Oecophylla smaragdina

Aggressive arboreal ants that use larval silk to weave together leaves to form their nesting cavities. A mature colony of Oecophylla smaragdina can entirely dominate a tree (sometimes several) with nests distributed throughout their heavily defended arboreal territory.

Distribution
Wetterer (2017) - The vast majority of O. smaragdina records come from areas with Tropical climates according to the Köppen-Geiger system: rainforest, monsoon, and savanna. However, >250 records come from areas classified on the map as having Subtropical climates, mostly in the Himalayan foothills of India and Nepal, southern China, northern Vietnam, and the southern coast of Queensland, Australia. Almost all these sites are classified as dry winter subtropical climate. A few O. smaragdina sites are classified as having Arid climates, all from warm semi-arid areas.

Distribution based on Regional Taxon Lists
Australasian Region: Australia. Indo-Australian Region: Borneo, Indonesia, Krakatau Islands, Malaysia, New Guinea, Philippines, Singapore. Oriental Region: Bangladesh, Cambodia, India, Laos, Nepal, Sri Lanka, Thailand. Palaearctic Region: China.

Biology
There is a webpage with a list of some recent publications about weaver ants. You can also read an overview of their biology from the a chapter in The Ants: The Weaver Ants (Hölldobler and Wilson 1990).

Peeters and Andersen (1989) - A few aggregations of dealate queens were collected in coastal regions of Northern Territory, Australia. Newly mated queens can cooperate to found new colonies, but only one survives in established colonies of ''0. smaragdina''.

Pinkalski et al. (2015) - Colonies nesting in mango trees (Mangifera indica) in Darwin, Australia were found to deposit significant amounts of nitrogen on their host trees via their waste. This deposition increased when the ants were provided access to additional sucrose resources.



Oecophylla smaragdina, together with Anoplolepis gracilipes and Dolichoderus thoracicus, is one of the most common ant species which tends honeydew-producing hemipterans in Indonesia. Fanani et al. (2020) examined the influence of these species on the introduced parasitoid Anagyrus lopezi, a species used to control the invasive cassava mealybug Phenacoccus manihoti (Hemiptera: Pseudococcidae). They found that when ants were absent the average time spent foraging by individual parasitoids was significantly longer (27.39 minutes) compared to when ants were present (2.47- 4.68 minutes). As a result, parasitoids spent less time in finding hosts and a longer time in handling hosts. This resulted in more oviposition activities and a 2-3 fold increase in parasitism and the number of wasps that emerged from their hosts.

Ethnoentomology
Bhotwate & Kumar (2020) report that selected rural and tribal groups within India mash live workers with salt, red chillies and mustard oil, and eat them with rice. They are reported to prevent gastritis and provide nutritive value.

Nomenclature

 *  smaragdina. Formica smaragdina Fabricius, 1775: 828 (q.) INDIA. Jerdon, 1851: 121 (w.m.); Wheeler, G.C. & Wheeler, J. 1953e: 176 (l.); Crozier, 1970: 115 (k.). Combination in Oecophylla: Smith, F. 1860b: 102. Senior synonym of viridis: Smith, F. 1857a: 53; Taylor & Brown, D.R. 1985: 127; of macra, zonata: Roger, 1863b: 10; Dalla Torre, 1893: 176; of virescens: Mayr, 1872: 143; Taylor & Brown, D.R. 1985: 127. Current subspecies: nominal plus fuscoides, gracilior, gracillima, selebensis, subnitida.
 * virescens. Formica virescens Fabricius, 1775: 392 (w.) AUSTRALIA. Smith, F. 1858b: 30 (m.). Combination in Oecophylla: Smith, F. 1860b: 102. Status as species: Dalla Torre, 1893: 177; Emery, 1921c: 102. Subspecies of smaragdina: Emery, 1887a: 242; Forel, 1915b: 95; Wheeler, W.M. 1922a: 228 (in key); Emery, 1925b: 52; Karavaiev, 1933a: 315. Junior synonym of smaragdina: Mayr, 1872: 143; Taylor & Brown, D.R. 1985: 127.
 * viridis. Formica viridis Kirby, W. 1819: 478 (w.) AUSTRALIA. Junior synonym of virescens: Roger, 1863b: 10; Dalla Torre, 1893: 177; Emery, 1925b: 52; of smaragdina: Smith, F. 1857a: 53; Taylor & Brown, D.R. 1985: 127.
 * macra. Formica macra Guérin-Méneville, 1831, pl. 8, fig. 1 (w.) "Offack". Junior synonym of virescens: Smith, F. 1858b: 29; of smaragdina: Roger, 1863b: 10; Dalla Torre, 1893: 176; Arnold, 1922: 609.
 * zonata. Formica zonata Guérin-Méneville, 1838: 205 (q.) "Port Praslin". Junior synonym of smaragdina: Roger, 1863b: 10; Dalla Torre, 1893: 176.

References based on Global Ant Biodiversity Informatics

 * Abdullah N. A., and R. Hashim. 2010. An inventory of ants (Hymenoptera: Formicidae) at Melawi Sandy Beach, Bachok, Kelantan. Malaysian Journal of Science 29: 67-72.
 * Abe T., S. Yamane, and K. Onoyama. Ants collected on the Krakatau Islands 100 years after the great eruptions. Biogeography 14: 65-75.
 * Alcantara M. J., S. Modi, T. C. Ling, J. Monkai, H. Xu, S. Huang, and A. Nakamura. 2019. Differences in geographic distribution of ant species (Hymenoptera: Formicidae) between forests and rubber plantations: a case study in Xishuangbanna, China, and a global meta-analysis. Myrmecological News 29: 135-145.
 * Amarasinghe H. E. 2010. Species composition and nesting habits of ants in a hill-country home garden in Sri Lanka. Asian Myrmecology 3: 9-20.
 * Andersen A. N. 1992. The rainforest ant fauna of the northern Kimberley region of western Australia (Hymenoptera: Formicidae). Journal of Australian Entomological Society 31: 187-192.
 * Andersen A. N. 1993. Ants as indicators of restoration success at a uranium mine in tropical Australia. Restoration Ecology 1(3): 156-167.
 * Andersen A. N., J. C. Z. Woinarski, and B. Hoffman. 2004. Biogeography of the ant fauna of the Tiwi Islands, in northern Australia's moonsoonal tropics. Australian Journal of Zoology 52: 97-110.
 * Andersen A. N., J. Lanoue, and I. Radford. 2010. The ant fauna of the remote Mitchell Falls area of tropical north-western Australia: Biogeography, environmental relationships and conservation significance. Journal of Insect Conservation 14: 647-661.
