Checklist of Cardiocondyla species

The following species and subspecies belong to the genus Cardiocondyla. Synonyms are listed under their senior names. For valid names only see Cardiocondyla species and for distribution information see Cardiocondyla species by Country.

A
allonivalis Seifert, 2022 (New Guinea)

argentea Seifert, 2022 (Indonesia)

argyrotricha Seifert, 2022 (Malaysia, Philippines)

atalanta Forel, 1915 (Australia, Lord Howe Island, New Guinea, Norfolk Island)

B
batesii Forel, 1894 (Algeria, Balearic Islands, Iberian Peninsula, Morocco, Portugal, Spain, Tunisia)

bicoronata Seifert, 2003 (Israel, Jordan, United Arab Emirates, Yemen)

brachyceps Seifert, 2003 (Afghanistan, Armenia, Azerbaijan, Georgia, Iran, Turkey)

britteni Crawley, 1920 (United Kingdom of Great Britain and Northern Ireland)

bulgarica Forel, 1892 (Bulgaria, Greece, North Macedonia, Turkey)

• Cardiocondyla elegans eleonorae Forel, 1911

C
carbonaria Forel, 1907 (India)

caspiense Seifert, 2023 (Iran)

compressa Seifert, 2017 (Australia)

cristata (Santschi, 1912) (Madagascar)

D
dalmatica Soudek, 1925 (Albania, Austria, Bosnia and Herzegovina, Croatia, Cyprus, Greece, Hungary, Iran, Italy, Montenegro, North Macedonia, Serbia, Slovakia, Slovenia, Turkey)

dalmaticoides Seifert, 2023 (Turkey)

E
elegans Emery, 1869 (France, Iberian Peninsula, Italy, Nepal, Portugal, Spain)

• Cardiocondyla elegans santschii Forel, 1905

• Cardiocondyla provincialis Bernard, 1956

• Xenometra gallica Bernard, 1957

emeryi Forel, 1881 (Angola, Anguilla, Aruba, Australia, Bahamas, Barbados, Benin, Bermuda, Borneo, Botswana, Brazil, British Virgin Islands, Cameroun, Cameroun, Canary Islands, Comoros, Cook Islands, Costa Rica, Cuba, Dominican Republic, Ecuador, Egypt, Fiji, French Polynesia, Galapagos Islands, Gambia, Ghana, Greater Antilles, Grenada, Guadeloupe, Haiti, Hawaii, Honduras, Indonesia, Iran, Israel, Ivory Coast, Kenya, Lesser Antilles, Madagascar, Malaysia, Mali, Mauritius, Mexico, Mozambique, Nepal, Netherlands Antilles, New Caledonia, New Guinea, Nigeria, Niue, Norfolk Island, Oman, Puerto Rico, Rwanda, Saint Helena, Samoa, Saudi Arabia, Seychelles, Socotra Archipelago, South Africa, Spain, Sri Lanka, Sudan, Switzerland, Thailand, Tonga, Turkey, Turks and Caicos Islands, Uganda, United Arab Emirates, United Republic of Tanzania, United States, Vietnam, Wallis and Futuna Islands, Yemen, Zimbabwe)

• Cardiocondyla emeryi mahdii Karavaiev, 1911

• Cardiocondyla emeryi rasalamae Forel, 1891

• Cardiocondyla mauritia Donisthorpe, 1946

• Cardiocondyla nuda nereis Wheeler, W.M., 1927

• Xenometra monilicornis Emery, 1917

emeryi fezzanensis Bernard, 1948 (Algeria)

excavata Seifert, 2022 (New Guinea)

F
fajumensis Forel, 1913 (Egypt, Iran, Sudan, Yemen, Zimbabwe)

• Cardiocondyla emeryi schatzmayri Finzi, 1936

• Cardiocondyla nilotica Weber, 1952

G
gallagheri Collingwood & Agosti, 1996 (Oman, United Arab Emirates)

gallilaeica Seifert, 2003 (Israel)

gibbosa Kuznetsov-Ugamsky, 1927 (China, Kazakhstan, Kyrgyzstan)

goa Seifert, 2003 (India)

goroka Seifert, 2022 (New Guinea)

H
humilis (Smith, F., 1858)

I
insutura Zhou, 2001 (China)

israelica Seifert, 2003 (Egypt, Israel)

itsukii Seifert, Okita & Heinze, 2017 (Bhutan, China, Hawaii, India, Indonesia, Japan, Kiribati, Nepal, Philippines, Réunion, Sri Lanka, Thailand)

