Wheeler, George Carlos (1897-1991)
BIOGRAPHICAL NOTE
George Carlos Wheeler (1897-1991) went to Texas Christian University in 1914 with the intention of becoming a high school language teacher. His long-time interest in biology, however, dominated his spare time and upon transferring to the Rice Institute in 1915, Wheeler began to enroll in biology classes. His first teacher and academic advisor was Julian Huxley, who strongly influenced Wheeler to become a professional biologist. When Wheeler completed his undergraduate work at the Rice Institute in 1918, he enlisted in the army and was sent to the Yale Army Laboratory School. After Armistice, he carried out his military service at Base Hospital in Camp Dodge, Iowa. In 1919, he started his graduate work in entomology at Harvard's Bussey Institute. After he received his PhD, Wheeler went on to teach at Syracuse University, the University of North Dakota and the Desert Research Institute (University of Nevada). In 1986, Wheeler authored a brief autobiography of his early career in biology entitled My Association with William Morton Wheeler. In this work, he describes vividly the reasons why he became a biologist, his studies and experiences at Rice Institute and his work at Harvard's Bussey Institute. Also included with George Wheeler's records is the memory book that he maintained during his stay at Rice Institute. In this scrapbook, Wheeler kept photographs of the biology staff, field trips and the campus, as well as newspaper and magazine clippings and memorabilia from social events.
ANT TAXONOMY
GEORGE CARLOS WHEELER, Doctor of Science in Applied Biology, Special Field, Economic Entomology. Thesis: "The Larvae of the Subfamily Dolichoderinae and Formicinae, with a General Consideration of the Developmental Stages of Ants."
PUBLICATIONS
- Deyrup, M.; Johnson, C.; Wheeler, G. C.; Wheeler, J. 1989. A preliminary list of the ants of Florida. Fla. Entomol. 72: 91-101.
- Wheeler, G. C. 1928. The larva of Leptanilla (Hym.: Formicidae). Psyche (Camb.) 35: 85-91.
- Wheeler, G. C. 1935. The larva of Allomerus (Hym.: Formicidae). Psyche (Camb.) 42: 92-98.
- Wheeler, G. C. 1938. Are ant larvae apodous? Psyche (Cambridge) 45:139-145. [1938-12]Stamp date in MCZ library: 1939.03.01.
- Wheeler, G. C. 1943. The larvae of the army ants. Ann. Entomol. Soc. Am. 36: 319-332.
- Wheeler, G. C. 1949 [1948]. The larvae of the fungus-growing ants. Am. Midl. Nat. 40: 664-689.
- Wheeler, G. C. 1950. Ant larvae of the subfamily Cerapachyinae. Psyche (Camb.) 57: 102-113.
- Wheeler, G. C. 1956. Myrmecological orthoepy and onomatology. Grand Forks, North Dakota: published by the author (printed by University of North Dakota Press), 22 pp. [1956]
- Wheeler, G. C.; Wheeler, E. W. 1930. Two new ants from Java. Psyche (Camb.) 37: 193-201.
- Wheeler, G. C.; Wheeler, E. W. 1934. New forms of Aphaenogaster treatae Forel from the southern United States (Hym.: Formicidae). Psyche (Camb.) 41: 6-12.
- Wheeler, G. C.; Wheeler, E. W. 1944. The ants of North Dakota. N. D. Hist. Q. 11: 231-271.
- Wheeler, G. C.; Wheeler, J. 1951. The ant larvae of the subfamily Dolichoderinae. Proc. Entomol. Soc. Wash. 53: 169-210.
- Wheeler, G. C.; Wheeler, J. 1952a. The ant larvae of the subfamily Ponerinae - Part I. Am. Midl. Nat. 48: 111-144.
- Wheeler, G. C.; Wheeler, J. 1952b. The ant larvae of the myrmicine tribe Crematogastrini. J. Wash. Acad. Sci. 42: 248-262.