 * Andersen A. N., M. Houadria, M. Berman, and M. van der Geest. Rainforest ants of the Tiwi Islands: a remarkable centr of endemism in Australia's monsoonal tropics. Insectes Sociaux 59: 433-441.
 * Andersen A. N., and B. D. Hoffmann. 2010. Conservation value of low fire frequency in tropical savannas: ants in monsoonal northern Australia. Austral Ecology 36(5): 597-503.
 * Andersen A. N., and H. Reichel. 1994. The ant (Hymenoptera: Formicidae) fauna of Holmes Jungle, a rainforest patch in the seasonal tropics of Australia's Northern Territory. Aust. Ent. Soc. 33. 153-158.
 * Andersen, Alan N., John C.Z. Woinarski and Ben D. Hoffman. 2004. Biogeography of the ant fauna of the Tiwi Islands, in northern Australia's monsoonal tropics. Australian Journal of Zoology 52: 97-110.
 * Asfiya W., R. Ubaidillah, and Sk. Yamane. 2008. Ants (Hymenoptera: Formicidae) of the Krakataus, and Sebesi and Sebuku islands. Treubia 36: 1-9.
 * Azuma N., K. Ogata, T. Kikuchi, and S. Higashi. 2006. Phylogeography of Asian weaver ants, Oecophylla smaragdina. Ecol. Res. 21: 126-136.
 * Basu P. 1994. Ecology of ground foraging ants in a tropical evergreen forest in Western Ghats, India. PhD Thesis, School of ecology and environmental sciences, Pondichery University, India. 155 pages.
 * Basu P., N. Tak, and A. K. Sanyal. 2013. Ants (insecta: Hymenoptera: Formicidae) of Bethuadahari wildlife sanctuary, Nadia, West Bengal, India. Rec. zool, Surv. India: 113(4): 17-22.
 * Bharti H., Y. P. Sharma, M. Bharti, and M. Pfeiffer. 2013. Ant species richness, endemicity and functional groups, along an elevational gradient in the Himalayas. Asian Myrmecology 5: 79-101.
 * Bharti H., Y. P. Sharma, and A. Kaur. 2009. Seasonal patterns of ants (Hymenoptera: Formicidae) in Punjab Shivalik. Halteres 1(1): 36-47.
 * Bharti H., and I. Kaur. 2011. SEM studies on immature stages of weaver ant Oecophylla smaragdina (Fabricius, 1775) (Hymenoptera: Formicidae) from India. Halteres 3: 16-25.
 * Bhoje P. M., K. Shilpa, and T. V. Sathe. 2014. Diversity of ants (Hymenoptera: Formicidae) from Kolhapur district of Maharashtra, India. Uttar Pradesh J. Zool. 34(1): 23-25.
 * Bluthgen N., G. Gebauer, and K. Fiedler. 2003. Disentangling a rainforest food web using stable isotopes: dietary diversity in a species-rich ant community. Oecologia 137: 426-435.
 * Borowiec L., and S. Salata. 2019. Next step in the invasion: Trichomyrmex mayri (Forel, 1902) new to the Philippines (Hymenoptera: Formicidae). Annals of the Upper Silesian Museum in Bytom Entomology 28(3): 1-3.
 * Brown E.S. 1959. Immature nutfall of coconuts in the Solomon Islands. II. Changes in ant populations, and their relation to vegetation. Bulletin of Entomological Research 50: 523-558.
 * CSIRO Collection
 * Chapman, J. W., and Capco, S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monogr. Inst. Sci. Technol. Manila 1: 1-327
 * Chen Z. L., S. Y. Zhou, D. D. Ye, Y. Chen, and C. W. Lu. 2013. Moleular phylogeny of the ant subfamily Formicinae (Hymenoptera, Formicidae) from China based on Mitochondrial genes. Sociobiology 60(2): 135-144.
 * Chhotani O. B., and P. K. Maiti. 1977. Contribution to the knowledge of Formicidae of the Andaman Islands. Zoological Survey of India 3(1): 17-20.
 * Cole A. C., Jr., and J. W. Jones Jr. 1948. A study of the weaver ant, Oecophylla smaragdina (Fab.). American Midland Naturalist 39: 641-651.
 * Collingwood C. A. 1962. Some ants (Hym. Formicidae) from north-east Asia. Entomologisk Tidskrift 83: 215-230.
 * Collingwood C.A. 1970. Formicidae (Hymenopter: Aculeata) of Nepal. Himalaya Khumbu Himal, 3: 371-388.
 * D'Cunha P., and V. M. Grover Nair. 2014. Ant fauna on the mangroves of Dakshina Kannada and Udupi districts, Karnataka, India. Journal of Ent. Res. 38(1): 59-66.
 * D'Cunha P., and V. Mala Grover Nair. 2013. Diversity and Distribution of Ant Fauna in Hejamadi Kodi Sandspit, Udupi District, Karnataka, India. Halteres 4: 33-47.
 * Dad J. M., S. A. Akbar, H. Bharti, and A. A. Wachkoo. 2019. Community structure and ant species diversity across select sites ofWestern Ghats, India. Acta Ecologica Sinica 39: 219–228.
 * Dahl F. 1901. Das Leben der Ameisen im Bismarck-Archipel, nach eigenen Beobachtungen vergleichend dargestellt. Mitt. Zool. Mus. Berl. 2: 1-70.
 * Davidson D. W., S. C. Cook, R. R. Snelling and T. H. Chua. 2003. Explaining the Abundance of Ants in Lowland Tropical Rainforest Canopies. Science 300: 969-972.
 * Davidson, D.W., J.-P. Lessard, C.R. Bernau and S.C. Cook. 2007. The Tropical Ant Mosaic in a Primary Bornean Rain Forest. Biotropica 39(4):468-475
 * Dhote J. 2012. Ant (Hymenoptera: Formicidae) records from Shri Shivaji science college campus, Amravati, India. Applied Research and Development Institute Journal 3(12): 114-117.
 * Dias R. K. S. 2002. Current knowledge on ants of Sri Lanka. ANeT Newsletter 4: 17- 21.
 * Dias R. K. S. 2006. Current taxonomic status of ants (Hymenoptera: Formicidae) in Sri Lanka. The Fauna of Sri Lanka: 43-52. Bambaradeniya, C.N.B. (Editor), 2006. Fauna of Sri Lanka: Status of Taxonomy, Research and Conservation. The World Conservation Union, Colombo, Sri Lanka & Government of Sri Lanka. viii + 308pp.
