Wheeler, William Morton (1865-1937)

Biographical Note
Born as the son of Julius Morton and Caroline Georgiana Wheeler (née Anderson) in Milwaukee, he was transferred from public school to a local German academy due to, in his own words, "persistently bad behavior". They had a small museum which Wheeler had studied since he was a child, and when Ward's Natural Science Establishment in early 1884 brought a collection of stuffed and skeletonized animals to the academy, to persuade the city fathers to purchase them, Wheeler volunteered to spend the nights in helping Ward to unpack and install the specimens. The latter was so impressed that he offered Wheeler a job in his Rochester, New York establishment. Here he identified birds and mammals, and later collections of shells, echinoderms and sponges. His shell catalogue was still in use by collectors in the late 1920s.

Wheeler was trained as an insect embryologist, having studied under Baur, Dohrn and Whitman, but he became the leading authority on the behaviour of social insects, achieving particular renown for his studies of social behaviour of ants. He was instrumental in the development of ethology and first popularized the term in a 1902 paper in Science.

He was a taxonomist of the highest order, and was responsible for the descriptions of innumerable species, among them Pogonomyrmex maricopa, the most venomous insect in the world. Professor Wheeler was curator of invertebrate zoology in the American Museum of Natural History, New York, from 1903 to 1908. He was elected to the National Academy of Sciences.

A close contact of the British myrmecologist and coleopterist Horace Donisthorpe, it was to Wheeler that Donisthorpe dedicated his first major book on ants in 1915. Donisthorpe and Wheeler also frequently exchanged specimens, leading the latter to first develop the idea that the Formicinae subfamily had its origins in North America.

He was professor of applied biology at Harvard University's Bussey Institute, which had one of the most highly regarded biology programs in the United States. One of his pupils there was Alfred Kinsey.

His work includes 467 titles

Wheeler began his study of ants while he was a professor of zoology at the University of Texas at Austin (1899–1903), and he greatly extended the scope of his research after becoming curator of invertebrate zoology at the American Museum of Natural History in New York City (1903–08). His investigations dealt particularly with ant taxonomy, morphology, and distribution as well as with ecology, habits, and social relations. He discovered that the social behaviour of ants was among the most complex in the insect world, leading him to use the ant colony as a behavioural analogy for human civilization. His findings were based on firsthand observations of ant species collected from all over the world, including Morocco, the Galapagos, and the Canary Islands.

Later in his career (1930), Wheeler made an important study of the biology of the antlion, the larvae of the neuropteran family Myrmeleontidae.

Student, German-English Academy, Milwaukee, ?-1883

Assistant, Ward's Natural Science Establishment, Rochester, 1884-1885

Teacher, German-English Academy, Milwaukee, 1885-1887

Researcher, Allis Lake Laboratory, 1886-1890

Director, Milwaukee Public Museum, 1887-1890

Summer research at Woods Hole, 1889-1892, 1894, 1898, 1900, etc.

Ph.D. student, Clark University, 1890-1892

Professor, Department of Zoology, University of Chicago, 1892-1899

Research in Münich, Naples, Liège, 1893-1894

Chairman, Department of Zoology, University of Texas, Austin, 1899-1903

American Museum of Natural History, 1903-1908

Harvard University, 1908-1937

Ant Taxonomy
Wheeler published 252 ant taxonomic papers during his lifetime. He published from 1900 until 1942.

Taxonomic Publications

 * Wheeler, W. M. 1900a. The female of Eciton sumichrasti Norton, with some notes on the habits of Texan Ecitons. Am. Nat. 34: 563-574


 * [[Media:1900b.pdf|Wheeler, W. M. 1900b. A study of some Texan Ponerinae. Biol. Bull. (Woods Hole) 2: 1-31 PDF]]


 * [[Media:1901a.pdf|Wheeler, W. M. 1901a. The compound and mixed nests of American ants. Part I. Observations on a new guest ant. Am. Nat. 35: 431-448 PDF]]


 * [[Media:WheelerW1901c.pdf|Wheeler, W. M. 1901g. The parasitic origin of macroërgates among ants. American Naturalist 35:877-886. [1901-11-18] PDF]]


 * [[Media:WheelerW1901a.pdf|Wheeler, W. M. 1901c. The compound and mixed nests of American ants. Part II. The known cases of social symbiosis among American ants. Am. Nat. 35: 513-539 PDF]]


 * [[Media:WheelerW1901b.pdf|Wheeler, W. M. 1901e. The compound and mixed nests of American ants. Part II (continued). Am. Nat. 35: 701-724 PDF]]


 * [[Media:1902a.pdf|Wheeler, W. M. 1902a. A new agricultural ant from Texas, with remarks on the known North American species. Am. Nat. 36: 85-100 PDF]]


 * Wheeler, W. M. 1902b. Formica fusca Linn. subsp. subpolita Mayr, var. perpilosa, n. var. Mem. Rev. Soc. Cient. "Antonio Alzate" 17: 141-142


 * [[Media:Wheeler 1902d.pdf|Wheeler, W. M. 1902d. New agricultural ants from Texas. Psyche (Camb.) 9: 387-393 PDF]]


 * [[Media:Wheeler 1902e.pdf|Wheeler, W. M. 1902e. An American Cerapachys, with remarks on the affinities of the Cerapachyinae. Biol. Bull. (Woods Hole) 3: 181-191 PDF]]


 * [[Media:Wheeler 1902f.pdf|Wheeler, W. M. 1902f. The occurrence of Formica cinerea Mayr and Formica rufibarbis Fabricius in America. Am. Nat. 36: 947-952 PDF]]


 * [[Media:Wheeler 1902g.pdf|Wheeler, W. M. 1902g. A consideration of S. B. Buckley's "North American Formicidae.". Trans. Tex. Acad. Sci. 4: 17-31 PDF]]


 * [[Media:Wheeler 1903a.pdf|Wheeler, W. M. 1903a. Erebomyrma, a new genus of hypogaeic ants from Texas. Biol. Bull. (Woods Hole) 4: 137-148 PDF]]


 * [[Media:Wheeler 1903c.pdf|Wheeler, W. M. 1903c. A decad of Texan Formicidae. Psyche (Camb.) 10: 93-111 PDF]]


 * [[Media:Wheeler 1903d.pdf|Wheeler, W. M. 1903d. A revision of the North American ants of the genus Leptothorax Mayr. Proc. Acad. Nat. Sci. Phila. 55: 215-260 PDF]]


 * [[Media:Wheeler 1903f.pdf|Wheeler, W. M. 1903f. The North American ants of the genus Stenamma sensu stricto. Psyche (Camb.) 10: 164-168 PDF]]


 * [[Media:WheelerW1902.pdf|Wheeler, W. M. 1903g. Extraordinary females in three species of Formica, with remarks on mutation in the Formicidae. Bull. Am. Mus. Nat. Hist. 19: 639-651 PDF]]


 * [[Media:Wheeler 1903h.pdf|Wheeler, W. M. 1903h. Some new gynandromorphous ants, with a review of the previously recorded cases. Bull. Am. Mus. Nat. Hist. 19: 653-683 PDF]]


 * [[Media:Wheeler 1903j.pdf|Wheeler, W. M. 1903j. Some notes on the habits of Cerapachys augustae. Psyche (Camb.) 10: 205-209 PDF]]


 * [[Media:Wheeler 1904a.pdf|Wheeler, W. M. 1904a. Three new genera of inquiline ants from Utah and Colorado. Bull. Am. Mus. Nat. Hist. 20: 1-17 PDF]]


