Lasius umbratus

Lasius umbratus nests in moist soil in open woodlands, early successional forests, forest edges, and in farm fields (Ellison et al., 2012). It is known to be a temporary social parasite. Queens found new colonies by infiltrating an established nest of another Lasius species (including, , , , , and ), killing the queen and using host workers to care for her initial brood.

Identification
Collingwood (1979). Clear yellow to reddish yellow; funiculus segments 2 to 4 slightly longer than wide. Scape elliptical in cross section. Petiole tapering to dorsal crest which is usually emarginate. Body surface and appendages covered in adpressed silvery pubescence. Longest hairs on gaster 0.06 mm to 0.11 mm, about half maximum hind tibial width. Erect hairs on genae, scapes and tibiae numerous. Length: 3.8-5.5 mm

Mackay and Mackay (2002) - The workers of this species are yellow or pale brown ants, which have small eyes (35 - 65 ommatidia). The length of the longest hairs on the posterior half of the first gastral tergum (excluding those along the posterior strip) are less than ½ the maximum width of the posterior tibia at mid-length. The hairs on the first gastral tergum are spaced far enough apart that the distances between the tips of most or all hairs are greater than their lengths.

Ellison et al. (2012) - Lasius umbratus is most easily confused with the similarly hairy Lasius subumbratus. However, the hairs on the gaster of L. umbratus are short and bristly – less than half as long as the hind tibia is wide – whereas the hairs on the gaster of L. subumbratus are longer than the hind tibia is wide. In addition, L. umbratus has a concave-topped petiole (viewed from the front or back), whereas L. subumbratus has a convex-topped petiole. An undescribed species of Lasius from New England is similar to L. umbratus except that the new species has erect hairs on its antennal scape and hind tibiae, whereas L. umbratus does not.

Distribution
Widespread over both Eurasia and North America. In eastern North America Lasius umbratus occurs from Nova Scotia south to the Gulf States. Westward, umbratus is abundant through North Dakota, as evidenced by the large numbers of collections made in many localities in that state by G. C. Wheeler and his students. It appears to be relatively common in the southern Rockies, but sparse to absent over most of the rest of western North America.

Distribution based on Regional Taxon Lists
Neotropical Region: Mexico. Palaearctic Region: Albania, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Japan, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Korea, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, United Kingdom of Great Britain and Northern Ireland.

Biology
Wilson (1955) - In the face of the revelation that umbratus has a common and hitherto poorly known Palaearctic sibling, Lasius rabaudi (=meridionalis Bondroit), the great mass of European literature pertaining to this species and its many synonyms cannot be accepted without major qualifications. It is in fact very probable that much of the literature deals with rabaudi instead of umbratus. Among the European authors, only Starcke (1937) seems to have fully realized the status and common occurrence of rabaudi and taken this taxonomic information into account in his ecological work. We are still very much in the dark as to whether the two species differ ecologically to any appreciable extent. In the following brief resume, reference to European literature on umbratus is made with the understanding that both species may be included.

In Europe, according to Gosswald (1932), Zimmermann (1934), Donisthorpe (1927), and others, umbratus is less common than the prominent members of Lasius s. s. It prefers dry areas and rarely nests in moist soil; Skwarra (1929) found it very rare on the wet Zehlau moors of East Prussia. It usually nests under rocks, but also occurs in rotting wood (including the timbers of houses), at the foot of trees, or in open ground. On occasion it builds mounds. Its preferred habitat is woodland, but it has also been taken along forest borders and in cultivated fields. In North America, where no sibling comparable to rabaudi is yet known, umbratus differs from the European population in that it prefers moist soil, but it still shows the same latitude in specific nesting sites. The majority of colonies have been taken under stones, while the rest have been taken in or about rotting logs and stumps. I do not know of any case of this ant building mounds or even nesting in the open soil in North America, as it (or rabaudi) has been known to do in Europe. In the northern U. S. east of the Mississippi River, umbratus is limited mostly to moist woodland, where it occurs under a wide variety of conditions of soil texture and insolation. In Alabama and Florida, at the southern extremity of the range, all of the several colonies recorded were found in rotting logs and stumps in swampland. In the western U.S., all of the collections with ecological data that I have examined were made under rocks in open forest and along or near forest borders. In New Mexico, A. C. Cole took this species between 7100 and 8000 feet, always under rocks but under variable conditions of soil moisture and vegetation, e.g. dry soil with scattered juniper and pine or oak and pine, moist soil in a clearing near a hardwood forest, and moist soil in an open grassy area.

As is the case in other species of Lasius, observations on the food habits of umbratus are entirely fragmentary and anecdotal. umbratus is generally thought to be subterranean and to subsist primarily on the excreta of aphids and coccids, since these insects are often found in great numbers in the galleries with the ants (cf. Donisthorpe, 1927, and Gregg, 1944). However, in Holland, Starcke (1937) has observed workers foraging aboveground at night and carrying insects to the nests presumably for use as food. Brown (pers. commun.) has also observed workers aboveground on cloudy days in Pennsylvania.

A number of nuptial flights recorded by Donisthorpe (1927), Crawley (1915), and Eidmann (1926) suggest a long flight season in Europe, extending from as early as August 8 (Crawley) to as late as October 7 (Eidmann). However, there is again no way of knowing whether these records might not represent the overlapping periods of the two species umbratus and rabaudi. Winged queens, determined by me as authentic umbratus, have been taken in Europe on the following dates: IV-4, V-11, VI-9, VI-12, VII-24, VII-31, VIII, VIII, VIII- (15-20), VIII-22, VIII-28, IX3, IX-10, IX-13, IX-16, IX-17, IX-29; these do not involve any apparent geographic trend and by themselves may indicate an unusually long flight period.

The situation in North America is somewhat similar. I have observed queens in flight in the environs of Boston, Mass., in September during two recent seasons. Lone dealate queens were found wandering above ground at Cambridge, Mass., on September 5, 1952, and October 2, 1953, and at Plantagenet, Ontario, on June 30, 1952. Dates on which winged forms have been taken alone or in nido cover the same period, as shown by the following randqm sample: VI-28, VII-8, VII-27, VIII-I, VIII-8, VIII-13, VIII-13, VIII-18, VIII-20, VIII-31, IX-I, IX-5, IX-9, IX-27, X-14, X-28, X-29. Since there are no known sibling species to complicate the picture in North America, the data here suggest that on this continent at least umbratus has an unusually long nuptial season.

There is no evidence to indicate that the reproductives of umbratus build aerial swarms during their nuptial flights, as do those of Lasius niger and Lasius brevicornis, although this does not preclude the possibility. Eidmann (1926) observed queens of umbratus (or rabaudi?) flying singly in Germany, and I have observed definitely determined umbratus queens flying singly on two occasions in the Boston area.

Donisthorpe (1927), Crawley (in Donisthorpe, ibid.), Gosswald (1938), and Holldobler (1953) have reported in detail on the colony founding behavior of "umbratus" and "Lasius mixtus". The normal hosts are niger and Lasius alienus. Under both field and laboratory conditions dealate umbratus queens attack host workers as they encounter them away from the nests, seizing them up the mandibles and carrying them about as they resume foraging. Their victims are usually killed by this treatment and may eventually be eaten. With the fulfillment of this Mordinstinkt, as Holldobler calls it, and the presumed acquisition of the host odor, the queens are ready and able to enter host colonies, although they may be subjected to further attack before acquiring final acceptance. Unfortunately, the authors who have witnessed this phenomenon failed to make a convincing distinction between umbratus and rabaudi, and specimens were not saved to allow corroborative determinations during the present revision.