 * Dias R. K. S., K. R. K. A. Kosgamage, and H. A. W. S. Peiris. 2012. The Taxonomy and Conservation Status of Ants (Order: Hymenoptera, Family: Formicidae) in Sri Lanka. In: The National Red List 2012 of Sri Lanka; Conservation Status of the Fauna and Flora. Weerakoon, D.K. & S. Wijesundara Eds., Ministry of Environment, Colombo, Sri Lanka. p11-19.
 * Dias R. K. S., and K. R. K. A. Kosgamage. Systematics and community composition of foraging workers ants (Family: Formicidae) collected from three habitats in a dry zone region of Sri Lanka. Proceedings of the Annual Research Symposium 2008. Faculty of Graduate Studies, University of Kelaniya.
 * Dias R. K. S., and K. R. K. Anuradha Kosgamage. 2012. Occurrence and species diversity of ground-dwelling worker ants (Family: Formicidae) in selected lands in the dry zone of Sri Lanka. J. Sci. Univ. Kelaniya 7: 55-72.
 * Dias R. K. S., and R. P. K. C. Rajapaksa. 2016. Geographic records of subfamilies, genera and species of ants (Hymenoptera: Formicidae) in the four climatic zones of Sri Lanka: a review. J. Sci. Univ. Kelaniya 11(2): 23-45.
 * Dias, R.K.S. 2006. Overview of ant research in Sri Lanka: 2000-2004. ANeT Newsletter 8:7-10
 * Donisthorpe H. 1942. Ants from the Colombo Museum Expedition to Southern India, September-October 1938. Annals and Magazine of Natural History (11)9: 449-461.
 * Donisthorpe, Horace. 1943. The Ants of Waigeu Island, North Dutch New Guinea. The Annals and Magazine of Natural History 11 (10): 433-475.
 * Emery C. 1887. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia. [part]. Ann. Mus. Civ. Stor. Nat. 24(4): 209-258.
 * Emery C. 1888. Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza . Formiche raccolte dal sig. Elio Modigliani in Sumatra e nell'isola Nias. Annali del Museo Civico di Storia Naturale 25: 528-534.
 * Emery C. 1893. Formicides de l'Archipel Malais. Revue Suisse de Zoologie 1: 187-229.
 * Emery C. 1893. Voyage de M. E. Simon à l'île de Ceylan (janvier-février 1892). Formicides. Annales de la Société Entomologique de France 62: 239-258.
 * Emery C. 1901. Ameisen gesammelt in Ceylon von Dr. W. Horn 1899. Deutsche Entomologische Zeitschrift 1901: 113-122.
 * Emery C. 1911. Formicidae. Résultats de l'expédition scientifique néerlandaise à la Nouvelle-Guinée en 1907 et 1909 sous les auspices de Dr. H. A. Lorentz. Nova Guin. 9: 249-259
 * Emery C. Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei. Annali del Museo Civico di Storia Naturale 40: 661-722.
 * Emery, C. "Catalogo delle formiche esistenti nelle collezioni del Museo Civico di Genova. Parte terza. Formiche della regione Indo-Malese e dell'Australia." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 4, no. 24 (1887): 209-258.
 * Emery, C. "Formiche raccolte da Elio Modigliani in Sumatra, Engano e Mentawei." Annali del Museo Civico di Storia Naturale Giacomo Doria (Genova) (2) 20, no. 40 (1900): 661-722.
 * Emery, C. "Formicides recoltes a Buitenzorg (Java), par M. Massart." Annales de la Société Entomologique de Belgique 40 (1896): 245-249.
 * Emery, C. "Voyage de MM. Bedot et Pictet dans l'Archipel Malais. Formicides de l'Archipel Malais." Revue Suisse de Zoologie 1 (1893): 187-229.
 * Fontanilla A. M., A. Nakamura, Z. Xu, M. Cao, R. L. Kitching, Y. Tang, and C. J. Burwell. 2019. Taxonomic and functional ant diversity along tropical, subtropical, and subalpine elevational transects in southwest China. Insects 10, 128; doi:10.3390/insects10050128
 * Forel A. 1885. Indian ants of the Indian Museum, Calcutta. J. Asiat. Soc. Bengal Part II Nat. Sci. 54: 176-182.
 * Forel A. 1886. Études myrmécologiques en 1886. Annales de la Société Entomologique de Belgique. 30: 131-215.
 * Forel A. 1886. Indian ants of the Indian Museum, Calcutta, No. 2. J. Asiat. Soc. Bengal Part II Nat. Sci. 55: 239-249.
 * Forel A. 1886. Indian ants of the Indian Museum, Calcutta, No. 2. Journal of the Asiatic Society of Bengal. Part II. Natural Science 55: 239-249.
 * Forel A. 1893. Nouvelles fourmis d'Australie et des Canaries. Ann. Soc. Entomol. Belg. 37: 454-466.
 * Forel A. 1894. Les Formicides de l'Empire des Indes et de Ceylan. Part IV. J. Bombay Nat. Hist. Soc. 8: 396-420.
 * Forel A. 1901. Formiciden aus dem Bismarck-Archipel, auf Grundlage des von Prof. Dr. F. Dahl gesammelten Materials. Mitt. Zool. Mus. Berl. 2: 4-37.
 * Forel A. 1903. Les fourmis des îles Andamans et Nicobares. Rapports de cette faune avec ses voisines. Rev. Suisse Zool. 11: 399-411.
 * Forel A. 1904. Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Ezheg. Zool. Muz. 8: 368-388.
 * Forel A. 1905. Ameisen aus Java. Gesammelt von Prof. Karl Kraepelin 1904. Mitt. Naturhist. Mus. Hambg. 22: 1-26.
 * Forel A. 1907. Ameisen von Madagaskar, den Comoren und Ostafrika. Wissenschaftliche Ergebnisse. Reise in Ostafrika 2: 75-92.
 * Forel A. 1907. Formiciden aus dem Naturhistorischen Museum in Hamburg. II. Teil. Neueingänge seit 1900. Mitt. Naturhist. Mus. Hambg. 24: 1-20.
 * Forel A. 1909. Ameisen aus Java und Krakatau beobachtet und gesammelt von Herrn Edward Jacobson. Notes Leyden Mus. 31: 221-232.
 * Forel A. 1911. Fourmis nouvelles ou intéressantes. Bull. Soc. Vaudoise Sci. Nat. 47: 331-400.
 * Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien ausgeführt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. II. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel-Reepen in den Jahren 1911-1912. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 36:1-148.
