei Donisthorpe, 1949 (New Guinea)

clarki Kohout, 2013 (Australia)

cleopatra Forel, 1902 (Australia)

cleophanes Smith, F., 1861 (Indonesia, Sulawesi)

• Polyrhachis vibidia Smith, F., 1861

clio Forel, 1902 (Australia)

clotho Forel, 1902 (Australia)

coerulescens Emery, 1897 (New Guinea)

coerulescens nigronitens Viehmeyer, 1914 (New Guinea)

coerulescens strigifrons Viehmeyer, 1914 (New Guinea)

cognata Kohout, 2008 (Indonesia, Sulawesi)

compressicornis Smith, F., 1860 (Indonesia, Sulawesi)

concava André, 1889 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Equatorial Guinea, Gabon, Ghana, Ivory Coast, Sierra Leone)

conciliata Kohout, 2013 (Australia)

conops Forel, 1901 (New Guinea)

consimilis Smith, F., 1858 (Australia)

conspicua Kohout, 2006 (New Guinea)

constricta Emery, 1897 (Australia)

constructor Smith, F., 1857 (Borneo, Indonesia, Malaysia)

contemta Mayr, 1876 (Australia)

continua Emery, 1887 (Indonesia, New Guinea, Sulawesi)

• Polyrhachis continua revocata Viehmeyer, 1913

convexa Roger, 1863 (China, India, Sri Lanka)

convexa isabellae Forel, 1908 (Sri Lanka, Sri Lanka)

cornihumera Xu, Z., 2002 (China)

cornuhumera Zhou & Huang, 2002 (China)

cornuta Stitz, 1910 (Democratic Republic of Congo)

corporaali Santschi, 1928 (India, Indonesia)

costulata Emery, 1897 (New Guinea)

cracenta Kohout, 2013 (Australia)

craddocki Bingham, 1903 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Myanmar)

crassispinosa Viehmeyer, 1914 (New Guinea)

crawleyi Forel, 1916 (Australia)

creusa Emery, 1897 (Australia, Indonesia, New Guinea, Philippines)

• Polyrhachis creusa chlorizans Forel, 1901

• Polyrhachis creusa distinguenda Karavaiev, 1927

• Polyrhachis hecuba Forel, 1902

croceiventris Emery, 1900 (New Guinea)

cryptoceroides Emery, 1887 (Borneo, Indonesia, Laos, Malaysia, Philippines, Sulawesi)

• Polyrhachis mystica Karavaiev, 1927

cubaensis Mayr, 1862 (Kenya, Mozambique, South Africa, Sudan, United Republic of Tanzania)

cupreata Emery, 1895 (Australia)

• Polyrhachis daemeli exlex Forel, 1915

curta André, 1890 (Cameroun, Congo, Democratic Republic of Congo, Ghana, Sierra Leone)

• Polyrhachis lyrifera Stitz, 1933

• Polyrhachis maynei Forel, 1911

curtospinosa Kohout, 2013 (Australia)

curvispina Forel, 1908 (Sri Lanka)

cyaniventris Smith, F., 1858 (Philippines)

• Polyrhachis cyaneus Mayr, 1862

cybele Wheeler, W.M., 1919 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines)

cydista Kohout, 2008 (Australia)

cyphonota Xu, Z., 1998 (China)

cyrus Forel, 1901 (Australia, New Guinea)

• Polyrhachis urania Forel, 1902

D
daemeli Mayr, 1876 (Australia)

dahlii Forel, 1901 (New Guinea, Solomon Islands)

danum Kohout, 2006 (Borneo, Indonesia, Malaysia)

daphne Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

darlingtoni Kohout, 2013 (Australia)

davydovi Karavaiev, 1927 (Indonesia)

debilis Emery, 1887 (China, New Guinea)

decellei Bolton, 1973 (Central African Republic, Ghana)

decemdentata André, 1889 (Cameroun, Congo, Equatorial Guinea, Gabon, Ghana, Guinea, Ivory Coast, Nigeria, Sierra Leone, Uganda)

• Polyrhachis decemdentata fernandensis Forel, 1901

• Polyrhachis decemdentata flavipes Stitz, 1910

• Polyrhachis decemdentata gustavi Emery, 1921

• Polyrhachis decemdentata tenuistriata Menozzi, 1933

deceptor Kohout, 2008 (Indonesia, Sulawesi)

decipiens Roger, 1863 (Indonesia)

• Polyrhachis restituta Viehmeyer, 1913

• Polyrhachis restituta conclusa Viehmeyer, 1913

decora Kohout, 2007 (New Guinea)

decumbens Kohout, 2006 (Australia, New Guinea)

delecta Kohout, 2006 (Australia)

delicata Crawley, 1915 (Australia)

• Polyrhachis lysistrata Santschi, 1920

demangei Santschi, 1910 (China, Vietnam)

denselineata Viehmeyer, 1914 (New Guinea)

dentata Donisthorpe, 1947 (New Guinea)

denticulata Karavaiev, 1927 (Australia, Indonesia, New Guinea)

dentihumera Xu, Z., 2002 (China)

derecyna Smith, F., 1871 (New Guinea)

• Polyrhachis dolomedes Smith, F. Smith, F., 1863

• Polyrhachis pseuadonyma Forel, 1886

• Polyrhachis taurus Donisthorpe, 1937

diana Wheeler, W.M., 1909 (Philippines)

diaphanta Smith, F., 1861 (Indonesia, Sulawesi)

dimoculata Kohout, 2013 (Borneo, Indonesia, Malaysia)

diotima Forel, 1911 (Borneo, Indonesia, Malaysia)

dispar Kohout, 2010 (Australia)

diversa Kohout, 2013 (Australia)

dives Smith, F., 1857 (Australia, Borneo, Cambodia, China, Guam, India, Indonesia, Japan, Krakatau Islands, Laos, Malaysia, Myanmar, New Guinea, Philippines, Singapore, Thailand, Vietnam)

• Polyrhachis affinis Smith, F., 1858

• Polyrhachis democles Smith, F., 1861

• Polyrhachis dives euclides Forel, 1913

• Polyrhachis dives siwiensis Santschi, 1932

• Polyrhachis exulans Clark, 1941

• Polyrhachis lucens Donisthorpe, 1947

• Polyrhachis mutiliae Smith, F., 1861

• Polyrhachis vicina Roger, 1863

dives belli Forel, 1912 (India)

dives rectispina Karavaiev, 1927 (Indonesia)

dohrni Forel, 1901 (New Guinea)

dolomedes Smith, F., 1863 (Borneo, Indonesia, Malaysia, New Guinea, Thailand)