During the falls of 1952 and 1953 I collected numerous dealate umbratus queens at Cambridge, Mass., and tried introducing them into colonies and colony fragments of Lasius pallitarsis, alienus, and neoniger, but never obtained a complete adoption and saw no evidence of the Mordinstinkt behavior. I have also worked on the theory that the umbratus may join recently fecundated host queens, since both host and parasite queens are often found in species-pure groups under rocks following nuptial flights. Various attempts to bring umbratus and neoniqer queens together, including placing them in the same chamber while chilled, have so far failed; the reason may be, however, that neoniger is not a natural host. I would like to suggest, on what admittedly constitutes negative evidence, that behavior in the population I studied may differ from that in the European populations. If true, this could be due either to geographic variation or to the fact that the European authors were using rabaudi instead of umbratus. Only additional research accompanied by careful determinations will settle the matter.

niger and alienus probably serve as hosts of umbratus in North America as they do in Europe. I have seen two mixed niger-umbratus nest series from Ute Park, New Mexico (A. C. Cole leg. and Coll., MCZ) and one alienus-umbratus series from Beatty, Pennsylvania (Schmitt leg.; MCZ). The Beatty umbratus are minimas. Buren (1944) found a single dealate “aphidicola" queen with a depauperate colony of "flavus nearcticus”, but to my knowledge there has been no additional evidence forthcoming that this or any other Chthonolasius uses species of Cautolasius as hosts.

Collingwood (1979) reporting on umbratus in Europe - This species nests under boulders, in tree stumps and at the base of old trees. Workers are subterranean and seldom or never seen above ground. Flight period from mid August to late September. Single queens found colonies by invasion of and adoption in Lasius niger, Lasius alienus or occasionally Lasius brunneus nests. In late summer dealate queens often wander over the surface of L. niger nests, sometimes carrying a dead L. niger worker as a prelude to securing adoption.

In New Mexico (Mackay and Mackay 2002) - Moist areas, ranging from pastures to mixed forests and pine forests. This species nests under stones and logs, tends Homoptera, and is a temporary social parasite of Lasius alienus, Lasius niger and Lasius neoniger. Reproductives were found in nests July and September, dealate females were found in September. It may form a plesiobiotic relationship with Temnothorax rugatulus.

Nomenclature

 *  umbratus. Formica umbrata Nylander, 1846b: 1048 (q.m.) FINLAND. Schenck, 1852: 59 (w.); Hauschteck, 1962: 219 (k.); Hung, 1969: 456 (k.). Combination in Lasius: Mayr, 1861: 50; in Donisthorpea: Donisthorpe, 1915d: 223; in Formicina: Emery, 1916b: 241; in Acanthomyops: Ruzsky, 1925b: 44; in Chthonolasius: Ruzsky, 1936: 91; in Lasius (Chthonolasius): Ruzsky, 1914a: 59; Müller, 1923: 129; Emery, 1925b: 233; Wilson, 1955a: 150. Senior synonym of belgarum: Wilson, 1955a: 150; Seifert, 1988a: 150; of hirtiscapus: Wilson, 1955a: 150; Seifert, 1990: 11; of affinoumbratus, aphidicola, epinotalis, exacutus, nyaradi, osakana, silvestrii and material of the unavailable names mixtoaffinis, mixtobicornis referred here: Wilson, 1955a: 150; of mixtoumbratus: Wilson, 1955a: 150; Kutter, 1977c: 14. See also: Stitz, 1939: 291; Bernard, 1967: 363; Kutter, 1977c: 232; Yamauchi, 1979: 168; Seifert, 1988a: 150; Atanassov & Dlussky, 1992: 245; Radchenko, 2007: 34.
 * exacutus. Lasius umbratus var. exacutus Ruzsky, 1902d: 15 (w.) RUSSIA. Combination in Acanthomyops: Ruzsky, 1925b: 44; in Lasius (Chthonolasius): Ruzsky, 1914a: 61; Emery, 1925b: 234. Subspecies of umbratus: Ruzsky, 1905b: 287. Raised to species: Ruzsky, 1925b: 44. Subspecies of bicornis: Menozzi, 1939a: 314 (in key). Junior synonym of umbratus: Wilson, 1955a: 150.
 * affinoumbratus. Lasius umbratus var. affinoumbratus Donisthorpe, 1914: 40 (w.) GREAT BRITAIN. Junior synonym of umbratus: Wilson, 1955a: 151.
 * belgarum. Formicina belgarum Bondroit, 1918: 31, fig. 17 (w.q.) BELGIUM. Combination in Lasius (Chthonolasius): Emery, 1925b: 234. Subspecies of umbratus: Stärcke, 1937: 57; of mixtus: Stitz, 1939: 299. Junior synonym of umbratus: Wilson, 1955a: 151; Seifert, 1988: 150.
 * silvestrii. Lasius (Chthonolasius) silvestrii Wheeler, W.M. 1928d: 120 (q.) JAPAN. Junior synonym of umbratus: Wilson, 1955a: 152.
 * hirtiscapus. Lasius umbratus var. hirtiscapus Stärcke, 1937: 43 (q.) CZECHOSLOVAKIA. Subspecies of mixtus: Stitz, 1939: 299. Junior synonym of umbratus: Wilson, 1955a: 152.
 * osakana. Lasius (Chthonolasius) silvestrii var. osakana Santschi, 1941: 278 (q.) JAPAN. Junior synonym of umbratus: Wilson, 1955a: 152.
 * nyaradi. Chthonolasius affinis var. nyaradi Röszler, 1943: 47 (w.q.) ROMANIA. Junior synonym of umbratus: Wilson, 1955a: 152.

Type Material
Wilson (1955) - LECTOTYPE. A dealate queen in the Helsinki Museum, selection by Starcke (1937).

Worker
Wilson (1955) - (1) Pilosity and pubescence as in queen. Maximum length of hairs of first gastric tergite anterior to the extreme posterior strip 0.06-0.08 mm., not exceeding one-half the maximum width of the hind tibia at midlength. Alitruncal and cephalic hairs with maximum length of about 0.11 mm.

(2) Petiole in frontal view tapering slightly from the widest point, just above the foramina, to the dorsal crest. The dorsal crest broad and very variable in outline, from flat or even feebly convex to deeply concave; the emargination rounded or angular, never as deep as in Lasius bicornis, i.e. the width (taken at the midpoint of the depth measurement) always exceeds the depth. Intranidal variation considerable; the petiole in a single series may range from flat to distinctly emarginate. In side view the scale is relatively narrow, its dorsal crest acute.

Dong (2017) - (Korean individuals) Body length 4~4.5mm, bright yellow ground color, and body largely hairy. Compound eyes relatively well-developed. Antennal scape protruded head occipital margin in full-face view. Antennal scape with appressed simple hairs densely. Anterior clypeal margin convex, not concaved. Thorax dorsally hairy. Propodeal slope straight and steep. Metapleural gland and propodeal spiracle relatively wide but partially obscured by a couple of setae. In profile view, petiole reversed-V shaped and midpoint of petiole slightly concave or straight when seen in frontal view. Dorsum of petiole with 7~10 filiform fine hairs.