 * [[Media:Wheeler 1904c.pdf|Wheeler, W. M. 1904c. The American ants of the subgenus Colobopsis. Bull. Am. Mus. Nat. Hist. 20: 139-158 PDF]]


 * [[Media:Wheeler 1904d.pdf|Wheeler, W. M. 1904d. A crustacean-eating ant (Leptogenys elongata Buckley). Biol. Bull. (Woods Hole) 6: 251-259 PDF]]


 * [[Media:Wheeler 1904e.pdf|Wheeler, W. M. 1904e. Ants from Catalina Island, California. Bull. Am. Mus. Nat. Hist. 20: 269-271 PDF]]


 * [[Media:Wheeler 1904f.pdf|Wheeler, W. M. 1904f. The ants of North Carolina. Bull. Am. Mus. Nat. Hist. 20: 299-306 PDF]]


 * [[Media:Wheeler 1904i.pdf|Wheeler, W. M. 1904i. A new type of social parasitism among ants. Bull. Am. Mus. Nat. Hist. 20: 347-375 PDF]]


 * [[Media:Wheeler 1905a.pdf|Wheeler, W. M. 1905a. An interpretation of the slave-making instincts in ants. Bull. Am. Mus. Nat. Hist. 21: 1-16 PDF]]


 * [[Media:Wheeler 1905c.pdf|Wheeler, W. M. 1905c. The ants of the Bahamas, with a list of the known West Indian species. Bull. Am. Mus. Nat. Hist. 21: 79-135 PDF]]


 * Wheeler, W. M. 1905d. New species of ''Formica'. Bull. Am. Mus. Nat. Hist. 21: 267-274


 * [[Media:Wheeler 1905g.pdf|Wheeler, W. M. 1905g. The North American ants of the genus Dolichoderus. Bull. Am. Mus. Nat. Hist. 21: 305-319 PDF]]


 * [[Media:Wheeler 1905h.pdf|Wheeler, W. M. 1905h. The North American ants of the genus Liometopum. Bull. Am. Mus. Nat. Hist. 21: 321-333 PDF]]


 * Wheeler 1905j.pdf


 * [[Media:Wheeler 1905l.pdf|Wheeler, W. M. 1905l. Ants from the summit of Mount Washington. Psyche (Cambridge) 12:111-114. [1905-12] PDF]]


 * [[Media:Wheeler 1906e.pdf|Wheeler, W. M. 1906e. On the founding of colonies by queen ants, with special reference to the parasitic and slave-making species. Bull. Am. Mus. Nat. Hist. 22: 33-105 PDF]]


 * [[Media:Wheeler 1906f.pdf|Wheeler, W. M. 1906f. New ants from New England. Psyche (Camb.) 13: 38-41 PDF]]


 * [[Media:Wheeler 1906g.pdf|Wheeler, W. M. 1906g. Fauna of New England. 7. List of the Formicidae. Occas. Pap. Boston Soc. Nat. Hist. 7: 1-24 PDF]]


 * [[Media:Wheeler 1906h.pdf|Wheeler, W. M. 1906h. The ants of Japan. Bull. Am. Mus. Nat. Hist. 22: 301-328 PDF]]


 * [[Media:Wheeler 1906i.pdf|Wheeler, W. M. 1906i. The ants of the Grand Cañon. Bull. Am. Mus. Nat. Hist. 22: 329-345 PDF]]


 * [[Media:Wheeler 1906j.pdf|Wheeler, W. M. 1906j. The ants of the Bermudas. Bull. Am. Mus. Nat. Hist. 22: 347-352 PDF]]


 * [[Media:Wheeler 1907b.pdf|Wheeler, W. M. 1907b. A collection of ants from British Honduras. Bull. Am. Mus. Nat. Hist. 23: 271-277 PDF]]


 * [[Media:Wheeler 1907c.pdf|Wheeler, W. M. 1907c. Notes on a new guest-ant, Leptothorax glacialis, and the varieties of Myrmica brevinodis Emery. Bull. Wis. Nat. Hist. Soc. 5: 70-83 PDF]]


 * [[Media:Wheeler 1907d.pdf|Wheeler, W. M. 1907d. The fungus-growing ants of North America. Bull. Am. Mus. Nat. Hist. 23: 669-807 PDF]]


 * Wheeler, W. M. 1908a. The ants of Porto Rico and the Virgin Islands. Bull. Am. Mus. Nat. Hist. 24: 117-158


 * Wheeler, W. M. 1908b. The ants of Jamaica. Bull. Am. Mus. Nat. Hist. 24: 159-163


 * Wheeler, W. M. 1908c. Ants from Moorea, Society Islands. Bull. Am. Mus. Nat. Hist. 24: 165-167


 * [[Media:WheelerW1908.pdf|Wheeler, W. M. 1908g. Honey ants, with a revision of the American Myrmecocysti. Bull. Am. Mus. Nat. Hist. 24: 345-397 PDF]]


 * [[Media:WheelerW1908h.pdf|Wheeler, W. M. 1908h. The ants of Texas, New Mexico and Arizona. (Part I.). Bull. Am. Mus. Nat. Hist. 24: 399-485 PDF]]


 * [[Media:Wheeler 1908i.pdf|Wheeler, W. M. 1908i. The ants of Casco Bay, Maine, with observations on two races of Formica sanguinea Latreille. Bull. Am. Mus. Nat. Hist. 24: 619-645 PDF]]


 * [[Media:Wheeler 1908k.pdf|Wheeler, W. M. 1908k. Comparative ethology of the European and North American ants. J. Psychol. Neurol. 13: 404-435 PDF]]


 * [[Media:Wheeler 1909a.pdf|Wheeler, W. M. 1909a. A small collection of ants from Victoria, Australia. J. N. Y. Entomol. Soc. 17: 25-29 PDF]]


 * [[Media:Wheeler 1909b.pdf|Wheeler, W. M. 1909b. Ants collected by Prof. F. Silvestri in Mexico. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 3: 228-238 PDF]]


 * [[Media:Wheeler 1909c.pdf|Wheeler, W. M. 1909c. Ants collected by Professor Filippo Silvestri in the Hawaiian Islands. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 3: 269-272 PDF]]


 * [[Media:WheelerW1909.pdf|Wheeler, W. M. 1909d. Ants of Formosa and the Philippines. Bull. Am. Mus. Nat. Hist. 26: 333-345 PDF]]


 * [[Media:Wheeler 1909e.pdf|Wheeler, W. M. 1909e. A decade of North American Formicidae. J. N. Y. Entomol. Soc. 17: 77-90 PDF]]


 * [[Media:Wheeler 1909f.pdf|Wheeler, W. M. 1909f. A new honey ant from California. J. N. Y. Entomol. Soc. 17: 98-99 PDF]]


 * [[Media:Wheeler 1909g.pdf|Wheeler, W. M. 1909g. Observations on some European ants. J. N. Y. Entomol. Soc. 17: 172-187 PDF]]


 * [[Media:Wheeler 1910a.pdf|Wheeler, W. M. 1910a. Small artificial ant-nests of novel patterns. Psyche (Cambridge) 17:73-75. [1910-04]PDF]]


 * Wheeler, W. M. 1910b. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp.