• Polyrhachis schang amboinae Santschi, 1928

• Polyrhachis schang gracilior Karavaiev, 1927

dorowi Kohout, 2009 (Australia)

dorsena Kohout, 2006 (New Guinea)

doudou Rigato, 2016 (Gabon)

dougcooki Kohout, 2013 (Australia)

dubia Rigato, 2016 (Cameroun)

durbanensis Forel, 1914 (South Africa)

durvillei Donisthorpe, 1938 (New Guinea)

E
edentula Emery, 1900 (Indonesia, Philippines)

edwardi Donisthorpe, 1948 (New Guinea)

electra Kohout, 2013 (Australia)

elegantula Kohout, 2013 (Australia)

elii Emery, 1900 (Borneo, Indonesia, Malaysia)

emeryana Mann, 1919 (Solomon Islands)

emmae Santschi, 1920 (Laos)

empesoi Kohout, 2006 (Philippines)

enigma Kohout, 2006 (New Guinea)

epinotalis Santschi, 1924 (Central African Republic, Democratic Republic of Congo, Equatorial Guinea, Gabon, Kenya, Rwanda, Uganda, Zambia)

equina Smith, F., 1857 (Borneo, Indonesia, Malaysia)

• Polyrhachis biloba Forel, 1911

erato Forel, 1902 (Australia)

• Polyrhachis aeschyle Forel, 1915

eremita Kohout, 1990 (Australia)

erosispina Emery, 1900 (Indonesia, New Guinea)

esarata Bolton, 1973 (Democratic Republic of Congo, Equatorial Guinea, Ghana)

escherichi Viehmeyer, 1914 (New Guinea)

esuriens Emery, 1897 (New Guinea, Philippines)

• Polyrhachis arcuspinosa Donisthorpe, 1941

• Polyrhachis juxtaspinosa Donisthorpe, 1949

etheli Chapman, 1963 (Philippines)

eudora Smith, F., 1860 (Indonesia)

eureka Kohout, 2013 (Australia)

euryala Smith, F., 1863 (Indonesia, New Guinea)

• Polyrhachis rastellata baduri Donisthorpe, 1941

• Polyrhachis rastellata torricelliana Viehmeyer, 1912

euryala javana Crawley, 1923 (Indonesia)

eurynota Emery, 1897 (New Guinea)

euterpe Forel, 1902 (Australia)

euthiacaena Zhou & Zheng, 1997 (China)

exarata Emery, 1887 (Indonesia, New Guinea)

excellens Viehmeyer, 1912 (New Guinea)

excisa Mayr, 1867 (Indonesia)

excitata Viehmeyer, 1913 (Indonesia, Sulawesi)

exercita (Walker, 1859) (Bangladesh, India, Sri Lanka)

• Polyrhachis clypeatus Mayr, 1862

• Polyrhachis indica Mayr, 1870

exercita lucidiventris Forel, 1907 (India)

exercita obtusisquama Forel, 1902 (India)

exercita rastrata Emery, 1889 (India, Myanmar)

exotica Kohout, 1987 (Philippines)

expressa Kohout, 2006 (Australia)

F
feehani Kohout, 2013 (Australia)

femorata Smith, F., 1858 (Australia)

• Camponotus emeryi Forel, 1879 Forel, 1879

fervens Smith, F., 1860 (Australia, Indonesia, New Guinea, Sulawesi)

• Polyrhachis bicolor Karavaiev Karavaiev, 1927

• Polyrhachis indocilis Santschi, 1928

• Polyrhachis kershawi Clark, 1930

• Polyrhachis valerus Smith, F., 1861

festina Kohout, 2008 (Indonesia, Sulawesi)

fisheri Rigato, 2016 (Gabon)

fissa Mayr, 1902 (Cameroun, Congo, Democratic Republic of Congo, Gabon, Ghana, Uganda)

• Polyrhachis bequaerti Wheeler, W.M., 1922

• Polyrhachis fissa ugandensis Arnold, 1954

flavibasis Clark, 1930 (Australia)

flavicornis Smith, F., 1857 (Borneo, Indonesia, Laos, Malaysia, Singapore, Thailand)

flavoflagellata Karavaiev, 1927 (Borneo, Indonesia, Krakatau Islands, Malaysia)

follicula Menozzi, 1926 (Philippines)

foreli Kohout, 1989 (Australia, New Guinea)

fornicata Emery, 1900 (Indonesia, Sulawesi)

fortis Emery, 1893 (Myanmar)

frauenfeldi Mayr, 1862 (Indonesia)

frauenfeldi sanguinea Forel, 1911 (Borneo, Indonesia, Malaysia)

fruhstorferi Emery, 1898 (Borneo, Indonesia, Malaysia, Philippines, Sulawesi)

fruhstorferi torta Santschi, 1928 (Indonesia)

• Polyrhachis fruhstorferi arcuata Karavaiev, 1927

fulakora Mann, 1919 (Solomon Islands)

fulgens Viehmeyer, 1912 (New Guinea)

furcata Smith, F., 1858 (Borneo, Cambodia, China, India, Indonesia, Malaysia, Myanmar, Philippines, Thailand)

• Polyrhachis furcata bankensis Forel, 1911

• Polyrhachis furcata pahangana Forel, 1911

• Polyrhachis furcata tenella Forel, 1902

furcula Emery, 1911 (New Guinea)

fuscipes Mayr, 1862 (Australia)

• Polyrhachis semipolita hestia Forel, 1911

G
gab Forel, 1879 (Australia, New Guinea)

• Polyrhachis gab tripellis Forel, 1915

gagates Smith, F., 1858 (Angola, Congo, Gabon, Kenya, Malawi, Mozambique, South Africa, Uganda, United Republic of Tanzania, Zambia, Zimbabwe)

• Polyrhachis gagates congolensis Santschi, 1910

• Polyrhachis gagates indefinita Forel, 1913

• Polyrhachis gagates obsidiana Emery, 1921

• Polyrhachis nigriseta Santschi, 1910

• Polyrhachis nigriseta clariseta Santschi, 1910

gamaii Santschi, 1917 (South Africa)

gazelle Kohout, 2013 (New Guinea)

geminata Mann, 1919 (Solomon Islands)

gentilis Kohout, 2007 (New Guinea)

geometrica Smith, F., 1859 (Indonesia, New Guinea)

gerstaeckeri Forel, 1886 (Kenya, United Republic of Tanzania)

• Polyrhachis cubaensis striolatorugosa Mayr, 1893

gestroi Emery, 1900 (Borneo, Indonesia, Malaysia)

gestroi rufiventris Forel, 1911 (Borneo, Indonesia, Malaysia)

• Polyrhachis gestroi moeschiella Forel, 1918

gibba Emery, 1901 (China, Philippines, Sulawesi)

gibbosa Forel, 1908 (Sri Lanka)

gibbula Rigato, 2016 (Kenya, United Republic of Tanzania)

glabrinotum Clark, 1930 (Australia)

glykera Forel, 1912 (Indonesia)

gobini Kohout, 2008 (Indonesia, Sulawesi)

goramensis Emery, 1887 (Indonesia)