- Wheeler, G. C.; Wheeler, J. 1952c. The ant larvae of the subfamily Ponerinae - Part II. Am. Midl. Nat. 48: 604-672.
- Wheeler, G. C.; Wheeler, J. 1953a [1952]. The ant larvae of the myrmicine tribe Myrmicini. Psyche (Camb.) 59: 105-125.
- Wheeler, G. C.; Wheeler, J. 1953b. The ant larvae of the myrmicine tribe Pheidolini (Hymenoptera, Formicidae). Proc. Entomol. Soc. Wash. 55: 49-84.
- Wheeler, G. C.; Wheeler, J. 1953c. The ant larvae of the subfamily Formicinae. Ann. Entomol. Soc. Am. 46: 126-171.
- Wheeler, G. C.; Wheeler, J. 1953d. The ant larvae of the myrmicine tribes Melissotarsini, Metaponini, Myrmicariini, and Cardiocondylini. J. Wash. Acad. Sci. 43: 185-189.
- Wheeler, G. C.; Wheeler, J. 1953e. The ant larvae of the subfamily Formicinae. Part II. Ann. Entomol. Soc. Am. 46: 175-217.
- Wheeler, G. C.; Wheeler, J. 1954a [1953]. The ant larvae of the myrmicine tribe Pheidologetini. Psyche (Camb.) 60: 129-147.
- Wheeler, G. C.; Wheeler, J. 1954b. The ant larvae of the myrmicine tribes Cataulacini and Cephalotini. J. Wash. Acad. Sci. 44: 149-157.
- Wheeler, G. C.; Wheeler, J. 1954c. The ant larvae of the myrmicine tribe Myrmecinini (Hymenoptera). Proc. Entomol. Soc. Wash. 56: 126-138.
- Wheeler, G. C.; Wheeler, J. 1954d. The ant larvae of the myrmicine tribes Meranoplini, Ochetomyrmicini and Tetramoriini. Am. Midl. Nat. 52: 443-452.
- Wheeler, G. C.; Wheeler, J. 1955a [1954]. The ant larvae of the myrmicine tribes Basicerotini and Dacetini. Psyche (Camb.) 61: 111-145.
- Wheeler, G. C.; Wheeler, J. 1955b. The ant larvae of the myrmicine tribe Leptothoracini. Ann. Entomol. Soc. Am. 48: 17-29.
- Wheeler, G. C.; Wheeler, J. 1955c. The ant larvae of the myrmicine tribe Solenopsidini. Am. Midl. Nat. 54: 119-141.
- Wheeler, G. C.; Wheeler, J. 1956. The ant larvae of the subfamily Pseudomyrmecinae (Hymenoptera: Formicidae). Ann. Entomol. Soc. Am. 49: 374-398.
- Wheeler, G. C.; Wheeler, J. 1957a. The larva of the ant genus Dacetinops Brown and Wilson. Breviora 78: 1-4.
- Wheeler, G. C.; Wheeler, J. 1957b. The larva of Simopelta (Hymenoptera: Formicidae). Proc. Entomol. Soc. Wash. 59: 191-194.
- Wheeler, G. C.; Wheeler, J. 1957c [1956]. Veromessor lobognathus in North Dakota (Hymenoptera: Formicidae). Psyche (Camb.) 63: 140-145.
- Wheeler, G. C.; Wheeler, J. 1959b. The larva of Paramyrmica (Hymenoptera: Formicidae). J. Tenn. Acad. Sci. 34: 219-220.
- Wheeler, G. C.; Wheeler, J. 1960a. The ant larvae of the subfamily Myrmicinae. Ann. Entomol. Soc. Am. 53: 98-110.
- Wheeler, G. C.; Wheeler, J. 1960b. Supplementary studies on the larvae of the Myrmicinae (Hymenoptera: Formicidae). Proc. Entomol. Soc. Wash. 62: 1-32.
- Wheeler, G.C.& J.Wheeler 1960c. Techniques for the Study of Ant Larvae. Psyche 67 (4):87-94.