Queen
Wilson (1955) - (1) Most of the body surface covered with abundant, relatively short, silvery yellow, predominantly erect hairs. The hairs on the first three gastric tergites with maximum length variable internidally, 0.05-0.11 mm., never more than one-half the maximum width of the hind tibiae at midlength, and often less than one-third; very variable in density, never less than 20 hairs visible above the dorsal profile of the first gastric segment seen in perfect side view and usually more than 30, but never dense enough to overlap one another extensively, Erect hairs forming a fringe on the dorsal crest of the petiole, their maximum length close to that of the gastric hairs. The longest hairs of the alitrunk are on the scutellum, maximum length 0.12-0.21 mm. Maximum length of scutal hairs 0.05-0.15 mm. (see also under geographic variation). Maximum length of cephalic hairs exclusive of those on the clypeus 0.09.-0.11 mm. Body hairs mostly straight or feebly curved, rarely strongly curved (on propodeum) and never sinuate. Standing hairs mayor may not be present on the appendages (see also under geographic variation). All of body and appendages densely covered with short, whitish pubescence which is completely appressed on the body and appressed to decumbent on the appendages; on the gaster it is often abundant enough to obscure partly the shining cuticular surface and to present a whitish overcast to the naked eye.

(2) HW ranging 1.40-1.82 mm. in 45 nest series measured (see also under geographic variation). SI of size extremes 75 and 81.

(3) Petiole in frontal view tapering gradually but distinctly from the level of maximum width (just above the foramina) to the dorsal crest, the width just ventral to the dorsolateral angles 0.9 X the maximum width or less but the frontal outline rarely subquadrate as in rabaudi. Dorsal crest extremely variable in shape, from very feebly concave or even straight to deeply concave with the emargination almost right-angular. The dorsolateral angles always broadly rounded. In side view the scale is narrow and with an acute dorsal crest.

(4) The scape short-elliptical to circular in cross-section, never conspicuously flattened, the minimum width at the midpoint 0.11 mm. or greater. The third funicular segment 1.0-1.5 X longer than broad.

(5) Body medium to dark brown, the appendages lighter, light to medium brown.

Male
Wilson (1955) - (1) Pilosity and pubescence essentially the same as in the queen and worker, except that hairs of the first three gastric tergites are more frequently subdecumbent-suberect. Despite this greater tendency toward obliqueness, the hairs of the first gastric tergite are still too sparse to show much overlap, and their maximum length (excluding those on the extreme posterior strips) ranges internidally 0.07-0.08 mm., or always less than 0.7 X the maximum width of the hind tibia at its midlength. Maximum length of scutellar and cephalic hairs (excluding those on the clypeus) 0.09 mm.

(2) HW 0.85-1.23 mm. in 15 nest series measured; SI of the extremes 62 and 66.

(3) Petiole in frontal view tapering dorsally as in queen and worker. Dorsal margin flat to deeply emarginate, the emargination rounded or angular, never greater than semicircular or rightangular. The scale in side view relatively thin, with an acute dorsal crest.