• Polyrhachis coronata Santschi, 1928

• Polyrhachis obsidiana Karavaiev, 1927

gracilior Forel, 1893 (India)

• Polyrhachis weberi Donisthorpe, 1943

grandis Donisthorpe, 1949 (New Guinea)

gravis Clark, 1930 (Australia)

greensladei Kohout, 1990 (New Guinea, Solomon Islands)

gressitti Kohout, 2007 (New Guinea)

gribodoi Emery, 1887 (Borneo, Indonesia, Malaysia)

grisescens Emery, 1895 (Jordan, Myanmar)

guerini Roger, 1863 (Australia, New Caledonia)

• Polyrhachis exul Emery, 1883

H
halidayi Emery, 1889 (China, India, Laos, Myanmar, Thailand, Vietnam)

hashimotoi Kohout, 2007 (Borneo, Indonesia, Malaysia)

hastata (Latreille, 1802) (Indonesia)

hauxwelli Bingham, 1903 (India, Jordan, Myanmar, Thailand)

hector Smith, F., 1857 (Borneo, India, Indonesia, Malaysia, Myanmar, Philippines, Singapore)

• Formica ruiginosa Le Guillou, 1842

• Polyrhachis abdominalis reversa André, 1896

• Polyrhachis achilles Forel, 1893

• Polyrhachis malignus Smith, F., 1858

heinlethii Forel, 1895 (Australia)

• Polyrhachis heinlethi sophiae Forel, 1902

heinlethii papuana Emery, 1900 (New Guinea)

hemiopticoides Mukerjee, 1930 (India, Laos, Thailand)

hera Forel, 1911 (Indonesia)

hermione Emery, 1895 (Australia)

• Polyrhachis daemeli sulcativentris Emery, 1915

hespera Kohout, 2013 (Australia)

hexacantha (Erichson, 1842) (Australia)

• Polyrhachis froggatti Forel, 1910

hilaris Kohout, 2008 (Indonesia, Sulawesi)

hippomanes Smith, F., 1861 (Borneo, India, Indonesia, Malaysia, Sulawesi, Thailand)

hippomanes boettcheri Stitz, 1925 (Borneo, Indonesia, Malaysia, Philippines)

hippomanes ceylonensis Emery, 1893 (China, India, Sri Lanka)

hirsuta Mayr, 1876 (Australia)

• Polyrhachis hirsuta quinquedentata Viehmeyer, 1925

hirta Viehmeyer, 1914 (New Guinea)

hispida Kohout, 2008 (Indonesia, Sulawesi)

hodgsoni Forel, 1902 (Borneo, Indonesia, Malaysia, Myanmar)

hoelldobleri Kohout, 2006 (Australia)

hoffmanni Kohout, 2013 (Australia)

hookeri Lowne, 1865 (Australia)

• Polyrhachis cataulacoidea Stitz, 1911

• Polyrhachis hookeri aerea Forel, 1902

horacei Hung, 1967 (New Guinea)

• Polyrhachis cheesmanae Donisthorpe Donisthorpe, 1943

horni Emery, 1901 (India, Sri Lanka)

hortensis Forel, 1913 (Indonesia, New Guinea)

hosei Donisthorpe, 1942 (Borneo, Indonesia, Malaysia)

hostilis Smith, F., 1859 (Indonesia, New Guinea)

• Polyrhachis hostilis hebes Donisthorpe, 1941

• Polyrhachis hostilis hirsuta Viehmeyer, 1914

• Polyrhachis hostilis intricata Forel, 1918

hostilis circumflexa Santschi, 1916 (New Guinea)

• Polyrhachis hostilis arcuata Stitz, 1911

hungi Bolton, 1974 (New Guinea)

• Polyrhachis nitens Donisthorpe Donisthorpe, 1944

hybosa Kohout, 2006 (New Guinea)

I
ignota Kohout, 1987 (Philippines)

illaudata Walker, 1859 (Bangladesh, Borneo, China, India, Indonesia, Krakatau Islands, Laos, Malaysia, Nepal, Philippines, Sri Lanka, Thailand)

• Polyrhachis duodentata Donisthorpe, 1942

• Polyrhachis latispinosa Donisthorpe, 1942

• Polyrhachis mayri Roger, 1863

illaudata intermedia Forel, 1886 (India)

illaudata pauperata Emery, 1889 (Myanmar)

illaudata proximomayri Emery, 1896 (Myanmar)

imitator Kohout, 2008 (Indonesia, Sulawesi)

impressa Kohout, 2007 (New Guinea)

incerta Kohout, 2008 (Australia)

inclusa Viehmeyer, 1912 (New Guinea, Sulawesi)

incognita Kohout, 2008 (Indonesia, Sulawesi)

inconspicua Emery, 1887 (Australia)

• Polyrhachis thalia Forel, 1902

inducta Kohout, 2006 (New Guinea)

inermis Smith, F., 1858 (Borneo, Indonesia, Malaysia, Philippines, Sulawesi, Thailand)

• Polyrhachis bryanti Donisthorpe, 1942

inflata Kohout, 2006 (New Guinea)

injinooi Kohout, 2013 (Australia)

insularis Emery, 1887 (Australia, New Guinea)

• Polyrhachis inconspicua subnitens Emery, 1895

integra Kohout, 2006 (New Guinea)

inusitata Kohout, 1989 (Australia)

io Forel, 1915 (Australia)

isacantha Emery, 1887 (Indonesia)

isolata Kohout, 2013 (Australia)

ithona Smith, F., 1860 (Indonesia, New Guinea, Solomon Islands)

J
jacksoniana Roger, 1863 (Australia)

jacobsoni Forel, 1909 (Indonesia)

javanica Mayr, 1867 (Borneo, Indonesia, Malaysia, Philippines)

jerdonii Forel, 1892 (Sri Lanka)

jianghuaensis Wang, C. & Wu, J., 1991 (China)

johnsoni Mann, 1919 (Solomon Islands)

jurii Karavaiev, 1935 (Cambodia)

K
kaipi Mann, 1919 (Solomon Islands)

kazuoi Kohout, 2008 (Indonesia, Sulawesi)

kebunraya Kohout, 2013 (Indonesia)

keratifera Karavaiev, 1927 (Indonesia)

khepra Bolton, 1973 (Cameroun, Congo, Ghana)

kohli Forel, 1916 (Democratic Republic of Congo)

kohouti Hoffmann, 2015 (Australia)

kokoda Kohout, 2007 (New Guinea)

kyawthani Kohout, 2006 (New Guinea)

L
labella Smith, F., 1860 (Indonesia)

labella brunneipes Wheeler, W.M., 1934 (Solomon Islands)

laboriosa Smith, F., 1858 (Angola, Cameroun, Congo, Democratic Republic of Congo, Equatorial Guinea, Ghana, Ivory Coast, Nigeria, Sierra Leone, Togo, Uganda)

