Myopopone castanea

These rare ants, the only species in the genus Myopopone, nest in rotten wood or under bark. Single colonies may be composed of several small nests scattered over a small area. They feed on large, soft-bodied insect larvae and may bring their larvae to food sources rather than attempt to move especially large prey back to their nest.

Identification
The mandibles are long and slender, with numerous (always more than 5) teeth which vary greatly in size and are scattered along the inner margins, and with a sharp, pointed tooth at the tip which is only slightly longer than the next longest tooth. The frontal lobes are large and extend well forward of the insertion point of the scapes, and when viewed from the front they cover the underlying clypeus and often form part of the front margin of the head. The antennae have the last few segments distinctly flattened in cross-section. The petiole has distinct front and upper faces but lacks a rear face, and its attachment to the gaster is broad and approximately the same height as the petiole so that the upper surfaces of petiole and gaster are separated by at most a shallow impression.

Although these ants are superficially similar to some Amblyopone, the presence of expanded and projecting frontal lobes and flattened tips of the antennae will separate these genera.

Distribution based on Regional Taxon Lists
Australasian Region: Australia. Indo-Australian Region: Borneo, Brunei Darussalam, Indonesia, Malaysia, New Guinea, Philippines, Singapore, Solomon Islands, Timor-Leste. Oriental Region: Cambodia, India, Laos, Myanmar, Nicobar Island, Sri Lanka, Thailand, Vietnam. Palaearctic Region: China.

Biology
Wilson (1958) - Both the Karema and Bisianumu collections consisted of workers found under the thick bark of large rotting logs on the floor of rain forests. At Bisianumu workers were clustered with larvae around two large, freshly killed cerambycid larvae on the same log. Since the beetle larvae were well separated from one another, and appeared to be too large for the ants to transport through the preformed galleries under the bark, it is inferred that the ants had transferred their own larvae to feed on the prey after the latter had been attacked and killed. The Myopopone are singularly clumsy and shy ants, and immediately commence searching for cover when exposed to light, abandoning their brood in the process. When handled, however, they are capable of inflicting a painful sting.

Ito (2010) - Workers kept with larvae in the laboratory frequently showed larval hemolymph feeding (hereafter LHF). This behavior is very similar to that of queens of Stigmatomma silvestrii Wheeler as reported by Masuko (1986). Workers pinched the larvae with their mandibles and licked hemolymph from the wounds. Workers most often pinched the dorsal integument of the upper abdomen of larvae. Many large larvae had scars which seem to be made by LHF. During 14 h of observations of an orphan colony with 25 workers, LHF was observed 38 times. Twelve of the 25 workers performed LHF. One worker monopolized more than 50% of LHF (21 times) and the others only showed the behavior one to three times. Among workers, we never observed aggressive behavior. The relationship between LHF and ovarian development could not be examined, because colony condition was unfortunately poor at the end of the observation. During the observation period, no egg-laying occurred.



Among amblyoponine ants, the occurrence of LHF has been reported in Amblyopone spp., Prionopelta kraepelini, Adetomyrma sp., Mystrium camillae and Onychomyrmex (Ito & Billen 1998; Masuko 2003; Saux et al. 2004; F. Ito, unpubl. data, 1989, 2002). Even though we still do not know whether queens of M. castanea perform LHF, all genera in this subfamily so far observed show LHF. Although LHF has been found in a handful of ant species outside of Amblyoponinae, e.g. a few species of Proceratium (Proceratiinae), a few species of Gnamptogenys (Ectatomminae), Leptanilla japonica and Leptanilla clypeata (Leptanillinae), and Calyptomyrmex sp. (Myrmiciinae) (Masuko 1986, 1989; Ito 2001; Ito & Gobin 2008; F. Ito, unpubl. data, 2001), this aberrant feeding mode is one of the important characteristics of the subfamily Amblyoponinae.