J
jacquemini Bernard, 1953 (Algeria)

K
kagutsuchi Terayama, 1999 (Borneo, China, Guam, Indonesia, Japan, Malaysia, Micronesia (Federated States of), Nepal, New Guinea, Northern Mariana Islands, Palau, Philippines, Singapore, Solomon Islands, Sri Lanka, Thailand, Vietnam)

kazanensis Terayama, 2013 (Japan)

koshewnikovi Ruzsky, 1902 (Afghanistan, China, Kazakhstan, Kyrgyzstan, Mongolia, Russian Federation, Turkmenistan, Uzbekistan)

kushanica Pisarski, 1967 (Afghanistan, Iran, Turkmenistan)

L
latifrons Seifert, 2022 (Indonesia, Malaysia, Philippines, Singapore, Thailand)

littoralis Seifert, 2003 (Kazakhstan)

longiceps Seifert, 2003 (Socotra Archipelago, Yemen)

longinoda Rigato, 2002 (United Republic of Tanzania)

luciae Rigato, 2002 (United Republic of Tanzania)

M
mauritanica Forel, 1890 (Afghanistan, Canary Islands, Egypt, Greece, Grenada, Iberian Peninsula, India, Indonesia, Iran, Iraq, Israel, Jordan, Libya, Malta, Morocco, Nepal, Netherlands Antilles, Oman, Pakistan, Portugal, Puerto Rico, Russian Federation, Saudi Arabia, Socotra Archipelago, Tunisia, Turkey, Ukraine, United Arab Emirates, United States)

• Cardiocondyla ectopia Snelling, R.R., 1974

• Cardiocondyla emeryi nitida Bernard, 1948

• Leptothorax caparica Henin, Paiva & Collingwood, 2001

melana Seifert, 2003 (Saudi Arabia, Yemen)

micropila Seifert, 2022 (Brunei Darussalam, Malaysia, Singapore, Thailand)

minutior Forel, 1899 (Australia, Barbados, Costa Rica, Dominican Republic, Ecuador, Galapagos Islands, Greater Antilles, Grenada, Guam, Guatemala, Hawaii, Honduras, India, Indonesia, Japan, Marshall Islands, Mexico, Micronesia (Federated States of), Nepal, New Guinea, New Zealand, Northern Mariana Islands, Oman, Socotra Archipelago, Sri Lanka, United States, Vietnam, Yemen)

• Cardiocondyla breviscapa Seifert, 2003

• Cardiocondyla tsukuyomi Terayama, 1999

monardi Santschi, 1930 (Angola)

N
nana Seifert, 2003 (Borneo, Indonesia, Malaysia)

neferka Bolton, 1982 (Benin, Cameroun, Ghana, Ivory Coast)

nigra Forel, 1905 (Algeria, Egypt, Iberian Peninsula, Iran, Israel, Malta, Morocco, Portugal, Spain, Tunisia, Turkey, Yemen)

• Cardiocondyla elegans torretassoi Finzi, 1936

nigrocerea Karavaiev, 1935 (Indonesia)

nivalis Mann, 1919 (Solomon Islands)

nuda (Mayr, 1866) (Australia, Fiji, Micronesia (Federated States of), New Guinea, Samoa, Solomon Islands, Tonga, Vanuatu, Wallis and Futuna Islands)

O
obscurior Wheeler, W.M., 1929 (American Samoa, Barbados, Bermuda, Brazil, Canary Islands, Comoros, Costa Rica, Fiji, Grenada, Guadeloupe, Guam, Israel, Mexico, Micronesia (Federated States of), Nepal, Netherlands Antilles, New Guinea, Northern Mariana Islands, Singapore, Taiwan, United States)

• Cardiocondyla bicolor Donisthorpe, 1930

opaca Seifert, 2003 (India)

opistopsis Seifert, 2003 (Kuwait)

P
papuana (Reiskind, 1965) (New Guinea)

paradoxa Emery, 1897 (New Guinea)

• Pheidole brevispinosa Donisthorpe, 1948

paranuda Seifert, 2003 (Australia)

parvinoda Forel, 1902 (India, Philippines, Thailand)

persiana Seifert, 2003 (Georgia, Iran, Israel)

pirata Seifert & Frohschammer, 2013 (Philippines)