References based on Global Ant Biodiversity Informatics

 * Agosti, D. and C.A. Collingwood. 1987. A provisional list of the Balkan ants (Hym. Formicidae) and a key to the worker caste. I. Synonymic list. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 60: 51-62
 * Aldawood AS, Sharaf MR (2011) Monomorium dryhimi sp. n., a new ant species (Hymenoptera, Formicidae) of the M. monomorium group from Saudi Arabia, with a key to the Arabian Monomorium monomorium-group. ZooKeys 106: 4754. doi: 10.3897/zookeys.106.139
 * AntArea. Accessed on February 5th 2014 at http://antarea.fr/fourmi/
 * Antarea (Personal Communication - Rumsais Blatrix- 27 April 2018)
 * Antarea (at www.antarea.fr on June 11th 2017)
 * ArtDatabanken Bugs (via GBIG)
 * Asociacion Iberica de Mirmecologia. 2014. List of species collected during the Taxomara 2014 Oviedo. Iberomyrmex 6: 23-24.
 * Assing V. 1989. Die Ameisenfauna (Hym.: Formicidae) nordwestdeutscher Calluna-Heiden. Drosera 89: 49-62.
 * Azuma M. 1955. A list of ants (Formicidae) from Hokkaido Is. Hyogo Biology 3:79-80.
 * Banert P, and B. Pisarski. 1972. Mrówki (Formicidae) Sudetów. Fragmenta Faunistica (Warsaw) 18: 345-359.
 * Baroni Urbani C., and C. A. Collingwood. 1976. A Numerical Analysis of the Distribution of British Formicidae (Hymenoptera, Aculeata). Verhandlungen der Naturforschenden Gesellschaft in Basel 85: 51-91.
 * Baroni Urbani C., and C. A. Collingwood. 1977. The zoogeography of ants (Hymenoptera, Formicidae) in Northern Europe. Acta Zoologica Fennica 152: 1-34.
 * Baroni Urbani, C.. "Catalogo delle specie di Formicidae d'Italia (Studi sulla mirmecofauna d'Italia X)." Memorie della Società Entomologica Italiana Volume 50 (1971): 5-287.
 * Barrett K. E. 1967. Ants in South Brittany. Entomologist's Record and Journal of Variation 79:112-116.
 * Barrett K. E. J. 1968b. The distribution of ants in central southern England. Transactions of the Society for British Entomology 17: 235-250.
 * Barrett K. E. J. 1970. Ants in France, 1968-69. Entomologist 103: 270-274.
 * Baugnee J. Y. 2003. Camponotus piceus (Leach, 1825), fourmi nouvelle pour la faune belge decouverte dans le parc naturel Viroin-Hermeton (Hymenoptera: Formicidae). Bulletin S. R. B. E./K. B. V. E. 139: 219-225.
 * Behr D., S. Lippke, and K. Colln. 1996. Zur kenntnis der ameisen von Koln (Hymenoptera, Formicidae). Decheniana-Beihefte (Bonn) 35: 215-232.
 * Behr D., and K. Colln. 1993. Zur ameisenfauna (Hymenoptera, Formicidae) von Gonnersdorf (Kr. Daun). Dendrocopos 20: 148-160.
 * Bernard F. 1959. Les fourmis de l'île de Port-Cros. Contribution à l'écologie des anciennes forêts méditerranéennes. Vie Milieu 9: 340-360.
 * Bernard F. 1967. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.
 * Bernard F. 1976. Écologie des fourmis des grès d'Annot, comparées à celles de la Provence calcaire. Annales du Muséum d'Histoire Naturelle de Nice 3: 33-54.
 * Berville L., M. Renucci, and E. Provost. 2012. Mise en place de protocoles de contrôle de la fourmi dArgentine (Linepithema humile) sur les îles de Port-Cros et de Porquerolles (Var, France). Sci. Rep. Port-Cros natl. Park, 26: 91-108.
 * Bezdecka P. 1996. The ants of Slovakia (Hymenoptera: Formicidae). Entomofauna carpathica 8: 108-114.
 * Bigot L. 1959. Complement a l'inventaire de la faune entomologique de la Camargue 93eme note). La Terre et la Vie 106 : 149-157.
 * Blacker N. C. and C. A. Collingwood. 2002. Some significant new records of ants (Hymenoptera: Formicidae) from the Salisbury area, south Wiltshire, England, with a key to the British species of Lasius. British Journal of Entomology and Natural History 15: 25-46
 * Blatrix R., C. Lebas, C. Galkowski, P. Wegnez, P. Pimenta, and D. Morichon. 2016. Vegetation cover and elevation drive diversity and composition of ant communities (Hymenoptera: Formicidae) in a Mediterranean ecosystem. – Myrmecological News 22: 119-127.
 * Boer P. 2019. Species list of the Netherlands. Accessed on January 22 2019 at http://www.nlmieren.nl/websitepages/specieslist.html
 * Boer P., W. Dekoninck, A. J. Van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten (Amsterdam) 63: 54-58.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. Lijst van mieren (Hymenoptera: Formicidae) van Belgie en Nederland, hun Nederlandse namen en hun voorkomen. Entomologische Berichten 63(3): 54-57.
 * Boer P., W. Dekoninck, A. J. van Loon, and F. Vankerkhoven. 2003. List of ants (Hymenoptera: Formicidae) of Belgium and The Netherlands, their status and Dutch vernacular names. Entomologische Berichten 63 (3): 54-58.
 * Boer P., and J. Noordijk. 2004. De ruige gaststeekmier Myrmica hirsuta nieuw voor Nederland (Hymenoptera: Formicidae). Ned. Faun. Meded. 20: 25-32.
 * Bonaric J. C. 1971. Contribution a l'etude systematique et ecologique des formicides du Bas-Languedoc. PhD thesis Universite des sciences et techniques du Languedoc, 175 pages.
 * Bonaric J. C. 1971. Étude systématique et ecologique des fourmis de lHérault (fin). Ann. Soc. Hortic. Hist. Nat. Hérault 111: 169-176.
 * Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
 * Borowiec L., and S. Salata. 2012. Ants of Greece - Checklist, comments and new faunistic data (Hymenoptera: Formicidae). Genus 23(4): 461-563.
 * Bourne R. A. 1973. A taxonomic study of the ant genus Lasius Fabricius in the British Isles (Hymenoptera: Formicidae). J. Entomol. Ser. B 42: 17-27.
 * Boven J. K. A. 1947. Liste de détermination des principales espèces de fourmis belges (Hymenoptera Formicidae). Bulletin et Annales de la Société Entomologique de Belgique 83: 163-190.
 * Bracko G. 2007. Checklist of the ants of Slovenia (Hymenoptera: Formicidae). Natura Sloveniae 9: 15-24
 * Bracko G., H. C. Wagner, A. Schulz, E. Gioahim, J. Maticic, and A. Tratnik. 2014. New investigation and a revised checklist of the ants (Hymenoptera: Formicidae) of the Republic of Macedonia. North-Western Journal of Zoology 10(1): 10-24.
 * Bracko, G. 2006. Review of the ant fauna (Hymenoptera:Formicidae) of Croatia. Acta Entomologica Slovenica 14(2): 131-156.
 * Bracko, G.. "Review of the ant fauna (Hymenoptera: Formicidae) of Croatia." Acta Entomologica Slovenica Vol 14 st (2006): 131-156.
 * Bujan J., A. Brigic, Z. Sedlar, and R. Sostaric. 2015. Progressive vegetation succession of fen habitats promotes the lack of habitat specialist ants. Insectes Sociaux 62: 415-422.
 * Carniel A. 1998. Ricerche sulla mirmecofauna delle Prealpi Orobiche (Lombardia) (Insecta, Hymenoptera, Formicidae). Atti. Mus. Civ. Stor. Nat. Morbegno 9: 29-39.
 * Casevitz-Weulersse J. 1990. Etude Systematique de la Myrmecofaune Corse (Hymenoptera: Formicidae), deuxieme partie. Bull. Mus. Natn. Hist. Nat. Paris. 4eme serie 12, section A(2): 415-442.
 * Casevitz-Weulersse J., and C. Galkowski. 2009. Liste actualisee des Fourmis de France (Hymenoptera, Formicidae). Bull. Soc. Entomol. Fr. 114: 475-510.
 * Casevitz-Weulersse J., and M. Prost. 1991. Fourmis de la Côte-d'Or présentes dans les collections du Muséum d'Histoire Naturelle de Dijon. Bulletin Scientifique de Bourgogne 44: 53-72.