 * Forel A. 1915. Results of Dr. E. Mjöbergs Swedish Scientific Expeditions to Australia 1910-13. 2. Ameisen. Ark. Zool. 9(16): 1-119
 * Forel, A. 1908. Fourmis de Ceylan et d'Égypte récoltées par le Prof. E. Bugnion. Lasius carniolicus. Fourmis de Kerguelen. Pseudandrie? Strongylognathus testaceus. Bull. Soc. Vaudoise Sci. Nat. 44: 1-22
 * General D. M., and G. D. Alpert. 2012. A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. Zookeys 200: 1-111.
 * Ghosh S. N., S. Sheela, B. G. Kundu, S. Roychowdhury, and R. N. Tiwari. 2006. Insecta: Hymenoptera: Formicidae. Pp. 369-398 in: Alfred, J. R. B. (ed.) 2006. Fauna of Arunachal Pradesh. (Part -2). [State Fauna Series 13.]. New Delhi: Zoological Survey of India, iv + 518 pp.
 * Ghosh S. N., S. Sheela, and B. G. Kundu. 2005. Ants (Hymenoptera: Formicidae) of Rabindra Sarovar, Kolkata. Records of the Zoological Survey of India. Occasional Paper 234: 1-40.
 * Greenslade P.J.M. 1970. Interspecific competition and frequency changes among ants in Solomon islands coconut Plantations. Journal of Applied Ecology 8(2): 323-352.
 * Greenslade P.J.M. 1971. Interspecific Competition and Frequency Changes Among Ants in Solomon Islands Coconut Plantations. Journal of Applied Ecology 8(2): 323-352
 * Greenslade P.J.M. 1971. Phenology of Three Ant Species In the Solomon Islands. Australian Journal of Entomology 10(4): 241252
 * Greenslade P.J.M. and Greenslade Penelope. 1977. Some Effects of Vegetation Cover and Disturbance on a Tropical Ant Fauna. Insectes Sociaux 24(2): 163-182
 * Greenslade P.J.M. and P. Greenslade. 1977. Some effects of vegetation cover and disturbance on a tropical ant fauna. Insectes Sociaux 24(2): 163-182.
 * Gumawardene, N.R., J.D. Majer and J.P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2:71-83
 * Gunawardene N. R., J. D. Majer, and J. P. Edirisinghe. 2008. Diversity and richness of ant species in a lowland wet forest reserve in Sri Lanka. Asian Myrmecology 2: 71-83.
 * Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
 * Hannan M.A. 2008. New ant records fro Bangladesh. Asian Myrmecology 2: 95-98.
 * Hashimoto Y., S. Yamane, and T. Itioka. 1997. A preliminary study on dietary habits of ants in a Bornean rain forest. Japanese Journal of Entomology 65(4): 688-695.
 * Hatter S. J. S., Sen, M. Nibedita, R. Mathew, and S. Sharma. 2004. Faunal diversity of Saipung wild life sanctuaryl Narpuh Reserve forest, laintia Hills, Meghalaya, Conservation Area Series 21 : 1-66.
 * Helms J. A., S. M. Helms, N. I. Fawzi, Tarjudin, F. Xaverius. 2017. Ant community of an Acacia mangium forest in Indonesian Borneo. Serangga 22(1): 147-159.
 * Herwina H., N. Nasir, Jumjunidang, and Yaherwandi. 2013. The composition of ant species on banana plants with Banana Bunchy-top Virus (BBTV) symptoms in West Sumatra, Indonesia. Asian Myrmecology 5: 151-161.
 * Herwina H., and K. Nakamura. 2007. Ant species diversity study using pitfall traps in a small yard in Bogor Botanic garden, West Java, Indonesia. Treubia 35: 99-116.
 * IZIKO South Africa Museum Collection
 * Imai H. T., C. Baroni Urbani, M. Kubota, G. P. Sharma, M. H. Narasimhanna, B. C. Das, A. K. Sharma, A. Sharma, G. B. Deodikar, V. G. Vaidya, and M. R. Rajasekarasetty. 1984. Karyological survey of Indian ants. Japanese Journal of Genetics 59: 1-32.
 * Ito, F.; Yamane, S.; Eguchi, K.; Noerdjito, W. A.; Kahono, S.; Tsuji, K.; Ohkawara, K.; Yamauchi, K.; Nishida, T.; Nakamura, K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10:379-404.
 * Jaitrong W., B. Guenard, E. P. Economo, N. Buddhakala, and S. Yamane. 2016. A checklist of known ant species of Laos (Hymenoptera: Formicidae). Asian Myrmecology 8: 1-32. DOI: 10.20362/am.008019
 * Jaitrong W., and T. Ting-Nga. 2005. Ant fauna of Peninsular Botanical Garden (Khao Chong), Trang Province, Southern Thailand (Hymenoptera: Formicidae). The Thailand Natural History Museum Journal 1(2): 137-147.
 * Jaitrong W.; Nabhitabhata, J. 2005. A list of known ant species of Thailand. The Thailand Natural History Museum Journal 1(1): 9-54.
 * Janda M., G. D. Alpert, M. L. Borowiec, E. P. Economo, P. Klimes, E. Sarnat, and S. O. Shattuck. 2011. Cheklist of ants described and recorded from New Guinea and associated islands. Available on http://www.newguineants.org/. Accessed on 24th Feb. 2011.
 * Janda M., and M. Konecna. 2011. Canopy assembalges of ants in a New Guinea rain forest. Journal of Tropical Ecology 27: 83-91.
 * Kaleeswaran B., S. Ezil, B. P. Ganesh, and S. Bhavatarini. 2008. Biodiversity and niches of Ants in Alagar hills, Tamil nadu Wildlife Biodiversity Conservation Published by Day publishing house pp 188-208.
 * Kami K.S., and S. E. Miller. 1998. Samoan insects and related arthropods: checklist and bibliography. Bishop Museum Technical Report 13, pp 121.
 * Kami KS & Miller SE. 1998. Samoan insects and related arthropods: checklist and bibliography. Bishop Museum Technical Report No. 13.
 * Karavaiev V. 1933. Ameisen aus dem Indo-Australischen Gebiet, VII. Konowia 11: 305-320.
 * Karavaiev V. 1935. Neue Ameisen aus dem Indo-Australischen Gebiet, nebst Revision einiger Formen. Treubia 15: 57-118.
 * Karmaly K. A.; S. Sumesh, T. P. Rabeesh, and L. Kishore. 2010. A checklist of ants of Thirunelli in Wayanad, Kerala. J. of the Bombay Natural History Society 107(1): 64-67.