- Wheeler, G. C.; Wheeler, J. 1963. The ants of North Dakota. Grand Forks, North Dakota: University of North Dakota Press, viii + 326 pp.
- Wheeler, G. C.; Wheeler, J. 1964a. The ant larvae of the subfamily Cerapachyinae: supplement. Proc. Entomol. Soc. Wash. 66: 65-71.
- Wheeler, G. C.; Wheeler, J. 1964b. The ant larvae of the subfamily Ponerinae: supplement. Ann. Entomol. Soc. Am. 57: 443-462.
- Wheeler, G. C.; Wheeler, J. 1964c. The ant larvae of the subfamily Dorylinae: supplement. Proc. Entomol. Soc. Wash. 66: 129-137.
- Wheeler, G. C.; Wheeler, J. 1965b. The ant larvae of the subfamily Leptanillinae (Hymenoptera, Formicidae). Psyche (Camb.) 72: 24-34.
- Wheeler, G. C.; Wheeler, J. 1966. Ant larva of the subfamily Dolichoderinae: supplement. Ann. Entomol. Soc. Am. 59: 726-732.
- Wheeler, G. C.; Wheeler, J. 1968a. The ant larvae of the subfamily Formicinae (Hymenoptera: Formicidae): supplement. Ann. Entomol. Soc. Am. 61: 205-222.
- Wheeler, G. C.; Wheeler, J. 1968b. The rediscovery of Manica parasitica (Hymenoptera:Formicidae). Pan-Pacific Entomologist 44:71-72.
- Wheeler, G. C.; Wheeler, J. 1969. The larva of Acanthognathus (Hymenoptera: Formicidae). Psyche (Camb.) 76: 110-113.
- Wheeler, G. C.; Wheeler, J. 1970a. The natural history of Manica (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 43:129-162.
- Wheeler, G.C. & J. Wheeler, 1970b. Additions to the natural history of Manica. Journal of the Kansas Entomological Society 43:363.
- Wheeler, G. C.; Wheeler, J. 1970c. Ant larvae of the subfamily Formicinae: second supplement. Ann. Entomol. Soc. Am. 63: 648-656.
- Wheeler, G. C.; Wheeler, J. 1971a [1970]. The larva of Apomyrma (Hymenoptera: Formicidae). Psyche (Camb.) 77: 276-279.
- Wheeler, G. C.; Wheeler, J. 1971b. Ant larvae of the subfamily Ponerinae: second supplement. Ann. Entomol. Soc. Am. 6 64: 1197-1217.
- Wheeler, G. C.; Wheeler, J. 1971c. The larvae of the ant genus Bothroponera (Hymenoptera: Formicidae). Proc. Entomol. Soc. Wash. 73: 386-394.
- Wheeler, G. C.; Wheeler, J. 1971d. Ant larvae of the subfamily Myrmeciinae (Hymenoptera: Formicidae). Pan-Pac. Entomol. 47: 245-256.
- Wheeler, G. C.; Wheeler, J. 1972a. The subfamilies of Formicidae. Proc. Entomol. Soc. Wash. 74: 35-45.
- Wheeler, G. C.; Wheeler, J. 1972b. Ant larvae of the subfamily Myrmicinae: second supplement on the tribes Myrmicini and Pheidolini. J. Ga. Entomol. Soc. 7: 233-246.
- Wheeler, G. C.; Wheeler, J. 1973a. The ant larvae of six tribes: second supplement (Hymenoptera: Formicidae: Myrmicinae). J. Ga. Entomol. Soc. 8: 27-39.
- Wheeler, G. C.; Wheeler, J. 1973b. Ant larvae of four tribes: second supplement (Hymenoptera: Formicidae: Myrmicinae). Psyche (Camb.) 80: 70-82.
- Wheeler, G. C.; Wheeler, J. 1973c. The ant larvae of the tribes Basicerotini and Dacetini: second supplement (Hymenoptera: Formicidae: Myrmicinae). Pan-Pac. Entomol. 49: 207-214.