 * Wheeler, W. M. 1910d. Three new genera of myrmicine ants from tropical America. Bull. Am. Mus. Nat. Hist. 28: 259-265


 * [[Media:Wheeler 1910e.pdf|Wheeler, W. M. 1910e. An aberrant Lasius from Japan. Biol. Bull. (Woods Hole) 19: 130-137 PDF]]


 * [[Media:Wheeler 1910f.pdf|Wheeler, W. M. 1910f. A new species of Aphomomyrmex from Borneo. Psyche (Camb.) 17: 131-135 PDF]]


 * [[Media:Wheeler 1910g.pdf|Wheeler, W. M. 1910g. The North American ants of the genus Camponotus Mayr. Ann. N. Y. Acad. Sci. 20: 295-354 PDF]]


 * [[Media:Wheeler 1910.pdf|Wheeler, W. M. 1910h. The North American forms of Lasius umbratus Nylander. Psyche (Camb.) 17: 235-243 PDF]]


 * [[Media:Wheeler 1910i.pdf|Wheeler, W. M. 1910i. The North American forms of Camponotus fallax Nylander. J. N. Y. Entomol. Soc. 18: 216-232 PDF]]


 * [[Media:Wheeler 1911a.pdf|Wheeler, W. M. 1911a. Additions to the ant-fauna of Jamaica. Bull. Am. Mus. Nat. Hist. 30: 21-29 PDF]]


 * [[Media:Wheeler 1911b.pdf|Wheeler, W. M. 1911b. Ants collected in Grenada, W. I. by Mr. C. T. Brues. Bull. Mus. Comp. Zool. 54: 167-172 PDF]]


 * [[Media:Wheeler 1911c.pdf|Wheeler, W. M. 1911c. Three formicid names which have been overlooked. Science (N. Y.) (n.s.) 33: 858-860 PDF]]


 * [[Media:Wheeler 1911d.pdf|Wheeler, W. M. 1911d. A new Camponotus from California. J. N. Y. Entomol. Soc. 19: 96-98 PDF]]


 * [[Media:Wheeler 1911e.pdf|Wheeler, W. M. 1911e. Two fungus-growing ants from Arizona. Psyche (Camb.) 18: 93-101 PDF]]


 * [[Media:Wheeler 1911g.pdf|Wheeler, W. M. 1911g. A list of the type species of the genera and subgenera of Formicidae. Ann. N. Y. Acad. Sci. 21: 157-175 PDF]]


 * [[Media:Wheeler 1911i.pdf|Wheeler, W. M. 1911i. Descriptions of some new fungus-growing ants from Texas, with Mr. C. G. Hartman's observations on their habits. J. N. Y. Entomol. Soc. 19: 245-255 PDF]]


 * [[Media:Wheeler 1911k.pdf|Wheeler, W. M. 1911k. Three new ants from Mexico and Central America. Psyche (Camb.) 18: 203-208 PDF]]


 * [[Media:Wheeler 1912a.pdf|Wheeler, W. M. 1912a. The ants of Guam. J. N. Y. Entomol. Soc. 20: 44-48 PDF]]


 * [[Media:Wheeler 1912b.pdf|Wheeler, W. M. 1912b. Notes on a mistletoe ant. J. N. Y. Entomol. Soc. 20: 130-133 PDF]]


 * [[Media:Wheeler 1912c.pdf|Wheeler, W. M. 1912c. New names for some ants of the genus Formica. Psyche (Camb.) 19: 90 PDF]]


 * [[Media:Wheeler 1912e.pdf|Wheeler, W. M. 1912e. Additions to our knowledge of the ants of the genus Myrmecocystus Wesmael. Psyche (Camb.) 19: 172-181 PDF]]


 * [[Media:Wheeler 1912f.pdf|Wheeler, W. M. 1912f. The male of Eciton vagans Olivier. Psyche (Camb.) 19: 206-207 PDF]]


 * [[Media:Wheeler 1913a.pdf|Wheeler, W. M. 1913a. Corrections and additions to "List of type species of the genera and subgenera of Formicidae". Ann. N. Y. Acad. Sci. 23: 77-83 PDF]]


 * [[Media:Wheeler 1913b.pdf|Wheeler, W. M. 1913b. The ants of Cuba. Bull. Mus. Comp. Zool. 54: 477-505 PDF]]


 * Wheeler, W. M. 1913c. [Untitled. Description of Iridomyrmex humilis Mayr.]. Pp. 27-29 in: Newell, W., Barber, T. C. The Argentine ant. U. S. Dep. Agric. Bur. Entomol. Bull. 122:1-98.


 * [[Media:Wheeler 1913d.pdf|Wheeler, W. M. 1913d. Ants collected in Georgia by Dr. J. C. Bradley and Mr. W. T. Davis. Psyche (Camb.) 20: 112-117 PDF]]


 * [[Media:Wheeler 1913e.pdf|Wheeler, W. M. 1913e. Ants collected in the West Indies. Bull. Am. Mus. Nat. Hist. 32: 239-244 PDF]]


 * [[Media:Wheeler 1913f.pdf|Wheeler, W. M. 1913f. Zoological results of the Abor Expedition, 1911-1912, XVII. Hymenoptera, II: Ants (Formicidae). Rec. Indian Mus. 8: 233-237 PDF]]


 * [[Media:Wheeler 1913i.pdf|Wheeler, W. M. 1913i. A revision of the ants of the genus Formica (Linné) Mayr. Bull. Mus. Comp. Zool. 53: 379-565 PDF]]


 * [[Media:Wheeler 1914a.pdf|Wheeler, W. M. 1914a. Gynandromorphous ants described during the decade 1903-1913. Am. Nat. 48: 49-56 PDF]]


 * [[Media:Wheeler 1914b.pdf|Wheeler, W. M. 1914b. Formica exsecta in Japan. Psyche (Camb.) 21: 26-27 PDF]]


 * [[Media:1914c.pdf|Wheeler, W. M. 1914c. Ants collected by W. M. Mann in the state of Hidalgo, Mexico. J. N. Y. Entomol. Soc. 22: 37-61 PDF]]


 * [[Media:Wheeler 1914.pdf|Wheeler, W. M. 1914d. Note on the habits of Liomyrmex. Psyche (Camb.) 21: 75-76 PDF]]


 * [[Media:Wheeler 1914e.pdf|Wheeler, W. M. 1914e. The American species of Myrmica allied to M. rubida Latreille. Psyche (Camb.) 21: 118-122 PDF]]


 * [[Media:Wheeler 1914f.pdf|Wheeler, W. M. 1914f. New and little known harvesting ants of the genus Pogonomyrmex. Psyche (Camb.) 21: 149-157 PDF]]


 * [[Media:Wheeler 1915a.pdf|Wheeler, W. M. 1915a. Neomyrma versus Oreomyrma. A correction. Psyche (Camb.) 22: 50 PDF]]


 * [[Media:Wheeler 1915b.pdf|Wheeler, W. M. 1915b. Some additions to the North American ant-fauna. Bull. Am. Mus. Nat. Hist. 34: 389-421 PDF]]


 * [[Media:WheelerW1915B.pdf|Wheeler, W. M. 1915c. Paranomopone, a new genus of ponerine ants from Queensland. Psyche (Camb.) 22: 117-120 PDF]]


 * [[Media:WheelerW1915A.pdf|Wheeler, W. M. 1915e. The Australian honey-ants of the genus Leptomyrmex Mayr. Proc. Am. Acad. Arts Sci. 51: 255-286 PDF]]