 * Casevitz-Weulersse, J.. "Contribution a la connaisance des fourmis de la Corse (Hymenoptera: Formicidae)." These de Doctorat Museum Nat (1989): 379pp.
 * Choi B.M., K. Ogata, and M. Terayama. 1993. Comparative studies of ant faunas of Korea and Japan. 1. Faunal comparison  among islands of Southern Korean and northern Kyushu, Japan. Bull. Biogeogr. Soc. Japan 48(1): 37-49.
 * Colindre L. 2015. Les fourmis en Picardie: bilan 2014 (Hymenoptera/ Formicidae). Entomologiste Picard 26, 15 pages.
 * Colindre L. 2017. Richess et utilite du cortege de fourmis en foret d'Ermenonville, Oise, Region Hauts-de-France. Association des Entomologistes de Picardie. 19 pages.
 * Collingwood C. A. 1955. Ants in S.W. Scotland. Entomologist's Record and Journal of Variation 67: 11-12.
 * Collingwood C. A. 1956. Ant hunting in France. Entomologist 89: 106-108.
 * Collingwood C. A. 1961. Ants in the Scottish Highlands. Scotish Naturalist 70: 12-21.
 * Collingwood C. A. 1971. A synopsis of the Formicidae of north Europe. Entomologist 104: 150-176
 * Collingwood C.A. 1955. Ants in S.W. Scotland. Entomol.Rec. 67: 11-12
 * Collingwood C.A. 1957. The Species of Ants of the Genus Lasius in Britain. Journal of the Society for British Entomology. 5: 204-214
 * Collingwood C.A. 1959. Ants in the Scottish Highlands. The Scottish Naturalist. 70: 12-21
 * Collingwood C.A. 1961. Ants in Finland. Entomol. Rec. 73: 190-195
 * Collingwood C.A. 1961. New Vice-County Records for British Ants. Entomologist. 73: 90-93
 * Collingwood C.A. and Satchell J.E. 1956. The Ants of the South Lake District. Journal of the Society for British Entomology. 5: 159-164
 * Collingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Trans. Soc. Br. Entomol. 13: 69-96
 * Collingwood, C. A. 1964. The Identification of British Ants (Hym. Formicidae). Transactions of the Society for British Entomology. 16:93-121.
 * Collingwood, C. A. 1974. A revised list of Norwegian ants (Hymenoptera: Formicidae). Norsk Entomologisk Tidsskrift 21: 31-35.
 * Collingwood, C. A., and I. H. H. Yarrow. "A survey of Iberian Formicidae." EOS (Revista española de entomología) 44 (1969): 53-101.
 * Collingwood, C. A.. "The Formicidae (Hymenoptera) of Fennoscandia and Denmark." Fauna Entomologica Scandinavica 8 (1979): 1-174.
 * Collingwood, C.A. 1958. A survey of Irish Formicidae. Proceedings of the Royal Irish Academy 59B:213-219
 * Consani M., and P. Zangheri. 1952. Fauna di Romagna. Imenotteri - Formicidi. Memorie della Societa Entomologica Italiana 31: 38-48.
 * Csosz S., B. Marko, K. Kiss, A. Tartally, and L. Galle. 2002. The ant fauna of the Ferto-Hansag National Park (Hymenoptera: Formicidae). In: Mahunka, S. (Ed.): The fauna of the Fert?-Hanság National Park. Hungarian Natural History Museum, Budapest, pp. 617-629.
 * Csősz S., B. Markó, and L. Gallé. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7: 55-62.
 * Czechowski W., A. Radchenko, W. Czechowska and K. Vepsäläinen. 2012. The ants of Poland with reference to the myrmecofauna of Europe. Fauna Poloniae 4. Warsaw: Natura Optima Dux Foundation, 1-496 pp
 * Dekoninck W., H. De Koninck, J. Y. Baugnee, and J. P. Maelfait. 2007. Ant biodiversity conservation in Belgian calcareous grasslands: active management is vital. Belg. J. Zool. 137 (2): 137-146.
 * Dekoninck W., P. Boer, and J. P. Maelfait. 2004. Lasius platythorax Seifert, 1991 as a host of several Chthonolasius species, with remarks on the colony foundation of the parasites (Hymenoptera: Formicidae). Myrmecologische Nachrichten 6: 5-8.
 * Della Santa E. 1994. Guide pour l'identification des principales espèces de fourmis de Suisse. Miscellanea Faunistica Helvetiae 3: 1-124.
 * Della Santa E. 1995. Fourmis de Provence. Faune Provence 16: 5-37.
 * Dewes E. 2005. Ameisenerfassung im Waldschutzgebiet Steinbachtal/Netzbachtal. Abh. Delattinia 31: 89-118.
 * Donisthorpe H. 1914. Myrmecophilous notes for 1913. Entomologist's Record and Journal of Variation 26: 37-45.
 * Downing H., and J. Clark. 2018. Ant biodiversity in the Northern Black Hills, South Dakota (Hymenoptera, Formicidae). Journal of the Kansas Entomological Society 91(2): 119-132.
 * Dubovikoff D. A., and Z. M. Yusupov. 2018. Family Formicidae - Ants. In Belokobylskij S. A. and A. S. Lelej: Annotated catalogue of the Hymenoptera of Russia. Proceedingss of the Zoological Institute of the Russian Academy of Sciences 6: 197-210.
 * Else G., B. Bolton, and G. Broad. 2016. Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea). Biodiversity Data Journal 4: e8050. doi: 10.3897/BDJ.4.e8050
 * Emery C. 1869. Enumerazione dei formicidi che rinvengonsi nei contorni di Napoli con descrizioni di specie nuove o meno conosciute. Ann. Accad. Aspir. Nat. Secunda Era 2: 1-26.
 * Emery C. 1878. Liste des fourmis de la collection de feu Camille van Volxem, avec la description d'une espèce nouvelle. Annales de la Société Entomologique de Belgique 21: viii-x.
 * Emery C. 1916. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47: 79-275.
 * Emery, C.. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte seconda. Formiche dell'Europa e delle regioni limitrofe in Africa e in Asia." Annali del Museo Civico di Storia Naturale 12 (1878): 43-59.
 * Emery, C.. "Liste des fourmis de la collection de feu Camille van Volxem, avec la description d'une espèce nouvelle." Ann. Soc. Entomol. Belg. 21 (1878): viii-x.
 * Entomological Society of Latvia. 2003. http://leb.daba.lv/Formicidae.htm (Accessed on December 1st 2013).
 * Espadaler, X.. "Formigues del Montseny." in: Terradas, J., Miralles, J. (eds.) El patrimoni biològic del Montseny Catàlegs d (1987): 101-103.
 * Field Museum Collection, Chicago, Illinois (C. Moreau)
 * Forel A. 1892. Die Ameisenfauna Bulgariens. (Nebst biologischen Beobachtungen.). 305-318.
 * Forel A. 1900. Fourmis du Japon. Nids en toile. Strongylognathus Huberi et voisins. Fourmilière triple. Cyphomymrex Wheeleri. Fourmis importées. Mitt. Schweiz. Entomol. Ges. 10: 267-287.
 * Forel, A.. "Ameisen aus den Sporaden, den Cykladen und Griechenland, gesammelt 1887 von Herrn von Oertzen." Berliner Entomologische Zeitschrift 32 (1889): 255-265.
 * Formidabel Database
 * Fowles, A.P. 1996. A provisional checklist of the invertebrates recorded from Wales. 2. Aculeate wasps, bees and ants (Hymenoptera: Aculeata). Countryside Council for Wales
 * Fukumoto S. 2009. Ants at the Korimoto Campus of Kagoshima University. Nature of Kagoshima 35: 67. (+Personal communication 19 May 20014)
 * GRETIA. 2017. Bilan annuel de l'enquete sur la repartition des fourmis armoricaines. 23 pages.
 * Gadeau de Kerville H. 1922. Materiaux pour la Faune des Hymenopteres de la Normandie. Bull. Soc. Amis Sc. Nat. Rouen 1916-1921, 1922: 217-225.
 * Galkowski C. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galkowski C., and C. Foin. 2013. Nouvelles données sur la répartition de Strongylognathus huberi Forel, 1874 (Hymenoptera, Formicidae) en France. Bulletin de la Société Linnéenne de Bordeaux (n.s.) 41: 167-174.
 * Galle L. 1972. Study of ant-populations in various grassland ecosystems. Acta Biologica Szeged 18(1-4): 159-164.