• Polyrhachis hortulana Arnold, 1955

• Polyrhachis laboriosa architecta Santschi, 1924

lachesis Forel, 1897 (Australia)

laciniata Emery, 1900 (New Guinea)

lacteipennis Smith, F., 1858 (Afghanistan, India, Iran, Israel, Nepal, Oman, Turkmenistan, United Arab Emirates)

• Polyrhachis simplex Mayr, 1862

lacteipennis obsoleta Forel, 1893 (India)

laevigata Smith, F., 1857 (Borneo, China, Indonesia, Malaysia)

laevissima Smith, F., 1858 (Bangladesh, Borneo, Cambodia, India, Indonesia, Laos, Malaysia, Myanmar, Thailand)

• Polyrhachis globularia Mayr, 1867

laevissima dichroa Forel, 1893 (India)

lama Kohout, 1994 (China, Indonesia, Philippines, Tibet)

lamellidens Smith, F., 1874 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Taiwan)

laminata Mayr, 1867 (Indonesia)

lanuginosa Santschi, 1910 (Cameroun, Congo)

• Polyrhachis lanuginosa conradti Santschi, 1923

• Polyrhachis lanuginosa felici Emery, 1925

• Polyrhachis lanuginosa santschii Emery, 1921

lata Emery, 1895 (Australia)

• Polyrhachis gab aegra Forel, 1915

latharis Bolton, 1973 (Congo, Ghana)

latinota Viehmeyer, 1912 (New Guinea)

latispina Emery, 1925 (Cameroun, Central African Republic, Democratic Republic of Congo, Equatorial Guinea, Gabon, Kenya, Uganda, United Republic of Tanzania)

• Polyrhachis atalanta Wheeler, W.M. Wheeler, W.M., 1922

• Polyrhachis iperpunctata Menozzi, 1942

latona Wheeler, W.M., 1909 (China, Japan, Taiwan, Thailand, Vietnam)

• Polyrhachis latona dorsorugosa Forel, 1913

latreillii (Guérin-Méneville, 1838) (Australia)

lauta Santschi, 1910 (Cameroun, Congo, Democratic Republic of Congo, Ghana, Uganda)

• Polyrhachis lauta laeta Emery, 1921

• Polyrhachis lauta localis Forel, 1913

leae Forel, 1913 (Australia)

leonidas Forel, 1901 (New Guinea)

leopoldi Santschi, 1932 (Indonesia)

lepida Kohout, 2006 (Borneo, Indonesia, Malaysia)

lestoni Bolton, 1973 (Central African Republic, Congo, Ghana, Kenya)

levior Roger, 1863 (Australia, China, Indonesia, New Guinea)

• Polyrhachis laevissimus Smith, F., 1859

leviuscula Viehmeyer, 1916 (Singapore)

• Polyrhachis schang laurae Menozzi, 1926

lilianae Forel, 1911 (Borneo, Indonesia, Malaysia)

limbata Emery, 1897 (New Guinea)

• Polyrhachis abrupta Karavaiev Karavaiev, 1927

• Polyrhachis verticalis Santschi, 1928

limitis Santschi, 1939 (Democratic Republic of Congo, United Republic of Tanzania)

linae Donisthorpe, 1938 (New Guinea)

litigiosa Emery, 1897 (New Guinea, Solomon Islands)

• Polyrhachis sericata pruinosa Karavaiev, 1927

• Polyrhachis sericata pruinosula Santschi, 1928

lombokensis Emery, 1898 (Indonesia, New Guinea)

longipes Smith, F., 1859 (Indonesia, New Guinea)

longiseta Rigato, 2016 (Cameroun, Congo)

loriai Emery, 1897 (New Guinea)

loweryi Kohout, 1990 (Australia)

lownei Forel, 1895 (Australia)

lucidula Emery, 1893 (Cambodia, China, Myanmar)

luctuosa Emery, 1921 (New Guinea)

lugens Mayr, 1867 (Borneo, Indonesia, Malaysia)

lumi Kohout, 2007 (New Guinea)

luteipes Rigato, 2016 (Cameroun)

luteogaster Kohout, 2012 (New Guinea)

• Polyrhachis alphea rufiventris Emery, 1911

lydiae Forel, 1902 (Australia)

M
maai Kohout, 2007 (New Guinea)

machaon Santschi, 1920 (Australia)

mackayi Donisthorpe, 1938 (Australia)

macropus Wheeler, W.M., 1916 (Australia)

• Polyrhachis longipes Wheeler, W.M. Wheeler, W.M., 1915

maculata Forel, 1915 (Australia)

magnifica Menozzi, 1926 (Philippines)

malaensis Mann, 1919 (Solomon Islands)

maliau Kohout, 2014 (Borneo, Indonesia, Malaysia)

mamba Kohout, 2007 (New Guinea)

manni Kohout, 2008 (Indonesia, Philippines, Sulawesi)

manusensis Kohout, 2013 (New Guinea)

marginata Smith, F., 1859 (Indonesia, New Guinea, Philippines)

• Polyrhachis rotundiceps Karavaiev, 1927

maryatiae Kohout, 2007 (Borneo, Indonesia, Malaysia)

masaokai Kohout, 2008 (Indonesia, Sulawesi)

medusa Forel, 1897 (Kenya, Mozambique, United Republic of Tanzania)

melanura Kohout, 2013 (Australia)

mellita Kohout, 2008 (Indonesia, Sulawesi)

melpomene Emery, 1897 (New Guinea)

• Polyrhachis dolichocephala Viehmeyer, 1914

menelas Forel, 1904 (India)

menozzii Karavaiev, 1927 (Australia, Indonesia, New Guinea)

mentor Forel, 1901 (New Guinea)

metella Smith, F., 1860 (Indonesia, New Guinea)

micans Mayr, 1876 (Australia)

militaris (Fabricius, 1782) (Angola, Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Equatorial Guinea, Gabon, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Liberia, Malawi, Mozambique, Nigeria, Sierra Leone, Sudan, Togo, Uganda, United Republic of Tanzania, Zambia)

• Polyrhachis militaris bruta Santschi, 1912

• Polyrhachis militaris calabarica Forel, 1907

• Polyrhachis militaris cupreopubescens Forel, 1879

• Polyrhachis militaris nkomoensis Santschi, 1924

• Polyrhachis militaris ssibangensis Forel, 1907

• Polyrhachis militaris striativentris Emery, 1892

mindanaensis Emery, 1923 (Philippines)

minima Kohout, 2007 (New Guinea)

mitrata Menozzi, 1932 (Borneo, Indonesia, Malaysia, Philippines)

mjobergi Forel, 1915 (Australia)