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Nomenclature

 * . Amblyopone castaneus Smith, F. 1860b: 105, pl. 1, fig. 6 (w.) INDONESIA (Bacan I.).
 * Type-material: 2 syntype workers.
 * Type-locality: Indonesia: Bachian (= Bacan) I., “Bac” (A.R. Wallace).
 * Type-depository: OXUM.
 * Mayr, 1867a: 90 (q.); Wheeler, G.C. & Wheeler, J. 1964b: 446 (l.).
 * Combination in Myopopone: Roger, 1862c: 292.
 * Status as species: Roger, 1862c: 292; Smith, F. 1863: 18; Roger, 1863b: 20; Mayr, 1867a: 90 (redescription); Smith, F. 1871a: 324; Emery, 1887b: 447; Emery, 1888a: 531; Dalla Torre, 1893: 15; Emery, 1897d: 547; Forel, 1900c: 54; Emery, 1900d: 661; Forel, 1901b: 5; Dahl, 1901: 18; Bingham, 1903: 33; Emery, 1911d: 26; Viehmeyer, 1912: 4; Forel, 1913k: 5; Mann, 1919: 281; Stitz, 1925: 110; Karavaiev, 1930a: 212; Donisthorpe, 1932c: 461; Menozzi, 1932d: 3; Wheeler, W.M. 1935g: 9; Donisthorpe, 1942c: 31; Chapman & Capco, 1951: 22; Wilson, 1958a: 144; Brown, 1960a: 213; Taylor & Brown, 1985: 35; Taylor, 1987a: 41; Bolton, 1995b: 270; Jaitrong & Nabhitabhata, 2005: 28; Mohanraj, et al. 2010: 6; Pfeiffer, et al. 2011: 32; Xu & He, 2011: 232 (redescription); Sarnat, et al. 2013: 69; Bharti, Guénard, et al. 2016: 18; Jaitrong, Guénard, et al. 2016: 22.
 * Senior synonym of bakeri: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of beccarii: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of bugnioni: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of maculata: Bingham, 1903: 33; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of moelleri: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of picea: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of proxima: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of rossi: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of rufula: Roger, 1862c: 292; Roger, 1863b: 20; Mayr, 1863: 430; Forel, 1900c: 54; Bingham, 1903: 33; Dalla Torre, 1893: 15; Emery, 1911d: 26; Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of similis: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of smithi: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of striatifrons: Brown, 1960a: 215; Bolton, 1995b: 270.
 * Senior synonym of wollastoni: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * Distribution: Australia, China, India (+ Andaman Is), Indonesia (Bacan, Irian Jaya, Java, Lombok, Sulawesi, Sumatra, Ternate), Laos, Malaysia (Peninsula, Sabah, Sarawak), Papua New Guinea, Philippines (Bataan, Leyte, Luzon, Negros, Palawan), Solomon Is, Sri Lanka, Thailand.
 * bakeri. Myopopone castanea var. bakeri Viehmeyer, 1916b: 283 (w.) PHILIPPINES (Leyte I.).
 * Type-material: holotype worker.
 * Type-locality: Philippines: Leyte, Tacloban (C.F. Baker).
 * Type-depository: MNHU.
 * Subspecies of castanea: Wheeler, W.M. & Chapman, 1925: 57; Chapman & Capco, 1951: 23; Baltazar, 1966: 233.
 * Subspecies of moelleri: Donisthorpe, 1942c: 31.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * beccarii. Myopopone beccarii Emery, 1887b: 447 (w.) INDONESIA (Ternate I.).
 * Type-material: syntype workers (number not stated).
 * Type-locality: Indonesia: Ternate I. (Beccari).
 * Type-depository: MSNG.
 * Subspecies of castanea: Emery, in Dalla Torre, 1893: 15; Emery, 1911d: 26; Donisthorpe, 1942c: 31; Chapman & Capco, 1951: 23.
 * Status as species: Wilson, 1958a: 143.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * bugnioni. Myopopone castanea var. bugnioni Forel, 1913k: 5 (footnote) (q.m.) SRI LANKA.
 * Type-material: syntype workers (number not stated), 1 syntype queen, 1 syntype male.
 * Type-locality: Sri Lanka (“Ceylon”): Peradeniya (Bugnion).
 * Type-depository: MHNG. NHMB.
 * Subspecies of moelleri: Forel, 1915a: 22; Donisthorpe, 1942c: 31.
 * Subspecies of castanea: Chapman & Capco, 1951: 23.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * maculata. Myopopone maculata Roger, 1861a: 50 (w.q.) SRI LANKA.
 * Type-material: syntype workers (number not stated), 1 syntype queen.
 * Type-locality: workers Sri Lanka (“Ceylon”), queen Indonesia: Bintam (= Sumatra, Riau Archipelago, Bintan I.?).
 * [Note: Roger gives no locality data for the worker in the original description; Sri Lanka is provided by Roger, 1863b: 20.]
 * Type-depository: MNHU.
 * Donisthorpe, 1942c: 29 (m.).
 * Status as species: Mayr, 1863: 430; Roger, 1863b: 20; Mayr, 1865: 73; Emery, 1895k: 456.