 * [[Media:Wheeler 1915f.pdf|Wheeler, W. M. 1915f. Two new genera of myrmicine ants from Brasil. Bull. Mus. Comp. Zool. 59: 483-491 PDF]]


 * [[Media:Wheeler 1915.pdf|Wheeler, W. M. 1915g. A new bog-inhabiting variety of Formica fusca L. Psyche (Camb.) 22: 203-206 PDF]]


 * [[Media:Wheeler 1915h.pdf|Wheeler, W. M. 1915h. Hymenoptera. [In "Scientific notes on an expedition into the north-western regions of South Australia".]. Trans. R. Soc. S. Aust. 39: 805-823 PDF]]


 * [[Media:Wheeler 1915i.pdf|Wheeler, W. M. 1915i [1914]. The ants of the Baltic Amber. Schr. Phys.-Ökon. Ges. Königsb. 55: 1-142 PDF]]


 * [[Media:WheelerW1916c.pdf|Wheeler, W. M. 1916a. The Australian ants of the genus Onychomyrmex. Bull. Mus. Comp. Zool. 60: 45-54 PDF]]


 * [[Media:Wheeler 1916b.pdf|Wheeler, W. M. 1916b. Four new and interesting ants from the mountains of Borneo and Luzon. Proc. N. Engl. Zool. Club 6: 9-18 PDF]]


 * [[Media:Wheeler 1916c.pdf|Wheeler, W. M. 1916c. Ants collected in British Guiana by the expedition of the American Museum of Natural History during 1911. Bull. Am. Mus. Nat. Hist. 35: 1-14 PDF]]


 * [[Media:Wheeler 1916f.pdf|Wheeler, W. M. 1916f. Ants collected in Trinidad by Professor Roland Thaxter, Mr. F. W. Urich, and others. Bull. Mus. Comp. Zool. 60: 323-330 PDF]]


 * [[Media:Wheeler 1916g.pdf|Wheeler, W. M. 1916g. Some new formicid names. Psyche (Camb.) 23: 40-41 PDF]]


 * [[Media:Wheeler 1916i.pdf|Wheeler, W. M. 1916i. Two new ants from Texas and Arizona. Proc. N. Engl. Zool. Club 6: 29-35 PDF]]


 * [[Media:Wheeler 1916i.pdf|Wheeler, W. M. 1916j. Note on the Brazilian fire-ant, Solenopsis saevissima F. Smith. Psyche (Camb.) 23: 142-143 PDF]]


 * [[Media:WheelerW1916d.pdf|Wheeler, W. M. 1916k. An anomalous blind worker ant. Psyche (Camb.) 23: 143-145 PDF]]


 * [[Media:WheelerW1916b.pdf|Wheeler, W. M. 1916m. Prodiscothyrea, a new genus of ponerine ants from Queensland. Trans. R. Soc. S. Aust. 40: 33-37 PDF]]


 * [[Media:WheelerW1916a.pdf|Wheeler, W. M. 1916n. The Australian ants of the genus Aphaenogaster Mayr. Trans. R. Soc. S. Aust. 40: 213-223 PDF]]


 * [[Media:WheelerW1916e.pdf|Wheeler, W. M. 1916o. Questions of nomenclature connected with the ant genus Lasius and its subgenera. Psyche (Camb.) 23: 168-173 'PDF]]


 * [[Media:WheelerW1916f.pdf|Wheeler, W. M. 1916p. Ants carried in a floating log from the Brazilian mainland to San Sebastian Island. Psyche (Camb.) 23: 180-183 PDF]]


 * [[Media:Wheeler 1916.pdf|Wheeler, W. M. 1916q. A phosphorescent ant. Psyche (Cambridge) 23:173-174. [1916-12] PDF]]


 * Wheeler, W. M. 1916r. Formicoidea. Formicidae. Pp. 577-601 in: Viereck, H. L. Guide to the insects of Connecticut. Part III. The Hymenoptera, or wasp-like insects, of Connecticut. Conn. State Geol. Nat. Hist. Surv. Bull. 22:1-824.


 * [[Media:WheelerW1917e.pdf|Wheeler, W. M. 1917a. The mountain ants of western North America. Proc. Am. Acad. Arts Sci. 52: 457-569 PDF]]


 * [[Media:WheelerW1919c.pdf|Wheeler, W. M. 1917b. The phylogenetic development of subapterous and apterous castes in the Formicidae. Proc. Natl. Acad. Sci. U. S. A. 3: 109-117 PDF]]


 * [[Media:WheelerW1917f.pdf|Wheeler, W. M. 1917c. The North American ants described by Asa Fitch. Psyche (Camb.) 24: 26-29 PDF]]


 * [[Media:Wheeler 1917d.pdf|Wheeler, W. M. 1917d. A new Malayan ant of the genus Prodiscothyrea. Psyche (Camb.) 24: 29-30 PDF]]


 * [[Media:WheelerW1917d.pdf|Wheeler, W. M. 1917e. The ants of Alaska. Bull. Mus. Comp. Zool. 61: 13-23 PDF]]


 * [[Media:WheelerW1917b.pdf|Wheeler, W. M. 1917f. The Australian ant-genus Myrmecorhynchus (Ern. André) and its position in the subfamily Camponotinae. Trans. R. Soc. S. Aust. 41: 14-19 PDF]]


 * [[Media:Wheeler 1917g.pdf|Wheeler, W. M. 1917g. Jamaican ants collected by Prof. C. T. Brues. Bull. Mus. Comp. Zool. 61: 457-471 PDF]]


 * [[Media:WheelerW1917g.pdf|Wheeler, W. M. 1917h. The temporary social parasitism of Lasius subumbratus Viereck. Psyche (Camb.) 24: 167-176 PDF]]


 * [[Media:Wheeler 1917.pdf|Wheeler, W. M. 1917i. Notes on the marriage flights of some Sonoran ants. Psyche (Cambridge) 24:177-180. [1917-12] PDF]]


 * [[Media:Wheeler 1917.pdf|Wheeler, W. M. 1917j. The pleometrosis of Myrmecocystus. Psyche (Cambridge) 24:180-182. [1917-12] PDF]]


 * [[Media:WheelerW1917d.pdf|Wheeler, W. M. 1917k. A list of Indiana ants. Proc. Indiana Acad. Sci. 26: 460-466 PDF]]


 * [[Media:WheelerW1918b.pdf|Wheeler, W. M. 1918a. The Australian ants of the ponerine tribe Cerapachyini. Proc. Am. Acad. Arts Sci. 53: 215-265 PDF]]


 * [[Media:WheelerW1918.pdf|Wheeler, W. M. 1918b. Ants collected in British Guiana by Mr. C. William Beebe. J. N. Y. Entomol. Soc. 26: 23-28 PDF]]


 * [[Media:WheelerW1918c.pdf|Wheeler, W. M. 1918c. A study of some ant larvae, with a consideration of the origin and meaning of the social habit among insects. Proc. Am. Philos. Soc. 57: 293-343 PDF]]


 * [[Media:WheelerW1918a.pdf|Wheeler, W. M. 1918d. The ants of the genus Opisthopsis Emery. Bull. Mus. Comp. Zool. 62: 341-362 PDF]]


 * [[Media:WheelerW1919.pdf|Wheeler, W. M. 1919a. Two gynandromorphous ants. Psyche (Camb.) 26: 1-8 PDF]]


 * [[Media:WheelerW1919e.pdf|Wheeler, W. M. 1919b. A new subspecies of Aphaenogaster treatae Forel. Psyche (Camb.) 26: 50 PDF]]