 * Katayama M., K. Kishimoto-Yamada, H. O. Tanaka, T. Endo, Y. Hashimoto, Sk. Yamane, and T. Itioka. 2015. Negative correlation between ant and spider abundances in the canopy of a Bornean tropical rain forest. Biotropica (in press).
 * Khoo Y.H. 1990. A note on the Formicidae (Hymenoptera) from pitfall traps at Ulu Kinchin, Pahang, Malaysia. Malayan Nature Journal 43: 290-293.
 * Klimes P., M. Janda, S. Ibalim, J. Kua, and V. Novotny. 2011. Experimental suppression of ants foraging on rainforest vegetation in New Guinea: testing methods for a whole-forest manipulation of insect communities. Ecological Entomology 36: 94-103.
 * Klimes P., P. Fibich, C. Idigel, and M. Rimandai. 2015. Disentangling the diversity of arboreal ant communities in tropical forest trees. PLoS ONE 10(2): e0117853. doi:10.1371/journal.pone.0117853
 * Kosgamage K. R. K. A., and R. K. S. Dias 2008. Systematics and community composition of Foraging worker ants ((Order: Hymenoptera, Family: Formicidae) collected from three habitats on a dry zone region of Sri Lanka.Proceedings of Postgraduate Symposium of Kelaniya University. 115pp.
 * Kumar D., and M. Archana. 2008. Ant community variation in urban and agricultural ecosystems in Vadodara District (Gujarat State), western India. Asian Myrmecology 2: 85-93.
 * Kutter H. 1932. Ameisen aus dem Museum zu Dresden. Mitt. Schweiz. Entomol. Ges. 15: 207-210.
 * Kutter H. 1933. Einige Ameisen von der Südküste von Neu-Britannien. Mitt. Schweiz. Entomol. Ges. 15: 471-474.
 * Latumahina F., M. Borovanska, N. S. Putra, and M. Janda. 2015. Ants of Ambon Island  diversity survey and checklist. ZooKeys 472: 4357.
 * Lopwichan S., and S. Khachonpisitsak. 2015. Ant diversity in Nong Tha Yu Arboretum, Si Racha District, Chon Buri Province. Proceedings The 7 th National Science Research Conference. 30-31 March 2015. Naresuan University.
 * Lu Z., K. Li, N. Zhang, and Y. Chen. 2017. Diversity and indicator species of leaf-litter ants in Eucalyptus grandis plantations and secondary natural forests. Forest Research 29(4): 576-580
 * Lu Z., and Y. Chen. 2016. Effects of habitat on ant functional groups: a case study of Luchun County, Yunnan Province, China. Chinese Journal of Eco-Agriculture 24(5): 801-810.
 * Lucky A., E. Sarnat, and L. Alonso. 2011. Ants of the Muller Range, Papua New Guinea, Chapter 10. In Richards, S. J. and Gamui, B. G. (editors). 2013. Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: surveying the biodiversity of Papua New Guineas sublime karst environments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA.
 * Lucky A., K. Sagata, and E. Sarnat. 2011. Ants of the Nakanai Mountains, East New Britain Province, Papua New Guinea, Chapter 1. In Richards, S. J. and Gamui, B. G. (editors). 2013. Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: surveying the biodiversity of Papua New Guineas sublime karst environments. RAP Bulletin of Biological Assessment 60. Conservation International. Arlington, VA.
 * Majer J. D., R. L. Kitching, B. E. Heterick, K. Hurley, and K. E. C. Brennan. 2001. North-south patterns within arboreal ant assembalages from rain forests in Eastern Australia. Biotropica 33(4): 643-661.
 * Mann William. 1916. The Ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology at Harvard College 63(7): 273-391
 * Mann, W.M. 1919. The ants of the British Solomon Islands. Bulletin of the Museum of Comparative Zoology of Harvard College 63: 273-391
 * Mathew R. 2003. On Formicidae (Insecta: Hymenoptera) of Nongkhyllem Wild Life Sanctuary, Ri-Bhoi District, Meghalaya. Records of the Zoological Survey of India 101:195-207.
 * Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
 * Menozzi, C. 1932. Formiche dell'Isola di Nias. Misc. Zool. Sumatr. 65: 1-13
 * Mezger D., and M. Pfeiffer. 2011. Influence of the arrival of Anoplolepis gracilipes (Hymenoptera: Formicidae) on the composition of an ant community in a clearing in Gunung Mulu National Park, Sarawak, Malaysia. Asian Myrmecology 4: 89-98.
 * Mohanraj P., M. Ali, and K. Veerakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay of Bengal). Journal of Insect Science 10: Article 172
 * Mohanraj, P., M. Ali and K. Veenakumari. 2010. Formicidae of the Andaman and Nicobar Islands (Indian Ocean: Bay Of Bengal). Journal of Insect Science 10:172.
 * Mukerjee D. 1930. Report on a collection of ants in the Indian Museum, Calcutta. J. Bombay Nat. Hist. Soc. 34: 149-163.
 * Mukherji D., and S. Ribeiro. 1925. On a collection of ants (Formicidae) from the Andaman Islands. Records of the Indian Museum 27: 205-209.
 * Mustafa N.A., H.M.W. Salim, C. Fletcher, A.R. Kassim, and M.D. Potts. 2011. Taxonomic and functional diversity of ants (Hymenoptera: Formicidae) in an upper hill dipterocarp forest in Peninsular Malaysia. The Raffles bulletin of Zoology 59(2): 181-194.
 * Musthak Ali T. M. 1982. Ant fauna (Hymenoptera: Formicidae) of Bangalore with observations on their nesting and foraging habits. Thesis Abstracts. Haryana Agricultural University 8: 370-371.
 * Narendra A., H. Gibb, and T. M. Ali. 2011. Structure of ant assemblages in Western Ghats, India: role of habitat, disturbance and introduced species. Insect Conservation and diversity 4(2): 132-141.
 * Nelson X.J., R. R. Jackson, S.D. Pollard, G.B. Edwards, A.T. Barrion. 2004. Predation by ants on jumping spiders (Araneae: Salticidae) in the Philippines. New Zealand Journal of Zoology 31: 45-56.
 * Offenberg J., N. Thi Thu Cuc, and D. Wiwatwitaya. 2013. The effectiveness of weaver ant (Oecophylla smaragdina) biocontrol in Southeast Asian citrus and mango. Asian Myrmecology 5: 139-149.