 * Subspecies of castanea: Emery, in Dalla Torre, 1893: 15; Emery, 1893f: 240; Emery, 1900d: 661; Forel, 1900c: 54; Forel, 1903d: 399; Forel, 1907a: 17; Forel, 1907e: 17; Emery, 1911d: 26; Wheeler, W.M. 1913e: 233; Wheeler, W.M. 1919e: 50; Santschi, 1920h: 158; Santschi, 1924c: 95; Wheeler, W.M. & Chapman, 1925: 57; Wheeler, W.M. 1929g: 29; Donisthorpe, 1942c: 29; Chapman & Capco, 1951: 23; Baltazar, 1966: 233.
 * Junior synonym of castanea: Bingham, 1903: 33; Brown, 1960a: 215; Bolton, 1995b: 270.
 * moelleri. Myopopone moelleri Bingham, 1903: 34 (q.) INDIA (Sikkim).
 * Type-material: holotype queen.
 * Type-locality: India: Sikkim, 7000 ft, at light (C.T. Bingham).
 * Type-depository: OXUM.
 * [Misspelled as mulleri by Santschi, 1924c: 95.]
 * Donisthorpe, 1942c: 30 (m.).
 * Status as species: Santschi, 1924c: 95; Stitz, 1925: 110; Santschi, 1932b: 11; Donisthorpe, 1942c: 31.
 * Subspecies of castanea: Emery, 1911d: 26; Menozzi, 1939a: 327; Chapman & Capco, 1951: 23.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * picea. Myopopone picea Donisthorpe, 1938e: 498 (w.) BORNEO (East Malaysia: Sarawak).
 * Type-material: holotype worker.
 * Type-locality: Malaysia: Borneo, Sarawak, Mt Dulit, junction of Rivers Tinjar and Lejok, 9.viii.1932 (Oxford Univ. Expd.).
 * Type-depository: BMNH.
 * Status as species: Donisthorpe, 1942c: 31; Chapman & Capco, 1951: 23.
 * Junior synonym of castanea: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * proxima. Myopopone castanea var. proxima Stitz, 1925: 110 (w.) PHILIPPINES (Luzon I., Palawan I.).
 * Type-material: syntype workers (number not stated, “a number”).
 * Type-localities: Philippines: Luzon, Limay (Boettcher), Luzon, Los Baños (Boettcher), Palawan, Binaluan (Boettcher).
 * Type-depository: MNHU.
 * Subspecies of castanea: Chapman & Capco, 1951: 23.
 * Junior synonym of bakeri: Donisthorpe, 1942c: 31.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * rossi. Myopopone rossi Donisthorpe, 1948b: 297 (w.) NEW GUINEA (Papua New Guinea).
 * Type-material: 24 syntype workers.
 * Type-locality: Papua New Guinea: Finschhafen, 20.iv.1944 (E.S. Ross).
 * Type-depository: CASC.
 * Status as species: Donisthorpe, 1948g: 131; Donisthorpe, 1949g: 403; Donisthorpe, 1950a: 338; Chapman & Capco, 1951: 23.
 * Junior synonym of castanea: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * rufula. Myopopone rufula Roger, 1861a: 52 (w.) INDONESIA (Bacan I.).
 * Type-material: 2 syntype workers.
 * Type-locality: Indonesia: Batchian (= Bacan) I.
 * Type-depository: MNHU.
 * As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 23.
 * Junior synonym of castanea: Roger, 1862c: 292; Mayr, 1863: 430; Roger, 1863b: 20; Forel, 1900c: 54; Bingham, 1903: 33; Dalla Torre, 1893: 15; Emery, 1911d: 26; Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * similis. Myopopone similis Donisthorpe, 1949b: 488 (q.) NEW GUINEA (Indonesia).
 * Type-material: holotype queen.
 * Type-locality: Indonesia: Irian Jaya (“Dutch New Guinea”), Maffin Bay, viii.1944 (E.S. Ross).
 * Type-depository: CASC.
 * Status as species: Chapman & Capco, 1951: 24.
 * Junior synonym of castanea: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.
 * smithi. Myopopone smithi Donisthorpe, 1947c: 577 (q.) NEW GUINEA (Papua New Guinea).
 * Type-material: holotype queen.
 * Type-locality: Papua New Guinea: Nazdab, Markham River valley, vi.1944 (K.V. Krombein).
 * Type-depository: USNM.
 * Junior synonym of castanea: Wilson, 1958a: 144; Bolton, 1995b: 270.
 * striatifrons. Myopopone moelleri var. striatifrons Stitz, 1925: 110 (q.) INDONESIA (Lombok I., Sumatra).
 * Type-material: 3 syntype queens.
 * Type-localities: Indonesia: Lombok (Fruhstorfer), and Indonesia: W Sumatra, Padang (Schoede).
 * Type-depository: MNHU.
 * As unavailable (infrasubspecific) name: Chapman & Capco, 1951: 23.
 * Subspecies of moelleri: Donisthorpe, 1942c: 31.
 * Junior synonym of castanea: Brown, 1960a: 215; Bolton, 1995b: 270.
 * wollastoni. Myopopone wollastoni Donisthorpe, 1942c: 29 (q.) NEW GUINEA (Indonesia).
 * Type-material: 2 syntype queens.
 * Type-locality: Indonesia: Irian Jaya, Mimika River, vii.1910 (A.F.R. Wollaston).
 * Type-depository: BMNH.
 * Status as species: Chapman & Capco, 1951: 24.
 * Junior synonym of castanea: Wilson, 1958a: 144; Brown, 1960a: 215; Bolton, 1995b: 270.