• Polyrhachis anguliceps Viehmeyer, 1925

modesta Smith, F., 1857 (Singapore)

moeschi Forel, 1912 (India, Indonesia)

moesta Emery, 1887 (China, Indonesia, Japan)

• Polyrhachis paromalus tobias Forel, 1911

mondoi Donisthorpe, 1938 (New Guinea)

monista Santschi, 1910 (Congo, Democratic Republic of Congo, Gabon, Ghana, Nigeria, Uganda)

montana Hung, 1970 (Borneo, Indonesia, Malaysia)

monteithi Kohout, 2006 (Australia)

monticola Kohout, 2007 (New Guinea)

muara Kohout, 2009 (Borneo, Brunei Darussalam, Indonesia, Malaysia)

mucronata Smith, F., 1859 (Australia, Indonesia, New Guinea, Philippines)

• Polyrhachis cyrtomyrmoides Donisthorpe, 1947

mucronata janthinogaster Emery, 1911 (New Guinea)

mucronata japensis Donisthorpe, 1941 (New Guinea)

muelleri Forel, 1893 (Borneo, Indonesia, Malaysia, Philippines, Singapore, Thailand)

murina Emery, 1893 (Borneo, Indonesia, Malaysia)

mutata Smith, F., 1858 (Borneo, India, Indonesia, Malaysia, Myanmar)

• Polyrhachis mutata ajax Forel, 1893

N
narendrani Karmaly, 2004 (India)

neglecta Kohout, 2008 (Indonesia, Sulawesi)

nepenthicola Kohout, 2013 (Malaysia)

neptunus Smith, F., 1865 (New Guinea, Philippines)

neuguinensis Kohout, 2013 (New Guinea)

nidificans (Jerdon, 1851) (India)

nigra Mayr, 1862 (Sri Lanka)

nigrescens Karavaiev, 1927 (Indonesia)

nigriceps Smith, F., 1863 (New Guinea)

• Polyrhachis atalanta Emery Emery, 1898

• Polyrhachis kirkae Donisthorpe, 1937

nigripes Emery, 1897 (Indonesia, New Guinea, Philippines)

nigrita Mayr, 1895 (Democratic Republic of Congo, Ghana, Uganda)

• Polyrhachis schoutedeni Santschi, 1919

nigropilosa Mayr, 1872 (Borneo, Indonesia, Malaysia, Philippines)

• Polyrhachis nigropilosa conophthalma Emery, 1900

nigropilosa polluta Menozzi, 1926 (Philippines)

nitens Donisthorpe, 1943 (New Guinea)

nitida Smith, F., 1857 (Borneo, Indonesia, Malaysia)

noesaensis Forel, 1915 (Borneo, Indonesia, Malaysia, Philippines)

nofra Bolton, 1975 (Solomon Islands)

nomo Donisthorpe, 1941 (New Guinea)

noonananti Kohout, 2013 (Thailand)

nourlangie Kohout, 2013 (Australia)

nudata Smith, F., 1860 (Indonesia, Sulawesi)

numeria Smith, F., 1861 (India, Indonesia, Sulawesi)

• Polyrhachis schizospina Karavaiev, 1927

O
obesior Viehmeyer, 1916 (Borneo, Indonesia, Malaysia, Philippines)

obliqua Stitz, 1911 (Solomon Islands)

obscura Forel, 1895 (Australia)

obtusa Emery, 1897 (Australia, Indonesia, New Guinea)

• Polyrhachis arcuata aruana Karavaiev, 1927

ochracea Karavaiev, 1927 (Borneo, Indonesia, Malaysia)

oedacantha Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia, Philippines)

ogatai Kohout, 2008 (Indonesia, Sulawesi)

olena Smith, F., 1861 (Indonesia, Sulawesi)

• Polyrhachis eurytus Smith, F., 1861

olybria Forel, 1912 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Philippines, Singapore, Thailand)

omissa Rigato, 2016 (Ethiopia, Kenya, Somalia, Yemen)

omyrmex (Donisthorpe, 1938) (New Guinea)

opacita Kohout, 2013 (Australia)

opalescens Clark, 1930 (Vanuatu)

ops Forel, 1907 (Australia)

orbihumera Xu, Z., 2002 (China)

ornata Mayr, 1876 (Australia)

• Polyrhachis chrysothorax Viehmeyer, 1925

• Polyrhachis humerosa Emery, 1921

orokana Kohout, 2007 (New Guinea)

orpheus Forel, 1911 (Indonesia, Philippines)

orsylla Smith, F., 1861 (Indonesia, Sulawesi)

orsylla javanensis Santschi, 1928 (Indonesia)

• Polyrhachis orsyllus javana Karavaiev, 1927

orsylla muscula Forel, 1901 (Indonesia)

orsylla ritsemai Mayr, 1883 (Indonesia, Krakatau Islands)

osae Mann, 1919 (Solomon Islands)

osiris Bolton, 1975 (Philippines)

otleti Forel, 1916 (Central African Republic, Democratic Republic of Congo, Ghana, Nigeria, United Republic of Tanzania)

P
pacifica Kohout, 2006 (Indonesia, Solomon Islands)

pagana Santschi, 1928 (India)

palaearctica Dietrich, 2004 (Jordan)

palawanensis Kohout, 2013 (Philippines)

pallescens Mayr, 1876 (Australia)

• Polyrhachis aurea depilis Emery, 1897

pallipes Donisthorpe, 1948 (New Guinea)

palmerae Kohout, 2013 (Australia)

parabiotica Chapman, 1963 (Philippines)

paracamponota Wang, C. & Wu, J., 1991 (China)

paromala Smith, F., 1863 (Indonesia)

parva Kohout, 2007 (New Guinea)

patiens Santschi, 1920 (Australia)

paxilla Smith, F., 1863 (Australia, Indonesia, New Guinea)

• Polyrhachis lachesis maeandrifera Emery, 1897

pelecta Kohout, 2007 (New Guinea)

pellita Menozzi, 1922 (Philippines)

• Polyrhachis bubalus Stitz, 1925

penelope Forel, 1895 (Australia, New Guinea)

peregrina Smith, F., 1860 (Indonesia, Sulawesi)

personata Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

phalerata Menozzi, 1926 (Borneo, Cambodia, Indonesia, Laos, Malaysia, Philippines, Thailand)

phidias Forel, 1910 (Central African Republic, Equatorial Guinea, United Republic of Tanzania)

philippinensis Smith, F., 1858 (Philippines)

phryne Forel, 1907 (Australia)