 * [[Media:WheelerW1919d.pdf|Wheeler, W. M. 1919c. The parasitic Aculeata, a study in evolution. Proceedings of the American Philosophical Society 58:1-40. [1919-06-11] PDF]]


 * [[Media:WheelerW1919g.pdf|Wheeler, W. M. 1919d. Expedition of the California Academy of Sciences to the Galapagos Islands, 1905-1906. XIV. The ants of the Galapagos Islands. Proc. Calif. Acad. Sci. (4)2(2 2: 259-297 PDF]]


 * [[Media:Wheeler 1919e.pdf|Wheeler, W. M. 1919e. Expedition of the California Academy of Sciences to the Galapagos Islands, 1905-1906. XV. The ants of Cocos Island. Proc. Calif. Acad. Sci. (4)2(2 2: 299-308 PDF]]


 * Wheeler, W. M. 1919f. The ants of Borneo. Bull. Mus. Comp. Zool. 63: 43-147


 * [[Media:WheelerW1919b.pdf|Wheeler, W. M. 1919g. The ant genus Lordomyrma Emery. Psyche (Camb.) 26: 97-106 PDF]]


 * [[Media:WheelerW1919h.pdf|Wheeler, W. M. 1919h. A new paper-making Crematogaster from the southeastern United States. Psyche (Camb.) 26: 107-112 PDF]]


 * [[Media:Wheeler 1919.pdf|Wheeler, W. M. 1919i. The ants of Tobago Island. Psyche (Cambridge) 26:113. [1919-08] PDF]]


 * [[Media:WheelerW1919a.pdf|Wheeler, W. M. 1919j. The ants of the genus Metapone Forel. Ann. Entomol. Soc. Am. 12: 173-191 PDF]]


 * [[Media:WheelerW1919i.pdf|Wheeler, W. M. 1919k. A singular neotropical ant (Pseudomyrma filiformis Fabricius). Psyche (Camb.) 26: 124-131 PDF]]


 * [[Media:WheelerW1920.pdf|Wheeler, W. M. 1920. The subfamilies of Formicidae, and other taxonomic notes. Psyche (Camb.) 27: 46-55 PDF]]


 * [[Media:WheelerW1921.pdf|Wheeler, W. M. 1921a. Professor Emery's subgenera of the genus Camponotus Mayr. Psyche (Camb.) 28: 16-19 PDF]]


 * [[Media:WheelerW1921a.pdf|Wheeler, W. M. 1921b. A new case of parabiosis and the "ant gardens" of British Guiana. Ecology 2: 89-103 PDF]]


 * [[Media:Wheeler 1921c.pdf|Wheeler, W. M. 1921c. Chinese ants. Bull. Mus. Comp. Zool. 64: 529-547 PDF]]


 * [[Media:WheelerW1921b.pdf|Wheeler, W. M. 1921d. Observations on army ants in British Guiana. Proc. Am. Acad. Arts Sci. 56: 291-328 PDF]]


 * [[Media:WheelerW1921c.pdf|Wheeler, W. M. 1921e. Chinese ants collected by Prof. C. W. Howard. Psyche (Camb.) 28: 110-115 PDF]]


 * [[Media:WheelerW1921d.pdf|Wheeler, W. M. 1921f. The Tachigalia ants. Zoologica (N. Y.) 3: 137-168 PDF]]


 * [[Media:Wheeler 1922a.pdf|Wheeler, W. M. 1922a. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. I. On the distribution of the ants of the Ethiopian and Malagasy regions. Bull. Am. Mus. Nat. Hist. 45: 13-37 PDF]]


 * Wheeler, W. M. 1922b. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. II. The ants collected by the American Museum Congo Expedition. Bull. Am. Mus. Nat. Hist. 45: 39-269


 * [[Media:WheelerW1922d.pdf|Wheeler, W. M. 1922c. Observations on Gigantiops destructor Fabricius and other leaping ants. Biol. Bull. (Woods Hole) 42: 185-201 PDF]]


 * [[Media:Wheeler WM 1922.pdf|Wheeler, W. M. 1922d. Ants of the genus Formica in the tropics. Psyche (Cambridge) 29:174-177. [1922-08] PDF]]


 * [[Media:WheelerW1922e.pdf|Wheeler, W. M. 1922e. The ants of Trinidad. Am. Mus. Novit. 45: 1-16 PDF]]


 * [[Media:WheelerW1922f.pdf|Wheeler, W. M. 1922f. A new genus and subgenus of Myrmicinae from tropical America. Am. Mus. Novit. 46: 1-6 PDF]]


 * [[Media:WheelerW1922c.pdf|Wheeler, W. M. 1922g. Neotropical ants of the genera Carebara, Tranopelta and Tranopeltoides, new genus. Am. Mus. Novit. 48: 1-14 PDF]]


 * [[Media:WheelerW1922B.pdf|Wheeler, W. M. 1922i. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VII. Keys to the genera and subgenera of ants. Bull. Am. Mus. Nat. Hist. 45: 631-710 PDF]]


 * [[Media:Wheeler 1922j.pdf|Wheeler, W. M. 1922j. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bull. Am. Mus. Nat. Hist. 45: 711-1004 PDF]]


 * Wheeler, W. M. 1922k. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bull. Am. Mus. Nat. Hist. 4 45: 1005-1055


 * [[Media:WheelerW1922.pdf|Wheeler, W. M. 1922l. Social life among the insects. Lecture IV - Ants, their development, castes, nesting and feeding habits. The Scientific Monthly 15:385-404. [1922-11] PDF]]


 * [[Media:WheelerW1922A.pdf|Wheeler, W. M. 1922n. Formicidae from Easter Island and Juan Fernandez. Pp. 317-319 in: Skottsberg, C. (ed.) 1921-1940. The natural history of Juan Fernandez and Easter Island. Vol. III. Zoology. Uppsala: Almqvist & Wiksells, 688 pp. PDF]]


 * [[Media:WheelerW1923.pdf|Wheeler, W. M. 1923a. Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn Dr. A. Roman in Amazonas 1914-1915. 7. Formicidae. Ark. Zool. 15(7 7: 1-6 PDF]]


 * Wheeler, W. M. 1923b. Social life among the insects. Lecture V. Parasitic ants and ant guests. Sci. Mon. 16: 5-33


 * [[Media:WheelerW1923d.pdf|Wheeler, W. M. 1923c. Chinese ants collected by Professor S. F. Light and Professor A. P. Jacot. Am. Mus. Novit. 69: 1-6 PDF]]


 * [[Media:WheelerW1923e.pdf|Wheeler, W. M. 1923d. Report on the ants collected by the Barbados-Antigua Expedition from the University of Iowa in 1918. Stud. Nat. Hist. Iowa Univ. 10(3 3: 3-9 PDF]]


 * [[Media:WheelerW1923c.pdf|Wheeler, W. M. 1923e. The occurrence of winged females in the ant genus Leptogenys Roger, with descriptions of new species. Am. Mus. Novit. 90: 1-16 PDF]]


 * [[Media:WheelerW1923b.pdf|Wheeler, W. M. 1923f. Ants of the genera Myopias and Acanthoponera. Psyche (Camb.) 30: 175-192 PDF]]


 * [[Media:WheelerW1924.pdf|Wheeler, W. M. 1924b. The Formicidae of the Harrison Williams Galapagos Expedition. Zoologica (N. Y.) 5: 101-122 PDF]]


