Dolichoderus quadripunctatus

This is a moderately common species in Greece, known from most provinces except Cyclades. It prefers shadow habitats, close to streams and rivers. In Achaia, it was observed in rural sites in urban areas and on a large plane tree (Borowiec & Salata, 2021). Pashaei Rad et al. (2018) found this species in Iran on the ground and on trees in a Caspian moist littoral area.

Distribution
This is an uncommon species although known from almost all Greek provinces except Cyclades (Borowiec et al., 2022).

Distribution based on Regional Taxon Lists
Palaearctic Region: Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, Iran, Liechtenstein, Lithuania, Luxembourg, Monaco, Montenegro, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine.

Biology
Czechowski et al. (2019) observed an alate queen of Dolichoderus quadripunctatus visiting aphids in a Polish garden. The gyne walked along a foraging trail to a colony of the aphid Panaphis juglandis where it imbibed honeydew excreted on the leaf by the aphids, after which it returned to the trail. This recurred during two more days, always a single alate gyne at a time; hence the total number of gynes, one or more, remains an open question. This feeding phenomenon is practically unknown in ants.

Nomenclature

 *  quadripunctatus. Formica quadripunctata Linnaeus, 1771: 541 (w.) FRANCE.
 * [Note: type-locality is now France, after Fabricius, 1775: 392, “Habitat in Alsatia”; the latter confirmed by Latreille, 1802c: 181, “Linnée avoit reçu cette espèce de l’Alsace”.]
 * [Misspelled as quatuorpunctata by Christ, 1791: 513; misspelled as 4 punctatus by Forel, 1895d: 227, and others.]
 * Latreille, 1802c: 179 (q.); Mayr, 1855: 380 (m.); Karavaiev, 1916: 497 (m.).
 * Combination in Tapinoma: Schenck, 1852: 129.
 * Combination in Hypoclinea: Mayr, 1855: 379; Mayr, 1863: 424; Roger, 1863b: 14.
 * Combination in Formica (Hypoclinea): Roger, 1859: 242.
 * Combination in Dolichoderus: Emery & Forel, 1879: 455.
 * Combination in D. (Hypoclinea): Emery, 1894c: 229.
 * Status as species: Fabricius, 1775: 392; Fabricius, 1782: 490; Fabricius, 1787: 308; Gmelin, 1790: 2799; Christ, 1791: 513; Olivier, 1792: 494; Fabricius, 1793: 355; Latreille, 1802c: 179; Walckenaer, 1802: 163; Fabricius, 1804: 403; Losana, 1834: 322; Mayr, 1855: 379 (redescription); Nylander, 1856b: 73; Smith, F. 1858b: 57; Roger, 1859: 242; Mayr, 1861: 40 (in key); Mayr, 1863: 424; Roger, 1863b: 14; Emery, 1869b: 11; Mayr, 1870b: 957 (in key); Forel, 1874: 59 (in key); André, 1874: 186 (in key); Emery, 1878b: 49; Emery & Forel, 1879: 455; André, 1882b: 226 (in key); Mayr, 1886d: 435 (in key); Forel, 1886e: clxvii; Nasonov, 1889: 27; Forel, 1892i: 307; Dalla Torre, 1893: 160; Emery, 1894c: 229; Forel, 1895d: 227; Ruzsky, 1902d: 18; Ruzsky, 1903b: 308; Ruzsky, 1905b: 468; Wasmann, 1906: 115 (in key); Bondroit, 1910: 489; Forel, 1911d: 350; Emery, 1913a: 11; Stitz, 1914: 79; Forel, 1915d: 41 (in key); Emery, 1916b: 210; Karavaiev, 1916: 497; Escherich, 1917: 329; Bondroit, 1918: 87; Menozzi, 1921: 32; Soudek, 1922: 58; Müller, 1923b: 108; Soudek, 1925b: 15; Santschi, 1926f: 287; Karavaiev, 1927c: 272 (in key); Lomnicki, 1928: 6; Kuznetsov-Ugamsky, 1929b: 42; Zimmermann, 1935: 40; Karavaiev, 1936: 165 (redescription); Teranishi, 1940: 19; Novák & Sadil, 1941: 95 (in key); Schmitz, 1950: 14; Consani & Zangheri, 1952: 42; Ceballos, 1956: 309; Bernard, 1967: 250 (redescription); Baroni Urbani, 1971c: 157; van Pisarski, 1975: 27; Boven, 1977: 124; Kutter, 1977c: 172; Arnol’di & Dlussky, 1978: 546 (in key); Collingwood, 1978: 86 (in key); Agosti & Collingwood, 1987b: 278 (in key); Atanassov & Dlussky, 1992: 186; Arakelian, 1994: 71; Radchenko, 1994b: 112 (in key); Shattuck, 1994: 63; Bolton, 1995b: 176; Czechowski, et al. 2002: 13; Seifert, 2007: 252; Casevitz-Weulersse & Galkowsky, 2009: 478; Lapeva-Gjonova, et al. 2010: 31; Boer, 2010: 14; Legakis, 2011: 23; Borowiec, L. & Salata, 2012: 491; Czechowski, et al. 2012: 63; Borowiec, L. 2014: 70 (see note in bibliography); Radchenko, 2016: 82.
 * Senior synonym of kratochvili: Radchenko, 2016: 82.
 * Senior synonym of unicolor: Arakelian, 1994: 72; Radchenko, 2016: 82.
 * kratochvili. Dolichoderus (Hypoclinea) quadripunctatus var. kratochvili Novak, V. 1941: 47, figs. 2-4 (w.) CZECHOSLOVAKIA.
 * Subspecies of quadripunctatus: Novák & Sadil, 1941: 95 (in key) (date of publication (31.xii).1941); Shattuck, 1994: 63.
 * Junior synonym of quadripunctatus: Radchenko, 2016: 82.
 * unicolor. Dolichoderus quadripunctatus var. unicolor Ruzsky, 1905b: 471 (w.q.) CAUCASUS.
 * Senior synonym of unicolor: Arakelian, 1994: 72; Radchenko, 2016: 82.

References based on Global Ant Biodiversity Informatics

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