• Polyrhachis sempronia Forel, 1907

• Polyrhachis sidnica perthensis Crawley, 1922

pilbara Kohout, 2013 (Australia)

piliventris Smith, F., 1858 (Singapore)

pilosa Donisthorpe, 1938 (Australia)

pirata Sorger & Zettel, 2009 (Philippines)

placida Kohout, 2013 (Australia)

planata Kohout, 2007 (New Guinea)

planoculata Kohout, 2013 (New Guinea)

plato Forel, 1911 (Indonesia)

platynota Stitz, 1933 (Borneo, Indonesia, Malaysia)

platyomma Emery, 1921 (Cameroun)

polymnia Forel, 1902 (Australia)

porcata Emery, 1921 (New Guinea)

pressa Mayr, 1862 (Borneo, Indonesia, Malaysia, Philippines)

procera Emery, 1897 (New Guinea)

prometheus Santschi, 1920 (Australia)

proxima Roger, 1863 (Cambodia, China, India, Indonesia, Krakatau Islands, Laos, Thailand, Vietnam)

pruinosa Mayr, 1872 (Borneo, Indonesia, Malaysia)

pseudothrinax Hung, 1967 (Australia)

pubescens Mayr, 1879 (Borneo, China, India, Indonesia, Malaysia, Philippines, Thailand)

pulleni Kohout, 2013 (New Guinea)

punctillata Roger, 1863 (China, India, Sri Lanka)

punctillata fergusoni Forel, 1902 (India)

punctillata smythiesii Forel, 1895 (India)

punctiventris Mayr, 1876 (Australia)

punjabi Bharti, 2003 (India)

pyrrhus Forel, 1910 (Australia)

Q
queenslandica Emery, 1895 (Australia, New Guinea)

• Polyrhachis dahli unisculpta Viehmeyer, 1914

R
radicicola Dahl, 1901 (New Guinea)

• Polyrhachis convexa Karavaiev Karavaiev, 1927

• Polyrhachis karawaiewi Santschi, 1928

ralumensis Forel, 1901 (New Guinea)

• Polyrhachis rastellata major Stitz, 1911

rastellata (Latreille, 1802) (Borneo, China, India, Indonesia, Malaysia, New Guinea, Philippines, Sri Lanka, Thailand)

• Polyrhachis rastellata ceylonensis Donisthorpe, 1938

rastellata congener Santschi, 1928 (Indonesia)

reclinata Emery, 1887 (Australia, New Guinea)

regesa Bolton, 1973 (Congo, Ghana)

regularis Mayr, 1867 (Indonesia)

reidi Kohout, 2007 (Borneo, Indonesia, Malaysia)

relucens (Latreille, 1802) (Indonesia)

rere Mann, 1919 (Solomon Islands)

retrorsa Emery, 1900 (Indonesia)

retusa Kohout, 2007 (New Guinea)

revoili André, 1887 (Somalia)

rhea Forel, 1911 (Indonesia)

ridleyi Forel, 1912 (Indonesia)

rixosa Smith, F., 1858 (Borneo, Indonesia, Malaysia, Philippines, Sulawesi, Thailand)

• Polyrhachis lycidas Smith, F., 1861

roberti Kohout, 2007 (New Guinea)

robsoni Kohout, 2006 (Australia)

robusta Kohout, 2013 (New Guinea)

romanovi Santschi, 1928 (Borneo, Indonesia, Malaysia)

• Polyrhachis acantha dichroa Karavaiev, 1927

roomi Kohout, 2007 (New Guinea)

rossi Donisthorpe, 1948 (New Guinea)

rotoccipita Xu, Z., 2002 (China)

rotumana Wilson & Taylor, 1967 (Fiji)

rowlandi Forel, 1910 (Australia)

rubigastrica Wu, J. & Wang, C., 1991 (China)

ruficornis Smith, F., 1857 (Borneo, Indonesia, Malaysia)

rufifemur Forel, 1907 (Australia)

• Polyrhachis terpsichore elegans Forel, 1910

rufipalpis Santschi, 1910 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Ghana, Uganda)

• Polyrhachis rufipalpis mayumbensis Forel, 1913

rufipes Smith, F., 1858 (Borneo, China, Indonesia, Laos, Malaysia, Myanmar, Philippines)

• Polyrhachis exasperata oblisa Forel, 1911

• Polyrhachis exasperatus Smith, F., 1861

• Polyrhachis phipsoni Forel, 1894

rufofemorata Smith, F., 1859 (Australia, Indonesia, New Guinea)

• Polyrhachis biroi Forel, 1907

• Polyrhachis biroi bidentata Stitz, 1912

• Polyrhachis biroi paprika Forel, 1911

• Polyrhachis merops Smith, F., 1860

rufofemorata atra Viehmeyer, 1914 (New Guinea)

rugifrons Smith, F., 1860 (Borneo, Indonesia, Malaysia, Sulawesi)

rupicapra Roger, 1863 (China, India, Sri Lanka)

rustica Kohout, 1990 (Australia)

rutila Kohout, 2006 (Australia)

S
saevissima Smith, F., 1860 (Borneo, India, Indonesia, Malaysia, Philippines, Sulawesi)

• Polyrhachis acantha Smith, F., 1860

• Polyrhachis acasta Smith, F., 1860

saevissima argentea Mayr, 1862 (Borneo, India, Indonesia, Malaysia, Philippines, Solomon Islands)

saevissima chrysophanes Emery, 1900 (Indonesia)

saevissima kerri Forel, 1911 (Thailand)

saevissima timorensis Forel, 1913 (Timor-Leste)

saigonensis Forel, 1886 (Philippines, Vietnam)

salebrosa Kohout, 2008 (Indonesia, Sulawesi)

salomo Forel, 1910 (Solomon Islands)

salomo hiram Forel, 1912 (Solomon Islands)

santschii Mann, 1919 (Solomon Islands)

scabra Kohout, 1987 (Philippines)

scapulata Santschi, 1932 (New Guinea)

schang Forel, 1879 (China, Philippines)

• Polyrhachis gracilis Emery, 1887

schang cnemidata Emery, 1900 (Indonesia)

schang parvicella Forel, 1911 (Indonesia)

schellerichae Dorow, 1996 (Malaysia)

schenckii Forel, 1886 (Australia, New Guinea)

schistacea (Gerstäcker, 1859) (Angola, Cameroun, Congo, Democratic Republic of Congo, Equatorial Guinea, Ethiopia, Gabon, Guinea, Ivory Coast, Kenya, Malawi, Mozambique, Nigeria, Sierra Leone, Somalia, South Africa, Sudan, Togo, Uganda, United Republic of Tanzania, Zimbabwe)

• Polyrhachis carinatus Smith, F., 1858

• Polyrhachis militaris cafrorum Forel, 1879

• Polyrhachis rugulosus Mayr, 1862

• Polyrhachis schistacea atrociliata Santschi, 1913

• Polyrhachis schistacea divina Forel, 1913

• Polyrhachis schistacea divinoides Arnold, 1924

• Polyrhachis schistacea fracta Santschi, 1914

• Polyrhachis schistacea gagatoides Santschi, 1913

schlueteri Forel, 1886 (Kenya, Mozambique, South Africa, United Republic of Tanzania, Zimbabwe)