 * [[Media:WheelerW1924e.pdf|Wheeler, W. M. 1924c. Ants of Krakatau and other islands in the Sunda Strait. Treubia 5: 239-258 PDF]]


 * [[Media:WheelerW1924b.pdf|Wheeler, W. M. 1924d. A gynandromorph of Tetramorium guineense Fabr. Psyche (Camb.) 31: 136-137 PDF]]


 * [[Media:WheelerW1924c.pdf|Wheeler, W. M. 1924e. On the ant-genus Chrysapace Crawley. Psyche (Camb.) 31: 224-225 PDF]]


 * [[Media:WheelerW1925.pdf|Wheeler, W. M. 1925a. Neotropical ants in the collections of the Royal Museum of Stockholm. Ark. Zool. 17A(8 8: 1-55 PDF]]


 * [[Media:WheelerW1924d.pdf|Wheeler, W. M. 1925b. Zoological results of the Swedish Expedition to Central Africa 1921. Insecta. 10. Formicidae. Ark. Zool. 17A(2 25: 1-3 PDF]]


 * [[Media:WheelerW1925a.pdf|Wheeler, W.M. 1925c. Obituary. Carlo Emery. (1925). Entomological News36: 318 - 320 PDF]]


 * [[Media:WheelerW1925c.pdf|Wheeler, W. M. 1925d. The finding of the queen of the army ant Eciton hamatum Fabricius. Biol. Bull. (Woods Hole) 49: 139-149 PDF]]


 * [[Media:WheelerW1925d.pdf|Wheeler, W. M. 1925e. A new guest-ant and other new Formicidae from Barro Colorado Island, Panama. Biol. Bull. (Woods Hole) 49: 150-181 PDF]]


 * [[Media:Wheeler 1926a.pdf|Wheeler, W. M. 1926a. Ants of the Balearic Islands. Folia Myrmecol. Termit. 1: 1-6 PDF]]


 * [[Media:WheelerW1927.pdf|Wheeler, W. M. 1927a. The occurrence of Formica fusca L. in Sumatra. Psyche (Camb.) 34: 40-41 PDF]]


 * [[Media:WheelerW1927g.pdf|Wheeler, W. M. 1927b. Burmese ants collected by Professor G. E. Gates. Psyche (Camb.) 34: 42-46 PDF]]


 * [[Media:WheelerW1927a.pdf|Wheeler, W. M. 1927c. Ants of the genus Amblyopone Erichson. Proc. Am. Acad. Arts Sci. 62: 1-29 PDF]]


 * [[Media:WheelerW1927f.pdf|Wheeler, W. M. 1927d. Chinese ants collected by Professor S. F. Light and Professor N. Gist Gee. Am. Mus. Novit. 255: 1-12 PDF]]


 * [[Media:WheelerW1927h.pdf|Wheeler, W. M. 1927e. A few ants from China and Formosa. Am. Mus. Novit. 259: 1-4 PDF]]


 * [[Media:WheelerW1927i.pdf|Wheeler, W. M. 1927f. The physiognomy of insects. Q. Rev. Biol. 2: 1-36 PDF]]


 * [[Media:Wheeler 1927g.pdf|Wheeler, W. M. 1927g. The ants of the Canary Islands. Proc. Am. Acad. Arts Sci. 62: 93-120 PDF]]


 * [[Media:WheelerW1927j.pdf|Wheeler, W. M. 1927h. Ants collected by Professor F. Silvestri in Indochina. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 20: 83-106 PDF]]


 * [[Media:WheelerW1927b.pdf|Wheeler, W. M. 1927i. The ants of Lord Howe Island and Norfolk Island. Proc. Am. Acad. Arts Sci. 62: 121-153 PDF]]


 * [[Media:Wheeler 1927j.pdf|Wheeler, W. M. 1927j. The occurrence of the pavement ant (Tetramorium caespitum L.) in Boston. Psyche (Cambridge) 34:164-165. [1927-08] PDF]]


 * [[Media:WheelerW1928.pdf|Wheeler, W. M. 1928a. Zatapinoma, a new genus of ants from India. Proc. N. Engl. Zool. Club 10: 19-23 PDF]]


 * [[Media:WheelerW1928a.pdf|Wheeler, W. M. 1928b. Mermis parasitism and intercastes among ants. J. Exp. Zool. 50: 165-237 PDF]]


 * [[Media:WheelerW1928b.pdf|Wheeler, W. M. 1928c. Ants collected by Professor F. Silvestri in China. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 22: 3-38 PDF]]


 * [[Media:WheelerW1928c.pdf|Wheeler, W. M. 1928d. Ants collected by Professor F. Silvestri in Japan and Korea. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 22: 96-125 PDF]]


 * [[Media:WheelerW1928d.pdf|Wheeler, W. M. 1928e. A new species of Probolomyrmex from Java. Psyche (Camb.) 35: 7-9 PDF]]


 * [[Media:Wheeler 1928f.pdf|Wheeler, W. M. 1928f. Ants of Nantucket Island, Mass. Psyche (Cambridge) 35:10-11. [1928-03] PDF]]


 * [[Media:WheelerW1929.pdf|Wheeler, W. M. 1929a. The identity of the ant genera Gesomyrmex Mayr and Dimorphomyrmex Ernest André. Psyche (Camb.) 36: 1-12 PDF]]


 * [[Media:WheelerW1929a.pdf|Wheeler, W. M. 1929b. Three new genera of ants from the Dutch East Indies. Am. Mus. Novit. 349: 1-8 PDF]]


 * [[Media:Wheeler 1929c.pdf|Wheeler, W. M. 1929c. Two Neotropical ants established in the United States. Psyche (Cambridge) 36:89-90. [1929-06] '''PDF]]


 * [[Media:WheelerW1929b.pdf|Wheeler, W. M. 1929d. Note on Gesomyrmex. Psyche (Camb.) 36: 91-92 PDF]]


 * [[Media:WheelerW1929c.pdf|Wheeler, W. M. 1929e. The ant genus Rhopalomastix. Psyche (Camb.) 36: 95-101 PDF]]


 * [[Media:WheelerW1929d.pdf|Wheeler, W. M. 1929f. A Camponotus mermithergate from Argentina. Psyche (Camb.) 36: 102-106 PDF]]


 * [[Media:WheelerW1929e.pdf|Wheeler, W. M. 1929g. Some ants from China and Manchuria. Am. Mus. Novit. 361: 1-11 PDF]]


 * [[Media:WheelerW1929f.pdf|Wheeler, W. M. 1929h. Ants collected by Professor F. Silvestri in Formosa, the Malay Peninsula and the Philippines. Boll. Lab. Zool. Gen. Agrar. R. Sc. Super. Agric. 24: 27-64 PDF]]


 * [[Media:Wheeler 1929i.pdf|Wheeler, W. M. 1929i. Two interesting Neotropical myrmecophytes (Cordia nodosa and C. alliodora). Pp. 342-353 in: Jordan, K., Horn, W. (eds.) Fourth International Congress of Entomology, Ithaca, August 1928. Volume II. Transactions. Naumburg a. Saale: G. Pätz, vii + 1037 pp. PDF]]


 * Wheeler, W. M. 1929j. Part II. Formicidae. Pp. 29-39 in: Wheeler, W. M., Bequaert, J. C. Amazonian myrmecophytes and their ants. Zool. Anz. 82:10-39.