 * Offenberg, J. 2007. The distribution of weaver ant pheremones on host trees. Insectes Sociaux 54:248-250
 * Offenberg, J., M.G. Nielsen, D.J. MacIntosh, S. Havanon and S. Aksornkoae. 2004. Evidence That Insect Herbivores Are Deterred by Ant Pheromones. Proceedings: Biological Sciences 271(6) :S433-S435
 * Offenberg, J., M.G. Nielsen, D.J. Macintosh,S. Aksornkoae and S. Havanon. 2006. Weaver Ants Increase Premature Loss of Leaves Used for Nest Construction in Rhizophora Trees. Biotropica 38(6):782-785.
 * Ogata K. 2005. Asian ant inventory and international networks. Report on Insect inventory Project in Tropic Asia TAIIV: 145-170.
 * Orivel J., P. Klimes, V. Novotny, and M. Leponce. 2018. Resource use and food preferences in understory ant communities along a complete elevational gradient in Papua New Guinea. Biotropica 10.1111/btp.12539
 * Overbeck, H. (1924). A List of Ants from Singapore and Neighbouring Places. Journal of the Malayan Branch of the Royal Asiatic Society 2 (1):25-40.
 * Pajni H. R., and R. K. Suri. 1978. First report on the Formicid fauna (Hymenoptera) of Chandigarh. Res. Bull. (Science) Punjab University 29: 5-12.
 * Parui A. K., S. Chatterjee, and P. Basu. 2015. Habitat characteristics shaping ant species assemblages in a mixed deciduous forest in Eastern India. Journal of Tropical Ecology: 1-14. doi:10.1017/S0266467415000036
 * Peng R., M. G. Nielsen, J. Offenberg, and D. Birkmose. 2013. Utilisation of multiple queens and pupae transplantation to boost early colony growth of weaver ants Oecophylla smaragdina. Asian Myrmecology 5: 177-184.
 * Pfeiffer M., D. Mezger, and J. Dyckmans. 2013. Trophic ecology of tropical leaf litter ants (Hymenoptera: Formicidae) - a stable isotope study in four types of Bornean rain forest. Myrmecological News 19: 31-41.
 * Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
 * Pfeiffer, M., H. Cheng Tuck, and T. Chong Lay. 2008. Exploring arboreal ant community composition and co-ccurrence patterns in plantations of oil palm Elaeis guineensis in Borneo and Peninsular Malaysia. Ecography 31(1): 21-32.
 * Philpott S.M., P. Bichier, R.A. Rice, and R. Greenberg. 2008. Biodiversity conservation, yield, and alternative products in coffee agroecosystems in Sumatra, Indonesia. Biodivers. Conserv. 17: 1805-1820. Data obtained from Stacy Philpott
 * Raci N., C. Sravanthy, C. Sammaiah, and M. Thirupahaiah. 2015. Biodiversity of ants (Insecta-Hymenoptera) in agroecosystem and grass land in Jammikunta, Karimnagar District, Telangana, India. Journal ofEnvironment 4(1): 11-16.
 * Rajagopal T., S. P. Sevarkodiyone, and A. Manimozhi. 2005. Ant diversity in some selected localities of Sattur Taluk, Virudhunagar district, Tamil Nadu. Zoos' Print Journal 20(6): 1887-1888.
 * Rajan P. D., M. Zacharias, and T. M. Mustak Ali. 2006. Insecta: Hymenoptera: Formicidae. Fauna of Biligiri Rangaswamy Temple Wildlife Sanctuary (Karnataka). Conservation Area Series, Zool. Surv. India.i-iv,27: 153-188.
 * Ran H., and S. Y. Zhou. 2012. Checklist of chinese ants: formicomorph subfamilies (Hymenoptera: Formicidae) II. Journal of Guangxi Normal University: Natural Science Edition 30(4): 81-91.
 * Rechinger, K. 1907. Botanische und zoologische Ergebnisse einer wissenschaftlichen Forschungreise nach den Samoa-Inseln, dem Neuguinea-Archipel und den Salomons-Inseln von März bis Dezember 1905. I. Teil. Wien: A. Hölder, 121 pp.
 * Rickson F.R., and M.M. Rickson. 1998. The cashew nut, Anacardium occidentale (Anacardiaceae), and its perennial association with ants: extrafloral nectary location and the potential for ant defense. American Journal of Botany 85(6): 835-849.
 * Rizali A., A. Rahim, B. Sahari, L.B. Prasetyo, and D. Buchori. 2011. Impact of invasive ant species in shaping ant community structure on small islands in Indonesia. Jurnal Biologi Indonesia 7(2): 221-230.
 * Rizali A., D. J. Lohman, D. Buchori, L. Budi Prasetyo, H. Triwidodo, M. M. Bos, S. Yamane, and C. H. Schulze. 2009. Ant communities on small tropical islands: effects of island size and isolation are obscured by habitat disturbance and tramp ant species. Journal of Biogeography 37(2): 229-236.
 * Rizali A., M. M. Bos, D. Buchori, Sk. Yamane, and C. H. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 15(2): 77-84.
 * Rizali A., M.M. Bos, D. Buchori, Sk. Yamane, C. Hans, and J. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 77-84.
 * Rizali A., Y. Clough, D. Buchori, M. L . A. Hosang, M. M. Bos, and T. Tscharntke. 2012. Long-term change of ant community structure in cacao agroforestry landscapes in Indonesia. Insect Conservation and Diversity doi: 10.1111/j.1752-4598.2012.00219.x
 * Robson Simon Ant Collection, 05-Sept-2014
 * Room P. M. 1975. Diversity and organization of the ground foraging ant faunas of forest, grassland and tree crops in Papua Nez Guinea. Aust. J. Zool. 23: 71-89.
 * Room, P.M. 1975. Relative Distributions of Ant Species in Cocoa Plantations in Papua New Guinea Relative Distributions of Ant Species in Cocoa Plantations in Papua New Guinea. Journal of Applied Ecology 12(1):47-61
 * Rubiana R., A. Rizali, L. H. Denmead, W. Alamsari, P. Hidaya, Pudjianto, D. Hindayana, Y. Clough, T. Tscharntke, and D. Buchori. 2015. Agricultural land use alters species composition but not species richness of ant communities. Asian Myrmecology 7,.
 * Santschi F. 1919. Cinq notes myrmécologiques. Bulletin de la Société Vaudoise des Sciences Naturelles 52: 325-350.
 * Santschi F. 1928. Formicidae (Fourmis). Insects Samoa. 5: 41-58.
 * Santschi F. 1928. Fourmis de Sumatra, récoltées par Mr. J. B. Corporaal. Tijdschrift voor Entomologie 71: 119-140.