Worker
Length 41/2 lines. Ferruginous; head wider than the thorax, slightly rounded at the sides, and emarginate behind, finely and distinctly punctured above, anteriorly it is longitudinally striated; the anterior margin fuscous; the antennae short and stout; the flagellum clavate; the mandibles with a row of short, stout, acute teeth on their inner margin; the head has a few scattered erect hairs. Thorax: the prothorax subglobose, strongly punctured in front; the mesothorax short and transverse; the metathorax oblong, parallel, and punctured; the apex transversely striated; the legs short, stout, and pubescent. Abdomen: the basal segment strongly punctured, the second and fullowing segments delicately and very sparingly so; the first and second segments deeply constricted at their margins; the apex pointed and pubescent, the pubescence ferruginous.

Xu and He (2011) - L 7.3-8.7, HL 1.57-1.90, HW 1.63-2.00, CI 104-105, SL 0.80-0.93, SI 47-49, ML 1.10-1.20, PW 1.03-1.30, AL 2.20-2.67, ED 0.15-0.18, PH 0.73-0.83, PL 1.03-1.20, LPI 69-73, DPW 0.83-1.03, DPI 81-86 (3 individuals measured).

Head in full face view rectangular, about as long as broad, slightly widen forward. Occipital margin evenly concave. Occipital corners roundly prominent. Lateral sides weakly convex. Clypeus narrow and transverse, anterior margin straight, with a row of about 6 minute denticles. Frontal lobes developed and relatively close together, concealed antennal sockets, protruding forward divergently and surpassed anterior margin of clypeus. Mandibles narrow and linear. Masticatory margin distinct, with 4 teeth, about 1/2 length of inner margin. Inner margin with 5 teeth. Frontal carinae short, extending backward to midline of head. Antennae short and stout, apexes of scapes reached to 1/2 of the distance from antennal sockets to occipital corners. Antennal clubs 6-segmented, incrassate and depressed. Eyes small, with about 15 facets, well behind the midline of the head. Ocelli absent.

In profile view, dorsum of alitrunk relatively straight, lateral sides nearly vertical. Pronotum long, weakly convex. Promesonotal suture and metanotal groove distinctly notched. Mesonotum very short and transverse. Dorsum of propodeum straight, longer than declivity, about as long as pronotum. Declivity roundly convex. Propodeal lobes small, rounded at apex. Propodeal spiracles elliptic and vertical, located at center of the lateral sides. In profile view, petiolar node nearly trapezoid, broadly attached to first gastral tergite. Dorsal face weakly convex, longer than anterior face. Anterior face weakly depressed. Ventral face weakly concave. Subpetiolar process small, nearly rectangular. In dorsal view, petiolar node broader than long, slightly widen backward, anterior margin straight, lateral sides moderately convex. Constriction between the two basal gastral segments distinct. Sting developed and extruding.