 * [[Media:WheelerW1930.pdf|Wheeler, W. M. 1930b. Formosan ants collected by Dr. R. Takahashi. Proc. N. Engl. Zool. Club 11: 93-106 PDF]]


 * [[Media:WheelerW1930h.pdf|Wheeler, W. M. 1930c. The ant Prenolepis imparis Say. Ann. Entomol. Soc. Am. 23: 1-26 PDF]]


 * [[Media:WheelerW1930i.pdf|Wheeler, W. M. 1930d. A second note on Gesomyrmex. Psyche (Camb.) 37: 35-40 PDF]]


 * [[Media:WheelerW1930a.pdf|Wheeler, W. M. 1930e. Two new genera of ants from Australia and the Philippines. Psyche (Camb.) 37: 41-47 PDF]]


 * [[Media:WheelerW1930d.pdf|Wheeler, W. M. 1930g. A new parasitic Crematogaster from Indiana. Psyche (Camb.) 37: 55-60 PDF]]


 * [[Media:WheelerW1930e.pdf|Wheeler, W. M. 1930h. A new Emeryella from Panama. Proc. N. Engl. Zool. Club 12: 9-13 PDF]]


 * [[Media:Wheeler 1930i.pdf|Wheeler, W. M. 1930i. [Untitled. Azteca xanthochroa Roger subsp. salti Wheeler, subsp. nov.]. Pp. 114-115 in: Wheeler, W. M., Darlington, P. J. Ant-tree notes from Rio Frio, Colombia. Psyche (Camb.) 37:107-117. PDF]]


 * [[Media:WheelerW1930f.pdf|Wheeler, W. M. 1930j. Philippine ants of the genus Aenictus with descriptions of the females of two species. J. N. Y. Entomol. Soc. 38: 193-212 PDF]]


 * [[Media:WheelerW1930-31.pdf|Wheeler, W. M. 1930k. A list of the known Chinese ants. Peking Nat. Hist. Bull. 5: 53-81 PDF]]


 * Wheeler, W. M. 1931a. Neotropical ants of the genus Xenomyrmex Forel. Rev. Entomol. (Rio J.) 1: 129-139


 * [[Media:Wheeler 1931b.pdf|Wheeler, W. M. 1931b. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bull. Mus. Comp. Zool. 72: 1-34 PDF]]


 * [[Media:Wheeler 1931c.pdf|Wheeler, W. M. 1931c. Concerning some ant gynandromorphs. Psyche (Cambridge) 38:80-85. [1931-09] PDF]]


 * [[Media:WheelerW1931.pdf|Wheeler, W. M. 1931d. The ant Camponotus (Myrmepomis) sericeiventris Guérin and its mimic. Psyche (Camb.) 38: 86-98 PDF]]


 * [[Media:WheelerW1932e.pdf|Wheeler, W. M. 1932a. A list of the ants of Florida with descriptions of new forms. J. N. Y. Entomol. Soc. 40: 1-17 PDF]]


 * [[Media:WheelerW1932.pdf|Wheeler, W. M. 1932b. Some attractions of the field study of ants. The Scientific Monthly 34:397-402. [1932-05] PDF]]


 * [[Media:WheelerW1932d.pdf|Wheeler, W. M. 1932d. An Australian Leptanilla. Psyche (Camb.) 39: 53-58 PDF]]


 * [[Media:WheelerW1932f.pdf|Wheeler, W. M. 1932e. Ants of the Marquesas Islands. Bull. Bernice P. Bishop Mus. 98: 155-163 PDF]]


 * [[Media:WheelerW1932c.pdf|Wheeler, W. M. 1932g. Ants from the Society Islands. Bull. Bernice P. Bishop Mus. 113: 13-19 PDF]]


 * [[Media:WheelerW1933e.pdf|Wheeler, W. M. 1933a. The Templeton Crocker Expedition of the California Academy of Sciences, 1932. No. 6. Formicidae of the Templeton Crocker Expedition. Proc. Calif. Acad. Sci. (4) 21: 57-64 PDF]]


 * [[Media:WheelerW1933a.pdf|Wheeler, W. M. 1933b. Mermis parasitism in some Australian and Mexican ants. Psyche (Camb.) 40: 20-31 PDF]]


 * [[Media:WheelerW1933b.pdf|Wheeler, W. M. 1933c. New ants from China and Japan. Psyche (Camb.) 40: 65-67 PDF]]


 * [[Media:WheelerW1933c.pdf|Wheeler, W. M. 1933d. A second parasitic Crematogaster. Psyche (Camb.) 40: 83-86 PDF]]


 * [[Media:WheelerW1933d.pdf|Wheeler, W. M. 1933e. A new Myrmoteras from Java. Proc. N. Engl. Zool. Club 13: 73-75 PDF]]


 * [[Media:Wheeler 1933f.pdf|Wheeler, W. M. 1933f. A new species of Ponera and other records of ants from the Marquesas Islands. Bull. Bernice P. Bishop Mus. 114: 141-144 PDF]]


 * [[Media:WheelerW1933G.pdf|Wheeler, W. M. 1933g. Three obscure genera of ponerine ants. Am. Mus. Novit. 672: 1-23 PDF]]


 * [[Media:WheelerW1933I.pdf|Wheeler, W. M. 1933i. Colony founding among ants, with an account of some primitive Australian species. Cambridge, Mass.: Harvard University Press, viii + 179 pp. PDF]]


 * [[Media:WheelerW1934d.pdf|Wheeler, W. M. 1934a. Formicidae of the Templeton Crocker Expedition, 1933. Proc. Calif. Acad. Sci. (4) 21: 173-181 PDF]]


 * [[Media:WheelerW1934a.pdf|Wheeler, W. M. 1934b. An Australian ant of the genus Leptothorax Mayr. Psyche (Camb.) 41: 60-62 PDF]]


 * [[Media:WheelerW1934b.pdf|Wheeler, W. M. 1934c. A second revision of the ants of the genus Leptomyrmex Mayr. Bull. Mus. Comp. Zool. 77: 69-118 PDF]]


 * [[Media:WheelerW1934c.pdf|Wheeler, W. M. 1934d. Contributions to the fauna of Rottnest Island, Western Australia. No. IX. The ants. J. R. Soc. West. Aust. 20: 137-163 PDF]]


 * [[Media:WheelerW1934.pdf|Wheeler, W. M. 1934e. Some aberrant species of Camponotus (Colobopsis) from the Fiji Islands. Ann. Entomol. Soc. Am. 27: 415-424 PDF]]


 * [[Media:WheelerW1934e.pdf|Wheeler, W. M. 1934f. Ants from the islands off the west coast of Lower California and Mexico. Pan-Pac. Entomol. 10: 132-144 PDF]]


 * [[Media:WheelerW1934f.pdf|Wheeler, W. M. 1934g. Neotropical ants collected by Dr. Elisabeth Skwarra and others. Bull. Mus. Comp. Zool. 77: 157-240 PDF]]


 * [[Media:Wheeler 1934h.pdf|Wheeler, W. M. 1934h. Some ants from the Bahama Islands. Psyche (Camb.) 41: 230-232 PDF]]


 * [[Media:WheelerW1935d.pdf|Wheeler, W. M. 1935a. Two new genera of myrmicine ants from Papua and the Philippines. Proc. N. Engl. Zool. Club 15: 1-9 PDF]]


 * [[Media:WheelerW1935e.pdf|Wheeler, W. M. 1935b. New ants from the Philippines. Psyche (Camb.) 42: 38-52 PDF]]