 * Galle L. 1973. Formicidae populations of the ecosystems in the environs of Tiszafured. Tiscia, Szeged 7: 59-68.
 * Galle L. 1981. The Formicoid fauna of the Hortobagy. Pp. 307-311 in: Mahunka, S. (ed.) 1981. The fauna of the Hortobágy National Park. Budapest: Akadémiai Kiadó, 415 pp.
 * Gallé L. 1991. Structure and succession of ant assemblages in a north European sand dune area. Holarctic Ecology 14: 31-37.
 * Gallé L., B. Markó, K. Kiss, E. Kovács, H. Dürgő, K. Kőváry, and S. Csősz. 2005. Ant fauna of Tisza river basin (Hymenoptera: Formicidae).  In: Gallé, L. (szerk.): Vegetation and Fauna of Tisza River Basin I. Tiscia Monograph Series 7; Szeged, pp. 149-197.
 * Garcia Garcia F., and A. D. Cuesta-Esgura. 2017. First catalogue of the ants of Burgos province, Spain (Hymenoptera: Formicidae). Boletín de la Sociedad Entomológica Aragonesa 60: 245–258.
 * Gaspare Charles. 1965. Étude myrmécologique d'une région naturelle de Belgique: la Famenne Survey des Fourmis de la Région (Hymenoptera, Formicidae). Institut agronomique de l'Etat a' Gembloux. 32(4): 427-434.
 * Giacalone I., and M. Moretti. 2001. Contributo alla conoscenza della mirmecofauna (Hymenoptera: Formicidae) dei castagneti al Sud delle Alpi (ticino, Svizzera). Bollettino della Societa ticinese di Scienze naturali 89(1-2): 51-60.
 * Gilev A. V., I. V. Kuzmin, V. A. Stolbov, and S. D. Sheikin. 2012. Materials on the fauna and ecology of ants (formicidae) Southern part of the Tyumen region. Tyumen State University Herald 6: 86-91.
 * Glaser F. 2009. Die Ameisen des Fürstentums Liechtenstein. (Hymenoptera, Formicidae). Amtlicher Lehrmittelverlag, Vaduz, 2009 (Naturkundliche Forschung im Fürstentum Liechtenstein; Bd. 26).
 * Glaser F., T. Kopf, and K. H. Steiberger. 2003. Ameisen (Hymenoptera: Formicidae) im Frastanzer Ried und den Illauen (Vorarlberg, Österreich) Artenspektrum, Gefährdung und Schutzempfehlungen. Vorarlberger Naturschau 13: 287-310.
 * Gouraud C. 2015. Bilan de l’année 2014 : Atlas des fourmis de Loire-Atlantique (Hymenoptera, Formicidae). Atlas des Formicidae de Loire-Atlantique, compte rendu de la première année d’étude (2014)
 * Grandi G. 1935. Contributi alla conoscenza degli Imenotteri Aculeati. XV. Boll. R. Ist. Entomol. Univ. Studi Bologna 8: 27-121.
 * Gratiashvili N., Barjadze S. 2008. Checklist of the ants (Formicidae Latreille, 1809) of Georgia. Proceedings of the Institute of Zoology (Tbilisi) 23: 130-146.
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Harada Y., K. Nishikubo, K. Matsumoto, M. Matsuda, Y. Inazawa, Y. Ozono, S. Koto, N. Kawaguchi, and S. Yamane. 2011. Ant fauna of Japanese beech (Fagus crenata) forests in southwestern Japan. Bull. Biogeogr. Soc. Japan 66: 115-127.
 * Hayashida K. 1957. Ecological distribution of ants in Sapporo and vicinity. (Preliminary report.). Journal of the Faculty of Science, Hokkaido University. Series VI. Zoology 13:173-177.
 * Hayashida K. 1961. Studies on the ecological distribution of ants in Sapporo and its vicinity (1 et 2). Insectes Sociaux 7: 125-162.
 * Hayashida K. 1972. Ecological survey on ants in Nakagawa Experiment Forest of Hokkaido University. Res. Bull. Exper. Forests, Coll. Agr., Hokkaido Univ. 29: 25-36.
 * Hayashida K., and S. Maeda. 1960. Studies on the ecological distribution of ants in Akkeshi. Journ. Sc. Hokkaido Univ., IV. Zool., 14 (3) : 305-319.
 * Hisamatsu M. 2004. List of Hymenoptera Recorded in Ibaraki Prefecture. Bulletin of Ibaraki Nature Museum 7: 125-164.
 * Holgersen H. 1944. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84: 165-203.
 * Ichikawa A. 1999. Records of ants observed from several localities of Osaka Prefecture, Japan, -1. Ari 23: 1-3.
 * Ichinose K. 1990. The Ant Fauna of the Tomakomai Experiment Forest, Hokkaido University (Hymenoptera: Formicidae) with Notes on the Nuptial Season. Research Bulletins of College Experiment Forests 47(1) 137-144.
 * Jakubzik A., H. Kinkler, and K. Colln. 2010. Aculeate Hymenoptera from a Humid Biotope in Leverkusen-Steinbüchel. Decheniana (Bonn) 163: 145158.
 * Jeffery H. G. 1931. The Formicidae (or ants) of the Isle of Wight. Proceedings of the Isle of Wight Natural History and Archaeological Society 2: 125-128.
 * Karaman M. G. 2009. An introduction to the ant fauna of Macedonia (Balkan Peninsula), a check list (Hymenoptera, Formicidae). Natura Montenegrina 8(3): 151-162.
 * Karaman M. G. 2011. A catalogue of the ants (Hymenoptera, Formicidae) of Montenegro. Podgorica: Catalogues 3, Volume 2, Montenegrin Academy of Sciences and Arts, 140 pp.
 * Karavaiev V. 1912. Ameisen aus dem paläarktischen Faunengebiete. Rus. Entomol. Obozr. 12: 581-596.
 * Kim B.J. 1986. A systematic study of ants in Island Ullungdo of Korea on the basis of external fine features. The journal of Natural Science 5(2): 84-94.
 * Kim B.J. 1996. Synonymic list and distribution of Formicidae (Hymenoptera) in Korea. Entomological Research Bulletin Supplement 169-196.
 * Kim G., and D. Lyu. 2012. Distribution of ants (Insecta, Hymenoptera) in Chiaksan Mountain, Prov Gangweon, Korea. Journal of Korean Nature 5(2): 127-129.
 * Kiran K., and N. Aktac. 2006. The vertical distribution of the ant fauna (Hymenoptera: Formicidae) of the Samanh Mountains, Turkey. Linzer Biol. Beitr. 38(2): 1105-1122.
 * Kofler A. 1995. Nachtrag zur Ameisenfauna Osttirols (Tirol, Österreich) (Hymenoptera: Formicidae). Myrmecologische Nachrichten 1: 14-25.
 * Kupianskaya A. N., Lelej, A.S., and Urbain, B. K. 2000. The Ants (Hymenoptera: Formicidae) of the Kuril Islands. Far Eastern Entomologist. 92:1-21.
 * Kupianskaya A. N., and A. S. Lelej. 2000. The ants (Hymenoptera, Formicidae) collected in the Habomai and Shikotan (Kuril Islands) in 1998.  Far Eastern entomologist 92: 22-24.
 * Kupianskaya, A. N., Lelej, A.S., and Urbain, B. K. 2000. The Ants (Hymenoptera: Formicidae) of the Kuril Islands. Far Eastern Entomologist. 92:1-21.
 * Kvamme T. 1982. Atlas of the Formicidae of Norway (Hymenoptera: Aculeata). Insecta Norvegiae 2: 1-56.
 * Kvamme T., and A. Wetas. 2010. Revidert liste over norske maur  Inkludert dialektiske navn og forslag til nye norske navn og forslag til norske navn. Norsk institutt for skog og landskap, Ås. 127 pp
 * Lameere A. 1892. Note sur les fourmis de la Belgique. Annales dr la Société Entomologique de Belgique 36: 61-69.
 * Lapeva-Gjonova, L., V. Antonova, A. G. Radchenko, and M. Atanasova. "Catalogue of the ants (Hymenoptera, Formicidae) of Bulgaria." ZooKeys 62 (2010): 1-124.
 * Le Moli F., and A. Zaccone. 1995. Ricerche sulla mirmecofauna del Cansiglio (Prealpi Carniche). Soc. Ven. Sc. Nat. 20: 33-52.
 * Lebas C., C. Galkowski, P. Wegnez, X. Espadaler, and R. Blatrix. 2015. The exceptional diversity of ants on mount Coronat (Pyrénées-Orientales), and Temnothorax gredosi(Hymenoptera, Formicidae) new to France. R.A.R.E., T. XXIV (1): 24  33
 * Legakis Collection Database
 * Lelej A. S. 2012. Annotated catalogue of the Insects of Russian Far East. Volume 1. Hymenoptera. Dalnauka: Vladivostok. 635 p.