Mandibles smooth, with sparse punctures. Clypeus densely transversely striate. Dorsum of head smooth, with sparse punctures, but densely longitudinally striate on anterior portion. Ventral face of head and frontal lobes sparsely longitudinally striate. Lateral sides of alitrunk densely longitudinally striate. Dorsum of alitrunk smooth, with sparse punctures. Posterior 2/3 of propodeal dorsum densely punctate, declivity transversely striate. Dorsal and anterior surfaces of petiole smooth, with sparse punctures, lateral sides longitudinally striate. Dorsum of peduncle transversely striate. Gaster smooth, with sparse punctures.

Head and body with sparse erect to subdecumbent hairs. Clypeus, propodeum, petiole, and gaster with abundant hairs. Gastral apex with dense hairs. Antennal flagella, propodeum, and dorsum of petiole with dense decumbent pubescence. Antennal scapes and hind tibiae with sparse subdecumbent hairs. Outer surfaces of tarsi and middle tibiae with strong spines. Color brownish red. Mandibles, clypeus, frontal lobes, and antennal flagella reddish brown. Eyes grey. Hairs golden yellow.

Queen
Xu and He (2011) - TL 13.7, HL 2.47, HW 2.63, CI 107, SL 1.17, SI 44, ML 1.60, PW 2.07, AL 4.33, ED 0.63, PH 1.27, PL 1.93, LPI 66, DPW 1.70, DPI 88 (1 individual measured). Head similar to the worker, in full face view slightly broader than long, not narrowed forward. Anterior margin of clypeus straight, without denticles. Eyes large, located at midpoints of lateral sides of head. With 3 ocelli. In profile view, alitrunk massive, dorsum nearly straight, mesonotum weakly convex. Propodeal dorsum shorter than declivity, the latter weakly convex. In dorsal view, mesonotum with a pair of oblique longitudinal furrows on anterior portion, and a longitudinal central furrow on posterior portion of scutum. Transverse groove between scutum and scutellum distinct and straight. Posterior margin of scutellum concave. Metanotum crescent, narrow and transverse. Petiole and gaster similar to the worker.

Sculptures similar to the worker, but occipital margin longitudinally striate, lateral sides of promesothorax longitudinal striate and abundantly punctured, propodeal dorsum densely punctuate. Pilosity similar to the worker, but anterior face of petiolar node without hairs, hind tibiae with sparse subdecumbent hairs and abundant decumbent pubescence. Color black. Mandibles, clypeus, apexes of frontal lobes, antennae, and coxae reddish brown. Tibiae, tarsi, and gastral apex brownish red. Ocelli light yellow. Hairs golden yellow.

