Cardiocondyla shuckardi

The most common and widespread endemic Cardiocondyla species in the Afrotropical region.

Identification
Bolton (1982) - C. shuckardi is recognised by its dimensions and extremely reduced propodeal armament. Other species in the region with reduced propodeal armament include Cardiocondyla monardi, Cardiocondyla sekhemka and Cardiocondyla nilotica. In the first of these the metanotal groove is absent and the pedicel segments are very elongate. C. sekhemka is a much smaller species with shorter scapes and a broader head, and nilotica has longer scapes than shuckardi and a narrower postpetiole.

Seifert (2003) - A member of the Cardiocondyla shuckardi group. Cardiocondyla shuckardi is treated here as a polymorphic Malagasy and SE African species. The occurrence of transitions between the different sculpture and pubescence types observed in the type specimens of C. shuckardi and Cardiocondyla shuckardoides and the high overall similarity of these taxa suggest conspecifity.

Distribution based on Regional Taxon Lists
Afrotropical Region: Botswana, Cameroun, Gabon, Ghana, Kenya, Mozambique, Namibia, Nigeria, Saudi Arabia, South Africa, Uganda, United Arab Emirates, Zimbabwe. Malagasy Region: Madagascar. Palaearctic Region: Kuwait.

Nomenclature

 * . Cardiocondyla shuckardi Forel, 1891b: 161 (w.) MADAGASCAR.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Madagascar: Imerina, Antananarivo (Camboué).
 * Type-depository: MHNG.
 * Subspecies of nuda: Forel, 1895c: 251; Emery, 1922e: 126.
 * Status as species: Dalla Torre, 1893: 71; Wheeler, W.M. 1922a: 1021; Santschi, 1932a: 387; Bolton, 1982: 316 (redescription); Collingwood, 1985: 257; Bolton, 1995b: 133; Collingwood & Agosti, 1996: 327; Rigato, 2002: 172 (in key); Seifert, 2003a: 257 (redescription); Collingwood, et al. 2011: 424; Borowiec, L. 2014: 48.
 * Senior synonym of brevispinosa Weber: Bolton, 1982: 316; Bolton, 1995b: 133.
 * Senior synonym of fusca: Bolton, 1982: 316; Bolton, 1995b: 133.
 * Senior synonym of sculptior: Bolton, 1982: 316; Bolton, 1995b: 133.
 * Senior synonym of shuckardoides: Seifert, 2003a: 257.
 * Senior synonym of wassmanni: Bolton, 1982: 316; Bolton, 1995b: 133.
 * Distribution: Botswana, Cameroon, Democratic Republic of Congo, Gabon, Ghana, Iran, Kuwait, Madagascar, Mozambique, Namibia, Nigeria, Saudi Arabia, South Africa, Uganda, United Arab Emirates, Yemen, Zimbabwe.
 * Current subspecies: nominal plus sculptinodis.
 * brevispinosa. Cardiocondyla brevispinosa Weber, 1952a: 6 (w.) DEMOCRATIC REPUBLIC OF CONGO.
 * Type-material: holotype worker.
 * Type-locality: Democratic Republic of Congo (“Belgian Congo”): Beni, 0°24’N, 29°24’E, 24.ii.1948, no. 2116 (N.A. Weber).
 * Type-depository: MCZC.
 * [Unresolved junior secondary homonym of Pheidole brevispinosa Donisthorpe, 1948d: 593 (Bolton, 1995b: 132).]
 * Junior synonym of shuckardi: Bolton, 1982: 316; Bolton, 1995b: 132.
 * fusca. Cardiocondyla fusca Weber, 1952a: 7 (w.) UGANDA.
 * Type-material: holotype worker.
 * Type-locality: Uganda: Jinja, edge of Ripon Falls, 15.viii.1939, no. 1495 (N.A. Weber).
 * Type-depository: MCZC.
 * Junior synonym of shuckardi: Bolton, 1982: 316; Bolton, 1995b: 132.
 * sculptior. Cardiocondyla wasmanni var. sculptior Santschi, 1926b: 241 (w.) GABON.
 * Type-material: holotype worker.
 * Type-locality: Gabon: Samkita (F. Faure).
 * Type-depository: NHMB.
 * As unavailable (infrasubspecific) name: Santschi, 1932a: 387.
 * Junior synonym of shuckardi: Bolton, 1982: 316; Bolton, 1995b: 133.
 * shuckardoides. Cardiocondyla nuda var. shuckardoides Forel, 1895c: 250 (w.) MADAGASCAR.
 * Type-material: syntype workers (number not stated).
 * Type-locality: Madagascar: Antananarivo (Sikora).
 * Type-depository: MHNG.
 * Subspecies of nuda: Wheeler, W.M. 1922a: 1021; Emery, 1922e: 126; Bolton, 1995b: 133.
 * Junior synonym of shuckardi: Seifert, 2003a: 257.
 * wasmanni. Cardiocondyla wassmanni Santschi, 1926b: 241 (w.) CAMEROON.
 * Type-material: holotype worker.
 * Type-locality: Cameroon: Gr. Batanga (Schwab).
 * Type-depository: NHMB.
 * [Justified emendation of spelling to wasmanni: Santschi, 1932a: 387.]
 * Subspecies of shuckardi: Santschi, 1932a: 387.
 * Junior synonym of shuckardi: Bolton, 1982: 316; Bolton, 1995b: 133.

Worker
Bolton (1982) - TL 2.0-2.6, HL 0.50-0.60, HW 0.38-0.46, CI 75-79, SL 0.36-0.45, SI 93-100, PW 0.27-0.35, AL 0.54-0.69 (35 measured).

Antennal scapes when laid back on the head in full-face view either just failing to reach or just reaching the occipital corners, never distinctly surpassing them; the scapes moderately long, SI > 90. Maximum diameter of eye 0.11-0.14, about 0.26-0.30 x HW and with 9-12 ommatidia in the longest row. Head always obviously longer than broad, CI < 80 in material examined. Pronotal corners in dorsal view broadly and evenly rounded. With the alitrunk in profile the promesonotal dorsum forming an even shallow convexity from front to back, sloping evenly into the metanotal groove. Metanotal groove impressed but the depth of the impression varying between samples. To some extent the apparent variation in depth is caused by the convexity of the propodeum behind the groove as in some cases it rises more steeply and is more convex than in others. Propodeal dorsum behind the convex portion sloping downwards posteriorly to the junction with the declivity. Propodeal armament very reduced, at best represented only by a pair of minute triangular denticles which may be acute or blunted, or by a pair of tubercles, or merely bluntly angular; never with developed teeth or spines. In dorsal view the propodeal armament scarcely visible, the length of each component constituting only a fraction of the distance separating their bases. Petiole node in dorsal view subglobular, usually broader than long but in some only about as broad as long. Postpetiole distinctly broader than long. In profile the petiole and postpetiole as in Fig. 1, the petiolar dorsum convex and somewhat variable in length. Sculpture of dorsal head and alitrunk usually of fine, very dense blanketing shagreening or granulation, but this may be reduced on tile alitrunk or even on the head, though less frequently on the latter than on the former. In extreme cases the dorsal alitrunk may be almost smooth. Hairs absent except on mouthparts and gastral apex. Colour varying from medium brown to blackish brown, sometimes black.

Type Material
Seifert (2003) - Syntype workers labelled by Forel “C. shuckardi Forel male, Imcrina Madagascar (Camboue)” and with a printed red label “Type”, Geneve. 1 syntype worker labelled by Forel “C. shuckardi Forel d Type Antananarivo (Camboue)”, MHN Geneve.

References based on Global Ant Biodiversity Informatics

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