• Polyrhachis schlueteri indigens Forel, 1914

• Polyrhachis schlueteri plebeia Santschi, 1914

schoopae Forel, 1902 (Australia)

schwiedlandi Forel, 1902 (Australia)

scissa (Roger, 1862) (India, Sri Lanka)

sculpta Emery, 1887 (New Guinea)

sculpturata Smith, F., 1860 (Borneo, India, Indonesia, Malaysia, Philippines, Sulawesi)

• Polyrhachis sumatrensis hamulata Emery, 1887

sculpturata javaniana Santschi, 1928 (Indonesia, Philippines)

• Polyrhachis sculpturata javana Stitz, 1925

sculpturata siamensis Mayr, 1879 (Thailand)

scutulata Smith, F., 1859 (Indonesia, New Guinea)

sedlaceki Kohout, 2006 (New Guinea)

seducta Kohout, 2013 (Australia)

selecta Forel, 1911 (Indonesia)

semiaurata Mayr, 1876 (Australia)

semiinermis Donisthorpe, 1941 (Philippines)

semiobscura Donisthorpe, 1944 (Australia, Indonesia, New Guinea)

semipolita André, 1896 (Australia)

semitestacea Emery, 1900 (New Guinea)

senilis Forel, 1902 (Australia)

• Polyrhachis comata Crawley, 1915

• Polyrhachis crawleyella Santschi, 1916

sericata (Guérin-Méneville, 1831) (New Guinea, Vanuatu)

• Formica grisea Le Guillou, 1842

sericata glabra Forel, 1911 (New Guinea)

sericata nitidissima Karavaiev, 1927 (Indonesia)

sericata nitidiventris Stitz, 1911 (New Guinea)

sericeopubescens Donisthorpe, 1941 (New Guinea)

• Polyrhachis conops cuspidata Stitz, 1911

setosa Kohout, 2006 (Solomon Islands)

sexspinosa (Latreille, 1802) (Australia, Indonesia, New Caledonia, New Guinea, Philippines)

• Formica argentata Fabricius, 1804

• Polyrhachis barnardi Clark, 1928

• Polyrhachis irritabilis Smith, F., 1859

• Polyrhachis sexspinosa sericea Karavaiev, 1927

shattucki Kohout, 2013 (Australia)

shixingensis Wu, J. & Wang, C., 1995 (China)

sidnica Mayr, 1866 (Australia)

• Polyrhachis quadricuspis Mayr, 1870

similis Viehmeyer, 1912 (New Guinea, Solomon Islands)

similis angustior Viehmeyer, 1912 (New Guinea)

simillima Emery, 1900 (Indonesia)

simpla Santschi, 1928 (Indonesia, New Guinea)

• Polyrhachis conops simplex Karavaiev, 1927

simulans Kohout, 2007 (New Guinea)

sinuata Kohout, 2013 (New Guinea)

smithersi Kohout, 2012 (Australia)

smithi Emery, 1901 (Indonesia, Sulawesi)

snellingi Kohout, 2007 (New Guinea)

sokolova Forel, 1902 (Australia, New Caledonia, New Guinea)

• Polyrhachis sokolova degener Forel, 1910

solivaga Menozzi, 1926 (Philippines)

solmsi Emery, 1887 (Indonesia)

• Polyrhachis alexandri Karavaiev, 1906

solmsi multicella Forel, 1911 (Indonesia)

sophocles Forel, 1908 (Sri Lanka)

sparaxes Smith, F., 1863 (Indonesia)

spengeli Forel, 1912 (Indonesia)

spinicola Forel, 1894 (Kenya, Mozambique, South Africa)

• Polyrhachis cubaensis gallicola Forel, 1894

spinifera Stitz, 1911 (New Guinea)

• Polyrhachis continua hirsutula Emery, 1911

spinigera Mayr, 1879 (India)

spinosa Mayr, 1867 (Indonesia)

spitteleri Forel, 1916 (Central African Republic, Democratic Republic of Congo, Ghana)

splendens Santschi, 1932 (New Guinea)

starri Kohout, 2013 (Philippines)

stigmatifera Kohout, 1990 (Indonesia)

stitzi Santschi, 1928 (New Guinea)

• Polyrhachis conops bismarckensis Karavaiev, 1927

storki Kohout, 2008 (Borneo, Indonesia, Malaysia, Sulawesi)

striata Mayr, 1862 (Borneo, China, India, Indonesia, Malaysia, Philippines, Thailand)

• Polyrhachis striata assamensis Forel, 1902

striatorugosa Mayr, 1862 (Borneo, India, Indonesia, Malaysia)

• Polyrhachis striatorugosa exophthalma Forel, 1913

stricta Kohout, 2013 (Australia)

strictifrons Emery, 1898 (Indonesia, Sulawesi)

strumosa Kohout, 2006 (New Guinea)

stylifera Karavaiev, 1935 (Cambodia)

subaenescens Viehmeyer, 1912 (New Guinea)

subcyanea Emery, 1897 (New Guinea)

subcyanea rotundinota Viehmeyer, 1914 (New Guinea)

subfossa Viehmeyer, 1913 (Indonesia, Sulawesi)

subfossoides Karavaiev, 1927 (Indonesia)

submarginata Rigato, 2016 (Central African Republic)

subpilosa Emery, 1895 (China, India, Myanmar)

subtridens Emery, 1900 (Indonesia)

sukarmani Kohout, 2007 (Borneo, Indonesia, Malaysia)

sulang Kohout, 2006 (Borneo, Indonesia, Malaysia)

sulawesiensis Kohout, 2008 (Indonesia, Sulawesi)

sulcata André, 1895 (Congo, Equatorial Guinea, Ghana)

sulcifera Kohout, 2007 (New Guinea)

sylvicola (Jerdon, 1851) (India)

T
tambourinensis Forel, 1915 (Australia)

tanami Kohout, 2013 (Australia)

tapini Kohout, 2013 (New Guinea)

taylori Kohout, 1988 (New Guinea)

templi Forel, 1902 (Australia)

tenebra Kohout, 2013 (Australia)

terminata Rigato, 2016 (United Republic of Tanzania)

terpsichore Forel, 1893 (Australia)

tersa Viehmeyer, 1914 (New Guinea)

textor Smith, F., 1857 (Malaysia, Thailand)

textor brunneogaster Donisthorpe, 1937 (India, Nicobar Island)

textor charpillioni Forel, 1911 (Indonesia)

textor hero Forel, 1913 (Indonesia)

thailandica Kohout, 2006 (Thailand)

thais Forel, 1910 (Australia)

thompsoni Bingham, 1903 (China, India, Myanmar)

thrinax Roger, 1863 (Bangladesh, China, India, Philippines, Sri Lanka, Thailand)

• Polyrhachis thrinax mucronis Donisthorpe, 1942

thrinax castanea Stitz, 1925 (Philippines)