References based on Global Ant Biodiversity Informatics

 * Brown W. L., Jr. 1960. Contributions toward a reclassification of the Formicidae. III. Tribe Amblyoponini (Hymenoptera). Bulletin of the Museum of Comparative Zoology 122: 143-230.
 * Bui T.V. 2002. Result of ant survey in Tam Dao National Park. Hoi Nghi Con Trung Hoc Toan Quoc, Ha Noi 495-498.
 * CSIRO Collection
 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Chapman, J.W. and S.R. Capco. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monographs of the Institute of Science and Technology (Manila) 1: 1- 327
 * Dahl F. 1901. Das Leben der Ameisen im Bismarck-Archipel, nach eigenen Beobachtungen vergleichend dargestellt. Mitt. Zool. Mus. Berl. 2: 1-70.
 * Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
 * Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
 * Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
 * Donisthorpe H. 1938. New species of ants and a new subgenus of Dolichoderus from various localities. Annals and Magazine of Natural History (11)2: 498-504.
 * Donisthorpe H. 1942. Myopopone wollastoni sp. n., with notes on other forms in the genus and descriptions of the males of two species (Hym., Formicidae). Entomologist's Monthly Magazine 78: 29-31.
 * Donisthorpe H. 1947. Ants from New Guinea, including new species and a new genus. Annals and Magazine of Natural History (11)13: 577-595.
 * Donisthorpe H. 1948. A fourth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)1: 131-143.
 * Donisthorpe H. 1948. A second instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (11)14: 297-317.
 * Donisthorpe H. 1949. A fifth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)1: 487-506.
 * Donisthorpe H. 1949. A seventh instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)2: 401-422.
 * Donisthorpe H. 1950. An eighth instalment of the Ross Collection of ants from New Guinea. Annals and Magazine of Natural History (12)3: 338-341.
 * Eguchi K., B. T. Viet, and S. Yamane. 2014. Generic Synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part IICerapachyinae, Aenictinae, Dorylinae, Leptanillinae, Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae. Zootaxa 3860: 001-046.
 * Eguchi K., T. V. Bui, S. Yamane, H. Okido, and K. Ogata. 2004. Ant faunas of Ba Vi and Tam Dao, North Vietnam (Insecta: Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 27: 77-98.
 * Eguchi K., and S. Yamane. 2003. Species diversity of ants (Hymenoptera, Formicidae) in a lowland rainforest, northwestern Borneo. New Entomol. 52(1,2): 49-59.
 * Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine). [concl.]. Ann. Mus. Civ. Stor. Nat. 25(5): 427-473.
 * Emery C. 1888. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza . Formiche raccolte dal sig. Elio Modigliani in Sumatra e nell'isola Nias. Annali del Museo Civico di Storia Naturale 25: 528-534.
 * Emery C. 1893. Voyage de M. E. Simon à l'île de Ceylan (janvier-février 1892). Formicides. Annales de la Société Entomologique de France 62: 239-258.
 * Emery C. 1911. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118: 1-125.
 * Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia (continuazione e fine)." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 5, no. 25 (1887): 427-473.
 * Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
 * Forel A. 1900. Les Formicides de l'Empire des Indes et de Ceylan. Part VI. J. Bombay Nat. Hist. Soc. 13: 52-65.
 * Forel A. 1901. Formiciden aus dem Bismarck-Archipel, auf Grundlage des von Prof. Dr. F. Dahl gesammelten Materials. Mitt. Zool. Mus. Berl. 2: 4-37.
 * Forel A. 1903. Les fourmis des îles Andamans et Nicobares. Rapports de cette faune avec ses voisines. Rev. Suisse Zool. 11: 399-411.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * [[Media:Forel 1913l.pdf|Forel A. 1913k. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.]]
 * Forel A. 1915. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43.
 * General D. M., and G. D. Alpert. 2012. A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. Zookeys 200: 1-111.
 * Ghosh S. N., S. Sheela, B. G. Kundu, S. Roychowdhury, and R. N. Tiwari. 2006. Insecta: Hymenoptera: Formicidae. Pp. 369-398 in: Alfred, J. R. B. (ed.) 2006. Fauna of Arunachal Pradesh. (Part -2). [State Fauna Series 13.]. New Delhi: Zoological Survey of India, iv + 518 pp.
 * Gumawardene, N.R., J.D. Majer and J.P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2:71-83
 * Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
 * Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
 * Janda M., G. D. Alpert, M. L. Borowiec, E. P. Economo, P. Klimes, E. Sarnat, and S. O. Shattuck. 2011. Cheklist of ants described and recorded from New Guinea and associated islands. Available on http://www.newguineants.org/. Accessed on 24th Feb. 2011.
 * Karavaiev V. 1926. Ameisen aus dem Indo-Australischen Gebiet. Treubia 8: 413-445.