 * [[Media:Wheeler 1935c.pdf|Wheeler, W. M. 1935c. Myrmecological notes. Psyche (Camb.) 42: 68-72 PDF]]


 * [[Media:Wheeler 1935d.pdf|Wheeler, W. M. 1935d. Ants of the genera Belonopelta Mayr and Simopelta Mann. Rev. Entomol. (Rio J.) 5: 8-19 PDF]]


 * [[Media:WheelerW1935b.pdf|Wheeler, W. M. 1935e. The Australian ant genus Mayriella Forel. Psyche (Camb.) 42: 151-160 PDF]]


 * [[Media:WheelerW1935f.pdf|Wheeler, W. M. 1935f. Ants of the genus Acropyga Roger, with description of a new species. J. N. Y. Entomol. Soc. 43: 321-329 PDF]]


 * [[Media:WheelerW1935c.pdf|Wheeler, W. M. 1935g. Check list of the ants of Oceania. Occas. Pap. Bernice P. Bishop Mus. 11(1 11: 1-56 PDF]]


 * [[Media:WheelerW1936a.pdf|Wheeler, W. M. 1936a. The Australian ant genus Froggattella. Am. Mus. Novit. 842: 1-11 PDF]]


 * [[Media:WheelerW1936b.pdf|Wheeler, W. M. 1936b. Binary anterior ocelli in ants. Biological Bulletin (Woods Hole) 70:185-192. [1936-04] PDF]]


 * [[Media:WheelerW1936.pdf|Wheeler, W. M. 1936c. Ants from Hispaniola and Mona Island. Bull. Mus. Comp. Zool. 80: 195-211 PDF]]


 * [[Media:Wheeler 1936.pdf|Wheeler, W. M. 1936d. A singular Crematogaster from Guatemala. Psyche (Camb.) 43: 40-48 PDF]]


 * Wheeler, W. M. 1936e. Ecological relations of ponerine and other ants to termites. Proc. Am. Acad. Arts Sci. 71: 159-243


 * [[Media:WheelerW1936e.pdf|Wheeler, W. M. 1936f. Notes on some aberrant Indonesian ants of the subfamily Formicinae. Tijdschr. Entomol. 79: 217-221 PDF]]


 * [[Media:WheelerW1936c.pdf|Wheeler, W. M. 1936g. Ants from the Society, Austral, Tuamotu and Mangareva Islands. Occas. Pap. Bernice P. Bishop Mus. 12(1 18: 3-17 PDF]]


 * Wheeler, W. M. 1937a. Additions to the ant-fauna of Krakatau and Verlaten Island. Treubia 16: 21-24


 * [[Media:WheelerW1937b.pdf|Wheeler, W. M. 1937b. Ants mostly from the mountains of Cuba. Bull. Mus. Comp. Zool. 81: 439-465 PDF]]


 * [[Media:WheelerW1937.pdf|Wheeler, W. M. 1937c. Mosaics and other anomalies among ants. Cambridge, Mass.: Harvard University Press, 95 pp. PDF]]


 * [[Media:WheelerW1938.pdf|Wheeler, W. M. 1938. Ants from the caves of Yucatan. Pp. 251-255 in: Pearse, A. S. Fauna of the caves of Yucatan. Carnegie Inst. Wash. Publ. 491:1-304. PDF]]


 * Wheeler, W. M. 1942. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90: 1-262


 * [[Media:WheelerW1920b.pdf|Wheeler, W. M.; Bailey, I. W. 1920. The feeding habits of pseudomyrmine and other ants. Trans. Am. Philos. Soc. (2) 22: 235-279 PDF]]


 * [[Media:WheelerW1929g.pdf|Wheeler, W. M.; Bequaert, J. C. 1929. Amazonian myrmecophytes and their ants. Zool. Anz. 82: 10-39 PDF]]


 * [[Media:Wheeler Chapman 1922.pdf|Wheeler, W. M.; Chapman, J. W. 1922. The mating of Diacamma. Psyche (Cambridge) 29:203-211. [1922-12] PDF]]


 * [[Media:WheelerW1925b.pdf|Wheeler, W. M.; Chapman, J. W. 1925. The ants of the Philippine Islands. Part I, Dorylinae and Ponerinae. Philipp. J. Sci. 28: 47-73 PDF]]


 * [[Media:WheelerW1934g.pdf|Wheeler, W. M.; Creighton, W. S. 1934. A study of the ant genera Novomessor and Veromessor. Proc. Am. Acad. Arts Sci. 69: 341-387 PDF]]


 * [[Media:WheelerW1930g.pdf|Wheeler, W. M.; Darlington, P. J., Jr. 1930. Ant-tree notes from Rio Frio, Colombia. Psyche (Camb.) 37: 107-117 PDF]]


 * [[Media:Wheeler & Gaige 1920.pdf|Wheeler, W. M.; Gaige, F. M. 1920. Euponera gilva (Roger), a rare North American ant. Psyche (Camb.) 27: 69-72 PDF]]


 * [[Media:Wheeler & Long 1901.pdf|Wheeler, W. M.; Long, W. H. 1901. The males of some Texan Ecitons. Am. Nat. 35: 157-173 PDF]]


 * [[Media:WheelerW1914.pdf|Wheeler, W. M.; Mann, W. M. 1914. The ants of Haiti. Bull. Am. Mus. Nat. Hist. 33: 1-61 PDF]]


 * [[Media:Wheeler & Mann 1916.pdf|Wheeler, W. M.; Mann, W. M. 1916. The ants of the Phillips Expedition to Palestine during 1914. Bull. Mus. Comp. Zool. 60: 167-174 PDF]]


 * Wheeler, W. M.; Mann, W. M. 1942a. [Untitled. Pseudomyrma picta Stitz var. heterogyna Wheeler and Mann, var. nov.]. Pp. 172-173 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942b. [Untitled. Pseudomyrma triplarina (Weddell) var. rurrenabaquensis Wheeler & Mann, var. nov.]. Pp. 188-189 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942c. [Untitled. Allomerus decemarticulatus Mayr subsp. novemarticulatus Wheeler & Mann, subsp. nov.]. P. 199 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942d. [Untitled. Allomerus decemarticulatus Mayr subsp. octoarticulatus Mayr var. exsanguis Wheeler and Mann, var. nov.]. P. 200 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942e. [Untitled. Allomerus decemarticulatus Mayr subsp. octoarticulatus Mayr var. angulatus Wheeler & Mann, var. nov.]. P. 201 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942f. [Untitled. Allomerus decemarticulatus Mayr subsp. octoarticulatus Mayr var. melanoticus Wheeler & Mann, var. nov.]. Pp. 202-203 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942g. [Untitled. Azteca brevicornis Mayr var. boliviana Wheeler & Mann, var. nov.]. P. 225 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942h. [Untitled. Azteca ulei Forel var. gagatina Wheeler and Mann var. nov.]. P. 246 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * Wheeler, W. M.; Mann, W. M. 1942i. [Untitled. Myrmelachista (Decamera) schumanni Emery var. cordincola Wheeler & Mann, var. nov.]. P. 255 in: Wheeler, W. M. Studies of Neotropical ant-plants and their ants. Bull. Mus. Comp. Zool. 90:1-262.


 * [[Media:WheelerW1902.pdf|Wheeler, W. M.; McClendon, J. F. 1903. Dimorphic queens in an American ant (Lasius latipes Walsh). Biol. Bull. (Woods Hole) 4: 149-163 PDF]]