• Polyrhachis thrinax castanella Santschi, 1928

thrinax inconstans Viehmeyer, 1916 (Singapore)

thrinax lancearia Forel, 1893 (India)

thrinax lucida Emery, 1894 (Malaysia)

• Polyrhachis thrinax lucidula Emery, 1893

thrinax overbecki Dorow, 1995 (Philippines, Singapore)

• Polyrhachis thrinax nigripes Viehmeyer, 1916

thusnelda Forel, 1902 (Australia)

tianjingshanensis Quin & Zhou, 2008 (China)

tibialis Smith, F., 1858 (Bangladesh, Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Thailand)

tibialis addax Santschi, 1928 (Indonesia)

tibialis caligata Emery, 1895 (Bangladesh, India, Indonesia, Myanmar, Vietnam)

tibialis completa Santschi, 1928 (Indonesia)

tibialis crassisquama Forel, 1913 (Malaysia)

tibialis nigricornis Santschi, 1928 (Indonesia)

tibialis orientalis Karavaiev, 1927 (Indonesia)

tibialis parsis Emery, 1900 (India)

tibialis pectita Santschi, 1928 (Sri Lanka)

tibialis robustior Karavaiev, 1927 (Indonesia)

tragos Stitz, 1925 (Philippines)

transiens Bolton, 1973 (Uganda)

trapezoidea Mayr, 1876 (Australia)

triaena Wheeler, W.M., 1919 (Borneo, Indonesia, Malaysia)

tricuspis André, 1887 (Indonesia)

• Polyrhachis ternatae Karavaiev, 1933

trina Donisthorpe, 1944 (New Guinea)

trispinosa Smith, F., 1861 (Indonesia, Sulawesi)

tristis Mayr, 1867 (Borneo, Indonesia, Malaysia)

trophima Smith, F., 1863 (Indonesia)

tschu Forel, 1879 (China)

tubericeps Forel, 1893 (Bangladesh, India)

tuberosa Kohout, 2006 (New Guinea)

tubifera Forel, 1902 (Australia)

tubifex Karavaiev, 1926 (Borneo, Indonesia, Malaysia, Philippines)

turneri Forel, 1895 (Australia)

tyrannica Smith, F., 1858 (Borneo, China, Indonesia, Malaysia)

U
ugiensis Mann, 1919 (Solomon Islands)

ulysses Forel, 1910 (Solomon Islands)

umboi Kohout, 2007 (New Guinea)

uncaria Kohout, 2013 (Australia)

undulata Kohout, 2006 (Solomon Islands)

unicornis Kohout, 2013 (Australia)

unicuspis Emery, 1898 (Indonesia, Sulawesi)

V
vanachterbergi Kohout, 2008 (Indonesia, Sulawesi)

varicolor Viehmeyer, 1916 (Singapore, Singapore, Thailand)

variegata Kohout, 2007 (New Guinea)

venus Forel, 1893 (Myanmar, Philippines)

vermiculosa Mayr, 1876 (Australia)

vernoni Kohout, 2013 (Australia)

vestita Smith, F., 1860 (Indonesia, Sulawesi)

• Polyrhachis vestita unicolor Emery, 1898

viehmeyeri Emery, 1921 (New Guinea)

vigilans Smith, F., 1858 (China, Vietnam)

• Polyrhachis pyrgops Viehmeyer, 1912

villipes Smith, F., 1857 (Borneo, India, Indonesia, Malaysia, Philippines, Thailand)

• Polyrhachis striata tritschleri Forel, 1912

• Polyrhachis sumatrensis Smith, F., 1858

villosa Emery, 1897 (New Guinea)

villosa pubiventris Viehmeyer, 1914 (New Guinea)

vindex Smith, F., 1857 (Borneo, Indonesia, Malaysia, Philippines)

viola Zettel, 2013 (Philippines)

violaceonigra Viehmeyer, 1914 (New Guinea)

viscosa Smith, F., 1858 (Burkina Faso, Democratic Republic of Congo, Eritrea, Ethiopia, Ghana, Kenya, Malawi, Nigeria, Somalia, South Africa, Sudan, Uganda, United Republic of Tanzania, Zimbabwe)

• Polyrhachis antinorii Emery, 1877

• Polyrhachis cubaensis imatongica Weber, 1943

• Polyrhachis viscosa spretula Santschi, 1923

vitalisi Santschi, 1920 (Laos)

volkarti Forel, 1916 (Democratic Republic of Congo)

W
wagneri Viehmeyer, 1914 (New Guinea)

waigeuensis Donisthorpe, 1943 (New Guinea)

wallacei Emery, 1887 (India, Indonesia, Sulawesi)

wallacei warburgi Forel, 1901 (Indonesia)

wamuki Kohout, 2007 (New Guinea)

wardi Kohout, 2007 (New Guinea)

watanasiti Kohout, 2013 (Thailand)

weiri Kohout, 2013 (Australia)

weissi Santschi, 1910 (Cameroun, Central African Republic, Congo, Democratic Republic of Congo, Ghana, Guinea, South Africa, Zambia)

• Polyrhachis natalensis Santschi, 1914

• Polyrhachis revoili balli Santschi, 1939

• Polyrhachis revoili conduensis Forel, 1915

• Polyrhachis revoili crassa Emery, 1921

• Polyrhachis revoili donisthorpei Forel, 1916

wellmani Forel, 1909 (Angola, Democratic Republic of Congo, Zambia)

wheeleri Mann, 1919 (Borneo, Indonesia, Malaysia, Solomon Islands)

widodoi Kohout, 2006 (Borneo, Indonesia, Malaysia)

wilmsi Forel, 1910 (Mozambique, Somalia, South Africa)

wilsoni Kohout, 2007 (New Guinea)

wolfi Forel, 1912 (China, Taiwan)

wroughtonii Forel, 1894 (India)

X
xanthippe Forel, 1911 (Sri Lanka)

xiphias Smith, F., 1863 (Indonesia, New Guinea)

Y
yarrabahensis Forel, 1915 (Australia, Indonesia)

yerburyi Forel, 1893 (Sri Lanka)

yorkana Forel, 1915 (Australia)

ypsilon Emery, 1887 (Borneo, Brunei Darussalam, Indonesia, Malaysia, Singapore)

• Polyrhachis synacantha Santschi, 1933

• Polyrhachis ypsilon victoris Santschi, 1925

Z
zhengi Zhou & Huang, 2002 (China)

zimmerae Clark, 1941 (Australia)

zopyra Smith, F., 1861 (Indonesia, Philippines, Sulawesi)

• Polyrhachis aurichalceus Mayr, 1862

• Polyrhachis moorei Donisthorpe, 1941

zopyra imbellis Emery, 1887 (Indonesia)

=Fossil Species=

A
annosa Wappler, Dlussky & Reuter, 2009