- Wheeler, G. C.; Wheeler, J. 1973d. Supplementary studies on ant larvae: Cerapachyinae, Pseudomyrmecinae and Myrmicinae. Psyche (Camb.) 80: 204-211.
- Wheeler, G. C.; Wheeler, J. 1973e. Ants of Deep Canyon. Riverside, Calif.: University of California, xiii + 162 pp.
- Wheeler, G. C.; Wheeler, J. 1974a. Ant larvae of the subfamily Formicinae: third supplement. J. Ga. Entomol. Soc. 9: 59-64.
- Wheeler, G. C.; Wheeler, J. 1974b [1973]. Ant larvae of the subfamily Dolichoderinae: second supplement (Hymenoptera: Formicidae). Pan-Pac. Entomol. 49: 396-401.
- Wheeler, G. C.; Wheeler, J. 1974c. Ant larvae of the subfamily Dorylinae: second supplement (Hymenoptera: Formicidae). J. Kans. Entomol. Soc. 47: 166-172.
- Wheeler, G. C.; Wheeler, J. 1974d. Ant larvae of the myrmicine tribe Attini: second supplement (Hymenoptera: Formicidae). Proc. Entomol. Soc. Wash. 76: 76-81.
- Wheeler, G. C.; Wheeler, J. 1974e. Supplementary studies on ant larvae: Teratomyrmex. Psyche (Camb.) 81: 38-41.
- Wheeler, G. C.; Wheeler, J. 1974f. Supplementary studies on ant larvae: Simopone and Turneria. J. N. Y. Entomol. Soc. 82: 103-105.
- Wheeler, G. C.; Wheeler, J. 1974g. Ant larvae of the subfamily Ponerinae: third supplement (Hymenoptera: Formicidae). Proc. Entomol. Soc. Wash. 76: 278-281.
- Wheeler, G. C.; Wheeler, J. 1976a. Supplementary studies on ant larvae: Ponerinae. Trans. Am. Entomol. Soc. 102: 41-64.
- Wheeler, G. C.; Wheeler, J. 1976b. Ant larvae: review and synthesis. Mem. Entomol. Soc. Wash. 7: 1-108.
- Wheeler, G. C.; Wheeler, J. 1977a. Supplementary studies on ant larvae: Myrmicinae. Trans. Am. Entomol. Soc. 103: 581-602.
- Wheeler, G. C.; Wheeler, J. 1977b. North Dakota ants updated. Reno, Nevada: published by the authors, 27 pp.
- Wheeler, G.C.; Wheeler, J. 1978. Mountain ants of Nevada. Great Basin Naturalist 38: 379-396.
- Wheeler, G. C.; Wheeler, J. 1980. Supplementary studies on ant larvae: Ponerinae, Myrmicinae and Formicinae. Trans. Am. Entomol. Soc. 106: 527-545.
- Wheeler, G. C.; Wheeler, J. 1982b. Supplementary studies on ant larvae: Formicinae (Hymenoptera: Formicidae). Psyche (Camb.) 89: 175-181.
- Wheeler, G. C.; Wheeler, J. 1983a. Supplementary studies on ant larvae: Myrmicinae. Trans. Am. Entomol. Soc. 108: 601-610.
- Wheeler, G. C.; Wheeler, J. 1984a. The larvae of the army ants (Hymenoptera: Formicidae): a revision. J. Kans. Entomol. Soc. 57: 263-275.
- Wheeler, G. C.; Wheeler, J. 1984b. Myrmecological orthoepy and onomatology. (2nd edition.). Grand Forks, North Dakota: published by the authors, 20 pp PDF
- Wheeler, G. C. and J. Wheeler. 1985a. A checklist of Texas ants. The Prairie Naturalist 17(2): 49-64.
- Wheeler, G. C.; Wheeler, J. 1985b. A simplified conspectus of the Formicidae. Trans. Am. Entomol. Soc. 111: 255-264.