R
rolandi Seifert, 2023 (China, Kazakhstan, Mongolia)

rugulosa Seifert, 2003 (Yemen)

S
sahlbergi Forel, 1913 (Afghanistan, Armenia, Azerbaijan, Georgia, Iran, Israel, Kazakhstan, Kyrgyzstan, Russian Federation, Tunisia, Turkey, Turkmenistan, Uzbekistan)

schulzi Seifert, 2022 (India)

sekhemka Bolton, 1982 (Ghana)

semiargentea Seifert, 2022 (Indonesia)

semirubra Seifert, 2003 (Turkey)

shagrinata Seifert, 2003 (India)

shuckardi Forel, 1891 (Botswana, Cameroun, Gabon, Ghana, Kenya, Kuwait, Madagascar, Mozambique, Namibia, Nigeria, Rwanda, Saudi Arabia, South Africa, Uganda, United Arab Emirates, Zimbabwe)

• Cardiocondyla brevispinosa Weber, 1952

• Cardiocondyla fusca Weber, 1952

• Cardiocondyla nuda shuckardoides Forel, 1895

• Cardiocondyla wasmanni sculptior Santschi, 1926

• Cardiocondyla wassmanni Santschi, 1926

shuckardi sculptinodis Santschi, 1913 (Madagascar)

sima Wheeler, W.M., 1935 (Malaysia, Philippines, Thailand)

stambuloffii Forel, 1892 (Armenia, Azerbaijan, Bulgaria, China, Georgia, Greece, Iran, Republic of Moldova, Romania, Russian Federation, Turkey, Ukraine)

• Cardiocondyla bogdanovi Ruzsky, 1905

• Cardiocondyla montandoni Santschi, 1912

• Cardiocondyla stambuloffi taurica Karavaiev, 1927

strigifrons Viehmeyer, 1922 (Indonesia, Japan, Malaysia, New Guinea, Philippines, Singapore)

subspina Seifert, 2022 (Indonesia)

sulcata Seifert, 2022 (Malaysia)

T
tenuifrons Seifert, 2003 (Jordan)

thoracica (Smith, F., 1859) (Australia, Indonesia, New Guinea)

tibetana Seifert, 2003 (China, Tibet)

tiwarii Ghosh, Sheela & Kundu, 2005 (India)

tjibodana Karavaiev, 1935 (Belize, Borneo, Guam, Indonesia, Kiribati, Krakatau Islands, Malaysia, Marshall Islands, Micronesia (Federated States of), New Guinea, Northern Mariana Islands, Palau, Philippines, Singapore)

U
ulianini Emery, 1889 (Afghanistan, Azerbaijan, China, Iran, Kazakhstan, Kyrgyzstan, Russian Federation, Saudi Arabia, Turkey, Ukraine)

• Cardiocondyla elegans schkaffi Arnoldi, 1933

unicalis Seifert, 2003 (Iran)

V
venustula Wheeler, W.M., 1908 (Barbados, Cuba, Dominican Republic, Greater Antilles, Haiti, Hawaii, Honduras, Mexico, Mozambique, Namibia, Puerto Rico, United States, Zimbabwe)

• Cardiocondyla badonei Arnold, 1926

• Cardiocondyla globinodis Stitz, 1923

verdensis Seifert, 2023 (Cape Verde)

W
weserka Bolton, 1982 (Cameroun)

wheeleri Viehmeyer, 1914 (New Guinea)

wroughtonii (Forel, 1890) (Australia, Borneo, Cambodia, China, Cuba, Guadeloupe, Hawaii, India, Indonesia, Israel, Japan, Kenya, Laos, Malaysia, Martinique, Mayotte, Mexico, Nepal, New Caledonia, New Guinea, Oman, Philippines, Saudi Arabia, Singapore, Somalia, Sri Lanka, Taiwan, Thailand, United Republic of Tanzania, United States, Vietnam)

• Cardiocondyla emeryi chlorotica Menozzi, 1930

• Cardiocondyla longispina Karavaiev, 1935

• Cardiocondyla wroughtoni bimaculata Wheeler, W.M., 1929

• Cardiocondyla wroughtonii hawaiensis Forel, 1899

• Cardiocondyla wroughtonii quadraticeps Forel, 1912

• Cardiocondyla yamauchii Terayama, 1999

Y
yemeni Collingwood & Agosti, 1996 (Oman, Yemen)

yoruba Rigato, 2002 (Nigeria)

Z
zoserka Bolton, 1982 (Nigeria)