 * Santschi F. 1933. Formicides des collections de S. A. R. le Prince Léopold de Belgique. Voyage aux Indes orientales, 1932. Bulletin du Musée Royal d'Histoire Naturelle de Belgique. 9(27): 1-3.
 * Schmidt F. A., C. R. Ribas, T. G. Sobrinho, R. Ubaidillah, J. H. Schereder, Y. Clough, and T. Tscharntke. 2017. Similar alpha and beta diversity changes in tropical ant communities, comparing savannas and rainforests in Brazil and Indonesia. Oecologia DOI 10.1007/s00442-017-3960-y
 * Sheela S. 2008. Handbook of Hymenoptera, Formicidae. Zoological Survey of India, 56 pages
 * Sheikh A. H., M. Manzoor, Y. A. Rather, and T. Jobiraj. 2019. Taxonomic study of ant (Formicidae : Hymenoptera) fauna of Dumna Nature Park, Jabalpur, Madhya Pradesh, India. Journal of Entomological Research 43(2): 203-212.
 * Sinu P. A., M. Nasser, and P. D. Rajan. 2006. Feeding fauna and foraging habits of tiger beetles found in agro-ecosystems in Western Ghats, India. Biotropica 38(4): 500-507.
 * Sitthicharoenchai D., and N. Chantarasawat. 2006. Ant species diversity in the establishing area for Advanced Technology Institute at Lai-Nan Sub-district, Wiang Sa district, Nan Province, Thailand. The Natural History Journal of Chulalongkorn University 6(2): 67-74
 * Smith F. 1863. Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the islands of Mysol, Ceram, Waigiou, Bouru and Timor. Journal and Proceedings of the Linnean Society of London. Zoology 7: 6-48.
 * Smith F. 1865. Descriptions of new species of hymenopterous insects from the islands of Sumatra, Sula, Gilolo, Salwatty, and New Guinea, collected by Mr. A. R. Wallace. Journal and Proceedings of the Linnean Society of London. Zoology 8: 61-94.
 * Smith, F. "Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace." Journal of the Proceedings of the Linnean Society of London, Zoology 2 (1857): 42-88.
 * Smith, Fr. "Catalogue of hymenopterous insects collected by Mr. A. R. Wallace in the Islands of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea." Journal of the Proceedings of the Linnean Society of London, Zoology 5 (1860): 93-143.
 * Snelling R. R. 1998. Insect Part 1: The social Hymenoptera. In Mack A. L. (Ed.) A Biological Assessment of the Lakekamu Basin, Papua New Guinea, RAP 9. 189 ppages
 * Snelling R. R. 2000. Ants of the Wapoga river area, Irian Jaya, Indonesia. In Mack, Andrew L. and Leeanne E. Alonso (eds.). 2000. A Biological Assessment of the Wapoga River Area of Northwestern Irian Jaya, Indonesia. RAP Bulletin of Biological Assessment 14, Conservation International, Washington, DC.
 * Song Y., Z. Xu, C. Li, N. Zhang, L. Zhang, H. Jiang, and F. Mo. 2013. An Analysis on the Ant Fauna of the Nangun river Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
 * Sonune B. V., and R. J. Chavan. 2016. Distribution and diversity of ants (Hymenoptera: Formicidae) around Gautala Autramghat Sanctuary, Aurangabad Maharashtra, India. Journal of Entomology and Zoology Studies 4(2): 361-364.
 * Stitz H. 1911. Australische Ameisen. (Neu-Guinea und Salomons-Inseln, Festland, Neu-Seeland). Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1911: 351-381.
 * Stitz H. 1912. Ameisen aus Ceram und Neu-Guinea. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1912: 498-514.
 * Sukimin S., M. Mohamed, and H. Aris. 2010. Ant diversity of Maliau Basin Conservation Area, Sabah, Malaysia. Journal of Tropical Biology and Conservation 6:89-101.
 * Sureh P. V., V. V. Sudheendrakumar, C. F. Binoy, G. Mathew, and T. C. Narendran. 1999. The macro Hymenopteran fauna of Parambikulam wildlife Sanctuary. Zoos' Print Journal 14(4): 1-2.
 * Tak N. 2000. Studies on ants (Formicidae) of Rajasthan - III. Banswara. Entomon 25:97-101.
 * Tak N. 2008. Ants of Rajasthan. Conserving Biodiversity of Rajasthan Zool. Surv. India. 149-155.
 * Tak N. 2009. Ants Formicidae of Rajasthan. Records of the Zoological Survey of India, Occasional Paper No. 288, iv, 46 p
 * Tak N., and N. S. Rathore. 2004. Insecta: Hymenoptera: Formicidae. State Fauna Series 8: Fauna of Gujarat. Zool. Surv. India. Pp. 161-183.
 * Tak N., and S. I. Kazmi. 2013. On some ants (Insecta: Hymenoptera: Formicidae) from Nagaland, India. Rec. zool. Surv. India: 113(1): 169-182.
 * Tak N., and S. L. Kazmi. 2011. On a collection of Insecta: Hymenoptera: Formicidae from Uttarakhand. Rec. zool. Surv. India : 111(2) : 39-49.
 * Tang J., Li S., Huang E., Zhang B. and Chen Y. 1995. Hymenoptera: Formicidae (1). Economic Insect Fauna of China 47: 1-133.
 * Thapa V. K. 2000. An Inventory of Nepal's Insects, Vol. III. IUCN Nepal, Kathmandu, xi + 475 pp.
 * Tiwari R. N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4): 1-96.
 * Tiwari R. N., B. G. Kundu, S. Roy Chowdhury, and S. N. Ghosh. 2003. Insecta: Hymenoptera: Formicidae. Fauna of Sikkim. Part 4. State Fauna Series. 9.Zool.Surv.India. i-iii, 1-512. Chapter pagination: 467-506.
 * Tiwari R.N., B.G. Kundu, S. Roychowdhury, S.N. Ghosh. 1999. Insecta: Hymenoptera: Formicidae. Pp. 211-294 in: Director; Zoological Survey of India (ed.) 1999. Fauna of West Bengal. Part 8. Insecta (Trichoptera, Thysanoptera, Neuroptera, Hymenoptera and Anoplura). Calcutta: Zoological Survey of India, iv + 442 pp.
 * Tiwari R.N., B.G. Kundu, S. Sheela, and S. N. Ghosh. 2004. Insecta: Hymenoptera: Formicidae. Alfred, J.R.B. [Ed.], Fauna of Manipur, part - 2 (insects). State fauna series 10. Zool.Surv.India.i-v, 1-625. Chapter pagination: 605-625.