 * Lillig M., and E. Dewes. 2015. The former Siegfried Line as habitats for ants (Hymenoptera: Formicidae). Abh. Delattinia 37: 117 - 136
 * Lomnicki J. 1928. Spis mrówek Lwowa i okolicy. Ksiegi Pamiatkowej (Lecia Gimn. IV Jana Dlugosza Lwowie) 50: 1-10.
 * Lorinczi G. 2011. Density and spatial pattern of nests in sub-Mediterranean ground-dwelling ant communities (Hymenoptera: Formicidae). Community Ecology 12(1): 51-57.
 * Maavara V. 1953. Ants of Estonian SSR. ABIKS loodusevaatlejale 10: 1-44.
 * Maeto K. and S. Sato. 2004. Impacts of forestry on ant species richness and composition in warm-temperate forests of Japan. Forest Ecology and Management 187: 213223.
 * Majzlan O., and P. Devan. 2009. Selected insect groups (Hymenoptera, Neuroptera, Mecoptera, Raphidioptera) of the Rokoš Massif (Strážovské vrchy Mts.). Rosalia (Nitra), 20, p. 63–70.
 * Malozemova L. A. 1972. Ants of steppe forests, their distribution by habitats, and perspectives of their utilization for protection of forests (north Kazakhstan). [In Russian.]. Zoologicheskii Zhurnal 51: 57-68.
 * Manabe K. 1994. Ants of the shrine forest in Fukuoka Prefecture (second report; ants of mountain zone). Ari 17: 8.
 * Markó B., B. Sipos, S. Csősz, K. Kiss, I. Boros, and L. Gallé. 2006. A comprehensive list of the ants of Romania (Hymenoptera: Formicidae). Myrmecologische Nachrichten 9: 65-76.
 * Maruyama M., F. M. Steiner, C. Stauffer, T. Akino, R. H. Crozier, and B. C. Schlick-Steiner. 2008. A DNA and morphology based phylogenetic framework of the ant genus Lasius with hypotheses for the evolution of social parasitism and fungiculture. BMC Evolutionary Biology 8:Article 237 (doi:10.1186/1471-2148-8-237).
 * Minato M., T. Kameyama, F. Ito, and T. Itino. 1996. A preliminary report of ant fauna in Gagawa Prefecture. Ari 20: 9-13.
 * Mizutani A. 1979. A myrmecofaunal survey at Hiyama Experiment Forest, Hokkaido University. Research Bulletin of the College Experiment Forests, Hokkaido University 36:509-516.
 * Mizutani A. 1979. A myrmecofaunal survey at Hiyama Experiment Forest, Hokkaido University. Research Bulletins of the College Experiment Forests Hokkaido University 36(2): 509-516.
 * Moretti M. P. Duelli, and M. K. Obrist. 2006. Biodiversity and resilience of arthropod communities after fire disturbance in temperate forests. Oecologia 149: 312327.
 * Morisita M. 1945. Ants of the southern part of Hokkaido, Japan. [In Japanese.] Mushi 16:21-28.
 * Nadig A. 1918. Alcune note sulla fauna dell'alta Valsesia. Formicidae. Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano 56: 331-341.
 * Neumeyer R., and B. Seifert. 2005. Commented check list of free living ant (Hymenoptera: Formicidae) species of Switzerland. Bulletin de la Societe Entomologique Suisse 78: 1-17.
 * Nielsen M. G. 2011. A check list of Danish ants and proposed common names. Ent. Meddr. 79: 13-18.
 * Noordijk, J., R. Morssinkhof, P. Boer, A. P. Schaffers, Th. Heijerman and K. V. Sýkora. 2008. How ants find each other; temporal and spatial patterns in nuptial flights. Insectes Sociaux 55(3):266-273.
 * Nylander, W.. "Synopsis des Formicides de France et d'Algérie." Annales des Sciences Naturelles, Zoologie (4)5 (1856): 51-109.
 * O'Rourke F. J. 1948. The distribution and general ecology of the Irish Formicidae. Proceedings of the Royal Irish Academy. Section B: Biological, Geological, and Chemical Science 52: 383-410.
 * Paik W.H. 1984. A checklist of Formicidae (Hymenoptera) of Korea. Korean J. Plant Prot. 23(3): 193-195.
 * Paraschivescu D. 1978. Elemente balcanice in mirmecofauna R. S. Romania. Nymphaea 6: 463- 474.
 * Park, Seong, Joon and Byung, and Kim, Jin. 2002. Faunal Comparison of Ants among Cheongsando and Other Islands of South Sea in Korea. Korean Jornal of Entomology. 32(1):7-12.
 * Pavlova N. S. 2014. To ant fauna (Hymenoptera, Formicidae) of the floodplain of Medvedista River (Saratov Province). Entomological and parasitological studies in the Volga region 11: 145-147.
 * Petal J. M. 1963. Faune des fourmis de la reserve de tourbiere en projet a Rakowskie Bagno pres de Frampol (voivodie de Lublin). Annales Universitatis Mariae Curie-Sk?odowska 58(7): 143-174.
 * Petrov I. Z., and C. A. Collingwood. 1992. Survey of the myrmecofauna (Formicidae, Hymenoptera) of Yugoslavia. Archives of Biological Sciences (Belgrade) 44: 79-91.
 * Punttila P., Y. Haila, J. Niemela, and T. Pajunen. 1994. Ant communities in fragments of old-growth taiga and managed surroundings. Ann. Zool. Fennici 31: 131-144.
 * Pusvaskyte O. 1979. Myrmecofauna of the Lituanian SSR. Acta Entomologica Lituanica 4: 99-105.
 * Radchenko A. G. 2007. The ants (Hymenoptera, Formicidae) in the collection of William Nylander. Fragmenta Faunistica (Warsaw) 50: 27-41.
 * Radchenko A., W. Czechowska, W. Czechowski, and E. Siedlar. 1999. Four species of the ant genus Lasius F. new to Poland, with additions to the records for previously reported species (Hymenoptera, Formicidae). Fragmenta Faunistica 42(11): 115-121.
 * Radchenko, A. 2005. Monographic revision of the ants (Hymenoptera, Formicidae) of North Korea. Annales Zoologici 55(2): 127-221.
 * Ran H., and S. Y. Zhou. 2012. Checklist of chinese ants: formicomorph subfamilies (Hymenoptera: Formicidae) II. Journal of Guangxi Normal University: Natural Science Edition 30(4): 81-91.
 * Rigato F., and R. Sciaky. 1989. Contributo alla conoscenza della mirmecofauna della Val Gesso (alpi Marittime) (Hymenoptera, Formicidae). Boll. Mus. Reg. Sci. Nat. Torino 7(2): 427-442.
 * Ruzsky M. 1916. On zoological research in Yeniseisk province, work of summer of 1915. Izv. Imp. Tomsk. Univ. 65 (3rd p part: 1-21.
 * Rzeszowski K., H. Babik, W. Czechowski, and B. Marko. 2013. Ants (Hymenoptera: Formicidae) of Chelmowa Gora in the Swietokrzyski National Park. Fragmenta Faunistica 56(1): 1-15.
 * Sakai H. 2002. Reproductive flight season of Japanese ants. Ari 26: 33-39.
 * Salata S., and L. Borowiec. 2018. Taxonomic and faunistic notes on Greek ants (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 27: 1-51.
 * Sato T., N. Tsurusaki, K. Hamaguchi, and K. Kinomura. 2010. Ant fauna of Tottori prefecture, Honshu, Japan. Bulletin of the Tottori Prefectural Museum 47: 27-44.
 * Schar S., G Talavera, X. Espadaler, J. D. Rana, A. A. Andersen, S. P. Cover, and R. Vila. 2018. Do Holarctic ant species exist? Trans-Beringian dispersal and homoplasy in the Formicidae. Journal of Biogeography 00: 1-12.
 * Schifani E., and A. Alicata. 2018. Exploring the myrmecofauna of Sicily: thirty-two new ant species recorded, including six new to Italy and many new aliens (Hymenoptera, Formicidae). Polish Journal of Entomology 87 (4): 323–348.
 * Schlick-Steiner B. C., and F. M. Steiner. 1999. Faunistisch-ökologische Untersuchungen an den freilebenden Ameisen (Hymenoptera: Formicidae) Wiens. Myrmecologische Nachrichten 3: 9-53.
 * Seifert B. 1994. Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3): 1-44.
 * Seifert B. 2007. Die Ameisen Mittel- und Nordeuropas. Tauer: lutra Verlags- und Vertriebsgesellschaft, 368 pp.