 * Karavaiev V. 1930. Ameisen von den Molukken und Neuguinea. (Ergebnisse der Sunda-Expedition der Notgemeinschaft der deutschen Wissenschaft 1929/30.). Zool. Anz. 92: 206-214.
 * Latumahina F., M. Borovanska, N. S. Putra, and M. Janda. 2015. Ants of Ambon Island  diversity survey and checklist. ZooKeys 472: 4357.
 * Mann W. M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology 63:273-391.
 * Mann William. 1916. The Ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology at Harvard College 63(7): 273-391
 * Mann, W.M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology of Harvard College 63: 273-391
 * Menozzi, C. 1932. Formiche dell'Isola di Nias. Misc. Zool. Sumatr. 65: 1-13
 * Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
 * Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
 * Ogata K. 2005. Asian ant inventory and international networks. Report on Insect inventory Project in Tropic Asia TAIIV: 145-170.
 * Pfeiffer M., D. Mezger, and J. Dyckmans. 2013. Trophic ecology of tropical leaf litter ants (Hymenoptera: Formicidae) - a stable isotope study in four types of Bornean rain forest. Myrmecological News 19: 31-41.
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
 * Santschi F. 1920. Fourmis d'Indo-Chine. Annales de la Société Entomologique de Belgique 60: 158-176.
 * Santschi F. 1924. Fourmis d'Indochine. Opuscules de l'Institut Scientifique de l'Indochine 3: 95-117
 * Santschi F. 1932. Résultats scientifiques du voyage aux Indes orientales néerlandaises de LL. AA. RR. le Prince et la Princesse Léopold de Belgique. Hymenoptera. Formicidae. Mémoires du Musée Royal d'Histoire Naturelle de Belgique. (2)4: 11-29.
 * Smith F. 1863. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Mysol, Ceram, Waigiou, Bouru and Timor. Journal and Proceedings of the Linnean Society of London. Zoology 7: 6-48.
 * Smith, Fr. "Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the Islands of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea." Journal of the Proceedings of the Linnean Society of London, Zoology 5 (1860): 93-143.
 * Snelling R. R. 1998. Insect Part 1: The social Hymenoptera. In Mack A. L. (Ed.) A Biological Assessment of the Lakekamu Basin, Papua New Guinea, RAP 9. 189 ppages
 * Snelling R. R. 2000. Ants of the Wapoga river area, Irian Jaya, Indonesia. In Mack, Andrew L. and Leeanne E. Alonso (eds.). 2000. A Biological Assessment of the Wapoga River Area of Northwestern Irian Jaya, Indonesia. RAP Bulletin of Biological Assessment 14, Conservation International, Washington, DC.
 * Stitz H. 1925. Ameisen von den Philippinen, den malayischen und ozeanischen Inseln. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1923: 110-136.
 * Taylor R. W. 1987. A checklist of the ants of Australia, New Caledonia and New Zealand (Hymenoptera: Formicidae). CSIRO (Commonwealth Scientific and Industrial Research Organization) Division of Entomology Report 41: 1-92.
 * Tiwari R. N., B. G. Kundu, S. Roy Chowdhury, and S. N. Ghosh. 2003. Insecta: Hymenoptera: Formicidae. Fauna of Sikkim. Part 4. State Fauna Series. 9.Zool.Surv.India. i-iii, 1-512. Chapter pagination: 467-506.
 * Tiwary R. N., D. K. Guha, and P. K. Maiti. 1977. New records of Ponerine ants from Arunachal Pradesh (Hymenoptera: Formicidae). Newsl Zool Surv India 3(3): 49-50
 * Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
 * Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
 * Wheeler W. M., and J. W. Chapman. 1925. The ants of the Philippine Islands. Part I, Dorylinae and Ponerinae. Philipp. J. Sci. 28: 47-73.
 * Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
 * Wheeler, William Morton.1935.Checklist of the Ants of Oceania.Occasional Papers 11(11): 3-56
 * Widihastuty, Tobing M. C., Marheni, and R. A. Kuswardani. 2018. Prey preference of Myopopone castanea (hymenoptera: formicidae) toward larvae Oryctes rhinoceros Linn (coleoptera: scarabidae). IOP Conf. Series: Earth and Environmental Science 122: 012120.
 * Wilson E. O. 1958. Studies on the ant fauna of Melanesia. I. The tribe Leptogenyini. II. The tribes Amblyoponini and Platythyreini. Bulletin of the Museum of Comparative Zoology 118: 101-153.
 * Wilson E. O. 1959. Some ecological characteristics of ants in New Guinea rain forests. Ecology 40: 437-447.
 * Wilson E.O. 1959. Adaptive shift and dispersal in a tropical ant fauna. Evolution 13(1): 122-144.
 * Wilson Edward O. 1959. Adaptive Shift and Dispersal in a Tropical Ant Fauna. Evolution 13(1): 122-144
 * Xu Z. H., and Q. J. He. 2011. Description of Myopopone castanea (Smith) (Hymenoptera: Formicidae) from Himalaya Region. Entomotaxonomia 33(3): 231-235.
 * Yamane S.; Bui T. V.; Ogata K.; Okido H.; Eguchi K. 2002. Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture Kyushu University 25: 51-62.
 * Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.