 * Tiwari, R.N. 1999. Taxonomic studies on ants of southern India (Insecta: Hymenoptera: Formicidae). Memoirs of the Zoological Survey of India 18(4):1-96
 * Trainor C.R. and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
 * Trainor C.R., and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
 * Varghese T. 2004. Taxonomic studies on ant genera of the Indian Institute of Science campus with notes on their nesting habits. Pp. 485-502 in : Rajmohana, K.; Sudheer, K.; Girish Kumar, P.; Santhosh, S. (eds.) 2004. Perspectives on biosystematics and biodiversity. Prof. T.C. Narendran commemoration volume. Kerala: Systematic Entomology Research Scholars Association, xxii + 666 pp.
 * Viehmeyer H. 1912. Ameisen aus Deutsch Neuguinea gesammelt von Dr. O. Schlaginhaufen. Nebst einem Verzeichnisse der papuanischen Arten. Abhandlungen und Berichte des Königlichen Zoologischen und Anthropologische-Ethnographischen Museums zu Dresden 14: 1-26.
 * Viehmeyer H. 1914. Ameisen aus Perak, Bali und Ceram (Hym.) (Freiburger Molukken-Expedition), gesammelt von E. Streesemann. Entomologische Mitteilungen. Berlin-Dahlem 3: 112-116.
 * Wang C. and Wu J. 1992. Ants of the Jianfengling forest region in Hainan Province (Hymenoptera: Formicidae). Scientia Silvae Sinicae 28: 561-564.
 * Wang W. R., S. Q. Zhang, and Z. H. Xu. 2012. A faunistic and taxonomic study of ants (Hymenoptera: Formicidae) in Shenzhen Municipality. Journal of Southwest Forestry University 32(1): 64-73.
 * Wang W., S. Zhang, and Z. Xu. 2012. Distribution Patterns of Ant Species in Shenzhen City. Journal of Southwest Forestry University 32(3): 69-74.
 * Watanasit S., S. Sonthichai, and N. Noon-anant. 2003. Preliminary survey of ants at Tarutao National Park, Southern Thailand. Songklanakarin J. Sci. Technol. 25(1) : 115-122
 * Way M. J., and B. Bolton. 1997. Competition between ants for coconut palm nesting sites. Journal of Natural History 31: 439-455.
 * Wetterer, James K. and Vargo, Donald Vargo L. 2003. Ants (Hymenoptera: Formicidae) of Samoa. Pacific Science. 57(4):409-419.
 * Wheeler W. M. 1909. Ants of Formosa and the Philippines. Bulletin of the American Museum of Natural History 26: 333-345.
 * Wheeler W. M. 1913. Zoological results of the Abor Expedition, 1911-1912, XVII. Hymenoptera, II: Ants (Formicidae). Records of the Indian Museum 8: 233-237.
 * Wheeler W. M. 1919. The ants of Borneo. Bulletin of the Museum of Comparative Zoology 63:43-147.
 * Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bulletin of the American Museum of Natural History 45: 711-1004
 * Wheeler W. M. 1927. Ants collected by Professor F. Silvestri in Indochina. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 20: 83-106.
 * Wheeler W. M. 1937. Additions to the ant-fauna of Krakatau and Verlaten Island. Treubia 16: 21-24.
 * Wheeler W.M. 1934. Formicidae of the Templeton Crocker expedition, 1933. Proceedings of the California Academy of Sciences 21(14): 173-181.
 * Wheeler W.M. 1935. Check list of the ants of Oceania. Occasional Papers of the Bernice Pauahi Bishop Museum 11(11):1-56.
 * Wheeler, William Morton. 1924. Ants of Krakatau and Other Islands in the Sunda Strait. Treubia. 5(1-3):1-20.
 * Wheeler, William Morton. 1934. Formicidae of the Templeton Crocker Expedition, 1933. California Academy of Sciences. 21(14):173-181.
 * Wheeler, William Morton.1935.Checklist of the Ants of Oceania.Occasional Papers 11(11): 3-56
 * Wilson E. O. 1959. Some ecological characteristics of ants in New Guinea rain forests. Ecology 40: 437-447.
 * Wu B., Y. Lu, G. Liang, and L. Zeng. 2010. Influence of the red inported fire ant, Solenopsis invicta Buren (Hymenoptera: Formicidae) on the diversity of ant communities in a newly infested longan orchard and grass areas nearby. Acta Ecologica Sinica 30(8): 2075-2083.
 * Xu Z. 1999. [An analysis on the ant fauna of the tropical rain forest in Xishuangbanna of China.] Zoological Research 20: 379-384.
 * Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
 * Yamane S., T. Itino, and A.R. Nona. 1996. Ground ant fauna in a Bornean dipterocarp forest. Raffles Bulletin of Zoology 44(1): 253-262.
 * Yamane S.; Bui T. V.; Ogata K.; Okido H.; Eguchi K. 2002. Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture Kyushu University 25: 51-62.
 * Yamane S.; Nona, A. R. 1994. Ants from Lambir Hills National Park, Sarawak. Pp. 222-226 in: Inoue, T.; Hamid, A. A. (eds.) 1994. Plant reproductive systems and animal seasonal dynamics. Long-term study of dipterocarp forests in Sarawak. Kyoto: Center for Ecological Research, Kyoto University, vii + 255 pp.
 * Yamane Sk. 2005. Krakatau in 1982, and the commencement of myrmecological research. The nature and Insects (Konchu to shizen) 40: 27-33.
 * Yukawa J., K. Ogata, S. Kamitani, T. Ueno, T. Partomihardjo, S. Kahono, and N. P. Oka. 1999. A preliminary report of the field survey in 1999 on Sulawesi, Indonesia. Bulletin of the Institute of Tropical Agriculture 22: 51-57.
 * Zhang N. N., Y. Q. Chen, Z. X. Lu, W. Zhang, and K. L. Li. 2013. Species diversity, community structure difference and indicator species of leaf-litter ants in rubber plantations and secondary natural forests in Yunnan, southwestern China. Acta Entomologica Sinica 56(11): 1314-1323.
 * Zhang R. J., L. W. Liang, and S. Y. Zhou. 2014. An analysis on the ant fauna of Nonggang Nature Reserve in Guangxi, China. Journal of Guangxi Normal university: Natural Science Edition 32(3): 86-93.
 * Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.
 * Zryanin V. A., and M. V. Mokrousov. 2015. Contribution to the ant fauna of Lombok Island. Proceedings of the 10th ANeT International Conference, 23-26 October 2015, University of Kelaniya, Sri Lanka. 34