 * Sellier Y., C. Galkowski, C. Lebas, and P. Wegnez. 2016. Découverte de Temnothorax pardoi (Tinaut, 1987) dans la réserve naturelle nationale du Pinail (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie 25(2): 106-113.
 * Shimana Y., and S. Yamane. 2009. Geogrpahical distribution of Technomyrmex brunneus Forel (Hymenoptera, Formicidae) in the western part of the mainland of Kagoshima, South Kyushu, Japan. Ari 32: 9-19.
 * Slipinski P., M. Zmihorski, and W. Czechowski. 2012. Species diversity and nestedness of ant assemblages in an urban environment. Eur. J. Entomol. 109: 197206.
 * Sonnenburg H. 2005. Die Ameisenfauna (Hymenoptera: Formicidae) Niedersachsens und Bremens. Braunschweiger Naturkundliche Schriften 7: 377-441.
 * Steiner F. M., S. Schödl, and B. C. Schlick-Steiner. 2002. Liste der Ameisen Österreichs (Hymenoptera: Formicidae), Stand Oktober 2002. Beiträge zur Entomofaunistik 3: 17-25.
 * Stukalyuk S. V. 2015. Structure of the ant assemblages (Hymenoptera, Formicidae) in the broad-leaved forests of Kiev. Entomological Review 95(3): 370–387.
 * Stumper R. 1953. Etudes myrmecologiques. XI. Fourmis luxembourgeoises. Bulletin Soc. Nat. luxemb. 57: 122-135.
 * Stärcke A. 1937. Retouches sur quelques fourmis d'Europe. II. Lasius groupe umbratus Nylander. Tijdschr. Entomol. 80: 38-72.
 * Tamura H., Y. Nakamura, K. Yamauchi, and T. Fujikawa. 1969. An ecological survey of soil fauna in Hidaka-Mombetsu, Southern Hokkaido. Journal of the Faculty of Science Hokkaido University, Zoology 17(1): 17-57.
 * Tausan I., M. M. Jerpel, I. R. Puscasu, C. Sadeanu, R. E. Brutatu, L. A. Radutiu, and V. Giurescu. 2012. Ant fauna (Hymenoptera: Formicidae) of Sibiu County (Transylvania, Romania). Brukenthal. Acta Musei 7(3): 499-520.
 * Terayama M. 1977. Checklist of the known ants of Saitama Prefecture. Insects and nature 12(4): 26-27
 * Terayama M. 1978. Ants of Bukozan. Nature and insects 13(4): 32-34.
 * Terayama M. 1992. Structure of ant communities in East Asia. A. Regional differences and species richness. Bulletin of the Bio-geographical Society of Japan 47: 1-31.
 * Terayama M. 1994. Ants of Okushiri-to Island, Hokkaido. Ari 18: 28-29.
 * Terayama M. 2001. Ants of the Institute for Nature study in Minato-ku, Tokyo: species composition, relative nest abundance, and nest density. Rept. Inst. Nat. Stu. 33: 289-300.
 * Terayama M., Choi, B.M., Kim, C.H. 1992. A check list of ants from Korea, with taxonomic notes. Bulletin of the Toho Gakuen 7:19-54.
 * Terayama M., K. Ogata, and B.M. Choi. 1994. Distribution records of ants in 47 prefectures of Japan. Ari (report of the Myrmecologists Society of Japan) 18: 5-17.
 * Terayama M., S. Kubota, and K. Eguchi. 2014. Encyclopedia of Japanese ants. Asakura Shoten: Tokyo, 278 pp.
 * Terayama M., and R. Sonobe. 2002. Ants from the Nasu Imperial Villa, Tochigi Prefecture, Japan. Flora and fauna of the Tochigi Prefectural Museum research report Nasu Imperial Villa 157-161.
 * Terayama M., and S. Kubota. 2002. Ants of Tokyo, Japan. ARI 26: 1-32.
 * Terayama. M. 1994. Ants of Okushiri-to Island, Hokkaido. ARI Reports of the Myrmecologists Society (Japan) 18: 28-29
 * Terayama. M. 2004. Geological and ecological distribution of Japanese ants communities. (translated from Japanese) Reports of the Saitama Prefecture Animal Research Association. 48:24
 * Terayama. M. 2004. Geological and ecological distribution of Japanese ants communities. (translated from Japanese) Reports of the Saitama Prefecture Animal Research Association. 48:30
 * Teruyama. M. 1988. Ant fauna of Saitama Prefecture, Japan. ARI Reports of the Myrmecologists Society (Japan) 16: 4-13
 * Togashi I., and M. Terayama. 2010. Ant fauna of Hegurajima and Nanatsujima Islands, Ishikawa Prefecture. Bull. Biogeoger. Soc. Japan 65: 195-196.
 * Togashi I., and M. Terayama. 2010. Ant fauna of Hegurajima and Nanatsujima islands, Ishikawa Prefecture. Bull. Biogeoger. Soc. Japan 65: 195-196.
 * Urbieta Balado A., A. Palanca Soler, C. Castan Lanaspa, and X. Espadaler. 1984. Ants in the supraforestal pastures of the Tendenera massif. Documents dEcologie Pyrénéenne III-IV: 223-224.
 * Vepsalainen K., H. Ikonene, and M. J. Koivula. 2008. The structure of ant assembalges in an urban area of Helsinki, southern Finland. Ann. Zool. Fennici 45: 109-127.
 * Verdcourt, B. 2004. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew XXXV. Miscellaneous Records. Kew Bulletin 59(4):639-649
 * Vesni? A. 2011: Revidirani sistematski prijegled mrava Bosne i Hercegovine. Unutar : S. Lelo (urednik),  Fauna  Bosne  i  Hercegovine    Biosistematski  pregledi.  7.  izmijenjeno  i popravljeno interno izdanje Udruenja za inventarizaciju i zatitu  ivotinja, Ilija, Kanton Sarajevo, pp: 205-207.
 * Vogrin, V.. "Prilog fauni Hymenoptera - Aculeata Jugoslavije." Zast. Bilja 31(suppl.) (1955): 1-74.
 * Wegnez P. 2017. Découverte de Myrmica lobicornis Nylander, 1846 et Lasius jensi Seifert, 1982, deux nouvelles espèces pour le Grand-Duché de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 153: 46-49.
 * Wegnez P. 2018. Premières decouvertes de Myrmica bibikoffi Kutter, 1963 et de Ponera testacea Emery, 1895, au Luxembourg (Hymenoptera: Formicidae)Bulletin de la Société royale belge d’Entomologie 154: 263–272.
 * Wegnez P., and M. Fichaux. 2015. Liste actualisee des especes de fourmis repertoriees au Grand-Duche de Luxembourg (Hymenoptera: Formicidae). Bulletin de la Société royale belge d’Entomologie 151: 150-165
 * Wheeler W. M. 1928. Ants collected by Professor F. Silvestri in Japan and Korea. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 22: 96-125.
 * Wilson E. O. 1955. A monographic revision of the ant genus Lasius. Bulletin of the Museum of Comparative Zoology 113: 1-201
 * Xia Yongjuan, and Zheng Zhemin. 1997. A survey of Formicidae form Xinjiang. Journal of Shaanxi Normal University 25(2): 64-66.
 * Yamane S., Y. Harada, and K. Eguchi. 2013. Classification and ecology of ants. Natural history of ants in Southern Kyushu. 200 pages
 * Yamauchi K. 1968. Additional Notes on the Ecological Distribution of Ants in Sapporo and the Vicinity . Journal of the Faculty of Science Hokaido University, series VI, Zoology 16(3): 382-395.
 * Yamauchi K. 1979. Taxonomical and ecological studies on the ant genus Lasius in Japan (Hymenoptera: Formicidae). I. Taxonomy. Sci. Rep. Fac. Educ. Gifu Univ. (Nat. Sci.) 6: 147-181.
 * Yoshimura M. 1998. Ants from Islands in Hokkaido, Northern Japan (No. 1, Risiri Island). Rishiri Studies 17:33-38.
 * Yoshimura M. 1999. Ants in the island of Hokkaido (Part 2, Part Rebun) Ants from Islands in Hokkaido, Northern Japan (No. 2, Rebun-Island). Rishiri Studies 18: 49-54.
 * Zhuytszyuan D. 2016. The ants (Hymenoptera, Formicidae) Nizhne-Svirsky reserve and their environmental features. Master's thesis Saint Petersburg State University.
 * Zryanin V. A., and T. A. Zryanina. 2007. New data on the ant fauna Hymenoptera, Formicidae in the middle Volga River Basin. Uspekhi Sovremennoi Biologii 127(2): 226-240.
 * de Haro, Andrés, and C. A. Collingwood. "Prospección mirmecológica en la Cordillera Ibérica." Orsis 6 (1991): 129-126.