- Wheeler, G. C.; Wheeler, J. 1986a [1985]. The larva of Dinoponera (Hymenoptera: Formicidae: Ponerinae). Psyche (Camb.) 92: 387-39.
- Wheeler, G. C.; Wheeler, J. 1986b [1985]. The larva of Proatta (Hymenoptera: Formicidae). Psyche (Camb.) 92: 447-450.
- Wheeler, G. C.; Wheeler, J. 1986c. Supplementary studies of ant larvae: Ponerinae. Trans. Am. Entomol. Soc. 112: 85-94.
- Wheeler, G. C.; Wheeler, J. 1986d. Supplementary studies on ant larvae: Formicinae (Hymenoptera: Formicidae). J. N. Y. Entomol. Soc. 94: 331-341.
- Wheeler, G. C.; Wheeler, J. 1986f. Supplementary studies on ant larvae: Myrmicinae (Hymenoptera: Formicidae). J. N. Y. Entomol. Soc. 94: 489-499.
- Wheeler, G. C.; Wheeler, J. 1986g. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp.
- Wheeler, G. C.; Wheeler, J. 1987a [1986]. Young larvae of Eciton (Hymenoptera: Formicidae: Dorylinae). Psyche (Camb.) 93: 341-349.
- Wheeler, G. C.; Wheeler, J. 1988b [1987]. Young larvae of Veromessor pergandei (Hymenoptera: Formicidae: Myrmicinae). Psyche (Camb.) 94: 303-307.
- Wheeler, G. C.; Wheeler, J. 1988c. The larva of Notostigma (Hymenoptera: Formicidae: Formicinae). J. N. Y. Entomol. Soc. 96: 355-358.
- Wheeler, G. C.; Wheeler, J. 1988e. A checklist of the ants of Wyoming (Hymenoptera: Formicidae). Insecta Mundi 2:231-239.
- Wheeler, G. C.; Wheeler, J. 1989a [1988]. Notes on ant larvae: Myrmicinae. Trans. Am. Entomol. Soc. 114: 319-327.
- Wheeler, G. C.; Wheeler, J. 1989b. Notes on ant larvae: Ponerinae. J. N. Y. Entomol. Soc. 97: 50-55.
- Wheeler, G. C.; Wheeler, J. 1989c [1988]. The larva of Leptanilla japonica, with notes on the genus (Hymenoptera: Formicidae: Leptanillinae). Psyche (Camb.) 95: 185-189.
- Wheeler, G. C.; Wheeler, J. 1988d. A checklist of the ants of Montana. Psyche (Cambridge) 95:101-114. [1988-12-20]
- Wheeler, G. C.; Wheeler, J. 1990a [1989]. Notes on ant larvae. Trans. Am. Entomol. Soc. 115: 457-473.
- Wheeler, G. C.; Wheeler, J. 1990b. Larvae of the formicine ant genus Polyrhachis. Trans. Am. Entomol. Soc. 116: 753-767.
- Wheeler, G. C.; Wheeler, J. 1991a. The larva of Blepharidatta (Hymenoptera: Formicidae). J. N. Y. Entomol. Soc. 99: 132-137.
- Wheeler, G. C.; Wheeler, J. 1991b. Instars of three ant species. Psyche (Camb.) 98: 89-99.
- Wheeler, G. C.; Wheeler, J.; Kannowski, P. B. 1994. Checkist of ants of Michigan (Hymenoptera: Formicidae). Great Lakes Entomol. 26: 297-310.
- Wheeler, G. C.; Wheeler, J.; Taylor, R. W. 1980. The larval and egg stages of the primitive ant Nothomyrmecia macrops Clark (Hymenoptera: Formicidae). J. Aust. Entomol. Soc. 19: 131-137
- Wilson, E. O.; Eisner, T.; Wheeler, G. C.; Wheeler, J. 1956. Aneuretus simoni Emery, a major link in ant evolution. Bulletin of the Museum of Comparative Zoology 115: 81-99.